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<document id="4D81CBF1FB61AA0A3768F80FEA317CA4" ID-CLB-Dataset="307388" ID-DOI="10.11646/zootaxa.5555.3.7" ID-GBIF-Dataset="72879b2f-282a-4ef7-862a-679550bad221" ID-ISSN="1175-5326" ID-Zenodo-Dep="14594832" ID-ZooBank="6228E828-0BE0-411B-B555-6D724EF43876" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="diego" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="diego" IM.treatments_approvedBy="diego" checkinTime="1735930107474" checkinUser="plazi" docAuthor="Fleck, Günther &amp; Mézière, Nicolas" docDate="2024" docId="D15A87AAFFD1FFB4FF670FD0EEB6F8F7" docLanguage="en" docName="zootaxa.5555.3.7.pdf" docOrigin="Zootaxa 5555 (3)" docSource="https://doi.org/10.11646/zootaxa.5555.3.7" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Aeschnosoma Selys Longchamps" docType="treatment" docVersion="3" lastPageNumber="448" masterDocId="2D63FFD2FFD4FFB1FFF00F0FEA32FF98" masterDocTitle="Aeschnosoma yelenae sp. nov. from French Guiana (Odonata: Corduliidae), with notes on its phylogenetic affinities and bionomics" masterLastPageNumber="450" masterPageNumber="443" pageNumber="448" updateTime="1736761821554" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="7FC8460E9223737522E73F3225750FD3">Aeschnosoma yelenae sp. nov. from French Guiana (Odonata: Corduliidae), with notes on its phylogenetic affinities and bionomics</mods:title>
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<mods:namePart id="0E52A7F42D0E2C963C46E6BA1AC0EAE9">Fleck, Günther</mods:namePart>
<mods:affiliation id="3199D588ABF5743E0E75F92FDFA1B9EF">07150 Lagorce, France</mods:affiliation>
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<mods:namePart id="F83D79F81ADCA4A1C22F77C4C56291BE">Mézière, Nicolas</mods:namePart>
<mods:affiliation id="CDF24317CBD36918CDA35A6109DCFB24">Cité Cadre COMILOG, Moanda, Gabon</mods:affiliation>
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<mods:date id="E880C0AF85F2D11C2B15C248530FD5CF">2024</mods:date>
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<treatment id="D15A87AAFFD1FFB4FF670FD0EEB6F8F7" ID-DOI="http://doi.org/10.5281/zenodo.14637361" ID-GBIF-Taxon="251994425" ID-Zenodo-Dep="14637361" LSID="urn:lsid:plazi:treatment:D15A87AAFFD1FFB4FF670FD0EEB6F8F7" httpUri="http://treatment.plazi.org/id/D15A87AAFFD1FFB4FF670FD0EEB6F8F7" lastPageNumber="448" pageId="5" pageNumber="448" scope_family="Corduliidae" scope_genus="Aeschnosoma" scope_order="Odonata">
<subSubSection id="11E96537FFD1FFB4FF670FD0EE31FF61" box="[151,1027,223,250]" pageId="5" pageNumber="448" type="nomenclature">
<paragraph id="594C36BCFFD1FFB4FF670FD0EE31FF61" blockId="5.[151,1027,223,250]" box="[151,1027,223,250]" pageId="5" pageNumber="448">
<heading id="020481D0FFD1FFB4FF670FD0EE31FF61" bold="true" box="[151,1027,223,250]" fontSize="11" level="1" pageId="5" pageNumber="448" reason="1">
<emphasis id="6B87EAAEFFD1FFB4FF670FD0EE31FF61" bold="true" box="[151,1027,223,250]" italics="true" pageId="5" pageNumber="448">
Differential diagnosis, and a key to males of
<taxonomicName id="9EF34D3FFFD1FFB4FD430FD0E9B5FF61" ID-CoL="PSY" authority="Selys Longchamps" authorityName="Selys Longchamps" box="[691,903,223,249]" class="Insecta" family="Corduliidae" genus="Aeschnosoma" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="448" phylum="Arthropoda" rank="genus">auripennis-group</taxonomicName>
of species
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="11E96537FFD1FFB4FF670E28EE4AFDA5" pageId="5" pageNumber="448" type="discussion">
<paragraph id="594C36BCFFD1FFB4FF670E28E91EFE49" blockId="5.[151,1436,295,574]" pageId="5" pageNumber="448">
The new species belongs to the genus
<taxonomicName id="9EF34D3FFFD1FFB4FDB30E28E8EFFED9" box="[579,733,295,321]" class="Insecta" family="Corduliidae" genus="Aeschnosoma" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="448" phylum="Arthropoda" rank="genus">
<emphasis id="6B87EAAEFFD1FFB4FDB30E28E8EFFED9" box="[579,733,295,321]" italics="true" pageId="5" pageNumber="448">Aeschnosoma</emphasis>
</taxonomicName>
according to the following diagnosis (which allows separation of the genus from all other New World genera of
<taxonomicName id="9EF34D3FFFD1FFB4FD370E44E979FEFD" box="[711,843,331,357]" class="Insecta" family="Corduliidae" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="448" phylum="Arthropoda" rank="family">Corduliidae</taxonomicName>
s.l.): 1) vertex bilobed, 2) all discoidal triangles and hypertriangles crossed, 3) FW subtriangle crossed, 4) FW discoidal field greatly widened distally (at the level of RP1/2 – RP3/4 fork), 5) FW subdiscoidal space with two or more crossveins (including CuP-crossing), 6) anal loop well developed, foot-shaped with distinct sole and midrib).
</paragraph>
<paragraph id="594C36BCFFD1FFB4FF370ED4EE4AFDA5" blockId="5.[151,1436,295,574]" pageId="5" pageNumber="448">
Fleck (2012) defined different species groups of
<taxonomicName id="9EF34D3FFFD1FFB4FD1E0ED4E9BAFE6D" box="[750,904,475,501]" class="Insecta" family="Corduliidae" genus="Aeschnosoma" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="448" phylum="Arthropoda" rank="genus">
<emphasis id="6B87EAAEFFD1FFB4FD1E0ED4E9BAFE6D" box="[750,904,475,501]" italics="true" pageId="5" pageNumber="448">Aeschnosoma</emphasis>
</taxonomicName>
; together with
<taxonomicName id="9EF34D3FFFD1FFB4FBC40ED4EEFEFE6D" authorityName="Geijskes" authorityYear="1970" box="[1076,1228,475,501]" class="Insecta" family="Corduliidae" genus="Aeschnosoma" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="448" phylum="Arthropoda" rank="species" species="auripennis">
<emphasis id="6B87EAAEFFD1FFB4FBC40ED4EEFEFE6D" box="[1076,1228,475,501]" italics="true" pageId="5" pageNumber="448">A. auripennis</emphasis>
</taxonomicName>
and
<taxonomicName id="9EF34D3FFFD1FFB4FAF40ED4EFA4FE6D" authorityName="Fleck" authorityYear="2012" box="[1284,1430,475,501]" class="Insecta" family="Corduliidae" genus="Aeschnosoma" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="448" phylum="Arthropoda" rank="species" species="heliophila">
<emphasis id="6B87EAAEFFD1FFB4FAF40ED4EFA4FE6D" box="[1284,1430,475,501]" italics="true" pageId="5" pageNumber="448">A. heliophila</emphasis>
</taxonomicName>
,
<taxonomicName id="9EF34D3FFFD1FFB4FF670EF0EB3EFD81" authorityName="Fleck &amp; Mézière" authorityYear="2024" box="[151,268,511,537]" class="Insecta" family="Corduliidae" genus="Aeschnosoma" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="448" phylum="Arthropoda" rank="species" species="yelenae">
<emphasis id="6B87EAAEFFD1FFB4FF670EF0EB3EFD81" box="[151,268,511,537]" italics="true" pageId="5" pageNumber="448">A. yelenae</emphasis>
</taxonomicName>
belongs to the
<taxonomicName id="9EF34D3FFFD1FFB4FE450D0FE81FFD81" authorityName="Geijskes" authorityYear="1970" box="[437,557,512,537]" class="Insecta" family="Corduliidae" genus="Aeschnosoma" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="448" phylum="Arthropoda" rank="species" species="auripennis">
<emphasis id="6B87EAAEFFD1FFB4FE450D0FE81FFD81" box="[437,557,512,537]" italics="true" pageId="5" pageNumber="448">auripennis</emphasis>
</taxonomicName>
-group (diagonosis of this group included in the key below). The presented key separates males of this group from congeners (female and larva of
<taxonomicName id="9EF34D3FFFD1FFB4FC720D2CE9C5FDA5" authorityName="Fleck &amp; Mézière" authorityYear="2024" box="[898,1015,547,573]" class="Insecta" family="Corduliidae" genus="Aeschnosoma" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="448" phylum="Arthropoda" rank="species" species="yelenae">
<emphasis id="6B87EAAEFFD1FFB4FC720D2CE9C5FDA5" box="[898,1015,547,573]" italics="true" pageId="5" pageNumber="448">A. yelenae</emphasis>
</taxonomicName>
unknown):
</paragraph>
</subSubSection>
<subSubSection id="11E96537FFD1FFB4FF670D67EFAEFB7D" pageId="5" pageNumber="448" type="key">
<key id="2D62618DFFD1FFB4FF670D67EFAEFB7D" pageId="5" pageNumber="448">
<keyStep id="E2072E19FFD1FFB4FF670D67EFAEFC3A" pageId="5" pageNumber="448">
<paragraph id="594C36BCFFD1FFB4FF670D67EFAEFC1D" blockId="5.[151,1437,616,931]" pageId="5" pageNumber="448">
<keyLead id="E2029589FFD1FFB4FF670D67EFAEFC1D" pageId="5" pageNumber="448">
1 clypeus, frons and side of thorax brown and lacking distinct and contrasting yellow markings; abdomen brown to dark brown and lacking yellow markings; proportionally very long legs (hind leg length ca 60% of HW length); similar unusual and unique posterior hamule [clearly bipartite with notably, in lateral view, 1) a well-developed inner lobe with apex bent backwards, 2) a distinctive outer lobe bearing a strong anterior spur and a moderately developed posterior hump, and 3) a posterior tuft of very long and strong setae] (cf text above,
<figureCitation id="C1C82A39FFD1FFB4FD750DD2E8F1FD6B" box="[645,707,733,755]" captionStart="FIGURES 1–5" captionStartId="1.[152,270,1937,1961]" captionTargetBox="[151,1436,741,1913]" captionTargetId="figure-166@1.[151,1436,741,1913]" captionTargetPageId="1" captionText="FIGURES 1–5. Aeschnosoma yelenae sp. nov. 1. General habitus, dorsal view. 2. General habitus, left lateral view. 3. Head, dorsal view. 4. Wings, dorsal view. 5. Secondary genitalia, left lateral view." figureDoi="http://doi.org/10.5281/zenodo.14594834" httpUri="https://zenodo.org/record/14594834/files/figure.png" pageId="5" pageNumber="448">Figs 5</figureCitation>
,
<figureCitation id="C1C82A39FFD1FFB4FD3F0DD1E8DAFD6B" box="[719,744,734,755]" captionStart="FIGURES 9–15. 9–10" captionStartId="4.[152,270,1734,1758]" captionTargetBox="[151,1436,181,1711]" captionTargetId="figure-27@4.[151,1436,181,1711]" captionTargetPageId="4" captionText="FIGURES 9–15. 9–10. Aeschnosoma yelenae sp. nov. Posterior abdominal segments and caudal appendages: 9—dorsal view, 10—left lateral view. 11. Aeschnosoma heliophila. Hamule posteriores in right lateral view. 12. Aeschnosoma auripennis. Vesica spermalis in right lateral view. 13–14. S10 and anal appendages in dorsal view: 13—Aeschnosoma auripennis, 14— Aeschnosoma heliophila. 15. Type locality of Aeschnosoma yelenae. The red ellipse depicts the spot where the holotype was flying and caught." figureDoi="http://doi.org/10.5281/zenodo.14594838" httpUri="https://zenodo.org/record/14594838/files/figure.png" pageId="5" pageNumber="448">11</figureCitation>
, and
<bibRefCitation id="3D624B4DFFD1FFB4FCED0DD2E9F4FD6B" author="Neiss, U. G. &amp; Fleck, G. &amp; Feitoza, L. A. &amp; Hamada, N." box="[797,966,733,755]" pageId="5" pageNumber="448" pagination="596 - 599" refId="ref4904" refString="Neiss, U. G., Fleck, G., Feitoza, L. A. &amp; Hamada, N. (2013) Description of the adult male of Aeschnosoma auripennis Geijskes, 1970 (Odonata: Corduliidae s. s.). Zootaxa, 3718 (6), 596 - 599. https: // doi. org / 10.11646 / zootaxa. 3718.6.8" type="journal article" year="2013">
Neiss
<emphasis id="6B87EAAEFFD1FFB4FCAA0DD1E9BEFD6A" box="[858,908,733,755]" italics="true" pageId="5" pageNumber="448">et al.</emphasis>
2013
</bibRefCitation>
); similarly derived and curious
<emphasis id="6B87EAAEFFD1FFB4FB0C0DD1EFAEFD6A" box="[1276,1436,733,755]" italics="true" pageId="5" pageNumber="448">vesica spermalis</emphasis>
[with enormously developed
<collectionCode id="3FE2AE79FFD1FFB4FE080DF4E83CFC88" box="[504,526,763,784]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="5" pageNumber="448" type="Museum">V</collectionCode>
1; small and straight
<collectionCode id="3FE2AE79FFD1FFB4FD200DF4E8D1FC88" box="[720,739,763,784]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="5" pageNumber="448" type="Museum">V</collectionCode>
2; elongated
<collectionCode id="3FE2AE79FFD1FFB4FCAC0DF4E942FC88" box="[860,880,763,784]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="5" pageNumber="448" type="Museum">V</collectionCode>
3, more than twice as long as wide;
<collectionCode id="3FE2AE79FFD1FFB4FB310DF4EEE0FC88" box="[1217,1234,763,784]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="5" pageNumber="448" type="Museum">V</collectionCode>
4 slightly shorter and lacking distinct flagellum and with well developed translucent membrane] (cf text above,
<figureCitation id="C1C82A39FFD1FFB4FBB00C17EEA4FCB6" box="[1088,1174,792,814]" captionStart="FIGURES 6–8" captionStartId="2.[152,270,1322,1346]" captionTargetBox="[151,1436,181,1298]" captionTargetId="figure-27@2.[151,1436,181,1298]" captionTargetPageId="2" captionText="FIGURES 6–8. Vesica spermalis: 6—a) ventral view (anterior on the left side) b) detail of the anterior half, 7—a) right lateral view (anterior on the left side) b) detail of the anterior half, 8—a) left lateral view (anterior on the right side) b) detail of the anterior half" figureDoi="http://doi.org/10.5281/zenodo.14594836" httpUri="https://zenodo.org/record/14594836/files/figure.png" pageId="5" pageNumber="448">Figs 6–8</figureCitation>
,
<figureCitation id="C1C82A39FFD1FFB4FB500C17EE88FCB5" box="[1184,1210,792,813]" captionStart="FIGURES 9–15. 9–10" captionStartId="4.[152,270,1734,1758]" captionTargetBox="[151,1436,181,1711]" captionTargetId="figure-27@4.[151,1436,181,1711]" captionTargetPageId="4" captionText="FIGURES 9–15. 9–10. Aeschnosoma yelenae sp. nov. Posterior abdominal segments and caudal appendages: 9—dorsal view, 10—left lateral view. 11. Aeschnosoma heliophila. Hamule posteriores in right lateral view. 12. Aeschnosoma auripennis. Vesica spermalis in right lateral view. 13–14. S10 and anal appendages in dorsal view: 13—Aeschnosoma auripennis, 14— Aeschnosoma heliophila. 15. Type locality of Aeschnosoma yelenae. The red ellipse depicts the spot where the holotype was flying and caught." figureDoi="http://doi.org/10.5281/zenodo.14594838" httpUri="https://zenodo.org/record/14594838/files/figure.png" pageId="5" pageNumber="448">12</figureCitation>
, and
<bibRefCitation id="3D624B4DFFD1FFB4FB1C0C17EFA0FCB5" author="Neiss, U. G. &amp; Fleck, G. &amp; Feitoza, L. A. &amp; Hamada, N." box="[1260,1426,792,814]" pageId="5" pageNumber="448" pagination="596 - 599" refId="ref4904" refString="Neiss, U. G., Fleck, G., Feitoza, L. A. &amp; Hamada, N. (2013) Description of the adult male of Aeschnosoma auripennis Geijskes, 1970 (Odonata: Corduliidae s. s.). Zootaxa, 3718 (6), 596 - 599. https: // doi. org / 10.11646 / zootaxa. 3718.6.8" type="journal article" year="2013">
Neiss
<emphasis id="6B87EAAEFFD1FFB4FAD80C16EF60FCB5" box="[1320,1362,792,814]" italics="true" pageId="5" pageNumber="448">et al</emphasis>
. 2013
</bibRefCitation>
); anal appendages of similar structure with cerci slightly divergent and straight in their proximal part and strongly bent inwards and somewhat downturned in their distal part, lacking indentation, distally flattened with dorsal membrane and distoventral tuft of strong setae (
<figureCitation id="C1C82A39FFD1FFB4FE8B0C7FE818FC1D" box="[379,554,880,902]" captionStart="FIGURES 9–15. 9–10" captionStartId="4.[152,270,1734,1758]" captionTargetBox="[151,1436,181,1711]" captionTargetId="figure-27@4.[151,1436,181,1711]" captionTargetPageId="4" captionText="FIGURES 9–15. 9–10. Aeschnosoma yelenae sp. nov. Posterior abdominal segments and caudal appendages: 9—dorsal view, 10—left lateral view. 11. Aeschnosoma heliophila. Hamule posteriores in right lateral view. 12. Aeschnosoma auripennis. Vesica spermalis in right lateral view. 13–14. S10 and anal appendages in dorsal view: 13—Aeschnosoma auripennis, 14— Aeschnosoma heliophila. 15. Type locality of Aeschnosoma yelenae. The red ellipse depicts the spot where the holotype was flying and caught." figureDoi="http://doi.org/10.5281/zenodo.14594838" httpUri="https://zenodo.org/record/14594838/files/figure.png" pageId="5" pageNumber="448">Figs. 9–10, 13–14</figureCitation>
).......................................................
<taxonomicName id="9EF34D3FFFD1FFB4FB240C7EEF09FC1E" authorityName="Geijskes" authorityYear="1970" box="[1236,1339,881,902]" class="Insecta" family="Corduliidae" genus="Aeschnosoma" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="448" phylum="Arthropoda" rank="species" species="auripennis">
<emphasis id="6B87EAAEFFD1FFB4FB240C7EEF09FC1E" box="[1236,1339,881,902]" italics="true" pageId="5" pageNumber="448">auripennis</emphasis>
</taxonomicName>
-group (2)
</keyLead>
</paragraph>
<paragraph id="594C36BCFFD1FFB4FF670C81EFAEFC3A" blockId="5.[151,1437,616,931]" box="[151,1436,909,931]" pageId="5" pageNumber="448">
<keyLead id="E2029589FFD1FFB4FF670C81EFAEFC3A" box="[151,1436,909,931]" pageId="5" pageNumber="448">
1’ not as above...........................................................................other
<taxonomicName id="9EF34D3FFFD1FFB4FAE80C82EFAEFC3A" box="[1304,1436,909,930]" class="Insecta" family="Corduliidae" genus="Aeschnosoma" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="448" phylum="Arthropoda" rank="genus">
<emphasis id="6B87EAAEFFD1FFB4FAE80C82EFAEFC3A" box="[1304,1436,909,930]" italics="true" pageId="5" pageNumber="448">Aeschnosoma</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="E2072E19FFD1FFB4FF670CC7EFAEFC63" pageId="5" pageNumber="448">
<paragraph id="594C36BCFFD1FFB4FF670CC7EFAFFC46" blockId="5.[151,1437,968,1019]" box="[151,1437,968,990]" pageId="5" pageNumber="448">
<keyLead id="E2029589FFD1FFB4FF670CC7EFAFFC46" box="[151,1437,968,990]" pageId="5" pageNumber="448">
<quantity id="9E0B9B59FFD1FFB4FF670CC7EAC7FC46" box="[151,245,968,990]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="5" pageNumber="448" unit="in" value="2.0">2 in</quantity>
dorsal view distal part of cercus distinctly bent at obtuse angle (
<figureCitation id="C1C82A39FFD1FFB4FCAE0CC7E991FC45" box="[862,931,968,990]" captionStart="FIGURES 9–15. 9–10" captionStartId="4.[152,270,1734,1758]" captionTargetBox="[151,1436,181,1711]" captionTargetId="figure-27@4.[151,1436,181,1711]" captionTargetPageId="4" captionText="FIGURES 9–15. 9–10. Aeschnosoma yelenae sp. nov. Posterior abdominal segments and caudal appendages: 9—dorsal view, 10—left lateral view. 11. Aeschnosoma heliophila. Hamule posteriores in right lateral view. 12. Aeschnosoma auripennis. Vesica spermalis in right lateral view. 13–14. S10 and anal appendages in dorsal view: 13—Aeschnosoma auripennis, 14— Aeschnosoma heliophila. 15. Type locality of Aeschnosoma yelenae. The red ellipse depicts the spot where the holotype was flying and caught." figureDoi="http://doi.org/10.5281/zenodo.14594838" httpUri="https://zenodo.org/record/14594838/files/figure.png" pageId="5" pageNumber="448">Fig. 13</figureCitation>
)..............................
<taxonomicName id="9EF34D3FFFD1FFB4FAEB0CC7EFAFFC46" authorityName="Geijskes" authorityYear="1970" box="[1307,1437,968,990]" class="Insecta" family="Corduliidae" genus="Aeschnosoma" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="448" phylum="Arthropoda" rank="species" species="auripennis">
<emphasis id="6B87EAAEFFD1FFB4FAEB0CC7EFAFFC46" box="[1307,1437,968,990]" italics="true" pageId="5" pageNumber="448">A. auripennis</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="594C36BCFFD1FFB4FF670CE9EFAEFC63" blockId="5.[151,1437,968,1019]" box="[151,1436,997,1019]" pageId="5" pageNumber="448">
<keyLead id="E2029589FFD1FFB4FF670CE9EFAEFC63" box="[151,1436,997,1019]" pageId="5" pageNumber="448">
2’ in dorsal view distal part of cercus distinctly bent at right angle (
<figureCitation id="C1C82A39FFD1FFB4FCBC0CEAE99CFC63" box="[844,942,997,1019]" captionStart="FIGURES 9–15. 9–10" captionStartId="4.[152,270,1734,1758]" captionTargetBox="[151,1436,181,1711]" captionTargetId="figure-27@4.[151,1436,181,1711]" captionTargetPageId="4" captionText="FIGURES 9–15. 9–10. Aeschnosoma yelenae sp. nov. Posterior abdominal segments and caudal appendages: 9—dorsal view, 10—left lateral view. 11. Aeschnosoma heliophila. Hamule posteriores in right lateral view. 12. Aeschnosoma auripennis. Vesica spermalis in right lateral view. 13–14. S10 and anal appendages in dorsal view: 13—Aeschnosoma auripennis, 14— Aeschnosoma heliophila. 15. Type locality of Aeschnosoma yelenae. The red ellipse depicts the spot where the holotype was flying and caught." figureDoi="http://doi.org/10.5281/zenodo.14594838" httpUri="https://zenodo.org/record/14594838/files/figure.png" pageId="5" pageNumber="448">Figs 9, 14</figureCitation>
)....................................... 3
</keyLead>
</paragraph>
</keyStep>
<keyStep id="E2072E19FFD1FFB4FF670B2FEFAEFB7D" pageId="5" pageNumber="448">
<paragraph id="594C36BCFFD1FFB4FF670B2FEFAEFBE8" blockId="5.[151,1437,1056,1137]" pageId="5" pageNumber="448">
<keyLead id="E2029589FFD1FFB4FF670B2FEFAEFBE8" pageId="5" pageNumber="448">
3 distal half of abdomen, including basal fourth of cerci, brown to orangish brown; epiproct mostly yellowish (except apex dark brown) (
<figureCitation id="C1C82A39FFD1FFB4FEC30B32EB48FBCB" box="[307,378,1085,1107]" captionStart="FIGURES 9–15. 9–10" captionStartId="4.[152,270,1734,1758]" captionTargetBox="[151,1436,181,1711]" captionTargetId="figure-27@4.[151,1436,181,1711]" captionTargetPageId="4" captionText="FIGURES 9–15. 9–10. Aeschnosoma yelenae sp. nov. Posterior abdominal segments and caudal appendages: 9—dorsal view, 10—left lateral view. 11. Aeschnosoma heliophila. Hamule posteriores in right lateral view. 12. Aeschnosoma auripennis. Vesica spermalis in right lateral view. 13–14. S10 and anal appendages in dorsal view: 13—Aeschnosoma auripennis, 14— Aeschnosoma heliophila. 15. Type locality of Aeschnosoma yelenae. The red ellipse depicts the spot where the holotype was flying and caught." figureDoi="http://doi.org/10.5281/zenodo.14594838" httpUri="https://zenodo.org/record/14594838/files/figure.png" pageId="5" pageNumber="448">Fig. 14</figureCitation>
); in dorsal view, distal bend of cercus occupying about 3/10 of length (
<figureCitation id="C1C82A39FFD1FFB4FBE50B32EE69FBCB" box="[1045,1115,1085,1107]" captionStart="FIGURES 9–15. 9–10" captionStartId="4.[152,270,1734,1758]" captionTargetBox="[151,1436,181,1711]" captionTargetId="figure-27@4.[151,1436,181,1711]" captionTargetPageId="4" captionText="FIGURES 9–15. 9–10. Aeschnosoma yelenae sp. nov. Posterior abdominal segments and caudal appendages: 9—dorsal view, 10—left lateral view. 11. Aeschnosoma heliophila. Hamule posteriores in right lateral view. 12. Aeschnosoma auripennis. Vesica spermalis in right lateral view. 13–14. S10 and anal appendages in dorsal view: 13—Aeschnosoma auripennis, 14— Aeschnosoma heliophila. 15. Type locality of Aeschnosoma yelenae. The red ellipse depicts the spot where the holotype was flying and caught." figureDoi="http://doi.org/10.5281/zenodo.14594838" httpUri="https://zenodo.org/record/14594838/files/figure.png" pageId="5" pageNumber="448">Fig. 14</figureCitation>
); in dorsal view; cercus with apex slightly but distinctly recurved (cerci J-shaped,
<figureCitation id="C1C82A39FFD1FFB4FD5A0B54E8DDFBE8" box="[682,751,1115,1137]" captionStart="FIGURES 9–15. 9–10" captionStartId="4.[152,270,1734,1758]" captionTargetBox="[151,1436,181,1711]" captionTargetId="figure-27@4.[151,1436,181,1711]" captionTargetPageId="4" captionText="FIGURES 9–15. 9–10. Aeschnosoma yelenae sp. nov. Posterior abdominal segments and caudal appendages: 9—dorsal view, 10—left lateral view. 11. Aeschnosoma heliophila. Hamule posteriores in right lateral view. 12. Aeschnosoma auripennis. Vesica spermalis in right lateral view. 13–14. S10 and anal appendages in dorsal view: 13—Aeschnosoma auripennis, 14— Aeschnosoma heliophila. 15. Type locality of Aeschnosoma yelenae. The red ellipse depicts the spot where the holotype was flying and caught." figureDoi="http://doi.org/10.5281/zenodo.14594838" httpUri="https://zenodo.org/record/14594838/files/figure.png" pageId="5" pageNumber="448">Fig. 14</figureCitation>
)..............................................
<taxonomicName id="9EF34D3FFFD1FFB4FAD00B54EFAEFBE8" authorityName="Fleck" authorityYear="2012" box="[1312,1436,1115,1136]" class="Insecta" family="Corduliidae" genus="Aeschnosoma" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="448" phylum="Arthropoda" rank="species" species="heliophila">
<emphasis id="6B87EAAEFFD1FFB4FAD00B54EFAEFBE8" box="[1312,1436,1115,1136]" italics="true" pageId="5" pageNumber="448">A. heliophila</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="594C36BCFFD1FFB4FF670B9AEFAEFB7D" blockId="5.[151,1437,1173,1254]" pageId="5" pageNumber="448">
<keyLead id="E2029589FFD1FFB4FF670B9AEFAEFB7D" pageId="5" pageNumber="448">
3’ distal half of abdomen, including proximal part of cerci, dark brown; epiproct dark brown (
<figureCitation id="C1C82A39FFD1FFB4FBAB0B9AEEF2FB32" box="[1115,1216,1173,1195]" captionStart="FIGURES 9–15. 9–10" captionStartId="4.[152,270,1734,1758]" captionTargetBox="[151,1436,181,1711]" captionTargetId="figure-27@4.[151,1436,181,1711]" captionTargetPageId="4" captionText="FIGURES 9–15. 9–10. Aeschnosoma yelenae sp. nov. Posterior abdominal segments and caudal appendages: 9—dorsal view, 10—left lateral view. 11. Aeschnosoma heliophila. Hamule posteriores in right lateral view. 12. Aeschnosoma auripennis. Vesica spermalis in right lateral view. 13–14. S10 and anal appendages in dorsal view: 13—Aeschnosoma auripennis, 14— Aeschnosoma heliophila. 15. Type locality of Aeschnosoma yelenae. The red ellipse depicts the spot where the holotype was flying and caught." figureDoi="http://doi.org/10.5281/zenodo.14594838" httpUri="https://zenodo.org/record/14594838/files/figure.png" pageId="5" pageNumber="448">Figs 9–10</figureCitation>
); in dorsal view, distal bend of cercus occupying about 2/10 of length (
<figureCitation id="C1C82A39FFD1FFB4FD540BBCE8D9FB50" box="[676,747,1203,1225]" captionStart="FIGURES 9–15. 9–10" captionStartId="4.[152,270,1734,1758]" captionTargetBox="[151,1436,181,1711]" captionTargetId="figure-27@4.[151,1436,181,1711]" captionTargetPageId="4" captionText="FIGURES 9–15. 9–10. Aeschnosoma yelenae sp. nov. Posterior abdominal segments and caudal appendages: 9—dorsal view, 10—left lateral view. 11. Aeschnosoma heliophila. Hamule posteriores in right lateral view. 12. Aeschnosoma auripennis. Vesica spermalis in right lateral view. 13–14. S10 and anal appendages in dorsal view: 13—Aeschnosoma auripennis, 14— Aeschnosoma heliophila. 15. Type locality of Aeschnosoma yelenae. The red ellipse depicts the spot where the holotype was flying and caught." figureDoi="http://doi.org/10.5281/zenodo.14594838" httpUri="https://zenodo.org/record/14594838/files/figure.png" pageId="5" pageNumber="448">Fig. 10</figureCitation>
); in dorsal view, cerci with apex not recurved (cerci hockey club-shaped,
<figureCitation id="C1C82A39FFD1FFB4FF120BDFEB15FB7D" box="[226,295,1232,1254]" captionStart="FIGURES 9–15. 9–10" captionStartId="4.[152,270,1734,1758]" captionTargetBox="[151,1436,181,1711]" captionTargetId="figure-27@4.[151,1436,181,1711]" captionTargetPageId="4" captionText="FIGURES 9–15. 9–10. Aeschnosoma yelenae sp. nov. Posterior abdominal segments and caudal appendages: 9—dorsal view, 10—left lateral view. 11. Aeschnosoma heliophila. Hamule posteriores in right lateral view. 12. Aeschnosoma auripennis. Vesica spermalis in right lateral view. 13–14. S10 and anal appendages in dorsal view: 13—Aeschnosoma auripennis, 14— Aeschnosoma heliophila. 15. Type locality of Aeschnosoma yelenae. The red ellipse depicts the spot where the holotype was flying and caught." figureDoi="http://doi.org/10.5281/zenodo.14594838" httpUri="https://zenodo.org/record/14594838/files/figure.png" pageId="5" pageNumber="448">Fig. 10</figureCitation>
)...............................................................................
<taxonomicName id="9EF34D3FFFD1FFB4FB160BDFEF7BFB7D" authorityName="Fleck &amp; Mézière" authorityYear="2024" box="[1254,1353,1232,1253]" class="Insecta" family="Corduliidae" genus="Aeschnosoma" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="448" phylum="Arthropoda" rank="species" species="yelenae" status="sp. nov.">
<emphasis id="6B87EAAEFFD1FFB4FB160BDFEF7BFB7D" box="[1254,1353,1232,1253]" italics="true" pageId="5" pageNumber="448">A. yelenae</emphasis>
</taxonomicName>
<emphasis id="6B87EAAEFFD1FFB4FAA00BDFEFAEFB7D" bold="true" box="[1360,1436,1232,1253]" pageId="5" pageNumber="448">
<taxonomicNameLabel id="70B457D5FFD1FFB4FAA00BDFEFAEFB7D" box="[1360,1436,1232,1253]" pageId="5" pageNumber="448" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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</key>
</subSubSection>
<subSubSection id="11E96537FFD1FFB4FF370A1BEEB6F8F7" pageId="5" pageNumber="448" type="discussion">
<paragraph id="594C36BCFFD1FFB4FF370A1BEB38F8BF" blockId="5.[151,1437,1300,1903]" pageId="5" pageNumber="448">
<taxonomicName id="9EF34D3FFFD1FFB4FF370A1BEB8CFAB6" authorityName="Fleck &amp; Mézière" authorityYear="2024" box="[199,446,1300,1326]" class="Insecta" family="Corduliidae" genus="Aeschnosoma" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="448" phylum="Arthropoda" rank="species" species="yelenae">
<emphasis id="6B87EAAEFFD1FFB4FF370A1BEB8CFAB6" box="[199,446,1300,1326]" italics="true" pageId="5" pageNumber="448">Aeschnosoma yelenae</emphasis>
</taxonomicName>
can be easily separated from its close relatives by the shape of its cerci. This species also exhibits more pronounced melanism, S10 and the epiproct are dark brown whereas S10 is orangish brown and the basal third of the epiproct distinctly yellowish in
<taxonomicName id="9EF34D3FFFD1FFB4FD4C0A52E97FFAEE" authorityName="Fleck" authorityYear="2012" box="[700,845,1372,1399]" class="Insecta" family="Corduliidae" genus="Aeschnosoma" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="448" phylum="Arthropoda" rank="species" species="heliophila">
<emphasis id="6B87EAAEFFD1FFB4FD4C0A52E97FFAEE" box="[700,845,1372,1399]" italics="true" pageId="5" pageNumber="448">A. heliophila</emphasis>
</taxonomicName>
and
<taxonomicName id="9EF34D3FFFD1FFB4FC730A52EE28FAEE" authorityName="Geijskes" authorityYear="1970" box="[899,1050,1373,1399]" class="Insecta" family="Corduliidae" genus="Aeschnosoma" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="448" phylum="Arthropoda" rank="species" species="auripennis">
<emphasis id="6B87EAAEFFD1FFB4FC730A52EE28FAEE" box="[899,1050,1373,1399]" italics="true" pageId="5" pageNumber="448">A. auripennis</emphasis>
</taxonomicName>
. The posterior hamule of the three species differs (compare
<figureCitation id="C1C82A39FFD1FFB4FE440A8FEBC5FA03" box="[436,503,1408,1435]" captionStart="FIGURES 1–5" captionStartId="1.[152,270,1937,1961]" captionTargetBox="[151,1436,741,1913]" captionTargetId="figure-166@1.[151,1436,741,1913]" captionTargetPageId="1" captionText="FIGURES 1–5. Aeschnosoma yelenae sp. nov. 1. General habitus, dorsal view. 2. General habitus, left lateral view. 3. Head, dorsal view. 4. Wings, dorsal view. 5. Secondary genitalia, left lateral view." figureDoi="http://doi.org/10.5281/zenodo.14594834" httpUri="https://zenodo.org/record/14594834/files/figure.png" pageId="5" pageNumber="448">Fig. 5</figureCitation>
with
<figureCitation id="C1C82A39FFD1FFB4FDC90A8FE8BBFA03" box="[569,649,1408,1435]" captionStart="FIGURES 9–15. 9–10" captionStartId="4.[152,270,1734,1758]" captionTargetBox="[151,1436,181,1711]" captionTargetId="figure-27@4.[151,1436,181,1711]" captionTargetPageId="4" captionText="FIGURES 9–15. 9–10. Aeschnosoma yelenae sp. nov. Posterior abdominal segments and caudal appendages: 9—dorsal view, 10—left lateral view. 11. Aeschnosoma heliophila. Hamule posteriores in right lateral view. 12. Aeschnosoma auripennis. Vesica spermalis in right lateral view. 13–14. S10 and anal appendages in dorsal view: 13—Aeschnosoma auripennis, 14— Aeschnosoma heliophila. 15. Type locality of Aeschnosoma yelenae. The red ellipse depicts the spot where the holotype was flying and caught." figureDoi="http://doi.org/10.5281/zenodo.14594838" httpUri="https://zenodo.org/record/14594838/files/figure.png" pageId="5" pageNumber="448">Fig. 11</figureCitation>
and
<figureCitation id="C1C82A39FFD1FFB4FD330A8FE937FA03" box="[707,773,1408,1435]" captionStart="FIGURES 1–5" captionStartId="1.[152,270,1937,1961]" captionTargetBox="[151,1436,741,1913]" captionTargetId="figure-166@1.[151,1436,741,1913]" captionTargetPageId="1" captionText="FIGURES 1–5. Aeschnosoma yelenae sp. nov. 1. General habitus, dorsal view. 2. General habitus, left lateral view. 3. Head, dorsal view. 4. Wings, dorsal view. 5. Secondary genitalia, left lateral view." figureDoi="http://doi.org/10.5281/zenodo.14594834" httpUri="https://zenodo.org/record/14594834/files/figure.png" pageId="5" pageNumber="448">
Fig.
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in
<bibRefCitation id="3D624B4DFFD1FFB4FCDC0A8FE9C0FA03" author="Neiss, U. G. &amp; Fleck, G. &amp; Feitoza, L. A. &amp; Hamada, N." box="[812,1010,1408,1435]" pageId="5" pageNumber="448" pagination="596 - 599" refId="ref4904" refString="Neiss, U. G., Fleck, G., Feitoza, L. A. &amp; Hamada, N. (2013) Description of the adult male of Aeschnosoma auripennis Geijskes, 1970 (Odonata: Corduliidae s. s.). Zootaxa, 3718 (6), 596 - 599. https: // doi. org / 10.11646 / zootaxa. 3718.6.8" type="journal article" year="2013">
Neiss
<emphasis id="6B87EAAEFFD1FFB4FC830A8EE994FA02" box="[883,934,1408,1434]" italics="true" pageId="5" pageNumber="448">et al</emphasis>
. 2013
</bibRefCitation>
): in
<taxonomicName id="9EF34D3FFFD1FFB4FBD50A8EEEA9FA02" authorityName="Fleck &amp; Mézière" authorityYear="2024" box="[1061,1179,1408,1435]" class="Insecta" family="Corduliidae" genus="Aeschnosoma" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="448" phylum="Arthropoda" rank="species" species="yelenae">
<emphasis id="6B87EAAEFFD1FFB4FBD50A8EEEA9FA02" box="[1061,1179,1408,1435]" italics="true" pageId="5" pageNumber="448">A. yelenae</emphasis>
</taxonomicName>
the anterior margin of the inner lobe is distinctly arched and easily visible in lateral view whereas it is distinctly less arched and slightly angled, and also less visible (basal part mostly hidden by outer lobe) in
<taxonomicName id="9EF34D3FFFD1FFB4FC4D0AC6EE67FA7A" authorityName="Geijskes" authorityYear="1970" box="[957,1109,1481,1507]" class="Insecta" family="Corduliidae" genus="Aeschnosoma" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="448" phylum="Arthropoda" rank="species" species="auripennis">
<emphasis id="6B87EAAEFFD1FFB4FC4D0AC6EE67FA7A" box="[957,1109,1481,1507]" italics="true" pageId="5" pageNumber="448">A. auripennis</emphasis>
</taxonomicName>
and
<taxonomicName id="9EF34D3FFFD1FFB4FB7C0AC6EF2FFA7A" authorityName="Fleck" authorityYear="2012" box="[1164,1309,1480,1507]" class="Insecta" family="Corduliidae" genus="Aeschnosoma" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="448" phylum="Arthropoda" rank="species" species="heliophila">
<emphasis id="6B87EAAEFFD1FFB4FB7C0AC6EF2FFA7A" box="[1164,1309,1480,1507]" italics="true" pageId="5" pageNumber="448">A. heliophila</emphasis>
</taxonomicName>
; its apex is rather narrow and rounded whereas it is distinctly larger and subrectangular in
<taxonomicName id="9EF34D3FFFD1FFB4FBF40AE2EEA7F99E" authorityName="Fleck" authorityYear="2012" box="[1028,1173,1516,1543]" class="Insecta" family="Corduliidae" genus="Aeschnosoma" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="448" phylum="Arthropoda" rank="species" species="heliophila">
<emphasis id="6B87EAAEFFD1FFB4FBF40AE2EEA7F99E" box="[1028,1173,1516,1543]" italics="true" pageId="5" pageNumber="448">A. heliophila</emphasis>
</taxonomicName>
in lateral view; the free visible posterior part of the inner lobe is directed distinctly caudad at the base in
<taxonomicName id="9EF34D3FFFD1FFB4FBC1091EEE95F9B2" authorityName="Fleck &amp; Mézière" authorityYear="2024" box="[1073,1191,1552,1579]" class="Insecta" family="Corduliidae" genus="Aeschnosoma" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="448" phylum="Arthropoda" rank="species" species="yelenae">
<emphasis id="6B87EAAEFFD1FFB4FBC1091EEE95F9B2" box="[1073,1191,1552,1579]" italics="true" pageId="5" pageNumber="448">A. yelenae</emphasis>
</taxonomicName>
whereas it is roughly perpendicular to the body axis in
<taxonomicName id="9EF34D3FFFD1FFB4FDE0093AE895F9D6" authorityName="Geijskes" authorityYear="1970" box="[528,679,1589,1615]" class="Insecta" family="Corduliidae" genus="Aeschnosoma" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="448" phylum="Arthropoda" rank="species" species="auripennis">
<emphasis id="6B87EAAEFFD1FFB4FDE0093AE895F9D6" box="[528,679,1589,1615]" italics="true" pageId="5" pageNumber="448">A. auripennis</emphasis>
</taxonomicName>
; in
<taxonomicName id="9EF34D3FFFD1FFB4FD23093AE97BF9D6" authorityName="Fleck &amp; Mézière" authorityYear="2024" box="[723,841,1588,1615]" class="Insecta" family="Corduliidae" genus="Aeschnosoma" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="448" phylum="Arthropoda" rank="species" species="yelenae">
<emphasis id="6B87EAAEFFD1FFB4FD23093AE97BF9D6" box="[723,841,1588,1615]" italics="true" pageId="5" pageNumber="448">A. yelenae</emphasis>
</taxonomicName>
the anterior spur and the posterior hump of the outer lobe are less developed than those of
<taxonomicName id="9EF34D3FFFD1FFB4FDB20956E8E7F9EA" authorityName="Fleck" authorityYear="2012" box="[578,725,1624,1651]" class="Insecta" family="Corduliidae" genus="Aeschnosoma" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="448" phylum="Arthropoda" rank="species" species="heliophila">
<emphasis id="6B87EAAEFFD1FFB4FDB20956E8E7F9EA" box="[578,725,1624,1651]" italics="true" pageId="5" pageNumber="448">A. heliophila</emphasis>
</taxonomicName>
, moreover the margin between these two structures is straight, whereas it is strongly concave in
<taxonomicName id="9EF34D3FFFD1FFB4FDFB0972E8AEF90E" authorityName="Fleck" authorityYear="2012" box="[523,668,1660,1687]" class="Insecta" family="Corduliidae" genus="Aeschnosoma" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="448" phylum="Arthropoda" rank="species" species="heliophila">
<emphasis id="6B87EAAEFFD1FFB4FDFB0972E8AEF90E" box="[523,668,1660,1687]" italics="true" pageId="5" pageNumber="448">A. heliophila</emphasis>
</taxonomicName>
; the anterior spur of
<taxonomicName id="9EF34D3FFFD1FFB4FC770972E9CEF90E" authorityName="Fleck &amp; Mézière" authorityYear="2024" box="[903,1020,1660,1687]" class="Insecta" family="Corduliidae" genus="Aeschnosoma" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="448" phylum="Arthropoda" rank="species" species="yelenae">
<emphasis id="6B87EAAEFFD1FFB4FC770972E9CEF90E" box="[903,1020,1660,1687]" italics="true" pageId="5" pageNumber="448">A. yelenae</emphasis>
</taxonomicName>
is longer and less curved than that of
<taxonomicName id="9EF34D3FFFD1FFB4FF6709AEEB1DF922" authorityName="Geijskes" authorityYear="1970" box="[151,303,1697,1723]" class="Insecta" family="Corduliidae" genus="Aeschnosoma" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="448" phylum="Arthropoda" rank="species" species="auripennis">
<emphasis id="6B87EAAEFFD1FFB4FF6709AEEB1DF922" box="[151,303,1697,1723]" italics="true" pageId="5" pageNumber="448">A. auripennis</emphasis>
</taxonomicName>
and posterior margin of the outer lobe is also less curved in the former.
<collectionCode id="3FE2AE79FFD1FFB4FBA909AEEE5FF923" box="[1113,1133,1697,1723]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="5" pageNumber="448" type="Museum">V</collectionCode>
4 of
<emphasis id="6B87EAAEFFD1FFB4FB5209AEEF6EF922" box="[1186,1372,1696,1722]" italics="true" pageId="5" pageNumber="448">vesica spermalis</emphasis>
has a small apical lateral flap in
<taxonomicName id="9EF34D3FFFD1FFB4FE4B09CAE81CF946" authorityName="Fleck &amp; Mézière" authorityYear="2024" box="[443,558,1732,1759]" class="Insecta" family="Corduliidae" genus="Aeschnosoma" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="448" phylum="Arthropoda" rank="species" species="yelenae">
<emphasis id="6B87EAAEFFD1FFB4FE4B09CAE81CF946" box="[443,558,1732,1759]" italics="true" pageId="5" pageNumber="448">A. yelenae</emphasis>
</taxonomicName>
, this flap is apparently absent or very poorly marked in
<taxonomicName id="9EF34D3FFFD1FFB4FB6709CAEF15F946" authorityName="Fleck" authorityYear="2012" box="[1175,1319,1732,1759]" class="Insecta" family="Corduliidae" genus="Aeschnosoma" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="448" phylum="Arthropoda" rank="species" species="heliophila">
<emphasis id="6B87EAAEFFD1FFB4FB6709CAEF15F946" box="[1175,1319,1732,1759]" italics="true" pageId="5" pageNumber="448">A. heliophila</emphasis>
</taxonomicName>
(character state unknown in the unique
<taxonomicName id="9EF34D3FFFD1FFB4FE2609E6E85EF89A" authorityName="Geijskes" authorityYear="1970" box="[470,620,1769,1795]" class="Insecta" family="Corduliidae" genus="Aeschnosoma" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="448" phylum="Arthropoda" rank="species" species="auripennis">
<emphasis id="6B87EAAEFFD1FFB4FE2609E6E85EF89A" box="[470,620,1769,1795]" italics="true" pageId="5" pageNumber="448">A. auripennis</emphasis>
</taxonomicName>
studied, a dry-stored specimen not allowing clear view of membrane fine structure).
</paragraph>
<paragraph id="594C36BCFFD1FFB4FF37083EEEB6F8F7" blockId="5.[151,1437,1300,1903]" pageId="5" pageNumber="448">
By virtue of their general appearance and the presence of cerci unusually bent at right angles (possible synapomorphy),
<taxonomicName id="9EF34D3FFFD1FFB4FEA5085AEBD4F8F6" authorityName="Fleck" authorityYear="2012" box="[341,486,1876,1903]" class="Insecta" family="Corduliidae" genus="Aeschnosoma" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="448" phylum="Arthropoda" rank="species" species="heliophila">
<emphasis id="6B87EAAEFFD1FFB4FEA5085AEBD4F8F6" box="[341,486,1876,1903]" italics="true" pageId="5" pageNumber="448">A. heliophila</emphasis>
</taxonomicName>
and
<taxonomicName id="9EF34D3FFFD1FFB4FDEC085AE881F8F6" authorityName="Geijskes" authorityYear="1970" box="[540,691,1877,1903]" class="Insecta" family="Corduliidae" genus="Aeschnosoma" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="448" phylum="Arthropoda" rank="species" species="auripennis">
<emphasis id="6B87EAAEFFD1FFB4FDEC085AE881F8F6" box="[540,691,1877,1903]" italics="true" pageId="5" pageNumber="448">A. auripennis</emphasis>
</taxonomicName>
are for the present considered sister taxa.
</paragraph>
</subSubSection>
</treatment>
</document>
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