194 lines
22 KiB
XML
194 lines
22 KiB
XML
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<mods:title id="3475B58EBCCB51B967DAE7A80675DC93">Filling the gaps: descriptions of unnamed species included in the latest molecular phylogeny of Pholcidae (Araneae)</mods:title>
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<mods:namePart id="CF7560371C3315FAF8CCED1C2A98292B">Huber, Bernhard A.</mods:namePart>
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<mods:namePart id="D2BA4B76E3AB5D50A725B832AEAE48F1">Carvalho, Leonardo S.</mods:namePart>
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<mods:title id="C4A57C514EFF24824EF4946995480A01">Zootaxa</mods:title>
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<mods:date id="D1DB1662BBCF115869969AFAA510070A">2019</mods:date>
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<taxonomicName id="ADBC4D4EFF9DFFD1FF11FEA94AD7FEDE" ID-CoL="62CGD" ID-ENA="280364" authority="Walckenaer, 1837" authorityName="Walckenaer" authorityYear="1837" box="[151,468,356,382]" class="Arachnida" family="Pholcidae" genus="Artema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
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<emphasis id="58C8EADFFF9DFFD1FF11FEA94AD7FEDE" bold="true" box="[151,468,356,382]" pageId="5" pageNumber="6">
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<emphasis id="58C8EADFFF9DFFD1FF11FEA94BF2FEDE" bold="true" box="[151,241,356,382]" italics="true" pageId="5" pageNumber="6">Artema</emphasis>
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<bibRefCitation id="0E2D4B3CFF9DFFD1FF7EFEA94AD7FEDE" author="Walckenaer, C. A." box="[248,468,356,382]" pageId="5" pageNumber="6" pagination="1 - 682" refId="ref39676" refString="Walckenaer, C. A. (1837) Histoire Naturelle des Insectes. Apteres. I. Libraire Encylopedique de Roret, Paris, pp. 1 - 682." type="book chapter" year="1837">Walckenaer, 1837</bibRefCitation>
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</emphasis>
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</taxonomicName>
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<subSubSection id="22A66546FF9DFFD1FF11FE664E6AFDBC" pageId="5" pageNumber="6" type="reference_group">
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<paragraph id="6A0336CDFF9DFFD1FF11FE664FA2FE60" blockId="5.[151,1385,426,541]" box="[151,1185,426,448]" pageId="5" pageNumber="6">
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<treatmentCitationGroup id="4AAC11E3FF9DFFD1FF11FE664FA2FE60" box="[151,1185,426,448]" pageId="5" pageNumber="6">
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<taxonomicName id="ADBC4D4EFF9DFFD1FF11FE664AD0FE60" ID-CoL="62CGD" authority="Walckenaer, 1837: 656" authorityName="Walckenaer" authorityPageNumber="656" authorityYear="1837" box="[151,467,426,448]" class="Arachnida" family="Pholcidae" genus="Artema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
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<emphasis id="58C8EADFFF9DFFD1FF11FE664BE1FE60" box="[151,226,427,448]" italics="true" pageId="5" pageNumber="6">Artema</emphasis>
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<bibRefCitation id="0E2D4B3CFF9DFFD1FF6EFE674AA3FE60" author="Walckenaer, C. A." box="[232,416,426,448]" pageId="5" pageNumber="6" pagination="1 - 682" refId="ref39676" refString="Walckenaer, C. A. (1837) Histoire Naturelle des Insectes. Apteres. I. Libraire Encylopedique de Roret, Paris, pp. 1 - 682." type="book chapter" year="1837">Walckenaer, 1837</bibRefCitation>
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: 656
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;
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<typeStatus id="B507886FFF9DFFD1FE5BFE66490AFE60" box="[477,521,427,448]" pageId="5" pageNumber="6">type</typeStatus>
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species:
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<taxonomicName id="ADBC4D4EFF9DFFD1FDE0FE664886FE60" ID-CoL="5VSKV" authority="Walckenaer, 1837" authorityName="Walckenaer" authorityYear="1837" box="[614,901,426,448]" class="Arachnida" family="Pholcidae" genus="Artema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="atlanta">
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<emphasis id="58C8EADFFF9DFFD1FDE0FE6649C9FE60" box="[614,714,426,448]" italics="true" pageId="5" pageNumber="6">A. atlanta</emphasis>
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<bibRefCitation id="0E2D4B3CFF9DFFD1FD56FE674886FE60" author="Walckenaer, C. A." box="[720,901,426,448]" pageId="5" pageNumber="6" pagination="1 - 682" refId="ref39676" refString="Walckenaer, C. A. (1837) Histoire Naturelle des Insectes. Apteres. I. Libraire Encylopedique de Roret, Paris, pp. 1 - 682." type="book chapter" year="1837">Walckenaer, 1837</bibRefCitation>
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(by subsequent monotypy).
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<paragraph id="6A0336CDFF9DFFD1FF11FE074F3CFE7F" blockId="5.[151,1385,426,541]" box="[151,1087,457,479]" pageId="5" pageNumber="6">
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<treatmentCitationGroup id="4AAC11E3FF9DFFD1FF11FE074F3CFE7F" box="[151,1087,457,479]" pageId="5" pageNumber="6">
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<taxonomicName id="ADBC4D4EFF9DFFD1FF11FE074BE1FE7F" ID-CoL="62CGD" authorityName="Walckenaer" authorityYear="1837" box="[151,226,458,479]" class="Arachnida" family="Pholcidae" genus="Artema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
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<emphasis id="58C8EADFFF9DFFD1FF11FE074BE1FE7F" box="[151,226,458,479]" italics="true" pageId="5" pageNumber="6">Artema</emphasis>
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—
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<treatmentCitation id="EB1D10DCFF9DFFD1FF7DFE044AAFFE7F" author="Simon, E." box="[251,428,457,479]" page="463" pageId="5" pageNumber="6" year="1893">
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<bibRefCitation id="0E2D4B3CFF9DFFD1FF7DFE044A7AFE7F" author="Simon, E." box="[251,377,457,479]" pageId="5" pageNumber="6" pagination="256 - 488" refId="ref39605" refString="Simon, E. (1893) Histoire Naturelle des Araignees. 1 (2). 2 nd edition. Roret, Paris, 233 pp. [pp. 256 - 488]" type="book chapter" year="1893">Simon 1893</bibRefCitation>
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: 463
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</treatmentCitation>
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.
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||
<treatmentCitation id="EB1D10DCFF9DFFD1FE30FE044994FE7F" author="Chrysanthus, O. F. M." box="[438,663,457,479]" page="92" pageId="5" pageNumber="6" year="1967">
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<bibRefCitation id="0E2D4B3CFF9DFFD1FE30FE044972FE7F" author="Chrysanthus, O. F. M." box="[438,625,457,479]" pageId="5" pageNumber="6" pagination="89 - 105" refId="ref37431" refString="Chrysanthus, O. F. M. (1967) Spiders from South New Guinea IX. Tijdschrift voor Entomologie, 110 (5), 89 - 105." type="journal article" year="1967">Chrysanthus 1967</bibRefCitation>
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||
: 92
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||
</treatmentCitation>
|
||
.
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||
<treatmentCitation id="EB1D10DCFF9DFFD1FD27FE044854FE7F" author="Brignoli, P. M." box="[673,855,457,479]" page="91" pageId="5" pageNumber="6" year="1981">
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<bibRefCitation id="0E2D4B3CFF9DFFD1FD27FE044832FE7F" author="Brignoli, P. M." box="[673,817,457,479]" pageId="5" pageNumber="6" pagination="90 - 100" refId="ref37292" refString="Brignoli, P. M. (1981) Studies on the Pholcidae, I. Notes on the genera Artema and Physocyclus (Araneae). Bulletin of the American Museum of Natural History, 170 (1), 90 - 100." type="journal article" year="1981">Brignoli 1981</bibRefCitation>
|
||
: 91
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||
</treatmentCitation>
|
||
.
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||
<treatmentCitation id="EB1D10DCFF9DFFD1FCE7FE044F38FE7F" ID-DOI="http://doi.org/10.5281/zenodo.6524915" ID-Zenodo-Dep="6524915" author="Aharon, S. & Huber, B. A. & Gavish-Regev, E." box="[865,1083,457,479]" httpUri="http://treatment.plazi.org/id/516F87BB130EBC70FD50B9E2FAF2ED91" page="3" pageId="5" pageNumber="6" year="2017">
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<bibRefCitation id="0E2D4B3CFF9DFFD1FCE7FE044F21FE7F" author="Aharon, S. & Huber, B. A. & Gavish-Regev, E." box="[865,1058,457,479]" pageId="5" pageNumber="6" pagination="1 - 57" refId="ref37173" refString="Aharon, S., Huber, B. A. & Gavish-Regev, E. (2017) Daddy-long-leg giants: revision of the spider genus Artema Walckenaer, 1837 (Araneae, Pholcidae). European Journal of Taxonomy, 376, 1 - 57. https: // doi. org / 10.5852 / ejt. 2017.376" type="journal article" year="2017">
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Aharon
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||
<emphasis id="58C8EADFFF9DFFD1FC35FE0748E5FE7F" box="[947,998,457,479]" italics="true" pageId="5" pageNumber="6">et al.</emphasis>
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2017
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</bibRefCitation>
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||
: 3
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||
</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="6A0336CDFF9DFFD1FF11FE254F8DFE5E" blockId="5.[151,1385,426,541]" box="[151,1166,488,510]" pageId="5" pageNumber="6">
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<treatmentCitationGroup id="4AAC11E3FF9DFFD1FF11FE254F8DFE5E" box="[151,1166,488,510]" pageId="5" pageNumber="6">
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<taxonomicName id="ADBC4D4EFF9DFFD1FF11FE254ABFFE5E" authority="Roewer, 1960: 42" authorityName="Roewer" authorityPageNumber="42" authorityYear="1960" box="[151,444,488,510]" class="Arachnida" family="Pholcidae" genus="Pholciella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
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<emphasis id="58C8EADFFF9DFFD1FF11FE254BFCFE5E" box="[151,255,488,510]" italics="true" pageId="5" pageNumber="6">Pholciella</emphasis>
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<treatmentCitation id="EB1D10DCFF9DFFD1FE83FE244ABFFE5E" author="Roewer, C. F." box="[261,444,488,510]" page="42" pageId="5" pageNumber="6" year="1960">
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<bibRefCitation id="0E2D4B3CFF9DFFD1FE83FE244A95FE5E" author="Roewer, C. F." box="[261,406,488,510]" pageId="5" pageNumber="6" pagination="3 - 53" refId="ref39495" refString="Roewer, C. F. (1960) Solifugen und Opilionoiden. Araneae Orthognathae, Haplogynae und Entelegynae. Goteborgs Kungliga Vetenskaps och Vitterhets Samhalles Handlingar, 8 (7), 3 - 53." type="journal article" year="1960">Roewer, 1960</bibRefCitation>
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: 42
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</treatmentCitation>
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</taxonomicName>
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||
;
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<typeStatus id="B507886FFF9DFFD1FE40FE244AF1FE5E" box="[454,498,489,510]" pageId="5" pageNumber="6">type</typeStatus>
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species:
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<taxonomicName id="ADBC4D4EFF9DFFD1FDC8FE24485EFE5E" authority="Roewer, 1960" authorityName="Roewer" authorityYear="1960" box="[590,861,488,510]" class="Arachnida" family="Pholcidae" genus="Pholciella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="ziaretana">
|
||
<emphasis id="58C8EADFFF9DFFD1FDC8FE2449C5FE5E" box="[590,710,489,510]" italics="true" pageId="5" pageNumber="6">P. ziaretana</emphasis>
|
||
<bibRefCitation id="0E2D4B3CFF9DFFD1FD4AFE24485EFE5E" author="Roewer, C. F." box="[716,861,488,510]" pageId="5" pageNumber="6" pagination="3 - 53" refId="ref39495" refString="Roewer, C. F. (1960) Solifugen und Opilionoiden. Araneae Orthognathae, Haplogynae und Entelegynae. Goteborgs Kungliga Vetenskaps och Vitterhets Samhalles Handlingar, 8 (7), 3 - 53." type="journal article" year="1960">Roewer, 1960</bibRefCitation>
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||
</taxonomicName>
|
||
; synonymized in
|
||
<bibRefCitation id="0E2D4B3CFF9DFFD1FB88FE254F8AFE5E" author="Huber, B. A." box="[1038,1161,488,510]" pageId="5" pageNumber="6" pagination="64 - 68" refId="ref38088" refString="Huber, B. A. (2009) Four new generic and 14 new specific synonymies in Pholcidae, and transfer of Pholcoides Roewer to Filistatidae (Araneae). Zootaxa, 1970, 64 - 68." type="journal article" year="2009">Huber 2009</bibRefCitation>
|
||
.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="6A0336CDFF9DFFD1FF11FDCA4E6AFDBC" blockId="5.[151,1385,426,541]" box="[151,1385,518,541]" pageId="5" pageNumber="6">
|
||
<emphasis id="58C8EADFFF9DFFD1FF11FDCA4BDDFDBC" box="[151,222,519,540]" italics="true" pageId="5" pageNumber="6">Coroia</emphasis>
|
||
<bibRefCitation id="0E2D4B3CFF9DFFD1FF62FDCB4AD9FDBC" author="Gonzalez-Sponga, M. A." box="[228,474,518,540]" pageId="5" pageNumber="6" pagination="99 - 109" refId="ref37755" refString="Gonzalez-Sponga, M. A. (2005) Aracnidos de Venezuela. Tres nuevos generos y cuatro nuevas especies de la familia Pholcidae (Araneae). Saber, Universidad de Oriente, Venezuela, 17, 99 - 109." type="journal article" year="2005">González-Sponga, 2005</bibRefCitation>
|
||
: 102;
|
||
<typeStatus id="B507886FFF9DFFD1FD9EFDCA4947FDBC" box="[536,580,519,540]" pageId="5" pageNumber="6">type</typeStatus>
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species:
|
||
<taxonomicName id="ADBC4D4EFF9DFFD1FD26FDCA4F03FDBC" authority="Gonzalez-Sponga, 2005" authorityName="Gonzalez-Sponga" authorityYear="2005" box="[672,1024,518,541]" class="Arachnida" family="Pholcidae" genus="Crossopriza" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="magna">
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||
<emphasis id="58C8EADFFF9DFFD1FD26FDCA4801FDBD" box="[672,770,519,541]" italics="true" pageId="5" pageNumber="6">C. magna</emphasis>
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||
<bibRefCitation id="0E2D4B3CFF9DFFD1FC8FFDCB4F03FDBC" author="Gonzalez-Sponga, M. A." box="[777,1024,518,540]" pageId="5" pageNumber="6" pagination="99 - 109" refId="ref37755" refString="Gonzalez-Sponga, M. A. (2005) Aracnidos de Venezuela. Tres nuevos generos y cuatro nuevas especies de la familia Pholcidae (Araneae). Saber, Universidad de Oriente, Venezuela, 17, 99 - 109." type="journal article" year="2005">González-Sponga, 2005</bibRefCitation>
|
||
</taxonomicName>
|
||
; synonymized in
|
||
<bibRefCitation id="0E2D4B3CFF9DFFD1FB37FDCB4E66FDBC" author="Huber, B. A. & Colmenares, P. A. & Ramirez, M. J." box="[1201,1381,518,541]" pageId="5" pageNumber="6" pagination="413 - 422" refId="ref38794" refString="Huber, B. A., Colmenares, P. A. & Ramirez, M. J. (2014) Fourteen new generic and ten new specific synonymies in Pholcidae (Araneae), and transfer of Mystes Bristowe to Filistatidae. Zootaxa, 3847 (3), 413 - 422. https: // doi. org / 10.11646 / zootaxa. 3847.3.5" type="journal article" year="2014">
|
||
Huber
|
||
<emphasis id="58C8EADFFF9DFFD1FB70FDC54E29FDBC" box="[1270,1322,518,541]" italics="true" pageId="5" pageNumber="6">et al.</emphasis>
|
||
2014
|
||
</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="22A66546FF9DFFD1FF11FD844E5EFC22" pageId="5" pageNumber="6" type="description">
|
||
<paragraph id="6A0336CDFF9DFFD1FF11FD844845FCB5" blockId="5.[151,1437,585,898]" pageId="5" pageNumber="6">
|
||
<emphasis id="58C8EADFFF9DFFD1FF11FD844BD9FDC2" bold="true" box="[151,218,585,610]" pageId="5" pageNumber="6">Notes</emphasis>
|
||
. This genus has been revised recently (
|
||
<bibRefCitation id="0E2D4B3CFF9DFFD1FD13FD84486DFDC1" author="Aharon, S. & Huber, B. A. & Gavish-Regev, E." box="[661,878,585,610]" pageId="5" pageNumber="6" pagination="1 - 57" refId="ref37173" refString="Aharon, S., Huber, B. A. & Gavish-Regev, E. (2017) Daddy-long-leg giants: revision of the spider genus Artema Walckenaer, 1837 (Araneae, Pholcidae). European Journal of Taxonomy, 376, 1 - 57. https: // doi. org / 10.5852 / ejt. 2017.376" type="journal article" year="2017">
|
||
Aharon
|
||
<emphasis id="58C8EADFFF9DFFD1FD74FD874828FDC2" box="[754,811,585,610]" italics="true" pageId="5" pageNumber="6">et al.</emphasis>
|
||
2017
|
||
</bibRefCitation>
|
||
), and the general diagnosis and description of the genus are still largely valid with the new species described below. Two of the new species have slightly more slender legs than previously described species, so male tibia 1 L/d ranges from 34–57; some specimens of
|
||
<taxonomicName id="ADBC4D4EFF9DFFD1FABBFD5F4E9FFD0A" authorityName="Huber & Carvalho" authorityYear="2019" box="[1341,1436,657,682]" class="Arachnida" family="Pholcidae" genus="Artema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="bahla">
|
||
<emphasis id="58C8EADFFF9DFFD1FABBFD5F4E9FFD0A" box="[1341,1436,657,682]" italics="true" pageId="5" pageNumber="6">A. bahla</emphasis>
|
||
</taxonomicName>
|
||
are relatively small, so carapace width ranges from 2.4–4.3; the position of the retrolateral trichobothrium on the male leg tibia 1 ranges from 6–10%; the pair of usually distinct light brown projections in front of the anterior epigynal plate is barely visible in the new species
|
||
<taxonomicName id="ADBC4D4EFF9DFFD1FD41FD33483CFCB6" authorityName="Huber & Carvalho" authorityYear="2019" box="[711,831,765,790]" class="Arachnida" family="Pholcidae" genus="Artema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="ghubrat">
|
||
<emphasis id="58C8EADFFF9DFFD1FD41FD33483CFCB6" box="[711,831,765,790]" italics="true" pageId="5" pageNumber="6">A. ghubrat</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="6A0336CDFF9DFFD1FF41FCEC4E5EFC22" blockId="5.[151,1437,585,898]" pageId="5" pageNumber="6">
|
||
The new species
|
||
<taxonomicName id="ADBC4D4EFF9DFFD1FE0EFCEF4902FC9A" authorityName="Huber & Carvalho" authorityYear="2019" box="[392,513,801,826]" class="Arachnida" family="Pholcidae" genus="Artema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="ghubrat">
|
||
<emphasis id="58C8EADFFF9DFFD1FE0EFCEF4902FC9A" box="[392,513,801,826]" italics="true" pageId="5" pageNumber="6">A. ghubrat</emphasis>
|
||
</taxonomicName>
|
||
seems to be the only (slightly) troglomorphic representative of
|
||
<taxonomicName id="ADBC4D4EFF9DFFD1FB49FCEC4BE9FCFE" authority="(Huber 2018 b)" authorityName="Huber" authorityYear="2018" baseAuthorityName="Huber" baseAuthorityYear="2018" class="Bivalvia" family="Veneridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Venerida" pageId="5" pageNumber="6" phylum="Mollusca" rank="subFamily" subFamily="Arteminae">
|
||
Arteminae (
|
||
<bibRefCitation id="0E2D4B3CFF9DFFD1FAD2FCEC4BE1FCFE" author="Huber, B. A." pageId="5" pageNumber="6" pagination="1 - 18" refId="ref38482" refString="Huber, B. A. (2018 b) Cave-dwelling pholcid spiders (Araneae, Pholcidae): a review. Subterranean Biology, 26, 1 - 18. https: // doi. org / 10.3897 / subtbiol. 26.26430" type="journal article" year="2018">Huber 2018b</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. It is paler than known congeners, never with dark marks on the abdomen, and has the most slender legs (tibia 1 L/d: 57; versus
|
||
<quantity id="AD449B28FF9DFFD1FE1BFCA74903FC22" box="[413,512,873,898]" metricMagnitude="-1" metricUnit="m" metricValue="9.652" metricValueMax="10.668" metricValueMin="8.636" pageId="5" pageNumber="6" unit="in" value="38.0" valueMax="42.0" valueMin="34.0">34–42 in</quantity>
|
||
most other species;
|
||
<quantity id="AD449B28FF9DFFD1FD62FCA4481DFC22" box="[740,798,873,898]" metricMagnitude="0" metricUnit="m" metricValue="1.2953999999999999" pageId="5" pageNumber="6" unit="in" value="51.0">51 in</quantity>
|
||
<taxonomicName id="ADBC4D4EFF9DFFD1FCA2FCA7488DFC22" authorityName="Huber & Carvalho" authorityYear="2019" box="[804,910,873,898]" class="Arachnida" family="Pholcidae" genus="Artema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="dhofar">
|
||
<emphasis id="58C8EADFFF9DFFD1FCA2FCA7488DFC22" box="[804,910,873,898]" italics="true" pageId="5" pageNumber="6">A. dhofar</emphasis>
|
||
</taxonomicName>
|
||
which is also largely restricted to caves).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |