387 lines
53 KiB
XML
387 lines
53 KiB
XML
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<mods:title id="EA76F1CB45029477ADA9DDD7E2353E33">A taxonomic revision of Proctoporus bolivianus Werner (Squamata: Gymnophthalmidae) with the description of three new species and resurrection of Proctoporus lacertus Stejneger</mods:title>
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<mods:namePart id="4DFCE986F509CA02865C63B467C5F836">Goicoechea, NoemÍ</mods:namePart>
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<mods:namePart id="2DC84BA4A1C0E41D41089562C0E7E789">Padial, José M.</mods:namePart>
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<mods:namePart id="B8A79112160A7D933E674BF257CB8C91">Chaparro, Juan Carlos</mods:namePart>
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<mods:namePart id="A14F24EFCBADF55AD4ABF9D69B67B5FA">Castroviejo-Fisher, Santiago</mods:namePart>
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<mods:namePart id="9FA5ED10D6A77FE35B6EF9860670152A">Riva, Ignacio De La</mods:namePart>
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<mods:title id="5F0D3B7C8589496D7499D5222B52F3AD">American Museum Novitates</mods:title>
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<mods:date id="C71656B6DEDDD440BEC5DF779097F2F0">2013</mods:date>
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<emphasis id="4AEC6F5215375311FE0C0DC79DDCFE34" bold="true" box="[466,724,404,431]" italics="true" pageId="20" pageNumber="21">Proctoporus kiziriani</emphasis>
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,
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,
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<emphasis id="4AEC6F5215375311FFA60DB29E61FE63" box="[120,361,480,504]" italics="true" pageId="20" pageNumber="21">Proctoporus bolivianus</emphasis>
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[Ca5 MNCN20610]:
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<bibRefCitation id="1C09CEB115375311FD8C0DB39C88FE63" author="Goicoechea, N. & J. M. Padial & J. C. Chaparro & S. Castroviejo-Fisher & I. De la Riva" box="[594,896,480,505]" pageId="20" pageNumber="21" pagination="953 - 964" refId="ref17160" refString="Goicoechea, N., J. M. Padial, J. C. Chaparro, S. Castroviejo-Fisher, and I. De la Riva. 2012. Molecular phylogenetics, species delimitation and biogeography of the Andean lizards of the genus Proctoporus (Squamata: Gymnophthalmidae). Molecular Phylogenetics and Evolution 65: 953 - 964." type="journal article" year="2012">Goicoechea et al., 2012: 953</bibRefCitation>
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.
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:
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(field number
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) (fig. 7),
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, from
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, (
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/
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, collected by
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,
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,
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<collectorName id="D56DD69615375311FB4D0E099E9CFD02" pageId="20" pageNumber="21">Santiago Castroviejo-Fisher</collectorName>
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, and
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<collectorName id="D56DD69615375311FE0A0E2C9DDEFD02" box="[468,726,639,665]" pageId="20" pageNumber="21">Juan Carlos Chaparro</collectorName>
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on
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.
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:
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(field numbers
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,
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<specimenCode id="283E1B3B15375311FCAD0EF79CA4FD24" box="[883,940,676,703]" collectionCode="MNCN" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="20" pageNumber="21" type="Museum">4605</specimenCode>
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,
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<specimenCode id="283E1B3B15375311FC650EF79CFCFD24" box="[955,1012,676,703]" collectionCode="MNCN" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="20" pageNumber="21" type="Museum">4606</specimenCode>
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)
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, and
|
||
<specimenCode id="283E1B3B15375311FFA60E999E11FD78" box="[120,281,713,740]" collectionCode="MNCN" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="20" pageNumber="21" type="Museum">MNCN 43673</specimenCode>
|
||
(field number
|
||
<specimenCode id="283E1B3B15375311FE600E999D5BFD7F" box="[446,595,713,740]" collectionCode="MNCN" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="20" pageNumber="21" type="Museum">MNCN 4608</specimenCode>
|
||
)
|
||
<specimenCount id="6E9E78C915375311FDBE0E9A9DE4FD78" box="[608,748,713,739]" pageId="20" pageNumber="21" type="female">adult female</specimenCount>
|
||
, same data as holotype;
|
||
</materialsCitation>
|
||
<materialsCitation id="C8F0B91D15375311FC2A0E999B7DFC5F" ID-GBIF-Occurrence="3446987321" collectingDate="2007-01-31" collectionCode="MNCN" collectorName="Ignacio De la Riva & Jose Manuel Padial & Santiago Castroviejo-Fisher & Juan Carlos Chaparro" country="Peru" county="Department" elevation="3368" latitude="-13.583445" location="Huancarayo" longLatPrecision="1" longitude="-71.03475" municipality="Province Quispicanchis" pageId="20" pageNumber="21" specimenCode="MHNC 5680, MNCN 4750, MNCN 44216, MNCN 4744, MNCN 44218-19, MNCN 4746, 4747, MHNC 5682-83, MNCN 4751, 4752, MHNC 5685, MNCN 4754, MNCN 44217, MNCN 4745, MNCN 44220-21, MNCN 4748, 4749" specimenCount="2" stateProvince="Cusco" typeStatus="paratype">
|
||
<specimenCode id="283E1B3B15375311FC2A0E999B8FFD7F" box="[1012,1159,713,740]" collectionCode="MHNC" country="0" httpUri="http://grbio.org/cool/25c9-tsqt" name="Museo de Historia Natural de Concepcion (Chile)" pageId="20" pageNumber="21">MHNC 5680</specimenCode>
|
||
(field number
|
||
<specimenCode id="283E1B3B15375311FF030EBC9E70FC91" box="[221,376,751,778]" collectionCode="MNCN" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="20" pageNumber="21" type="Museum">MNCN 4750</specimenCode>
|
||
),
|
||
<specimenCode id="283E1B3B15375311FE440EBC9D49FC92" box="[410,577,751,777]" collectionCode="MNCN" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="20" pageNumber="21" type="Museum">MNCN 44216</specimenCode>
|
||
(field number
|
||
<specimenCode id="283E1B3B15375311FD2B0EBC9C98FC92" box="[757,912,751,777]" collectionCode="MNCN" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="20" pageNumber="21" type="Museum">MNCN 4744</specimenCode>
|
||
),
|
||
<specimenCode id="283E1B3B15375311FC6C0EBC9B8BFC92" box="[946,1155,751,777]" collectionCode="MNCN" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="20" pageNumber="21" type="Museum">MNCN 44218–19</specimenCode>
|
||
(field numbers
|
||
<specimenCode id="283E1B3B15375311FF3F0F479E7DFCB5" box="[225,373,788,814]" collectionCode="MNCN" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="20" pageNumber="21" type="Museum">MNCN 4746</specimenCode>
|
||
,
|
||
<specimenCode id="283E1B3B15375311FEA00F479EBEFCB5" box="[382,438,788,814]" collectionCode="MNCN" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="20" pageNumber="21" type="Museum">4747</specimenCode>
|
||
)
|
||
<specimenCount id="6E9E78C915375311FE1D0F479D4BFCB5" box="[451,579,788,814]" pageId="20" pageNumber="21" type="adult">adult males</specimenCount>
|
||
, and
|
||
<specimenCode id="283E1B3B15375311FDA20F479C31FCB4" box="[636,825,788,815]" collectionCode="MHNC" country="0" httpUri="http://grbio.org/cool/25c9-tsqt" name="Museo de Historia Natural de Concepcion (Chile)" pageId="20" pageNumber="21">MHNC 5682–83</specimenCode>
|
||
(field numbers
|
||
<specimenCode id="283E1B3B15375311FC380F479B72FCB4" box="[998,1146,788,815]" collectionCode="MNCN" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="20" pageNumber="21" type="Museum">MNCN 4751</specimenCode>
|
||
,
|
||
<specimenCode id="283E1B3B15375311FB5A0F479BB4FCB4" box="[1156,1212,788,815]" collectionCode="MNCN" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="20" pageNumber="21" type="Museum">4752</specimenCode>
|
||
),
|
||
<specimenCode id="283E1B3B15375311FFA60F6A9E18FCCF" box="[120,272,825,852]" collectionCode="MHNC" country="0" httpUri="http://grbio.org/cool/25c9-tsqt" name="Museo de Historia Natural de Concepcion (Chile)" pageId="20" pageNumber="21">MHNC 5685</specimenCode>
|
||
(field number
|
||
<specimenCode id="283E1B3B15375311FE1E0F6A9D50FCCF" box="[448,600,825,852]" collectionCode="MNCN" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="20" pageNumber="21" type="Museum">MNCN 4754</specimenCode>
|
||
),
|
||
<specimenCode id="283E1B3B15375311FDB00F6A9C1AFCC8" box="[622,786,825,851]" collectionCode="MNCN" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="20" pageNumber="21" type="Museum">MNCN 44217</specimenCode>
|
||
(field number
|
||
<specimenCode id="283E1B3B15375311FC1C0F6A9B52FCCF" box="[962,1114,825,852]" collectionCode="MNCN" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="20" pageNumber="21" type="Museum">MNCN 4745</specimenCode>
|
||
),
|
||
<specimenCode id="283E1B3B15375311FBAF0F6A9FEEFCE2" collectionCode="MNCN" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="20" pageNumber="21" type="Museum">MNCN 44220–21</specimenCode>
|
||
(field numbers
|
||
<specimenCode id="283E1B3B15375311FE450F0C9D3AFCE2" box="[411,562,863,889]" collectionCode="MNCN" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="20" pageNumber="21" type="Museum">MNCN 4748</specimenCode>
|
||
,
|
||
<specimenCode id="283E1B3B15375311FDE30F0C9D7DFCE2" box="[573,629,863,889]" collectionCode="MNCN" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="20" pageNumber="21" type="Museum">4749</specimenCode>
|
||
),
|
||
<specimenCount id="6E9E78C915375311FD570F0C9C29FCE2" box="[649,801,863,889]" pageId="20" pageNumber="21" type="adult">adult females</specimenCount>
|
||
, from
|
||
<location id="7D47E59B15375311FCB20F0C9CFEFCE2" LSID="urn:lsid:plazi:treatment:F031025615375312FE0C0DC79DC1FB96:7D47E59B15375311FCB20F0C9CFEFCE2" box="[876,1014,863,889]" country="Peru" county="Department" latitude="-13.583445" longLatPrecision="1" longitude="-71.03475" municipality="Province Quispicanchis" name="Huancarayo" pageId="20" pageNumber="21" stateProvince="Cusco">Huancarayo</location>
|
||
,
|
||
<location id="7D47E59B15375311FBDF0F0C9BCDFCE2" LSID="urn:lsid:plazi:treatment:F031025615375312FE0C0DC79DC1FB96:7D47E59B15375311FBDF0F0C9BCDFCE2" box="[1025,1221,863,889]" country="Peru" county="Department" latitude="-13.583445" longLatPrecision="1" longitude="-71.03475" municipality="Province Quispicanchis" name="Marcapata Valley" pageId="20" pageNumber="21" stateProvince="Cusco">Marcapata Valley</location>
|
||
,
|
||
<collectingMunicipality id="9843293A15375311FFA60FD79E8BFC05" box="[120,387,900,926]" pageId="20" pageNumber="21">Province Quispicanchis</collectingMunicipality>
|
||
,
|
||
<collectingCounty id="9146CBCC15375311FE500FD69D10FC04" box="[398,536,901,927]" pageId="20" pageNumber="21">Department</collectingCounty>
|
||
of
|
||
<collectingRegion id="BA5C7DA215375311FDE20FD79D8CFC05" box="[572,644,900,926]" country="Peru" name="Cusco" pageId="20" pageNumber="21">Cusco</collectingRegion>
|
||
,
|
||
<collectingCountry id="008FF3D015375311FD510FD69DCEFC04" box="[655,710,901,927]" name="Peru" pageId="20" pageNumber="21">Peru</collectingCountry>
|
||
,
|
||
<quantity id="BF601EA515375311FD0E0FD79C2EFC04" box="[720,806,900,927]" metricMagnitude="3" metricUnit="m" metricValue="3.368" pageId="20" pageNumber="21" unit="m" value="3368.0">
|
||
<elevation id="F3B5547315375311FD0E0FD79C2EFC04" box="[720,806,900,927]" metricMagnitude="3" metricUnit="m" metricValue="3.368" pageId="20" pageNumber="21" unit="m" value="3368.0">3368 m</elevation>
|
||
</quantity>
|
||
, collected by
|
||
<collectorName id="D56DD69615375311FC600FD79B98FC05" box="[958,1168,900,927]" hasSafeAffix="true" pageId="20" pageNumber="21">Ignacio De la Riva</collectorName>
|
||
,
|
||
<collectorName id="D56DD69615375311FB440FD79E15FC58" pageId="20" pageNumber="21">José Manuel Padial</collectorName>
|
||
,
|
||
<collectorName id="D56DD69615375311FEF90FFA9D56FC58" box="[295,606,937,963]" pageId="20" pageNumber="21">Santiago Castroviejo-Fisher</collectorName>
|
||
, and
|
||
<collectorName id="D56DD69615375311FD440FF99C99FC58" box="[666,913,937,964]" pageId="20" pageNumber="21">Juan Carlos Chaparro</collectorName>
|
||
on
|
||
<date id="0C26958015375311FC620FF99B7DFC5F" box="[956,1141,938,964]" pageId="20" pageNumber="21" value="2007-01-31">
|
||
<collectingDate id="1C626C6815375311FC620FF99B7DFC5F" box="[956,1141,938,964]" pageId="20" pageNumber="21" value="2007-01-31">31 January 2007</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="3082E0CB15375310FF760F9C9D4EFAC0" lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21" type="diagnosis">
|
||
<paragraph id="7827B34015375311FF760F9C9B4FFA32" blockId="20.[120,1228,526,1785]" pageId="20" pageNumber="21">
|
||
DIAGNOSIS: (1) Frontonasal length usually equal to frontal length; (2) nasoloreal suture present; (3) three supraoculars; (4) 3–4 superciliaries, first not expanded onto dorsal surface of head; (5) postoculars two; (6) palpebral disc made up of a single, undivided scale; (7) supralabials anterior to the posteroventral angle of the subocular four; (8) two pairs of genials in contact (9) dorsal body scales quadrangular, slightly keeled; (10) transverse rows of dorsals
|
||
<quantity id="BF601EA515375311FBC608379B76FBE5" box="[1048,1150,1124,1151]" metricMagnitude="-1" metricUnit="m" metricValue="9.398" metricValueMax="9.906" metricValueMin="8.89" pageId="20" pageNumber="21" unit="in" value="37.0" valueMax="39.0" valueMin="35.0">35–39 in</quantity>
|
||
males and
|
||
<quantity id="BF601EA515375311FF7508D99E19FB38" box="[171,273,1161,1188]" metricMagnitude="0" metricUnit="m" metricValue="1.0032999999999999" metricValueMax="1.0413999999999999" metricValueMin="0.9652" pageId="20" pageNumber="21" unit="in" value="39.5" valueMax="41.0" valueMin="38.0">38–41 in</quantity>
|
||
females; (11) transverse ventral rows
|
||
<quantity id="BF601EA515375311FD1908D99C25FB38" box="[711,813,1161,1188]" metricMagnitude="-1" metricUnit="m" metricValue="5.715" metricValueMax="5.842" metricValueMin="5.588" pageId="20" pageNumber="21" unit="in" value="22.5" valueMax="23.0" valueMin="22.0">22–23 in</quantity>
|
||
males and
|
||
<quantity id="BF601EA515375311FC6C08D99B10FB38" box="[946,1048,1161,1188]" metricMagnitude="-1" metricUnit="m" metricValue="5.842" metricValueMax="6.096" metricValueMin="5.588" pageId="20" pageNumber="21" unit="in" value="23.0" valueMax="24.0" valueMin="22.0">22–24 in</quantity>
|
||
females; (12) a continuous series of small lateral scales separating dorsals from ventrals; (13) posterior cloacal plate made up of six scales in both sexes; (14) anterior preanal plate scales paired; (15) femoral pores per hind limb in males 5–7, absent in females; (16) preanal pores absent; (17) subdigital lamellae on toe IV 16–20; subdigital lamellae on toe V 7–13; (18) limbs not overlapping when adpressed against body in adults; (19) pentadactyl; digits clawed; (20) dorsum brown or pale brown with dorsolateral pale stripes bordered by a discontinuous dark line on neck and body; lateral ocelli usually present in both sexes; ventral surfaces black, with pale stippling.
|
||
</paragraph>
|
||
<paragraph id="7827B34015375310FF7609E79D4EFAC0" blockId="20.[120,1228,526,1785]" lastBlockId="21.[120,1228,1084,1782]" lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21">
|
||
Specimens of
|
||
<taxonomicName id="BF98C8C315375311FE8B09E69D47FA55" authorityName="Goicoechea & Padial & Chaparro & Castroviejo-Fisher & Riva" authorityYear="2013" box="[341,591,1460,1486]" class="Reptilia" family="Gymnophthalmidae" genus="Proctoporus" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="kiziriani">
|
||
<emphasis id="4AEC6F5215375311FE8B09E69D47FA55" box="[341,591,1460,1486]" italics="true" pageId="20" pageNumber="21">Proctoporus kiziriani</emphasis>
|
||
</taxonomicName>
|
||
show the presence of an undivided palpebral disc, which identifies them as members of the genus
|
||
<taxonomicName id="BF98C8C315375311FD4009899C2FFA68" authorityName="Tschudi" authorityYear="1845" box="[670,807,1498,1523]" class="Reptilia" family="Gymnophthalmidae" genus="Proctoporus" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="21" phylum="Chordata" rank="genus">
|
||
<emphasis id="4AEC6F5215375311FD4009899C2FFA68" box="[670,807,1498,1523]" italics="true" pageId="20" pageNumber="21">Proctoporus</emphasis>
|
||
</taxonomicName>
|
||
, as opposed to
|
||
<taxonomicName id="BF98C8C315375311FC0409899B2EFA68" box="[986,1062,1498,1523]" class="Reptilia" family="Gymnophthalmidae" genus="Riama" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="21" phylum="Chordata" rank="genus">
|
||
<emphasis id="4AEC6F5215375311FC0409899B2EFA68" box="[986,1062,1498,1523]" italics="true" pageId="20" pageNumber="21">Riama</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="BF98C8C315375311FB80098A9BC0FA68" authorityName="Doan & Castoe" authorityYear="2005" box="[1118,1224,1497,1523]" class="Reptilia" family="Gymnophthalmidae" genus="Petracola" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="21" phylum="Chordata" rank="genus">
|
||
<emphasis id="4AEC6F5215375311FB80098A9BC0FA68" box="[1118,1224,1497,1523]" italics="true" pageId="20" pageNumber="21">Petracola</emphasis>
|
||
</taxonomicName>
|
||
(Doan and Castoe, 2005;
|
||
<bibRefCitation id="1C09CEB115375311FE6A09AC9D45F982" author="Uzzell, T. M." box="[436,589,1535,1561]" pageId="20" pageNumber="21" pagination="1 - 39" refId="ref17410" refString="Uzzell, T. M. 1970. Teiid lizards of the genus Proctoporus from Bolivia and Peru. Postilla 142: 1 - 39." type="journal article" year="1970">Uzzell, 1970</bibRefCitation>
|
||
).
|
||
<taxonomicName id="BF98C8C315375311FDBA0A539C56F982" authorityName="Goicoechea & Padial & Chaparro & Castroviejo-Fisher & Riva" authorityYear="2013" box="[612,862,1535,1561]" class="Reptilia" family="Gymnophthalmidae" genus="Proctoporus" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="kiziriani">
|
||
<emphasis id="4AEC6F5215375311FDBA0A539C56F982" box="[612,862,1535,1561]" italics="true" pageId="20" pageNumber="21">Proctoporus kiziriani</emphasis>
|
||
</taxonomicName>
|
||
can be distinguished from
|
||
<taxonomicName id="BF98C8C315375311FB6B0A539FE7F9A5" class="Reptilia" family="Gymnophthalmidae" genus="Proctoporus" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="pachyurus">
|
||
<emphasis id="4AEC6F5215375311FB6B0A539FE7F9A5" italics="true" pageId="20" pageNumber="21">P. pachyurus</emphasis>
|
||
</taxonomicName>
|
||
by having three supraoculars not fused with superciliaries (four supraoculars in
|
||
<taxonomicName id="BF98C8C315375311FB6B0A769FE7F9F8" class="Reptilia" family="Gymnophthalmidae" genus="Proctoporus" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="pachyurus">
|
||
<emphasis id="4AEC6F5215375311FB6B0A769FE7F9F8" italics="true" pageId="20" pageNumber="21">P. pachyurus</emphasis>
|
||
</taxonomicName>
|
||
, first fused with first superciliary) and by having 35–41 transverse dorsal scale rows (
|
||
<quantity id="BF601EA515375311FF5E0A3C9FE3F912" box="[128,235,1647,1673]" metricMagnitude="0" metricUnit="m" metricValue="1.3589" metricValueMax="1.524" metricValueMin="1.1938" pageId="20" pageNumber="21" unit="in" value="53.5" valueMax="60.0" valueMin="47.0">47–60 in</quantity>
|
||
<taxonomicName id="BF98C8C315375311FF2A0A239E80F912" box="[244,392,1647,1673]" class="Reptilia" family="Gymnophthalmidae" genus="Proctoporus" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="pachyurus">
|
||
<emphasis id="4AEC6F5215375311FF2A0A239E80F912" box="[244,392,1647,1673]" italics="true" pageId="20" pageNumber="21">P. pachyurus</emphasis>
|
||
</taxonomicName>
|
||
); from
|
||
<taxonomicName id="BF98C8C315375311FE3B0A239D66F912" authorityName="Doan & Castoe" authorityYear="2003" box="[485,622,1647,1673]" class="Reptilia" family="Gymnophthalmidae" genus="Proctoporus" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="sucullucu">
|
||
<emphasis id="4AEC6F5215375311FE3B0A239D66F912" box="[485,622,1647,1673]" italics="true" pageId="20" pageNumber="21">P. sucullucu</emphasis>
|
||
</taxonomicName>
|
||
by having limbs not overlapping when adpressed, first supraocular not fused with first superciliary (fused in
|
||
<taxonomicName id="BF98C8C315375311FCF90AC69CA6F935" authorityName="Doan & Castoe" authorityYear="2003" box="[807,942,1684,1710]" class="Reptilia" family="Gymnophthalmidae" genus="Proctoporus" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="sucullucu">
|
||
<emphasis id="4AEC6F5215375311FCF90AC69CA6F935" box="[807,942,1684,1710]" italics="true" pageId="20" pageNumber="21">P. sucullucu</emphasis>
|
||
</taxonomicName>
|
||
), and loreal scale not in contact with supralabials (loreal in contact with second and third supralabials in
|
||
<taxonomicName id="BF98C8C315375311FBEE0AE99BBFF948" authorityName="Doan & Castoe" authorityYear="2003" box="[1072,1207,1721,1747]" class="Reptilia" family="Gymnophthalmidae" genus="Proctoporus" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="sucullucu">
|
||
<emphasis id="4AEC6F5215375311FBEE0AE99BBFF948" box="[1072,1207,1721,1747]" italics="true" pageId="20" pageNumber="21">P. sucullucu</emphasis>
|
||
</taxonomicName>
|
||
); from
|
||
<taxonomicName id="BF98C8C315375311FF670AB39E41F962" box="[185,329,1759,1785]" class="Reptilia" family="Gymnophthalmidae" genus="Proctoporus" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="bolivianus">
|
||
<emphasis id="4AEC6F5215375311FF670AB39E41F962" box="[185,329,1759,1785]" italics="true" pageId="20" pageNumber="21">P. bolivianus</emphasis>
|
||
</taxonomicName>
|
||
by having frontonasal length equal to frontal and first supraocular not fused with first superciliary; from
|
||
<taxonomicName id="BF98C8C315365310FE13086E9D5CFBCD" authorityName="Doan & Castoe" authorityYear="2003" box="[461,596,1085,1110]" class="Reptilia" family="Gymnophthalmidae" genus="Proctoporus" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="unsaacae">
|
||
<emphasis id="4AEC6F5215365310FE13086E9D5CFBCD" box="[461,596,1085,1110]" italics="true" pageId="21" pageNumber="22">P. unsaacae</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="BF98C8C315365310FD4A086E9C16FBCD" baseAuthorityName="Boettger" baseAuthorityYear="1891" box="[660,798,1084,1110]" class="Reptilia" family="Gymnophthalmidae" genus="Proctoporus" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="guentheri">
|
||
<emphasis id="4AEC6F5215365310FD4A086E9C16FBCD" box="[660,798,1084,1110]" italics="true" pageId="21" pageNumber="22">P. guentheri</emphasis>
|
||
</taxonomicName>
|
||
by not having the first superciliary expanded onto dorsal surface of the head; from
|
||
<taxonomicName id="BF98C8C315365310FD7908319C97FBE0" authorityName="Goicoechea & Padial & Chaparro & Castroviejo-Fisher & Riva" authorityYear="2013" box="[679,927,1121,1147]" class="Reptilia" family="Gymnophthalmidae" genus="Proctoporus" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="carabaya">
|
||
<emphasis id="4AEC6F5215365310FD7908319C97FBE0" box="[679,927,1121,1147]" italics="true" pageId="21" pageNumber="22">Proctoporus carabaya</emphasis>
|
||
</taxonomicName>
|
||
by having first supraocular not fused with first superciliary and by the presence of dorsolateral stripes; from
|
||
<taxonomicName id="BF98C8C315365310FB9F08DB9BC0FB3A" authorityName="Goicoechea & Padial & Chaparro & Castroviejo-Fisher & Riva" authorityYear="2013" box="[1089,1224,1159,1185]" class="Reptilia" family="Gymnophthalmidae" genus="Proctoporus" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="iridescens">
|
||
<emphasis id="4AEC6F5215365310FB9F08DB9BC0FB3A" box="[1089,1224,1159,1185]" italics="true" pageId="21" pageNumber="22">P. iridescens</emphasis>
|
||
</taxonomicName>
|
||
by having four supralabials anterior to the posteroventral angle of the subocular (three in
|
||
<taxonomicName id="BF98C8C315365310FB6B08FE9FEFFB70" authorityName="Goicoechea & Padial & Chaparro & Castroviejo-Fisher & Riva" authorityYear="2013" class="Reptilia" family="Gymnophthalmidae" genus="Proctoporus" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="iridescens">
|
||
<emphasis id="4AEC6F5215365310FB6B08FE9FEFFB70" italics="true" pageId="21" pageNumber="22">P. iridescens</emphasis>
|
||
</taxonomicName>
|
||
) and two pairs of genials in contact (three pairs in
|
||
<taxonomicName id="BF98C8C315365310FC9708819CDCFB70" authorityName="Goicoechea & Padial & Chaparro & Castroviejo-Fisher & Riva" authorityYear="2013" box="[841,980,1233,1259]" class="Reptilia" family="Gymnophthalmidae" genus="Proctoporus" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="iridescens">
|
||
<emphasis id="4AEC6F5215365310FC9708819CDCFB70" box="[841,980,1233,1259]" italics="true" pageId="21" pageNumber="22">P. iridescens</emphasis>
|
||
</taxonomicName>
|
||
); from
|
||
<taxonomicName id="BF98C8C315365310FBF108819BACFB70" authorityName=", Stejneger" authorityYear="1913" box="[1071,1188,1233,1259]" class="Reptilia" family="Gymnophthalmidae" genus="Proctoporus" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="lacertus">
|
||
<emphasis id="4AEC6F5215365310FBF108819BACFB70" box="[1071,1188,1233,1259]" italics="true" pageId="21" pageNumber="22">P. lacertus</emphasis>
|
||
</taxonomicName>
|
||
by the presence of loreal scales (absent in
|
||
<taxonomicName id="BF98C8C315365310FD9308AB9DCCFA8A" authorityName=", Stejneger" authorityYear="1913" box="[589,708,1271,1297]" class="Reptilia" family="Gymnophthalmidae" genus="Proctoporus" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="lacertus">
|
||
<emphasis id="4AEC6F5215365310FD9308AB9DCCFA8A" box="[589,708,1271,1297]" italics="true" pageId="21" pageNumber="22">P. lacertus</emphasis>
|
||
</taxonomicName>
|
||
) and dorsolateral stripes; from
|
||
<taxonomicName id="BF98C8C315365310FB9F08AB9BAAFA8A" baseAuthorityName="Uzzell" baseAuthorityYear="1969" box="[1089,1186,1272,1297]" class="Reptilia" family="Gymnophthalmidae" genus="Proctoporus" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="xestus">
|
||
<emphasis id="4AEC6F5215365310FB9F08AB9BAAFA8A" box="[1089,1186,1272,1297]" italics="true" pageId="21" pageNumber="22">P. xestus</emphasis>
|
||
</taxonomicName>
|
||
by the lack of prefrontal scales and having keeled dorsal scales (smooth in
|
||
<taxonomicName id="BF98C8C315365310FC18094E9B2EFAAD" baseAuthorityName="Uzzell" baseAuthorityYear="1969" box="[966,1062,1309,1334]" class="Reptilia" family="Gymnophthalmidae" genus="Proctoporus" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="xestus">
|
||
<emphasis id="4AEC6F5215365310FC18094E9B2EFAAD" box="[966,1062,1309,1334]" italics="true" pageId="21" pageNumber="22">P.xestus</emphasis>
|
||
</taxonomicName>
|
||
); and from
|
||
<taxonomicName id="BF98C8C315365310FB6B094E9FC7FAC0" baseAuthorityName="Chavez, Siu-ting, Duran & Venegas" baseAuthorityYear="2011" class="Reptilia" family="Gymnophthalmidae" genus="Proctoporus" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="chasqui">
|
||
<emphasis id="4AEC6F5215365310FB6B094E9FC7FAC0" italics="true" pageId="21" pageNumber="22">P. chasqui</emphasis>
|
||
</taxonomicName>
|
||
by the lack of prefrontal scales.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="2CE7E3C815365310FFA4085C9CD5FBBE" ID-DOI="http://doi.org/10.5281/zenodo.4565898" ID-Zenodo-Dep="4565898" box="[122,989,1037,1061]" httpUri="https://zenodo.org/record/4565898/files/figure.png" pageId="21" pageNumber="22" startId="21.[122,159,1039,1060]" targetBox="[120,1224,224,1024]" targetPageId="21">
|
||
<paragraph id="7827B34015365310FFA4085C9CD5FBBE" blockId="21.[122,989,1037,1061]" box="[122,989,1037,1061]" pageId="21" pageNumber="22">
|
||
FIG. 7. Holotype of
|
||
<taxonomicName id="BF98C8C315365310FE9B085D9D1FFBBE" authorityName="Goicoechea & Padial & Chaparro & Castroviejo-Fisher & Riva" authorityYear="2013" box="[325,535,1037,1061]" class="Reptilia" family="Gymnophthalmidae" genus="Proctoporus" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="kiziriani">
|
||
<emphasis id="4AEC6F5215365310FE9B085D9D1FFBBE" box="[325,535,1037,1061]" italics="true" pageId="21" pageNumber="22">Proctoporus kiziriani</emphasis>
|
||
</taxonomicName>
|
||
(MHNC 5366 snout-vent length, 59.1 mm).
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="3082E0CB15365312FF7609349C42FCB6" lastPageId="23" lastPageNumber="24" pageId="21" pageNumber="22" type="description">
|
||
<paragraph id="7827B34015365313FF7609349EF1F9A7" blockId="21.[120,1228,1084,1782]" lastBlockId="22.[120,1225,225,1783]" lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22">
|
||
DESCRIPTION OF
|
||
<typeStatus id="A7230DE215365310FEAE09349EE5FA1A" box="[368,493,1383,1409]" pageId="21" pageNumber="22" type="holotype">HOLOTYPE</typeStatus>
|
||
: Adult male, snout-vent length (SVL)
|
||
<quantity id="BF601EA515365310FC7A09349B04FA1A" box="[932,1036,1383,1410]" metricMagnitude="-2" metricUnit="m" metricValue="5.91" pageId="21" pageNumber="22" unit="mm" value="59.1">59.1 mm</quantity>
|
||
, regenerated tail
|
||
<quantity id="BF601EA515365310FFA609DF9FE9FA3C" box="[120,225,1420,1447]" metricMagnitude="-2" metricUnit="m" metricValue="5.26" pageId="21" pageNumber="22" unit="mm" value="52.6">52.6 mm</quantity>
|
||
; head scales smooth, without striations or rugosities; rostral scale wider (
|
||
<quantity id="BF601EA515365310FBF609DF9B8CFA3C" box="[1064,1156,1420,1447]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="21" pageNumber="22" unit="mm" value="1.7">1.7 mm</quantity>
|
||
) than tall (
|
||
<quantity id="BF601EA515365310FF7509E19E02FA57" box="[171,266,1458,1484]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="21" pageNumber="22" unit="mm" value="0.9">0.9 mm</quantity>
|
||
), meeting supralabials on either side at a point above the height of supralabials and becoming higher medially, in contact with frontonasal, nasals, and first supralabials; frontonasal longer than wide, widest posteriorly, in contact with rostral, nasals, anteriormost superciliary, and frontal; prefrontals absent; frontal longer than wide, pentagonal, not in contact with anteriormost superciliary, in contact with frontonasal, first two supraoculars, and frontoparietals; frontoparietals pentagonal, in contact with frontal, second and third supraoculars, parietals, and interparietal; supraoculars three, all in contact with superciliaries, third in contact with frontoparietals, parietal, and postocular; interparietal longer than wide, pentagonal, in contact with frontoparietals anteriorly, parietals laterally, and occipitals posteriorly; parietals polygonal, with posterior sutures in contact with occipital, lateral sutures diagonally in contact with temporal, third supraocular, and frontoparietal; occipitals five, smaller than parietals, three median smaller than laterals. Nasal entire with no separate loreal scale, longer than high, in contact with rostral, frontonasal, first and second supralabials, first superciliary, and frenocular; nostril situated in anterior third of nasal scale; four superciliaries, first not expanded onto dorsal surface of head; two preoculars, first in contact with first superciliary, frenocular, and nasoloreal scales, second in contact with frenocular and first subocular; frenocular in contact with third supralabial, second preoculars, first subocular, and nasoloreal scales; palpebral disc made up of a single transparent scale; suboculars three; postoculars two; temporals smooth, glossy, polygonal; supratympanic temporals two; supralabials anterior to the posteroventral angle of the subocular four. Mental wider (
|
||
<quantity id="BF601EA515355313FBBC0E619BC8FDD7" box="[1122,1216,562,588]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="22" pageNumber="23" unit="mm" value="1.9">1.9 mm</quantity>
|
||
) than long (
|
||
<quantity id="BF601EA515355313FF280E049E5CFDEA" box="[246,340,599,625]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="22" pageNumber="23" unit="mm" value="1.3">1.3 mm</quantity>
|
||
), in contact with first infralabial and postmental posteriorly; postmental single, pentagonal, in contact with mental, first and second infralabials, and first pair of genials; two pairs of genials in contact, anterior pair in contact with second and third infralabials; second genials in contact with third and fourth infralabials laterally; one pair of chin shields, separated by irregular pregulars; gular scale rows five; collar fold slightly distinct, concealing two rows of small scales; lateral neck scales round, smooth. Dorsals rectangular, longer than wide, juxtaposed, with single keel, in 35 transverse rows; longitudinal dorsal scale rows 23 at midbody; continuous lateral scale series, two to three scales wide, smaller than dorsals; reduced scales at limb-insertion regions present; transverse ventral scale rows 22; longitudinal ventral scale rows at midbody 20; anterior preanal plate scales paired; posterior preanal plate scales six, lateralmost scales small; scales on tail rectangular, juxtaposed; dorsal and dorsolateral caudal scales not keeled, ventral and ventrolateral caudal scales smooth; midventral subcaudal scales wider than adjacent scales, almost square. Limbs pentadactyl; digits clawed; dorsal brachial scales polygonal, subequal in size, subimbricate, smooth; ventral brachial scales roundish, subimbricate, smooth; antebrachial scales polygonal, subequal in size, smooth, ventral antebrachial scales smallest; dorsal manus scales polygonal, smooth, subimbricate; palmar scales small, rounded, subimbricate, domelike; dorsal scales on fingers smooth, quadrangular, covering dorsal half of digit, overhanging subdigital scales, three on finger I, five on II, seven on III, 10 on IV, seven on V; scales on anterodorsal surface of thigh large, polygonal, smooth, subimbricate; scales on posterior surface of thigh small, rounded, juxtaposed; scales on ventral surface of thigh large, rounded, flat, smooth; femoral pores seven; preanal pores absent; scales on anterior surface of crus polygonal, smooth, juxtaposed, decreasing in size distally; scales on anterodorsal surface of crus rounded, juxtaposed; scales on ventral surface of crus large, smooth, flat, subimbricate; scales on dorsal surface of toes quadrangular, smooth, overhanging supradigital lamellae, two on toe I, three on II, eight on III, 10 on IV, seven on V; subdigital lamellae single distally, double proximally, three on toe I, seven on II, nine on III, 19 on IV, 11 on V; limbs not overlapping when adpressed against the body, separated by eight to nine dorsal scale lengths.
|
||
</paragraph>
|
||
<paragraph id="7827B34015355312FF760A149D67FE23" blockId="22.[120,1225,225,1783]" lastBlockId="23.[120,1224,227,1037]" lastPageId="23" lastPageNumber="24" pageId="22" pageNumber="23">COLORATION IN PRESERVATIVE: Dorsal surface of head dark brown (fig. 7); lateral surface of head like dorsal surface, with light mottling; ventral surface of head cream with black spots stippling on each scale. Gular region like head but with denser stippling per scale, forming longitudinal clumps. Dorsal surface of body same color as head, with dark brown spots that form four stripes extending from occiput to just posterior to forelimb insertions, with cream color between stripes; flanks with same coloration as dorsum, fading to lighter brown near venter and with a series of ocelli with dark borders and cream centers. This series of ocelli begins anterior to forelimb insertion, near tympanum, and ends near hind-limb insertion. Ventral surface of body with black ground color, scales with black stippling. Forearms and hind limbs with same background color as dorsum. Dorsal and ventral tail coloration like that of body and gray on the regenerated portion.</paragraph>
|
||
<paragraph id="7827B34015345312FF760D909EFCFDB3" blockId="23.[120,1224,227,1037]" pageId="23" pageNumber="24">COLORATION IN LIFE: Dorsal, lateral, and ventral surfaces are similar to coloration in preservative, but with orange coloration along flanks, forelimbs and tail, not expanding into ventral region of the body (fig. 3C).</paragraph>
|
||
<paragraph id="7827B34015345312FF760E609C42FCB6" blockId="23.[120,1224,227,1037]" pageId="23" pageNumber="24">VARIATION: Coloration is similar among all the specimens, with some having a lighter dorsum and more distinct longitudinal stripes. Stippling on the venter is more distinct or grayer in some specimens and nearly absent in others. The presence of ocelli is also variable. Little sexual dimorphism is observed; the primary difference is that males have wider heads (relative to head length; see table 3). The number of femoral pores is also a sexually dimorphic character, with males possessing 5–7 femoral pores per leg and females lacking them. Variation of morphometric and meristic characters is shown in table 3.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="3082E0CB15345312FF760F6A9BA4FC05" pageId="23" pageNumber="24" type="distribution">
|
||
<paragraph id="7827B34015345312FF760F6A9DB6FCE3" blockId="23.[120,1224,227,1037]" pageId="23" pageNumber="24">
|
||
DISTRIBUTION: This species is known from montane forests of Marcapata Valley, in the department of
|
||
<collectingRegion id="BA5C7DA215345312FEF20F0D9E70FCE3" box="[300,376,862,888]" country="Peru" name="Cusco" pageId="23" pageNumber="24">Cusco</collectingRegion>
|
||
, southeastern
|
||
<collectingCountry id="008FF3D015345312FDF90F0D9D57FCE3" box="[551,607,862,888]" name="Peru" pageId="23" pageNumber="24">Peru</collectingCountry>
|
||
(fig. 9).
|
||
</paragraph>
|
||
<paragraph id="7827B34015345312FF760FD09BA4FC05" blockId="23.[120,1224,227,1037]" box="[168,1196,899,926]" pageId="23" pageNumber="24">HABITAT AND ECOLOGY: Individuals were found during the day under logs and stones.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="3082E0CB15345312FF760FFA9DC1FB96" pageId="23" pageNumber="24" type="etymology">
|
||
<paragraph id="7827B34015345312FF760FFA9DC1FB96" blockId="23.[120,1224,227,1037]" pageId="23" pageNumber="24">ETYMOLOGY: We dedicate this species to our colleague and friend David A. Kizirian (American Museum of Natural History), as a tribute for his outstanding contribution to the knowledge of gymnophthalmid lizard systematics.</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |