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<document id="B2225FAAAFF6D2B22509E435401C0CB7" ID-CLB-Dataset="27622" ID-DOI="10.11646/zootaxa.4546.1.1" ID-GBIF-Dataset="aa93de4b-c9ed-4b3e-a1c2-3f5d90faba57" ID-ISSN="1175-5326" ID-Zenodo-Dep="2619020" ID-ZooBank="D2C9F49A-9B76-40AE-9A60-CAE9B99BA547" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553964486275" checkinUser="plazi" docAuthor="Huber, Bernhard A. &amp; Carvalho, Leonardo S." docDate="2019" docId="E21587DBFFA9FFE5FF11FF5A48D7FC92" docLanguage="en" docName="zootaxa.4546.1.1.pdf" docOrigin="Zootaxa 4546 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Arenita Huber &amp; Carvalho 2019, gen. n." docType="treatment" docVersion="16" lastPageNumber="50" masterDocId="1E2CFFA3FF98FFD4FF86FFCD4B03FFA0" masterDocTitle="Filling the gaps: descriptions of unnamed species included in the latest molecular phylogeny of Pholcidae (Araneae)" masterLastPageNumber="96" masterPageNumber="1" pageNumber="50" updateTime="1698716013903" updateUser="ExternalLinkService">
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<mods:title id="DA71DD3F3469D6D5DC6388D8130F9402">Filling the gaps: descriptions of unnamed species included in the latest molecular phylogeny of Pholcidae (Araneae)</mods:title>
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<mods:namePart id="DFF05A1893A1DEB99DBB7554E3C8ADAD">Huber, Bernhard A.</mods:namePart>
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<mods:namePart id="17FDA3C490E79D074E5351F05D5F7E22">Carvalho, Leonardo S.</mods:namePart>
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<mods:date id="695719316B53865DD0B873D2943B0222">2019</mods:date>
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<mods:number id="EE23662D950345CFED1DBED22E462E03">2019-01-22</mods:number>
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<mods:number id="E6B3CBE1FC3DC80AAA5D96AC21BF4069">4546</mods:number>
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<mods:classification id="BDA6773E0E4091E55E80D851E53C19A9">journal article</mods:classification>
<mods:identifier id="FEFD55D52CAD80AA707EB266FE76BDBD" type="CLB-Dataset">27622</mods:identifier>
<mods:identifier id="114E4D45B5664C5021B288E9A8727D6A" type="DOI">10.11646/zootaxa.4546.1.1</mods:identifier>
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<treatment id="E21587DBFFA9FFE5FF11FF5A48D7FC92" ID-DOI="http://doi.org/10.5281/zenodo.5449729" ID-GBIF-Taxon="154531212" ID-Zenodo-Dep="5449729" LSID="urn:lsid:plazi:treatment:E21587DBFFA9FFE5FF11FF5A48D7FC92" httpUri="http://treatment.plazi.org/id/E21587DBFFA9FFE5FF11FF5A48D7FC92" lastPageNumber="50" pageId="49" pageNumber="50">
<subSubSection id="22A66546FFA9FFE5FF11FF5A4943FF12" box="[151,576,151,178]" pageId="49" pageNumber="50" type="nomenclature">
<paragraph id="6A0336CDFFA9FFE5FF11FF5A4943FF12" blockId="49.[151,576,151,178]" box="[151,576,151,178]" pageId="49" pageNumber="50">
<heading id="314B81A1FFA9FFE5FF11FF5A4943FF12" bold="true" box="[151,576,151,178]" fontSize="11" level="1" pageId="49" pageNumber="50" reason="1">
<emphasis id="58C8EADFFFA9FFE5FF11FF5A4943FF12" bold="true" box="[151,576,151,178]" pageId="49" pageNumber="50">
<taxonomicName id="ADBC4D4EFFA9FFE5FF11FF5A4AE5FF12" ID-CoL="32J3" authority="Huber &amp; Carvalho" authorityName="Huber &amp; Carvalho" authorityYear="2019" box="[151,486,151,178]" class="Arachnida" family="Pholcidae" genus="Arenita" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="49" pageNumber="50" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="58C8EADFFFA9FFE5FF11FF5A4BF1FF11" bold="true" box="[151,242,151,177]" italics="true" pageId="49" pageNumber="50">Arenita</emphasis>
Huber &amp; Carvalho
</taxonomicName>
<taxonomicNameLabel id="43FB57A4FFA9FFE5FE68FF554943FF12" box="[494,576,152,178]" pageId="49" pageNumber="50" rank="genus">gen. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="22A66546FFA9FFE5FF11FF134925FF54" box="[151,550,222,245]" pageId="49" pageNumber="50" type="materials_examined">
<paragraph id="6A0336CDFFA9FFE5FF11FF134925FF54" blockId="49.[151,550,222,245]" box="[151,550,222,245]" pageId="49" pageNumber="50">
<emphasis id="58C8EADFFFA9FFE5FF11FF134A1FFF54" bold="true" box="[151,284,222,244]" pageId="49" pageNumber="50">
<typeStatus id="B507886FFFA9FFE5FF11FF134BCFFF54" box="[151,204,222,244]" pageId="49" pageNumber="50">Type</typeStatus>
species
</emphasis>
.
<taxonomicName id="ADBC4D4EFFA9FFE5FEAFFF124AE9FF54" authority="Huber &amp; Carvalho, 2019" authorityName="Huber &amp; Carvalho" authorityYear="2019" box="[297,490,222,244]" class="Arachnida" family="Pholcidae" genus="Arenita" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="fazendinha" status="sp. nov.">
<emphasis id="58C8EADFFFA9FFE5FEAFFF124AE9FF54" box="[297,490,222,244]" italics="true" pageId="49" pageNumber="50">Arenita fazendinha</emphasis>
</taxonomicName>
<taxonomicNameLabel id="43FB57A4FFA9FFE5FE76FF124925FF54" box="[496,550,223,244]" pageId="49" pageNumber="50" rank="species">sp. n.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="22A66546FFA9FFE5FF11FEEC49B3FEFE" pageId="49" pageNumber="50" type="etymology">
<paragraph id="6A0336CDFFA9FFE5FF11FEEC49B3FEFE" blockId="49.[151,1436,289,818]" pageId="49" pageNumber="50">
<emphasis id="58C8EADFFFA9FFE5FF11FEEC4A15FE9A" bold="true" box="[151,278,289,314]" pageId="49" pageNumber="50">Etymology</emphasis>
. The genus name is derived from the Latin
<emphasis id="58C8EADFFFA9FFE5FD7BFEEF483EFE99" box="[765,829,290,313]" italics="true" pageId="49" pageNumber="50">arena</emphasis>
= sand, and refers to the small size of the
<typeStatus id="B507886FFFA9FFE5FA92FEEF4E47FE9A" box="[1300,1348,290,314]" pageId="49" pageNumber="50">type</typeStatus>
species comparable to a grain of sand; gender feminine.
</paragraph>
</subSubSection>
<subSubSection id="22A66546FFA9FFE5FF41FEA448D7FC92" pageId="49" pageNumber="50" type="diagnosis">
<paragraph id="6A0336CDFFA9FFE5FF41FEA44F88FE22" blockId="49.[151,1436,289,818]" box="[199,1163,361,386]" pageId="49" pageNumber="50">
<emphasis id="58C8EADFFFA9FFE5FF41FEA44ACFFE22" bold="true" box="[199,460,361,386]" pageId="49" pageNumber="50">Diagnosis/Description</emphasis>
. See diagnosis and description of single known species below.
</paragraph>
<paragraph id="6A0336CDFFA9FFE5FF41FE4048D7FC92" blockId="49.[151,1436,289,818]" pageId="49" pageNumber="50">
<emphasis id="58C8EADFFFA9FFE5FF41FE404A09FE06" bold="true" box="[199,266,397,422]" pageId="49" pageNumber="50">Notes</emphasis>
. The recent molecular phylogeny of
<taxonomicName id="ADBC4D4EFFA9FFE5FD2DFE4048FFFE06" authority="(Eberle et al. 2018)" authorityName="Eberle" authorityYear="2018" baseAuthorityName="Eberle" baseAuthorityYear="2018" box="[683,1020,397,422]" class="Arachnida" family="Pholcidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="49" pageNumber="50" phylum="Arthropoda" rank="family">
Pholcidae (
<bibRefCitation id="0E2D4B3CFFA9FFE5FCAEFE4048F7FE06" author="Eberle, J. &amp; Dimitrov, D. &amp; Valdez-Mondragon, A. &amp; Huber, B. A." box="[808,1012,397,422]" pageId="49" pageNumber="50" pagination="141" refId="ref37668" refString="Eberle, J., Dimitrov, D., Valdez-Mondragon, A. &amp; Huber, B. A. (2018) Microhabitat change drives diversification in pholcid spiders. BMC Evolutionary Biology, 18, 141. https: // doi. org / 10.1186 / s 12862 - 018 - 1244 - 8" type="journal article" year="2018">
Eberle
<emphasis id="58C8EADFFFA9FFE5FCFCFE4348B0FE06" box="[890,947,397,422]" italics="true" pageId="49" pageNumber="50">et al.</emphasis>
2018
</bibRefCitation>
)
</taxonomicName>
placed this genus/species as sister to an undescribed species from
<collectingCountry id="12AB765DFFA9FFE5FE6EFE7C4954FE6A" box="[488,599,433,458]" name="Argentina" pageId="49" pageNumber="50">Argentina</collectingCountry>
(“Gen.n. MACN270”), and both among ‘basal’
<taxonomicName id="ADBC4D4EFFA9FFE5FB0DFE7C4E20FE6A" authorityName="Simon" authorityYear="1893" box="[1163,1315,433,458]" class="Arachnida" family="Pholcidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="49" pageNumber="50" phylum="Arthropoda" rank="subFamily" subFamily="Modisiminae">Modisiminae</taxonomicName>
. However, relationships among ‘basal’
<taxonomicName id="ADBC4D4EFFA9FFE5FE55FE184964FE4E" authorityName="Simon" authorityYear="1893" box="[467,615,469,494]" class="Arachnida" family="Pholcidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="49" pageNumber="50" phylum="Arthropoda" rank="subFamily" subFamily="Modisiminae">Modisiminae</taxonomicName>
were poorly resolved, so the closest relatives of
<taxonomicName id="ADBC4D4EFFA9FFE5FB0EFE1B4FD8FE4D" authority="Huber &amp; Carvalho, 2019" authorityName="Huber &amp; Carvalho" authorityYear="2019" box="[1160,1243,470,493]" class="Arachnida" family="Pholcidae" genus="Arenita" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="49" pageNumber="4" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="58C8EADFFFA9FFE5FB0EFE1B4FD8FE4D" box="[1160,1243,470,493]" italics="true" pageId="49" pageNumber="50">Arenita</emphasis>
</taxonomicName>
(beyond “Gen.n. MACN270”) remain unknown. The new species described below and “Gen.n. MACN270” are unusual among
<taxonomicName id="ADBC4D4EFFA9FFE5FF11FDD04A28FD96" authorityName="Simon" authorityYear="1893" box="[151,299,541,566]" class="Arachnida" family="Pholcidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="49" pageNumber="50" phylum="Arthropoda" rank="subFamily" subFamily="Modisiminae">Modisiminae</taxonomicName>
in lacking an apophysis on the male palpal coxa, in having the PME and ALE very close together, in lacking a thoracic furrow; and in having the plesiomorphic set of ALS spigots (i.e. with 7–8 spigots on each ALS, see below; this character was not studied in “Gen.n. MACN270”). The latter character is only shared by the mysterious genus
<taxonomicName id="ADBC4D4EFFA9FFE5FEE6FD4449F0FD02" authority="Simon, 1893" authorityName="Simon" authorityYear="1893" box="[352,755,649,674]" class="Arachnida" family="Pholcidae" genus="Priscula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="49" pageNumber="50" phylum="Arthropoda" rank="genus">
<emphasis id="58C8EADFFFA9FFE5FEE6FD444ABCFD02" box="[352,447,649,674]" italics="true" pageId="49" pageNumber="50">Priscula</emphasis>
<bibRefCitation id="0E2D4B3CFFA9FFE5FE42FD444950FD02" author="Simon, E." box="[452,595,649,674]" pageId="49" pageNumber="50" pagination="256 - 488" refId="ref39605" refString="Simon, E. (1893) Histoire Naturelle des Araignees. 1 (2). 2 nd edition. Roret, Paris, 233 pp. [pp. 256 - 488]" type="book chapter" year="1893">Simon, 1893</bibRefCitation>
(
<bibRefCitation id="0E2D4B3CFFA9FFE5FDE7FD4449E8FD02" author="Huber, B. A." box="[609,747,649,674]" pageId="49" pageNumber="50" pagination="1 - 348" refId="ref37855" refString="Huber, B. A. (2000) New World pholcid spiders (Araneae: Pholcidae): a revision at generic level. Bulletin of the American Museum of Natural History, 254, 1 - 348. https: // doi. org / 10.1206 / 0003 - 0090 (2000) 254 &lt;0001: nwpsap&gt; 2.0. co; 2" type="journal article" year="2000">Huber 2000</bibRefCitation>
)
</taxonomicName>
, whose position within
<taxonomicName id="ADBC4D4EFFA9FFE5FC7CFD444F8DFD02" authorityName="Simon" authorityYear="1893" box="[1018,1166,649,674]" class="Arachnida" family="Pholcidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="49" pageNumber="50" phylum="Arthropoda" rank="subFamily" subFamily="Modisiminae">Modisiminae</taxonomicName>
is far from clear (Huber
<emphasis id="58C8EADFFFA9FFE5FF11FD634BD2FD66" box="[151,209,685,710]" italics="true" pageId="49" pageNumber="50">et al.</emphasis>
2018). At the same time,
<taxonomicName id="ADBC4D4EFFA9FFE5FE7AFD6349D1FD66" authorityName="Huber &amp; Carvalho" authorityYear="2019" box="[508,722,685,710]" class="Arachnida" family="Pholcidae" genus="Arenita" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="fazendinha">
<emphasis id="58C8EADFFFA9FFE5FE7AFD6349D1FD66" box="[508,722,685,710]" italics="true" pageId="49" pageNumber="50">Arenita fazendinha</emphasis>
</taxonomicName>
has several highly distinctive characters (see diagnosis below) that are not present in “Gen.n. MACN270”, justifying the creation of a new monotypic genus. The undescribed species “Gen.n. MACN270” will be treated in a separate paper together with further undescribed congeneric species present in Argentinean collections (M.A. Izquierdo, unpubl. data).
</paragraph>
</subSubSection>
</treatment>
</document>
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