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<document ID-CLB-Dataset="6052" ID-DOI="http://dx.doi.org/10.3897/zookeys.1044.64082" ID-GBIF-Dataset="8f5916e6-bf2b-4bdc-9a34-210a8db2dd81" ID-PMC="PMC8222218" ID-Pensoft-Pub="1313-2970-1044-479" ID-Pensoft-UUID="B3EEA3D804B75B92B442D83BD05CA0AB" ID-PubMed="34183883" ID-ZooBank="93F442EAB4BB4BD8AC2A676CA52788D0" ModsDocID="1313-2970-1044-479" checkinTime="1623875385211" checkinUser="pensoft" docAuthor="Casale, Achille" docDate="2021" docId="E633EE75AE01535CA7301C970C8844BB" docLanguage="en" docName="ZooKeys 1044: 479-510" docOrigin="ZooKeys 1044" docPubDate="2021-06-16" docSource="http://dx.doi.org/10.3897/zookeys.1044.64082" docTitle="Calleida solitaria Casale 2021, sp. nov." docType="treatment" docUuid="2C830D71-E6EF-4842-8103-E15383F00626" docUuidSource="ZooBank" docVersion="6" id="B3EEA3D804B75B92B442D83BD05CA0AB" lastPageNumber="479" masterDocId="B3EEA3D804B75B92B442D83BD05CA0AB" masterDocTitle="Biodiversity in tropical rainforests: Calleida Dejean, 1825 at the BIOLAT Biological Station, Rio Manu, Peru, with descriptions of seven new species (Coleoptera Carabidae, Lebiini). Part 1" masterLastPageNumber="510" masterPageNumber="479" pageNumber="479" updateTime="1732751963166" updateUser="ExternalLinkService">
<mods:mods id="10B1F61698F05EE1D0BF3EC2A97204CD" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="364ADE1BC1CDD36320A013B342BB4881">Biodiversity in tropical rainforests: Calleida Dejean, 1825 at the BIOLAT Biological Station, Rio Manu, Peru, with descriptions of seven new species (Coleoptera Carabidae, Lebiini). Part 1</mods:title>
</mods:titleInfo>
<mods:name id="8BCB173B0567ED9B0FE716DBC59EFB53" type="personal">
<mods:role id="73A5A4CE33554295DC6B829952C26D99">
<mods:roleTerm id="0C0A2F89729F88A32A273AA63F3F9B80">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="50E12721BAD3086279BD56D4DAAAFF08">Casale, Achille</mods:namePart>
<mods:affiliation id="B5C50FC66C7D357C920BAB6D10312F85">University of Sassari, Italy (Zoology). Private: Corso Raffaello 12, 10126 Torino, Italy</mods:affiliation>
<mods:nameIdentifier id="ACCD1F7240682C2B579FDB9EF881AADB" type="email">a_casale@libero.it</mods:nameIdentifier>
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<mods:typeOfResource id="7C59E651C82BC8DF255ADBE37D06398E">text</mods:typeOfResource>
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<mods:title id="CB3B1CF5754D4721ADE445E595186FB0">ZooKeys</mods:title>
</mods:titleInfo>
<mods:part id="356684B590116DF1A77F3DFAC3484D53">
<mods:date id="501FDE01745854A07938B53E7418F076">2021</mods:date>
<mods:detail id="8220D60D7F2B8D5D3EC8A4A7BEB386FA" type="pubDate">
<mods:number id="B8773CD2117AF375B411C19DB673B39A">2021-06-16</mods:number>
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<mods:detail id="117ED2B613467A9ABCF9AB0D4E107140" type="volume">
<mods:number id="D9DE352457E340B25F95BDE0871A1622">1044</mods:number>
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<mods:start id="C8AA5859F73800FFC9972EFA5CA7097D">479</mods:start>
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<mods:location id="CA7CCFFC5C12AA60AE8F542A9DF30685">
<mods:url id="3E40CB30FD8B81CCA636C7056C64C754">http://dx.doi.org/10.3897/zookeys.1044.64082</mods:url>
</mods:location>
<mods:classification id="4BD340F42D269FC80AD66721D18CFB91">journal article</mods:classification>
<mods:identifier id="52C3C1F88CFB712EF901887885DB4909" type="DOI">http://dx.doi.org/10.3897/zookeys.1044.64082</mods:identifier>
<mods:identifier id="A1A6246D9E0FBB82F42A7922D90F2295" type="Pensoft-Pub">1313-2970-1044-479</mods:identifier>
<mods:identifier id="4698BFC3E9EC46E961CE377A87ACADF3" type="ZooBank">93F442EAB4BB4BD8AC2A676CA52788D0</mods:identifier>
<mods:identifier id="9FFB6B7448BA856B146D1210F967EC64" type="Pensoft-UUID">B3EEA3D804B75B92B442D83BD05CA0AB</mods:identifier>
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<treatment id="E633EE75AE01535CA7301C970C8844BB" ID-GBIF-Taxon="182753973" LSID="urn:lsid:zoobank.org:act:2C830D71-E6EF-4842-8103-E15383F00626" httpUri="http://treatment.plazi.org/id/E633EE75AE01535CA7301C970C8844BB" lastPageNumber="479" pageId="0" pageNumber="479" scope_family="Carabidae" scope_order="Coleoptera" scope_tribe="Lebiini">
<subSubSection id="40A608E6E2B1316CA151044D78630283" pageId="0" pageNumber="479" type="nomenclature">
<paragraph id="4904DD7D58EDD4905543F97E417E70DA" pageId="0" pageNumber="479">
<taxonomicName id="56F20305B7C9A4E2923DF60666A96D5D" ID-CoL="8MFC6" LSID="http://zoobank.org/2C830D71-E6EF-4842-8103-E15383F00626" authority="Casale, 2021" authorityName="Casale" authorityYear="2021" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Calleida solitaria" order="Coleoptera" pageId="0" pageNumber="479" phylum="Arthropoda" rank="species" species="solitaria" status="sp. nov.">Calleida solitaria</taxonomicName>
<taxonomicNameLabel id="11BFBEA8D89A2F4CA3A6A7CD9E708F93" pageId="0" pageNumber="479">sp. nov.</taxonomicNameLabel>
<figureCitation id="9188801B4329B11D395998B51BD6072B" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Calleida solitaria sp. nov., male holotype, habitus." figureDoi="10.3897/zookeys.1044.64082.figure4" httpUri="https://binary.pensoft.net/fig/554595" pageId="0" pageNumber="479">Figures 4</figureCitation>
<figureCitation id="F55DF446B284AC239B65F3689B5999E0" captionStart="Figures 810" captionStartId="F7" captionText="Figures 8 - 10. Median lobe of aedeagus and abdominal segment IX of Calleida species 8 C. solitaria sp. nov., male holotype 9 C. manuensis sp. nov., male holotype 10 C. anomala sp. nov., male holotype A left lateral aspect B dorsal aspect C abdominal segment IX. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1044.64082.figures8-10" httpUri="https://binary.pensoft.net/fig/554598" pageId="0" pageNumber="479">, 8</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="E482351D11EC22F18AB8EF57675CEB66" pageId="0" pageNumber="479" type="type locality">
<paragraph id="D31E3D57815F7D1941BF48E6A73C4FFF" pageId="0" pageNumber="479">Type locality.</paragraph>
<paragraph id="97CEC94BA314BE48DECAEC7C20D7D581" pageId="0" pageNumber="479">
Peru, Madre de Dios: Rio Manu, Pakitza, 356 m
<geoCoordinate id="EAC2BBA0AEFD42E18B625889B356F462" degrees="12" direction="south" minutes="07" orientation="latitude" precision="925" value="-12.116667">12°07'S</geoCoordinate>
,
<geoCoordinate id="B4B96D405E3A075C4778DAFAF1321C77" degrees="70" direction="west" minutes="58" orientation="longitude" precision="925" value="-70.96667">70°58'W</geoCoordinate>
.
</paragraph>
</subSubSection>
<subSubSection id="9D1DAF815EE750BBCC2FE05566100813" pageId="0" pageNumber="479" type="type material">
<paragraph id="859D3F0979DE1FF06C5C6868D9F6D734" pageId="0" pageNumber="479">Type material.</paragraph>
<paragraph id="8DD79E30063FAEEB1F88A6FF799A576C" pageId="0" pageNumber="479">
<emphasis id="AD7E2D6B0B654114403764D51E62C8EB" bold="true" italics="true" pageId="0" pageNumber="479">Holotype</emphasis>
, male: &quot;PERU, MADRE DE DIOS, Pakitza, 14-20 Oct 90 T.L. Erwin Coll.
<geoCoordinate id="DBDC748A9BDE48A234E1DC2B84195429" degrees="12" direction="south" minutes="07" orientation="latitude" precision="925" value="-12.116667">12°07'S</geoCoordinate>
,
<geoCoordinate id="D61E25C7EC509B6FEB366A454BDC4152" degrees="70" direction="west" minutes="58" orientation="longitude" precision="925" value="-70.96667">70°58'W</geoCoordinate>
, at lab lights&quot;, &quot;BIOLAT/COLE 000006017&quot; (NMNH)
</paragraph>
</subSubSection>
<subSubSection id="905EFE5DB6BFCDDF27E80D3C0B0AF957" pageId="0" pageNumber="479" type="specific epithet">
<paragraph id="F7CF40CA9A07D7E2C1BB08648AD5981F" pageId="0" pageNumber="479">Specific epithet.</paragraph>
<paragraph id="7EAE64A3826150543835B905EF020FD5" pageId="0" pageNumber="479">
Feminine adjective, from the Latin
<emphasis id="6AEA28925436A5AA39C258CBF8DBB9E6" italics="true" pageId="0" pageNumber="479">solitarius, -a</emphasis>
(lonely), which indicates that this new species is known from a single individual only, flying at night.
</paragraph>
<caption id="AA40A5E915311CA1D599A8C637207AB1" doi="10.3897/zookeys.1044.64082.figure4" httpUri="https://binary.pensoft.net/fig/554595" pageId="0" pageNumber="479" start="Figure 4" startId="F4">
<paragraph id="D94980B7275F263B5E3D268AEB85C18F" pageId="0" pageNumber="479">
<emphasis id="DCD1D6415408D38ED652A36130550BEA" bold="true" pageId="0" pageNumber="479">Figure 4.</emphasis>
<taxonomicName id="4D2B5FCF11EF183F0679F48FD44709D2" authorityName="Casale" authorityYear="2021" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Calleida solitaria" order="Coleoptera" pageId="0" pageNumber="479" phylum="Arthropoda" rank="species" species="solitaria">
<emphasis id="BD6F01DB8B26288B4FB78ECE5A24DFFB" italics="true" pageId="0" pageNumber="479">Calleida solitaria</emphasis>
</taxonomicName>
sp. nov., male holotype, habitus.
</paragraph>
</caption>
<caption id="F1C8BB3A2697AB3B5ED1303D33E5C5D6" doi="10.3897/zookeys.1044.64082.figure5" httpUri="https://binary.pensoft.net/fig/554596" pageId="0" pageNumber="479" start="Figure 5" startId="F5">
<paragraph id="5809DC3C8926F0C4644BA7EDCAF743A0" pageId="0" pageNumber="479">
<emphasis id="8D1B964654E5679130D9CE549E540AC5" bold="true" pageId="0" pageNumber="479">Figure 5.</emphasis>
<taxonomicName id="A87C9544E4184D5851284D07348C2F13" authorityName="Casale" authorityYear="2021" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Calleida manuensis" order="Coleoptera" pageId="0" pageNumber="479" phylum="Arthropoda" rank="species" species="manuensis">
<emphasis id="3CC756676C09D8EA408DE170D4553DFF" italics="true" pageId="0" pageNumber="479">Calleida manuensis</emphasis>
</taxonomicName>
sp. nov., male holotype, habitus.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="9BA3ABB2A25B5EA10A83914D88D14495" pageId="0" pageNumber="479" type="diagnosis">
<paragraph id="16CBAE6FA0EE777C987BB606521D6A43" pageId="0" pageNumber="479">Diagnosis.</paragraph>
<paragraph id="B039D99B71772B36F6B779AE724FEC16" pageId="0" pageNumber="479">
With the character states of the genus
<taxonomicName id="647C22DC675BBD063C32D4DDFB39B28D" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Calleida" order="Coleoptera" pageId="0" pageNumber="479" phylum="Arthropoda" rank="genus">
<emphasis id="E307E94AD4CAF3F3A46CF1ADB64906F4" italics="true" pageId="0" pageNumber="479">Calleida</emphasis>
</taxonomicName>
in the narrow sense, but characterized by the peculiar combination of the following morphological features: small-medium sized (TL: mm 7.1); body brown, darkened on head and disc of pronotum; antennae, tibiae, tarsi, and abdominal segments rufous. Pronotum cordiform; elytra elongate, depressed, beaded at apex, with deep and deeply punctate striae. Abdominal sternum VII with one seta on each side in males. Male genitalia as in Fig.
<figureCitation id="64128702EF99A16A35A878942CE77074" captionStart="Figures 810" captionStartId="F7" captionText="Figures 8 - 10. Median lobe of aedeagus and abdominal segment IX of Calleida species 8 C. solitaria sp. nov., male holotype 9 C. manuensis sp. nov., male holotype 10 C. anomala sp. nov., male holotype A left lateral aspect B dorsal aspect C abdominal segment IX. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1044.64082.figures8-10" httpUri="https://binary.pensoft.net/fig/554598" pageId="0" pageNumber="479">8</figureCitation>
. Female unknown.
</paragraph>
</subSubSection>
<subSubSection id="558CD0E49094C1C5F703DEF05FDB62B6" pageId="0" pageNumber="479" type="description">
<paragraph id="E56C3BE3AE7CD8EBCF2185FADD7A3CB3" pageId="0" pageNumber="479">Description.</paragraph>
<paragraph id="5CAFDA469CC63B43D327FBD6B85C66D5" pageId="0" pageNumber="479">
General features as in Fig.
<figureCitation id="C8B57B557D4C99D3ABBCE45FA1D904F0" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Calleida solitaria sp. nov., male holotype, habitus." figureDoi="10.3897/zookeys.1044.64082.figure4" httpUri="https://binary.pensoft.net/fig/554595" pageId="0" pageNumber="479">4</figureCitation>
. Small-medium sized: L: 7.6 mm; TL: 7.1 mm.
</paragraph>
<paragraph id="6CB77F3F46AD52558C2573549119587F" pageId="0" pageNumber="479">
<emphasis id="8E6479C140D819B0794F1E0B7A34281B" bold="true" italics="true" pageId="0" pageNumber="479">Color</emphasis>
: head dark brown, without reddish spots on frons; antennomeres dark rufous. Prothorax and pterothorax brown; pronotum dark reddish at sides. Elytra brown, with slight metallic bronze reflection; tibiae, tarsi, and abdominal segments dark rufous.
</paragraph>
<paragraph id="5670628AB33F02D1B2EF4C1DE20A0830" pageId="0" pageNumber="479">
<emphasis id="AF0F6248FB9C9D76C248812557463F05" bold="true" italics="true" pageId="0" pageNumber="479">Luster and microsculpture</emphasis>
: head and pronotum moderately glossy, with generally effaced cuticular microlines, hardly visible as reticulate meshes; elytra moderately glossy, with cuticular microlines evident in form of transverse pattern.
</paragraph>
<paragraph id="C323B4CE491B4DFC9DB931E386414EC4" pageId="0" pageNumber="479">
<emphasis id="8ACCAEBD2165DE5138484D015760AB15" bold="true" italics="true" pageId="0" pageNumber="479">Head</emphasis>
: wide, transverse, with evident, deep neck constriction; genae very swollen, contiguous with the posterior margin of eyes; frontal furrows deep, connected posteriorly with the supra-orbital keels; eyes very large and prominent; two supraorbital setae on each side.
</paragraph>
<caption id="A0E0170D51960F86151CA1F1E25D7F5C" doi="10.3897/zookeys.1044.64082.figures6-7" httpUri="https://binary.pensoft.net/fig/554597" pageId="0" pageNumber="479" start="Figures 6, 7" startId="F6">
<paragraph id="5A7470E5D5E899A2379C7E55B9CDC991" pageId="0" pageNumber="479">
<emphasis id="BE114C80F545BEB9277EF2623E35123A" bold="true" pageId="0" pageNumber="479">Figures 6, 7.</emphasis>
<taxonomicName id="FBBBC3ACCF8142145A523C7809FACA7D" authorityName="Casale" authorityYear="2021" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Calleida anomala" order="Coleoptera" pageId="0" pageNumber="479" phylum="Arthropoda" rank="species" species="anomala">
<emphasis id="25F791F94DE56F31B156DEE6A17B83D0" italics="true" pageId="0" pageNumber="479">Calleida anomala</emphasis>
</taxonomicName>
sp. nov.
<emphasis id="927014249AD5BF6A5FCA74BE2C272189" bold="true" pageId="0" pageNumber="479">6</emphasis>
male holotype, habitus
<emphasis id="5E6AE0FE90C4F5C29ABB3757C41FBF76" bold="true" pageId="0" pageNumber="479">7</emphasis>
idem, head and pronotum.
</paragraph>
</caption>
<paragraph id="529FEC33042D9C5F4A2FC8A7FD31B80B" pageId="0" pageNumber="479">
<emphasis id="A82DC096F53242A3409CE1712DA2A6B3" bold="true" italics="true" pageId="0" pageNumber="479">Prothorax</emphasis>
: transverse-cordiform (ratio PL/PW: 8.82), with lateral margins moderately sinuate in the posterior third and constricted to the base. Lateral furrows narrow, punctate; lateral reflection slight. Disc convex, with superficial transverse wrinkles. Median furrow deep, reaching the posterior margin. Anterior angles rounded, not prominent; basal angles obtuse but evident. Base almost straight, beaded. One paramedial seta and one basolateral seta on each side present.
</paragraph>
<paragraph id="67CB3780F367F20D0D39294E4997A584" pageId="0" pageNumber="479">
<emphasis id="5C973F558A94D99AF0893B1C5917DBF7" bold="true" italics="true" pageId="0" pageNumber="479">Elytra</emphasis>
: elongate (ratio EL/EW: 1.8), depressed, slightly widened at the apical third; striae deep, deeply punctate; intervals slightly convex. Post-humeral sinuation inconspicuous, pre-apical callosity absent. Apex beaded, oblique, with the outer angle fully rounded and sutural angle prominent. Interval 3 with two discal and one apical setiferous puncture; umbilicate series of 14 large foveate punctures along stria 8, widely interrupted in the middle.
</paragraph>
<paragraph id="BCD97C6DDAA1E70CA08B7C7237D32821" pageId="0" pageNumber="479">
<emphasis id="F60A90EB52B6325F3E78AA28931A012F" bold="true" italics="true" pageId="0" pageNumber="479">Hind wings</emphasis>
: fully developed.
</paragraph>
<paragraph id="761605784E2CF8CE82C888A129F022E9" pageId="0" pageNumber="479">
<emphasis id="5D34863179908CE89A9E6F070FAE2444" bold="true" italics="true" pageId="0" pageNumber="479">Legs</emphasis>
: slender. Metatarsomeres 1-3 superficially grooved dorsally; metatarsomere 4 deeply bilobed, its lobes long and narrow. Tarsal claws denticulate, each with five long teeth on the inner side.
</paragraph>
<paragraph id="7059686314F1F7C6E8B49F6DAFCDDD5A" pageId="0" pageNumber="479">
<emphasis id="C77D3D502D9E73FC2162D14E0A38695E" bold="true" italics="true" pageId="0" pageNumber="479">Abdominal sterna</emphasis>
: sternum VII with one seta on each side in males (unknown, but probably two setae on each side in females).
</paragraph>
<paragraph id="535FA0F59E571C750B09369C9EC06641" pageId="0" pageNumber="479">
<emphasis id="AC2CF3D61A91C3AF2E2077B22D036850" bold="true" italics="true" pageId="0" pageNumber="479">Male genitalia</emphasis>
: median lobe of aedeagus (Fig.
<figureCitation id="43F5192478622C6EFFA233E524CF017C" captionStart="Figures 810" captionStartId="F7" captionText="Figures 8 - 10. Median lobe of aedeagus and abdominal segment IX of Calleida species 8 C. solitaria sp. nov., male holotype 9 C. manuensis sp. nov., male holotype 10 C. anomala sp. nov., male holotype A left lateral aspect B dorsal aspect C abdominal segment IX. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1044.64082.figures8-10" httpUri="https://binary.pensoft.net/fig/554598" pageId="0" pageNumber="479">8</figureCitation>
) very elongate and slender, with basal lobe markedly differentiated. Endophallus with a long flagellum, twisted at base.
</paragraph>
<paragraph id="D6576523978A5EFE9343B82F59C0CB9C" pageId="0" pageNumber="479">
<emphasis id="16AF58BE9EBB79F6891C24C13E3274DB" bold="true" italics="true" pageId="0" pageNumber="479">Female genitalia</emphasis>
: unknown.
</paragraph>
</subSubSection>
<subSubSection id="B3EBBDF1281A4952877D14CD60A9A4D7" pageId="0" pageNumber="479" type="distribution">
<paragraph id="CEF8360E01D006EC49505A87AB6BF790" pageId="0" pageNumber="479">Geographical distribution and habitat.</paragraph>
<paragraph id="59563DC4828FDFD828701A0BE18B1D4C" pageId="0" pageNumber="479">Known so far from the type locality only. It is probably a nocturnal forest-dwelling species at lower altitudes. The single male individual of this taxon was obtained in October, at the lights of the Biolab in Pakitza.</paragraph>
</subSubSection>
<subSubSection id="E875B6AEDF89A40061F6F6CEE81E293B" pageId="0" pageNumber="479" type="relationships">
<paragraph id="A572EDBC6DD51391F688C7D0FB801CCF" pageId="0" pageNumber="479">Relationships.</paragraph>
<paragraph id="AE0323A69EAB81144D38EDAE368E8CC5" pageId="0" pageNumber="479">
For its basic morphological features of shape, size, and characters of the median lobe of aedeagus and endophallus, this species seems to be close to the species of the
<emphasis id="A86EA7E87E3A531061C6CF3752FFC0AD" italics="true" pageId="0" pageNumber="479">cordicollis</emphasis>
(=
<emphasis id="C7CDE76E69323CE10DAB952A460BB52D" italics="true" pageId="0" pageNumber="479">decora</emphasis>
) species group in the sense of Casale in
<bibRefCitation id="242A194C5D3F91B8C590DB24D0C9689A" DOI="https://doi.org/10.1649/0010-065X(2002)056[0071:CMCNSC]2.0.CO;2" author="Desender, K" journalOrPublisher="Bulletin de la Societe Imperiale des naturalistes de Moscou" pageId="0" pageNumber="479" refId="B9" refString="Desender, K, Casale, A, Baert, L, Maelfait, J-P, Verdyck, P, 2002. Calleida migratoria Casale, new species (Coleoptera: Carabidae), a newly introduced ground beetle in the Galapagos Islands, Ecuador. The Coleopterists Bulletin 56(1): 71-78. https://doi.org/10.1649/0010-065X(2002)056[0071:CMCNSC]2.0.CO;2" title="Calleida migratoria Casale, new species (Coleoptera: Carabidae), a newly introduced ground beetle in the Galapagos Islands, Ecuador. The Coleopterists Bulletin 56 (1): 71 - 78." url="https://doi.org/10.1649/0010-065X(2002)056[0071:CMCNSC]2.0.CO;2" year="2002">Desender et al (2002)</bibRefCitation>
, from which it is markedly distinct by the different pattern of color on the dorsal side (mostly metallic in the
<emphasis id="782B6511C11DC7D5F71710F7D80580A1" italics="true" pageId="0" pageNumber="479">cordicollis</emphasis>
group), and by the different number of setae on the abdominal sternum VII (two in male in
<taxonomicName id="D63B7C77649C546B025C643F3070F321" class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="C. solitaria" order="Coleoptera" pageId="0" pageNumber="479" phylum="Arthropoda" rank="species" species="solitaria">
<emphasis id="0C59779AC1071D830D07B4DB5B71A714" italics="true" pageId="0" pageNumber="479">C. solitaria</emphasis>
</taxonomicName>
, three in species of the
<emphasis id="73D87AD50270E5E938C60E3320E992E8" italics="true" pageId="0" pageNumber="479">cordicollis</emphasis>
group).
</paragraph>
<caption id="EAEE44B970F2EF68865BBDAE49C7EC96" doi="10.3897/zookeys.1044.64082.figures8-10" httpUri="https://binary.pensoft.net/fig/554598" pageId="0" pageNumber="479" start="Figures 810" startId="F7">
<paragraph id="ABADDAE123E58C798B12FF409BF52C7C" pageId="0" pageNumber="479">
<emphasis id="E5575E02C046F5AE5B547179EC5060D5" bold="true" pageId="0" pageNumber="479">Figures 8-10.</emphasis>
Median lobe of aedeagus and abdominal segment IX of
<taxonomicName id="BAE77C77909C885ADB1B753A0F0826DC" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Calleida" order="Coleoptera" pageId="0" pageNumber="479" phylum="Arthropoda" rank="genus">
<emphasis id="0F47943A842FBD79B79B2FE1F7C3BF2D" italics="true" pageId="0" pageNumber="479">Calleida</emphasis>
</taxonomicName>
species
<emphasis id="B68228777EEC5EDBDE78FB0843A04109" bold="true" pageId="0" pageNumber="479">8</emphasis>
<taxonomicName id="FC0555152F0B02B69EB16A0948556E9C" class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="C. solitaria" order="Coleoptera" pageId="0" pageNumber="479" phylum="Arthropoda" rank="species" species="solitaria">
<emphasis id="74E0772985544A8E0687A3A0B4E3CACD" italics="true" pageId="0" pageNumber="479">C. solitaria</emphasis>
</taxonomicName>
sp. nov., male holotype
<emphasis id="0DFF78DAA65490570C56B10200E710E4" bold="true" pageId="0" pageNumber="479">9</emphasis>
<taxonomicName id="1FF49EE1BA5A41FC2F3D9C95D3D81D6A" class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="C. manuensis" order="Coleoptera" pageId="0" pageNumber="479" phylum="Arthropoda" rank="species" species="manuensis">
<emphasis id="FFFAECFA3292207584983C5A0638AE67" italics="true" pageId="0" pageNumber="479">C. manuensis</emphasis>
</taxonomicName>
sp. nov., male holotype
<emphasis id="ED17B02922F480B240C2B09F6F2B93B9" bold="true" pageId="0" pageNumber="479">10</emphasis>
<taxonomicName id="A9A7B9106C2BE3BC0A5898D8CC9F2F0E" class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="C. anomala" order="Coleoptera" pageId="0" pageNumber="479" phylum="Arthropoda" rank="species" species="anomala">
<emphasis id="989EA3DB4C12AF0F47546DA744538BE4" italics="true" pageId="0" pageNumber="479">C. anomala</emphasis>
</taxonomicName>
sp. nov., male holotype
<emphasis id="F31B4DF419C0755619AF6DF786099C5F" bold="true" pageId="0" pageNumber="479">A</emphasis>
left lateral aspect
<emphasis id="3591288C00B961F1C307B5AEF753DF24" bold="true" pageId="0" pageNumber="479">B</emphasis>
dorsal aspect
<emphasis id="7D7179905369D2E60D02A1882B7701DE" bold="true" pageId="0" pageNumber="479">C</emphasis>
abdominal segment IX. Scale bar: 1 mm.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>