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<document ID-CLB-Dataset="6049" ID-DOI="http://dx.doi.org/10.3897/zookeys.1044.63775" ID-GBIF-Dataset="e6957b3b-f228-4d6a-b430-35d41cdb2caa" ID-PMC="PMC8222308" ID-Pensoft-Pub="1313-2970-1044-375" ID-Pensoft-UUID="EBD9E81E5EC55AA491952682787793A4" ID-PubMed="34183881" ID-ZooBank="D50CC77C2E6941FDA9BD395B025C43AE" ModsDocID="1313-2970-1044-375" checkinTime="1623875406767" checkinUser="pensoft" docAuthor="Will, Kipling &amp; Gueorguiev, Borislav" docDate="2021" docId="BA3D575E698E5A2698798036B5AA7E67" docLanguage="en" docName="ZooKeys 1044: 375-425" docOrigin="ZooKeys 1044" docPubDate="2021-06-16" docSource="http://dx.doi.org/10.3897/zookeys.1044.63775" docTitle="Coptocarpus erwini Will &amp; Guéorguiev 2021, sp. nov." docType="treatment" docUuid="0C0F90B7-37C7-48DF-B2C6-4F5E55325E1C" docUuidSource="ZooBank" docVersion="6" id="EBD9E81E5EC55AA491952682787793A4" lastPageNumber="375" masterDocId="EBD9E81E5EC55AA491952682787793A4" masterDocTitle="Phylogenetic systematics of the genera of Thryptocerina Jeannel, 1949 and new species from New Caledonia (Coleoptera, Carabidae, Oodini)" masterLastPageNumber="425" masterPageNumber="375" pageNumber="375" updateTime="1732931690188" updateUser="ExternalLinkService">
<mods:mods id="A1D8D70111FB46CAE3C201BF9D232D98" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="94FE6AC83223B60A1379A4F42D4BE142">
<mods:title id="961F8BCD750D589558FB8188F5B6D17D">Phylogenetic systematics of the genera of Thryptocerina Jeannel, 1949 and new species from New Caledonia (Coleoptera, Carabidae, Oodini)</mods:title>
</mods:titleInfo>
<mods:name id="287B5E149FDEE639885F24ACC7D68C6F" type="personal">
<mods:role id="1AC62E8B23944F07D6F7DE2610F85AAE">
<mods:roleTerm id="160EB4E2C0766512B816881D25CE6265">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5631AF838AC4C32B0120207A83B3C215">Will, Kipling</mods:namePart>
<mods:nameIdentifier id="C61B272F1CFACEE6A9C69793FFB9D467" type="ORCID">https://orcid.org/0000-0002-7056-9011</mods:nameIdentifier>
<mods:affiliation id="F96C160035466C7F709B49E320A0CABA">University of California Berkeley, Essig Museum of Entomology. Berkeley, California, USA</mods:affiliation>
<mods:nameIdentifier id="643D23CE550B7B76C774B50C8E45106C" type="email">kipwill@berkeley.edu</mods:nameIdentifier>
</mods:name>
<mods:name id="C11CA83F3A877AE9AE2D1A77963B74F9" type="personal">
<mods:role id="CC7B290235F9C4FE8DCBEC3A4281EF7F">
<mods:roleTerm id="5C2C18F46CE2A901B91D132064A6EA3E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="60A6B33817374BEEF34D56FA44AF6918">Gueorguiev, Borislav</mods:namePart>
<mods:affiliation id="8D256B7BA39F9F8F4002749B58A70A3F">National Museum of Natural History, 1 Blvd. Tsar Osvoboditel, 1000 Sofia, Bulgaria</mods:affiliation>
</mods:name>
<mods:typeOfResource id="3AF092EA791FF0744C04B71C86A7D47B">text</mods:typeOfResource>
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<mods:title id="683476393AABC6228334603E91109ADE">ZooKeys</mods:title>
</mods:titleInfo>
<mods:part id="688BD6183DE52B361FFEFF40A0A66EFF">
<mods:date id="BF945EFE859D941467FACA66CC0C1AC7">2021</mods:date>
<mods:detail id="4E4A3B95279FAAF70A4C5A87C7E73038" type="pubDate">
<mods:number id="B519E18750FD763B38F6D5970345F91B">2021-06-16</mods:number>
</mods:detail>
<mods:detail id="60A0084EFD9616C3C466CB1A4B383C44" type="volume">
<mods:number id="2671D1909E76036957F54BC4B709B6B4">1044</mods:number>
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<mods:start id="B5D9E00250F2B8555C01275DD8C591FE">375</mods:start>
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<mods:url id="6E43575A7CCFB9E3F2F3D012C4EF12A8">http://dx.doi.org/10.3897/zookeys.1044.63775</mods:url>
</mods:location>
<mods:classification id="A2F924532F4E352922820F678DBCEFC5">journal article</mods:classification>
<mods:identifier id="69245750E28DC48192C9813AF08C2F12" type="DOI">http://dx.doi.org/10.3897/zookeys.1044.63775</mods:identifier>
<mods:identifier id="E8FF6C203660E69BFA684BC4A68DAE28" type="Pensoft-Pub">1313-2970-1044-375</mods:identifier>
<mods:identifier id="C48905FBBEC81B700F4CC59BDC7360DA" type="ZooBank">D50CC77C2E6941FDA9BD395B025C43AE</mods:identifier>
<mods:identifier id="2FC6B0B8919165F63CFEB451E2F85D51" type="Pensoft-UUID">EBD9E81E5EC55AA491952682787793A4</mods:identifier>
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<treatment id="BA3D575E698E5A2698798036B5AA7E67" ID-GBIF-Taxon="182754025" LSID="urn:lsid:zoobank.org:act:0C0F90B7-37C7-48DF-B2C6-4F5E55325E1C" httpUri="http://treatment.plazi.org/id/BA3D575E698E5A2698798036B5AA7E67" lastPageNumber="375" pageId="0" pageNumber="375" scope_family="Carabidae" scope_order="Coleoptera" scope_subTribe="Thryptocerina" scope_tribe="Oodini">
<subSubSection id="37F53EE38F77D1F9CAB198A28D352B17" pageId="0" pageNumber="375" type="nomenclature">
<paragraph id="CAA38B9B7007E443BDA3207125AA67EE" pageId="0" pageNumber="375">
<taxonomicName id="46F14C2D931CA03C1306C08DD6B41FF0" ID-CoL="8M8TL" LSID="http://zoobank.org/0C0F90B7-37C7-48DF-B2C6-4F5E55325E1C" authority="Will &amp; Guéorguiev, 2021" authorityName="Will &amp; Guéorguiev" authorityYear="2021" class="Insecta" family="Carabidae" genus="Coptocarpus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coptocarpus erwini" order="Coleoptera" pageId="0" pageNumber="375" phylum="Arthropoda" rank="species" species="erwini" status="sp. nov.">Coptocarpus erwini</taxonomicName>
<taxonomicNameLabel id="299357153F86371E07270E659F1E0D2C" pageId="0" pageNumber="375">sp. nov.</taxonomicNameLabel>
<figureCitation id="CAF881569129087AB69007D4BF3E898A" captionStart="Figures 9–12" captionStartId="F3" captionText="Figures 9 - 12. Female reproductive tract of 9 Coptocarpus grossus Erwin (dorsal view, Margaret River, Western Australia, Australia) 10 Coptocarpus sp. group 6 (ventral view, Mt. Lewis, Queensland, Australia) 11 C. erwini sp. nov. (ventral view, Mt. Humboldt, New Caledonia) 12 C. cyllodinus (Fauvel) (ventral view, Riviere Bleue, New Caledonia). Abbreviations: at, atrium; bc, bursa copulatrix; co, common oviduct; rc, receptaculum; sg, spermatheca gland; smc, seminal canal; spc, spermathecal canal. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1044.63775.figures9-12" httpUri="https://binary.pensoft.net/fig/554050" pageId="0" pageNumber="375">Figs 11</figureCitation>
<figureCitation id="8958F95C20EC873FD19A8CF119DEA497" captionStart="Figures 18–21" captionStartId="F6" captionText="Figures 18 - 21. Dorsal habitus of 18 C. erwini sp. nov. 19 holotype of Coptocarpus cyllodinus, with labels 20 C. amieuensis sp. nov. 21 C. lescheni sp. nov. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.1044.63775.figures18-21" httpUri="https://binary.pensoft.net/fig/554053" pageId="0" pageNumber="375">, 18</figureCitation>
<figureCitation id="66206B97DCBD34600063F8177A3C4F15" captionStart="Figures 22–31" captionStartId="F7" captionText="Figures 22 - 31. Aedeagus, dorsal view of 22 Coptocarpus microps sp. nov. 23 C. erwini sp. nov. 24 C. cyllodinus (Fauvel) 25 C. amieuensis sp. nov. 26 C. magnus sp. nov. 27 Adelopomorpha tethys sp. nov. 28 A. tuberculata sp. nov. 29 A. glabra Heller 30 Adelopomorpha glabra group male from Mt. Koghis 31 Adelopomorpha glabra group male from Gelima." figureDoi="10.3897/zookeys.1044.63775.figures22-31" httpUri="https://binary.pensoft.net/fig/554054" pageId="0" pageNumber="375">, 23</figureCitation>
<figureCitation id="71B749F9371E12C4BC81CEADCEC1D875" captionStart="Figures 32–40" captionStartId="F8" captionText="Figures 32 - 40. Aedeagus, left lateral view of 32 Coptocarpus microps sp. nov. 33 C. erwini sp. nov. 34 C. cyllodinus (Fauvel) 35 C. amieuensis sp. nov. 36 C. magnus sp. nov. 37 Adelopomorpha tethys sp. nov. 38 A. tuberculata sp. nov. 39 A. glabra Heller 40 right lateral view of tip with arrow indicating the dorsal tooth of C. magnus." figureDoi="10.3897/zookeys.1044.63775.figures32-40" httpUri="https://binary.pensoft.net/fig/554055" pageId="0" pageNumber="375">, 33</figureCitation>
<figureCitation id="1C4EBBF3DA002987867C96FF2047889A" captionStart="Figures 46, 47" captionStartId="F12" captionText="Figures 46, 47. Humeral region of left elytron of 46 Coptocarpus erwini sp. nov. 47 C. cyllodinus (Fauvel). Arrow indicates the submarginal carina." figureDoi="10.3897/zookeys.1044.63775.figures46-47" httpUri="https://binary.pensoft.net/fig/554059" pageId="0" pageNumber="375">, 46</figureCitation>
<figureCitation id="975F3DDA9AAE68D107AC214D234B9A02" captionStart="Figures 52, 53" captionStartId="F15" captionText="Figures 52, 53. Maps of localities of New Caledonian Oodini." figureDoi="10.3897/zookeys.1044.63775.figures52-53" httpUri="https://binary.pensoft.net/fig/554062" pageId="0" pageNumber="375">, 53</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="61CECE9C6DAFDCB5988EF687FD1D8C62" pageId="0" pageNumber="375" type="materials_examined">
<paragraph id="96F1CBD861AA5CB9C5FEB47A6333FEFB" pageId="0" pageNumber="375">Material examined.</paragraph>
<paragraph id="9F04C5B6B8A451FB99F89C5B0F092134" pageId="0" pageNumber="375">
<materialsCitation id="A00F626AAA8BE57BBF3D4C2E2C3D50CE" collectingDate="2002-11-06" collectingDateMax="2002-11-07" collectingDateMin="2002-11-06" country="NEW CALEDONIA" elevation="1400" latitude="-21.883333" location="New Caledonia" longLatPrecision="1262" longitude="166.4" specimenCount="♂" typeStatus="Holotype">
<emphasis id="6DCCA51A0F1AC1159EBC7DF59D5A4F42" bold="true" italics="true" pageId="0" pageNumber="375">
<typeStatus id="A0E0850A99D33B17325A464CC17EDF61">Holotype</typeStatus>
</emphasis>
:
<location id="E71689646C14480C9493B6FD15B38945" LSID="urn:lsid:plazi:treatment:BA3D575E698E5A2698798036B5AA7E67:E71689646C14480C9493B6FD15B38945" country="NEW CALEDONIA" latitude="-21.883333" longLatPrecision="1262" longitude="166.4" name="New Caledonia">New Caledonia</location>
•
<specimenCount id="551296E2FDF4C6C4D1C5A4603602C14B">♂</specimenCount>
&quot;
<collectingCountry id="CA0A84E6D57D273FC662E64B4973939F" name="New Caledonia">NEW CALEDONIA</collectingCountry>
11138
<geoCoordinate id="E2D33D810AC485A07E4D83542BD2BFC2" degrees="21" direction="south" minutes="53" orientation="latitude" precision="925" value="-21.883333">21°53'S</geoCoordinate>
x
<geoCoordinate id="52BC1E20EF50F38AB76E81D0255E195F" degrees="166" direction="east" minutes="24" orientation="longitude" precision="925" value="166.4">166°24'E</geoCoordinate>
,
<quantity id="795D38E9008CF794442ADAF1A94C553D" metricMagnitude="3" metricUnit="m" metricValue="1.4" unit="m" value="1400.0">
<elevation id="FEF4CEFF2A70904D387C3E41F43C61F7" metricMagnitude="3" metricUnit="m" metricValue="1.4" unit="m" value="1400.0">1400m</elevation>
</quantity>
Mt. Humboldt, moss forest,
<collectingDate id="6B8368333F33B34542C58D28BFAA9190" value="2002-11-06" valueMax="2002-11-07" valueMin="2002-11-06">6-7 Nov2002</collectingDate>
, Monteith &amp; Burwell, pyreth, trees &amp; logs&quot;
</materialsCitation>
<typeStatus id="8379C6EDC002E6A64ACA5E48273E9773">Holotype</typeStatus>
pinned, with genitalia in a separate microvial, abdominal ventrites and leg glued to point. Source collection QM, deposited MNHN. Type locality as given on label.
</paragraph>
<paragraph id="F97A195323C464DBDBD5B2D1F24BFFB2" pageId="0" pageNumber="375">
<materialsCitation id="3ADAC0CC324C73E850CB8DD42387A5AE" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" location="New Caledonia" specimenCount="♀" typeStatus="Paratypes">
<emphasis id="698CB1FFE05040CEA62C760DB1BCF4D6" bold="true" italics="true" pageId="0" pageNumber="375">
<typeStatus id="E15221A1EC7C5C45784DDFA3219E2012">Paratypes</typeStatus>
</emphasis>
:
<location id="CDD50F9BA5DA7FA8C67E707D09FB11BD" LSID="urn:lsid:plazi:treatment:BA3D575E698E5A2698798036B5AA7E67:CDD50F9BA5DA7FA8C67E707D09FB11BD" name="New Caledonia">New Caledonia</location>
•
<specimenCount id="D17B3CEC16D02F3FC82A901C943FC4DC">♀</specimenCount>
</materialsCitation>
;
<materialsCitation id="0D644574C3D19BD493ED17E3E335F1D1" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" country="New Caledonia" elevation="1400" latitude="-21.883333" location="New Caledonia" longLatPrecision="1262" longitude="166.4" specimenCount="♀" typeStatus="paratype">
same data as holotype [QM] •
<specimenCount id="4B872BF4E724C2039CFD586E820B49DB">♀</specimenCount>
</materialsCitation>
;
<materialsCitation id="C8FDD145C5C3E210E7F0571F357C88F1" collectingDate="1997-02-23" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="Dr. J." country="New Caledonia" elevation="1400" isTailEnumeration="true" latitude="-21.883333" location="Noumea Mt." longLatPrecision="1262" longitude="166.4" specimenCount="♂" typeStatus="paratype">
&quot;NEW-CALEDONIE
<location id="0BCADBE870D6A37B6DB2EE1B5F37ABF6" LSID="urn:lsid:plazi:treatment:BA3D575E698E5A2698798036B5AA7E67:0BCADBE870D6A37B6DB2EE1B5F37ABF6" country="New Caledonia" latitude="-21.883333" longLatPrecision="1262" longitude="166.4" name="Noumea Mt.">
<normalizedToken id="BEDFE59517B1A428D659410EF699DA8C" originalValue="Nouméa">Noumea</normalizedToken>
Mt.
</location>
Koghi //
<collectingDate id="C590B290EB0E906ADC1793A1648B6771" value="1997-02-23">23.II.1997</collectingDate>
leg.
<collectorName id="4580E12BEFA66E6F66C2AA322A8F2363">Dr. J.</collectorName>
BALOGH&quot; [ZSM] •
<specimenCount id="8C91E5205EE5F9852DBCE5BF4B7BEB87">♂</specimenCount>
</materialsCitation>
;
<materialsCitation id="7948340531BA20B3BA1A205ED2F9ADDF" collectingDate="2002-11-05" collectingDateMax="2002-11-08" collectingDateMin="2002-11-05" country="NEW CALEDONIA" county="Night" elevation="1350" location="Sx" longitude="166.41667" municipality="Mt Humboldt" specimenCount="1" typeStatus="paratype">
&quot;
<collectingCountry id="21A330EE7FDD086DF1521760DBB935FB" name="New Caledonia">NEW CALEDONIA</collectingCountry>
11131 21°53'
<location id="D2B30F4BFB467FE44E32177E9AC8F990" LSID="urn:lsid:plazi:treatment:BA3D575E698E5A2698798036B5AA7E67:D2B30F4BFB467FE44E32177E9AC8F990" country="NEW CALEDONIA" county="Night" longitude="166.41667" municipality="Mt Humboldt" name="Sx">Sx</location>
<geoCoordinate id="6F733AE02D898D587FFF8D2146CDB670" degrees="166" direction="east" minutes="25" orientation="longitude" precision="925" value="166.41667">166°25'E</geoCoordinate>
,
<quantity id="42865E2652CCB634BAEC759E117F2C37" metricMagnitude="3" metricUnit="m" metricValue="1.35" unit="m" value="1350.0">
<elevation id="4CE8B54E885AB181CCC423E5E6C937A2" metricMagnitude="3" metricUnit="m" metricValue="1.35" unit="m" value="1350.0">1350 m</elevation>
</quantity>
<collectingMunicipality id="E690A8D464C4C40D4FEBBE7AE9DF1FA2">Mt Humboldt</collectingMunicipality>
refuge,
<collectingCounty id="2C0A80D87BE67EE43D1F5C8B16BE6DEE">Night</collectingCounty>
collecting,
<collectingDate id="C9CB6D4741120552B943D5E44697DF2B" value="2002-11-05" valueMax="2002-11-08" valueMin="2002-11-05">5-8 Nov 2002</collectingDate>
, Burwell, Monteith &amp; Wright&quot; [QM]
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A29806151254516CBF24D43A0C6CD5A5" pageId="0" pageNumber="375" type="diagnosis">
<paragraph id="F964DE70D01AB84A8FC8028240947AD8" pageId="0" pageNumber="375">Diagnosis.</paragraph>
<paragraph id="0679A9737911AB7ADA1E5D50434DC136" pageId="0" pageNumber="375">
<taxonomicName id="4D8A19F97383B79186D23FFD2DDD9DF6" authorityName="Will &amp; Guéorguiev" authorityYear="2021" class="Insecta" family="Carabidae" genus="Coptocarpus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coptocarpus erwini" order="Coleoptera" pageId="0" pageNumber="375" phylum="Arthropoda" rank="species" species="erwini">
<emphasis id="F5665735FAAF18935070723EBEF0BE8C" italics="true" pageId="0" pageNumber="375">Coptocarpus erwini</emphasis>
</taxonomicName>
,
<taxonomicName id="5DFC52F2D1724FDFA845150789E7C010" lsidName="C. lescheni" pageId="0" pageNumber="375" rank="species" species="lescheni">
<emphasis id="F6CF9EB8198EB5A423DF4F0ECE25F720" italics="true" pageId="0" pageNumber="375">C. lescheni</emphasis>
</taxonomicName>
, and
<taxonomicName id="7696DE65CBBF0C6BDC360F3194F07930" lsidName="C. magnus" pageId="0" pageNumber="375" rank="species" species="magnus">
<emphasis id="4C1BAB000F273EE9E11F7E7A503C6FAF" italics="true" pageId="0" pageNumber="375">C. magnus</emphasis>
</taxonomicName>
all have completely impressed elytral striae. Of these three only
<taxonomicName id="23281CFABCB6EFC1B3AFE8103CDF08EE" lsidName="C. erwini" pageId="0" pageNumber="375" rank="species" species="erwini">
<emphasis id="256321B517DA1C94452AEFE20F31710B" italics="true" pageId="0" pageNumber="375">C. erwini</emphasis>
</taxonomicName>
lacks both the elytral submarginal carina (Fig.
<figureCitation id="25C5F65A33EDC415C33A0936C58E1822" captionStart="Figures 46, 47" captionStartId="F12" captionText="Figures 46, 47. Humeral region of left elytron of 46 Coptocarpus erwini sp. nov. 47 C. cyllodinus (Fauvel). Arrow indicates the submarginal carina." figureDoi="10.3897/zookeys.1044.63775.figures46-47" httpUri="https://binary.pensoft.net/fig/554059" pageId="0" pageNumber="375">46</figureCitation>
) and the apicolateral setae of the clypeus (present in
<taxonomicName id="059F281B4019A821C0B8FAF2DA39D20A" lsidName="C. magnus" pageId="0" pageNumber="375" rank="species" species="magnus">
<emphasis id="B2F0DCB3D265E5929470D8B1414E2A82" italics="true" pageId="0" pageNumber="375">C. magnus</emphasis>
</taxonomicName>
). In
<taxonomicName id="84DF5A90024595098E92BDC69F0D7B3A" lsidName="C. erwini" pageId="0" pageNumber="375" rank="species" species="erwini">
<emphasis id="5BD7A80718A738598E5180568CBFFB11" italics="true" pageId="0" pageNumber="375">C. erwini</emphasis>
</taxonomicName>
the antennae are pubescent from antennomere 4 (pubescent from antennomere 2 in
<taxonomicName id="C67007B598459B532D49921D21430809" lsidName="C. lescheni" pageId="0" pageNumber="375" rank="species" species="lescheni">
<emphasis id="26B5A9FCF43F240C89A1B02A0ECACCF8" italics="true" pageId="0" pageNumber="375">C. lescheni</emphasis>
</taxonomicName>
).
</paragraph>
<caption id="1F77E37698DC73260DC28E551EDD8A74" doi="10.3897/zookeys.1044.63775.figures32-40" httpUri="https://binary.pensoft.net/fig/554055" pageId="0" pageNumber="375" start="Figures 32–40" startId="F8">
<paragraph id="2492A7428EE305C04E8D9C0397516813" pageId="0" pageNumber="375">
<emphasis id="F0D41939D583BCB5077678F2D6894D63" bold="true" pageId="0" pageNumber="375">Figures 32-40.</emphasis>
Aedeagus, left lateral view of
<emphasis id="BA6273577541F459172A892630852596" bold="true" pageId="0" pageNumber="375">32</emphasis>
<taxonomicName id="0987CF513CC5B52416254565548F929F" authorityName="Will &amp; Guéorguiev" authorityYear="2021" class="Insecta" family="Carabidae" genus="Coptocarpus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coptocarpus microps" order="Coleoptera" pageId="0" pageNumber="375" phylum="Arthropoda" rank="species" species="microps">
<emphasis id="6CEB8C4CB450A855E57DF01132B33709" italics="true" pageId="0" pageNumber="375">Coptocarpus microps</emphasis>
</taxonomicName>
sp. nov.
<emphasis id="64F11068A05F2E22D0794BA586FAE195" bold="true" pageId="0" pageNumber="375">33</emphasis>
<taxonomicName id="80841E0FADE1DF1E7F8248CE6956CEF0" lsidName="C. erwini" pageId="0" pageNumber="375" rank="species" species="erwini">
<emphasis id="501C06AD1CFE8F5BE5751ED68F2D085C" italics="true" pageId="0" pageNumber="375">C. erwini</emphasis>
</taxonomicName>
sp. nov.
<emphasis id="2725803B821D3728903933B85E2091B5" bold="true" pageId="0" pageNumber="375">34</emphasis>
<taxonomicName id="D487B85A9480258F47791F08078BDCFE" lsidName="C. cyllodinus" pageId="0" pageNumber="375" rank="species" species="cyllodinus">
<emphasis id="521D035CE634D959B56F52737CE7DA9C" italics="true" pageId="0" pageNumber="375">C. cyllodinus</emphasis>
</taxonomicName>
(Fauvel)
<emphasis id="D42A0EE22BAE12DD5ABC17B466F113FF" bold="true" pageId="0" pageNumber="375">35</emphasis>
<taxonomicName id="C0F7106237B8E1022F5F205117F2BC52" lsidName="C. amieuensis" pageId="0" pageNumber="375" rank="species" species="amieuensis">
<emphasis id="D9907B4C6E49A8A542172F30F5E2071B" italics="true" pageId="0" pageNumber="375">C. amieuensis</emphasis>
</taxonomicName>
sp. nov.
<emphasis id="4D723B11431AAB3D7CCDE9FDF5B1D6E1" bold="true" pageId="0" pageNumber="375">36</emphasis>
<taxonomicName id="8971DE4F4C6687B11A61E56685231A80" lsidName="C. magnus" pageId="0" pageNumber="375" rank="species" species="magnus">
<emphasis id="33BEF7B93B2C6A6E4FADB07DC3B0183D" italics="true" pageId="0" pageNumber="375">C. magnus</emphasis>
</taxonomicName>
sp. nov.
<emphasis id="91C50034D46C82DB9CE708AB6B9410B8" bold="true" pageId="0" pageNumber="375">37</emphasis>
<taxonomicName id="93192459E1E58BAEAB385FD6EE7FB1C5" authorityName="Will &amp; Guéorguiev" authorityYear="2021" class="Insecta" family="Carabidae" genus="Adelopomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Adelopomorpha tethys" order="Coleoptera" pageId="0" pageNumber="375" phylum="Arthropoda" rank="species" species="tethys">
<emphasis id="DE0C8C2EE319F8DB707D0FCE10B52808" italics="true" pageId="0" pageNumber="375">Adelopomorpha tethys</emphasis>
</taxonomicName>
sp. nov.
<emphasis id="AF0C140831C3F136CB6CEE53CFF13F98" bold="true" pageId="0" pageNumber="375">38</emphasis>
<taxonomicName id="380633ED1DA71CF5CA7B2649C6FAB0D6" lsidName="A. tuberculata" pageId="0" pageNumber="375" rank="species" species="tuberculata">
<emphasis id="CDC0AD3207158DAAFCF91E6037F7B6C5" italics="true" pageId="0" pageNumber="375">A. tuberculata</emphasis>
</taxonomicName>
sp. nov.
<emphasis id="332385F887335CEE59CA0562628CBD0F" bold="true" pageId="0" pageNumber="375">39</emphasis>
<taxonomicName id="3B995AAF79266FA4CFFEF6BA1FBE6C20" lsidName="A. glabra" pageId="0" pageNumber="375" rank="species" species="glabra">
<emphasis id="C3A90A29F7550B2A1AB8CFA6844ABBB5" italics="true" pageId="0" pageNumber="375">A. glabra</emphasis>
</taxonomicName>
Heller
<emphasis id="FE5C97D7179C92CFAAC41CC804C350E1" bold="true" pageId="0" pageNumber="375">40</emphasis>
right lateral view of tip with arrow indicating the dorsal tooth of
<taxonomicName id="9CE8176E15D021A98406FDC802EE07F2" lsidName="C. magnus" pageId="0" pageNumber="375" rank="species" species="magnus">
<emphasis id="8777E237B206F4BF51D3D741EBDA7921" italics="true" pageId="0" pageNumber="375">C. magnus</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="487E98C44FE6CBB3C06AC1D6D39F1506" pageId="0" pageNumber="375" type="description">
<paragraph id="5CF5253DED130B46CA516AC39964AC40" pageId="0" pageNumber="375">Description.</paragraph>
<paragraph id="828D785B82D21E94CF4307974F218264" pageId="0" pageNumber="375">
<emphasis id="20DEA382190171F94F746F3AD86512E2" bold="true" pageId="0" pageNumber="375">
<emphasis id="A7AAEFDB3E5AFCB390E6C1DCBC73947E" italics="true" pageId="0" pageNumber="375">Habitus</emphasis>
.
</emphasis>
Medium sized, BL: 8.55 mm and BW: 3.65 mm (range, BL: 8.35-8.80 mm, BW 3.55-3.65 mm) ovate, convex body.
<emphasis id="3F660458FE402EC406639092311EAC6C" bold="true" pageId="0" pageNumber="375">
<emphasis id="23691345C3D8E4FC0DAB6CA840A408BB" italics="true" pageId="0" pageNumber="375">Color and luster</emphasis>
.
</emphasis>
Head and pronotum reddish black, elytra black to dark reddish black; ventral surface of body reddish brown to black; legs deep reddish brown, tarsomeres paler brown. Abdominal ventrites reddish black medially and apical edge of ventrite 6; antennae and palpi brown. Integument moderately dull, without spectral iridescence except irregularly ventrally.
<emphasis id="A1788D0E00FEEB62BACA5BF3784D82EE" bold="true" pageId="0" pageNumber="375">
<emphasis id="B0C02F14EFE9B16080A15C8BD1C5FAC7" italics="true" pageId="0" pageNumber="375">Microsculpture and punctation</emphasis>
.
</emphasis>
Dorsal surface of head, pronotum, and elytra with prominent isodiametric meshes; ventral surface with scarcely-visible sculpticells or mostly sculpticells not apparent except on ventrite 6 and apically on prosternum. Head lacking macropunctation or wrinkles on vertex; head, pronotum and elytra with scattered micropunctation; abdominal ventrites 1-3 laterally with coarse punctures, becoming finer and less extensive on 3; ventrites 3-5 very shallowly wrinkled laterally, 4 and 5 smooth medially, and 6 smooth throughout.
<emphasis id="9075869F0E9C1659A91937B2154A5087" bold="true" pageId="0" pageNumber="375">
<emphasis id="E49B295A0362A09CB99760E2C741C53A" italics="true" pageId="0" pageNumber="375">Chaetotaxy</emphasis>
.
</emphasis>
Labrum with six setae, each in its own socket, lateral setae longer than medial four, which are set more closely to each other than to the lateral setae. Clypeal setae absent. Single supraorbital seta absent. Elytron with two discal setiferous punctures in interval 3 (
<normalizedToken id="8E37791B6EA0CF06C0767C4F6027BB9E" originalValue="some×">somex</normalizedToken>
represented by only a puncture). Abdominal ventrites 3-5 with ambulatory setae.
<emphasis id="55BEF5DE71F403D612988333CFC48ECA" bold="true" pageId="0" pageNumber="375">
<emphasis id="CE98A836E91B9770538CACA36F389CF4" italics="true" pageId="0" pageNumber="375">Head</emphasis>
.
</emphasis>
Approximately 1/3 as wide as pronotum (Suppl. material 2: Table S1). Eye size moderate, eye somewhat protruded, EyW/HW: 1.30 (range 1.24-1.30). Labrum with anterior margin deeply emarginate. Frontoclypeal sutures not evident. Antenna moderate length, with last segment nearly reaching pronotal base and pubescence starting from antennomere 4. Last labial palpomere elongate, blunt at apex, length ~ equal to penultimate one. Mentum tooth without paramedial border.
<emphasis id="10E4EF7078D940814E69BBE4BEFF6354" bold="true" pageId="0" pageNumber="375">
<emphasis id="80EAFA95B1715E48A07CD2C3F1DCFC2C" italics="true" pageId="0" pageNumber="375">Thorax</emphasis>
.
</emphasis>
Pronotum elongate, 1 1/2
<normalizedToken id="CF0C64046321CEA262571C9A52F59C88" originalValue="×">x</normalizedToken>
wider than long (PW/PL: 1.54, (range, 1.48-1.54)); width at apex 2 2/3
<normalizedToken id="6C569D8B778C0B7F810863197166500B" originalValue="×">x</normalizedToken>
less than at widest point (PW/PA: 2.37 (range, 2.22-2.41)). Disc with midline fine, well impressed, and ended well before apical and basal margins, without apical transverse impression; anterior angles scarcely convex, anterior margin shallowly concave, submarginal sulcus present near angles, lacking in middle 3/5. Prosternal process acuminate, bordered throughout. Mesosternum with single, low medial tubercle. Metepisternum 1 1/3
<normalizedToken id="B1259138E295DC5FEDA9473D359162CE" originalValue="×">x</normalizedToken>
wider than long, with lateral margin produced posteriorly, coadunation with epipleuron along entire length.
<emphasis id="B57824A0C3DF7B40586D144AC3941E94" bold="true" pageId="0" pageNumber="375">
<emphasis id="905900523C90E05D6B96E32D0BDB3237" italics="true" pageId="0" pageNumber="375">Elytra</emphasis>
.
</emphasis>
Approximately 1/3 longer than wide (EL/EW: 1.27 (range 1.25-1.30)). Basal margin forming prominent, sharp tooth at shoulder, ended medially at level of stria 3. Humeral submarginal carina absent. Elytral apical sinuation not evident. Disc with all striae impressed; parascutellar striole absent or marked by a small, shallow remnant. Elytral intervals flat; granulation in marginal furrow discontinuous, interrupted broadly at midlength.
<emphasis id="61808600596BAAF01F062A2534B541CF" bold="true" pageId="0" pageNumber="375">
<emphasis id="DD6E8EA44C85B148F2D8F54D68BDAD00" italics="true" pageId="0" pageNumber="375">Legs</emphasis>
.
</emphasis>
Mesotibia slightly curved and moderately dilated apically. Basomesotarsus and basometatarsus glabrous dorsally; basomesotarsus in male constricted basally, apically flattened, with 11 or 12 short, stout setae ventrally and many smaller setae scattered on lateral portions. In female, basomesotarsus cylindrical and with scattered, fine dorsal setae and rows of moderately thicker ventral setae.
<emphasis id="7D9618E5D502551310C6C8A2A9FC414B" bold="true" pageId="0" pageNumber="375">
<emphasis id="37BD0DDD780D5E0AB90AAD31A658D130" italics="true" pageId="0" pageNumber="375">Female genitalia</emphasis>
.
</emphasis>
Gonocoxite 2 elongate. Spermatheca narrow and long, coiled in distal 1/2, undifferentiated; spermathecal gland connected near medial 1/3 of spermatheca.
<emphasis id="57A05CB54BDD1A74AAFEE3A80AB377BB" bold="true" pageId="0" pageNumber="375">
<emphasis id="AC554D85B72F4E86111C883E4A3D7615" italics="true" pageId="0" pageNumber="375">Male genitalia</emphasis>
.
</emphasis>
Median lobe of aedeagus in lateral view long, almost straight, with thin apex slightly flexed ventrally on the left lateral edge (Fig.
<figureCitation id="2936DD00BFC0EEB60A2F39364BA7FFD7" captionStart="Figures 32–40" captionStartId="F8" captionText="Figures 32 - 40. Aedeagus, left lateral view of 32 Coptocarpus microps sp. nov. 33 C. erwini sp. nov. 34 C. cyllodinus (Fauvel) 35 C. amieuensis sp. nov. 36 C. magnus sp. nov. 37 Adelopomorpha tethys sp. nov. 38 A. tuberculata sp. nov. 39 A. glabra Heller 40 right lateral view of tip with arrow indicating the dorsal tooth of C. magnus." figureDoi="10.3897/zookeys.1044.63775.figures32-40" httpUri="https://binary.pensoft.net/fig/554055" pageId="0" pageNumber="375">33</figureCitation>
); lobe in dorsal view with apical lamella well bent to right, with right side deeply concave and left side straight (Fig.
<figureCitation id="9053CBC682933C7183F1FE7332285A31" captionStart="Figures 22–31" captionStartId="F7" captionText="Figures 22 - 31. Aedeagus, dorsal view of 22 Coptocarpus microps sp. nov. 23 C. erwini sp. nov. 24 C. cyllodinus (Fauvel) 25 C. amieuensis sp. nov. 26 C. magnus sp. nov. 27 Adelopomorpha tethys sp. nov. 28 A. tuberculata sp. nov. 29 A. glabra Heller 30 Adelopomorpha glabra group male from Mt. Koghis 31 Adelopomorpha glabra group male from Gelima." figureDoi="10.3897/zookeys.1044.63775.figures22-31" httpUri="https://binary.pensoft.net/fig/554054" pageId="0" pageNumber="375">23</figureCitation>
); basal bulb forming edge dorsally with evident crest; sclerotized portion of endophallus visible in repose with one evident sclerite.
</paragraph>
</subSubSection>
<subSubSection id="BAA8CD81391C9FE3BD6145D38481692A" pageId="0" pageNumber="375" type="etymology">
<paragraph id="989FED23BEA384B57184C54C487E2585" pageId="0" pageNumber="375">Etymology.</paragraph>
<paragraph id="13A5CCDAE16992E95D627913459A7D15" pageId="0" pageNumber="375">
The specific epithet
<taxonomicName id="EA8C2B3CB6D0DA08A8E0D734575D8905" authorityName="Will &amp; Guéorguiev" authorityYear="2021" class="Insecta" family="Carabidae" genus="Coptocarpus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coptocarpus erwini" order="Coleoptera" pageId="0" pageNumber="375" phylum="Arthropoda" rank="species" species="erwini">
<emphasis id="8FC2E195EB6DD84A8E76C12B85C2CE8B" italics="true" pageId="0" pageNumber="375">Coptocarpus erwini</emphasis>
</taxonomicName>
is treated as a noun in the genitive case and is in honor of Terry Erwin and his amazing, life-long contribution to carabidology.
</paragraph>
<caption id="EF5227FD77AB74868D6E79808DAC9C99" doi="10.3897/zookeys.1044.63775.figures41-42" httpUri="https://binary.pensoft.net/fig/554056" pageId="0" pageNumber="375" start="Figures 41, 42" startId="F9">
<paragraph id="3F0740842A4332B61049EABD49D014D6" pageId="0" pageNumber="375">
<emphasis id="17231066EC4DD3A5511898C9BFD77E2E" bold="true" pageId="0" pageNumber="375">Figures 41, 42.</emphasis>
Dorsal habitus of
<emphasis id="DE119388ED58A85C76D796F0CEE09C15" bold="true" pageId="0" pageNumber="375">41</emphasis>
<taxonomicName id="719270764CB863C716E7208B0B688C20" authorityName="Will &amp; Guéorguiev" authorityYear="2021" class="Insecta" family="Carabidae" genus="Adelopomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Adelopomorpha tuberculata" order="Coleoptera" pageId="0" pageNumber="375" phylum="Arthropoda" rank="species" species="tuberculata">
<emphasis id="6CBD5597502E81EB0AFC7577695EAA40" italics="true" pageId="0" pageNumber="375">Adelopomorpha tuberculata</emphasis>
</taxonomicName>
<emphasis id="A85A50B6A9C95EDB776BBF4B8311C0D0" bold="true" pageId="0" pageNumber="375">42</emphasis>
holotype of
<taxonomicName id="578DEDDB9C7D50BB3E58966152CBF339" lsidName="A. glabra" pageId="0" pageNumber="375" rank="species" species="glabra">
<emphasis id="1BFAFC4B37B35DE534AB8E3F339A404C" italics="true" pageId="0" pageNumber="375">A. glabra</emphasis>
</taxonomicName>
Heller, with labels. Images of
<taxonomicName id="46380EEAB807C847FC614857A1A3796F" lsidName="A. glabra" pageId="0" pageNumber="375" rank="species" species="glabra">
<emphasis id="E5D1936D0E5810944A46939D2EC20CCF" italics="true" pageId="0" pageNumber="375">A. glabra</emphasis>
</taxonomicName>
by O.
<normalizedToken id="E24AAB4764979D5048A656D9A8A0E2F2" originalValue="Jäger">Jaeger</normalizedToken>
, reproduced with permission from MTKD.
</paragraph>
</caption>
<caption id="BE5088110CD35DBF23881AA7D64576F9" doi="10.3897/zookeys.1044.63775.figures43-44" httpUri="https://binary.pensoft.net/fig/554057" pageId="0" pageNumber="375" start="Figures 43, 44" startId="F10">
<paragraph id="C67408772EB2940B591056B0525227E4" pageId="0" pageNumber="375">
<emphasis id="80942B6D6CB52A9D9AB1CC6BB574F735" bold="true" pageId="0" pageNumber="375">Figures 43, 44.</emphasis>
Forebody, left dorsolateral view of
<emphasis id="2E99120B0EC4F58311D76A86CA4100B7" bold="true" pageId="0" pageNumber="375">43</emphasis>
<taxonomicName id="F5E811759E7E3CF81B73B32440B8A219" authorityName="Heller" authorityYear="1916" class="Insecta" family="Carabidae" genus="Adelopomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Adelopomorpha glabra" order="Coleoptera" pageId="0" pageNumber="375" phylum="Arthropoda" rank="species" species="glabra">
<emphasis id="51DFA804C4759AEFA2CC1D89291BAEDF" italics="true" pageId="0" pageNumber="375">Adelopomorpha glabra</emphasis>
</taxonomicName>
Heller
<emphasis id="BE285D091F45DF3D454E1E3D9CC1B7B0" bold="true" pageId="0" pageNumber="375">44</emphasis>
<taxonomicName id="0DB72D2BBB7029AC4F624E6F100C46AF" authorityName="Heller" authorityYear="1916" class="Insecta" family="Carabidae" genus="Adelopomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Adelopomorpha glabra" order="Coleoptera" pageId="0" pageNumber="375" phylum="Arthropoda" rank="species" species="glabra">
<emphasis id="BDFB11F973D800DC14ACCF66D65EBB1B" italics="true" pageId="0" pageNumber="375">Adelopomorpha glabra</emphasis>
</taxonomicName>
group male from Gelima.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>
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