treatments-xml/data/AF/30/BA/AF30BA35FF9FFF9813F6FA65DA124FCC.xml

<document id="F9378468C2F2D515E25E2EC0C1EF4BDE" ID-CLB-Dataset="25396" ID-DOI="10.11646/zootaxa.4674.3.2" ID-GBIF-Dataset="c8e7a489-a454-48da-86c1-5f6a20f98470" ID-ISSN="1175-5326" ID-Zenodo-Dep="3460178" ID-ZooBank="6E3DF207-741B-4589-8DB3-4A3BFD16B11C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1569395014923" checkinUser="plazi" docAuthor="Johnson, Jeffrey W. &amp; Wilmer, Jessica Worthington" docDate="2019" docId="AF30BA35FF9FFF9813F6FA65DA124FCC" docLanguage="en" docName="zootaxa.4674.3.2.pdf" docOrigin="Zootaxa 4674 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Epinephelus fuscomarginatus Johnson &amp; Wilmer 2019, sp. nov." docType="treatment" docVersion="6" lastPageNumber="346" masterDocId="5309C24DFF9CFF891361FF94D8074D03" masterDocTitle="Epinephelus fuscomarginatus (Perciformes: Epinephelidae), a new species of grouper from off the Great Barrier Reef, Australia" masterLastPageNumber="348" masterPageNumber="329" pageNumber="332" updateTime="1698756413827" updateUser="ExternalLinkService">
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<mods:title id="5F6736B559681C042AD87DC8703A99E5">Epinephelus fuscomarginatus (Perciformes: Epinephelidae), a new species of grouper from off the Great Barrier Reef, Australia</mods:title>
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<mods:namePart id="DA57FC3AF9C21CDF4BCC0AB2A433BF57">Johnson, Jeffrey W.</mods:namePart>
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<mods:namePart id="6BF57816F5209CFB0991A3DA9DCC99E9">Wilmer, Jessica Worthington</mods:namePart>
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<mods:title id="B644CBA0870290F4D6F31083A9A816CA">Zootaxa</mods:title>
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<mods:date id="8CDBA06004301845F3990413B38F69D4">2019</mods:date>
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<mods:number id="B47AA6F619AACD96653D7CB30115A253">2019-09-25</mods:number>
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<taxonomicName id="E09970A0FF9FFF8A13F6FA65DA054B08" authority="Johnson &amp; Wilmer, 2019" authorityName="Johnson &amp; Wilmer" authorityYear="2019" box="[151,514,1521,1548]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="332" phylum="Chordata" rank="species" species="fuscomarginatus" status="sp. nov.">
<emphasis id="15EDD731FF9FFF8A13F6FA65DA054B08" bold="true" box="[151,514,1521,1548]" italics="true" pageId="3" pageNumber="332">Epinephelus fuscomarginatus</emphasis>
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<taxonomicNameLabel id="0EDE6A4AFF9FFF8A1168FA65DA604B08" box="[521,615,1521,1547]" pageId="3" pageNumber="332" rank="species">sp. nov.</taxonomicNameLabel>
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<paragraph id="27260B23FF9FFF8A13F6F9ADDA614B50" blockId="3.[151,614,1593,1655]" box="[151,614,1593,1619]" pageId="3" pageNumber="332">New English name: Darkmargin Grouper</paragraph>
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<subSubSection id="6F8358A8FF9FFF8A13F6F9C9DA644B74" box="[151,611,1629,1655]" pageId="3" pageNumber="332" type="description">
<paragraph id="27260B23FF9FFF8A13F6F9C9DA644B74" blockId="3.[151,614,1593,1655]" box="[151,611,1629,1655]" pageId="3" pageNumber="332">
<figureCitation id="BFA217A6FF9FFF8A13F6F9C9D9074B74" box="[151,256,1629,1655]" captionStart="FIGURE 1" captionStartId="4.[151,250,1916,1941]" captionTargetBox="[183,1398,315,1886]" captionTargetId="figure@4.[177,1411,309,1892]" captionTargetPageId="4" captionText="FIGURE 1. Maximum clade credibility Bayesian tree of species of Epinephelus and outgroup taxa. Clade support is indicated by posterior probabilities ≥ 0.95 above the branch and bootstrap values ≥ 70 representing a topologically identical best maxi- mum likelihood tree are below the branch. Scale represents average number of substitutions per site." figureDoi="http://doi.org/10.5281/zenodo.3460180" httpUri="https://zenodo.org/record/3460180/files/figure.png" pageId="3" pageNumber="332">Figures 1</figureCitation>
, 
<figureCitation id="BFA217A6FF9FFF8A126FF9C9D91B4B74" box="[270,284,1629,1655]" captionStart="FIGURE 2" captionStartId="5.[151,250,1697,1722]" captionTargetBox="[151,1436,181,1666]" captionTargetId="figure@5.[151,1436,181,1671]" captionTargetPageId="5" captionText="FIGURE 2. Type specimens of Epinephelus fuscomarginatus sp. nov. A. Holotype, QM I.40945 (GenBank MK693011), 564 mm SL, Capricorn Channel, south of Swain Reefs, Qld, Australia, preserved; B. Holotype, fresh; C. Paratype, CSIRO H.8398- 01 (GenBank MK693013), 456 mm SL, same data as holotype, fresh." figureDoi="http://doi.org/10.5281/zenodo.3460182" httpUri="https://zenodo.org/record/3460182/files/figure.png" pageId="3" pageNumber="332">2</figureCitation>
A–C, 3A–D, 4A; 
<tableCitation id="6A1B3E98FF9FFF8A1283F9C9DA584B74" box="[482,607,1629,1655]" captionStart-0="TABLE 1" captionStart-1="TABLE 2" captionStart-2="TABLE 3" captionStart-3="TABLE 4" captionStartId-0="8.[151,244,152,179]" captionStartId-1="12.[151,244,152,179]" captionStartId-2="14.[151,244,152,179]" captionStartId-3="17.[151,244,764,791]" captionTargetBox-0="[163,1359,194,1978]" captionTargetBox-1="[163,1406,245,1097]" captionTargetBox-2="[163,1423,266,2011]" captionTargetBox-3="[163,1398,878,1467]" captionTargetPageId-0="8" captionTargetPageId-1="12" captionTargetPageId-2="14" captionText-0="TABLE 1. Details of all sequences analysed in this study, including Genbank/BOLD accession numbers." captionText-1="TABLE 2. Estimates of mean sequence divergence within (on diagonal) and between (below diagonal) species." captionText-2="TABLE 3. Meristic and morphological values for type specimens of Epinephelus fuscomarginatus sp. nov. and comparative material examined of E. epistictus, E. cf epistictus and E. magniscuttis (measurements as percentage of standard length). * Data for largest specimen of E. magniscuttis is from Nakamura et al. (2018)." captionText-3="TABLE 4. Selected meristic data, morphometric ratios and morphological characters for Epinephelus fuscomarginatus sp. nov., E. epistictus, E. cf epistictus and E. magniscuttis from material examined. * Data for largest specimen of E. magniscuttis based on Nakamura et al. (2018)." httpUri-0="http://table.plazi.org/id/73E65BABFF94FF8113F6FF0CDD374DB0" httpUri-1="http://table.plazi.org/id/73E65BABFF90FF8513F6FF0CDD7E4DB0" httpUri-2="http://table.plazi.org/id/73E65BABFF92FF8713F6FF0CDCCD4DF8" httpUri-3="http://table.plazi.org/id/73E65BABFF8DFF9813F6FD68DAE74E5C" pageId="3" pageNumber="332" tableUuid-0="73E65BABFF94FF8113F6FF0CDD374DB0" tableUuid-1="73E65BABFF90FF8513F6FF0CDD7E4DB0" tableUuid-2="73E65BABFF92FF8713F6FF0CDCCD4DF8" tableUuid-3="73E65BABFF8DFF9813F6FD68DAE74E5C">Tables 1–4</tableCitation>
.
</paragraph>
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<paragraph id="27260B23FF9FFF8A13F6F931DA204BE0" blockId="3.[151,1437,1700,2015]" pageId="3" pageNumber="332">
<materialsCitation id="97F1017EFF9FFF8A13F6F931DA204BE0" ID-GBIF-Occurrence="2423596363" collectingDate="2017-08-07" collectionCode="QM" collectorName="A. Lancaster" county="Qld" elevation="230" latitude="-22.88" location="Capricorn Channel" longLatPrecision="125" longitude="152.68" municipality="Swain Reefs" pageId="3" pageNumber="332" specimenCount="1" typeStatus="holotype">
<emphasis id="15EDD731FF9FFF8A13F6F931D90C4BBC" bold="true" box="[151,267,1701,1727]" pageId="3" pageNumber="332">
<typeStatus id="F822B581FF9FFF8A13F6F931D9014BBC" box="[151,262,1701,1727]" pageId="3" pageNumber="332" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectionCode id="418893E6FF9FFF8A1275F931D9464BBC" box="[276,321,1701,1727]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="3" pageNumber="332" type="Museum">QM</collectionCode>
I. 
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, 
<location id="22465DF8FF9FFF8A117FF931DAF04BBC" LSID="urn:lsid:plazi:treatment:AF30BA35FF9FFF9813F6FA65DA124FCC:22465DF8FF9FFF8A117FF931DAF04BBC" box="[542,759,1701,1727]" county="Qld" latitude="-22.88" longLatPrecision="125" longitude="152.68" municipality="Swain Reefs" name="Capricorn Channel" pageId="3" pageNumber="332">Capricorn Channel</location>
, south of 
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, 
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, dropline, 
<collectorName id="8A6C6EF5FF9FFF8A1260F95DD9954BE0" box="[257,402,1737,1763]" pageId="3" pageNumber="332">A. Lancaster</collectorName>
, 
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<collectingDate id="4363D40BFF9FFF8A12FCF95DDA234BE0" box="[413,548,1737,1763]" pageId="3" pageNumber="332" value="2017-08-07">7 Aug 2017</collectingDate>
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.
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</paragraph>
<paragraph id="27260B23FF9FFF8A13A6F979DC0F4A4C" blockId="3.[151,1437,1700,2015]" pageId="3" pageNumber="332">
<materialsCitation id="97F1017EFF9FFF8A13A6F979DB034A28" ID-GBIF-Occurrence="2423596366" collectingDate="2017-12-11" collectionCode="CSIRO" collectorName="S. Campbell" county="Qld" elevation="220" latitude="-22.88" location="Capricorn Channel" longLatPrecision="125" longitude="152.68" municipality="Swain Reefs" pageId="3" pageNumber="332" specimenCount="1" typeStatus="paratype">
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<typeStatus id="F822B581FF9FFF8A13A6F979D9464A04" box="[199,321,1773,1799]" pageId="3" pageNumber="332" type="paratype">Paratypes</typeStatus>
.
</emphasis>
(n = 4) 
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H.8398-01, 
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, 
<location id="22465DF8FF9FFF8A1060F979DBDC4A04" LSID="urn:lsid:plazi:treatment:AF30BA35FF9FFF9813F6FA65DA124FCC:22465DF8FF9FFF8A1060F979DBDC4A04" box="[769,987,1773,1799]" county="Qld" latitude="-22.88" longLatPrecision="125" longitude="152.68" municipality="Swain Reefs" name="Capricorn Channel" pageId="3" pageNumber="332">Capricorn Channel</location>
, south of 
<collectingMunicipality id="C7429159FF9FFF8A1731F979DCE44A04" box="[1104,1251,1773,1799]" pageId="3" pageNumber="332">Swain Reefs</collectingMunicipality>
, 
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<quantity id="E061A6C6FF9FFF8A1245F885D96B4A28" box="[292,364,1809,1835]" metricMagnitude="2" metricUnit="m" metricValue="2.2" pageId="3" pageNumber="332" unit="m" value="220.0">
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, dropline, 
<collectorName id="8A6C6EF5FF9FFF8A1281F885DA6A4A28" box="[480,621,1809,1835]" pageId="3" pageNumber="332">S. Campbell</collectorName>
, 
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; 
<materialsCitation id="97F1017EFF9FFF8A106EF885DD044A28" ID-GBIF-Occurrence="2423596361" box="[783,1283,1809,1835]" collectingDate="2017-08-07" collectionCode="QM" collectorName="A. Lancaster" county="Qld" elevation="230" latitude="-22.88" location="Capricorn Channel" longLatPrecision="125" longitude="152.68" municipality="Swain Reefs" pageId="3" pageNumber="332" specimenCount="1" typeStatus="paratype">
<collectionCode id="418893E6FF9FFF8A106EF885DB3B4A28" box="[783,828,1809,1835]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="3" pageNumber="332" type="Museum">QM</collectionCode>
I. 
<quantity id="E061A6C6FF9FFF8A102FF885DC054A28" box="[846,1026,1809,1835]" metricMagnitude="4" metricUnit="m" metricValue="4.0942408" pageId="3" pageNumber="332" unit="mm" value="4.0942408E7">40942, 408 mm</quantity>
, same data as holotype
</materialsCitation>
; 
<materialsCitation id="97F1017EFF9FFF8A166DF885D9FB4A4C" ID-GBIF-Occurrence="2423596368" collectingDate="2017-08-07" collectionCode="QM" collectorName="A. Lancaster" county="Qld" elevation="230" latitude="-22.88" location="Capricorn Channel" longLatPrecision="125" longitude="152.68" municipality="Swain Reefs" pageId="3" pageNumber="332" specimenCount="1" typeStatus="paratype">
<collectionCode id="418893E6FF9FFF8A166DF885DD3E4A28" box="[1292,1337,1809,1835]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="3" pageNumber="332" type="Museum">QM</collectionCode>
I. 
<quantity id="E061A6C6FF9FFF8A162DF885D8F14A4C" metricMagnitude="4" metricUnit="m" metricValue="4.0943502" pageId="3" pageNumber="332" unit="mm" value="4.0943502E7">40943, 502 mm</quantity>
, same data as holotype
</materialsCitation>
; 
<materialsCitation id="97F1017EFF9FFF8A1166F8A1DC024A4C" ID-GBIF-Occurrence="2423596364" box="[519,1029,1844,1871]" collectingDate="2017-08-07" collectionCode="QM" collectorName="A. Lancaster" county="Qld" elevation="230" latitude="-22.88" location="Capricorn Channel" longLatPrecision="125" longitude="152.68" municipality="Swain Reefs" pageId="3" pageNumber="332" specimenCount="1" typeStatus="paratype">
<collectionCode id="418893E6FF9FFF8A1166F8A1DA334A4C" box="[519,564,1845,1871]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="3" pageNumber="332" type="Museum">QM</collectionCode>
I. 
<quantity id="E061A6C6FF9FFF8A1129F8A1DAF94A4C" box="[584,766,1844,1871]" metricMagnitude="4" metricUnit="m" metricValue="4.0944528" pageId="3" pageNumber="332" unit="mm" value="4.0944528E7">40944, 528 mm</quantity>
, same data as holotype
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.
</paragraph>
</subSubSection>
<subSubSection id="6F8358A8FF9FFF8D13A6F8CCDAB24C1C" lastPageId="4" lastPageNumber="333" pageId="3" pageNumber="332" type="diagnosis">
<paragraph id="27260B23FF9FFF8D13A6F8CCDAB24C1C" blockId="3.[151,1437,1700,2015]" lastBlockId="4.[151,1437,153,287]" lastPageId="4" lastPageNumber="333" pageId="3" pageNumber="332">
<emphasis id="15EDD731FF9FFF8A13A6F8CCD9454A70" bold="true" box="[199,322,1880,1907]" pageId="3" pageNumber="332">Diagnosis.</emphasis>
A species of 
<taxonomicName id="E09970A0FF9FFF8A12B8F8CDDA644A70" authorityName="Bloch" authorityYear="1793" box="[473,611,1881,1907]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="332" phylum="Chordata" rank="genus">
<emphasis id="15EDD731FF9FFF8A12B8F8CDDA644A70" box="[473,611,1881,1907]" italics="true" pageId="3" pageNumber="332">Epinephelus</emphasis>
</taxonomicName>
with dorsal-fin rays XI, 14; anal-fin rays III, 8; pectoral-fin rays 17; caudal-fin rounded; lateral-line scales 60–67; lateral-scale series 101–116; gill rakers 9–10 + 16–19 = 25–28; midlateral part of lower jaw with 2 rows of teeth; vomer v-shaped, each arm with 2–3 rows of villiform teeth; palatines narrow, with 2–4 rows of villiform teeth; angle of preopercle broadly rounded, with 4–9 small non-prominent serrae; body depth 3.0– 
<quantity id="E061A6C6FF98FF8D1271FF0DD9564DB0" box="[272,337,153,179]" metricMagnitude="-2" metricUnit="m" metricValue="8.636" pageId="4" pageNumber="333" unit="in" value="3.4">3.4 in</quantity>
SL; fourth dorsal-fin spine longest, 3.3–4.0 in HL; and colouration including broad dark brown margins to the soft dorsal, anal and caudal fins, no dark spots on the head, body, or fins at any known size, and in subadults two faint pale brown bars radiating from the eye to the edge of the opercle, and diffuse irregular brown wavy bars and blotches on the sides of the body.
</paragraph>
</subSubSection>
<caption id="73E65BABFF98FF8D13F6F8E8DC964ADE" ID-DOI="http://doi.org/10.5281/zenodo.3460180" ID-Zenodo-Dep="3460180" httpUri="https://zenodo.org/record/3460180/files/figure.png" pageId="4" pageNumber="333" startId="4.[151,250,1916,1941]" targetBox="[183,1398,315,1886]" targetPageId="4">
<paragraph id="27260B23FF98FF8D13F6F8E8DC964ADE" blockId="4.[151,1436,1916,2014]" pageId="4" pageNumber="333">
<emphasis id="15EDD731FF98FF8D13F6F8E8D9144A96" bold="true" box="[151,275,1916,1941]" pageId="4" pageNumber="333">FIGURE 1.</emphasis>
Maximum clade credibility Bayesian tree of species of 
<taxonomicName id="E09970A0FF98FF8D102DF8E9DBCE4A96" authorityName="Bloch" authorityYear="1793" box="[844,969,1917,1941]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="333" phylum="Chordata" rank="genus">
<emphasis id="15EDD731FF98FF8D102DF8E9DBCE4A96" box="[844,969,1917,1941]" italics="true" pageId="4" pageNumber="333">Epinephelus</emphasis>
</taxonomicName>
and outgroup taxa. Clade support is indicated by posterior probabilities ≥ 0.95 above the branch and bootstrap values ≥ 70 representing a topologically identical best maxi- mum likelihood tree are below the branch. Scale represents average number of substitutions per site.
</paragraph>
</caption>
<caption id="73E65BABFF99FF8C13F6F935DB5E4A01" ID-DOI="http://doi.org/10.5281/zenodo.3460182" ID-Zenodo-Dep="3460182" httpUri="https://zenodo.org/record/3460182/files/figure.png" pageId="5" pageNumber="334" startId="5.[151,250,1697,1722]" targetBox="[151,1436,181,1666]" targetPageId="5">
<paragraph id="27260B23FF99FF8C13F6F935DB5E4A01" blockId="5.[151,1436,1697,1794]" pageId="5" pageNumber="334">
<emphasis id="15EDD731FF99FF8C13F6F935D9134BB9" bold="true" box="[151,276,1697,1722]" pageId="5" pageNumber="334">FIGURE 2.</emphasis>
Type specimens of 
<taxonomicName id="E09970A0FF99FF8C1283F935DB084BBA" authority="Johnson &amp; Wilmer, 2019" authorityName="Johnson &amp; Wilmer" authorityYear="2019" box="[482,783,1697,1721]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="334" phylum="Chordata" rank="species" species="fuscomarginatus" status="sp. nov.">
<emphasis id="15EDD731FF99FF8C1283F935DB084BBA" box="[482,783,1697,1721]" italics="true" pageId="5" pageNumber="334">Epinephelus fuscomarginatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0EDE6A4AFF99FF8C1077F936DB614BBA" box="[790,870,1698,1721]" pageId="5" pageNumber="334" rank="species">sp. nov.</taxonomicNameLabel>
A. Holotype, QM I.40945 (GenBank 
<accessionNumber id="38CA96C0FF99FF8C178DF935DD634BB9" box="[1260,1380,1697,1722]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MK693011" pageId="5" pageNumber="334">MK693011</accessionNumber>
), 564 mm SL, Capricorn Channel, south of Swain Reefs, Qld, Australia, preserved; B. Holotype, fresh; C. Paratype, CSIRO H.8398- 01 (GenBank 
<accessionNumber id="38CA96C0FF99FF8C1244F97DD9994A01" box="[293,414,1769,1794]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MK693013" pageId="5" pageNumber="334">MK693013</accessionNumber>
), 456 mm SL, same data as holotype, fresh.
</paragraph>
</caption>
<subSubSection id="6F8358A8FF99FF8213A6F8BBD9844ADC" lastPageId="11" lastPageNumber="340" pageId="5" pageNumber="334" type="description">
<paragraph id="27260B23FF99FF8C13A6F8BBDA834AB5" blockId="5.[151,1437,1839,1974]" pageId="5" pageNumber="334">
<emphasis id="15EDD731FF99FF8C13A6F8BBD95E4A49" bold="true" box="[199,345,1839,1866]" pageId="5" pageNumber="334">Description.</emphasis>
Morphometrics and meristics are presented in 
<tableCitation id="6A1B3E98FF99FF8C1001F8A4DBDC4A49" box="[864,987,1840,1866]" captionStart-0="TABLE 3" captionStart-1="TABLE 4" captionStartId-0="14.[151,244,152,179]" captionStartId-1="17.[151,244,764,791]" captionTargetBox-0="[163,1423,266,2011]" captionTargetBox-1="[163,1398,878,1467]" captionTargetPageId-0="14" captionText-0="TABLE 3. Meristic and morphological values for type specimens of Epinephelus fuscomarginatus sp. nov. and comparative material examined of E. epistictus, E. cf epistictus and E. magniscuttis (measurements as percentage of standard length). * Data for largest specimen of E. magniscuttis is from Nakamura et al. (2018)." captionText-1="TABLE 4. Selected meristic data, morphometric ratios and morphological characters for Epinephelus fuscomarginatus sp. nov., E. epistictus, E. cf epistictus and E. magniscuttis from material examined. * Data for largest specimen of E. magniscuttis based on Nakamura et al. (2018)." httpUri-0="http://table.plazi.org/id/73E65BABFF92FF8713F6FF0CDCCD4DF8" httpUri-1="http://table.plazi.org/id/73E65BABFF8DFF9813F6FD68DAE74E5C" pageId="5" pageNumber="334" tableUuid-0="73E65BABFF92FF8713F6FF0CDCCD4DF8" tableUuid-1="73E65BABFF8DFF9813F6FD68DAE74E5C">Tables 3–4</tableCitation>
. Dorsal-fin rays XI, 14; anal-fin rays III, 8; pectoral-fin rays 17, upper ray unbranched, others including lowermost branched; pelvic-fin rays I, 5; principal caudal-fin rays 17; lateral-line scales 65 (60–67); lateral-scale series 102 (101–116); gill rakers on first arch 9 + 17 = 26 (9–10 + 16–19 = 25–28); vertebrae 24.
</paragraph>
<caption id="73E65BABFF9AFF8F13F6F8E2D9B74AF8" ID-DOI="http://doi.org/10.5281/zenodo.3460184" ID-Zenodo-Dep="3460184" httpUri="https://zenodo.org/record/3460184/files/figure.png" pageId="6" pageNumber="335" startId="6.[151,250,1910,1935]" targetBox="[445,1136,181,1885]" targetPageId="6">
<paragraph id="27260B23FF9AFF8F13F6F8E2D9B74AF8" blockId="6.[151,1437,1910,2043]" pageId="6" pageNumber="335">
<emphasis id="15EDD731FF9AFF8F13F6F8E2D9144A8C" bold="true" box="[151,275,1910,1935]" pageId="6" pageNumber="335">FIGURE 3.</emphasis>
Fin colouration in type specimens of 
<taxonomicName id="E09970A0FF9AFF8F11F2F8E3DBB94A8C" authority="Johnson &amp; Wilmer, 2019" authorityName="Johnson &amp; Wilmer" authorityYear="2019" box="[659,958,1911,1935]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="335" phylum="Chordata" rank="species" species="fuscomarginatus" status="sp. nov.">
<emphasis id="15EDD731FF9AFF8F11F2F8E3DBB94A8C" box="[659,958,1911,1935]" italics="true" pageId="6" pageNumber="335">Epinephelus fuscomarginatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0EDE6A4AFF9AFF8F10A4F8E3DC134A8D" box="[965,1044,1911,1934]" pageId="6" pageNumber="335" rank="species">sp. nov.</taxonomicNameLabel>
A. Spinous and soft dorsal fin in holotype, QM I.40945 (GenBank 
<accessionNumber id="38CA96C0FF9AFF8F12A9F80EDA384AB0" box="[456,575,1946,1971]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MK693011" pageId="6" pageNumber="335">MK693011</accessionNumber>
), 564 mm SL, fresh; B. Spinous and soft dorsal fin in paratype, QM I.40942 (Gen- Bank 
<accessionNumber id="38CA96C0FF9AFF8F13B2F82AD94B4AD4" box="[211,332,1982,2007]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MK693009" pageId="6" pageNumber="335">MK693009</accessionNumber>
), 408 mm SL, fresh; C. Anal fin in holotype, QM I.40945, 564 mm SL, fresh; D. Caudal fin in holotype, QM I.40945, 564 mm SL, fresh.
</paragraph>
</caption>
<caption id="73E65BABFF9BFF8E13F6F8C0DAC34AFE" ID-DOI="http://doi.org/10.5281/zenodo.3460186" ID-Zenodo-Dep="3460186" httpUri="https://zenodo.org/record/3460186/files/figure.png" pageId="7" pageNumber="336" startId="7.[151,250,1876,1901]" targetBox="[223,1373,181,1840]" targetPageId="7">
<paragraph id="27260B23FF9BFF8E13F6F8C0DAC34AFE" blockId="7.[151,1437,1876,2045]" pageId="7" pageNumber="336">
<emphasis id="15EDD731FF9BFF8E13F6F8C0D9144A6E" bold="true" box="[151,275,1876,1901]" pageId="7" pageNumber="336">FIGURE 4.</emphasis>
Comparison of similar-sized subadults of 
<taxonomicName id="E09970A0FF9BFF8E11A5F8C1DBE84A6E" authority="Johnson &amp; Wilmer, 2019" authorityName="Johnson &amp; Wilmer" authorityYear="2019" box="[708,1007,1877,1901]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="336" phylum="Chordata" rank="species" species="fuscomarginatus" status="sp. nov.">
<emphasis id="15EDD731FF9BFF8E11A5F8C1DBE84A6E" box="[708,1007,1877,1901]" italics="true" pageId="7" pageNumber="336">Epinephelus fuscomarginatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0EDE6A4AFF9BFF8E1097F8C1DC404A6F" box="[1014,1095,1877,1900]" pageId="7" pageNumber="336" rank="species">sp. nov.</taxonomicNameLabel>
, 
<taxonomicName id="E09970A0FF9BFF8E1730F8C2DCC94A6E" authorityName="in Heemstra &amp; Randall" authorityYear="1993" box="[1105,1230,1878,1901]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="336" phylum="Chordata" rank="species" species="epistictus">
<emphasis id="15EDD731FF9BFF8E1730F8C2DCC94A6E" box="[1105,1230,1878,1901]" italics="true" pageId="7" pageNumber="336">E. epistictus</emphasis>
</taxonomicName>
, 
<emphasis id="15EDD731FF9BFF8E17BBF8C2DCEE4A6E" box="[1242,1257,1878,1901]" italics="true" pageId="7" pageNumber="336">E</emphasis>
. cf 
<taxonomicName id="E09970A0FF9BFF8E1670F8C2DD764A6E" authorityName="in Heemstra &amp; Randall" authorityYear="1993" box="[1297,1393,1878,1901]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="336" phylum="Chordata" rank="species" species="epistictus">
<emphasis id="15EDD731FF9BFF8E1670F8C2DD764A6E" box="[1297,1393,1878,1901]" italics="true" pageId="7" pageNumber="336">epistictus</emphasis>
</taxonomicName>
and 
<taxonomicName id="E09970A0FF9BFF8E13F6F8EED9324A92" box="[151,309,1914,1937]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="336" phylum="Chordata" rank="species" species="magniscuttis">
<emphasis id="15EDD731FF9BFF8E13F6F8EED9324A92" box="[151,309,1914,1937]" italics="true" pageId="7" pageNumber="336">E. magniscuttis</emphasis>
</taxonomicName>
. A. 
<taxonomicName id="E09970A0FF9BFF8E1200F8EEDA204A92" authority="Johnson &amp; Wilmer, 2019" authorityName="Johnson &amp; Wilmer" authorityYear="2019" box="[353,551,1913,1937]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="336" phylum="Chordata" rank="species" species="fuscomarginatus" status="sp. nov.">
<emphasis id="15EDD731FF9BFF8E1200F8EED9774A92" box="[353,368,1914,1937]" italics="true" pageId="7" pageNumber="336">E</emphasis>
. 
<emphasis id="15EDD731FF9BFF8E121EF8EDDA204A92" box="[383,551,1913,1937]" italics="true" pageId="7" pageNumber="336">fuscomarginatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0EDE6A4AFF9BFF8E114FF8EDDA874A93" box="[558,640,1913,1936]" pageId="7" pageNumber="336" rank="species">sp. nov.</taxonomicNameLabel>
, QM I.40942 (GenBank 
<accessionNumber id="38CA96C0FF9BFF8E10E0F8ECDBFD4A92" box="[897,1018,1912,1937]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MK693009" pageId="7" pageNumber="336">MK693009</accessionNumber>
), 408 mm, paratype, fresh; B. 
<taxonomicName id="E09970A0FF9BFF8E1655F8EED8B24AB6" authorityName="in Heemstra &amp; Randall" authorityYear="1993" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="336" phylum="Chordata" rank="species" species="epistictus">
<emphasis id="15EDD731FF9BFF8E1655F8EEDD444A92" box="[1332,1347,1914,1937]" italics="true" pageId="7" pageNumber="336">E</emphasis>
. 
<emphasis id="15EDD731FF9BFF8E1633F8EED8B24AB6" italics="true" pageId="7" pageNumber="336">epistictus</emphasis>
</taxonomicName>
, KAUM I.97319, 376 mm SL, off Mi-shima Island, Hagi, Yamaguchi, Sea of Japan, preserved; C. 
<emphasis id="15EDD731FF9BFF8E17CCF80ADCBB4AB6" box="[1197,1212,1950,1973]" italics="true" pageId="7" pageNumber="336">E</emphasis>
. cf 
<taxonomicName id="E09970A0FF9BFF8E1784F80ADD424AB6" authorityName="in Heemstra &amp; Randall" authorityYear="1993" box="[1253,1349,1950,1973]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="336" phylum="Chordata" rank="species" species="epistictus">
<emphasis id="15EDD731FF9BFF8E1784F80ADD424AB6" box="[1253,1349,1950,1973]" italics="true" pageId="7" pageNumber="336">epistictus</emphasis>
</taxonomicName>
, CSIRO CA.2266, 406 mm SL, Arafura Sea, NT, Australia, fresh (Photo: CSIRO); D. 
<taxonomicName id="E09970A0FF9BFF8E10C2F856DC384ADA" box="[931,1087,1986,2009]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="336" phylum="Chordata" rank="species" species="magniscuttis">
<emphasis id="15EDD731FF9BFF8E10C2F856DC384ADA" box="[931,1087,1986,2009]" italics="true" pageId="7" pageNumber="336">E. magniscuttis</emphasis>
</taxonomicName>
, WAM P.30510–007, 419 mm SL, off Ribbon Reefs, Qld, Australia, fresh (Photo: WAM).
</paragraph>
</caption>
<caption id="73E65BABFF94FF8113F6FF0CDD374DB0" ID-Table-UUID="73E65BABFF94FF8113F6FF0CDD374DB0" box="[151,1328,152,179]" httpUri="http://table.plazi.org/id/73E65BABFF94FF8113F6FF0CDD374DB0" pageId="8" pageNumber="337" startId="8.[151,244,152,179]" targetBox="[163,1359,194,1978]" targetIsTable="true" targetPageId="8">
<paragraph id="27260B23FF94FF8113F6FF0CDD374DB0" blockId="8.[151,1328,152,179]" box="[151,1328,152,179]" pageId="8" pageNumber="337">
<emphasis id="15EDD731FF94FF8113F6FF0CD9164DB0" bold="true" box="[151,273,152,179]" pageId="8" pageNumber="337">TABLE 1.</emphasis>
Details of all sequences analysed in this study, including Genbank/BOLD accession numbers.
</paragraph>
</caption>
<paragraph id="27260B23FF94FF8113C2FF56DBF54AB9" pageId="8" pageNumber="337">
<table id="5599F983FF94007613C2FF56DD484AB9" box="[163,1359,194,1978]" colsContinueIn="9.[163,1383,206,1980]" gridcols="3" gridrows="31" pageId="8" pageNumber="337">
<tr id="99A90961FF94007613C2FF56DD484DD8" box="[163,1359,194,219]" gridrow="0" pageId="8" pageNumber="337">
<th id="DA78601DFF94007613C2FF56DA444DD8" box="[163,579,194,219]" gridcol="0" gridrow="0" pageId="8" pageNumber="337">Species</th>
<th id="DA78601DFF9400761113FF56DB814DD8" box="[626,902,194,219]" gridcol="1" gridrow="0" pageId="8" pageNumber="337">Genbank/BOLD</th>
<th id="DA78601DFF94007610A7FF56DD484DD8" box="[966,1359,194,219]" gridcol="2" gridrow="0" pageId="8" pageNumber="337">Country</th>
</tr>
<tr id="99A90961FF94007613C2FF79DD484C05" box="[163,1359,237,262]" gridrow="1" pageId="8" pageNumber="337">
<th id="DA78601DFF94007613C2FF79DA444C05" box="[163,579,237,262]" gridcol="0" gridrow="1" pageId="8" pageNumber="337">
<emphasis id="15EDD731FF94FF8113C2FF79D9924C06" box="[163,405,237,261]" italics="true" pageId="8" pageNumber="337">
<taxonomicName id="E09970A0FF94FF8113C2FF79D98C4C06" baseAuthorityName="Bleeker" baseAuthorityYear="1855" box="[163,395,237,261]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="337" phylum="Chordata" rank="species" species="bontoides">Epinephelus bontoides</taxonomicName>
*
</emphasis>
</th>
<td id="DA78601DFF9400761113FF79DB814C05" box="[626,902,237,262]" gridcol="1" gridrow="1" pageId="8" pageNumber="337">
<accessionNumber id="38CA96C0FF94FF811113FF79DADD4C05" box="[626,730,237,262]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/JN312975" pageId="8" pageNumber="337">JN312975</accessionNumber>
/FOAM409-10
</td>
<td id="DA78601DFF94007610A7FF79DD484C05" box="[966,1359,237,262]" gridcol="2" gridrow="1" pageId="8" pageNumber="337">Indonesia: West Java</td>
</tr>
<tr id="99A90961FF94007613C2FE83DD484C33" box="[163,1359,279,304]" gridrow="2" pageId="8" pageNumber="337" rowspan-1="1" rowspan-2="1">
<th id="DA78601DFF94007613C2FE83DA444C33" box="[163,579,279,304]" gridcol="0" gridrow="2" pageId="8" pageNumber="337">
*Species ID in Genbank: 
<taxonomicName id="E09970A0FF94FF8112C6FE8DDA444C33" box="[423,579,281,304]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="337" phylum="Chordata" rank="species" species="magniscuttis">
<emphasis id="15EDD731FF94FF8112C6FE8DDA444C33" box="[423,579,281,304]" italics="true" pageId="8" pageNumber="337">E. magniscuttis</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="99A90961FF94007613C2FEF9DD484C85" box="[163,1359,365,390]" gridrow="3" pageId="8" pageNumber="337">
<th id="DA78601DFF94007613C2FEF9DA444C85" box="[163,579,365,390]" gridcol="0" gridrow="3" pageId="8" pageNumber="337">
<taxonomicName id="E09970A0FF94FF8113C2FEF9D9274C86" authorityName="Bloch" authorityYear="1793" box="[163,288,365,389]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="337" phylum="Chordata" rank="genus">
<emphasis id="15EDD731FF94FF8113C2FEF9D9274C86" box="[163,288,365,389]" italics="true" pageId="8" pageNumber="337">Epinephelus</emphasis>
</taxonomicName>
cf 
<taxonomicName id="E09970A0FF94FF811221FEFAD9A74C86" authorityName="in Heemstra &amp; Randall" authorityYear="1993" box="[320,416,366,389]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="337" phylum="Chordata" rank="species" species="epistictus">
<emphasis id="15EDD731FF94FF811221FEFAD9A74C86" box="[320,416,366,389]" italics="true" pageId="8" pageNumber="337">epistictus</emphasis>
</taxonomicName>
</th>
<td id="DA78601DFF9400761113FEF9DB814C85" box="[626,902,365,390]" gridcol="1" gridrow="3" pageId="8" pageNumber="337">
<accessionNumber id="38CA96C0FF94FF811113FEF9DAE04C85" box="[626,743,365,390]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/HM422400" pageId="8" pageNumber="337">HM422400</accessionNumber>
/FOAMI760-09
</td>
<td id="DA78601DFF94007610A7FEF9DD484C85" box="[966,1359,365,390]" gridcol="2" gridrow="3" pageId="8" pageNumber="337">Indonesia: West Java</td>
</tr>
<tr id="99A90961FF94007613C2FE0EDD484CB0" box="[163,1359,410,435]" gridrow="4" pageId="8" pageNumber="337">
<th id="DA78601DFF94007613C2FE0EDA444CB0" box="[163,579,410,435]" gridcol="0" gridrow="4" pageId="8" pageNumber="337">
<taxonomicName id="E09970A0FF94FF8113C2FE0FD9274CB0" authorityName="Bloch" authorityYear="1793" box="[163,288,411,435]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="337" phylum="Chordata" rank="genus">
<emphasis id="15EDD731FF94FF8113C2FE0FD9274CB0" box="[163,288,411,435]" italics="true" pageId="8" pageNumber="337">Epinephelus</emphasis>
</taxonomicName>
cf 
<taxonomicName id="E09970A0FF94FF811221FE08D9A74CB0" authorityName="in Heemstra &amp; Randall" authorityYear="1993" box="[320,416,412,435]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="337" phylum="Chordata" rank="species" species="epistictus">
<emphasis id="15EDD731FF94FF811221FE08D9A74CB0" box="[320,416,412,435]" italics="true" pageId="8" pageNumber="337">epistictus</emphasis>
</taxonomicName>
</th>
<td id="DA78601DFF9400761113FE0EDB814CB0" box="[626,902,410,435]" gridcol="1" gridrow="4" pageId="8" pageNumber="337">
<accessionNumber id="38CA96C0FF94FF811113FE0FDADC4CB0" box="[626,731,411,435]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/JN312980" pageId="8" pageNumber="337">JN312980</accessionNumber>
/FOAM414-10
</td>
<td id="DA78601DFF94007610A7FE0EDD484CB0" box="[966,1359,410,435]" gridcol="2" gridrow="4" pageId="8" pageNumber="337">Indonesia: West Java</td>
</tr>
<tr id="99A90961FF94007613C2FE51DD484CDD" box="[163,1359,453,478]" gridrow="5" pageId="8" pageNumber="337">
<th id="DA78601DFF94007613C2FE51DA444CDD" box="[163,579,453,478]" gridcol="0" gridrow="5" pageId="8" pageNumber="337">
<taxonomicName id="E09970A0FF94FF8113C2FE52D9274CDD" authorityName="Bloch" authorityYear="1793" box="[163,288,454,478]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="337" phylum="Chordata" rank="genus">
<emphasis id="15EDD731FF94FF8113C2FE52D9274CDD" box="[163,288,454,478]" italics="true" pageId="8" pageNumber="337">Epinephelus</emphasis>
</taxonomicName>
cf 
<taxonomicName id="E09970A0FF94FF811221FE52D9A74CDE" authorityName="in Heemstra &amp; Randall" authorityYear="1993" box="[320,416,454,477]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="337" phylum="Chordata" rank="species" species="epistictus">
<emphasis id="15EDD731FF94FF811221FE52D9A74CDE" box="[320,416,454,477]" italics="true" pageId="8" pageNumber="337">epistictus</emphasis>
</taxonomicName>
</th>
<td id="DA78601DFF9400761113FE51DB814CDD" box="[626,902,453,478]" gridcol="1" gridrow="5" pageId="8" pageNumber="337">
<accessionNumber id="38CA96C0FF94FF811113FE51DADD4CDD" box="[626,730,453,478]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/JN312981" pageId="8" pageNumber="337">JN312981</accessionNumber>
/FOAM415-10
</td>
<td id="DA78601DFF94007610A7FE51DD484CDD" box="[966,1359,453,478]" gridcol="2" gridrow="5" pageId="8" pageNumber="337">Indonesia: West Java</td>
</tr>
<tr id="99A90961FF94007613C2FE64DD484F30" box="[163,1359,496,563]" gridrow="6" pageId="8" pageNumber="337">
<th id="DA78601DFF94007613C2FE64DA444F30" box="[163,579,496,563]" gridcol="0" gridrow="6" pageId="8" pageNumber="337">
<taxonomicName id="E09970A0FF94FF8113C2FE64D9274F0B" authorityName="Bloch" authorityYear="1793" box="[163,288,496,520]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="337" phylum="Chordata" rank="genus">
<emphasis id="15EDD731FF94FF8113C2FE64D9274F0B" box="[163,288,496,520]" italics="true" pageId="8" pageNumber="337">Epinephelus</emphasis>
</taxonomicName>
cf 
<emphasis id="15EDD731FF94FF811221FE65D9274F30" italics="true" pageId="8" pageNumber="337">
<taxonomicName id="E09970A0FF94FF811221FE65D9A74F0B" authorityName="in Heemstra &amp; Randall" authorityYear="1993" box="[320,416,497,520]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="337" phylum="Chordata" rank="species" species="epistictus">epistictus</taxonomicName>
<taxonomicName id="E09970A0FF94FF8113C2FD8FD9274F30" authorityName="Bloch" authorityYear="1793" box="[163,288,539,563]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="337" phylum="Chordata" rank="genus">Epinephelus</taxonomicName>
</emphasis>
cf 
<taxonomicName id="E09970A0FF94FF811221FD88D9A74F30" authorityName="in Heemstra &amp; Randall" authorityYear="1993" box="[320,416,540,563]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="337" phylum="Chordata" rank="species" species="epistictus">
<emphasis id="15EDD731FF94FF811221FD88D9A74F30" box="[320,416,540,563]" italics="true" pageId="8" pageNumber="337">epistictus</emphasis>
</taxonomicName>
</th>
<td id="DA78601DFF9400761113FE64DB814F30" box="[626,902,496,563]" gridcol="1" gridrow="6" pageId="8" pageNumber="337">
<accessionNumber id="38CA96C0FF94FF811113FE64DADC4F0A" box="[626,731,496,521]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/JN312982" pageId="8" pageNumber="337">JN312982</accessionNumber>
/FOAM416-10 
<accessionNumber id="38CA96C0FF94FF811113FD8FDADC4F30" box="[626,731,539,563]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/JN312984" pageId="8" pageNumber="337">JN312984</accessionNumber>
/FOAM418-10
</td>
<td id="DA78601DFF94007610A7FE64DD484F30" box="[966,1359,496,563]" gridcol="2" gridrow="6" pageId="8" pageNumber="337">Indonesia: West Java Indonesia: West Java</td>
</tr>
<tr id="99A90961FF94007613C2FDD1DD484F5D" box="[163,1359,581,606]" gridrow="7" pageId="8" pageNumber="337">
<th id="DA78601DFF94007613C2FDD1DA444F5D" box="[163,579,581,606]" gridcol="0" gridrow="7" pageId="8" pageNumber="337">
<taxonomicName id="E09970A0FF94FF8113C2FDD2D9274F5D" authorityName="Bloch" authorityYear="1793" box="[163,288,582,606]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="337" phylum="Chordata" rank="genus">
<emphasis id="15EDD731FF94FF8113C2FDD2D9274F5D" box="[163,288,582,606]" italics="true" pageId="8" pageNumber="337">Epinephelus</emphasis>
</taxonomicName>
cf 
<taxonomicName id="E09970A0FF94FF811221FDD2D9A74F5E" authorityName="in Heemstra &amp; Randall" authorityYear="1993" box="[320,416,582,605]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="337" phylum="Chordata" rank="species" species="epistictus">
<emphasis id="15EDD731FF94FF811221FDD2D9A74F5E" box="[320,416,582,605]" italics="true" pageId="8" pageNumber="337">epistictus</emphasis>
</taxonomicName>
</th>
<td id="DA78601DFF9400761113FDD1DB814F5D" box="[626,902,581,606]" gridcol="1" gridrow="7" pageId="8" pageNumber="337">
<accessionNumber id="38CA96C0FF94FF811113FDD1DAE14F5D" box="[626,742,581,606]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KM226254" pageId="8" pageNumber="337">KM226254</accessionNumber>
(GenBank)
</td>
<td id="DA78601DFF94007610A7FDD1DD484F5D" box="[966,1359,581,606]" gridcol="2" gridrow="7" pageId="8" pageNumber="337">India</td>
</tr>
<tr id="99A90961FF94007613C2FDE4DD484FB0" box="[163,1359,624,691]" gridrow="8" pageId="8" pageNumber="337">
<th id="DA78601DFF94007613C2FDE4DA444FB0" box="[163,579,624,691]" gridcol="0" gridrow="8" pageId="8" pageNumber="337">
<taxonomicName id="E09970A0FF94FF8113C2FDE4D9274F8B" authorityName="Bloch" authorityYear="1793" box="[163,288,624,648]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="337" phylum="Chordata" rank="genus">
<emphasis id="15EDD731FF94FF8113C2FDE4D9274F8B" box="[163,288,624,648]" italics="true" pageId="8" pageNumber="337">Epinephelus</emphasis>
</taxonomicName>
cf 
<emphasis id="15EDD731FF94FF811221FDE5D9274FB0" italics="true" pageId="8" pageNumber="337">
<taxonomicName id="E09970A0FF94FF811221FDE5D9A74F8B" authorityName="in Heemstra &amp; Randall" authorityYear="1993" box="[320,416,625,648]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="337" phylum="Chordata" rank="species" species="epistictus">epistictus</taxonomicName>
<taxonomicName id="E09970A0FF94FF8113C2FD0FD9274FB0" authorityName="Bloch" authorityYear="1793" box="[163,288,667,691]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="337" phylum="Chordata" rank="genus">Epinephelus</taxonomicName>
</emphasis>
cf 
<taxonomicName id="E09970A0FF94FF811221FD08D9A74FB0" authorityName="in Heemstra &amp; Randall" authorityYear="1993" box="[320,416,668,691]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="337" phylum="Chordata" rank="species" species="epistictus">
<emphasis id="15EDD731FF94FF811221FD08D9A74FB0" box="[320,416,668,691]" italics="true" pageId="8" pageNumber="337">epistictus</emphasis>
</taxonomicName>
</th>
<td id="DA78601DFF9400761113FDE4DB814FB0" box="[626,902,624,691]" gridcol="1" gridrow="8" pageId="8" pageNumber="337">
<accessionNumber id="38CA96C0FF94FF811113FDE4DAEF4F8A" box="[626,744,624,649]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KM226255" pageId="8" pageNumber="337">KM226255</accessionNumber>
(GenBank) 
<accessionNumber id="38CA96C0FF94FF811113FD0EDAE14FB0" box="[626,742,666,691]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KM226256" pageId="8" pageNumber="337">KM226256</accessionNumber>
(GenBank)
</td>
<td id="DA78601DFF94007610A7FDE4DD484FB0" box="[966,1359,624,691]" gridcol="2" gridrow="8" pageId="8" pageNumber="337">India India</td>
</tr>
<tr id="99A90961FF94007613C2FD51DD484FDD" box="[163,1359,709,734]" gridrow="9" pageId="8" pageNumber="337">
<th id="DA78601DFF94007613C2FD51DA444FDD" box="[163,579,709,734]" gridcol="0" gridrow="9" pageId="8" pageNumber="337">
<taxonomicName id="E09970A0FF94FF8113C2FD52D9274FDD" authorityName="Bloch" authorityYear="1793" box="[163,288,710,734]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="337" phylum="Chordata" rank="genus">
<emphasis id="15EDD731FF94FF8113C2FD52D9274FDD" box="[163,288,710,734]" italics="true" pageId="8" pageNumber="337">Epinephelus</emphasis>
</taxonomicName>
cf 
<taxonomicName id="E09970A0FF94FF811221FD52D9A74FDE" authorityName="in Heemstra &amp; Randall" authorityYear="1993" box="[320,416,710,733]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="337" phylum="Chordata" rank="species" species="epistictus">
<emphasis id="15EDD731FF94FF811221FD52D9A74FDE" box="[320,416,710,733]" italics="true" pageId="8" pageNumber="337">epistictus</emphasis>
</taxonomicName>
</th>
<td id="DA78601DFF9400761113FD51DB814FDD" box="[626,902,709,734]" gridcol="1" gridrow="9" pageId="8" pageNumber="337">
<accessionNumber id="38CA96C0FF94FF811113FD51DAE14FDD" box="[626,742,709,734]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KM226257" pageId="8" pageNumber="337">KM226257</accessionNumber>
(GenBank)
</td>
<td id="DA78601DFF94007610A7FD51DD484FDD" box="[966,1359,709,734]" gridcol="2" gridrow="9" pageId="8" pageNumber="337">India</td>
</tr>
<tr id="99A90961FF94007613C2FD64DD484E30" box="[163,1359,752,819]" gridrow="10" pageId="8" pageNumber="337">
<th id="DA78601DFF94007613C2FD64DA444E30" box="[163,579,752,819]" gridcol="0" gridrow="10" pageId="8" pageNumber="337">
<taxonomicName id="E09970A0FF94FF8113C2FD64D9274E0B" authorityName="Bloch" authorityYear="1793" box="[163,288,752,776]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="337" phylum="Chordata" rank="genus">
<emphasis id="15EDD731FF94FF8113C2FD64D9274E0B" box="[163,288,752,776]" italics="true" pageId="8" pageNumber="337">Epinephelus</emphasis>
</taxonomicName>
cf 
<emphasis id="15EDD731FF94FF811221FD65D9274E30" italics="true" pageId="8" pageNumber="337">
<taxonomicName id="E09970A0FF94FF811221FD65D9A74E0B" authorityName="in Heemstra &amp; Randall" authorityYear="1993" box="[320,416,753,776]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="337" phylum="Chordata" rank="species" species="epistictus">epistictus</taxonomicName>
<taxonomicName id="E09970A0FF94FF8113C2FC8FD9274E30" authorityName="Bloch" authorityYear="1793" box="[163,288,795,819]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="337" phylum="Chordata" rank="genus">Epinephelus</taxonomicName>
</emphasis>
cf 
<taxonomicName id="E09970A0FF94FF811221FC88D9A74E30" authorityName="in Heemstra &amp; Randall" authorityYear="1993" box="[320,416,796,819]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="337" phylum="Chordata" rank="species" species="epistictus">
<emphasis id="15EDD731FF94FF811221FC88D9A74E30" box="[320,416,796,819]" italics="true" pageId="8" pageNumber="337">epistictus</emphasis>
</taxonomicName>
</th>
<td id="DA78601DFF9400761113FD64DB814E30" box="[626,902,752,819]" gridcol="1" gridrow="10" pageId="8" pageNumber="337">
<accessionNumber id="38CA96C0FF94FF811113FD64DAE14E0A" box="[626,742,752,777]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KM226258" pageId="8" pageNumber="337">KM226258</accessionNumber>
(GenBank) 
<accessionNumber id="38CA96C0FF94FF811113FC8EDAE14E30" box="[626,742,794,819]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KM226259" pageId="8" pageNumber="337">KM226259</accessionNumber>
(GenBank)
</td>
<td id="DA78601DFF94007610A7FD64DD484E30" box="[966,1359,752,819]" gridcol="2" gridrow="10" pageId="8" pageNumber="337">India India</td>
</tr>
<tr id="99A90961FF94007613C2FCD1DD484E5D" box="[163,1359,837,862]" gridrow="11" pageId="8" pageNumber="337">
<th id="DA78601DFF94007613C2FCD1DA444E5D" box="[163,579,837,862]" gridcol="0" gridrow="11" pageId="8" pageNumber="337">
<taxonomicName id="E09970A0FF94FF8113C2FCD2D9274E5D" authorityName="Bloch" authorityYear="1793" box="[163,288,838,862]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="337" phylum="Chordata" rank="genus">
<emphasis id="15EDD731FF94FF8113C2FCD2D9274E5D" box="[163,288,838,862]" italics="true" pageId="8" pageNumber="337">Epinephelus</emphasis>
</taxonomicName>
cf 
<taxonomicName id="E09970A0FF94FF811221FCD2D9A74E5E" authorityName="in Heemstra &amp; Randall" authorityYear="1993" box="[320,416,838,861]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="337" phylum="Chordata" rank="species" species="epistictus">
<emphasis id="15EDD731FF94FF811221FCD2D9A74E5E" box="[320,416,838,861]" italics="true" pageId="8" pageNumber="337">epistictus</emphasis>
</taxonomicName>
</th>
<td id="DA78601DFF9400761113FCD1DB814E5D" box="[626,902,837,862]" gridcol="1" gridrow="11" pageId="8" pageNumber="337">
<accessionNumber id="38CA96C0FF94FF811113FCD1DAE14E5D" box="[626,742,837,862]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KM226260" pageId="8" pageNumber="337">KM226260</accessionNumber>
(GenBank)
</td>
<td id="DA78601DFF94007610A7FCD1DD484E5D" box="[966,1359,837,862]" gridcol="2" gridrow="11" pageId="8" pageNumber="337">India</td>
</tr>
<tr id="99A90961FF94007613C2FCE4DD484EB0" box="[163,1359,880,947]" gridrow="12" pageId="8" pageNumber="337">
<th id="DA78601DFF94007613C2FCE4DA444EB0" box="[163,579,880,947]" gridcol="0" gridrow="12" pageId="8" pageNumber="337">
<taxonomicName id="E09970A0FF94FF8113C2FCE4D9274E8B" authorityName="Bloch" authorityYear="1793" box="[163,288,880,904]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="337" phylum="Chordata" rank="genus">
<emphasis id="15EDD731FF94FF8113C2FCE4D9274E8B" box="[163,288,880,904]" italics="true" pageId="8" pageNumber="337">Epinephelus</emphasis>
</taxonomicName>
cf 
<emphasis id="15EDD731FF94FF811221FCE5D9274EB0" italics="true" pageId="8" pageNumber="337">
<taxonomicName id="E09970A0FF94FF811221FCE5D9A74E8B" authorityName="in Heemstra &amp; Randall" authorityYear="1993" box="[320,416,881,904]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="337" phylum="Chordata" rank="species" species="epistictus">epistictus</taxonomicName>
<taxonomicName id="E09970A0FF94FF8113C2FC0FD9274EB0" authorityName="Bloch" authorityYear="1793" box="[163,288,923,947]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="337" phylum="Chordata" rank="genus">Epinephelus</taxonomicName>
</emphasis>
cf 
<taxonomicName id="E09970A0FF94FF811221FC08D9A74EB0" authorityName="in Heemstra &amp; Randall" authorityYear="1993" box="[320,416,924,947]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="337" phylum="Chordata" rank="species" species="epistictus">
<emphasis id="15EDD731FF94FF811221FC08D9A74EB0" box="[320,416,924,947]" italics="true" pageId="8" pageNumber="337">epistictus</emphasis>
</taxonomicName>
</th>
<td id="DA78601DFF9400761113FCE4DB814EB0" box="[626,902,880,947]" gridcol="1" gridrow="12" pageId="8" pageNumber="337">
<accessionNumber id="38CA96C0FF94FF811113FCE4DAE54E8A" box="[626,738,880,905]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KU366471" pageId="8" pageNumber="337">KU366471</accessionNumber>
(GenBank) 
<accessionNumber id="38CA96C0FF94FF811113FC0EDAE14EB0" box="[626,742,922,947]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MK693007" pageId="8" pageNumber="337">MK693007</accessionNumber>
(GenBank)
</td>
<td id="DA78601DFF94007610A7FCE4DD484EB0" box="[966,1359,880,947]" gridcol="2" gridrow="12" pageId="8" pageNumber="337">India Philippines: off Iloilo, Panay Island</td>
</tr>
<tr id="99A90961FF94007613C2FC64DD48490A" box="[163,1359,1008,1033]" gridrow="13" pageId="8" pageNumber="337">
<th id="DA78601DFF94007613C2FC64DA44490A" box="[163,579,1008,1033]" gridcol="0" gridrow="13" pageId="8" pageNumber="337">
<taxonomicName id="E09970A0FF94FF8113C2FC64D981490B" authorityName="in Heemstra &amp; Randall" authorityYear="1993" box="[163,390,1008,1032]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="337" phylum="Chordata" rank="species" species="epistictus">
<emphasis id="15EDD731FF94FF8113C2FC64D981490B" box="[163,390,1008,1032]" italics="true" pageId="8" pageNumber="337">Epinephelus epistictus</emphasis>
</taxonomicName>
</th>
<td id="DA78601DFF9400761113FC64DB81490A" box="[626,902,1008,1033]" gridcol="1" gridrow="13" pageId="8" pageNumber="337">
<accessionNumber id="38CA96C0FF94FF811113FC64DAE5490B" box="[626,738,1008,1032]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KU893048" pageId="8" pageNumber="337">KU893048</accessionNumber>
(GenBank)
</td>
<td id="DA78601DFF94007610A7FC64DD48490A" box="[966,1359,1008,1033]" gridcol="2" gridrow="13" pageId="8" pageNumber="337">Taiwan</td>
</tr>
<tr id="99A90961FF94007613C2FB8EDD484930" box="[163,1359,1050,1075]" gridrow="14" pageId="8" pageNumber="337">
<th id="DA78601DFF94007613C2FB8EDA444930" box="[163,579,1050,1075]" gridcol="0" gridrow="14" pageId="8" pageNumber="337">
<taxonomicName id="E09970A0FF94FF8113C2FB8FD9814930" authorityName="in Heemstra &amp; Randall" authorityYear="1993" box="[163,390,1051,1075]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="337" phylum="Chordata" rank="species" species="epistictus">
<emphasis id="15EDD731FF94FF8113C2FB8FD9814930" box="[163,390,1051,1075]" italics="true" pageId="8" pageNumber="337">Epinephelus epistictus</emphasis>
</taxonomicName>
</th>
<td id="DA78601DFF9400761113FB8EDB814930" box="[626,902,1050,1075]" gridcol="1" gridrow="14" pageId="8" pageNumber="337">
<accessionNumber id="38CA96C0FF94FF811113FB8EDAE44930" box="[626,739,1050,1075]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KU943501" pageId="8" pageNumber="337">KU943501</accessionNumber>
(GenBank)
</td>
<td id="DA78601DFF94007610A7FB8EDD484930" box="[966,1359,1050,1075]" gridcol="2" gridrow="14" pageId="8" pageNumber="337">Taiwan</td>
</tr>
<tr id="99A90961FF94007613C2FBD1DD48495D" box="[163,1359,1093,1118]" gridrow="15" pageId="8" pageNumber="337">
<th id="DA78601DFF94007613C2FBD1DA44495D" box="[163,579,1093,1118]" gridcol="0" gridrow="15" pageId="8" pageNumber="337">
<taxonomicName id="E09970A0FF94FF8113C2FBD2D981495E" authorityName="in Heemstra &amp; Randall" authorityYear="1993" box="[163,390,1094,1118]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="337" phylum="Chordata" rank="species" species="epistictus">
<emphasis id="15EDD731FF94FF8113C2FBD2D981495E" box="[163,390,1094,1118]" italics="true" pageId="8" pageNumber="337">Epinephelus epistictus</emphasis>
</taxonomicName>
</th>
<td id="DA78601DFF9400761113FBD1DB81495D" box="[626,902,1093,1118]" gridcol="1" gridrow="15" pageId="8" pageNumber="337">
<accessionNumber id="38CA96C0FF94FF811113FBD1DAE5495D" box="[626,738,1093,1118]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KU943517" pageId="8" pageNumber="337">KU943517</accessionNumber>
(GenBank)
</td>
<td id="DA78601DFF94007610A7FBD1DD48495D" box="[966,1359,1093,1118]" gridcol="2" gridrow="15" pageId="8" pageNumber="337">Taiwan</td>
</tr>
<tr id="99A90961FF94007613C2FBE4DD48498A" box="[163,1359,1136,1161]" gridrow="16" pageId="8" pageNumber="337">
<th id="DA78601DFF94007613C2FBE4DA44498A" box="[163,579,1136,1161]" gridcol="0" gridrow="16" pageId="8" pageNumber="337">
<taxonomicName id="E09970A0FF94FF8113C2FBE4D981498B" authorityName="in Heemstra &amp; Randall" authorityYear="1993" box="[163,390,1136,1160]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="337" phylum="Chordata" rank="species" species="epistictus">
<emphasis id="15EDD731FF94FF8113C2FBE4D981498B" box="[163,390,1136,1160]" italics="true" pageId="8" pageNumber="337">Epinephelus epistictus</emphasis>
</taxonomicName>
</th>
<td id="DA78601DFF9400761113FBE4DB81498A" box="[626,902,1136,1161]" gridcol="1" gridrow="16" pageId="8" pageNumber="337">
<accessionNumber id="38CA96C0FF94FF811113FBE4DAE5498A" box="[626,738,1136,1161]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KU943535" pageId="8" pageNumber="337">KU943535</accessionNumber>
(GenBank)
</td>
<td id="DA78601DFF94007610A7FBE4DD48498A" box="[966,1359,1136,1161]" gridcol="2" gridrow="16" pageId="8" pageNumber="337">Taiwan</td>
</tr>
<tr id="99A90961FF94007613C2FB0EDD4849B0" box="[163,1359,1178,1203]" gridrow="17" pageId="8" pageNumber="337">
<th id="DA78601DFF94007613C2FB0EDA4449B0" box="[163,579,1178,1203]" gridcol="0" gridrow="17" pageId="8" pageNumber="337">
<taxonomicName id="E09970A0FF94FF8113C2FB0FD98149B0" authorityName="in Heemstra &amp; Randall" authorityYear="1993" box="[163,390,1179,1203]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="337" phylum="Chordata" rank="species" species="epistictus">
<emphasis id="15EDD731FF94FF8113C2FB0FD98149B0" box="[163,390,1179,1203]" italics="true" pageId="8" pageNumber="337">Epinephelus epistictus</emphasis>
</taxonomicName>
</th>
<td id="DA78601DFF9400761113FB0EDB8149B0" box="[626,902,1178,1203]" gridcol="1" gridrow="17" pageId="8" pageNumber="337">
<accessionNumber id="38CA96C0FF94FF811113FB0EDAD249B0" box="[626,725,1178,1203]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/FJ237768" pageId="8" pageNumber="337">FJ237768</accessionNumber>
(GenBank)
</td>
<td id="DA78601DFF94007610A7FB0EDD4849B0" box="[966,1359,1178,1203]" gridcol="2" gridrow="17" pageId="8" pageNumber="337">Taiwan</td>
</tr>
<tr id="99A90961FF94007613C2FB51DD4849DD" box="[163,1359,1221,1246]" gridrow="18" pageId="8" pageNumber="337">
<th id="DA78601DFF94007613C2FB51DA4449DD" box="[163,579,1221,1246]" gridcol="0" gridrow="18" pageId="8" pageNumber="337">
<taxonomicName id="E09970A0FF94FF8113C2FB52D98149DE" authorityName="in Heemstra &amp; Randall" authorityYear="1993" box="[163,390,1222,1246]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="337" phylum="Chordata" rank="species" species="epistictus">
<emphasis id="15EDD731FF94FF8113C2FB52D98149DE" box="[163,390,1222,1246]" italics="true" pageId="8" pageNumber="337">Epinephelus epistictus</emphasis>
</taxonomicName>
</th>
<td id="DA78601DFF9400761113FB51DB8149DD" box="[626,902,1221,1246]" gridcol="1" gridrow="18" pageId="8" pageNumber="337">
<accessionNumber id="38CA96C0FF94FF811113FB51DAE549DD" box="[626,738,1221,1246]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MF185489" pageId="8" pageNumber="337">MF185489</accessionNumber>
(GenBank)
</td>
<td id="DA78601DFF94007610A7FB51DD4849DD" box="[966,1359,1221,1246]" gridcol="2" gridrow="18" pageId="8" pageNumber="337">China: Xiamen, Fujian</td>
</tr>
<tr id="99A90961FF94007613C2FB64DD48480A" box="[163,1359,1264,1289]" gridrow="19" pageId="8" pageNumber="337">
<th id="DA78601DFF94007613C2FB64DA44480A" box="[163,579,1264,1289]" gridcol="0" gridrow="19" pageId="8" pageNumber="337">
<taxonomicName id="E09970A0FF94FF8113C2FB64D981480B" authorityName="in Heemstra &amp; Randall" authorityYear="1993" box="[163,390,1264,1288]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="337" phylum="Chordata" rank="species" species="epistictus">
<emphasis id="15EDD731FF94FF8113C2FB64D981480B" box="[163,390,1264,1288]" italics="true" pageId="8" pageNumber="337">Epinephelus epistictus</emphasis>
</taxonomicName>
</th>
<td id="DA78601DFF9400761113FB64DB81480A" box="[626,902,1264,1289]" gridcol="1" gridrow="19" pageId="8" pageNumber="337">
<accessionNumber id="38CA96C0FF94FF811113FB64DAE5480A" box="[626,738,1264,1289]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MF185490" pageId="8" pageNumber="337">MF185490</accessionNumber>
(GenBank)
</td>
<td id="DA78601DFF94007610A7FB64DD48480A" box="[966,1359,1264,1289]" gridcol="2" gridrow="19" pageId="8" pageNumber="337">China: Haikou, Hainan</td>
</tr>
<tr id="99A90961FF94007613C2FA8EDD484830" box="[163,1359,1306,1331]" gridrow="20" pageId="8" pageNumber="337">
<th id="DA78601DFF94007613C2FA8EDA444830" box="[163,579,1306,1331]" gridcol="0" gridrow="20" pageId="8" pageNumber="337">
<taxonomicName id="E09970A0FF94FF8113C2FA8FD9814830" authorityName="in Heemstra &amp; Randall" authorityYear="1993" box="[163,390,1307,1331]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="337" phylum="Chordata" rank="species" species="epistictus">
<emphasis id="15EDD731FF94FF8113C2FA8FD9814830" box="[163,390,1307,1331]" italics="true" pageId="8" pageNumber="337">Epinephelus epistictus</emphasis>
</taxonomicName>
</th>
<td id="DA78601DFF9400761113FA8EDB814830" box="[626,902,1306,1331]" gridcol="1" gridrow="20" pageId="8" pageNumber="337">
<accessionNumber id="38CA96C0FF94FF811113FA8EDAE54830" box="[626,738,1306,1331]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MF185491" pageId="8" pageNumber="337">MF185491</accessionNumber>
(GenBank)
</td>
<td id="DA78601DFF94007610A7FA8EDD484830" box="[966,1359,1306,1331]" gridcol="2" gridrow="20" pageId="8" pageNumber="337">China: Xiamen, Fujian</td>
</tr>
<tr id="99A90961FF94007613C2FAD1DD48485D" box="[163,1359,1349,1374]" gridrow="21" pageId="8" pageNumber="337">
<th id="DA78601DFF94007613C2FAD1DA44485D" box="[163,579,1349,1374]" gridcol="0" gridrow="21" pageId="8" pageNumber="337">
<taxonomicName id="E09970A0FF94FF8113C2FAD2D981485E" authorityName="in Heemstra &amp; Randall" authorityYear="1993" box="[163,390,1350,1374]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="337" phylum="Chordata" rank="species" species="epistictus">
<emphasis id="15EDD731FF94FF8113C2FAD2D981485E" box="[163,390,1350,1374]" italics="true" pageId="8" pageNumber="337">Epinephelus epistictus</emphasis>
</taxonomicName>
</th>
<td id="DA78601DFF9400761113FAD1DB81485D" box="[626,902,1349,1374]" gridcol="1" gridrow="21" pageId="8" pageNumber="337">
<accessionNumber id="38CA96C0FF94FF811113FAD1DAE5485D" box="[626,738,1349,1374]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MF185492" pageId="8" pageNumber="337">MF185492</accessionNumber>
(GenBank)
</td>
<td id="DA78601DFF94007610A7FAD1DD48485D" box="[966,1359,1349,1374]" gridcol="2" gridrow="21" pageId="8" pageNumber="337">China: Sanya, Hainan</td>
</tr>
<tr id="99A90961FF94007613C2FAE4DD48488A" box="[163,1359,1392,1417]" gridrow="22" pageId="8" pageNumber="337">
<th id="DA78601DFF94007613C2FAE4DA44488A" box="[163,579,1392,1417]" gridcol="0" gridrow="22" pageId="8" pageNumber="337">
<taxonomicName id="E09970A0FF94FF8113C2FAE4D981488B" authorityName="in Heemstra &amp; Randall" authorityYear="1993" box="[163,390,1392,1416]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="337" phylum="Chordata" rank="species" species="epistictus">
<emphasis id="15EDD731FF94FF8113C2FAE4D981488B" box="[163,390,1392,1416]" italics="true" pageId="8" pageNumber="337">Epinephelus epistictus</emphasis>
</taxonomicName>
</th>
<td id="DA78601DFF9400761113FAE4DB81488A" box="[626,902,1392,1417]" gridcol="1" gridrow="22" pageId="8" pageNumber="337">
<accessionNumber id="38CA96C0FF94FF811113FAE4DAE5488A" box="[626,738,1392,1417]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MF185493" pageId="8" pageNumber="337">MF185493</accessionNumber>
(GenBank)
</td>
<td id="DA78601DFF94007610A7FAE4DD48488A" box="[966,1359,1392,1417]" gridcol="2" gridrow="22" pageId="8" pageNumber="337">China: Sanya, Hainan</td>
</tr>
<tr id="99A90961FF94007613C2FA0EDD4848B0" box="[163,1359,1434,1459]" gridrow="23" pageId="8" pageNumber="337">
<th id="DA78601DFF94007613C2FA0EDA4448B0" box="[163,579,1434,1459]" gridcol="0" gridrow="23" pageId="8" pageNumber="337">
<taxonomicName id="E09970A0FF94FF8113C2FA0FD98148B0" authorityName="in Heemstra &amp; Randall" authorityYear="1993" box="[163,390,1435,1459]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="337" phylum="Chordata" rank="species" species="epistictus">
<emphasis id="15EDD731FF94FF8113C2FA0FD98148B0" box="[163,390,1435,1459]" italics="true" pageId="8" pageNumber="337">Epinephelus epistictus</emphasis>
</taxonomicName>
</th>
<td id="DA78601DFF9400761113FA0EDB8148B0" box="[626,902,1434,1459]" gridcol="1" gridrow="23" pageId="8" pageNumber="337">MK639008 (GenBank)</td>
<td id="DA78601DFF94007610A7FA0EDD4848B0" box="[966,1359,1434,1459]" gridcol="2" gridrow="23" pageId="8" pageNumber="337">Taiwan: Kaohsiung</td>
</tr>
<tr id="99A90961FF94007613C2FA64DD484B25" box="[163,1359,1520,1574]" gridrow="24" pageId="8" pageNumber="337">
<th id="DA78601DFF94007613C2FA64DA444B25" box="[163,579,1520,1574]" gridcol="0" gridrow="24" pageId="8" pageNumber="337">
<taxonomicName id="E09970A0FF94FF8113C2FA64D9C94B0B" authority="Johnson &amp; Wilmer, 2019" authorityName="Johnson &amp; Wilmer" authorityYear="2019" box="[163,462,1520,1544]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="337" phylum="Chordata" rank="species" species="fuscomarginatus" status="sp. nov.">
<emphasis id="15EDD731FF94FF8113C2FA64D9C94B0B" box="[163,462,1520,1544]" italics="true" pageId="8" pageNumber="337">Epinephelus fuscomarginatus</emphasis>
</taxonomicName>
<emphasis id="15EDD731FF94FF8112B5FA65DA224B0A" bold="true" box="[468,549,1521,1545]" pageId="8" pageNumber="337">
<taxonomicNameLabel id="0EDE6A4AFF94FF8112B5FA65DA224B0A" box="[468,549,1521,1545]" pageId="8" pageNumber="337" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</th>
<td id="DA78601DFF9400761113FA64DB814B25" box="[626,902,1520,1574]" gridcol="1" gridrow="24" pageId="8" pageNumber="337">
<accessionNumber id="38CA96C0FF94FF811113FA64DAE14B0A" box="[626,742,1520,1545]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MK693009" pageId="8" pageNumber="337">MK693009</accessionNumber>
(GenBank)
</td>
<td id="DA78601DFF94007610A7FA64DD484B25" box="[966,1359,1520,1574]" gridcol="2" gridrow="24" pageId="8" pageNumber="337">Australia: Capricorn Channel, south of Swain Reefs, Queensland</td>
</tr>
<tr id="99A90961FF94007613C2F9A1DD484B6F" box="[163,1359,1589,1644]" gridrow="25" pageId="8" pageNumber="337">
<th id="DA78601DFF94007613C2F9A1DA444B6F" box="[163,579,1589,1644]" gridcol="0" gridrow="25" pageId="8" pageNumber="337">
<taxonomicName id="E09970A0FF94FF8113C2F9A2D9C94B4D" authority="Johnson &amp; Wilmer, 2019" authorityName="Johnson &amp; Wilmer" authorityYear="2019" box="[163,462,1590,1614]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="337" phylum="Chordata" rank="species" species="fuscomarginatus" status="sp. nov.">
<emphasis id="15EDD731FF94FF8113C2F9A2D9C94B4D" box="[163,462,1590,1614]" italics="true" pageId="8" pageNumber="337">Epinephelus fuscomarginatus</emphasis>
</taxonomicName>
<emphasis id="15EDD731FF94FF8112B5F9A2DA224B4D" bold="true" box="[468,549,1590,1614]" pageId="8" pageNumber="337">
<taxonomicNameLabel id="0EDE6A4AFF94FF8112B5F9A2DA224B4D" box="[468,549,1590,1614]" pageId="8" pageNumber="337" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</th>
<td id="DA78601DFF9400761113F9A1DB814B6F" box="[626,902,1589,1644]" gridcol="1" gridrow="25" pageId="8" pageNumber="337">
<accessionNumber id="38CA96C0FF94FF811113F9A1DAE14B4D" box="[626,742,1589,1614]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MK693010" pageId="8" pageNumber="337">MK693010</accessionNumber>
(GenBank)
</td>
<td id="DA78601DFF94007610A7F9A1DD484B6F" box="[966,1359,1589,1644]" gridcol="2" gridrow="25" pageId="8" pageNumber="337">Australia: Capricorn Channel, south of Swain Reefs, Queensland</td>
</tr>
<tr id="99A90961FF94007613C2F9EFDD484BB2" box="[163,1359,1659,1713]" gridrow="26" pageId="8" pageNumber="337">
<th id="DA78601DFF94007613C2F9EFDA444BB2" box="[163,579,1659,1713]" gridcol="0" gridrow="26" pageId="8" pageNumber="337">
<taxonomicName id="E09970A0FF94FF8113C2F9E8D9C94B97" authority="Johnson &amp; Wilmer, 2019" authorityName="Johnson &amp; Wilmer" authorityYear="2019" box="[163,462,1660,1684]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="337" phylum="Chordata" rank="species" species="fuscomarginatus" status="sp. nov.">
<emphasis id="15EDD731FF94FF8113C2F9E8D9C94B97" box="[163,462,1660,1684]" italics="true" pageId="8" pageNumber="337">Epinephelus fuscomarginatus</emphasis>
</taxonomicName>
<emphasis id="15EDD731FF94FF8112B5F9E8DA224B97" bold="true" box="[468,549,1660,1684]" pageId="8" pageNumber="337">
<taxonomicNameLabel id="0EDE6A4AFF94FF8112B5F9E8DA224B97" box="[468,549,1660,1684]" pageId="8" pageNumber="337" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</th>
<td id="DA78601DFF9400761113F9EFDB814BB2" box="[626,902,1659,1713]" gridcol="1" gridrow="26" pageId="8" pageNumber="337">
<accessionNumber id="38CA96C0FF94FF811113F9EFDAE14B97" box="[626,742,1659,1684]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MK693011" pageId="8" pageNumber="337">MK693011</accessionNumber>
(GenBank)
</td>
<td id="DA78601DFF94007610A7F9EFDD484BB2" box="[966,1359,1659,1713]" gridcol="2" gridrow="26" pageId="8" pageNumber="337">Australia: Capricorn Channel, south of Swain Reefs, Queensland</td>
</tr>
<tr id="99A90961FF94007613C2F954DD484BF4" box="[163,1359,1728,1783]" gridrow="27" pageId="8" pageNumber="337">
<th id="DA78601DFF94007613C2F954DA444BF4" box="[163,579,1728,1783]" gridcol="0" gridrow="27" pageId="8" pageNumber="337">
<taxonomicName id="E09970A0FF94FF8113C2F955D9C94BDA" authority="Johnson &amp; Wilmer, 2019" authorityName="Johnson &amp; Wilmer" authorityYear="2019" box="[163,462,1729,1753]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="337" phylum="Chordata" rank="species" species="fuscomarginatus" status="sp. nov.">
<emphasis id="15EDD731FF94FF8113C2F955D9C94BDA" box="[163,462,1729,1753]" italics="true" pageId="8" pageNumber="337">Epinephelus fuscomarginatus</emphasis>
</taxonomicName>
<emphasis id="15EDD731FF94FF8112B5F955DA224BDA" bold="true" box="[468,549,1729,1753]" pageId="8" pageNumber="337">
<taxonomicNameLabel id="0EDE6A4AFF94FF8112B5F955DA224BDA" box="[468,549,1729,1753]" pageId="8" pageNumber="337" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</th>
<td id="DA78601DFF9400761113F954DB814BF4" box="[626,902,1728,1783]" gridcol="1" gridrow="27" pageId="8" pageNumber="337">
<accessionNumber id="38CA96C0FF94FF811113F954DAE14BDA" box="[626,742,1728,1753]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MK693012" pageId="8" pageNumber="337">MK693012</accessionNumber>
(GenBank)
</td>
<td id="DA78601DFF94007610A7F954DD484BF4" box="[966,1359,1728,1783]" gridcol="2" gridrow="27" pageId="8" pageNumber="337">Australia: Capricorn Channel, south of Swain Reefs, Queensland</td>
</tr>
<tr id="99A90961FF94007613C2F892DD484A3F" box="[163,1359,1798,1852]" gridrow="28" pageId="8" pageNumber="337">
<th id="DA78601DFF94007613C2F892DA444A3F" box="[163,579,1798,1852]" gridcol="0" gridrow="28" pageId="8" pageNumber="337">
<taxonomicName id="E09970A0FF94FF8113C2F893D9C94A1C" authority="Johnson &amp; Wilmer, 2019" authorityName="Johnson &amp; Wilmer" authorityYear="2019" box="[163,462,1799,1823]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="337" phylum="Chordata" rank="species" species="fuscomarginatus" status="sp. nov.">
<emphasis id="15EDD731FF94FF8113C2F893D9C94A1C" box="[163,462,1799,1823]" italics="true" pageId="8" pageNumber="337">Epinephelus fuscomarginatus</emphasis>
</taxonomicName>
<emphasis id="15EDD731FF94FF8112B5F893DA224A1C" bold="true" box="[468,549,1799,1823]" pageId="8" pageNumber="337">
<taxonomicNameLabel id="0EDE6A4AFF94FF8112B5F893DA224A1C" box="[468,549,1799,1823]" pageId="8" pageNumber="337" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</th>
<td id="DA78601DFF9400761113F892DB814A3F" box="[626,902,1798,1852]" gridcol="1" gridrow="28" pageId="8" pageNumber="337">
<accessionNumber id="38CA96C0FF94FF811113F892DAE04A1C" box="[626,743,1798,1823]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MK693013" pageId="8" pageNumber="337">MK693013</accessionNumber>
(GenBank)
</td>
<td id="DA78601DFF94007610A7F892DD484A3F" box="[966,1359,1798,1852]" gridcol="2" gridrow="28" pageId="8" pageNumber="337">Australia: Capricorn Channel, south of Swain Reefs, Queensland</td>
</tr>
<tr id="99A90961FF94007613C2F8E2DD484A8C" box="[163,1359,1910,1935]" gridrow="29" pageId="8" pageNumber="337">
<th id="DA78601DFF94007613C2F8E2DA444A8C" box="[163,579,1910,1935]" gridcol="0" gridrow="29" pageId="8" pageNumber="337">
<taxonomicName id="E09970A0FF94FF8113C2F8E3D98A4A8C" baseAuthorityName="non Fowler" baseAuthorityYear="1904" box="[163,397,1911,1935]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="337" phylum="Chordata" rank="species" species="heniochus">
<emphasis id="15EDD731FF94FF8113C2F8E3D98A4A8C" box="[163,397,1911,1935]" italics="true" pageId="8" pageNumber="337">Epinephelus heniochus</emphasis>
</taxonomicName>
</th>
<td id="DA78601DFF9400761113F8E2DB814A8C" box="[626,902,1910,1935]" gridcol="1" gridrow="29" pageId="8" pageNumber="337">
<accessionNumber id="38CA96C0FF94FF811113F8E2DAE54A8C" box="[626,738,1910,1935]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KY371468" pageId="8" pageNumber="337">KY371468</accessionNumber>
(GenBank)
</td>
<td id="DA78601DFF94007610A7F8E2DD484A8C" box="[966,1359,1910,1935]" gridcol="2" gridrow="29" pageId="8" pageNumber="337">China: Beibu Gulf</td>
</tr>
<tr id="99A90961FF94007613C2F835DD484AB9" box="[163,1359,1953,1978]" gridrow="30" pageId="8" pageNumber="337">
<th id="DA78601DFF94007613C2F835DA444AB9" box="[163,579,1953,1978]" gridcol="0" gridrow="30" pageId="8" pageNumber="337">
<taxonomicName id="E09970A0FF94FF8113C2F836D98A4AB9" baseAuthorityName="non Fowler" baseAuthorityYear="1904" box="[163,397,1954,1978]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="337" phylum="Chordata" rank="species" species="heniochus">
<emphasis id="15EDD731FF94FF8113C2F836D98A4AB9" box="[163,397,1954,1978]" italics="true" pageId="8" pageNumber="337">Epinephelus heniochus</emphasis>
</taxonomicName>
</th>
<td id="DA78601DFF9400761113F835DB814AB9" box="[626,902,1953,1978]" gridcol="1" gridrow="30" pageId="8" pageNumber="337">
<accessionNumber id="38CA96C0FF94FF811113F835DADD4AB9" box="[626,730,1953,1978]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/JN208617" pageId="8" pageNumber="337">JN208617</accessionNumber>
(GenBank)
</td>
<td id="DA78601DFF94007610A7F835DD484AB9" box="[966,1359,1953,1978]" gridcol="2" gridrow="30" pageId="8" pageNumber="337">N/A</td>
</tr>
</table>
</paragraph>
<paragraph id="27260B23FF94FF8117C4F859DD8D4AE7" blockId="8.[1189,1418,1996,2020]" box="[1189,1418,1996,2020]" pageId="8" pageNumber="337">
<tableNote id="DA7F0AADFF94FF8117C4F859DD8D4AE7" box="[1189,1418,1996,2020]" pageId="8" pageNumber="337" targetBox="[163,1359,194,1978]" targetPageId="8">...Continued next page</tableNote>
</paragraph>
<caption id="73E65BABFF95FF8013C2FF37D9A24DBC" box="[163,421,163,191]" pageId="9" pageNumber="338" startId="9.[163,256,163,190]" targetBox="[163,1383,206,1980]" targetIsTable="true" targetPageId="9">
<paragraph id="27260B23FF95FF8013C2FF37D9A24DBC" blockId="9.[163,421,163,191]" box="[163,421,163,191]" pageId="9" pageNumber="338">
<tableNote id="DA7F0AADFF95FF8013C2FF37D9A24DBC" box="[163,421,163,191]" pageId="9" pageNumber="338" targetBox="[163,1383,206,1980]" targetPageId="9">
<emphasis id="15EDD731FF95FF8013C2FF37D91B4DBD" bold="true" box="[163,284,163,190]" pageId="9" pageNumber="338">TABLE 1.</emphasis>
(
<emphasis id="15EDD731FF95FF801252FF32D99A4DBD" box="[307,413,166,190]" italics="true" pageId="9" pageNumber="338">Continued</emphasis>
)
</tableNote>
</paragraph>
</caption>
<paragraph id="27260B23FF95FF8013C2FF5ADC404ABF" pageId="9" pageNumber="338">
<table id="5599F983FF95007613C2FF5ADD604ABF" box="[163,1383,206,1980]" colsContinueFrom="8.[163,1359,194,1978]" colsContinueIn="10.[163,1209,206,1340]" gridcols="3" gridrows="6" pageId="9" pageNumber="338">
<tr id="99A90961FF95007613C2FF5ADD604DE4" box="[163,1383,206,231]" gridrow="0" pageId="9" pageNumber="338">
<th id="DA78601DFF95007613C2FF5ADA244DE4" box="[163,547,206,231]" gridcol="0" gridrow="0" pageId="9" pageNumber="338">Species</th>
<th id="DA78601DFF9500761113FF5ADB614DE4" box="[626,870,206,231]" gridcol="1" gridrow="0" pageId="9" pageNumber="338">Genbank/BOLD</th>
<th id="DA78601DFF95007610A7FF5ADD604DE4" box="[966,1383,206,231]" gridcol="2" gridrow="0" pageId="9" pageNumber="338">Country</th>
</tr>
<tr id="99A90961FF95007613C2FF6DDD604C3F" box="[163,1383,249,316]" gridrow="1" pageId="9" pageNumber="338">
<th id="DA78601DFF95007613C2FF6DDA244C3F" box="[163,547,249,316]" gridcol="0" gridrow="1" pageId="9" pageNumber="338">
<emphasis id="15EDD731FF95FF8013C2FF6DD98A4C3F" italics="true" pageId="9" pageNumber="338">
<taxonomicName id="E09970A0FF95FF8013C2FF6DD98A4C12" baseAuthorityName="non Fowler" baseAuthorityYear="1904" box="[163,397,249,273]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="338" phylum="Chordata" rank="species" species="heniochus">Epinephelus heniochus</taxonomicName>
<taxonomicName id="E09970A0FF95FF8013C2FEB0D98A4C3F" baseAuthorityName="non Fowler" baseAuthorityYear="1904" box="[163,397,292,316]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="338" phylum="Chordata" rank="species" species="heniochus">Epinephelus heniochus</taxonomicName>
</emphasis>
</th>
<td id="DA78601DFF9500761113FF6DDB614C3F" box="[626,870,249,316]" gridcol="1" gridrow="1" pageId="9" pageNumber="338">
<accessionNumber id="38CA96C0FF95FF801113FF6DDADD4C11" box="[626,730,249,274]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/JN208618" pageId="9" pageNumber="338">JN208618</accessionNumber>
(GenBank) 
<accessionNumber id="38CA96C0FF95FF801113FEB7DAE64C3F" box="[626,737,291,316]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KR863512" pageId="9" pageNumber="338">KR863512</accessionNumber>
(GenBank)
</td>
<td id="DA78601DFF95007610A7FF6DDD604C3F" box="[966,1383,249,316]" gridcol="2" gridrow="1" pageId="9" pageNumber="338">N/A Malaysia</td>
</tr>
<tr id="99A90961FF95007613C2FEEDDD604964" box="[163,1383,377,1127]" gridrow="2" pageId="9" pageNumber="338">
<th id="DA78601DFF95007613C2FEEDDA244964" box="[163,547,377,1127]" gridcol="0" gridrow="2" pageId="9" pageNumber="338">
<emphasis id="15EDD731FF95FF8013C2FEEDD9A54964" italics="true" pageId="9" pageNumber="338">
<taxonomicName id="E09970A0FF95FF8013C2FEEDD9A54C92" baseAuthorityName="Temminck &amp; Schlegel" baseAuthorityYear="1843" box="[163,418,377,401]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="338" phylum="Chordata" rank="species" species="latifasciatus">Epinephelus latifasciatus</taxonomicName>
<taxonomicName id="E09970A0FF95FF8013C2FE30D9A54CBF" baseAuthorityName="Temminck &amp; Schlegel" baseAuthorityYear="1843" box="[163,418,420,444]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="338" phylum="Chordata" rank="species" species="latifasciatus">Epinephelus latifasciatus</taxonomicName>
<taxonomicName id="E09970A0FF95FF8013C2FE5BD9A54CE4" baseAuthorityName="Temminck &amp; Schlegel" baseAuthorityYear="1843" box="[163,418,463,487]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="338" phylum="Chordata" rank="species" species="latifasciatus">Epinephelus latifasciatus</taxonomicName>
<taxonomicName id="E09970A0FF95FF8013C2FE6DD9A54F12" baseAuthorityName="Temminck &amp; Schlegel" baseAuthorityYear="1843" box="[163,418,505,529]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="338" phylum="Chordata" rank="species" species="latifasciatus">Epinephelus latifasciatus</taxonomicName>
<taxonomicName id="E09970A0FF95FF8013C2FDB0D9A54F3F" baseAuthorityName="Temminck &amp; Schlegel" baseAuthorityYear="1843" box="[163,418,548,572]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="338" phylum="Chordata" rank="species" species="latifasciatus">Epinephelus latifasciatus</taxonomicName>
<taxonomicName id="E09970A0FF95FF8013C2FDDBD9A54F64" baseAuthorityName="Temminck &amp; Schlegel" baseAuthorityYear="1843" box="[163,418,591,615]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="338" phylum="Chordata" rank="species" species="latifasciatus">Epinephelus latifasciatus</taxonomicName>
<taxonomicName id="E09970A0FF95FF8013C2FDEDD9A54F92" baseAuthorityName="Temminck &amp; Schlegel" baseAuthorityYear="1843" box="[163,418,633,657]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="338" phylum="Chordata" rank="species" species="latifasciatus">Epinephelus latifasciatus</taxonomicName>
<taxonomicName id="E09970A0FF95FF8013C2FD30D9A54FBF" baseAuthorityName="Temminck &amp; Schlegel" baseAuthorityYear="1843" box="[163,418,676,700]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="338" phylum="Chordata" rank="species" species="latifasciatus">Epinephelus latifasciatus</taxonomicName>
<taxonomicName id="E09970A0FF95FF8013C2FD5BD9A54FE4" baseAuthorityName="Temminck &amp; Schlegel" baseAuthorityYear="1843" box="[163,418,719,743]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="338" phylum="Chordata" rank="species" species="latifasciatus">Epinephelus latifasciatus</taxonomicName>
<taxonomicName id="E09970A0FF95FF8013C2FD6DD9A54E12" baseAuthorityName="Temminck &amp; Schlegel" baseAuthorityYear="1843" box="[163,418,761,785]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="338" phylum="Chordata" rank="species" species="latifasciatus">Epinephelus latifasciatus</taxonomicName>
<taxonomicName id="E09970A0FF95FF8013C2FCB0D9A54E3F" baseAuthorityName="Temminck &amp; Schlegel" baseAuthorityYear="1843" box="[163,418,804,828]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="338" phylum="Chordata" rank="species" species="latifasciatus">Epinephelus latifasciatus</taxonomicName>
<taxonomicName id="E09970A0FF95FF8013C2FCDBD9A54E64" baseAuthorityName="Temminck &amp; Schlegel" baseAuthorityYear="1843" box="[163,418,847,871]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="338" phylum="Chordata" rank="species" species="latifasciatus">Epinephelus latifasciatus</taxonomicName>
<taxonomicName id="E09970A0FF95FF8013C2FCEDD9A54E92" baseAuthorityName="Temminck &amp; Schlegel" baseAuthorityYear="1843" box="[163,418,889,913]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="338" phylum="Chordata" rank="species" species="latifasciatus">Epinephelus latifasciatus</taxonomicName>
<taxonomicName id="E09970A0FF95FF8013C2FC30D9A54EBF" baseAuthorityName="Temminck &amp; Schlegel" baseAuthorityYear="1843" box="[163,418,932,956]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="338" phylum="Chordata" rank="species" species="latifasciatus">Epinephelus latifasciatus</taxonomicName>
<taxonomicName id="E09970A0FF95FF8013C2FC5BD9A54EE4" baseAuthorityName="Temminck &amp; Schlegel" baseAuthorityYear="1843" box="[163,418,975,999]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="338" phylum="Chordata" rank="species" species="latifasciatus">Epinephelus latifasciatus</taxonomicName>
<taxonomicName id="E09970A0FF95FF8013C2FC6DD9A54912" baseAuthorityName="Temminck &amp; Schlegel" baseAuthorityYear="1843" box="[163,418,1017,1041]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="338" phylum="Chordata" rank="species" species="latifasciatus">Epinephelus latifasciatus</taxonomicName>
<taxonomicName id="E09970A0FF95FF8013C2FBB0D9A5493F" baseAuthorityName="Temminck &amp; Schlegel" baseAuthorityYear="1843" box="[163,418,1060,1084]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="338" phylum="Chordata" rank="species" species="latifasciatus">Epinephelus latifasciatus</taxonomicName>
<taxonomicName id="E09970A0FF95FF8013C2FBDBD9A54964" baseAuthorityName="Temminck &amp; Schlegel" baseAuthorityYear="1843" box="[163,418,1103,1127]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="338" phylum="Chordata" rank="species" species="latifasciatus">Epinephelus latifasciatus</taxonomicName>
</emphasis>
</th>
<td id="DA78601DFF9500761113FEEDDB614964" box="[626,870,377,1127]" gridcol="1" gridrow="2" pageId="9" pageNumber="338">
<accessionNumber id="38CA96C0FF95FF801113FEEDDAD84C91" box="[626,735,377,402]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/EU014216" pageId="9" pageNumber="338">EU014216</accessionNumber>
(GenBank) 
<accessionNumber id="38CA96C0FF95FF801113FE30DAD84CBF" box="[626,735,420,444]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/EU014217" pageId="9" pageNumber="338">EU014217</accessionNumber>
(GenBank) 
<accessionNumber id="38CA96C0FF95FF801113FE5BDAD84CE4" box="[626,735,463,487]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/EU014218" pageId="9" pageNumber="338">EU014218</accessionNumber>
(GenBank) 
<accessionNumber id="38CA96C0FF95FF801113FE6DDAD84F12" box="[626,735,505,529]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/EU014219" pageId="9" pageNumber="338">EU014219</accessionNumber>
(GenBank) 
<accessionNumber id="38CA96C0FF95FF801113FDB7DAE54F3F" box="[626,738,547,572]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/HQ564415" pageId="9" pageNumber="338">HQ564415</accessionNumber>
(GenBank) 
<accessionNumber id="38CA96C0FF95FF801113FDDADAE14F64" box="[626,742,590,615]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KM226269" pageId="9" pageNumber="338">KM226269</accessionNumber>
(GenBank) 
<accessionNumber id="38CA96C0FF95FF801113FDEDDAE14F91" box="[626,742,633,658]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KM226270" pageId="9" pageNumber="338">KM226270</accessionNumber>
(GenBank) 
<accessionNumber id="38CA96C0FF95FF801113FD37DAE14FBF" box="[626,742,675,700]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KM656845" pageId="9" pageNumber="338">KM656845</accessionNumber>
(GenBank) KT8355689 (GenBank) 
<accessionNumber id="38CA96C0FF95FF801113FD6DDAE54E11" box="[626,738,761,786]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KU668639" pageId="9" pageNumber="338">KU668639</accessionNumber>
(GenBank) 
<accessionNumber id="38CA96C0FF95FF801113FCB0DAE54E3F" box="[626,738,804,828]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KU943473" pageId="9" pageNumber="338">KU943473</accessionNumber>
(GenBank) 
<accessionNumber id="38CA96C0FF95FF801113FCDADAE54E64" box="[626,738,846,871]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KU943485" pageId="9" pageNumber="338">KU943485</accessionNumber>
(GenBank) 
<accessionNumber id="38CA96C0FF95FF801113FCEDDAE54E91" box="[626,738,889,914]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KU943506" pageId="9" pageNumber="338">KU943506</accessionNumber>
(GenBank) 
<accessionNumber id="38CA96C0FF95FF801113FC37DAE54EBF" box="[626,738,931,956]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KU943540" pageId="9" pageNumber="338">KU943540</accessionNumber>
(GenBank) 
<accessionNumber id="38CA96C0FF95FF801113FC5BDAE54EE4" box="[626,738,975,999]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KY371470" pageId="9" pageNumber="338">KY371470</accessionNumber>
(GenBank) 
<accessionNumber id="38CA96C0FF95FF801113FC6DDAE54911" box="[626,738,1017,1042]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MF185521" pageId="9" pageNumber="338">MF185521</accessionNumber>
(GenBank) 
<accessionNumber id="38CA96C0FF95FF801113FBB7DAE5493F" box="[626,738,1059,1084]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MF185522" pageId="9" pageNumber="338">MF185522</accessionNumber>
(GenBank) 
<accessionNumber id="38CA96C0FF95FF801113FBDADAE54964" box="[626,738,1102,1127]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MF185523" pageId="9" pageNumber="338">MF185523</accessionNumber>
(GenBank)
</td>
<td id="DA78601DFF95007610A7FEEDDD604964" box="[966,1383,377,1127]" gridcol="2" gridrow="2" pageId="9" pageNumber="338">India India India India Indonesia: Bali India India Taiwan India N/A Taiwan Taiwan Taiwan Taiwan China China: Hong Kong China: Sanya, Hainan China: Xiamen, Fujian</td>
</tr>
<tr id="99A90961FF95007613C2FB37DD60483F" box="[163,1383,1187,1340]" gridrow="3" pageId="9" pageNumber="338">
<th id="DA78601DFF95007613C2FB37DA24483F" box="[163,547,1187,1340]" gridcol="0" gridrow="3" pageId="9" pageNumber="338">
<emphasis id="15EDD731FF95FF8013C2FB30D9B44812" italics="true" pageId="9" pageNumber="338">
<taxonomicName id="E09970A0FF95FF8013C2FB30D9A149BF" box="[163,422,1188,1212]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="338" phylum="Chordata" rank="species" species="magniscuttis">Epinephelus magniscuttis</taxonomicName>
<taxonomicName id="E09970A0FF95FF8013C2FB5BD9A149E5" box="[163,422,1231,1255]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="338" phylum="Chordata" rank="species" species="magniscuttis">Epinephelus magniscuttis</taxonomicName>
<taxonomicName id="E09970A0FF95FF8013C2FB6DD9AE4812" box="[163,425,1273,1297]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="338" phylum="Chordata" rank="species" species="magniscuttis">Epinephelus magniscuttis</taxonomicName>
*
</emphasis>
*Species ID in Genbank: 
<taxonomicName id="E09970A0FF95FF8012C6FAB1DA24483F" authorityName="in Heemstra &amp; Randall" authorityYear="1993" box="[423,547,1317,1340]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="338" phylum="Chordata" rank="species" species="epistictus">
<emphasis id="15EDD731FF95FF8012C6FAB1DA24483F" box="[423,547,1317,1340]" italics="true" pageId="9" pageNumber="338">E. epistictus</emphasis>
</taxonomicName>
</th>
<td id="DA78601DFF9500761113FB37DB61483F" box="[626,870,1187,1340]" gridcol="1" gridrow="3" pageId="9" pageNumber="338">
<accessionNumber id="38CA96C0FF95FF801113FB37DAE149BF" box="[626,742,1187,1212]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MH800836" pageId="9" pageNumber="338">MH800836</accessionNumber>
(GenBank) 
<accessionNumber id="38CA96C0FF95FF801113FB5ADAE149E4" box="[626,742,1230,1255]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MK693014" pageId="9" pageNumber="338">MK693014</accessionNumber>
(GenBank) 
<accessionNumber id="38CA96C0FF95FF801113FB6DDAE24812" box="[626,741,1273,1297]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KU893048" pageId="9" pageNumber="338">KU893048</accessionNumber>
* (GenBank)
</td>
<td id="DA78601DFF95007610A7FB37DD60483F" box="[966,1383,1187,1340]" gridcol="2" gridrow="3" pageId="9" pageNumber="338">Japan Australia: off Augusta, Western Australia Taiwan</td>
</tr>
<tr id="99A90961FF95007613C2FAEDDD604A3F" box="[163,1383,1401,1852]" gridrow="4" pageId="9" pageNumber="338">
<th id="DA78601DFF95007613C2FAEDDA244A3F" box="[163,547,1401,1852]" gridcol="0" gridrow="4" pageId="9" pageNumber="338">
<emphasis id="15EDD731FF95FF8013C2FAEDD9794A3F" italics="true" pageId="9" pageNumber="338">
<taxonomicName id="E09970A0FF95FF8013C2FAEDD9794892" baseAuthorityName="Valenciennes" baseAuthorityYear="1833" box="[163,382,1401,1425]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="338" phylum="Chordata" rank="species" species="morrhua">Epinephelus morrhua</taxonomicName>
<taxonomicName id="E09970A0FF95FF8013C2FA30D97948BF" baseAuthorityName="Valenciennes" baseAuthorityYear="1833" box="[163,382,1444,1468]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="338" phylum="Chordata" rank="species" species="morrhua">Epinephelus morrhua</taxonomicName>
<taxonomicName id="E09970A0FF95FF8013C2FA5BD97948E4" baseAuthorityName="Valenciennes" baseAuthorityYear="1833" box="[163,382,1487,1511]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="338" phylum="Chordata" rank="species" species="morrhua">Epinephelus morrhua</taxonomicName>
<taxonomicName id="E09970A0FF95FF8013C2FA6DD9794B12" baseAuthorityName="Valenciennes" baseAuthorityYear="1833" box="[163,382,1529,1553]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="338" phylum="Chordata" rank="species" species="morrhua">Epinephelus morrhua</taxonomicName>
<taxonomicName id="E09970A0FF95FF8013C2F9B0D9794B3F" baseAuthorityName="Valenciennes" baseAuthorityYear="1833" box="[163,382,1572,1596]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="338" phylum="Chordata" rank="species" species="morrhua">Epinephelus morrhua</taxonomicName>
<taxonomicName id="E09970A0FF95FF8013C2F9DBD9794B64" baseAuthorityName="Valenciennes" baseAuthorityYear="1833" box="[163,382,1615,1639]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="338" phylum="Chordata" rank="species" species="morrhua">Epinephelus morrhua</taxonomicName>
<taxonomicName id="E09970A0FF95FF8013C2F9EDD9794B92" baseAuthorityName="Valenciennes" baseAuthorityYear="1833" box="[163,382,1657,1681]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="338" phylum="Chordata" rank="species" species="morrhua">Epinephelus morrhua</taxonomicName>
<taxonomicName id="E09970A0FF95FF8013C2F930D9794BBF" baseAuthorityName="Valenciennes" baseAuthorityYear="1833" box="[163,382,1700,1724]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="338" phylum="Chordata" rank="species" species="morrhua">Epinephelus morrhua</taxonomicName>
<taxonomicName id="E09970A0FF95FF8013C2F95BD9794BE4" baseAuthorityName="Valenciennes" baseAuthorityYear="1833" box="[163,382,1743,1767]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="338" phylum="Chordata" rank="species" species="morrhua">Epinephelus morrhua</taxonomicName>
<taxonomicName id="E09970A0FF95FF8013C2F96DD9794A12" baseAuthorityName="Valenciennes" baseAuthorityYear="1833" box="[163,382,1785,1809]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="338" phylum="Chordata" rank="species" species="morrhua">Epinephelus morrhua</taxonomicName>
<taxonomicName id="E09970A0FF95FF8013C2F8B0D9794A3F" baseAuthorityName="Valenciennes" baseAuthorityYear="1833" box="[163,382,1828,1852]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="338" phylum="Chordata" rank="species" species="morrhua">Epinephelus morrhua</taxonomicName>
</emphasis>
</th>
<td id="DA78601DFF9500761113FAEDDB614A3F" box="[626,870,1401,1852]" gridcol="1" gridrow="4" pageId="9" pageNumber="338">
<accessionNumber id="38CA96C0FF95FF801113FAEDDAE54891" box="[626,738,1401,1426]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/DQ107896" pageId="9" pageNumber="338">DQ107896</accessionNumber>
(GenBank) 
<accessionNumber id="38CA96C0FF95FF801113FA30DAE548BF" box="[626,738,1444,1468]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/DQ107897" pageId="9" pageNumber="338">DQ107897</accessionNumber>
(GenBank) 
<accessionNumber id="38CA96C0FF95FF801113FA5BDAE748E4" box="[626,736,1487,1511]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/EU392188" pageId="9" pageNumber="338">EU392188</accessionNumber>
(GenBank) 
<accessionNumber id="38CA96C0FF95FF801113FA6DDAD84B12" box="[626,735,1529,1553]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/EU392189" pageId="9" pageNumber="338">EU392189</accessionNumber>
(GenBank) 
<accessionNumber id="38CA96C0FF95FF801113F9B0DAE14B3F" box="[626,742,1572,1596]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KM077930" pageId="9" pageNumber="338">KM077930</accessionNumber>
(GenBank) 
<accessionNumber id="38CA96C0FF95FF801113F9DADAE14B64" box="[626,742,1614,1639]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KM226284" pageId="9" pageNumber="338">KM226284</accessionNumber>
(GenBank) 
<accessionNumber id="38CA96C0FF95FF801113F9EDDAE14B91" box="[626,742,1657,1682]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KM226285" pageId="9" pageNumber="338">KM226285</accessionNumber>
(GenBank) 
<accessionNumber id="38CA96C0FF95FF801113F937DAE54BBF" box="[626,738,1699,1724]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KU943486" pageId="9" pageNumber="338">KU943486</accessionNumber>
(GenBank) 
<accessionNumber id="38CA96C0FF95FF801113F95BDAE54BE4" box="[626,738,1743,1767]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KY371488" pageId="9" pageNumber="338">KY371488</accessionNumber>
(GenBank) WLIND133 (BOLD) WLIND134 (BOLD)
</td>
<td id="DA78601DFF95007610A7FAEDDD604A3F" box="[966,1383,1401,1852]" gridcol="2" gridrow="4" pageId="9" pageNumber="338">Australia: Queensland N/A India India New Caledonia India India Taiwan China: north South China Sea India India</td>
</tr>
<tr id="99A90961FF95007613C2F8EDDD604ABF" box="[163,1383,1913,1980]" gridrow="5" pageId="9" pageNumber="338">
<th id="DA78601DFF95007613C2F8EDDA244ABF" box="[163,547,1913,1980]" gridcol="0" gridrow="5" pageId="9" pageNumber="338">
<emphasis id="15EDD731FF95FF8013C2F8EDD9A04ABF" italics="true" pageId="9" pageNumber="338">
<taxonomicName id="E09970A0FF95FF8013C2F8EDD9A04A92" baseAuthorityName="Temminck &amp; Schlegel" baseAuthorityYear="1842" box="[163,423,1913,1937]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="338" phylum="Chordata" rank="species" species="poecilonotus">Epinephelus poecilonotus</taxonomicName>
<taxonomicName id="E09970A0FF95FF8013C2F830D9A04ABF" baseAuthorityName="Temminck &amp; Schlegel" baseAuthorityYear="1842" box="[163,423,1956,1980]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="338" phylum="Chordata" rank="species" species="poecilonotus">Epinephelus poecilonotus</taxonomicName>
</emphasis>
</th>
<td id="DA78601DFF9500761113F8EDDB614ABF" box="[626,870,1913,1980]" gridcol="1" gridrow="5" pageId="9" pageNumber="338">DSLAF241 (BOLD) DSLAF245 (BOLD)</td>
<td id="DA78601DFF95007610A7F8EDDD604ABF" box="[966,1383,1913,1980]" gridcol="2" gridrow="5" pageId="9" pageNumber="338">South Africa South Africa</td>
</tr>
</table>
</paragraph>
<paragraph id="27260B23FF95FF8017C4F85BDD8D4AE5" blockId="9.[1189,1418,1999,2023]" box="[1189,1418,1999,2023]" pageId="9" pageNumber="338">
<tableNote id="DA7F0AADFF95FF8017C4F85BDD8D4AE5" box="[1189,1418,1999,2023]" pageId="9" pageNumber="338" targetBox="[163,1383,206,1980]" targetPageId="9">...Continued next page</tableNote>
</paragraph>
<caption id="73E65BABFF96FF8313C2FF37D9A24DBC" box="[163,421,163,191]" pageId="10" pageNumber="339" startId="10.[163,256,163,190]" targetBox="[163,1209,206,1340]" targetIsTable="true" targetPageId="10">
<paragraph id="27260B23FF96FF8313C2FF37D9A24DBC" blockId="10.[163,421,163,191]" box="[163,421,163,191]" pageId="10" pageNumber="339">
<tableNote id="DA7F0AADFF96FF8313C2FF37D9A24DBC" box="[163,421,163,191]" pageId="10" pageNumber="339" targetBox="[163,1209,206,1340]" targetPageId="10">
<emphasis id="15EDD731FF96FF8313C2FF37D91B4DBD" bold="true" box="[163,284,163,190]" pageId="10" pageNumber="339">TABLE 1.</emphasis>
(
<emphasis id="15EDD731FF96FF831252FF32D99A4DBD" box="[307,413,166,190]" italics="true" pageId="10" pageNumber="339">Continued</emphasis>
)
</tableNote>
</paragraph>
</caption>
<paragraph id="27260B23FF96FF8313C2FF5ADC9D483F" pageId="10" pageNumber="339">
<table id="5599F983FF96007613C2FF5ADCBE483F" box="[163,1209,206,1340]" colsContinueFrom="9.[163,1383,206,1980]" gridcols="3" gridrows="5" pageId="10" pageNumber="339">
<tr id="99A90961FF96007613C2FF5ADCBE4DE4" box="[163,1209,206,231]" gridrow="0" pageId="10" pageNumber="339">
<th id="DA78601DFF96007613C2FF5AD9A04DE4" box="[163,423,206,231]" gridcol="0" gridrow="0" pageId="10" pageNumber="339">Species</th>
<th id="DA78601DFF9600761113FF5ADB5A4DE4" box="[626,861,206,231]" gridcol="1" gridrow="0" pageId="10" pageNumber="339">Genbank/BOLD</th>
<th id="DA78601DFF96007610A7FF5ADCBE4DE4" box="[966,1209,206,231]" gridcol="2" gridrow="0" pageId="10" pageNumber="339">Country</th>
</tr>
<tr id="99A90961FF96007613C2FF6DDCBE4C11" box="[163,1209,249,274]" gridrow="1" pageId="10" pageNumber="339">
<th id="DA78601DFF96007613C2FF6DD9A04C11" box="[163,423,249,274]" gridcol="0" gridrow="1" pageId="10" pageNumber="339">
<taxonomicName id="E09970A0FF96FF8313C2FF6DD9A04C12" baseAuthorityName="Temminck &amp; Schlegel" baseAuthorityYear="1842" box="[163,423,249,273]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="339" phylum="Chordata" rank="species" species="poecilonotus">
<emphasis id="15EDD731FF96FF8313C2FF6DD9A04C12" box="[163,423,249,273]" italics="true" pageId="10" pageNumber="339">Epinephelus poecilonotus</emphasis>
</taxonomicName>
</th>
<td id="DA78601DFF9600761113FF6DDB5A4C11" box="[626,861,249,274]" gridcol="1" gridrow="1" pageId="10" pageNumber="339">DSLAG1400 (BOLD)</td>
<td id="DA78601DFF96007610A7FF6DDCBE4C11" box="[966,1209,249,274]" gridcol="2" gridrow="1" pageId="10" pageNumber="339">South Africa</td>
</tr>
<tr id="99A90961FF96007613C2FEB7DCBE4FE4" box="[163,1209,291,743]" gridrow="2" pageId="10" pageNumber="339">
<th id="DA78601DFF96007613C2FEB7D9A04FE4" box="[163,423,291,743]" gridcol="0" gridrow="2" pageId="10" pageNumber="339">
<emphasis id="15EDD731FF96FF8313C2FEB0D9A04FE4" italics="true" pageId="10" pageNumber="339">
<taxonomicName id="E09970A0FF96FF8313C2FEB0D9A04C3F" baseAuthorityName="Temminck &amp; Schlegel" baseAuthorityYear="1842" box="[163,423,292,316]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="339" phylum="Chordata" rank="species" species="poecilonotus">Epinephelus poecilonotus</taxonomicName>
<taxonomicName id="E09970A0FF96FF8313C2FEDBD9A04C64" baseAuthorityName="Temminck &amp; Schlegel" baseAuthorityYear="1842" box="[163,423,335,359]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="339" phylum="Chordata" rank="species" species="poecilonotus">Epinephelus poecilonotus</taxonomicName>
<taxonomicName id="E09970A0FF96FF8313C2FEEDD9A04C92" baseAuthorityName="Temminck &amp; Schlegel" baseAuthorityYear="1842" box="[163,423,377,401]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="339" phylum="Chordata" rank="species" species="poecilonotus">Epinephelus poecilonotus</taxonomicName>
<taxonomicName id="E09970A0FF96FF8313C2FE30D9A04CBF" baseAuthorityName="Temminck &amp; Schlegel" baseAuthorityYear="1842" box="[163,423,420,444]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="339" phylum="Chordata" rank="species" species="poecilonotus">Epinephelus poecilonotus</taxonomicName>
<taxonomicName id="E09970A0FF96FF8313C2FE5BD9A04CE4" baseAuthorityName="Temminck &amp; Schlegel" baseAuthorityYear="1842" box="[163,423,463,487]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="339" phylum="Chordata" rank="species" species="poecilonotus">Epinephelus poecilonotus</taxonomicName>
<taxonomicName id="E09970A0FF96FF8313C2FE6DD9A04F12" baseAuthorityName="Temminck &amp; Schlegel" baseAuthorityYear="1842" box="[163,423,505,529]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="339" phylum="Chordata" rank="species" species="poecilonotus">Epinephelus poecilonotus</taxonomicName>
<taxonomicName id="E09970A0FF96FF8313C2FDB0D9A04F3F" baseAuthorityName="Temminck &amp; Schlegel" baseAuthorityYear="1842" box="[163,423,548,572]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="339" phylum="Chordata" rank="species" species="poecilonotus">Epinephelus poecilonotus</taxonomicName>
<taxonomicName id="E09970A0FF96FF8313C2FDDBD9A04F64" baseAuthorityName="Temminck &amp; Schlegel" baseAuthorityYear="1842" box="[163,423,591,615]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="339" phylum="Chordata" rank="species" species="poecilonotus">Epinephelus poecilonotus</taxonomicName>
<taxonomicName id="E09970A0FF96FF8313C2FDEDD9A04F92" baseAuthorityName="Temminck &amp; Schlegel" baseAuthorityYear="1842" box="[163,423,633,657]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="339" phylum="Chordata" rank="species" species="poecilonotus">Epinephelus poecilonotus</taxonomicName>
<taxonomicName id="E09970A0FF96FF8313C2FD30D9A04FBF" baseAuthorityName="Temminck &amp; Schlegel" baseAuthorityYear="1842" box="[163,423,676,700]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="339" phylum="Chordata" rank="species" species="poecilonotus">Epinephelus poecilonotus</taxonomicName>
<taxonomicName id="E09970A0FF96FF8313C2FD5BD9A04FE4" baseAuthorityName="Temminck &amp; Schlegel" baseAuthorityYear="1842" box="[163,423,719,743]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="339" phylum="Chordata" rank="species" species="poecilonotus">Epinephelus poecilonotus</taxonomicName>
</emphasis>
</th>
<td id="DA78601DFF9600761113FEB7DB5A4FE4" box="[626,861,291,743]" gridcol="1" gridrow="2" pageId="10" pageNumber="339">
DSLAG1401 (BOLD) 
<accessionNumber id="38CA96C0FF96FF831113FEDADAE54C64" box="[626,738,334,359]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/GU804952" pageId="10" pageNumber="339">GU804952</accessionNumber>
(GenBank) 
<accessionNumber id="38CA96C0FF96FF831113FEEDDAE54C91" box="[626,738,377,402]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/GU805042" pageId="10" pageNumber="339">GU805042</accessionNumber>
(GenBank) 
<accessionNumber id="38CA96C0FF96FF831113FE37DAE54CBF" box="[626,738,419,444]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/HQ945797" pageId="10" pageNumber="339">HQ945797</accessionNumber>
(GenBank) 
<accessionNumber id="38CA96C0FF96FF831113FE5ADAD24CE4" box="[626,725,462,487]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/JF493454" pageId="10" pageNumber="339">JF493454</accessionNumber>
(GenBank) 
<accessionNumber id="38CA96C0FF96FF831113FE6DDAD94F11" box="[626,734,505,530]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KF489586" pageId="10" pageNumber="339">KF489586</accessionNumber>
(GenBank) 
<accessionNumber id="38CA96C0FF96FF831113FDB7DAD94F3F" box="[626,734,547,572]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KF929856" pageId="10" pageNumber="339">KF929856</accessionNumber>
(GenBank) 
<accessionNumber id="38CA96C0FF96FF831113FDDADAE14F64" box="[626,742,590,615]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KM226286" pageId="10" pageNumber="339">KM226286</accessionNumber>
(GenBank) 
<accessionNumber id="38CA96C0FF96FF831113FDEDDAE14F91" box="[626,742,633,658]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KM226287" pageId="10" pageNumber="339">KM226287</accessionNumber>
(GenBank) TZSAL642 (BOLD) TZSAL643 (BOLD)
</td>
<td id="DA78601DFF96007610A7FEB7DCBE4FE4" box="[966,1209,291,743]" gridcol="2" gridrow="2" pageId="10" pageNumber="339">South Africa South Africa South Africa South Africa South Africa South Africa Seychelles India India South Africa South Africa</td>
</tr>
<tr id="99A90961FF96007613C2FCB7DCBE4964" box="[163,1209,803,1127]" gridrow="3" pageId="10" pageNumber="339">
<th id="DA78601DFF96007613C2FCB7D9A04964" box="[163,423,803,1127]" gridcol="0" gridrow="3" pageId="10" pageNumber="339">
<emphasis id="15EDD731FF96FF8313C2FCB0D97D4964" italics="true" pageId="10" pageNumber="339">
<taxonomicName id="E09970A0FF96FF8313C2FCB0D97D4E3F" baseAuthorityName="Day" baseAuthorityYear="1868" box="[163,378,804,828]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="339" phylum="Chordata" rank="species" species="radiatus">Epinephelus radiatus</taxonomicName>
<taxonomicName id="E09970A0FF96FF8313C2FCDBD97D4E64" baseAuthorityName="Day" baseAuthorityYear="1868" box="[163,378,847,871]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="339" phylum="Chordata" rank="species" species="radiatus">Epinephelus radiatus</taxonomicName>
<taxonomicName id="E09970A0FF96FF8313C2FCEDD97D4E92" baseAuthorityName="Day" baseAuthorityYear="1868" box="[163,378,889,913]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="339" phylum="Chordata" rank="species" species="radiatus">Epinephelus radiatus</taxonomicName>
<taxonomicName id="E09970A0FF96FF8313C2FC30D97D4EBF" baseAuthorityName="Day" baseAuthorityYear="1868" box="[163,378,932,956]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="339" phylum="Chordata" rank="species" species="radiatus">Epinephelus radiatus</taxonomicName>
<taxonomicName id="E09970A0FF96FF8313C2FC5BD97D4EE4" baseAuthorityName="Day" baseAuthorityYear="1868" box="[163,378,975,999]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="339" phylum="Chordata" rank="species" species="radiatus">Epinephelus radiatus</taxonomicName>
<taxonomicName id="E09970A0FF96FF8313C2FC6DD97D4912" baseAuthorityName="Day" baseAuthorityYear="1868" box="[163,378,1017,1041]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="339" phylum="Chordata" rank="species" species="radiatus">Epinephelus radiatus</taxonomicName>
<taxonomicName id="E09970A0FF96FF8313C2FBB0D97D493F" baseAuthorityName="Day" baseAuthorityYear="1868" box="[163,378,1060,1084]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="339" phylum="Chordata" rank="species" species="radiatus">Epinephelus radiatus</taxonomicName>
<taxonomicName id="E09970A0FF96FF8313C2FBDBD97D4964" baseAuthorityName="Day" baseAuthorityYear="1868" box="[163,378,1103,1127]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="339" phylum="Chordata" rank="species" species="radiatus">Epinephelus radiatus</taxonomicName>
</emphasis>
</th>
<td id="DA78601DFF9600761113FCB7DB5A4964" box="[626,861,803,1127]" gridcol="1" gridrow="3" pageId="10" pageNumber="339">
<accessionNumber id="38CA96C0FF96FF831113FCB7DADD4E3F" box="[626,730,803,828]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/JQ681383" pageId="10" pageNumber="339">JQ681383</accessionNumber>
(GenBank) 
<accessionNumber id="38CA96C0FF96FF831113FCDADADD4E64" box="[626,730,846,871]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/JQ681405" pageId="10" pageNumber="339">JQ681405</accessionNumber>
(GenBank) 
<accessionNumber id="38CA96C0FF96FF831113FCEDDAE14E91" box="[626,742,889,914]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KM226295" pageId="10" pageNumber="339">KM226295</accessionNumber>
(GenBank) 
<accessionNumber id="38CA96C0FF96FF831113FC37DAE14EBF" box="[626,742,931,956]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KM226296" pageId="10" pageNumber="339">KM226296</accessionNumber>
(GenBank) 
<accessionNumber id="38CA96C0FF96FF831113FC5ADAE14EE4" box="[626,742,974,999]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KM226297" pageId="10" pageNumber="339">KM226297</accessionNumber>
(GenBank) 
<accessionNumber id="38CA96C0FF96FF831113FC6DDAD84911" box="[626,735,1017,1042]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KT835687" pageId="10" pageNumber="339">KT835687</accessionNumber>
(GenBank) 
<accessionNumber id="38CA96C0FF96FF831113FBB7DAD8493F" box="[626,735,1059,1084]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KT835688" pageId="10" pageNumber="339">KT835688</accessionNumber>
(GenBank) 
<accessionNumber id="38CA96C0FF96FF831113FBDADAE54964" box="[626,738,1102,1127]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KU943526" pageId="10" pageNumber="339">KU943526</accessionNumber>
(GenBank)
</td>
<td id="DA78601DFF96007610A7FCB7DCBE4964" box="[966,1209,803,1127]" gridcol="2" gridrow="3" pageId="10" pageNumber="339">China: South China Sea China: South China Sea India India India India India Taiwan</td>
</tr>
<tr id="99A90961FF96007613C2FB37DCBE483F" box="[163,1209,1187,1340]" gridrow="4" pageId="10" pageNumber="339">
<th id="DA78601DFF96007613C2FB37D9A0483F" box="[163,423,1187,1340]" gridcol="0" gridrow="4" pageId="10" pageNumber="339">
<emphasis id="15EDD731FF96FF8313C2FB30D960483F" italics="true" pageId="10" pageNumber="339">
<taxonomicName id="E09970A0FF96FF8313C2FB30D96049BF" authorityName="Randall &amp; Allen" authorityYear="1987" box="[163,359,1188,1212]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="339" phylum="Chordata" rank="species" species="stictus">Epinephelus stictus</taxonomicName>
<taxonomicName id="E09970A0FF96FF8313C2FB5BD96049E5" authorityName="Randall &amp; Allen" authorityYear="1987" box="[163,359,1231,1255]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="339" phylum="Chordata" rank="species" species="stictus">Epinephelus stictus</taxonomicName>
<taxonomicName id="E09970A0FF96FF8313C2FB6DD9604812" authorityName="Randall &amp; Allen" authorityYear="1987" box="[163,359,1273,1297]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="339" phylum="Chordata" rank="species" species="stictus">Epinephelus stictus</taxonomicName>
<taxonomicName id="E09970A0FF96FF8313C2FAB0D960483F" authorityName="Randall &amp; Allen" authorityYear="1987" box="[163,359,1316,1340]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="339" phylum="Chordata" rank="species" species="stictus">Epinephelus stictus</taxonomicName>
</emphasis>
</th>
<td id="DA78601DFF9600761113FB37DB5A483F" box="[626,861,1187,1340]" gridcol="1" gridrow="4" pageId="10" pageNumber="339">
<accessionNumber id="38CA96C0FF96FF831113FB37DAE049BF" box="[626,743,1187,1212]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MK693015" pageId="10" pageNumber="339">MK693015</accessionNumber>
(GenBank) 
<accessionNumber id="38CA96C0FF96FF831113FB5ADAE149E4" box="[626,742,1230,1255]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MK693016" pageId="10" pageNumber="339">MK693016</accessionNumber>
(GenBank) 
<accessionNumber id="38CA96C0FF96FF831113FB6DDAE14811" box="[626,742,1273,1298]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MK693017" pageId="10" pageNumber="339">MK693017</accessionNumber>
(GenBank) 
<accessionNumber id="38CA96C0FF96FF831113FAB7DAE1483F" box="[626,742,1315,1340]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MK693018" pageId="10" pageNumber="339">MK693018</accessionNumber>
(GenBank)
</td>
<td id="DA78601DFF96007610A7FB37DCBE483F" box="[966,1209,1187,1340]" gridcol="2" gridrow="4" pageId="10" pageNumber="339">Australia: Timor Sea Australia: Timor Sea Australia: Timor Sea Australia: Timor Sea</td>
</tr>
</table>
</paragraph>
<paragraph id="27260B23FF96FF8313A6FAF9DD154B16" blockId="10.[151,1437,1389,2014]" pageId="10" pageNumber="339">Body moderately deep, depth less than head length, proportionately greater with increasing length, 3.0 (3.1–3.4) in SL; body moderately compressed, greatest width 1.7 (1.6–1.8) in body depth; dorsal profile of head evenly convex, head length 2.5 (2.4–2.5) in SL; snout bluntly rounded, its length 3.9 (3.8–4.3) in HL; orbital diameter 5.5 (5.6–6.4) in HL; eyes directed laterally, interorbital space moderately convex, least width 6.0 (5.8–7.0) in HL; preorbital depth 9.2 (8.8–9.9) in HL; caudal-peduncle depth 3.7 (3.7–4.0) in HL; caudal-peduncle length 2.4 (2.5–2.6) in HL.</paragraph>
<paragraph id="27260B23FF96FF8313A6F989D9104ADD" blockId="10.[151,1437,1389,2014]" pageId="10" pageNumber="339">Mouth large, oblique, forming an angle of about 40° to the horizontal, lower jaw protruding beyond tip of upper jaw; upper jaw extending to vertical from just beyond posterior margin of pupil (just beyond posterior margin of pupil to just beyond posterior margin of eye), upper-jaw length 2.2 (2.2–2.4) in HL; posterior edge of maxilla straight (straight to slightly rounded), with narrowly rounded corners, maxilla width 8.4 (7.4–8.1) in HL. Pair of moderate canine teeth anteriorly in each jaw, those in upper jaw with gap 2.1 (1.5–2.0) times wider than gap in those in lower jaw. Teeth in lower jaw anteriorly in small patch 3–4 (3–5) teeth wide on either side of symphysis behind pair of more enlarged canines, followed posteriorly on each side by 2 rows of subequal teeth extending to rear of jaw; upper jaw with larger anterior tooth patch in 3–4 (3–5) rows, followed posteriorly by single row of enlarged teeth distally and about 3 rows of much smaller teeth mesially. Vomer v-shaped, with 2 (2–3) rows of villiform teeth; palatines narrow, with 3 (2–4) rows of villiform teeth. Tongue large, with broadly rounded tip. Longest gill raker at angle, slightly shorter than (subequal to shorter than) longest gill filament. Nostrils round, posterior nostril largest, 1.6 (1.4–1.6) times width of anterior nostril. Anterior nostril with membranous flap, apex of flap over posterior portion of rim.</paragraph>
<paragraph id="27260B23FF97FF8213A6FF0DD8F64C64" blockId="11.[151,1437,153,2015]" pageId="11" pageNumber="340">Opercle with three flat spines; uppermost small, barely exposed from skin and scales, middle and lower spines much larger and distinctly exposed, middle spine longest, equidistant between upper and lower spines, tip extending further posteriorly than other spines. Upper edge of opercular membrane slightly convex, forming rounded point posteriorly. Preopercle broadly rounded at angle, with 8 small (4–9 small to moderate), non-prominent serrae, lower edge of preopercle smooth, hind edge with numerous minute serrae. Margins of subopercle and interopercle smooth.</paragraph>
<paragraph id="27260B23FF97FF8213A6FEE5D9094F18" blockId="11.[151,1437,153,2015]" pageId="11" pageNumber="340">Lateral line slightly arched over pectoral region, then becoming parallel to contour of back and straightening on peduncle. Scales cycloid on head, thorax, pectoral-fin base, abdomen and anterodorsally on body; ctenoid elsewhere on body. No auxiliary scales on body and no scales on maxilla. Minute scales on basal third of pectoral fins, basal outer half of two inner rays of pelvic fins, basal two-thirds of soft dorsal and anal fins and basal three-fourths of caudal fin.</paragraph>
<paragraph id="27260B23FF97FF8213A6FDB1DB464FF0" blockId="11.[151,1437,153,2015]" pageId="11" pageNumber="340">Origin of dorsal fin above vertical from axil of pectoral fin (axil of pectoral fin to midway between axil of pectoral fin and upper corner of opercle); predorsal length 2.7 (2.6–2.8) in SL; membranes of spinous dorsal fin deeply incised; dorsal-fin spines progressively longer to fourth spine, then gradually decreasing in length to penultimate; first spine about half length of second, 8.4 (7.9–9.9) in HL; fourth spine longest, 3.7 (3.3–4.0) in HL; last dorsal-fin spine longer than penultimate, 5.5 (4.8–5.3) in HL; fifth (fifth to seventh) soft dorsal-fin ray longest, 3.1 (3.0–3.2) in HL. All dorsal and anal-fin rays branched, the last to base.</paragraph>
<paragraph id="27260B23FF97FF8213A6FD69D96F4934" blockId="11.[151,1437,153,2015]" pageId="11" pageNumber="340">Origin of anal fin below base of second (first or second) soft dorsal-fin ray, preanal length 1.4 (1.4–1.5) in SL; first anal-fin spine short, 1.9 (1.9–2.2) in length of second spine; second anal-fin spine slightly shorter (usually slightly shorter) than third spine, 1.2 (1.0–1.2) in length of third spine, third anal-fin spine 4.8 (4.6–5.1) in HL; third soft anal-fin ray longest, length 2.8 (2.8–3.0) in HL. Caudal fin rounded, length 5.5 (5.1–5.4) in SL. Pectoral fins rounded, eighth (seventh or eighth) ray longest, 1.9 (1.7–2.0) in SL, rays branched except short uppermost and lowermost rays. Origin of pelvic fins below vertical from upper base of pectoral fins, prepelvic length 2.7 (2.7–2.9) in SL; pelvic-fin spine reaching slightly more than half distance to tip of first ray; fourth soft pelvic-fin ray longest, much shorter than longest pectoral-fin ray, length 2.5 (2.4–2.6) in HL; first pelvic-fin ray firmly attached by membrane to abdomen.</paragraph>
<paragraph id="27260B23FF97FF8213F6FBD5DB374B50" blockId="11.[151,1437,153,2015]" pageId="11" pageNumber="340">
<emphasis id="15EDD731FF97FF8213F6FBD5D96B4958" box="[151,364,1089,1115]" italics="true" pageId="11" pageNumber="340">Colour when fresh.</emphasis>
Based on fresh specimens, ground colour of upper head and body in 
<typeStatus id="F822B581FF97FF821700FBD5DCC44958" box="[1121,1219,1089,1115]" pageId="11" pageNumber="340" type="holotype">holotype</typeStatus>
(fig. 2B) and larger 
<typeStatus id="F822B581FF97FF8213F6FBF1D905497C" box="[151,258,1125,1151]" pageId="11" pageNumber="340" type="paratype">paratypes</typeStatus>
uniformly dark brown; head below preopercles, thorax and abdomen pale pinkish brown. Faint broad darker brown diagonal bar from rear margin of eye to edge of opercle at third opercular spine. Smallest 
<typeStatus id="F822B581FF97FF82165DFB1DDD9B49A0" box="[1340,1436,1161,1187]" pageId="11" pageNumber="340" type="paratype">paratype</typeStatus>
(fig. 4A) with bar more distinct and with slightly narrower second bar from lower margin of eye crossing middle of preopercle to lower edge of opercle. Smallest 
<typeStatus id="F822B581FF97FF8211FDFB45DAFB49E8" box="[668,764,1233,1259]" pageId="11" pageNumber="340" type="paratype">paratype</typeStatus>
also with series of broad irregular diffuse wavy brown bars and blotches on sides of body, two longitudinally on upper body extending from upper opercle to below middle of soft dorsal fin and another two transversely between soft dorsal and anal fins. Spinous dorsal-fin membrane pale grey-brown in basal half, diffuse darker grey area medially, followed above adjacent to rear of each spine by dark yellowish brown pigmentation, portion of membrane posteroventral to darkly pigmented area pale and semitransparent (fig. 3A). Basal two-thirds of soft dorsal fin pale grey-brown, distal third with dark olive to honey-brown membrane (distal third dark brown in smallest 
<typeStatus id="F822B581FF97FF8211CEFA3DDB1548C0" box="[687,786,1449,1475]" pageId="11" pageNumber="340" type="paratype">paratype</typeStatus>
, see fig. 3B), contrasting with irregular submarginal pale semitransparent sections to individual rays. Basal half of anal fin pale grey, distal half dusky (fig. 3C). Caudal fin pale grey-brown on basal two-thirds, distal two-thirds diffusely dark olive-brown (fig. 3D), dark grey-brown in smallest 
<typeStatus id="F822B581FF97FF82139BF981D9594B2C" box="[250,350,1557,1583]" pageId="11" pageNumber="340" type="paratype">paratype</typeStatus>
. Pectoral fin rays faintly brownish, membrane semitranslucent. Pelvic fins mostly pale and semitranslucent, distal half faintly dusky, leading edges whitish.
</paragraph>
<paragraph id="27260B23FF97FF8213A6F9C9DB2B4A4C" blockId="11.[151,1437,153,2015]" pageId="11" pageNumber="340">
<emphasis id="15EDD731FF97FF8213A6F9C9D9934B74" box="[199,404,1629,1655]" italics="true" pageId="11" pageNumber="340">Colour in alcohol.</emphasis>
Head and body of 
<typeStatus id="F822B581FF97FF821104F9C9DAC04B74" box="[613,711,1629,1655]" pageId="11" pageNumber="340" type="holotype">holotype</typeStatus>
(fig. 2A) and largest two 
<typeStatus id="F822B581FF97FF821085F9C9DC484B74" box="[996,1103,1629,1655]" pageId="11" pageNumber="340" type="paratype">paratypes</typeStatus>
(
<quantity id="E061A6C6FF97FF82173CF9C8DCF44B74" box="[1117,1267,1628,1655]" metricMagnitude="-1" metricUnit="m" metricValue="5.15" metricValueMax="5.279999999999999" metricValueMin="5.02" pageId="11" pageNumber="340" unit="mm" value="515.0" valueMax="528.0" valueMin="502.0">502–528 mm</quantity>
SL) dark greyish brown. Smallest 
<typeStatus id="F822B581FF97FF8212E0F915D9E64B98" box="[385,481,1665,1691]" pageId="11" pageNumber="340" type="paratype">paratype</typeStatus>
(
<quantity id="E061A6C6FF97FF821293F915DA554B98" box="[498,594,1665,1691]" metricMagnitude="-1" metricUnit="m" metricValue="4.08" pageId="11" pageNumber="340" unit="mm" value="408.0">408 mm</quantity>
SL) with head and body more pale brown and including two faint darker brown bars radiating diagonally from rear of eye to margin of opercle and a series of irregular broad diffuse brown wavy bars and blotches on the sides of the body. Second smallest 
<typeStatus id="F822B581FF97FF8210E7F95DDBE14BE0" box="[902,998,1737,1763]" pageId="11" pageNumber="340" type="paratype">paratype</typeStatus>
(
<quantity id="E061A6C6FF97FF821097F95CDC514BE0" box="[1014,1110,1736,1763]" metricMagnitude="-1" metricUnit="m" metricValue="4.56" pageId="11" pageNumber="340" unit="mm" value="456.0">456 mm</quantity>
SL) with bars on head faint, those on body barely discernible. Spinous dorsal fin membrane pale greyish brown below, an irregular darker grey cross band in midsection, and distal triangular portion semitranslucent. Soft dorsal, anal and caudal fins with irregular broad dusky margins. Paired fins pale yellowish brown.
</paragraph>
<paragraph id="27260B23FF97FF8213A6F8CDD9844ADC" blockId="11.[151,1437,153,2015]" pageId="11" pageNumber="340">
<emphasis id="15EDD731FF97FF8213A6F8CDD9A74A70" bold="true" box="[199,416,1881,1907]" pageId="11" pageNumber="340">Molecular results.</emphasis>
The phylogenetic analyses indicate that 
<taxonomicName id="E09970A0FF97FF82100CF8CEDC404A70" authorityName="Johnson &amp; Wilmer" authorityYear="2019" box="[877,1095,1881,1907]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="340" phylum="Chordata" rank="species" species="fuscomarginatus">
<emphasis id="15EDD731FF97FF82100CF8CEDC404A70" box="[877,1095,1881,1907]" italics="true" pageId="11" pageNumber="340">E. fuscomarginatus</emphasis>
</taxonomicName>
is most closely related to, but distinct from 
<taxonomicName id="E09970A0FF97FF82124AF8EAD9D04A94" box="[299,471,1918,1943]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="340" phylum="Chordata" rank="species" species="magniscuttis">
<emphasis id="15EDD731FF97FF82124AF8EAD9D04A94" box="[299,471,1918,1943]" italics="true" pageId="11" pageNumber="340">E. magniscuttis</emphasis>
</taxonomicName>
, 
<taxonomicName id="E09970A0FF97FF821282F8EADA6C4A94" authorityName="in Heemstra &amp; Randall" authorityYear="1993" box="[483,619,1918,1943]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="340" phylum="Chordata" rank="species" species="epistictus">
<emphasis id="15EDD731FF97FF821282F8EADA6C4A94" box="[483,619,1918,1943]" italics="true" pageId="11" pageNumber="340">E. epistictus</emphasis>
</taxonomicName>
and 
<emphasis id="15EDD731FF97FF8211FEF8EADAB04A94" box="[671,695,1918,1943]" italics="true" pageId="11" pageNumber="340">E.</emphasis>
cf 
<taxonomicName id="E09970A0FF97FF8211B9F8EADB454A94" authorityName="in Heemstra &amp; Randall" authorityYear="1993" box="[728,834,1918,1943]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="340" phylum="Chordata" rank="species" species="epistictus">
<emphasis id="15EDD731FF97FF8211B9F8EADB454A94" box="[728,834,1918,1943]" italics="true" pageId="11" pageNumber="340">epistictus</emphasis>
</taxonomicName>
, with average genetic sequence divergences of 1.10%, 2.70% and 3.80%, respectively. There was no intraspecific divergence among the five samples of 
<taxonomicName id="E09970A0FF97FF821789F836D8C14ADC" authorityName="Johnson &amp; Wilmer" authorityYear="2019" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="340" phylum="Chordata" rank="species" species="fuscomarginatus">
<emphasis id="15EDD731FF97FF821789F836D8C14ADC" italics="true" pageId="11" pageNumber="340">E. fuscomarginatus</emphasis>
</taxonomicName>
(fig. 1; 
<tableCitation id="6A1B3E98FF97FF821241F851D9724ADC" box="[288,373,1989,2015]" captionStart="TABLE 2" captionStartId="12.[151,244,152,179]" captionTargetBox="[163,1406,245,1097]" captionText="TABLE 2. Estimates of mean sequence divergence within (on diagonal) and between (below diagonal) species." httpUri="http://table.plazi.org/id/73E65BABFF90FF8513F6FF0CDD7E4DB0" pageId="11" pageNumber="340" tableUuid="73E65BABFF90FF8513F6FF0CDD7E4DB0">Table 2</tableCitation>
).
</paragraph>
</subSubSection>
<caption id="73E65BABFF90FF8513F6FF0CDD7E4DB0" ID-Table-UUID="73E65BABFF90FF8513F6FF0CDD7E4DB0" box="[151,1401,152,180]" httpUri="http://table.plazi.org/id/73E65BABFF90FF8513F6FF0CDD7E4DB0" pageId="12" pageNumber="341" startId="12.[151,244,152,179]" targetBox="[163,1406,245,1097]" targetIsTable="true" targetPageId="12">
<paragraph id="27260B23FF90FF8513F6FF0CDD7E4DB0" blockId="12.[151,1401,152,180]" box="[151,1401,152,180]" pageId="12" pageNumber="341">
<emphasis id="15EDD731FF90FF8513F6FF0CD9174DB0" bold="true" box="[151,272,152,179]" pageId="12" pageNumber="341">TABLE 2.</emphasis>
Estimates of mean sequence divergence within (on diagonal) and between (below diagonal) species.
</paragraph>
</caption>
<paragraph id="27260B23FF90FF8511C3FEBEDD714946" pageId="12" pageNumber="341">
<table id="5599F983FF90007613C2FF61DD79494A" box="[163,1406,245,1097]" gridcols="12" gridrows="13" pageId="12" pageNumber="341">
<tr id="99A90961FF90007613C2FF61DD794C40" box="[163,1406,245,323]" gridrow="0" pageId="12" pageNumber="341" rowspan-0="1" rowspan-1="1" rowspan-10="1" rowspan-11="1" rowspan-2="1" rowspan-3="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<th id="DA78601DFF90007611E4FF61DACC4C40" box="[645,715,245,323]" gridcol="4" gridrow="0" pageId="12" pageNumber="341">sp. nov.</th>
</tr>
<tr id="99A90961FF90007613C2FEDEDD794F0D" box="[163,1406,330,526]" gridrow="1" pageId="12" pageNumber="341" rowspan-0="1">
<td id="DA78601DFF9000761208FEDED9A84F0D" box="[361,431,330,526]" gridcol="1" gridrow="1" pageId="12" pageNumber="341">
<emphasis id="15EDD731FF90FF8512E7FE6FD9984CF8" box="[390,415,504,526]" italics="true" pageId="12" pageNumber="341">E.</emphasis>
cf 
<taxonomicName id="E09970A0FF90FF8512E7FEECD9984CDB" authorityName="in Heemstra &amp; Randall" authorityYear="1993" box="[390,415,376,472]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="341" phylum="Chordata" rank="species" species="epistictus">
<emphasis id="15EDD731FF90FF8512E7FEECD9984CDB" box="[390,415,376,472]" italics="true" pageId="12" pageNumber="341">epistictus</emphasis>
</taxonomicName>
</td>
<td id="DA78601DFF90007612A8FEDEDA084F0D" box="[457,527,330,526]" gridcol="2" gridrow="1" pageId="12" pageNumber="341">
<taxonomicName id="E09970A0FF90FF851285FE6FD9FA4CF1" authorityName="in Heemstra &amp; Randall" authorityYear="1993" box="[484,509,402,526]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="341" phylum="Chordata" rank="species" species="epistictus">
<emphasis id="15EDD731FF90FF851285FE6FD9FA4CF1" box="[484,509,402,526]" italics="true" pageId="12" pageNumber="341">E. epistictus</emphasis>
</taxonomicName>
</td>
<td id="DA78601DFF9000761147FEDEDA6C4F0D" box="[550,619,330,526]" gridcol="3" gridrow="1" pageId="12" pageNumber="341">
<taxonomicName id="E09970A0FF90FF851123FE6FDA5C4CF1" box="[578,603,370,526]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="341" phylum="Chordata" rank="species" species="magniscuttis">
<emphasis id="15EDD731FF90FF851123FE6FDA5C4CF1" box="[578,603,370,526]" italics="true" pageId="12" pageNumber="341">E. magniscuttis</emphasis>
</taxonomicName>
</td>
<td id="DA78601DFF90007611E4FEDEDACC4F0D" box="[645,715,330,526]" gridcol="4" gridrow="1" pageId="12" pageNumber="341">
<taxonomicName id="E09970A0FF90FF8511C3FE6FDABC4CF1" authorityName="Johnson &amp; Wilmer" authorityYear="2019" box="[674,699,330,526]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="341" phylum="Chordata" rank="species" species="fuscomarginatus">
<emphasis id="15EDD731FF90FF8511C3FE6FDABC4CF1" box="[674,699,330,526]" italics="true" pageId="12" pageNumber="341">E. fuscomarginatus</emphasis>
</taxonomicName>
</td>
<td id="DA78601DFF9000761184FEDEDB2C4F0D" box="[741,811,330,526]" gridcol="5" gridrow="1" pageId="12" pageNumber="341">
<taxonomicName id="E09970A0FF90FF851060FE6FDB1D4CF1" baseAuthorityName="non Fowler" baseAuthorityYear="1904" box="[769,794,395,526]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="341" phylum="Chordata" rank="species" species="heniochus">
<emphasis id="15EDD731FF90FF851060FE6FDB1D4CF1" box="[769,794,395,526]" italics="true" pageId="12" pageNumber="341">E. heniochus</emphasis>
</taxonomicName>
</td>
<td id="DA78601DFF9000761025FEDEDB8D4F0D" box="[836,906,330,526]" gridcol="6" gridrow="1" pageId="12" pageNumber="341">
<taxonomicName id="E09970A0FF90FF85103EFE6FDB7F4CF2" baseAuthorityName="Valenciennes" baseAuthorityYear="1833" box="[863,888,409,526]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="341" phylum="Chordata" rank="species" species="morrhua">
<emphasis id="15EDD731FF90FF85103EFE6FDB7F4CF2" box="[863,888,409,526]" italics="true" pageId="12" pageNumber="341">E. morrhua</emphasis>
</taxonomicName>
</td>
<td id="DA78601DFF90007610C3FEDEDBEF4F0D" box="[930,1000,330,526]" gridcol="7" gridrow="1" pageId="12" pageNumber="341">
<taxonomicName id="E09970A0FF90FF8510A7FE6FDBD84CF2" baseAuthorityName="Temminck &amp; Schlegel" baseAuthorityYear="1842" box="[966,991,368,526]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="341" phylum="Chordata" rank="species" species="poecilonotus">
<emphasis id="15EDD731FF90FF8510A7FE6FDBD84CF2" box="[966,991,368,526]" italics="true" pageId="12" pageNumber="341">E. poecilonotus</emphasis>
</taxonomicName>
</td>
<td id="DA78601DFF9000761770FEDEDC504F0D" box="[1041,1111,330,526]" gridcol="8" gridrow="1" pageId="12" pageNumber="341">
<taxonomicName id="E09970A0FF90FF851755FE6FDC4A4CF2" baseAuthorityName="Day" baseAuthorityYear="1868" box="[1076,1101,413,526]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="341" phylum="Chordata" rank="species" species="radiatus">
<emphasis id="15EDD731FF90FF851755FE6FDC4A4CF2" box="[1076,1101,413,526]" italics="true" pageId="12" pageNumber="341">E. radiatus</emphasis>
</taxonomicName>
</td>
<td id="DA78601DFF900076171CFEDEDCC34F0D" box="[1149,1220,330,526]" gridcol="9" gridrow="1" pageId="12" pageNumber="341">
<taxonomicName id="E09970A0FF90FF85171FFE6FDC904CF1" baseAuthorityName="Temminck &amp; Schlegel" baseAuthorityYear="1843" box="[1150,1175,374,526]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="341" phylum="Chordata" rank="species" species="latifasciatus">
<emphasis id="15EDD731FF90FF85171FFE6FDC904CF1" box="[1150,1175,374,526]" italics="true" pageId="12" pageNumber="341">E. latifasciatus</emphasis>
</taxonomicName>
</td>
<td id="DA78601DFF900076178DFEDEDD344F0D" box="[1260,1331,330,526]" gridcol="10" gridrow="1" pageId="12" pageNumber="341">
<taxonomicName id="E09970A0FF90FF85166BFE6FDD244CF1" authorityName="Randall &amp; Allen" authorityYear="1987" box="[1290,1315,433,526]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="341" phylum="Chordata" rank="species" species="stictus">
<emphasis id="15EDD731FF90FF85166BFE6FDD244CF1" box="[1290,1315,433,526]" italics="true" pageId="12" pageNumber="341">E. stictus</emphasis>
</taxonomicName>
</td>
<td id="DA78601DFF900076162EFEDEDD794F0D" box="[1359,1406,330,526]" gridcol="11" gridrow="1" pageId="12" pageNumber="341">
<taxonomicName id="E09970A0FF90FF851604FE6FDD794CF1" baseAuthorityName="Bleeker" baseAuthorityYear="1855" box="[1381,1406,399,526]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="341" phylum="Chordata" rank="species" species="bontoides">
<emphasis id="15EDD731FF90FF851604FE6FDD794CF1" box="[1381,1406,399,526]" italics="true" pageId="12" pageNumber="341">E. bontoides</emphasis>
</taxonomicName>
</td>
</tr>
<tr id="99A90961FF90007613C2FDBCDD794F42" box="[163,1406,552,577]" gridrow="2" pageId="12" pageNumber="341" rowspan-10="1" rowspan-11="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<th id="DA78601DFF90007613C2FDBCD94C4F42" box="[163,331,552,577]" gridcol="0" gridrow="2" pageId="12" pageNumber="341">
<emphasis id="15EDD731FF90FF8513C2FDBED8BE4F42" box="[163,185,554,577]" italics="true" pageId="12" pageNumber="341">E.</emphasis>
cf 
<taxonomicName id="E09970A0FF90FF8513B8FDBED93E4F42" authorityName="in Heemstra &amp; Randall" authorityYear="1993" box="[217,313,554,577]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="341" phylum="Chordata" rank="species" species="epistictus">
<emphasis id="15EDD731FF90FF8513B8FDBED93E4F42" box="[217,313,554,577]" italics="true" pageId="12" pageNumber="341">epistictus</emphasis>
</taxonomicName>
</th>
<td id="DA78601DFF9000761208FDBCD9A84F42" box="[361,431,552,577]" gridcol="1" gridrow="2" pageId="12" pageNumber="341">0.20%</td>
</tr>
<tr id="99A90961FF90007613C2FDC3DD794F6C" box="[163,1406,599,623]" gridrow="3" pageId="12" pageNumber="341" rowspan-10="1" rowspan-11="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<th id="DA78601DFF90007613C2FDC3D94C4F6C" box="[163,331,599,623]" gridcol="0" gridrow="3" pageId="12" pageNumber="341">
<taxonomicName id="E09970A0FF90FF8513C2FDCCD9184F6C" authorityName="in Heemstra &amp; Randall" authorityYear="1993" box="[163,287,600,623]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="341" phylum="Chordata" rank="species" species="epistictus">
<emphasis id="15EDD731FF90FF8513C2FDCCD9184F6C" box="[163,287,600,623]" italics="true" pageId="12" pageNumber="341">E. epistictus</emphasis>
</taxonomicName>
</th>
<td id="DA78601DFF9000761208FDC3D9A84F6C" box="[361,431,599,623]" gridcol="1" gridrow="3" pageId="12" pageNumber="341">3.70%</td>
<td id="DA78601DFF90007612A8FDC3DA084F6C" box="[457,527,599,623]" gridcol="2" gridrow="3" pageId="12" pageNumber="341">0.10%</td>
</tr>
<tr id="99A90961FF90007613C2FD11DD794F9E" box="[163,1406,645,669]" gridrow="4" pageId="12" pageNumber="341" rowspan-10="1" rowspan-11="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<th id="DA78601DFF90007613C2FD11D94C4F9E" box="[163,331,645,669]" gridcol="0" gridrow="4" pageId="12" pageNumber="341">
<taxonomicName id="E09970A0FF90FF8513C2FD12D9384F9E" box="[163,319,646,669]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="341" phylum="Chordata" rank="species" species="magniscuttis">
<emphasis id="15EDD731FF90FF8513C2FD12D9384F9E" box="[163,319,646,669]" italics="true" pageId="12" pageNumber="341">E. magniscuttis</emphasis>
</taxonomicName>
</th>
<td id="DA78601DFF9000761208FD11D9A84F9E" box="[361,431,645,669]" gridcol="1" gridrow="4" pageId="12" pageNumber="341">3.70%</td>
<td id="DA78601DFF90007612A8FD11DA084F9E" box="[457,527,645,669]" gridcol="2" gridrow="4" pageId="12" pageNumber="341">2.40%</td>
<td id="DA78601DFF9000761147FD11DA6C4F9E" box="[550,619,645,669]" gridcol="3" gridrow="4" pageId="12" pageNumber="341">0.20%</td>
</tr>
<tr id="99A90961FF90007613C2FD25DD794E01" box="[163,1406,689,770]" gridrow="5" pageId="12" pageNumber="341" rowspan-10="1" rowspan-11="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<th id="DA78601DFF90007613C2FD25D94C4E01" box="[163,331,689,770]" gridcol="0" gridrow="5" pageId="12" pageNumber="341">
<taxonomicName id="E09970A0FF90FF8513C2FD25D94C4FE6" authority="Johnson &amp; Wilmer, 2019" authorityName="Johnson &amp; Wilmer" authorityYear="2019" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="341" phylum="Chordata" rank="species" species="fuscomarginatus" status="sp. nov.">
<emphasis id="15EDD731FF90FF8513C2FD25D94C4FE6" italics="true" pageId="12" pageNumber="341">E. fuscomarginatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0EDE6A4AFF90FF8513C8FD7FD8FF4E01" box="[169,248,747,770]" pageId="12" pageNumber="341" rank="species">sp. nov.</taxonomicNameLabel>
</th>
<td id="DA78601DFF9000761208FD25D9A84E01" box="[361,431,689,770]" gridcol="1" gridrow="5" pageId="12" pageNumber="341">3.80%</td>
<td id="DA78601DFF90007612A8FD25DA084E01" box="[457,527,689,770]" gridcol="2" gridrow="5" pageId="12" pageNumber="341">2.70%</td>
<td id="DA78601DFF9000761147FD25DA6C4E01" box="[550,619,689,770]" gridcol="3" gridrow="5" pageId="12" pageNumber="341">1.10%</td>
<td id="DA78601DFF90007611E4FD25DACC4E01" box="[645,715,689,770]" gridcol="4" gridrow="5" pageId="12" pageNumber="341">0.00%</td>
</tr>
<tr id="99A90961FF90007613C2FC82DD794E2D" box="[163,1406,790,814]" gridrow="6" pageId="12" pageNumber="341" rowspan-10="1" rowspan-11="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<th id="DA78601DFF90007613C2FC82D94C4E2D" box="[163,331,790,814]" gridcol="0" gridrow="6" pageId="12" pageNumber="341">
<taxonomicName id="E09970A0FF90FF8513C2FC83D9214E2D" baseAuthorityName="non Fowler" baseAuthorityYear="1904" box="[163,294,790,814]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="341" phylum="Chordata" rank="species" species="heniochus">
<emphasis id="15EDD731FF90FF8513C2FC83D9214E2D" box="[163,294,790,814]" italics="true" pageId="12" pageNumber="341">E. heniochus</emphasis>
</taxonomicName>
</th>
<td id="DA78601DFF9000761208FC82D9A84E2D" box="[361,431,790,814]" gridcol="1" gridrow="6" pageId="12" pageNumber="341">7.00%</td>
<td id="DA78601DFF90007612A8FC82DA084E2D" box="[457,527,790,814]" gridcol="2" gridrow="6" pageId="12" pageNumber="341">7.30%</td>
<td id="DA78601DFF9000761147FC82DA6C4E2D" box="[550,619,790,814]" gridcol="3" gridrow="6" pageId="12" pageNumber="341">6.70%</td>
<td id="DA78601DFF90007611E4FC82DACC4E2D" box="[645,715,790,814]" gridcol="4" gridrow="6" pageId="12" pageNumber="341">6.50%</td>
<td id="DA78601DFF9000761184FC82DB2C4E2D" box="[741,811,790,814]" gridcol="5" gridrow="6" pageId="12" pageNumber="341">0.20%</td>
</tr>
<tr id="99A90961FF90007613C2FCD0DD794E5F" box="[163,1406,836,860]" gridrow="7" pageId="12" pageNumber="341" rowspan-10="1" rowspan-11="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<th id="DA78601DFF90007613C2FCD0D94C4E5F" box="[163,331,836,860]" gridcol="0" gridrow="7" pageId="12" pageNumber="341">
<taxonomicName id="E09970A0FF90FF8513C2FCD1D9104E5F" baseAuthorityName="Valenciennes" baseAuthorityYear="1833" box="[163,279,836,860]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="341" phylum="Chordata" rank="species" species="morrhua">
<emphasis id="15EDD731FF90FF8513C2FCD1D9104E5F" box="[163,279,836,860]" italics="true" pageId="12" pageNumber="341">E. morrhua</emphasis>
</taxonomicName>
</th>
<td id="DA78601DFF9000761208FCD0D9A84E5F" box="[361,431,836,860]" gridcol="1" gridrow="7" pageId="12" pageNumber="341">5.10%</td>
<td id="DA78601DFF90007612A8FCD0DA084E5F" box="[457,527,836,860]" gridcol="2" gridrow="7" pageId="12" pageNumber="341">5.70%</td>
<td id="DA78601DFF9000761147FCD0DA6C4E5F" box="[550,619,836,860]" gridcol="3" gridrow="7" pageId="12" pageNumber="341">6.10%</td>
<td id="DA78601DFF90007611E4FCD0DACC4E5F" box="[645,715,836,860]" gridcol="4" gridrow="7" pageId="12" pageNumber="341">6.20%</td>
<td id="DA78601DFF9000761184FCD0DB2C4E5F" box="[741,811,836,860]" gridcol="5" gridrow="7" pageId="12" pageNumber="341">7.80%</td>
<td id="DA78601DFF9000761025FCD0DB8D4E5F" box="[836,906,836,860]" gridcol="6" gridrow="7" pageId="12" pageNumber="341">0.90%</td>
</tr>
<tr id="99A90961FF90007613C2FCE6DD794E89" box="[163,1406,882,906]" gridrow="8" pageId="12" pageNumber="341" rowspan-10="1" rowspan-11="1" rowspan-8="1" rowspan-9="1">
<th id="DA78601DFF90007613C2FCE6D94C4E89" box="[163,331,882,906]" gridcol="0" gridrow="8" pageId="12" pageNumber="341">
<taxonomicName id="E09970A0FF90FF8513C2FCE7D9474E89" baseAuthorityName="Temminck &amp; Schlegel" baseAuthorityYear="1842" box="[163,320,882,906]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="341" phylum="Chordata" rank="species" species="poecilonotus">
<emphasis id="15EDD731FF90FF8513C2FCE7D9474E89" box="[163,320,882,906]" italics="true" pageId="12" pageNumber="341">E. poecilonotus</emphasis>
</taxonomicName>
</th>
<td id="DA78601DFF9000761208FCE6D9A84E89" box="[361,431,882,906]" gridcol="1" gridrow="8" pageId="12" pageNumber="341">4.30%</td>
<td id="DA78601DFF90007612A8FCE6DA084E89" box="[457,527,882,906]" gridcol="2" gridrow="8" pageId="12" pageNumber="341">5.00%</td>
<td id="DA78601DFF9000761147FCE6DA6C4E89" box="[550,619,882,906]" gridcol="3" gridrow="8" pageId="12" pageNumber="341">5.00%</td>
<td id="DA78601DFF90007611E4FCE6DACC4E89" box="[645,715,882,906]" gridcol="4" gridrow="8" pageId="12" pageNumber="341">5.50%</td>
<td id="DA78601DFF9000761184FCE6DB2C4E89" box="[741,811,882,906]" gridcol="5" gridrow="8" pageId="12" pageNumber="341">8.50%</td>
<td id="DA78601DFF9000761025FCE6DB8D4E89" box="[836,906,882,906]" gridcol="6" gridrow="8" pageId="12" pageNumber="341">4.00%</td>
<td id="DA78601DFF90007610C3FCE6DBEF4E89" box="[930,1000,882,906]" gridcol="7" gridrow="8" pageId="12" pageNumber="341">0.50%</td>
</tr>
<tr id="99A90961FF90007613C2FC37DD794EB8" box="[163,1406,931,955]" gridrow="9" pageId="12" pageNumber="341" rowspan-10="1" rowspan-11="1" rowspan-9="1">
<th id="DA78601DFF90007613C2FC37D94C4EB8" box="[163,331,931,955]" gridcol="0" gridrow="9" pageId="12" pageNumber="341">
<taxonomicName id="E09970A0FF90FF8513C2FC37D9144EB8" baseAuthorityName="Day" baseAuthorityYear="1868" box="[163,275,931,955]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="341" phylum="Chordata" rank="species" species="radiatus">
<emphasis id="15EDD731FF90FF8513C2FC37D9144EB8" box="[163,275,931,955]" italics="true" pageId="12" pageNumber="341">E. radiatus</emphasis>
</taxonomicName>
</th>
<td id="DA78601DFF9000761208FC37D9A84EB8" box="[361,431,931,955]" gridcol="1" gridrow="9" pageId="12" pageNumber="341">4.40%</td>
<td id="DA78601DFF90007612A8FC37DA084EB8" box="[457,527,931,955]" gridcol="2" gridrow="9" pageId="12" pageNumber="341">4.30%</td>
<td id="DA78601DFF9000761147FC37DA6C4EB8" box="[550,619,931,955]" gridcol="3" gridrow="9" pageId="12" pageNumber="341">4.70%</td>
<td id="DA78601DFF90007611E4FC37DACC4EB8" box="[645,715,931,955]" gridcol="4" gridrow="9" pageId="12" pageNumber="341">4.60%</td>
<td id="DA78601DFF9000761184FC37DB2C4EB8" box="[741,811,931,955]" gridcol="5" gridrow="9" pageId="12" pageNumber="341">8.10%</td>
<td id="DA78601DFF9000761025FC37DB8D4EB8" box="[836,906,931,955]" gridcol="6" gridrow="9" pageId="12" pageNumber="341">4.00%</td>
<td id="DA78601DFF90007610C3FC37DBEF4EB8" box="[930,1000,931,955]" gridcol="7" gridrow="9" pageId="12" pageNumber="341">2.70%</td>
<td id="DA78601DFF9000761770FC37DC504EB8" box="[1041,1111,931,955]" gridcol="8" gridrow="9" pageId="12" pageNumber="341">1.40%</td>
</tr>
<tr id="99A90961FF90007613C2FC44DD794EEE" box="[163,1406,976,1005]" gridrow="10" pageId="12" pageNumber="341" rowspan-10="1" rowspan-11="1">
<th id="DA78601DFF90007613C2FC44D94C4EEE" box="[163,331,976,1005]" gridcol="0" gridrow="10" pageId="12" pageNumber="341">
<taxonomicName id="E09970A0FF90FF8513C2FC40D93C4EE8" baseAuthorityName="Temminck &amp; Schlegel" baseAuthorityYear="1843" box="[163,315,979,1003]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="341" phylum="Chordata" rank="species" species="latifasciatus">
<emphasis id="15EDD731FF90FF8513C2FC40D93C4EE8" box="[163,315,979,1003]" italics="true" pageId="12" pageNumber="341">E. latifasciatus</emphasis>
</taxonomicName>
</th>
<td id="DA78601DFF9000761208FC44D9A84EEE" box="[361,431,976,1005]" gridcol="1" gridrow="10" pageId="12" pageNumber="341">12.60%</td>
<td id="DA78601DFF90007612A8FC44DA084EEE" box="[457,527,976,1005]" gridcol="2" gridrow="10" pageId="12" pageNumber="341">13.40%</td>
<td id="DA78601DFF9000761147FC44DACC4EEE" box="[550,715,976,1005]" colspan="2" colspanRight="1" gridcol="3" gridrow="10" pageId="12" pageNumber="341">12.90% 12.60%</td>
<td id="DA78601DFF9000761184FC44DB2C4EEE" box="[741,811,976,1005]" gridcol="5" gridrow="10" pageId="12" pageNumber="341">13.70%</td>
<td id="DA78601DFF9000761025FC44DB8D4EEE" box="[836,906,976,1005]" gridcol="6" gridrow="10" pageId="12" pageNumber="341">11.60%</td>
<td id="DA78601DFF90007610C3FC44DBEF4EEE" box="[930,1000,976,1005]" gridcol="7" gridrow="10" pageId="12" pageNumber="341">12.10%</td>
<td id="DA78601DFF9000761770FC44DC504EEE" box="[1041,1111,976,1005]" gridcol="8" gridrow="10" pageId="12" pageNumber="341">11.60%</td>
<td id="DA78601DFF900076171CFC44DCC34EEE" box="[1149,1220,976,1005]" gridcol="9" gridrow="10" pageId="12" pageNumber="341">0.80%</td>
</tr>
<tr id="99A90961FF90007613C2FB95DD794918" box="[163,1406,1025,1051]" gridrow="11" pageId="12" pageNumber="341" rowspan-11="1">
<th id="DA78601DFF90007613C2FB95D94C4918" box="[163,331,1025,1051]" gridcol="0" gridrow="11" pageId="12" pageNumber="341">
<taxonomicName id="E09970A0FF90FF8513C2FB96D907491A" authorityName="Randall &amp; Allen" authorityYear="1987" box="[163,256,1026,1049]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="341" phylum="Chordata" rank="species" species="stictus">
<emphasis id="15EDD731FF90FF8513C2FB96D907491A" box="[163,256,1026,1049]" italics="true" pageId="12" pageNumber="341">E. stictus</emphasis>
</taxonomicName>
</th>
<td id="DA78601DFF9000761208FB95D9A84918" box="[361,431,1025,1051]" gridcol="1" gridrow="11" pageId="12" pageNumber="341">12.50%</td>
<td id="DA78601DFF90007612A8FB95DA084918" box="[457,527,1025,1051]" gridcol="2" gridrow="11" pageId="12" pageNumber="341">11.50%</td>
<td id="DA78601DFF9000761147FB95DA6C4918" box="[550,619,1025,1051]" gridcol="3" gridrow="11" pageId="12" pageNumber="341">11.80%</td>
<td id="DA78601DFF90007611E4FB95DACC4918" box="[645,715,1025,1051]" gridcol="4" gridrow="11" pageId="12" pageNumber="341">12.40%</td>
<td id="DA78601DFF9000761184FB95DB2C4918" box="[741,811,1025,1051]" gridcol="5" gridrow="11" pageId="12" pageNumber="341">12.60%</td>
<td id="DA78601DFF9000761025FB95DB8D4918" box="[836,906,1025,1051]" gridcol="6" gridrow="11" pageId="12" pageNumber="341">11.20%</td>
<td id="DA78601DFF90007610C3FB95DBEF4918" box="[930,1000,1025,1051]" gridcol="7" gridrow="11" pageId="12" pageNumber="341">12.00%</td>
<td id="DA78601DFF9000761770FB95DC504918" box="[1041,1111,1025,1051]" gridcol="8" gridrow="11" pageId="12" pageNumber="341">12.10%</td>
<td id="DA78601DFF900076171CFB95DCC34918" box="[1149,1220,1025,1051]" gridcol="9" gridrow="11" pageId="12" pageNumber="341">14.50%</td>
<td id="DA78601DFF900076178DFB95DD344918" box="[1260,1331,1025,1051]" gridcol="10" gridrow="11" pageId="12" pageNumber="341">0.30%</td>
</tr>
<tr id="99A90961FF90007613C2FBBBDD79494A" box="[163,1406,1071,1097]" gridrow="12" pageId="12" pageNumber="341">
<th id="DA78601DFF90007613C2FBBBD94C494A" box="[163,331,1071,1097]" gridcol="0" gridrow="12" pageId="12" pageNumber="341">
<taxonomicName id="E09970A0FF90FF8513C2FBA4D9254944" baseAuthorityName="Bleeker" baseAuthorityYear="1855" box="[163,290,1071,1095]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="341" phylum="Chordata" rank="species" species="bontoides">
<emphasis id="15EDD731FF90FF8513C2FBA4D9254944" box="[163,290,1071,1095]" italics="true" pageId="12" pageNumber="341">E. bontoides</emphasis>
</taxonomicName>
</th>
<td id="DA78601DFF9000761208FBBBD9A8494A" box="[361,431,1071,1097]" gridcol="1" gridrow="12" pageId="12" pageNumber="341">15.90%</td>
<td id="DA78601DFF90007612A8FBBBDA08494A" box="[457,527,1071,1097]" gridcol="2" gridrow="12" pageId="12" pageNumber="341">15.60%</td>
<td id="DA78601DFF9000761147FBBBDACC494A" box="[550,715,1071,1097]" colspan="2" colspanRight="1" gridcol="3" gridrow="12" pageId="12" pageNumber="341">15.30% 14.60%</td>
<td id="DA78601DFF9000761184FBBBDB2C494A" box="[741,811,1071,1097]" gridcol="5" gridrow="12" pageId="12" pageNumber="341">15.60%</td>
<td id="DA78601DFF9000761025FBBBDB8D494A" box="[836,906,1071,1097]" gridcol="6" gridrow="12" pageId="12" pageNumber="341">14.20%</td>
<td id="DA78601DFF90007610C3FBBBDBEF494A" box="[930,1000,1071,1097]" gridcol="7" gridrow="12" pageId="12" pageNumber="341">14.60%</td>
<td id="DA78601DFF9000761770FBBBDC50494A" box="[1041,1111,1071,1097]" gridcol="8" gridrow="12" pageId="12" pageNumber="341">15.40%</td>
<td id="DA78601DFF900076171CFBBBDCC3494A" box="[1149,1220,1071,1097]" gridcol="9" gridrow="12" pageId="12" pageNumber="341">15.90%</td>
<td id="DA78601DFF900076178DFBBBDD34494A" box="[1260,1331,1071,1097]" gridcol="10" gridrow="12" pageId="12" pageNumber="341">13.30%</td>
<td id="DA78601DFF900076162EFBBBDD79494A" box="[1359,1406,1071,1097]" gridcol="11" gridrow="12" pageId="12" pageNumber="341">N/A</td>
</tr>
</table>
</paragraph>
<subSubSection id="6F8358A8FF90FF8513A6FBEEDA2049B4" pageId="12" pageNumber="341" type="etymology">
<paragraph id="27260B23FF90FF8513A6FBEEDA2049B4" blockId="12.[151,1437,1145,2036]" pageId="12" pageNumber="341">
<emphasis id="15EDD731FF90FF8513A6FBEED9494997" bold="true" box="[199,334,1146,1172]" pageId="12" pageNumber="341">Etymology.</emphasis>
Derived from the Latin 
<emphasis id="15EDD731FF90FF85113EFBEEDA9D4997" box="[607,666,1146,1172]" italics="true" pageId="12" pageNumber="341">fusco</emphasis>
for brown and 
<emphasis id="15EDD731FF90FF851026FBEEDBC14990" box="[839,966,1146,1171]" italics="true" pageId="12" pageNumber="341">marginatus</emphasis>
for margin, in reference to the broad dark brown margins to the unpaired fins.
</paragraph>
</subSubSection>
<subSubSection id="6F8358A8FF90FF8513A6FB55D96348D4" pageId="12" pageNumber="341" type="distribution">
<paragraph id="27260B23FF90FF8513A6FB55D96348D4" blockId="12.[151,1437,1145,2036]" pageId="12" pageNumber="341">
<emphasis id="15EDD731FF90FF8513A6FB55DA2449DF" bold="true" box="[199,547,1217,1244]" pageId="12" pageNumber="341">Distribution and abundance.</emphasis>
The 
<typeStatus id="F822B581FF90FF851100FB56DA9649DF" box="[609,657,1218,1244]" pageId="12" pageNumber="341">type</typeStatus>
specimens were collected by dropline from the Capricorn Channel, south of the Swain Reefs group, Qld (
<geoCoordinate id="42AD6DE4FF90FF851127FB71DABE4803" box="[582,697,1253,1280]" degrees="22" direction="south" minutes="52.8" orientation="latitude" pageId="12" pageNumber="341" precision="92" value="-22.88">22°52.8’S</geoCoordinate>
<geoCoordinate id="42AD6DE4FF90FF8511A0FB71DB424803" box="[705,837,1253,1280]" degrees="152" direction="east" minutes="40.8" orientation="longitude" pageId="12" pageNumber="341" precision="92" value="152.68">152°40.8’E</geoCoordinate>
), at depths of 
<quantity id="E061A6C6FF90FF851086FB72DC6D4803" box="[999,1130,1254,1280]" metricMagnitude="2" metricUnit="m" metricValue="2.25" metricValueMax="2.3" metricValueMin="2.2" pageId="12" pageNumber="341" unit="m" value="225.0" valueMax="230.0" valueMin="220.0">
220– 
<specimenCount id="319FC0AAFF90FF85177EFB72DC6D4803" box="[1055,1130,1254,1280]" pageId="12" pageNumber="341" type="generic">230 m</specimenCount>
</quantity>
. 
<taxonomicName id="E09970A0FF90FF851714FB72D8C14820" authorityName="Johnson &amp; Wilmer" authorityYear="2019" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="341" phylum="Chordata" rank="species" species="fuscomarginatus">
<emphasis id="15EDD731FF90FF851714FB72D8C14820" italics="true" pageId="12" pageNumber="341">Epinephelus fuscomarginatus</emphasis>
</taxonomicName>
appears to have a scattered distribution within the area south of the Swain Reefs and east of the Capricorn- Bunker group. Professional fishers have reported that it can reliably be caught at a number of sites in this area, but is nowhere common. Our examination of catches of groupers from commercial deep dropline fishing off the east coast of 
<collectingRegion id="E55DC5C1FF90FF8513D7FAE1D93D488C" box="[182,314,1397,1423]" country="Australia" name="Queensland" pageId="12" pageNumber="341">Queensland</collectingRegion>
over the past 30 years have not resulted in any additional photographs, reports, or specimens of this species outside of this area and there are no other confirmed records of the species in 
<collectingCountry id="5F8E4BB3FF90FF851732FA0DDCBA48B0" box="[1107,1213,1433,1459]" name="Australia" pageId="12" pageNumber="341">Australia</collectingCountry>
, or elsewhere in the Indo-west Pacific.
</paragraph>
</subSubSection>
<subSubSection id="6F8358A8FF90FF9813A6FA75DA124FCC" lastPageId="17" lastPageNumber="346" pageId="12" pageNumber="341" type="discussion">
<paragraph id="27260B23FF90FF8413A6FA75D8DD4C1C" blockId="12.[151,1437,1145,2036]" lastBlockId="13.[151,1437,153,719]" lastPageId="13" lastPageNumber="342" pageId="12" pageNumber="341">
<emphasis id="15EDD731FF90FF8513A6FA75D94B48FF" bold="true" box="[199,332,1505,1532]" pageId="12" pageNumber="341">Discussion.</emphasis>
In the process of comparing specimens of 
<taxonomicName id="E09970A0FF90FF851050FA76DBBD48F8" authorityName="in Heemstra &amp; Randall" authorityYear="1993" box="[817,954,1506,1531]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="341" phylum="Chordata" rank="species" species="epistictus">
<emphasis id="15EDD731FF90FF851050FA76DBBD48F8" box="[817,954,1506,1531]" italics="true" pageId="12" pageNumber="341">E. epistictus</emphasis>
</taxonomicName>
to the new species, differences in colouration were noted between populations of 
<taxonomicName id="E09970A0FF90FF85113DF992DAE14B1C" authorityName="in Heemstra &amp; Randall" authorityYear="1993" box="[604,742,1542,1567]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="341" phylum="Chordata" rank="species" species="epistictus">
<emphasis id="15EDD731FF90FF85113DF992DAE14B1C" box="[604,742,1542,1567]" italics="true" pageId="12" pageNumber="341">E. epistictus</emphasis>
</taxonomicName>
from the Sea of 
<collectingCountry id="5F8E4BB3FF90FF8510C5F992DBE34B23" box="[932,996,1542,1568]" name="Japan" pageId="12" pageNumber="341">Japan</collectingCountry>
and East 
<collectingCountry id="5F8E4BB3FF90FF851732F991DC914B1C" box="[1107,1174,1541,1567]" name="China" pageId="12" pageNumber="341">China</collectingCountry>
Sea, versus those from the Indian Ocean east to the Indo-Australian Archipelago. As previously reported by 
<bibRefCitation id="430876D2FF90FF85173BF9BDDD9F4B47" author="Randall, J. E. &amp; Heemstra, P. C." box="[1114,1432,1577,1604]" pageId="12" pageNumber="341" pagination="1 - 332" refId="ref12975" refString="Randall, J. E. &amp; Heemstra, P. C. (1991) Revision of Indo-Pacific groupers (Perciformes: Serranidae: Epinephelinae), with descriptions of five new species. Indo-Pacific Fishes, 20, 1 - 332." type="journal article" year="1991">Randall &amp; Heemstra (1991)</bibRefCitation>
, 
<bibRefCitation id="430876D2FF90FF8513F6F9DAD9D34B6B" author="Heemstra, P. C. &amp; Randall, J. E." box="[151,468,1613,1640]" pageId="12" pageNumber="341" refId="ref12210" refString="Heemstra, P. C. &amp; Randall, J. E. (1993) Groupers of the World (Family Serranidae, Subfamily Epinephelinae); an annotated and illustrated catalogue of the grouper, rockcod, hind, coral grouper and lyretail species known to date. FAO Species Catalog 16. FAO, Rome, 382 pp." type="book" year="1993">Heemstra &amp; Randall (1993)</bibRefCitation>
and 
<bibRefCitation id="430876D2FF90FF85116EF9D9DAE34B6B" author="Craig, M. T. &amp; Sadovy de Mitcheson, Y. J. &amp; Heemstra, P. C." box="[527,740,1613,1640]" pageId="12" pageNumber="341" refId="ref11830" refString="Craig, M. T., Sadovy de Mitcheson, Y. J. &amp; Heemstra, P. C. (2011) Groupers of the world. A field and market guide. NISC Pty Ltd, Grahamstown, 356 pp." type="book" year="2011">
Craig 
<emphasis id="15EDD731FF90FF851137F9DADA8E4B6B" box="[598,649,1614,1640]" italics="true" pageId="12" pageNumber="341">et al</emphasis>
. (2011)
</bibRefCitation>
, specimens from the northwestern Pacific usually have more numerous small dark spots on the soft dorsal and caudal fins and on the head (e.g. fig. 4B (
<collectingCountry id="5F8E4BB3FF90FF8517E5F9E6DCCE4B8F" box="[1156,1225,1650,1676]" name="Japan" pageId="12" pageNumber="341">Japan</collectingCountry>
), fig. 5A (Taiwan); 
<bibRefCitation id="430876D2FF90FF8513F6F902D90B4BB3" author="Lee, S. - C." box="[151,268,1686,1712]" pageId="12" pageNumber="341" pagination="1 - 72" refId="ref12501" refString="Lee, S. - C. (1990) A revision of the serranid fish (family Serranidae) of Taiwan. Journal of Taiwan Museum, 43 (2), 1 - 72." type="journal article" year="1990">Lee, 1990</bibRefCitation>
: fig. 51 (Taiwan); 
<bibRefCitation id="430876D2FF90FF8512BDF901DAC74BB3" author="Masuda, H. &amp; Amaoka, K. &amp; Araga, C. &amp; Uyeno, T. &amp; Yoshino, T." box="[476,704,1685,1712]" pageId="12" pageNumber="341" refId="ref12594" refString="Masuda, H., Amaoka, K., Araga, C., Uyeno, T. &amp; Yoshino, T. (1984) The fishes of the Japanese Archipelago. Vols. 1 - 2. Tokai University Press, Tokyo, 437 pp." type="book" year="1984">
Masuda 
<emphasis id="15EDD731FF90FF85115CF902DA694BB3" box="[573,622,1686,1712]" italics="true" pageId="12" pageNumber="341">et al</emphasis>
., 1984
</bibRefCitation>
: pl. 117A (
<collectingCountry id="5F8E4BB3FF90FF85105AF902DB784BB3" box="[827,895,1686,1712]" name="Japan" pageId="12" pageNumber="341">Japan</collectingCountry>
)), than do other populations from other regions, which often have no, or only a few spots that are mostly confined to the dorsal-fin base (e.g. fig. 4C (Arafura Sea, 
<collectingCountry id="5F8E4BB3FF90FF8513F6F949D9044BFB" box="[151,259,1757,1784]" name="Australia" pageId="12" pageNumber="341">Australia</collectingCountry>
); 
<bibRefCitation id="430876D2FF90FF851278F949DA564BFB" author="Randall, J. E. &amp; Heemstra, P. C." box="[281,593,1757,1784]" pageId="12" pageNumber="341" pagination="1 - 332" refId="ref12975" refString="Randall, J. E. &amp; Heemstra, P. C. (1991) Revision of Indo-Pacific groupers (Perciformes: Serranidae: Epinephelinae), with descriptions of five new species. Indo-Pacific Fishes, 20, 1 - 332." type="journal article" year="1991">Randall &amp; Heemstra, 1991</bibRefCitation>
: pl. 11D (
<collectingCountry id="5F8E4BB3FF90FF8511A7F949DB1C4BF4" box="[710,795,1757,1783]" name="India" pageId="12" pageNumber="341">Cochin</collectingCountry>
, 
<collectingCountry id="5F8E4BB3FF90FF851048F949DB604BFB" box="[809,871,1757,1784]" name="India" pageId="12" pageNumber="341">India</collectingCountry>
); 
<bibRefCitation id="430876D2FF90FF85101CF949DC264BFB" author="Randall, J. E." box="[893,1057,1757,1784]" pageId="12" pageNumber="341" refId="ref12894" refString="Randall, J. E. (1995) Coastal fishes of Oman. Crawford House Publishing Pty Ltd, Bathurst, Australia, 439 pp." type="book" year="1995">Randall, 1995</bibRefCitation>
: pl. 297 (
<collectingCountry id="5F8E4BB3FF90FF8517F0F949DCE94BF4" box="[1169,1262,1757,1783]" name="Bahrain" pageId="12" pageNumber="341">Bahrain</collectingCountry>
, Persian Gulf); 
<bibRefCitation id="430876D2FF90FF8513F6F895D96A4A1F" author="White, W. T. &amp; Last, P. R. &amp; Dharmadi &amp; Faizah, R. &amp; Chodrijah, U. &amp; Prisantoso, B. I. &amp; Pogonoski, J. J. &amp; Puckridge, M. &amp; Blaber, S. J. M." box="[151,365,1793,1820]" pageId="12" pageNumber="341" refId="ref13559" refString="White, W. T., Last, P. R., Dharmadi, Faizah, R., Chodrijah, U., Prisantoso, B. I., Pogonoski, J. J., Puckridge, M. &amp; Blaber, S. J. M. (2013) Market Fishes of Indonesia. ACIAR Monograph 155. Australian Centre for International Agricultural Research, Canberra, 438 pp." type="book" year="2013">
White 
<emphasis id="15EDD731FF90FF851384F896D91E4A1F" box="[229,281,1794,1820]" italics="true" pageId="12" pageNumber="341">et al</emphasis>
., 2013
</bibRefCitation>
: fig. 51.28 (
<collectingCountry id="5F8E4BB3FF90FF85129AF895DA694A1F" box="[507,622,1793,1820]" name="Indonesia" pageId="12" pageNumber="341">Indonesia</collectingCountry>
)). Higher lateral-line scale counts were also recorded for northwestern Pacific specimens (63–70 (
<bibRefCitation id="430876D2FF90FF8512AEF8B1DB004A43" author="Randall, J. E. &amp; Heemstra, P. C." box="[463,775,1829,1856]" pageId="12" pageNumber="341" pagination="1 - 332" refId="ref12975" refString="Randall, J. E. &amp; Heemstra, P. C. (1991) Revision of Indo-Pacific groupers (Perciformes: Serranidae: Epinephelinae), with descriptions of five new species. Indo-Pacific Fishes, 20, 1 - 332." type="journal article" year="1991">Randall &amp; Heemstra, 1991</bibRefCitation>
) and 68–71 (
<bibRefCitation id="430876D2FF90FF8510C5F8B2DC634A43" author="Katayama, M." box="[932,1124,1829,1856]" pageId="12" pageNumber="341" refId="ref12417" refString="Katayama, M. (1960) Fauna Japonica. Vol. 1. Serranidae (Pisces). Biogeographical Society of Japan (Tokyo), 189 pp." type="book" year="1960">Katayama, 1960</bibRefCitation>
), versus 55–66 (
<bibRefCitation id="430876D2FF90FF851642F8B1D94E4A67" author="Randall, J. E. &amp; Heemstra, P. C." pageId="12" pageNumber="341" pagination="1 - 332" refId="ref12975" refString="Randall, J. E. &amp; Heemstra, P. C. (1991) Revision of Indo-Pacific groupers (Perciformes: Serranidae: Epinephelinae), with descriptions of five new species. Indo-Pacific Fishes, 20, 1 - 332." type="journal article" year="1991">Randall &amp; Heemstra, 1991</bibRefCitation>
) from elsewhere, which accords with the counts presented here (
<tableCitation id="6A1B3E98FF90FF851776F8DDDC6A4A67" box="[1047,1133,1865,1892]" captionStart="TABLE 3" captionStartId="14.[151,244,152,179]" captionTargetBox="[163,1423,266,2011]" captionText="TABLE 3. Meristic and morphological values for type specimens of Epinephelus fuscomarginatus sp. nov. and comparative material examined of E. epistictus, E. cf epistictus and E. magniscuttis (measurements as percentage of standard length). * Data for largest specimen of E. magniscuttis is from Nakamura et al. (2018)." httpUri="http://table.plazi.org/id/73E65BABFF92FF8713F6FF0CDCCD4DF8" pageId="12" pageNumber="341" tableUuid="73E65BABFF92FF8713F6FF0CDCCD4DF8">Table 3</tableCitation>
). These morphological differences were supported by the molecular results, which indicated a genetic divergence of 3.70% (
<tableCitation id="6A1B3E98FF90FF8517BAF8F9DD374A8B" box="[1243,1328,1901,1928]" captionStart="TABLE 2" captionStartId="12.[151,244,152,179]" captionTargetBox="[163,1406,245,1097]" captionText="TABLE 2. Estimates of mean sequence divergence within (on diagonal) and between (below diagonal) species." httpUri="http://table.plazi.org/id/73E65BABFF90FF8513F6FF0CDD7E4DB0" pageId="12" pageNumber="341" tableUuid="73E65BABFF90FF8513F6FF0CDD7E4DB0">Table 2</tableCitation>
) between samples available for the two populations and that each population formed a distinct, strongly supported clade (fig. 1). Consequently, the two populations are treated separately here, with those from the Indian Ocean referred to herein as 
<emphasis id="15EDD731FF90FF851260F84ED9154AF0" box="[257,274,2010,2035]" italics="true" pageId="12" pageNumber="341">E</emphasis>
. cf 
<taxonomicName id="E09970A0FF90FF85125AF84ED9A24AF0" authorityName="in Heemstra &amp; Randall" authorityYear="1993" box="[315,421,2010,2035]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="341" phylum="Chordata" rank="species" species="epistictus">
<emphasis id="15EDD731FF90FF85125AF84ED9A24AF0" box="[315,421,2010,2035]" italics="true" pageId="12" pageNumber="341">epistictus</emphasis>
</taxonomicName>
. 
<taxonomicName id="E09970A0FF90FF8512D3F84EDAF14AF7" baseAuthorityName="Boulenger" baseAuthorityYear="1888" box="[434,758,2010,2036]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="341" phylum="Chordata" rank="species" species="praeopercularis">
<emphasis id="15EDD731FF90FF8512D3F84EDAF14AF7" box="[434,758,2010,2036]" italics="true" pageId="12" pageNumber="341">Epinephelus praeopercularis</emphasis>
</taxonomicName>
may be an available name for the Indian Ocean form, should populations from the western Indian Ocean prove to be consistent with others from throughout the broader Indian Ocean region. However, genetic samples and specimens from near the 
<typeStatus id="F822B581FF91FF8410D6FF29DBE04DD4" box="[951,999,189,215]" pageId="13" pageNumber="342">type</typeStatus>
locality (Gulf of 
<collectingCountry id="5F8E4BB3FF91FF8417CFFF29DCF04DD4" box="[1198,1271,189,215]" name="Oman" pageId="13" pageNumber="342">Oman</collectingCountry>
), or from elsewhere in the western Indian Ocean were unavailable, so resolution of this complex was outside of the scope of this study.
</paragraph>
<paragraph id="27260B23FF91FF8413A6FEBDDC644FCC" blockId="13.[151,1437,153,719]" pageId="13" pageNumber="342">
<taxonomicName id="E09970A0FF91FF8413A6FEBDDA144C40" authorityName="Johnson &amp; Wilmer" authorityYear="2019" box="[199,531,297,323]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="342" phylum="Chordata" rank="species" species="fuscomarginatus">
<emphasis id="15EDD731FF91FF8413A6FEBDDA144C40" box="[199,531,297,323]" italics="true" pageId="13" pageNumber="342">Epinephelus fuscomarginatus</emphasis>
</taxonomicName>
(fig. 2A–C; 3A–D; 4A) is most similar to 
<taxonomicName id="E09970A0FF91FF841093FEBEDC7C4C40" authorityName="in Heemstra &amp; Randall" authorityYear="1993" box="[1010,1147,298,323]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="342" phylum="Chordata" rank="species" species="epistictus">
<emphasis id="15EDD731FF91FF841093FEBEDC7C4C40" box="[1010,1147,298,323]" italics="true" pageId="13" pageNumber="342">E. epistictus</emphasis>
</taxonomicName>
(fig. 4B, 5A), 
<emphasis id="15EDD731FF91FF841643FEBEDD344C40" box="[1314,1331,298,323]" italics="true" pageId="13" pageNumber="342">E</emphasis>
. cf 
<taxonomicName id="E09970A0FF91FF84163FFEBED8CB4C64" authorityName="in Heemstra &amp; Randall" authorityYear="1993" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="342" phylum="Chordata" rank="species" species="epistictus">
<emphasis id="15EDD731FF91FF84163FFEBED8CB4C64" italics="true" pageId="13" pageNumber="342">epistictus</emphasis>
</taxonomicName>
(fig. 4C) and 
<taxonomicName id="E09970A0FF91FF841208FEDADA114C64" box="[361,534,334,359]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="342" phylum="Chordata" rank="species" species="magniscuttis">
<emphasis id="15EDD731FF91FF841208FEDADA114C64" box="[361,534,334,359]" italics="true" pageId="13" pageNumber="342">E. magniscuttis</emphasis>
</taxonomicName>
(fig. 4D, 6A–B; 
<bibRefCitation id="430876D2FF91FF8411B5FED9DBD74C64" author="Nakamura, J. &amp; Takayama, M. &amp; Worthington Wilmer, J. &amp; Johnson, J. W. &amp; Motomura, H." box="[724,976,333,359]" pageId="13" pageNumber="342" pagination="225 - 228" refId="ref12706" refString="Nakamura, J., Takayama, M., Worthington Wilmer, J., Johnson, J. W. &amp; Motomura, H. (2018) First Japanese record of the Speckled Grouper Epinephelus magniscuttis (Perciformes: Serranidae) from the Osumi Islands. Species Diversity, 23 (2), 225 - 228. https: // doi. org / 10.12782 / specdiv. 23.225" type="journal article" year="2018">
Nakamura 
<emphasis id="15EDD731FF91FF841031FEDADB864C64" box="[848,897,333,359]" italics="true" pageId="13" pageNumber="342">et al</emphasis>
., 2018
</bibRefCitation>
, fig. 1), sharing very similar morphology and meristic formulae. It differs most obviously from all of the latter in lacking any small dark spots on the head, body, or fins at any known size. It also has irregular broad dark margins to the soft dorsal, anal and caudal fins (absent in the latter) and lacks pale or whitish outer margins to the soft dorsal and anal fins, or to the upper and lower corners of the caudal fin (usually present in 
<emphasis id="15EDD731FF91FF8411E9FE4ADA9E4CF4" box="[648,665,478,503]" italics="true" pageId="13" pageNumber="342">E</emphasis>
. cf 
<taxonomicName id="E09970A0FF91FF8411A4FE4ADB284CF4" authorityName="in Heemstra &amp; Randall" authorityYear="1993" box="[709,815,478,503]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="342" phylum="Chordata" rank="species" species="epistictus">
<emphasis id="15EDD731FF91FF8411A4FE4ADB284CF4" box="[709,815,478,503]" italics="true" pageId="13" pageNumber="342">epistictus</emphasis>
</taxonomicName>
). Colouration of juveniles is unknown, but that of subadults (fig. 4A) differs markedly from 
<taxonomicName id="E09970A0FF91FF841126FD96DAD74F18" authorityName="in Heemstra &amp; Randall" authorityYear="1993" box="[583,720,514,539]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="342" phylum="Chordata" rank="species" species="epistictus">
<emphasis id="15EDD731FF91FF841126FD96DAD74F18" box="[583,720,514,539]" italics="true" pageId="13" pageNumber="342">E. epistictus</emphasis>
</taxonomicName>
(fig. 4B), 
<emphasis id="15EDD731FF91FF841024FD96DB514F18" box="[837,854,514,539]" italics="true" pageId="13" pageNumber="342">E</emphasis>
. cf 
<taxonomicName id="E09970A0FF91FF8410E1FD96DBED4F18" authorityName="in Heemstra &amp; Randall" authorityYear="1993" box="[896,1002,514,539]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="342" phylum="Chordata" rank="species" species="epistictus">
<emphasis id="15EDD731FF91FF8410E1FD96DBED4F18" box="[896,1002,514,539]" italics="true" pageId="13" pageNumber="342">epistictus</emphasis>
</taxonomicName>
(fig. 4C) and 
<taxonomicName id="E09970A0FF91FF8417E6FD96DD334F18" box="[1159,1332,514,539]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="342" phylum="Chordata" rank="species" species="magniscuttis">
<emphasis id="15EDD731FF91FF8417E6FD96DD334F18" box="[1159,1332,514,539]" italics="true" pageId="13" pageNumber="342">E. magniscuttis</emphasis>
</taxonomicName>
(fig. 4D) by the presence on the sides of the body of a series of broad irregular diffuse wavy brown bars and blotches (absent in all of the latter). Reports by commercial fishers (S. Campbell pers. comm., 2018) indicate that smaller specimens of 
<taxonomicName id="E09970A0FF91FF8413D6FDFAD9954F84" authorityName="Johnson &amp; Wilmer" authorityYear="2019" box="[183,402,621,647]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="342" phylum="Chordata" rank="species" species="fuscomarginatus">
<emphasis id="15EDD731FF91FF8413D6FDFAD9954F84" box="[183,402,621,647]" italics="true" pageId="13" pageNumber="342">E. fuscomarginatus</emphasis>
</taxonomicName>
of around 
<quantity id="E061A6C6FF91FF841173FDF9DA854F84" box="[530,642,621,647]" metricMagnitude="-1" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="13" pageNumber="342" unit="cm" value="35.0" valueMax="40.0" valueMin="30.0">30–40 cm</quantity>
TL, which are caught and released due to minimum size restrictions, also typically have broad irregular diffuse wavy brown bars and blotches on the sides. Furthermore, in common with larger fish these smaller individuals also lack any dark spots on the head, body, or fins.
</paragraph>
<caption id="73E65BABFF91FF8413F6F82DDD854AF5" ID-DOI="http://doi.org/10.5281/zenodo.3460188" ID-Zenodo-Dep="3460188" httpUri="https://zenodo.org/record/3460188/files/figure.png" pageId="13" pageNumber="342" startId="13.[151,250,1977,2002]" targetBox="[151,1435,745,1943]" targetPageId="13">
<paragraph id="27260B23FF91FF8413F6F82DDD854AF5" blockId="13.[151,1436,1977,2038]" pageId="13" pageNumber="342">
<emphasis id="15EDD731FF91FF8413F6F82DD9154AD1" bold="true" box="[151,274,1977,2002]" pageId="13" pageNumber="342">FIGURE 5.</emphasis>
A. 
<taxonomicName id="E09970A0FF91FF841252F82EDA114AD1" authorityName="in Heemstra &amp; Randall" authorityYear="1993" box="[307,534,1978,2002]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="342" phylum="Chordata" rank="species" species="epistictus">
<emphasis id="15EDD731FF91FF841252F82EDA114AD1" box="[307,534,1978,2002]" italics="true" pageId="13" pageNumber="342">Epinephelus epistictus</emphasis>
</taxonomicName>
, KAUM I.113180 (GenBank MK639008), 224 mm SL, off Ke-Zai-Liao, Kaohsiung, Taiwan, fresh (Photo: KAUM); B. 
<taxonomicName id="E09970A0FF91FF8412B8F84ADAC44AF5" baseAuthorityName="non Fowler" baseAuthorityYear="1904" box="[473,707,2014,2038]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="342" phylum="Chordata" rank="species" species="heniochus">
<emphasis id="15EDD731FF91FF8412B8F84ADAC44AF5" box="[473,707,2014,2038]" italics="true" pageId="13" pageNumber="342">Epinephelus heniochus</emphasis>
</taxonomicName>
, KAUM I.23282, 86.9 mm, Gulf of Thailand, fresh (Photo: KAUM).
</paragraph>
</caption>
<caption id="73E65BABFF92FF8713F6FF0CDCCD4DF8" ID-Table-UUID="73E65BABFF92FF8713F6FF0CDCCD4DF8" httpUri="http://table.plazi.org/id/73E65BABFF92FF8713F6FF0CDCCD4DF8" pageId="14" pageNumber="343" startId="14.[151,244,152,179]" targetBox="[163,1423,266,2011]" targetIsTable="true" targetPageId="14">
<paragraph id="27260B23FF92FF8713F6FF0CDCCD4DF8" blockId="14.[151,1436,152,252]" pageId="14" pageNumber="343">
<emphasis id="15EDD731FF92FF8713F6FF0CD9144DB0" bold="true" box="[151,275,152,179]" pageId="14" pageNumber="343">TABLE 3.</emphasis>
Meristic and morphological values for 
<typeStatus id="F822B581FF92FF8711BEFF0DDB084DB0" box="[735,783,153,179]" pageId="14" pageNumber="343">type</typeStatus>
specimens of 
<taxonomicName id="E09970A0FF92FF8710D8FF0DDD004DB0" authority="Johnson &amp; Wilmer, 2019" authorityName="Johnson &amp; Wilmer" authorityYear="2019" box="[953,1287,153,179]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="343" phylum="Chordata" rank="species" species="fuscomarginatus" status="sp. nov.">
<emphasis id="15EDD731FF92FF8710D8FF0DDD004DB0" box="[953,1287,153,179]" italics="true" pageId="14" pageNumber="343">Epinephelus fuscomarginatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0EDE6A4AFF92FF871670FF0DDD6C4DB0" box="[1297,1387,153,179]" pageId="14" pageNumber="343" rank="species">sp. nov.</taxonomicNameLabel>
and comparative material examined of 
<taxonomicName id="E09970A0FF92FF87117BFF2ADAA54DD4" authorityName="in Heemstra &amp; Randall" authorityYear="1993" box="[538,674,190,215]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="343" phylum="Chordata" rank="species" species="epistictus">
<emphasis id="15EDD731FF92FF87117BFF2ADAA54DD4" box="[538,674,190,215]" italics="true" pageId="14" pageNumber="343">E. epistictus</emphasis>
</taxonomicName>
, 
<emphasis id="15EDD731FF92FF8711CEFF2ADAC74DD4" box="[687,704,190,215]" italics="true" pageId="14" pageNumber="343">E</emphasis>
. cf 
<taxonomicName id="E09970A0FF92FF871188FF2ADB544DD4" authorityName="in Heemstra &amp; Randall" authorityYear="1993" box="[745,851,190,215]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="343" phylum="Chordata" rank="species" species="epistictus">
<emphasis id="15EDD731FF92FF871188FF2ADB544DD4" box="[745,851,190,215]" italics="true" pageId="14" pageNumber="343">epistictus</emphasis>
</taxonomicName>
and 
<taxonomicName id="E09970A0FF92FF8710E6FF2ADC344DD4" box="[903,1075,190,215]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="343" phylum="Chordata" rank="species" species="magniscuttis">
<emphasis id="15EDD731FF92FF8710E6FF2ADC344DD4" box="[903,1075,190,215]" italics="true" pageId="14" pageNumber="343">E. magniscuttis</emphasis>
</taxonomicName>
(measurements as percentage of standard length). * Data for largest specimen of 
<taxonomicName id="E09970A0FF92FF8711D2FF76DB674DF8" box="[691,864,226,251]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="343" phylum="Chordata" rank="species" species="magniscuttis">
<emphasis id="15EDD731FF92FF8711D2FF76DB674DF8" box="[691,864,226,251]" italics="true" pageId="14" pageNumber="343">E. magniscuttis</emphasis>
</taxonomicName>
is from 
<bibRefCitation id="430876D2FF92FF8710DFFF75DCC14DF8" author="Nakamura, J. &amp; Takayama, M. &amp; Worthington Wilmer, J. &amp; Johnson, J. W. &amp; Motomura, H." box="[958,1222,225,251]" pageId="14" pageNumber="343" pagination="225 - 228" refId="ref12706" refString="Nakamura, J., Takayama, M., Worthington Wilmer, J., Johnson, J. W. &amp; Motomura, H. (2018) First Japanese record of the Speckled Grouper Epinephelus magniscuttis (Perciformes: Serranidae) from the Osumi Islands. Species Diversity, 23 (2), 225 - 228. https: // doi. org / 10.12782 / specdiv. 23.225" type="journal article" year="2018">
Nakamura 
<emphasis id="15EDD731FF92FF871758FF76DC6C4DF8" box="[1081,1131,225,251]" italics="true" pageId="14" pageNumber="343">et al</emphasis>
. (2018)
</bibRefCitation>
.
</paragraph>
</caption>
<paragraph id="27260B23FF92FF8712A0FE98DD694AD9" pageId="14" pageNumber="343">
<table id="5599F983FF92007613C2FE9EDD884AD8" box="[163,1423,266,2011]" gridcols="6" gridrows="43" pageId="14" pageNumber="343">
<tr id="99A90961FF92007613C2FE9EDD884C20" box="[163,1423,266,291]" gridrow="0" pageId="14" pageNumber="343" rowspan-0="1">
<th id="DA78601DFF92007612A0FE9EDA824C20" box="[449,645,266,291]" gridcol="1" gridrow="0" pageId="14" pageNumber="343">
<taxonomicName id="E09970A0FF92FF8712A0FE98DA824C20" authorityName="Johnson &amp; Wilmer" authorityYear="2019" box="[449,645,267,291]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="343" phylum="Chordata" rank="species" species="fuscomarginatus">
<emphasis id="15EDD731FF92FF8712A0FE98DA824C20" box="[449,645,267,291]" italics="true" pageId="14" pageNumber="343">E. fuscomarginatus</emphasis>
</taxonomicName>
</th>
<th id="DA78601DFF92007611FDFE9EDB584C20" box="[668,863,266,291]" gridcol="2" gridrow="0" pageId="14" pageNumber="343">
<taxonomicName id="E09970A0FF92FF8711FDFE98DB584C20" authorityName="Johnson &amp; Wilmer" authorityYear="2019" box="[668,863,267,291]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="343" phylum="Chordata" rank="species" species="fuscomarginatus">
<emphasis id="15EDD731FF92FF8711FDFE98DB584C20" box="[668,863,267,291]" italics="true" pageId="14" pageNumber="343">E. fuscomarginatus</emphasis>
</taxonomicName>
</th>
<th id="DA78601DFF92007610E6FE9EDC044C20" box="[903,1027,266,291]" gridcol="3" gridrow="0" pageId="14" pageNumber="343">
<taxonomicName id="E09970A0FF92FF8710E6FE98DC044C20" authorityName="in Heemstra &amp; Randall" authorityYear="1993" box="[903,1027,268,291]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="343" phylum="Chordata" rank="species" species="epistictus">
<emphasis id="15EDD731FF92FF8710E6FE98DC044C20" box="[903,1027,268,291]" italics="true" pageId="14" pageNumber="343">E. epistictus</emphasis>
</taxonomicName>
</th>
<th id="DA78601DFF9200761756FE9EDCCA4C20" box="[1079,1229,266,291]" gridcol="4" gridrow="0" pageId="14" pageNumber="343">
<emphasis id="15EDD731FF92FF871756FE98DC414C20" box="[1079,1094,268,291]" italics="true" pageId="14" pageNumber="343">E</emphasis>
. cf 
<taxonomicName id="E09970A0FF92FF87170CFE98DCCA4C20" authorityName="in Heemstra &amp; Randall" authorityYear="1993" box="[1133,1229,268,291]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="343" phylum="Chordata" rank="species" species="epistictus">
<emphasis id="15EDD731FF92FF87170CFE98DCCA4C20" box="[1133,1229,268,291]" italics="true" pageId="14" pageNumber="343">epistictus</emphasis>
</taxonomicName>
</th>
<th id="DA78601DFF9200761792FE9EDD884C20" box="[1267,1423,266,291]" gridcol="5" gridrow="0" pageId="14" pageNumber="343">
<taxonomicName id="E09970A0FF92FF871792FE98DD884C20" box="[1267,1423,268,291]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="343" phylum="Chordata" rank="species" species="magniscuttis">
<emphasis id="15EDD731FF92FF871792FE98DD884C20" box="[1267,1423,268,291]" italics="true" pageId="14" pageNumber="343">E. magniscuttis</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="99A90961FF92007613C2FEA6DD884C4F" box="[163,1423,306,332]" gridrow="1" pageId="14" pageNumber="343" rowspan-0="1">
<td id="DA78601DFF92007612A0FEA6DA824C4F" box="[449,645,306,332]" gridcol="1" gridrow="1" pageId="14" pageNumber="343">Holotype</td>
<td id="DA78601DFF92007611FDFEA6DB584C4F" box="[668,863,306,332]" gridcol="2" gridrow="1" pageId="14" pageNumber="343">Paratypes</td>
<td id="DA78601DFF92007610E6FEA6DC044C4F" box="[903,1027,306,332]" gridcol="3" gridrow="1" pageId="14" pageNumber="343">non-types</td>
<td id="DA78601DFF9200761756FEA6DCCA4C4F" box="[1079,1229,306,332]" gridcol="4" gridrow="1" pageId="14" pageNumber="343">non-types</td>
<td id="DA78601DFF9200761792FEA6DD884C4F" box="[1267,1423,306,332]" gridcol="5" gridrow="1" pageId="14" pageNumber="343">non-types</td>
</tr>
<tr id="99A90961FF92007613C2FECFDD884C77" box="[163,1423,347,372]" gridrow="2" pageId="14" pageNumber="343" rowspan-0="1">
<td id="DA78601DFF92007612A0FECFDA824C77" box="[449,645,347,372]" gridcol="1" gridrow="2" pageId="14" pageNumber="343">QM I.40945</td>
<td id="DA78601DFF92007611FDFECFDB584C77" box="[668,863,347,372]" gridcol="2" gridrow="2" pageId="14" pageNumber="343">4</td>
<td id="DA78601DFF92007610E6FECFDC044C77" box="[903,1027,347,372]" gridcol="3" gridrow="2" pageId="14" pageNumber="343">3</td>
<td id="DA78601DFF9200761756FECFDCCA4C77" box="[1079,1229,347,372]" gridcol="4" gridrow="2" pageId="14" pageNumber="343">8</td>
<td id="DA78601DFF9200761792FECFDD884C77" box="[1267,1423,347,372]" gridcol="5" gridrow="2" pageId="14" pageNumber="343">4*</td>
</tr>
<tr id="99A90961FF92007613C2FE17DD884C9F" box="[163,1423,387,412]" gridrow="3" pageId="14" pageNumber="343">
<th id="DA78601DFF92007613C2FE17D9AE4C9F" box="[163,425,387,412]" gridcol="0" gridrow="3" pageId="14" pageNumber="343">Standard length (mm)</th>
<td id="DA78601DFF92007612A0FE17DA824C9F" box="[449,645,387,412]" gridcol="1" gridrow="3" pageId="14" pageNumber="343">564</td>
<td id="DA78601DFF92007611FDFE17DB584C9F" box="[668,863,387,412]" gridcol="2" gridrow="3" pageId="14" pageNumber="343">408–528</td>
<td id="DA78601DFF92007610E6FE17DC044C9F" box="[903,1027,387,412]" gridcol="3" gridrow="3" pageId="14" pageNumber="343">190–376</td>
<td id="DA78601DFF9200761756FE17DCCA4C9F" box="[1079,1229,387,412]" gridcol="4" gridrow="3" pageId="14" pageNumber="343">134–406</td>
<td id="DA78601DFF9200761792FE17DD884C9F" box="[1267,1423,387,412]" gridcol="5" gridrow="3" pageId="14" pageNumber="343">419–613</td>
</tr>
<tr id="99A90961FF92007613C2FE3FDD884CC7" box="[163,1423,427,452]" gridrow="4" pageId="14" pageNumber="343">
<th id="DA78601DFF92007613C2FE3FD9AE4CC7" box="[163,425,427,452]" gridcol="0" gridrow="4" pageId="14" pageNumber="343">Total length</th>
<td id="DA78601DFF92007612A0FE3FDA824CC7" box="[449,645,427,452]" gridcol="1" gridrow="4" pageId="14" pageNumber="343">662</td>
<td id="DA78601DFF92007611FDFE3FDB584CC7" box="[668,863,427,452]" gridcol="2" gridrow="4" pageId="14" pageNumber="343">490–626</td>
<td id="DA78601DFF92007610E6FE3FDC044CC7" box="[903,1027,427,452]" gridcol="3" gridrow="4" pageId="14" pageNumber="343">236–459</td>
<td id="DA78601DFF9200761756FE3FDCCA4CC7" box="[1079,1229,427,452]" gridcol="4" gridrow="4" pageId="14" pageNumber="343">165–502</td>
<td id="DA78601DFF9200761792FE3FDD884CC7" box="[1267,1423,427,452]" gridcol="5" gridrow="4" pageId="14" pageNumber="343">507–739</td>
</tr>
<tr id="99A90961FF92007613C2FE47DD884CEF" box="[163,1423,467,492]" gridrow="5" pageId="14" pageNumber="343">
<th id="DA78601DFF92007613C2FE47D9AE4CEF" box="[163,425,467,492]" gridcol="0" gridrow="5" pageId="14" pageNumber="343">Dorsal-fin rays</th>
<td id="DA78601DFF92007612A0FE47DA824CEF" box="[449,645,467,492]" gridcol="1" gridrow="5" pageId="14" pageNumber="343">XI/14</td>
<td id="DA78601DFF92007611FDFE47DB584CEF" box="[668,863,467,492]" gridcol="2" gridrow="5" pageId="14" pageNumber="343">XI/14</td>
<td id="DA78601DFF92007610E6FE47DC044CEF" box="[903,1027,467,492]" gridcol="3" gridrow="5" pageId="14" pageNumber="343">XI/14–15</td>
<td id="DA78601DFF9200761756FE47DCCA4CEF" box="[1079,1229,467,492]" gridcol="4" gridrow="5" pageId="14" pageNumber="343">XI/14–15</td>
<td id="DA78601DFF9200761792FE47DD884CEF" box="[1267,1423,467,492]" gridcol="5" gridrow="5" pageId="14" pageNumber="343">XI/14–15</td>
</tr>
<tr id="99A90961FF92007613C2FE68DD884F16" box="[163,1423,508,533]" gridrow="6" pageId="14" pageNumber="343">
<th id="DA78601DFF92007613C2FE68D9AE4F16" box="[163,425,508,533]" gridcol="0" gridrow="6" pageId="14" pageNumber="343">Anal-fin rays</th>
<td id="DA78601DFF92007612A0FE68DA824F16" box="[449,645,508,533]" gridcol="1" gridrow="6" pageId="14" pageNumber="343">III/8</td>
<td id="DA78601DFF92007611FDFE68DB584F16" box="[668,863,508,533]" gridcol="2" gridrow="6" pageId="14" pageNumber="343">III/8</td>
<td id="DA78601DFF92007610E6FE68DC044F16" box="[903,1027,508,533]" gridcol="3" gridrow="6" pageId="14" pageNumber="343">III/8</td>
<td id="DA78601DFF9200761756FE68DCCA4F16" box="[1079,1229,508,533]" gridcol="4" gridrow="6" pageId="14" pageNumber="343">III/8</td>
<td id="DA78601DFF9200761792FE68DD884F16" box="[1267,1423,508,533]" gridcol="5" gridrow="6" pageId="14" pageNumber="343">III/8</td>
</tr>
<tr id="99A90961FF92007613C2FDB0DD884F3E" box="[163,1423,548,573]" gridrow="7" pageId="14" pageNumber="343">
<th id="DA78601DFF92007613C2FDB0D9AE4F3E" box="[163,425,548,573]" gridcol="0" gridrow="7" pageId="14" pageNumber="343">Pectoral-fin rays</th>
<td id="DA78601DFF92007612A0FDB0DA824F3E" box="[449,645,548,573]" gridcol="1" gridrow="7" pageId="14" pageNumber="343">17</td>
<td id="DA78601DFF92007611FDFDB0DB584F3E" box="[668,863,548,573]" gridcol="2" gridrow="7" pageId="14" pageNumber="343">17</td>
<td id="DA78601DFF92007610E6FDB0DC044F3E" box="[903,1027,548,573]" gridcol="3" gridrow="7" pageId="14" pageNumber="343">18</td>
<td id="DA78601DFF9200761756FDB0DCCA4F3E" box="[1079,1229,548,573]" gridcol="4" gridrow="7" pageId="14" pageNumber="343">18–19</td>
<td id="DA78601DFF9200761792FDB0DD884F3E" box="[1267,1423,548,573]" gridcol="5" gridrow="7" pageId="14" pageNumber="343">18</td>
</tr>
<tr id="99A90961FF92007613C2FDD8DD884F66" box="[163,1423,588,613]" gridrow="8" pageId="14" pageNumber="343">
<th id="DA78601DFF92007613C2FDD8D9AE4F66" box="[163,425,588,613]" gridcol="0" gridrow="8" pageId="14" pageNumber="343">Lateral-line scales</th>
<td id="DA78601DFF92007612A0FDD8DA824F66" box="[449,645,588,613]" gridcol="1" gridrow="8" pageId="14" pageNumber="343">65</td>
<td id="DA78601DFF92007611FDFDD8DB584F66" box="[668,863,588,613]" gridcol="2" gridrow="8" pageId="14" pageNumber="343">60–67</td>
<td id="DA78601DFF92007610E6FDD8DC044F66" box="[903,1027,588,613]" gridcol="3" gridrow="8" pageId="14" pageNumber="343">69–71</td>
<td id="DA78601DFF9200761756FDD8DCCA4F66" box="[1079,1229,588,613]" gridcol="4" gridrow="8" pageId="14" pageNumber="343">61–67</td>
<td id="DA78601DFF9200761792FDD8DD884F66" box="[1267,1423,588,613]" gridcol="5" gridrow="8" pageId="14" pageNumber="343">58–62</td>
</tr>
<tr id="99A90961FF92007613C2FDE0DD884F8E" box="[163,1423,628,653]" gridrow="9" pageId="14" pageNumber="343">
<th id="DA78601DFF92007613C2FDE0D9AE4F8E" box="[163,425,628,653]" gridcol="0" gridrow="9" pageId="14" pageNumber="343">Lateral-scale series</th>
<td id="DA78601DFF92007612A0FDE0DA824F8E" box="[449,645,628,653]" gridcol="1" gridrow="9" pageId="14" pageNumber="343">102</td>
<td id="DA78601DFF92007611FDFDE0DB584F8E" box="[668,863,628,653]" gridcol="2" gridrow="9" pageId="14" pageNumber="343">101–116</td>
<td id="DA78601DFF92007610E6FDE0DC044F8E" box="[903,1027,628,653]" gridcol="3" gridrow="9" pageId="14" pageNumber="343">–</td>
<td id="DA78601DFF9200761756FDE0DCCA4F8E" box="[1079,1229,628,653]" gridcol="4" gridrow="9" pageId="14" pageNumber="343">–</td>
<td id="DA78601DFF9200761792FDE0DD884F8E" box="[1267,1423,628,653]" gridcol="5" gridrow="9" pageId="14" pageNumber="343">–</td>
</tr>
<tr id="99A90961FF92007613C2FD09DD884FD0" box="[163,1423,669,723]" gridrow="10" pageId="14" pageNumber="343">
<th id="DA78601DFF92007613C2FD09D9AE4FD0" box="[163,425,669,723]" gridcol="0" gridrow="10" pageId="14" pageNumber="343">Gill rakers</th>
<td id="DA78601DFF92007612A0FD09DA824FD0" box="[449,645,669,723]" gridcol="1" gridrow="10" pageId="14" pageNumber="343">9+17 =26</td>
<td id="DA78601DFF92007611FDFD09DB584FD0" box="[668,863,669,723]" gridcol="2" gridrow="10" pageId="14" pageNumber="343">9–10+16–19 =25–28</td>
<td id="DA78601DFF92007610E6FD09DC044FD0" box="[903,1027,669,723]" gridcol="3" gridrow="10" pageId="14" pageNumber="343">8–9+16 =24–25</td>
<td id="DA78601DFF9200761756FD09DCCA4FD0" box="[1079,1229,669,723]" gridcol="4" gridrow="10" pageId="14" pageNumber="343">8–10+15–17 =23–27</td>
<td id="DA78601DFF9200761792FD09DD884FD0" box="[1267,1423,669,723]" gridcol="5" gridrow="10" pageId="14" pageNumber="343">8–9+15–16 =23–25</td>
</tr>
<tr id="99A90961FF92007613C2FD76DD884FF8" box="[163,1423,738,763]" gridrow="11" pageId="14" pageNumber="343">
<th id="DA78601DFF92007613C2FD76D9AE4FF8" box="[163,425,738,763]" gridcol="0" gridrow="11" pageId="14" pageNumber="343">Body depth</th>
<td id="DA78601DFF92007612A0FD76DA824FF8" box="[449,645,738,763]" gridcol="1" gridrow="11" pageId="14" pageNumber="343">33.2</td>
<td id="DA78601DFF92007611FDFD76DB584FF8" box="[668,863,738,763]" gridcol="2" gridrow="11" pageId="14" pageNumber="343">29.8–32.6</td>
<td id="DA78601DFF92007610E6FD76DC044FF8" box="[903,1027,738,763]" gridcol="3" gridrow="11" pageId="14" pageNumber="343">31.3–34.3</td>
<td id="DA78601DFF9200761756FD76DCCA4FF8" box="[1079,1229,738,763]" gridcol="4" gridrow="11" pageId="14" pageNumber="343">31.8–35.1</td>
<td id="DA78601DFF9200761792FD76DD884FF8" box="[1267,1423,738,763]" gridcol="5" gridrow="11" pageId="14" pageNumber="343">33.8–35.5</td>
</tr>
<tr id="99A90961FF92007613C2FC9EDD884E20" box="[163,1423,778,803]" gridrow="12" pageId="14" pageNumber="343">
<th id="DA78601DFF92007613C2FC9ED9AE4E20" box="[163,425,778,803]" gridcol="0" gridrow="12" pageId="14" pageNumber="343">Body width</th>
<td id="DA78601DFF92007612A0FC9EDA824E20" box="[449,645,778,803]" gridcol="1" gridrow="12" pageId="14" pageNumber="343">19.1</td>
<td id="DA78601DFF92007611FDFC9EDB584E20" box="[668,863,778,803]" gridcol="2" gridrow="12" pageId="14" pageNumber="343">16.9–19.7</td>
<td id="DA78601DFF92007610E6FC9EDC044E20" box="[903,1027,778,803]" gridcol="3" gridrow="12" pageId="14" pageNumber="343">15.6–18.3</td>
<td id="DA78601DFF9200761756FC9EDCCA4E20" box="[1079,1229,778,803]" gridcol="4" gridrow="12" pageId="14" pageNumber="343">16.2–19.4</td>
<td id="DA78601DFF9200761792FC9EDD884E20" box="[1267,1423,778,803]" gridcol="5" gridrow="12" pageId="14" pageNumber="343">16.7–22.2</td>
</tr>
<tr id="99A90961FF92007613C2FCA7DD884E4F" box="[163,1423,819,844]" gridrow="13" pageId="14" pageNumber="343">
<th id="DA78601DFF92007613C2FCA7D9AE4E4F" box="[163,425,819,844]" gridcol="0" gridrow="13" pageId="14" pageNumber="343">Head length</th>
<td id="DA78601DFF92007612A0FCA7DA824E4F" box="[449,645,819,844]" gridcol="1" gridrow="13" pageId="14" pageNumber="343">39.6</td>
<td id="DA78601DFF92007611FDFCA7DB584E4F" box="[668,863,819,844]" gridcol="2" gridrow="13" pageId="14" pageNumber="343">40.8–42.1</td>
<td id="DA78601DFF92007610E6FCA7DC044E4F" box="[903,1027,819,844]" gridcol="3" gridrow="13" pageId="14" pageNumber="343">40.1–43.0</td>
<td id="DA78601DFF9200761756FCA7DCCA4E4F" box="[1079,1229,819,844]" gridcol="4" gridrow="13" pageId="14" pageNumber="343">41.9–45.4</td>
<td id="DA78601DFF9200761792FCA7DD884E4F" box="[1267,1423,819,844]" gridcol="5" gridrow="13" pageId="14" pageNumber="343">41.2–42.5</td>
</tr>
<tr id="99A90961FF92007613C2FCCFDD884E77" box="[163,1423,859,884]" gridrow="14" pageId="14" pageNumber="343">
<th id="DA78601DFF92007613C2FCCFD9AE4E77" box="[163,425,859,884]" gridcol="0" gridrow="14" pageId="14" pageNumber="343">Snout length</th>
<td id="DA78601DFF92007612A0FCCFDA824E77" box="[449,645,859,884]" gridcol="1" gridrow="14" pageId="14" pageNumber="343">10.3</td>
<td id="DA78601DFF92007611FDFCCFDB584E77" box="[668,863,859,884]" gridcol="2" gridrow="14" pageId="14" pageNumber="343">9.8–10.7</td>
<td id="DA78601DFF92007610E6FCCFDC044E77" box="[903,1027,859,884]" gridcol="3" gridrow="14" pageId="14" pageNumber="343">8.5–9.8</td>
<td id="DA78601DFF9200761756FCCFDCCA4E77" box="[1079,1229,859,884]" gridcol="4" gridrow="14" pageId="14" pageNumber="343">8.6–11.0</td>
<td id="DA78601DFF9200761792FCCFDD884E77" box="[1267,1423,859,884]" gridcol="5" gridrow="14" pageId="14" pageNumber="343">10.0–11.1</td>
</tr>
<tr id="99A90961FF92007613C2FC17DD884E9F" box="[163,1423,899,924]" gridrow="15" pageId="14" pageNumber="343">
<th id="DA78601DFF92007613C2FC17D9AE4E9F" box="[163,425,899,924]" gridcol="0" gridrow="15" pageId="14" pageNumber="343">Orbit diameter</th>
<td id="DA78601DFF92007612A0FC17DA824E9F" box="[449,645,899,924]" gridcol="1" gridrow="15" pageId="14" pageNumber="343">7.2</td>
<td id="DA78601DFF92007611FDFC17DB584E9F" box="[668,863,899,924]" gridcol="2" gridrow="15" pageId="14" pageNumber="343">6.6–7.3</td>
<td id="DA78601DFF92007610E6FC17DC044E9F" box="[903,1027,899,924]" gridcol="3" gridrow="15" pageId="14" pageNumber="343">6.4–8.7</td>
<td id="DA78601DFF9200761756FC17DCCA4E9F" box="[1079,1229,899,924]" gridcol="4" gridrow="15" pageId="14" pageNumber="343">7.0–9.2</td>
<td id="DA78601DFF9200761792FC17DD884E9F" box="[1267,1423,899,924]" gridcol="5" gridrow="15" pageId="14" pageNumber="343">6.0–7.9</td>
</tr>
<tr id="99A90961FF92007613C2FC3FDD884EC7" box="[163,1423,939,964]" gridrow="16" pageId="14" pageNumber="343">
<th id="DA78601DFF92007613C2FC3FD9AE4EC7" box="[163,425,939,964]" gridcol="0" gridrow="16" pageId="14" pageNumber="343">Preorbital depth</th>
<td id="DA78601DFF92007612A0FC3FDA824EC7" box="[449,645,939,964]" gridcol="1" gridrow="16" pageId="14" pageNumber="343">4.3</td>
<td id="DA78601DFF92007611FDFC3FDB584EC7" box="[668,863,939,964]" gridcol="2" gridrow="16" pageId="14" pageNumber="343">4.2–4.8</td>
<td id="DA78601DFF92007610E6FC3FDC044EC7" box="[903,1027,939,964]" gridcol="3" gridrow="16" pageId="14" pageNumber="343">3.9–4.4</td>
<td id="DA78601DFF9200761756FC3FDCCA4EC7" box="[1079,1229,939,964]" gridcol="4" gridrow="16" pageId="14" pageNumber="343">3.1–4.0</td>
<td id="DA78601DFF9200761792FC3FDD884EC7" box="[1267,1423,939,964]" gridcol="5" gridrow="16" pageId="14" pageNumber="343">3.6–4.0</td>
</tr>
<tr id="99A90961FF92007613C2FC40DD884EEE" box="[163,1423,980,1005]" gridrow="17" pageId="14" pageNumber="343">
<th id="DA78601DFF92007613C2FC40D9AE4EEE" box="[163,425,980,1005]" gridcol="0" gridrow="17" pageId="14" pageNumber="343">Interorbital width</th>
<td id="DA78601DFF92007612A0FC40DA824EEE" box="[449,645,980,1005]" gridcol="1" gridrow="17" pageId="14" pageNumber="343">6.6</td>
<td id="DA78601DFF92007611FDFC40DB584EEE" box="[668,863,980,1005]" gridcol="2" gridrow="17" pageId="14" pageNumber="343">5.9–7.3</td>
<td id="DA78601DFF92007610E6FC40DC044EEE" box="[903,1027,980,1005]" gridcol="3" gridrow="17" pageId="14" pageNumber="343">5.9–7.3</td>
<td id="DA78601DFF9200761756FC40DCCA4EEE" box="[1079,1229,980,1005]" gridcol="4" gridrow="17" pageId="14" pageNumber="343">6.7–6.8</td>
<td id="DA78601DFF9200761792FC40DD884EEE" box="[1267,1423,980,1005]" gridcol="5" gridrow="17" pageId="14" pageNumber="343">6.7–7.6</td>
</tr>
<tr id="99A90961FF92007613C2FC68DD884916" box="[163,1423,1020,1045]" gridrow="18" pageId="14" pageNumber="343">
<th id="DA78601DFF92007613C2FC68D9AE4916" box="[163,425,1020,1045]" gridcol="0" gridrow="18" pageId="14" pageNumber="343">Maxilla width</th>
<td id="DA78601DFF92007612A0FC68DA824916" box="[449,645,1020,1045]" gridcol="1" gridrow="18" pageId="14" pageNumber="343">4.7</td>
<td id="DA78601DFF92007611FDFC68DB584916" box="[668,863,1020,1045]" gridcol="2" gridrow="18" pageId="14" pageNumber="343">5.0–5.7</td>
<td id="DA78601DFF92007610E6FC68DC044916" box="[903,1027,1020,1045]" gridcol="3" gridrow="18" pageId="14" pageNumber="343">5.1–5.4</td>
<td id="DA78601DFF9200761756FC68DCCA4916" box="[1079,1229,1020,1045]" gridcol="4" gridrow="18" pageId="14" pageNumber="343">5.2–5.8</td>
<td id="DA78601DFF9200761792FC68DD884916" box="[1267,1423,1020,1045]" gridcol="5" gridrow="18" pageId="14" pageNumber="343">4.8–5.6</td>
</tr>
<tr id="99A90961FF92007613C2FBB0DD88493E" box="[163,1423,1060,1085]" gridrow="19" pageId="14" pageNumber="343">
<th id="DA78601DFF92007613C2FBB0D9AE493E" box="[163,425,1060,1085]" gridcol="0" gridrow="19" pageId="14" pageNumber="343">Upper jaw length</th>
<td id="DA78601DFF92007612A0FBB0DA82493E" box="[449,645,1060,1085]" gridcol="1" gridrow="19" pageId="14" pageNumber="343">17.7</td>
<td id="DA78601DFF92007611FDFBB0DB58493E" box="[668,863,1060,1085]" gridcol="2" gridrow="19" pageId="14" pageNumber="343">17.4–18.6</td>
<td id="DA78601DFF92007610E6FBB0DC04493E" box="[903,1027,1060,1085]" gridcol="3" gridrow="19" pageId="14" pageNumber="343">18.6–20.2</td>
<td id="DA78601DFF9200761756FBB0DCCA493E" box="[1079,1229,1060,1085]" gridcol="4" gridrow="19" pageId="14" pageNumber="343">19.1–19.8</td>
<td id="DA78601DFF9200761792FBB0DD88493E" box="[1267,1423,1060,1085]" gridcol="5" gridrow="19" pageId="14" pageNumber="343">18.7–19.5</td>
</tr>
<tr id="99A90961FF92007613C2FBD8DD884966" box="[163,1423,1100,1125]" gridrow="20" pageId="14" pageNumber="343">
<th id="DA78601DFF92007613C2FBD8D9AE4966" box="[163,425,1100,1125]" gridcol="0" gridrow="20" pageId="14" pageNumber="343">Diameter anterior nostril</th>
<td id="DA78601DFF92007612A0FBD8DA824966" box="[449,645,1100,1125]" gridcol="1" gridrow="20" pageId="14" pageNumber="343">0.6</td>
<td id="DA78601DFF92007611FDFBD8DB584966" box="[668,863,1100,1125]" gridcol="2" gridrow="20" pageId="14" pageNumber="343">0.6–0.8</td>
<td id="DA78601DFF92007610E6FBD8DC044966" box="[903,1027,1100,1125]" gridcol="3" gridrow="20" pageId="14" pageNumber="343">0.7–0.9</td>
<td id="DA78601DFF9200761756FBD8DCCA4966" box="[1079,1229,1100,1125]" gridcol="4" gridrow="20" pageId="14" pageNumber="343">0.5–0.8</td>
<td id="DA78601DFF9200761792FBD8DD884966" box="[1267,1423,1100,1125]" gridcol="5" gridrow="20" pageId="14" pageNumber="343">0.5–0.9</td>
</tr>
<tr id="99A90961FF92007613C2FBE1DD88498D" box="[163,1423,1141,1166]" gridrow="21" pageId="14" pageNumber="343">
<th id="DA78601DFF92007613C2FBE1D9AE498D" box="[163,425,1141,1166]" gridcol="0" gridrow="21" pageId="14" pageNumber="343">Diameter posterior nostril</th>
<td id="DA78601DFF92007612A0FBE1DA82498D" box="[449,645,1141,1166]" gridcol="1" gridrow="21" pageId="14" pageNumber="343">1.0</td>
<td id="DA78601DFF92007611FDFBE1DB58498D" box="[668,863,1141,1166]" gridcol="2" gridrow="21" pageId="14" pageNumber="343">0.8–1.1</td>
<td id="DA78601DFF92007610E6FBE1DC04498D" box="[903,1027,1141,1166]" gridcol="3" gridrow="21" pageId="14" pageNumber="343">1.0–1.2</td>
<td id="DA78601DFF9200761756FBE1DCCA498D" box="[1079,1229,1141,1166]" gridcol="4" gridrow="21" pageId="14" pageNumber="343">0.7–1.1</td>
<td id="DA78601DFF9200761792FBE1DD88498D" box="[1267,1423,1141,1166]" gridcol="5" gridrow="21" pageId="14" pageNumber="343">1.1–1.6</td>
</tr>
<tr id="99A90961FF92007613C2FB09DD8849B5" box="[163,1423,1181,1206]" gridrow="22" pageId="14" pageNumber="343">
<th id="DA78601DFF92007613C2FB09D9AE49B5" box="[163,425,1181,1206]" gridcol="0" gridrow="22" pageId="14" pageNumber="343">Caudal peduncle depth</th>
<td id="DA78601DFF92007612A0FB09DA8249B5" box="[449,645,1181,1206]" gridcol="1" gridrow="22" pageId="14" pageNumber="343">10.6</td>
<td id="DA78601DFF92007611FDFB09DB5849B5" box="[668,863,1181,1206]" gridcol="2" gridrow="22" pageId="14" pageNumber="343">10.2–11.2</td>
<td id="DA78601DFF92007610E6FB09DC0449B5" box="[903,1027,1181,1206]" gridcol="3" gridrow="22" pageId="14" pageNumber="343">10.5–11.9</td>
<td id="DA78601DFF9200761756FB09DCCA49B5" box="[1079,1229,1181,1206]" gridcol="4" gridrow="22" pageId="14" pageNumber="343">11.2–11.7</td>
<td id="DA78601DFF9200761792FB09DD8849B5" box="[1267,1423,1181,1206]" gridcol="5" gridrow="22" pageId="14" pageNumber="343">10.8–11.3</td>
</tr>
<tr id="99A90961FF92007613C2FB51DD8849DD" box="[163,1423,1221,1246]" gridrow="23" pageId="14" pageNumber="343">
<th id="DA78601DFF92007613C2FB51D9AE49DD" box="[163,425,1221,1246]" gridcol="0" gridrow="23" pageId="14" pageNumber="343">Caudal peduncle length</th>
<td id="DA78601DFF92007612A0FB51DA8249DD" box="[449,645,1221,1246]" gridcol="1" gridrow="23" pageId="14" pageNumber="343">16.6</td>
<td id="DA78601DFF92007611FDFB51DB5849DD" box="[668,863,1221,1246]" gridcol="2" gridrow="23" pageId="14" pageNumber="343">15.7–17.1</td>
<td id="DA78601DFF92007610E6FB51DC0449DD" box="[903,1027,1221,1246]" gridcol="3" gridrow="23" pageId="14" pageNumber="343">15.4–17.4</td>
<td id="DA78601DFF9200761756FB51DCCA49DD" box="[1079,1229,1221,1246]" gridcol="4" gridrow="23" pageId="14" pageNumber="343">15.9–16.8</td>
<td id="DA78601DFF9200761792FB51DD8849DD" box="[1267,1423,1221,1246]" gridcol="5" gridrow="23" pageId="14" pageNumber="343">17.1–21.0</td>
</tr>
<tr id="99A90961FF92007613C2FB79DD884805" box="[163,1423,1261,1286]" gridrow="24" pageId="14" pageNumber="343">
<th id="DA78601DFF92007613C2FB79D9AE4805" box="[163,425,1261,1286]" gridcol="0" gridrow="24" pageId="14" pageNumber="343">Predorsal length</th>
<td id="DA78601DFF92007612A0FB79DA824805" box="[449,645,1261,1286]" gridcol="1" gridrow="24" pageId="14" pageNumber="343">36.7</td>
<td id="DA78601DFF92007611FDFB79DB584805" box="[668,863,1261,1286]" gridcol="2" gridrow="24" pageId="14" pageNumber="343">35.3–37.9</td>
<td id="DA78601DFF92007610E6FB79DC044805" box="[903,1027,1261,1286]" gridcol="3" gridrow="24" pageId="14" pageNumber="343">35.3–39.5</td>
<td id="DA78601DFF9200761756FB79DCCA4805" box="[1079,1229,1261,1286]" gridcol="4" gridrow="24" pageId="14" pageNumber="343">35.8–37.4</td>
<td id="DA78601DFF9200761792FB79DD884805" box="[1267,1423,1261,1286]" gridcol="5" gridrow="24" pageId="14" pageNumber="343">35.3–38.9</td>
</tr>
<tr id="99A90961FF92007613C2FA82DD88482C" box="[163,1423,1302,1327]" gridrow="25" pageId="14" pageNumber="343">
<th id="DA78601DFF92007613C2FA82D9AE482C" box="[163,425,1302,1327]" gridcol="0" gridrow="25" pageId="14" pageNumber="343">Preanal length</th>
<td id="DA78601DFF92007612A0FA82DA82482C" box="[449,645,1302,1327]" gridcol="1" gridrow="25" pageId="14" pageNumber="343">70.7</td>
<td id="DA78601DFF92007611FDFA82DB58482C" box="[668,863,1302,1327]" gridcol="2" gridrow="25" pageId="14" pageNumber="343">68.7–70.8</td>
<td id="DA78601DFF92007610E6FA82DC04482C" box="[903,1027,1302,1327]" gridcol="3" gridrow="25" pageId="14" pageNumber="343">68.4–69.4</td>
<td id="DA78601DFF9200761756FA82DCCA482C" box="[1079,1229,1302,1327]" gridcol="4" gridrow="25" pageId="14" pageNumber="343">68.0–70.0</td>
<td id="DA78601DFF9200761792FA82DD88482C" box="[1267,1423,1302,1327]" gridcol="5" gridrow="25" pageId="14" pageNumber="343">65.2–67.9</td>
</tr>
<tr id="99A90961FF92007613C2FAAADD884854" box="[163,1423,1342,1367]" gridrow="26" pageId="14" pageNumber="343">
<th id="DA78601DFF92007613C2FAAAD9AE4854" box="[163,425,1342,1367]" gridcol="0" gridrow="26" pageId="14" pageNumber="343">Prepelvic length</th>
<td id="DA78601DFF92007612A0FAAADA824854" box="[449,645,1342,1367]" gridcol="1" gridrow="26" pageId="14" pageNumber="343">36.5</td>
<td id="DA78601DFF92007611FDFAAADB584854" box="[668,863,1342,1367]" gridcol="2" gridrow="26" pageId="14" pageNumber="343">35.1–37.1</td>
<td id="DA78601DFF92007610E6FAAADC044854" box="[903,1027,1342,1367]" gridcol="3" gridrow="26" pageId="14" pageNumber="343">33.9–38.4</td>
<td id="DA78601DFF9200761756FAAADCCA4854" box="[1079,1229,1342,1367]" gridcol="4" gridrow="26" pageId="14" pageNumber="343">36.4–38.8</td>
<td id="DA78601DFF9200761792FAAADD884854" box="[1267,1423,1342,1367]" gridcol="5" gridrow="26" pageId="14" pageNumber="343">33.3–39.6</td>
</tr>
<tr id="99A90961FF92007613C2FAF2DD88487C" box="[163,1423,1382,1407]" gridrow="27" pageId="14" pageNumber="343">
<th id="DA78601DFF92007613C2FAF2D9AE487C" box="[163,425,1382,1407]" gridcol="0" gridrow="27" pageId="14" pageNumber="343">Dorsal-fin base</th>
<td id="DA78601DFF92007612A0FAF2DA82487C" box="[449,645,1382,1407]" gridcol="1" gridrow="27" pageId="14" pageNumber="343">52.5</td>
<td id="DA78601DFF92007611FDFAF2DB58487C" box="[668,863,1382,1407]" gridcol="2" gridrow="27" pageId="14" pageNumber="343">51.6–54.4</td>
<td id="DA78601DFF92007610E6FAF2DC04487C" box="[903,1027,1382,1407]" gridcol="3" gridrow="27" pageId="14" pageNumber="343">50.5–54.2</td>
<td id="DA78601DFF9200761756FAF2DCCA487C" box="[1079,1229,1382,1407]" gridcol="4" gridrow="27" pageId="14" pageNumber="343">53.0–54.9</td>
<td id="DA78601DFF9200761792FAF2DD88487C" box="[1267,1423,1382,1407]" gridcol="5" gridrow="27" pageId="14" pageNumber="343">53.0–55.4</td>
</tr>
<tr id="99A90961FF92007613C2FA1ADD8848A4" box="[163,1423,1422,1447]" gridrow="28" pageId="14" pageNumber="343">
<th id="DA78601DFF92007613C2FA1AD9AE48A4" box="[163,425,1422,1447]" gridcol="0" gridrow="28" pageId="14" pageNumber="343">First dorsal spine</th>
<td id="DA78601DFF92007612A0FA1ADA8248A4" box="[449,645,1422,1447]" gridcol="1" gridrow="28" pageId="14" pageNumber="343">4.7</td>
<td id="DA78601DFF92007611FDFA1ADB5848A4" box="[668,863,1422,1447]" gridcol="2" gridrow="28" pageId="14" pageNumber="343">4.1–5.3</td>
<td id="DA78601DFF92007610E6FA1ADC0448A4" box="[903,1027,1422,1447]" gridcol="3" gridrow="28" pageId="14" pageNumber="343">4.6–6.0</td>
<td id="DA78601DFF9200761756FA1ADCCA48A4" box="[1079,1229,1422,1447]" gridcol="4" gridrow="28" pageId="14" pageNumber="343">5.7–6.7</td>
<td id="DA78601DFF9200761792FA1ADD8848A4" box="[1267,1423,1422,1447]" gridcol="5" gridrow="28" pageId="14" pageNumber="343">5.2–5.9</td>
</tr>
<tr id="99A90961FF92007613C2FA22DD8848CC" box="[163,1423,1462,1487]" gridrow="29" pageId="14" pageNumber="343">
<th id="DA78601DFF92007613C2FA22D9AE48CC" box="[163,425,1462,1487]" gridcol="0" gridrow="29" pageId="14" pageNumber="343">Second dorsal spine</th>
<td id="DA78601DFF92007612A0FA22DA8248CC" box="[449,645,1462,1487]" gridcol="1" gridrow="29" pageId="14" pageNumber="343">8.7</td>
<td id="DA78601DFF92007611FDFA22DB5848CC" box="[668,863,1462,1487]" gridcol="2" gridrow="29" pageId="14" pageNumber="343">8.2–9.4</td>
<td id="DA78601DFF92007610E6FA22DC0448CC" box="[903,1027,1462,1487]" gridcol="3" gridrow="29" pageId="14" pageNumber="343">9.9–10.9</td>
<td id="DA78601DFF9200761756FA22DCCA48CC" box="[1079,1229,1462,1487]" gridcol="4" gridrow="29" pageId="14" pageNumber="343">10.9–12.0</td>
<td id="DA78601DFF9200761792FA22DD8848CC" box="[1267,1423,1462,1487]" gridcol="5" gridrow="29" pageId="14" pageNumber="343">8.5–11.1</td>
</tr>
<tr id="99A90961FF92007613C2FA4BDD8848FB" box="[163,1423,1503,1528]" gridrow="30" pageId="14" pageNumber="343">
<th id="DA78601DFF92007613C2FA4BD9AE48FB" box="[163,425,1503,1528]" gridcol="0" gridrow="30" pageId="14" pageNumber="343">Third dorsal spine</th>
<td id="DA78601DFF92007612A0FA4BDA8248FB" box="[449,645,1503,1528]" gridcol="1" gridrow="30" pageId="14" pageNumber="343">10.0</td>
<td id="DA78601DFF92007611FDFA4BDB5848FB" box="[668,863,1503,1528]" gridcol="2" gridrow="30" pageId="14" pageNumber="343">10.2–11.9</td>
<td id="DA78601DFF92007610E6FA4BDC0448FB" box="[903,1027,1503,1528]" gridcol="3" gridrow="30" pageId="14" pageNumber="343">11.7–12.8</td>
<td id="DA78601DFF9200761756FA4BDCCA48FB" box="[1079,1229,1503,1528]" gridcol="4" gridrow="30" pageId="14" pageNumber="343">13.3–14.5</td>
<td id="DA78601DFF9200761792FA4BDD8848FB" box="[1267,1423,1503,1528]" gridcol="5" gridrow="30" pageId="14" pageNumber="343">11.7–12.9</td>
</tr>
<tr id="99A90961FF92007613C2F993DD884B23" box="[163,1423,1543,1568]" gridrow="31" pageId="14" pageNumber="343">
<th id="DA78601DFF92007613C2F993D9AE4B23" box="[163,425,1543,1568]" gridcol="0" gridrow="31" pageId="14" pageNumber="343">Fourth dorsal spine</th>
<td id="DA78601DFF92007612A0F993DA824B23" box="[449,645,1543,1568]" gridcol="1" gridrow="31" pageId="14" pageNumber="343">10.7</td>
<td id="DA78601DFF92007611FDF993DB584B23" box="[668,863,1543,1568]" gridcol="2" gridrow="31" pageId="14" pageNumber="343">10.2–12.5</td>
<td id="DA78601DFF92007610E6F993DC044B23" box="[903,1027,1543,1568]" gridcol="3" gridrow="31" pageId="14" pageNumber="343">11.7–13.4</td>
<td id="DA78601DFF9200761756F993DCCA4B23" box="[1079,1229,1543,1568]" gridcol="4" gridrow="31" pageId="14" pageNumber="343">12.8–14.1</td>
<td id="DA78601DFF9200761792F993DD884B23" box="[1267,1423,1543,1568]" gridcol="5" gridrow="31" pageId="14" pageNumber="343">12.0–13.0</td>
</tr>
<tr id="99A90961FF92007613C2F9BBDD884B4B" box="[163,1423,1583,1608]" gridrow="32" pageId="14" pageNumber="343">
<th id="DA78601DFF92007613C2F9BBD9AE4B4B" box="[163,425,1583,1608]" gridcol="0" gridrow="32" pageId="14" pageNumber="343">Last dorsal spine</th>
<td id="DA78601DFF92007612A0F9BBDA824B4B" box="[449,645,1583,1608]" gridcol="1" gridrow="32" pageId="14" pageNumber="343">7.3</td>
<td id="DA78601DFF92007611FDF9BBDB584B4B" box="[668,863,1583,1608]" gridcol="2" gridrow="32" pageId="14" pageNumber="343">7.7–8.7</td>
<td id="DA78601DFF92007610E6F9BBDC044B4B" box="[903,1027,1583,1608]" gridcol="3" gridrow="32" pageId="14" pageNumber="343">9.0–10.7</td>
<td id="DA78601DFF9200761756F9BBDCCA4B4B" box="[1079,1229,1583,1608]" gridcol="4" gridrow="32" pageId="14" pageNumber="343">9.2–11.5</td>
<td id="DA78601DFF9200761792F9BBDD884B4B" box="[1267,1423,1583,1608]" gridcol="5" gridrow="32" pageId="14" pageNumber="343">7.5–9.5</td>
</tr>
<tr id="99A90961FF92007613C2F9C3DD884B72" box="[163,1423,1623,1649]" gridrow="33" pageId="14" pageNumber="343">
<th id="DA78601DFF92007613C2F9C3D9AE4B72" box="[163,425,1623,1649]" gridcol="0" gridrow="33" pageId="14" pageNumber="343">Longest dorsal ray</th>
<td id="DA78601DFF92007612A0F9C3DA824B72" box="[449,645,1623,1649]" gridcol="1" gridrow="33" pageId="14" pageNumber="343">12.8</td>
<td id="DA78601DFF92007611FDF9C3DB584B72" box="[668,863,1623,1649]" gridcol="2" gridrow="33" pageId="14" pageNumber="343">12.9–13.6</td>
<td id="DA78601DFF92007610E6F9C3DC044B72" box="[903,1027,1623,1649]" gridcol="3" gridrow="33" pageId="14" pageNumber="343">14.9–17.2</td>
<td id="DA78601DFF9200761756F9C3DCCA4B72" box="[1079,1229,1623,1649]" gridcol="4" gridrow="33" pageId="14" pageNumber="343">13.9–16.7</td>
<td id="DA78601DFF9200761792F9C3DD884B72" box="[1267,1423,1623,1649]" gridcol="5" gridrow="33" pageId="14" pageNumber="343">13.0–14.7</td>
</tr>
<tr id="99A90961FF92007613C2F914DD884B9A" box="[163,1423,1664,1689]" gridrow="34" pageId="14" pageNumber="343">
<th id="DA78601DFF92007613C2F914D9AE4B9A" box="[163,425,1664,1689]" gridcol="0" gridrow="34" pageId="14" pageNumber="343">Anal-fin base</th>
<td id="DA78601DFF92007612A0F914DA824B9A" box="[449,645,1664,1689]" gridcol="1" gridrow="34" pageId="14" pageNumber="343">14.9</td>
<td id="DA78601DFF92007611FDF914DB584B9A" box="[668,863,1664,1689]" gridcol="2" gridrow="34" pageId="14" pageNumber="343">14.9–16.5</td>
<td id="DA78601DFF92007610E6F914DC044B9A" box="[903,1027,1664,1689]" gridcol="3" gridrow="34" pageId="14" pageNumber="343">16.5–17.0</td>
<td id="DA78601DFF9200761756F914DCCA4B9A" box="[1079,1229,1664,1689]" gridcol="4" gridrow="34" pageId="14" pageNumber="343">15.4–17.4</td>
<td id="DA78601DFF9200761792F914DD884B9A" box="[1267,1423,1664,1689]" gridcol="5" gridrow="34" pageId="14" pageNumber="343">15.7–16.3</td>
</tr>
<tr id="99A90961FF92007613C2F93CDD884BC2" box="[163,1423,1704,1729]" gridrow="35" pageId="14" pageNumber="343">
<th id="DA78601DFF92007613C2F93CD9AE4BC2" box="[163,425,1704,1729]" gridcol="0" gridrow="35" pageId="14" pageNumber="343">First anal spine</th>
<td id="DA78601DFF92007612A0F93CDA824BC2" box="[449,645,1704,1729]" gridcol="1" gridrow="35" pageId="14" pageNumber="343">3.5</td>
<td id="DA78601DFF92007611FDF93CDB584BC2" box="[668,863,1704,1729]" gridcol="2" gridrow="35" pageId="14" pageNumber="343">3.5–4.0</td>
<td id="DA78601DFF92007610E6F93CDC044BC2" box="[903,1027,1704,1729]" gridcol="3" gridrow="35" pageId="14" pageNumber="343">4.0–4.9</td>
<td id="DA78601DFF9200761756F93CDCCA4BC2" box="[1079,1229,1704,1729]" gridcol="4" gridrow="35" pageId="14" pageNumber="343">3.8–5.9</td>
<td id="DA78601DFF9200761792F93CDD884BC2" box="[1267,1423,1704,1729]" gridcol="5" gridrow="35" pageId="14" pageNumber="343">3.2–4.5</td>
</tr>
<tr id="99A90961FF92007613C2F944DD884BEA" box="[163,1423,1744,1769]" gridrow="36" pageId="14" pageNumber="343">
<th id="DA78601DFF92007613C2F944D9AE4BEA" box="[163,425,1744,1769]" gridcol="0" gridrow="36" pageId="14" pageNumber="343">Second anal spine</th>
<td id="DA78601DFF92007612A0F944DA824BEA" box="[449,645,1744,1769]" gridcol="1" gridrow="36" pageId="14" pageNumber="343">6.8</td>
<td id="DA78601DFF92007611FDF944DB584BEA" box="[668,863,1744,1769]" gridcol="2" gridrow="36" pageId="14" pageNumber="343">7.4–8.4</td>
<td id="DA78601DFF92007610E6F944DC044BEA" box="[903,1027,1744,1769]" gridcol="3" gridrow="36" pageId="14" pageNumber="343">9.2–11.1</td>
<td id="DA78601DFF9200761756F944DCCA4BEA" box="[1079,1229,1744,1769]" gridcol="4" gridrow="36" pageId="14" pageNumber="343">8.0–11.6</td>
<td id="DA78601DFF9200761792F944DD884BEA" box="[1267,1423,1744,1769]" gridcol="5" gridrow="36" pageId="14" pageNumber="343">6.5–8.8</td>
</tr>
<tr id="99A90961FF92007613C2F96CDD884A12" box="[163,1423,1784,1809]" gridrow="37" pageId="14" pageNumber="343">
<th id="DA78601DFF92007613C2F96CD9AE4A12" box="[163,425,1784,1809]" gridcol="0" gridrow="37" pageId="14" pageNumber="343">Third anal spine</th>
<td id="DA78601DFF92007612A0F96CDA824A12" box="[449,645,1784,1809]" gridcol="1" gridrow="37" pageId="14" pageNumber="343">8.2</td>
<td id="DA78601DFF92007611FDF96CDB584A12" box="[668,863,1784,1809]" gridcol="2" gridrow="37" pageId="14" pageNumber="343">8.0–9.2</td>
<td id="DA78601DFF92007610E6F96CDC044A12" box="[903,1027,1784,1809]" gridcol="3" gridrow="37" pageId="14" pageNumber="343">9.8–11.2</td>
<td id="DA78601DFF9200761756F96CDCCA4A12" box="[1079,1229,1784,1809]" gridcol="4" gridrow="37" pageId="14" pageNumber="343">9.8–12.3</td>
<td id="DA78601DFF9200761792F96CDD884A12" box="[1267,1423,1784,1809]" gridcol="5" gridrow="37" pageId="14" pageNumber="343">7.8–9.5</td>
</tr>
<tr id="99A90961FF92007613C2F8B5DD884A39" box="[163,1423,1825,1850]" gridrow="38" pageId="14" pageNumber="343">
<th id="DA78601DFF92007613C2F8B5D9AE4A39" box="[163,425,1825,1850]" gridcol="0" gridrow="38" pageId="14" pageNumber="343">Longest anal soft ray</th>
<td id="DA78601DFF92007612A0F8B5DA824A39" box="[449,645,1825,1850]" gridcol="1" gridrow="38" pageId="14" pageNumber="343">14.1</td>
<td id="DA78601DFF92007611FDF8B5DB584A39" box="[668,863,1825,1850]" gridcol="2" gridrow="38" pageId="14" pageNumber="343">13.6–14.9</td>
<td id="DA78601DFF92007610E6F8B5DC044A39" box="[903,1027,1825,1850]" gridcol="3" gridrow="38" pageId="14" pageNumber="343">15.7–18.5</td>
<td id="DA78601DFF9200761756F8B5DCCA4A39" box="[1079,1229,1825,1850]" gridcol="4" gridrow="38" pageId="14" pageNumber="343">14.8–17.8</td>
<td id="DA78601DFF9200761792F8B5DD884A39" box="[1267,1423,1825,1850]" gridcol="5" gridrow="38" pageId="14" pageNumber="343">13.6–16.1</td>
</tr>
<tr id="99A90961FF92007613C2F8DDDD884A61" box="[163,1423,1865,1890]" gridrow="39" pageId="14" pageNumber="343">
<th id="DA78601DFF92007613C2F8DDD9AE4A61" box="[163,425,1865,1890]" gridcol="0" gridrow="39" pageId="14" pageNumber="343">Caudal-fin length</th>
<td id="DA78601DFF92007612A0F8DDDA824A61" box="[449,645,1865,1890]" gridcol="1" gridrow="39" pageId="14" pageNumber="343">18.1</td>
<td id="DA78601DFF92007611FDF8DDDB584A61" box="[668,863,1865,1890]" gridcol="2" gridrow="39" pageId="14" pageNumber="343">18.4–19.8</td>
<td id="DA78601DFF92007610E6F8DDDC044A61" box="[903,1027,1865,1890]" gridcol="3" gridrow="39" pageId="14" pageNumber="343">23.1–24.2</td>
<td id="DA78601DFF9200761756F8DDDCCA4A61" box="[1079,1229,1865,1890]" gridcol="4" gridrow="39" pageId="14" pageNumber="343">21.9–24.0</td>
<td id="DA78601DFF9200761792F8DDDD884A61" box="[1267,1423,1865,1890]" gridcol="5" gridrow="39" pageId="14" pageNumber="343">19.2–22.1</td>
</tr>
<tr id="99A90961FF92007613C2F8E5DD884A89" box="[163,1423,1905,1930]" gridrow="40" pageId="14" pageNumber="343">
<th id="DA78601DFF92007613C2F8E5D9AE4A89" box="[163,425,1905,1930]" gridcol="0" gridrow="40" pageId="14" pageNumber="343">Pectoral-fin length</th>
<td id="DA78601DFF92007612A0F8E5DA824A89" box="[449,645,1905,1930]" gridcol="1" gridrow="40" pageId="14" pageNumber="343">21.2</td>
<td id="DA78601DFF92007611FDF8E5DB584A89" box="[668,863,1905,1930]" gridcol="2" gridrow="40" pageId="14" pageNumber="343">21.1–24.2</td>
<td id="DA78601DFF92007610E6F8E5DC044A89" box="[903,1027,1905,1930]" gridcol="3" gridrow="40" pageId="14" pageNumber="343">22.3–25.5</td>
<td id="DA78601DFF9200761756F8E5DCCA4A89" box="[1079,1229,1905,1930]" gridcol="4" gridrow="40" pageId="14" pageNumber="343">22.3–24.9</td>
<td id="DA78601DFF9200761792F8E5DD884A89" box="[1267,1423,1905,1930]" gridcol="5" gridrow="40" pageId="14" pageNumber="343">20.3–23.9</td>
</tr>
<tr id="99A90961FF92007613C2F80DDD884AB1" box="[163,1423,1945,1970]" gridrow="41" pageId="14" pageNumber="343">
<th id="DA78601DFF92007613C2F80DD9AE4AB1" box="[163,425,1945,1970]" gridcol="0" gridrow="41" pageId="14" pageNumber="343">Pelvic-fin length</th>
<td id="DA78601DFF92007612A0F80DDA824AB1" box="[449,645,1945,1970]" gridcol="1" gridrow="41" pageId="14" pageNumber="343">15.8</td>
<td id="DA78601DFF92007611FDF80DDB584AB1" box="[668,863,1945,1970]" gridcol="2" gridrow="41" pageId="14" pageNumber="343">16.1–17.8</td>
<td id="DA78601DFF92007610E6F80DDC044AB1" box="[903,1027,1945,1970]" gridcol="3" gridrow="41" pageId="14" pageNumber="343">17.5–20.6</td>
<td id="DA78601DFF9200761756F80DDCCA4AB1" box="[1079,1229,1945,1970]" gridcol="4" gridrow="41" pageId="14" pageNumber="343">17.0–19.9</td>
<td id="DA78601DFF9200761792F80DDD884AB1" box="[1267,1423,1945,1970]" gridcol="5" gridrow="41" pageId="14" pageNumber="343">15.1–17.9</td>
</tr>
<tr id="99A90961FF92007613C2F856DD884AD8" box="[163,1423,1986,2011]" gridrow="42" pageId="14" pageNumber="343">
<th id="DA78601DFF92007613C2F856D9AE4AD8" box="[163,425,1986,2011]" gridcol="0" gridrow="42" pageId="14" pageNumber="343">Pelvic spine length</th>
<td id="DA78601DFF92007612A0F856DA824AD8" box="[449,645,1986,2011]" gridcol="1" gridrow="42" pageId="14" pageNumber="343">8.0</td>
<td id="DA78601DFF92007611FDF856DB584AD8" box="[668,863,1986,2011]" gridcol="2" gridrow="42" pageId="14" pageNumber="343">8.8–9.2</td>
<td id="DA78601DFF92007610E6F856DC044AD8" box="[903,1027,1986,2011]" gridcol="3" gridrow="42" pageId="14" pageNumber="343">10.3–11.4</td>
<td id="DA78601DFF9200761756F856DCCA4AD8" box="[1079,1229,1986,2011]" gridcol="4" gridrow="42" pageId="14" pageNumber="343">10.3–11.8</td>
<td id="DA78601DFF9200761792F856DD884AD8" box="[1267,1423,1986,2011]" gridcol="5" gridrow="42" pageId="14" pageNumber="343">8.4–10.2</td>
</tr>
</table>
</paragraph>
<paragraph id="27260B23FF93FF8613A6FF0DDBF24F84" blockId="15.[151,1437,153,2015]" pageId="15" pageNumber="344">
There are only minor differences in counts and proportional measurements between 
<taxonomicName id="E09970A0FF93FF861715FF0EDD4B4DB0" authorityName="Johnson &amp; Wilmer" authorityYear="2019" box="[1140,1356,153,179]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="344" phylum="Chordata" rank="species" species="fuscomarginatus">
<emphasis id="15EDD731FF93FF861715FF0EDD4B4DB0" box="[1140,1356,153,179]" italics="true" pageId="15" pageNumber="344">E. fuscomarginatus</emphasis>
</taxonomicName>
and its closest congeners. All five 
<typeStatus id="F822B581FF93FF8612A7FF29D9F14DD4" box="[454,502,189,215]" pageId="15" pageNumber="344">type</typeStatus>
specimens have 14 soft dorsal-fin rays and 17 pectoral fin rays, whereas 
<emphasis id="15EDD731FF93FF86164FFF2AD8BF4DF8" italics="true" pageId="15" pageNumber="344">
P. 
<taxonomicName id="E09970A0FF93FF86162BFF2AD8BF4DF8" authorityName="in Heemstra &amp; Randall" authorityYear="1993" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="344" phylum="Chordata" rank="species" species="epistictus">epistictus</taxonomicName>
</emphasis>
, 
<emphasis id="15EDD731FF93FF8613A4FF76D8D14DF8" box="[197,214,226,251]" italics="true" pageId="15" pageNumber="344">E</emphasis>
. cf 
<taxonomicName id="E09970A0FF93FF86139EFF76D96E4DF8" authorityName="in Heemstra &amp; Randall" authorityYear="1993" box="[255,361,226,251]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="344" phylum="Chordata" rank="species" species="epistictus">
<emphasis id="15EDD731FF93FF86139EFF76D96E4DF8" box="[255,361,226,251]" italics="true" pageId="15" pageNumber="344">epistictus</emphasis>
</taxonomicName>
and 
<taxonomicName id="E09970A0FF93FF8612FFFF76DA4D4DF8" box="[414,586,226,251]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="344" phylum="Chordata" rank="species" species="magniscuttis">
<emphasis id="15EDD731FF93FF8612FFFF76DA4D4DF8" box="[414,586,226,251]" italics="true" pageId="15" pageNumber="344">E. magniscuttis</emphasis>
</taxonomicName>
have 14–15 (usually 15) soft dorsal-fin rays and 17–19 (usually 18) pectoral fin rays (
<tableCitation id="6A1B3E98FF93FF86139BFE91D9564C1C" box="[250,337,261,287]" captionStart="TABLE 2" captionStartId="12.[151,244,152,179]" captionTargetBox="[163,1406,245,1097]" captionText="TABLE 2. Estimates of mean sequence divergence within (on diagonal) and between (below diagonal) species." httpUri="http://table.plazi.org/id/73E65BABFF90FF8513F6FF0CDD7E4DB0" pageId="15" pageNumber="344" tableUuid="73E65BABFF90FF8513F6FF0CDD7E4DB0">Table 2</tableCitation>
; 
<bibRefCitation id="430876D2FF93FF86123DFE91DA254C23" author="Katayama, M." box="[348,546,261,288]" pageId="15" pageNumber="344" refId="ref12417" refString="Katayama, M. (1960) Fauna Japonica. Vol. 1. Serranidae (Pisces). Biogeographical Society of Japan (Tokyo), 189 pp." type="book" year="1960">Katayama (1960)</bibRefCitation>
; 
<bibRefCitation id="430876D2FF93FF86114FFE91DAAB4C1C" author="Lee, S. - C." box="[558,684,261,287]" pageId="15" pageNumber="344" pagination="1 - 72" refId="ref12501" refString="Lee, S. - C. (1990) A revision of the serranid fish (family Serranidae) of Taiwan. Journal of Taiwan Museum, 43 (2), 1 - 72." type="journal article" year="1990">Lee (1990)</bibRefCitation>
; 
<bibRefCitation id="430876D2FF93FF8611D6FE91DBFC4C1C" author="Randall, J. E. &amp; Heemstra, P. C." box="[695,1019,261,287]" pageId="15" pageNumber="344" pagination="1 - 332" refId="ref12975" refString="Randall, J. E. &amp; Heemstra, P. C. (1991) Revision of Indo-Pacific groupers (Perciformes: Serranidae: Epinephelinae), with descriptions of five new species. Indo-Pacific Fishes, 20, 1 - 332." type="journal article" year="1991">Randall &amp; Heemstra (1991))</bibRefCitation>
. Lateral-line scale counts in 
<taxonomicName id="E09970A0FF93FF861658FE92D9114C40" authorityName="Johnson &amp; Wilmer" authorityYear="2019" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="344" phylum="Chordata" rank="species" species="fuscomarginatus">
<emphasis id="15EDD731FF93FF861658FE92D9114C40" italics="true" pageId="15" pageNumber="344">E. fuscomarginatus</emphasis>
</taxonomicName>
are similar to 
<taxonomicName id="E09970A0FF93FF8612D7FEBEDA384C40" authorityName="in Heemstra &amp; Randall" authorityYear="1993" box="[438,575,298,323]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="344" phylum="Chordata" rank="species" species="epistictus">
<emphasis id="15EDD731FF93FF8612D7FEBEDA384C40" box="[438,575,298,323]" italics="true" pageId="15" pageNumber="344">E. epistictus</emphasis>
</taxonomicName>
and 
<emphasis id="15EDD731FF93FF861115FEBEDA824C40" box="[628,645,298,323]" italics="true" pageId="15" pageNumber="344">E</emphasis>
. cf 
<taxonomicName id="E09970A0FF93FF8611CEFEBEDB1E4C40" authorityName="in Heemstra &amp; Randall" authorityYear="1993" box="[687,793,298,323]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="344" phylum="Chordata" rank="species" species="epistictus">
<emphasis id="15EDD731FF93FF8611CEFEBEDB1E4C40" box="[687,793,298,323]" italics="true" pageId="15" pageNumber="344">epistictus</emphasis>
</taxonomicName>
, but are somewhat higher than in 
<taxonomicName id="E09970A0FF93FF8617F2FEBEDD474C40" box="[1171,1344,298,323]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="344" phylum="Chordata" rank="species" species="magniscuttis">
<emphasis id="15EDD731FF93FF8617F2FEBEDD474C40" box="[1171,1344,298,323]" italics="true" pageId="15" pageNumber="344">E. magniscuttis</emphasis>
</taxonomicName>
(60–67, versus 55–62 (
<tableCitation id="6A1B3E98FF93FF86125DFED9D9934C64" box="[316,404,333,359]" captionStart="TABLE 2" captionStartId="12.[151,244,152,179]" captionTargetBox="[163,1406,245,1097]" captionText="TABLE 2. Estimates of mean sequence divergence within (on diagonal) and between (below diagonal) species." httpUri="http://table.plazi.org/id/73E65BABFF90FF8513F6FF0CDD7E4DB0" pageId="15" pageNumber="344" tableUuid="73E65BABFF90FF8513F6FF0CDD7E4DB0">Table 2</tableCitation>
; 
<bibRefCitation id="430876D2FF93FF8612C0FED9DAD44C64" author="Randall, J. E. &amp; Heemstra, P. C." box="[417,723,333,359]" pageId="15" pageNumber="344" pagination="1 - 332" refId="ref12975" refString="Randall, J. E. &amp; Heemstra, P. C. (1991) Revision of Indo-Pacific groupers (Perciformes: Serranidae: Epinephelinae), with descriptions of five new species. Indo-Pacific Fishes, 20, 1 - 332." type="journal article" year="1991">Randall &amp; Heemstra, 1991</bibRefCitation>
). The range of total and lower arch gill-rakers for 
<taxonomicName id="E09970A0FF93FF86166AFEDAD8EE4C88" authorityName="Johnson &amp; Wilmer" authorityYear="2019" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="344" phylum="Chordata" rank="species" species="fuscomarginatus">
<emphasis id="15EDD731FF93FF86166AFEDAD8EE4C88" italics="true" pageId="15" pageNumber="344">E. fuscomarginatus</emphasis>
</taxonomicName>
is slightly higher among our material examined than in the other three species (
<tableCitation id="6A1B3E98FF93FF861712FEE5DCCC4C88" box="[1139,1227,369,395]" captionStart="TABLE 2" captionStartId="12.[151,244,152,179]" captionTargetBox="[163,1406,245,1097]" captionText="TABLE 2. Estimates of mean sequence divergence within (on diagonal) and between (below diagonal) species." httpUri="http://table.plazi.org/id/73E65BABFF90FF8513F6FF0CDD7E4DB0" pageId="15" pageNumber="344" tableUuid="73E65BABFF90FF8513F6FF0CDD7E4DB0">Table 2</tableCitation>
). However, values for lower rakers presented by 
<bibRefCitation id="430876D2FF93FF861284FE01DB1C4CAC" author="Randall, J. E. &amp; Heemstra, P. C." box="[485,795,405,431]" pageId="15" pageNumber="344" pagination="1 - 332" refId="ref12975" refString="Randall, J. E. &amp; Heemstra, P. C. (1991) Revision of Indo-Pacific groupers (Perciformes: Serranidae: Epinephelinae), with descriptions of five new species. Indo-Pacific Fishes, 20, 1 - 332." type="journal article" year="1991">Randall &amp; Heemstra (1991)</bibRefCitation>
for 
<taxonomicName id="E09970A0FF93FF86102BFE02DBD44CAC" authorityName="in Heemstra &amp; Randall" authorityYear="1993" box="[842,979,406,431]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="344" phylum="Chordata" rank="species" species="epistictus">
<emphasis id="15EDD731FF93FF86102BFE02DBD44CAC" box="[842,979,406,431]" italics="true" pageId="15" pageNumber="344">E. epistictus</emphasis>
</taxonomicName>
(15–18) and 
<taxonomicName id="E09970A0FF93FF861709FE02DD134CAC" box="[1128,1300,406,431]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="344" phylum="Chordata" rank="species" species="magniscuttis">
<emphasis id="15EDD731FF93FF861709FE02DD134CAC" box="[1128,1300,406,431]" italics="true" pageId="15" pageNumber="344">E. magniscuttis</emphasis>
</taxonomicName>
(16–17) are slightly higher, hence we regard the apparent variation as unreliable given the relatively low number of specimens examined. It should also be noted that in large specimens of all the above species the lowermost rakers are typically reduced to rudiments or flattened platelets that are often closely associated with their neighbour. Accurate counts are thereby difficult and minor variations in lower gill raker counts should be treated with caution, especially when obtained from different observers. In specimens greater than 
<quantity id="E061A6C6FF93FF861025FDDDDBA54F60" box="[836,930,585,611]" metricMagnitude="-1" metricUnit="m" metricValue="4.0" pageId="15" pageNumber="344" unit="mm" value="400.0">400 mm</quantity>
SL 
<taxonomicName id="E09970A0FF93FF8610AEFDDEDCAE4F60" authorityName="Johnson &amp; Wilmer" authorityYear="2019" box="[975,1193,585,611]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="344" phylum="Chordata" rank="species" species="fuscomarginatus">
<emphasis id="15EDD731FF93FF8610AEFDDEDBE74F60" box="[975,992,586,611]" italics="true" pageId="15" pageNumber="344">E</emphasis>
. 
<emphasis id="15EDD731FF93FF86108EFDDDDCAE4F60" box="[1007,1193,585,611]" italics="true" pageId="15" pageNumber="344">fuscomarginatus</emphasis>
</taxonomicName>
has a shallower body depth than 
<taxonomicName id="E09970A0FF93FF861275FDFAD9C64F84" box="[276,449,622,647]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="344" phylum="Chordata" rank="species" species="magniscuttis">
<emphasis id="15EDD731FF93FF861275FDFAD9C64F84" box="[276,449,622,647]" italics="true" pageId="15" pageNumber="344">E. magniscuttis</emphasis>
</taxonomicName>
(depth in SL 3.0–3.4, versus 2.8–3.0, 
<tableCitation id="6A1B3E98FF93FF86100AFDF9DBCD4F84" box="[875,970,621,647]" captionStart="TABLE 3" captionStartId="14.[151,244,152,179]" captionTargetBox="[163,1423,266,2011]" captionText="TABLE 3. Meristic and morphological values for type specimens of Epinephelus fuscomarginatus sp. nov. and comparative material examined of E. epistictus, E. cf epistictus and E. magniscuttis (measurements as percentage of standard length). * Data for largest specimen of E. magniscuttis is from Nakamura et al. (2018)." httpUri="http://table.plazi.org/id/73E65BABFF92FF8713F6FF0CDCCD4DF8" pageId="15" pageNumber="344" tableUuid="73E65BABFF92FF8713F6FF0CDCCD4DF8">Tables 3</tableCitation>
, 
<tableCitation id="6A1B3E98FF93FF8610B6FDF9DBEF4F84" box="[983,1000,621,647]" captionStart="TABLE 4" captionStartId="17.[151,244,764,791]" captionTargetBox="[163,1398,878,1467]" captionText="TABLE 4. Selected meristic data, morphometric ratios and morphological characters for Epinephelus fuscomarginatus sp. nov., E. epistictus, E. cf epistictus and E. magniscuttis from material examined. * Data for largest specimen of E. magniscuttis based on Nakamura et al. (2018)." httpUri="http://table.plazi.org/id/73E65BABFF8DFF9813F6FD68DAE74E5C" pageId="15" pageNumber="344" tableUuid="73E65BABFF8DFF9813F6FD68DAE74E5C">4</tableCitation>
).
</paragraph>
<paragraph id="27260B23FF93FF8613A6FD05DD504958" blockId="15.[151,1437,153,2015]" pageId="15" pageNumber="344">
<bibRefCitation id="430876D2FF93FF8613A6FD05DA074FA8" author="Randall, J. E. &amp; Heemstra, P. C." box="[199,512,657,683]" pageId="15" pageNumber="344" pagination="1 - 332" refId="ref12975" refString="Randall, J. E. &amp; Heemstra, P. C. (1991) Revision of Indo-Pacific groupers (Perciformes: Serranidae: Epinephelinae), with descriptions of five new species. Indo-Pacific Fishes, 20, 1 - 332." type="journal article" year="1991">Randall &amp; Heemstra (1991)</bibRefCitation>
provided ratios for longest dorsal-fin spine in HL and pelvic fin in HL in 
<taxonomicName id="E09970A0FF93FF861622FD06D8F34FCC" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="344" phylum="Chordata" rank="species" species="magniscuttis">
<emphasis id="15EDD731FF93FF861622FD06D8F34FCC" italics="true" pageId="15" pageNumber="344">E. magniscuttis</emphasis>
</taxonomicName>
as 2.45–3.0 and 2.1–2.35, respectively. This data has also been quoted by several subsequent authors (e.g. 
<bibRefCitation id="430876D2FF93FF8613F6FD4DD9CD4FF0" author="Heemstra, P. C. &amp; Randall, J. E." box="[151,458,729,755]" pageId="15" pageNumber="344" refId="ref12210" refString="Heemstra, P. C. &amp; Randall, J. E. (1993) Groupers of the World (Family Serranidae, Subfamily Epinephelinae); an annotated and illustrated catalogue of the grouper, rockcod, hind, coral grouper and lyretail species known to date. FAO Species Catalog 16. FAO, Rome, 382 pp." type="book" year="1993">Heemstra &amp; Randall, 1993</bibRefCitation>
; 
<bibRefCitation id="430876D2FF93FF8612B6FD4DDAA54FF0" author="Craig, M. T. &amp; Sadovy de Mitcheson, Y. J. &amp; Heemstra, P. C." box="[471,674,729,755]" pageId="15" pageNumber="344" refId="ref11830" refString="Craig, M. T., Sadovy de Mitcheson, Y. J. &amp; Heemstra, P. C. (2011) Groupers of the world. A field and market guide. NISC Pty Ltd, Grahamstown, 356 pp." type="book" year="2011">
Craig 
<emphasis id="15EDD731FF93FF86117CFD4EDA574FF0" box="[541,592,729,755]" italics="true" pageId="15" pageNumber="344">et al</emphasis>
., 2011
</bibRefCitation>
). It suggests significant differences between 
<taxonomicName id="E09970A0FF93FF8617FCFD4EDD4B4FF0" box="[1181,1356,730,755]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="344" phylum="Chordata" rank="species" species="magniscuttis">
<emphasis id="15EDD731FF93FF8617FCFD4EDD4B4FF0" box="[1181,1356,730,755]" italics="true" pageId="15" pageNumber="344">E. magniscuttis</emphasis>
</taxonomicName>
and 
<taxonomicName id="E09970A0FF93FF8616E5FD4ED9564E14" authorityName="Johnson &amp; Wilmer" authorityYear="2019" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="344" phylum="Chordata" rank="species" species="fuscomarginatus">
<emphasis id="15EDD731FF93FF8616E5FD4ED9564E14" italics="true" pageId="15" pageNumber="344">E. fuscomarginatus</emphasis>
</taxonomicName>
, with longest dorsal-fin spine in HL 3.3–4.0 and pelvic fin in HL 2.4–2.6 applying in the latter. However, Randall &amp; Heemstra’s data for 
<taxonomicName id="E09970A0FF93FF86110CFCB6DB1C4E38" box="[621,795,802,827]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="344" phylum="Chordata" rank="species" species="magniscuttis">
<emphasis id="15EDD731FF93FF86110CFCB6DB1C4E38" box="[621,795,802,827]" italics="true" pageId="15" pageNumber="344">E. magniscuttis</emphasis>
</taxonomicName>
is at odds with that presented here (3.3–3.5 and 2.4–2.8, respectively, 
<tableCitation id="6A1B3E98FF93FF86124BFCD1D9784E5C" box="[298,383,837,863]" captionStart="TABLE 4" captionStartId="17.[151,244,764,791]" captionTargetBox="[163,1398,878,1467]" captionText="TABLE 4. Selected meristic data, morphometric ratios and morphological characters for Epinephelus fuscomarginatus sp. nov., E. epistictus, E. cf epistictus and E. magniscuttis from material examined. * Data for largest specimen of E. magniscuttis based on Nakamura et al. (2018)." httpUri="http://table.plazi.org/id/73E65BABFF8DFF9813F6FD68DAE74E5C" pageId="15" pageNumber="344" tableUuid="73E65BABFF8DFF9813F6FD68DAE74E5C">Table 4</tableCitation>
), where no significant difference in these ratios is indicated between similar-sized individuals of the two species. Differences between our data for 
<taxonomicName id="E09970A0FF93FF861181FCFEDB8B4E80" box="[736,908,874,899]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="344" phylum="Chordata" rank="species" species="magniscuttis">
<emphasis id="15EDD731FF93FF861181FCFEDB8B4E80" box="[736,908,874,899]" italics="true" pageId="15" pageNumber="344">E. magniscuttis</emphasis>
</taxonomicName>
and that of 
<bibRefCitation id="430876D2FF93FF86176EFCFDDD424E80" author="Randall, J. E. &amp; Heemstra, P. C." box="[1039,1349,873,899]" pageId="15" pageNumber="344" pagination="1 - 332" refId="ref12975" refString="Randall, J. E. &amp; Heemstra, P. C. (1991) Revision of Indo-Pacific groupers (Perciformes: Serranidae: Epinephelinae), with descriptions of five new species. Indo-Pacific Fishes, 20, 1 - 332." type="journal article" year="1991">Randall &amp; Heemstra (1991)</bibRefCitation>
may be most readily explained by their significantly smaller range of specimens examined (
<quantity id="E061A6C6FF93FF861729FC19DCD84EA4" box="[1096,1247,909,935]" metricMagnitude="-1" metricUnit="m" metricValue="2.75" metricValueMax="4.18" metricValueMin="1.3199999999999998" pageId="15" pageNumber="344" unit="mm" value="275.0" valueMax="418.0" valueMin="132.0">132–418 mm</quantity>
SL, versus 
<quantity id="E061A6C6FF93FF861605FC19D8F24EC8" metricMagnitude="-1" metricUnit="m" metricValue="5.16" metricValueMax="6.13" metricValueMin="4.1899999999999995" pageId="15" pageNumber="344" unit="mm" value="516.0" valueMax="613.0" valueMin="419.0">419– 613 mm</quantity>
SL in this study). Minor variations in several proportional measurements between 
<taxonomicName id="E09970A0FF93FF8617FDFC26DD714EC8" authorityName="Johnson &amp; Wilmer" authorityYear="2019" box="[1180,1398,945,971]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="344" phylum="Chordata" rank="species" species="fuscomarginatus">
<emphasis id="15EDD731FF93FF8617FDFC26DD714EC8" box="[1180,1398,945,971]" italics="true" pageId="15" pageNumber="344">E. fuscomarginatus</emphasis>
</taxonomicName>
, 
<taxonomicName id="E09970A0FF93FF8616E5FC26D9064EEC" authorityName="in Heemstra &amp; Randall" authorityYear="1993" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="344" phylum="Chordata" rank="species" species="epistictus">
<emphasis id="15EDD731FF93FF8616E5FC26D9064EEC" italics="true" pageId="15" pageNumber="344">E. epistictus</emphasis>
</taxonomicName>
and 
<emphasis id="15EDD731FF93FF861254FC42D9414EEC" box="[309,326,982,1007]" italics="true" pageId="15" pageNumber="344">E</emphasis>
. cf 
<taxonomicName id="E09970A0FF93FF86120EFC42D9DE4EEC" authorityName="in Heemstra &amp; Randall" authorityYear="1993" box="[367,473,982,1007]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="344" phylum="Chordata" rank="species" species="epistictus">
<emphasis id="15EDD731FF93FF86120EFC42D9DE4EEC" box="[367,473,982,1007]" italics="true" pageId="15" pageNumber="344">epistictus</emphasis>
</taxonomicName>
(
<tableCitation id="6A1B3E98FF93FF861286FC41DA5F4EEC" box="[487,600,981,1007]" captionStart-0="TABLE 3" captionStart-1="TABLE 4" captionStartId-0="14.[151,244,152,179]" captionStartId-1="17.[151,244,764,791]" captionTargetBox-0="[163,1423,266,2011]" captionTargetBox-1="[163,1398,878,1467]" captionTargetPageId-0="14" captionText-0="TABLE 3. Meristic and morphological values for type specimens of Epinephelus fuscomarginatus sp. nov. and comparative material examined of E. epistictus, E. cf epistictus and E. magniscuttis (measurements as percentage of standard length). * Data for largest specimen of E. magniscuttis is from Nakamura et al. (2018)." captionText-1="TABLE 4. Selected meristic data, morphometric ratios and morphological characters for Epinephelus fuscomarginatus sp. nov., E. epistictus, E. cf epistictus and E. magniscuttis from material examined. * Data for largest specimen of E. magniscuttis based on Nakamura et al. (2018)." httpUri-0="http://table.plazi.org/id/73E65BABFF92FF8713F6FF0CDCCD4DF8" httpUri-1="http://table.plazi.org/id/73E65BABFF8DFF9813F6FD68DAE74E5C" pageId="15" pageNumber="344" tableUuid-0="73E65BABFF92FF8713F6FF0CDCCD4DF8" tableUuid-1="73E65BABFF8DFF9813F6FD68DAE74E5C">Table 3–4</tableCitation>
) are not regarded as reliable diagnostically, due to wide variation in lengths of specimens examined between the species. Specimens of 
<taxonomicName id="E09970A0FF93FF86104DFC6EDBB24910" authorityName="in Heemstra &amp; Randall" authorityYear="1993" box="[812,949,1018,1043]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="344" phylum="Chordata" rank="species" species="epistictus">
<emphasis id="15EDD731FF93FF86104DFC6EDBB24910" box="[812,949,1018,1043]" italics="true" pageId="15" pageNumber="344">E. epistictus</emphasis>
</taxonomicName>
and 
<emphasis id="15EDD731FF93FF86108BFC6EDBFC4910" box="[1002,1019,1018,1043]" italics="true" pageId="15" pageNumber="344">E</emphasis>
. cf 
<taxonomicName id="E09970A0FF93FF861744FC6EDC884910" authorityName="in Heemstra &amp; Randall" authorityYear="1993" box="[1061,1167,1018,1043]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="344" phylum="Chordata" rank="species" species="epistictus">
<emphasis id="15EDD731FF93FF861744FC6EDC884910" box="[1061,1167,1018,1043]" italics="true" pageId="15" pageNumber="344">epistictus</emphasis>
</taxonomicName>
<quantity id="E061A6C6FF93FF8617F4FC6DDD2D4910" box="[1173,1322,1017,1043]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="15" pageNumber="344" unit="mm" value="500.0" valueMax="600.0" valueMin="400.0">400–600 mm</quantity>
SL would be required for examination to determine if there are significant differences in proportional measurements between the species, but specimens of this size are generally rare or absent in international ichthyological collections.
</paragraph>
<paragraph id="27260B23FF93FF8613A6FBF1DA724A28" blockId="15.[151,1437,153,2015]" pageId="15" pageNumber="344">
Among other species of 
<taxonomicName id="E09970A0FF93FF8612BBFBF1DA63497C" authorityName="Bloch" authorityYear="1793" box="[474,612,1125,1151]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="344" phylum="Chordata" rank="genus">
<emphasis id="15EDD731FF93FF8612BBFBF1DA63497C" box="[474,612,1125,1151]" italics="true" pageId="15" pageNumber="344">Epinephelus</emphasis>
</taxonomicName>
with a relatively uniform body colouration and few distinctive markings, 
<taxonomicName id="E09970A0FF93FF8613F6FB1ED97749A0" authorityName="Johnson &amp; Wilmer" authorityYear="2019" box="[151,368,1161,1187]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="344" phylum="Chordata" rank="species" species="fuscomarginatus">
<emphasis id="15EDD731FF93FF8613F6FB1ED97749A0" box="[151,368,1161,1187]" italics="true" pageId="15" pageNumber="344">E. fuscomarginatus</emphasis>
</taxonomicName>
differs from 
<taxonomicName id="E09970A0FF93FF861162FB1EDA9349A0" baseAuthorityName="non Fowler" baseAuthorityYear="1904" box="[515,660,1161,1187]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="344" phylum="Chordata" rank="species" species="heniochus">
<emphasis id="15EDD731FF93FF861162FB1EDA9349A0" box="[515,660,1161,1187]" italics="true" pageId="15" pageNumber="344">E. heniochus</emphasis>
</taxonomicName>
(fig. 5B) by higher lateral-line scale and gill raker counts (60–67 and 9–10 + 16–19 = 25–28, versus 54–62 and 7–9 + 14–16 = 21–25, respectively), a much larger maximum size (at least 
<quantity id="E061A6C6FF93FF8613F6FB44D8F249E8" box="[151,245,1232,1259]" metricMagnitude="-1" metricUnit="m" metricValue="5.64" pageId="15" pageNumber="344" unit="mm" value="564.0">564 mm</quantity>
SL, versus 
<quantity id="E061A6C6FF93FF86121AFB44D9DE49E8" box="[379,473,1232,1259]" metricMagnitude="-1" metricUnit="m" metricValue="3.5" pageId="15" pageNumber="344" unit="mm" value="350.0">350 mm</quantity>
SL), the lack of any small dark spots on the head and body at any known size (small yellowish brown dots sometimes present on the head and body, but often absent in large specimens and those from northwestern 
<collectingCountry id="5F8E4BB3FF93FF86124FFA8DD99F4830" box="[302,408,1305,1331]" name="Australia" pageId="15" pageNumber="344">Australia</collectingCountry>
, in 
<taxonomicName id="E09970A0FF93FF8612DFFA8EDA494830" baseAuthorityName="non Fowler" baseAuthorityYear="1904" box="[446,590,1305,1331]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="344" phylum="Chordata" rank="species" species="heniochus">
<emphasis id="15EDD731FF93FF8612DFFA8EDA494830" box="[446,590,1305,1331]" italics="true" pageId="15" pageNumber="344">E. heniochus</emphasis>
</taxonomicName>
) and spinous dorsal-fin membrane dark brown adjacent to distal tip of spine and semitransparent posteriorly, versus distal part of membrane pale yellowish in 
<taxonomicName id="E09970A0FF93FF861721FAAADCD44854" baseAuthorityName="non Fowler" baseAuthorityYear="1904" box="[1088,1235,1341,1367]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="344" phylum="Chordata" rank="species" species="heniochus">
<emphasis id="15EDD731FF93FF861721FAAADCD44854" box="[1088,1235,1341,1367]" italics="true" pageId="15" pageNumber="344">E. heniochus</emphasis>
</taxonomicName>
. Another grouper lacking distinctive markings, 
<taxonomicName id="E09970A0FF93FF861285FAF5DAEC4878" baseAuthorityName="Randall, Hoese &amp; Last" baseAuthorityYear="1991" box="[484,747,1377,1403]" class="Actinopterygii" family="Serranidae" genus="Hyporthodus" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="344" phylum="Chordata" rank="species" species="perplexus">
<emphasis id="15EDD731FF93FF861285FAF5DAEC4878" box="[484,747,1377,1403]" italics="true" pageId="15" pageNumber="344">Hyporthodus perplexus</emphasis>
</taxonomicName>
, known only from a single 
<quantity id="E061A6C6FF93FF861742FAF4DC874878" box="[1059,1152,1376,1403]" metricMagnitude="-1" metricUnit="m" metricValue="4.65" pageId="15" pageNumber="344" unit="mm" value="465.0">465 mm</quantity>
SL specimen (
<bibRefCitation id="430876D2FF93FF861649FAF5D8F8489C" author="Randall, J. E. &amp; Hoese, D. F. &amp; Last, P. R." pageId="15" pageNumber="344" pagination="25 - 31" refId="ref13019" refString="Randall, J. E., Hoese, D. F. &amp; Last, P. R. (1993) On the status of the Australian serranid fishes Epinephelus ergastularius Whitley and E. thompsoni Whitley. Records of the Australian Museum, 45, 25 - 31. https: // doi. org / 10.3853 / j. 0067 - 1975.45.1993.127" type="journal article" year="1993">
Randall 
<emphasis id="15EDD731FF93FF8616E9FAF6D8AA489C" italics="true" pageId="15" pageNumber="344">et al</emphasis>
., 1993
</bibRefCitation>
), has fewer soft dorsal-fin rays and lateral-line scales (13 and 52, versus 14 and 60–67, respectively), a greater number pectoral-fin rays (18, versus 17), a deeper body (depth 
<quantity id="E061A6C6FF93FF8610CFFA3DDBE948C0" box="[942,1006,1449,1475]" metricMagnitude="-2" metricUnit="m" metricValue="7.366" pageId="15" pageNumber="344" unit="in" value="2.9">2.9 in</quantity>
HL, versus 3.0–3.4) and a proportionately wider posterior nostril (about twice diameter of anterior nostril, versus 1.3–1.7). Despite a large amount of fishing effort over many years around the region of the 
<typeStatus id="F822B581FF93FF861071FA65DB474B08" box="[784,832,1521,1547]" pageId="15" pageNumber="344">type</typeStatus>
locality of 
<taxonomicName id="E09970A0FF93FF8610A2FA66DC534B08" baseAuthorityName="Randall, Hoese &amp; Last" baseAuthorityYear="1991" box="[963,1108,1521,1547]" class="Actinopterygii" family="Serranidae" genus="Hyporthodus" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="344" phylum="Chordata" rank="species" species="perplexus">
<emphasis id="15EDD731FF93FF8610A2FA66DC534B08" box="[963,1108,1521,1547]" italics="true" pageId="15" pageNumber="344">H. perplexus</emphasis>
</taxonomicName>
(northeast of Cape Moreton, Qld, 
<collectingCountry id="5F8E4BB3FF93FF8613AFF981D93F4B2C" box="[206,312,1557,1583]" name="Australia" pageId="15" pageNumber="344">Australia</collectingCountry>
, in 
<quantity id="E061A6C6FF93FF861201F981D9E54B2C" box="[352,482,1557,1583]" metricMagnitude="2" metricUnit="m" metricValue="1.325" metricValueMax="1.3699999999999999" metricValueMin="1.28" pageId="15" pageNumber="344" unit="m" value="132.5" valueMax="137.0" valueMin="128.0">128–137 m</quantity>
), no specimens or photographs of it or 
<taxonomicName id="E09970A0FF93FF8610FBF982DC744B2C" authorityName="Johnson &amp; Wilmer" authorityYear="2019" box="[922,1139,1557,1583]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="344" phylum="Chordata" rank="species" species="fuscomarginatus">
<emphasis id="15EDD731FF93FF8610FBF982DC744B2C" box="[922,1139,1557,1583]" italics="true" pageId="15" pageNumber="344">E. fuscomarginatus</emphasis>
</taxonomicName>
have been reported by the numerous commercial and recreational fishers that frequent this area, or the intervening area north to off the Capricorn-Bunker Group where is latter is known. Other deep water species, 
<taxonomicName id="E09970A0FF93FF8610CCF9CADC2B4B74" baseAuthorityName="Valenciennes" baseAuthorityYear="1833" box="[941,1068,1629,1655]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="344" phylum="Chordata" rank="species" species="morrhua">
<emphasis id="15EDD731FF93FF8610CCF9CADC2B4B74" box="[941,1068,1629,1655]" italics="true" pageId="15" pageNumber="344">E. morrhua</emphasis>
</taxonomicName>
, 
<taxonomicName id="E09970A0FF93FF861758F9CADCE14B74" baseAuthorityName="Temminck &amp; Schlegel" baseAuthorityYear="1842" box="[1081,1254,1629,1655]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="344" phylum="Chordata" rank="species" species="poecilonotus">
<emphasis id="15EDD731FF93FF861758F9CADCE14B74" box="[1081,1254,1629,1655]" italics="true" pageId="15" pageNumber="344">E. poecilonotus</emphasis>
</taxonomicName>
and 
<taxonomicName id="E09970A0FF93FF86167BF9CADD924B74" baseAuthorityName="Day" baseAuthorityYear="1868" box="[1306,1429,1629,1655]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="344" phylum="Chordata" rank="species" species="radiatus">
<emphasis id="15EDD731FF93FF86167BF9CADD924B74" box="[1306,1429,1629,1655]" italics="true" pageId="15" pageNumber="344">E. radiatus</emphasis>
</taxonomicName>
, share similar fin ray, gill raker, lateral-line scale and lateral tooth row counts, but are readily distinguished by the distinctive markings on the sides of their bodies. They also have fewer, more enlarged serrae on the angle of the preopercle (2–5 distinctly enlarged spines, versus 4–9 non-prominent serrae in 
<taxonomicName id="E09970A0FF93FF861779F95EDCF54BE0" authorityName="Johnson &amp; Wilmer" authorityYear="2019" box="[1048,1266,1737,1763]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="344" phylum="Chordata" rank="species" species="fuscomarginatus">
<emphasis id="15EDD731FF93FF861779F95EDCF54BE0" box="[1048,1266,1737,1763]" italics="true" pageId="15" pageNumber="344">E. fuscomarginatus</emphasis>
</taxonomicName>
) and generally have longer heads (HL 
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SL, versus 
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<taxonomicName id="E09970A0FF93FF861077F97ADBE84A04" authorityName="Johnson &amp; Wilmer" authorityYear="2019" box="[790,1007,1773,1799]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="344" phylum="Chordata" rank="species" species="fuscomarginatus">
<emphasis id="15EDD731FF93FF861077F97ADBE84A04" box="[790,1007,1773,1799]" italics="true" pageId="15" pageNumber="344">E. fuscomarginatus</emphasis>
</taxonomicName>
) and deeper bodies (depth 
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SL, versus 3.0– 
<quantity id="E061A6C6FF93FF861227F885D9824A28" box="[326,389,1809,1835]" metricMagnitude="-2" metricUnit="m" metricValue="8.636" pageId="15" pageNumber="344" unit="in" value="3.4">3.4 in</quantity>
<taxonomicName id="E09970A0FF93FF8612EDF886DA624A28" authorityName="Johnson &amp; Wilmer" authorityYear="2019" box="[396,613,1809,1835]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="344" phylum="Chordata" rank="species" species="fuscomarginatus">
<emphasis id="15EDD731FF93FF8612EDF886DA624A28" box="[396,613,1809,1835]" italics="true" pageId="15" pageNumber="344">E. fuscomarginatus</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="27260B23FF93FF9913A6F8A1DD8F4F49" blockId="15.[151,1437,153,2015]" lastBlockId="16.[151,1437,163,586]" lastPageId="16" lastPageNumber="345" pageId="15" pageNumber="344">
During examination of available specimens and photographs of 
<taxonomicName id="E09970A0FF93FF8610EAF8A2DC304A4C" box="[907,1079,1846,1871]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="344" phylum="Chordata" rank="species" species="magniscuttis">
<emphasis id="15EDD731FF93FF8610EAF8A2DC304A4C" box="[907,1079,1846,1871]" italics="true" pageId="15" pageNumber="344">E. magniscuttis</emphasis>
</taxonomicName>
, considerable intraspecific variation was noted in the size and number of spots between similar-sized individuals from differing localities. Spots in specimens from Western Australia (WAM P.26610-001, 
<quantity id="E061A6C6FF93FF861051F8E8DB894A94" box="[816,910,1916,1943]" metricMagnitude="-1" metricUnit="m" metricValue="5.58" pageId="15" pageNumber="344" unit="mm" value="558.0">558 mm</quantity>
SL, WAM P.34888-001, 
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SL (fig. 6A) and images of four fish 
<quantity id="E061A6C6FF93FF8612C3F835DA164AB8" box="[418,529,1953,1979]" metricMagnitude="-1" metricUnit="m" metricValue="6.5" metricValueMax="7.0" metricValueMin="6.0" pageId="15" pageNumber="344" unit="cm" value="65.0" valueMax="70.0" valueMin="60.0">60–70 cm</quantity>
TL (e.g. fig. 6B) supplied by C. Buckingham from off Exmouth, WA, in 
<quantity id="E061A6C6FF93FF861624F834DD884AB8" box="[1349,1423,1952,1979]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="15" pageNumber="344" unit="m" value="250.0">250 m</quantity>
), although distributed in roughly similar areas of the body and fins, were much smaller and more sparse, especially on the head, soft dorsal and caudal fins, than in a specimen from 
<collectingCountry id="5F8E4BB3FF8CFF99100DFF30DBA84DBD" box="[876,943,164,190]" name="Japan" pageId="16" pageNumber="345">Japan</collectingCountry>
, KAUM I. 
<quantity id="E061A6C6FF8CFF991742FF37DCD04DBD" box="[1059,1239,163,190]" metricMagnitude="4" metricUnit="m" metricValue="7.356061299999999" pageId="16" pageNumber="345" unit="mm" value="7.3560613E7">73560, 613 mm</quantity>
SL (
<bibRefCitation id="430876D2FF8CFF99166AFF30D8FB4DE1" author="Nakamura, J. &amp; Takayama, M. &amp; Worthington Wilmer, J. &amp; Johnson, J. W. &amp; Motomura, H." pageId="16" pageNumber="345" pagination="225 - 228" refId="ref12706" refString="Nakamura, J., Takayama, M., Worthington Wilmer, J., Johnson, J. W. &amp; Motomura, H. (2018) First Japanese record of the Speckled Grouper Epinephelus magniscuttis (Perciformes: Serranidae) from the Osumi Islands. Species Diversity, 23 (2), 225 - 228. https: // doi. org / 10.12782 / specdiv. 23.225" type="journal article" year="2018">
Nakamura 
<emphasis id="15EDD731FF8CFF9916E9FF31D8AA4DE1" italics="true" pageId="16" pageNumber="345">et al</emphasis>
., 2018
</bibRefCitation>
, fig. 1). The Western Australian specimen that was genetically sampled (WAM P.34888-001) differed only by a single base pair to the Japanese specimen across the DNA barcoding fragment. As noted, the level of sequence divergence difference between 
<taxonomicName id="E09970A0FF8CFF99129AFE85DAD04C29" authorityName="Johnson &amp; Wilmer" authorityYear="2019" box="[507,727,272,298]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="345" phylum="Chordata" rank="species" species="fuscomarginatus">
<emphasis id="15EDD731FF8CFF99129AFE85DAD04C29" box="[507,727,272,298]" italics="true" pageId="16" pageNumber="345">E. fuscomarginatus</emphasis>
</taxonomicName>
and 
<taxonomicName id="E09970A0FF8CFF991070FE85DBC64C29" box="[785,961,273,298]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="345" phylum="Chordata" rank="species" species="magniscuttis">
<emphasis id="15EDD731FF8CFF991070FE85DBC64C29" box="[785,961,273,298]" italics="true" pageId="16" pageNumber="345">E. magniscuttis</emphasis>
</taxonomicName>
is low (1.10%). The number of available 
<taxonomicName id="E09970A0FF8CFF9913F6FEA1D9404C4D" box="[151,327,309,334]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="345" phylum="Chordata" rank="species" species="magniscuttis">
<emphasis id="15EDD731FF8CFF9913F6FEA1D9404C4D" box="[151,327,309,334]" italics="true" pageId="16" pageNumber="345">E. magniscuttis</emphasis>
</taxonomicName>
sequences is very small (n = 3), including one specimen from 
<collectingCountry id="5F8E4BB3FF8CFF99174BFEA0DC7B4C4D" box="[1066,1148,308,334]" name="Taiwan" pageId="16" pageNumber="345">Taiwan</collectingCountry>
(
<accessionNumber id="38CA96C0FF8CFF9917EFFEA0DD0A4C4D" box="[1166,1293,308,334]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KU893048" pageId="16" pageNumber="345">KU893048</accessionNumber>
) taken from GenBank identified as 
<taxonomicName id="E09970A0FF8CFF9912FFFECDDA2E4C71" authorityName="in Heemstra &amp; Randall" authorityYear="1993" box="[414,553,345,370]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="345" phylum="Chordata" rank="species" species="epistictus">
<emphasis id="15EDD731FF8CFF9912FFFECDDA2E4C71" box="[414,553,345,370]" italics="true" pageId="16" pageNumber="345">E. epistictus</emphasis>
</taxonomicName>
, but with only another single base pair difference from other 
<taxonomicName id="E09970A0FF8CFF99178FFECDDD9A4C71" box="[1262,1437,345,370]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="345" phylum="Chordata" rank="species" species="magniscuttis">
<emphasis id="15EDD731FF8CFF99178FFECDDD9A4C71" box="[1262,1437,345,370]" italics="true" pageId="16" pageNumber="345">E. magniscuttis</emphasis>
</taxonomicName>
sequences (
<tableCitation id="6A1B3E98FF8CFF991276FEE8D96C4C95" box="[279,363,380,406]" captionStart="TABLE 1" captionStartId="8.[151,244,152,179]" captionTargetBox="[163,1359,194,1978]" captionText="TABLE 1. Details of all sequences analysed in this study, including Genbank/BOLD accession numbers." httpUri="http://table.plazi.org/id/73E65BABFF94FF8113F6FF0CDD374DB0" pageId="16" pageNumber="345" tableUuid="73E65BABFF94FF8113F6FF0CDD374DB0">Table 1</tableCitation>
, fig. 1). Further sequences from 
<taxonomicName id="E09970A0FF8CFF9911B9FEE9DB834C95" box="[728,900,381,406]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="345" phylum="Chordata" rank="species" species="magniscuttis">
<emphasis id="15EDD731FF8CFF9911B9FEE9DB834C95" box="[728,900,381,406]" italics="true" pageId="16" pageNumber="345">E. magniscuttis</emphasis>
</taxonomicName>
may provide greater genetic resolution between the two species. 
<bibRefCitation id="430876D2FF8CFF99123BFE34DAA54CB9" author="Iswarya Deepti, V. &amp; Sujatha, K. &amp; Kehdkar, G. D." box="[346,674,416,442]" pageId="16" pageNumber="345" pagination="659 - 663" refId="ref12352" refString="Iswarya Deepti, V., Sujatha, K. &amp; Kehdkar, G. D. (2018) DNA barcoding of five species of groupers (Pisces: Serranidae) off Wisakhapatnam, central eastern coast of India. Mitochondrial DNA Part A, 29 (5), 659 - 663. https: // doi. org / 10.1080 / 24701394.2017.1339188" type="journal article" year="2018">
Iswarya Deepti 
<emphasis id="15EDD731FF8CFF991173FE35DA4A4CB9" box="[530,589,416,442]" italics="true" pageId="16" pageNumber="345">et al.</emphasis>
(2018)
</bibRefCitation>
include three specimens identified as 
<taxonomicName id="E09970A0FF8CFF991700FE35DD164CB9" box="[1121,1297,417,442]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="345" phylum="Chordata" rank="species" species="magniscuttis">
<emphasis id="15EDD731FF8CFF991700FE35DD164CB9" box="[1121,1297,417,442]" italics="true" pageId="16" pageNumber="345">E. magniscuttis</emphasis>
</taxonomicName>
in their genetic (COI) identification of Indian grouper species, however, these sequences were not publically available (either through GenBank or BOLD), so could not be incorporated in this study. Other reports of 
<taxonomicName id="E09970A0FF8CFF991712FE7DDD274F01" box="[1139,1312,489,514]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="345" phylum="Chordata" rank="species" species="magniscuttis">
<emphasis id="15EDD731FF8CFF991712FE7DDD274F01" box="[1139,1312,489,514]" italics="true" pageId="16" pageNumber="345">E. magniscuttis</emphasis>
</taxonomicName>
from 
<collectingCountry id="5F8E4BB3FF8CFF991602FE7CDD9A4F01" box="[1379,1437,488,514]" name="India" pageId="16" pageNumber="345">India</collectingCountry>
(e.g. 
<bibRefCitation id="430876D2FF8CFF9913B1FD98D9AA4F25" author="Sujatha, K. &amp; Padmavathi, P. &amp; Iswarya Deepti, V. A. &amp; Shrikanya, K. V. L." box="[208,429,524,550]" pageId="16" pageNumber="345" pagination="341 - 343" refId="ref13303" refString="Sujatha, K., Padmavathi, P., Iswarya Deepti, V. A. &amp; Shrikanya, K. V. L. (2008) Epinephelus magniscuttis Postel, Fourmanoir and Gueze, 1963 - new record from Indian waters. Indian Journal of Fisheries, 55 (4), 341 - 343." type="journal article" year="2008">
Sujatha 
<emphasis id="15EDD731FF8CFF99124AFD99D95A4F25" box="[299,349,524,550]" italics="true" pageId="16" pageNumber="345">et al</emphasis>
., 2008
</bibRefCitation>
; 
<bibRefCitation id="430876D2FF8CFF9912D8FD98DA924F25" author="Sujatha, K. &amp; Shrikanya, K. V. L. &amp; Iswarya Deepti, V. A." box="[441,661,523,550]" pageId="16" pageNumber="345" pagination="18 - 33" refId="ref13228" refString="Sujatha, K., Shrikanya, K. V. L. &amp; Iswarya Deepti, V. A. (2015) Species diversity and some aspects of reproductive biology and life history of groupers (Pisces: Serranidae: Epinephelinae) off the central eastern coast of India. Marine Biology Research, 11 (1), 18 - 33. https: // doi. org / 10.1080 / 17451000.2014.949271" type="journal article" year="2015">
Sujatha 
<emphasis id="15EDD731FF8CFF991175FD99DA414F25" box="[532,582,524,550]" italics="true" pageId="16" pageNumber="345">et al</emphasis>
., 2015
</bibRefCitation>
) illustrate specimens that are doubtfully distinct from 
<emphasis id="15EDD731FF8CFF991797FD99DD004F25" box="[1270,1287,525,550]" italics="true" pageId="16" pageNumber="345">E</emphasis>
. cf 
<taxonomicName id="E09970A0FF8CFF991653FD99DD9B4F25" authorityName="in Heemstra &amp; Randall" authorityYear="1993" box="[1330,1436,525,550]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="345" phylum="Chordata" rank="species" species="epistictus">
<emphasis id="15EDD731FF8CFF991653FD99DD9B4F25" box="[1330,1436,525,550]" italics="true" pageId="16" pageNumber="345">epistictus</emphasis>
</taxonomicName>
and other authors (e.g. 
<bibRefCitation id="430876D2FF8CFF9912F8FDA4DA7D4F49" author="Basheer, V. S. &amp; Vineesh, N. &amp; Bineesh, K. K. &amp; Kumar, R. G. &amp; Mohitha, C. &amp; Venu, S. &amp; Kathirvelpandian, A. &amp; Gopalakrishnan, A. &amp; Jena, J. K." box="[409,634,560,586]" pageId="16" pageNumber="345" pagination="451 - 457" refId="ref11554" refString="Basheer, V. S., Vineesh, N., Bineesh, K. K., Kumar, R. G., Mohitha, C., Venu, S., Kathirvelpandian, A., Gopalakrishnan, A. &amp; Jena, J. K. (2017) Mitochondrial signatures for identification of grouper species from Indian waters. Mitochondrial DNA Part A, 28 (4), 451 - 457. https: // doi. org / 10.3109 / 19401736.2015.1137899" type="journal article" year="2017">
Basheer 
<emphasis id="15EDD731FF8CFF99129BFDA5DA2B4F49" box="[506,556,560,586]" italics="true" pageId="16" pageNumber="345">et al</emphasis>
., 2017
</bibRefCitation>
) have indicated that they are likely misidentifications of 
<taxonomicName id="E09970A0FF8CFF991799FDA5DD864F49" authorityName="in Heemstra &amp; Randall" authorityYear="1993" box="[1272,1409,561,586]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="345" phylum="Chordata" rank="species" species="epistictus">
<emphasis id="15EDD731FF8CFF991799FDA5DD864F49" box="[1272,1409,561,586]" italics="true" pageId="16" pageNumber="345">E. epistictus</emphasis>
</taxonomicName>
.
</paragraph>
<caption id="73E65BABFF8CFF9913F6F950DD254A02" ID-DOI="http://doi.org/10.5281/zenodo.3460190" ID-Zenodo-Dep="3460190" httpUri="https://zenodo.org/record/3460190/files/figure.png" pageId="16" pageNumber="345" startId="16.[151,250,1732,1757]" targetBox="[168,1437,617,1701]" targetPageId="16">
<paragraph id="27260B23FF8CFF9913F6F950DD254A02" blockId="16.[151,1436,1732,1793]" pageId="16" pageNumber="345">
<emphasis id="15EDD731FF8CFF9913F6F950D9154BDE" bold="true" box="[151,274,1732,1757]" pageId="16" pageNumber="345">FIGURE 6.</emphasis>
<taxonomicName id="E09970A0FF8CFF991279F951DA1C4BDE" box="[280,539,1733,1757]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="345" phylum="Chordata" rank="species" species="magniscuttis">
<emphasis id="15EDD731FF8CFF991279F951DA1C4BDE" box="[280,539,1733,1757]" italics="true" pageId="16" pageNumber="345">Epinephelus magniscuttis</emphasis>
</taxonomicName>
A. WAM P.34888–001 (GenBank 
<accessionNumber id="38CA96C0FF8CFF99101BF950DBF44BDE" box="[890,1011,1732,1757]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MK693014" pageId="16" pageNumber="345">MK693014</accessionNumber>
), 577 mm SL, off Augusta, WA, Australia, fresh (Photo: M. Allen); B. ca 60 cm TL, off Exmouth, WA, Australia, fresh, not retained (Photo: C. Buckingham).
</paragraph>
</caption>
<paragraph id="27260B23FF8CFF9813A6F8BADBF04C40" blockId="16.[151,1437,1838,2046]" lastBlockId="17.[151,1437,152,719]" lastPageId="17" lastPageNumber="346" pageId="16" pageNumber="345">
<emphasis id="15EDD731FF8CFF9913A6F8BAD9DB4A4A" bold="true" box="[199,476,1838,1865]" pageId="16" pageNumber="345">Comparative materials</emphasis>
: 
<taxonomicName id="E09970A0FF8CFF99128DF8BBDAEF4A4A" authorityName="in Heemstra &amp; Randall" authorityYear="1993" box="[492,744,1839,1865]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="345" phylum="Chordata" rank="species" species="epistictus">
<emphasis id="15EDD731FF8CFF99128DF8BBDAEF4A4A" box="[492,744,1839,1865]" italics="true" pageId="16" pageNumber="345">Epinephelus epistictus</emphasis>
</taxonomicName>
KAUM I. 
<quantity id="E061A6C6FF8CFF99103BF8BADC154A49" box="[858,1042,1838,1866]" metricMagnitude="4" metricUnit="m" metricValue="3.452019" pageId="16" pageNumber="345" unit="mm" value="3.452019E7">34520, 190 mm</quantity>
, East 
<collectingCountry id="5F8E4BB3FF8CFF991736F8BBDC9D4A4A" box="[1111,1178,1839,1865]" name="China" pageId="16" pageNumber="345">China</collectingCountry>
Sea, trawl, 
<date id="53272DE3FF8CFF991640F8A4DD904A4A" box="[1313,1431,1838,1866]" pageId="16" pageNumber="345" value="1965-05">May 1965</date>
; KAUM I. 
<quantity id="E061A6C6FF8CFF991262F8C7D9B84A6D" box="[259,447,1875,1902]" metricMagnitude="4" metricUnit="m" metricValue="9.7319376" pageId="16" pageNumber="345" unit="mm" value="9.7319376E7">97319, 376 mm</quantity>
, off Mi-shima Island, Hagi, 
<collectingRegion id="E55DC5C1FF8CFF991069F8C7DB8E4A6E" box="[776,905,1875,1901]" country="Japan" name="Yamaguchi" pageId="16" pageNumber="345">Yamaguchi</collectingRegion>
, 
<collectingCountry id="5F8E4BB3FF8CFF9910F6F8C0DBDD4A6D" box="[919,986,1876,1902]" name="Japan" pageId="16" pageNumber="345">Japan</collectingCountry>
, 
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<geoCoordinate id="42AD6DE4FF8CFF991714F8C7DD014A6E" box="[1141,1286,1875,1901]" degrees="131" direction="east" minutes="09.19" orientation="longitude" pageId="16" pageNumber="345" precision="9" value="131.15317">131°09.19’E</geoCoordinate>
, line fishing, 
<date id="53272DE3FF8CFF9913F6F8E3D91D4A92" box="[151,282,1910,1937]" pageId="16" pageNumber="345" value="2015-10-06">6 Oct 2015</date>
; KAUM I. 
<date id="53272DE3FF8CFF9912F0F8E3D9EF4A92" box="[401,488,1911,1937]" pageId="16" pageNumber="345" value="80-11-31">113180</date>
<quantity id="E061A6C6FF8CFF991290F8E3DA564A91" box="[497,593,1911,1938]" metricMagnitude="-1" metricUnit="m" metricValue="2.2399999999999998" pageId="16" pageNumber="345" unit="mm" value="224.0">224 mm</quantity>
, off Ke-Zai-Liao, Ziguan District, 
<collectingRegion id="E55DC5C1FF8CFF9910BFF8E3DC5D4A92" box="[990,1114,1911,1937]" country="Taiwan" name="Kaohsiung" pageId="16" pageNumber="345">Kaohsiung</collectingRegion>
, 
<collectingCountry id="5F8E4BB3FF8CFF991707F8E3DCBC4A92" box="[1126,1211,1911,1937]" name="Taiwan" pageId="16" pageNumber="345">Taiwan</collectingCountry>
, K. Koeda, H. Hata &amp; N. Muto, 
<date id="53272DE3FF8CFF991242F80ED9AB4AB6" box="[291,428,1946,1974]" pageId="16" pageNumber="345" value="2018-03-05">5 Mar 2018</date>
. 
<taxonomicName id="E09970A0FF8CFF9912D6F80FDA464AB6" authorityName="Bloch" authorityYear="1793" box="[439,577,1947,1973]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="345" phylum="Chordata" rank="genus">
<emphasis id="15EDD731FF8CFF9912D6F80FDA464AB6" box="[439,577,1947,1973]" italics="true" pageId="16" pageNumber="345">Epinephelus</emphasis>
</taxonomicName>
cf 
<taxonomicName id="E09970A0FF8CFF991107F808DAD74AB6" authorityName="in Heemstra &amp; Randall" authorityYear="1993" box="[614,720,1948,1973]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="345" phylum="Chordata" rank="species" species="epistictus">
<emphasis id="15EDD731FF8CFF991107F808DAD74AB6" box="[614,720,1948,1973]" italics="true" pageId="16" pageNumber="345">epistictus</emphasis>
</taxonomicName>
CSIRO CA. 
<quantity id="E061A6C6FF8CFF99103EF80FDC004AB5" box="[863,1031,1947,1974]" metricMagnitude="3" metricUnit="m" metricValue="2.266406" pageId="16" pageNumber="345" unit="mm" value="2266406.0">2266, 406 mm</quantity>
, Arafura Sea, NT, 
<collectingCountry id="5F8E4BB3FF8CFF9917B8F80FDD444AB6" box="[1241,1347,1947,1973]" name="Australia" pageId="16" pageNumber="345">Australia</collectingCountry>
, 
<geoCoordinate id="42AD6DE4FF8CFF99162FF80FDD9B4AB6" box="[1358,1436,1947,1973]" degrees="9" direction="south" minutes="37" orientation="latitude" pageId="16" pageNumber="345" precision="925" value="-9.616667">9°37’S</geoCoordinate>
<geoCoordinate id="42AD6DE4FF8CFF9913F6F82BD9014ADA" box="[151,262,1983,2009]" degrees="133" direction="east" minutes="06" orientation="longitude" pageId="16" pageNumber="345" precision="925" value="133.1">133°06’E</geoCoordinate>
, trawl, 
<quantity id="E061A6C6FF8CFF991239F82BD9DD4AD9" box="[344,474,1983,2010]" metricMagnitude="2" metricUnit="m" metricValue="1.28" metricValueMax="1.3" metricValueMin="1.26" pageId="16" pageNumber="345" unit="m" value="128.0" valueMax="130.0" valueMin="126.0">126–130 m</quantity>
, CSIRO, 
<date id="53272DE3FF8CFF991127F82BDAB94ADA" box="[582,702,1983,2009]" pageId="16" pageNumber="345" value="1981-07-02">2 Jul 1981</date>
; CSIRO H.7306-03, 
<quantity id="E061A6C6FF8CFF9910CBF82BDC0E4AD9" box="[938,1033,1983,2010]" metricMagnitude="-1" metricUnit="m" metricValue="1.89" pageId="16" pageNumber="345" unit="mm" value="189.0">189 mm</quantity>
, Pelabuhanratu, south coast of 
<collectingRegion id="E55DC5C1FF8CFF991604F854D8CC4AFD" country="Indonesia" name="Jawa Barat" pageId="16" pageNumber="345">West Java</collectingRegion>
, 
<collectingCountry id="5F8E4BB3FF8CFF9913B8F877D94E4AFE" box="[217,329,2019,2045]" name="Indonesia" pageId="16" pageNumber="345">Indonesia</collectingCountry>
, 
<geoCoordinate id="42AD6DE4FF8CFF991237F877D9A24AFE" box="[342,421,2019,2045]" degrees="7" direction="south" minutes="02" orientation="latitude" pageId="16" pageNumber="345" precision="925" value="-7.0333333">7°02’S</geoCoordinate>
<geoCoordinate id="42AD6DE4FF8CFF9912CFF877DA1C4AFE" box="[430,539,2019,2045]" degrees="106" direction="east" minutes="32" orientation="longitude" pageId="16" pageNumber="345" precision="925" value="106.53333">106°32’E</geoCoordinate>
, W. White, 
<date id="53272DE3FF8CFF9911C4F877DB3F4AFE" box="[677,824,2019,2045]" pageId="16" pageNumber="345" value="2010-10-28">28 Oct 2010</date>
; CSIRO H.7306-04, 
<quantity id="E061A6C6FF8CFF99174BF877DC8C4AFD" box="[1066,1163,2019,2046]" metricMagnitude="-1" metricUnit="m" metricValue="1.91" pageId="16" pageNumber="345" unit="mm" value="191.0">191 mm</quantity>
, same data as previous; CSIRO H.7306-05, 
<quantity id="E061A6C6FF8DFF98121DFF0DD9DA4DB0" box="[380,477,153,179]" metricMagnitude="-1" metricUnit="m" metricValue="2.1" pageId="17" pageNumber="346" unit="mm" value="210.0">210 mm</quantity>
, same data as previous; CSIRO H.7407-02, 
<quantity id="E061A6C6FF8DFF9810BDFF0DDC394DB0" box="[988,1086,153,179]" metricMagnitude="-1" metricUnit="m" metricValue="1.9899999999999998" pageId="17" pageNumber="346" unit="mm" value="199.0">199 mm</quantity>
, Pelabuhanratu, south coast of 
<collectingRegion id="E55DC5C1FF8DFF9813F6FF29D90E4DD4" box="[151,265,189,215]" country="Indonesia" name="Jawa Barat" pageId="17" pageNumber="346">West Java</collectingRegion>
, 
<collectingCountry id="5F8E4BB3FF8DFF981275FF29D9834DD4" box="[276,388,189,215]" name="Indonesia" pageId="17" pageNumber="346">Indonesia</collectingCountry>
, 
<geoCoordinate id="42AD6DE4FF8DFF9812EEFF29D9DA4DD4" box="[399,477,189,215]" degrees="7" direction="south" minutes="02" orientation="latitude" pageId="17" pageNumber="346" precision="925" value="-7.0333333">7°02’S</geoCoordinate>
<geoCoordinate id="42AD6DE4FF8DFF981282FF29DA564DD4" box="[483,593,189,215]" degrees="106" direction="east" minutes="32" orientation="longitude" pageId="17" pageNumber="346" precision="925" value="106.53333">106°32’E</geoCoordinate>
, W. White, 
<date id="53272DE3FF8DFF9811B2FF29DB664DD4" box="[723,865,189,215]" pageId="17" pageNumber="346" value="2010-10-10">10 Oct 2010</date>
; MZB PB 
<quantity id="E061A6C6FF8DFF9810B0FF29DC604DD4" box="[977,1127,189,215]" metricMagnitude="2" metricUnit="m" metricValue="3.36177" pageId="17" pageNumber="346" unit="mm" value="336177.0">336, 177 mm</quantity>
, Pelabuhanratu, south coast of 
<collectingRegion id="E55DC5C1FF8DFF9813D7FF75D92E4DF8" box="[182,297,225,251]" country="Indonesia" name="Jawa Barat" pageId="17" pageNumber="346">West Java</collectingRegion>
, 
<collectingCountry id="5F8E4BB3FF8DFF981254FF75D9A24DF8" box="[309,421,225,251]" name="Indonesia" pageId="17" pageNumber="346">Indonesia</collectingCountry>
, 
<geoCoordinate id="42AD6DE4FF8DFF9812D0FF75D9F84DF8" box="[433,511,225,251]" degrees="7" direction="south" minutes="02" orientation="latitude" pageId="17" pageNumber="346" precision="925" value="-7.0333333">7°02’S</geoCoordinate>
<geoCoordinate id="42AD6DE4FF8DFF981167FF75DA734DF8" box="[518,628,225,251]" degrees="106" direction="east" minutes="32" orientation="longitude" pageId="17" pageNumber="346" precision="925" value="106.53333">106°32’E</geoCoordinate>
, W. White, 
<date id="53272DE3FF8DFF981198FF75DB8E4DF8" box="[761,905,225,251]" pageId="17" pageNumber="346" value="2010-10-28">28 Oct 2010</date>
; KAUM I. 
<quantity id="E061A6C6FF8DFF98109EFF74DCB14DF8" box="[1023,1206,224,251]" metricMagnitude="4" metricUnit="m" metricValue="1.2451146" pageId="17" pageNumber="346" unit="mm" value="1.2451146E7">12451, 146 mm</quantity>
, Off Kota Kinabalu, 
<collectingRegion id="E55DC5C1FF8DFF9813F6FE91D8D94C1C" box="[151,222,261,287]" country="Malaysia" name="Sabah" pageId="17" pageNumber="346">Sabah</collectingRegion>
, 
<collectingCountry id="5F8E4BB3FF8DFF98138BFE91D9544C1C" box="[234,339,261,287]" name="Malaysia" pageId="17" pageNumber="346">Malaysia</collectingCountry>
, 
<geoCoordinate id="42AD6DE4FF8DFF981201FE91D9B44C1C" box="[352,435,261,287]" degrees="6" direction="north" minutes="00" orientation="latitude" pageId="17" pageNumber="346" precision="925" value="6.0">6°00’N</geoCoordinate>
<geoCoordinate id="42AD6DE4FF8DFF9812DBFE91DA204C1C" box="[442,551,261,287]" degrees="116" direction="east" minutes="07" orientation="longitude" pageId="17" pageNumber="346" precision="925" value="116.11667">116°07’E</geoCoordinate>
, KAUM, 
<date id="53272DE3FF8DFF9811F9FE90DB204C1C" box="[664,807,260,287]" pageId="17" pageNumber="346" value="2008-10-15">15 Oct 2008</date>
; NTM S.12902–028, 
<quantity id="E061A6C6FF8DFF98177CFE91DC7A4C1C" box="[1053,1149,261,287]" metricMagnitude="-1" metricUnit="m" metricValue="1.34" pageId="17" pageNumber="346" unit="mm" value="134.0">134 mm</quantity>
, NNE of Cape Don, Arafura Sea, NT, 
<collectingCountry id="5F8E4BB3FF8DFF981250FEBDD99C4C40" box="[305,411,297,323]" name="Australia" pageId="17" pageNumber="346">Australia</collectingCountry>
, 
<geoCoordinate id="42AD6DE4FF8DFF9812C7FEBDDA054C40" box="[422,514,297,323]" degrees="19" direction="south" minutes="19" orientation="latitude" pageId="17" pageNumber="346" precision="925" value="-19.316668">19°19’S</geoCoordinate>
<geoCoordinate id="42AD6DE4FF8DFF981168FEBDDA704C40" box="[521,631,297,323]" degrees="132" direction="east" minutes="49" orientation="longitude" pageId="17" pageNumber="346" precision="925" value="132.81667">132°49’E</geoCoordinate>
, 
<quantity id="E061A6C6FF8DFF9811E2FEBDDACB4C40" box="[643,716,297,323]" metricMagnitude="2" metricUnit="m" metricValue="1.43" pageId="17" pageNumber="346" unit="m" value="143.0">143 m</quantity>
, R. Williams, 
<date id="53272DE3FF8DFF98100AFEBDDBF54C40" box="[875,1010,297,323]" pageId="17" pageNumber="346" value="1990-11-07">7 Nov 1990</date>
.
</paragraph>
<paragraph id="27260B23FF8DFF9813A6FED9D8D04CF4" blockId="17.[151,1437,152,719]" pageId="17" pageNumber="346">
<taxonomicName id="E09970A0FF8DFF9813A6FED9D9C94C64" baseAuthorityName="non Fowler" baseAuthorityYear="1904" box="[199,462,333,359]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="346" phylum="Chordata" rank="species" species="heniochus">
<emphasis id="15EDD731FF8DFF9813A6FED9D9C94C64" box="[199,462,333,359]" italics="true" pageId="17" pageNumber="346">Epinephelus heniochus</emphasis>
</taxonomicName>
<materialsCitation id="97F1017EFF8DFF9812B8FED9DD904C64" ID-GBIF-Occurrence="2423596353" box="[473,1431,332,359]" collectionCode="KAUM" country="Malaysia" latitude="6.0" location="Kota Kinabalu" longLatPrecision="1305" longitude="116.11667" pageId="17" pageNumber="346" specimenCount="1" stateProvince="Sabah">
<collectionCode id="418893E6FF8DFF9812B8FED9DA284C64" box="[473,559,333,359]" country="Japan" httpUri="http://grbio.org/cool/r700-376n" name="Kagoshima University Museum" pageId="17" pageNumber="346">KAUM</collectionCode>
I.22013, 
<quantity id="E061A6C6FF8DFF9811C0FED8DB0C4C64" box="[673,779,332,359]" metricMagnitude="-2" metricUnit="m" metricValue="9.55" pageId="17" pageNumber="346" unit="mm" value="95.5">95.5 mm</quantity>
, off 
<location id="22465DF8FF8DFF981024FED9DBF74C64" LSID="urn:lsid:plazi:treatment:AF30BA35FF9FFF9813F6FA65DA124FCC:22465DF8FF8DFF981024FED9DBF74C64" box="[837,1008,333,359]" country="Malaysia" latitude="6.0" longLatPrecision="1305" longitude="116.11667" name="Kota Kinabalu" pageId="17" pageNumber="346" stateProvince="Sabah">Kota Kinabalu</location>
, 
<collectingRegion id="E55DC5C1FF8DFF98109EFED9DC414C64" box="[1023,1094,333,359]" country="Malaysia" name="Sabah" pageId="17" pageNumber="346">Sabah</collectingRegion>
, 
<collectingCountry id="5F8E4BB3FF8DFF981734FED9DCB94C64" box="[1109,1214,333,359]" name="Malaysia" pageId="17" pageNumber="346">Malaysia</collectingCountry>
, 
<geoCoordinate id="42AD6DE4FF8DFF9817ACFED9DD274C64" box="[1229,1312,333,359]" degrees="6" direction="north" minutes="00" orientation="latitude" pageId="17" pageNumber="346" precision="925" value="6.0">6°00’N</geoCoordinate>
<geoCoordinate id="42AD6DE4FF8DFF98164AFED9DD904C64" box="[1323,1431,333,359]" degrees="116" direction="east" minutes="07" orientation="longitude" pageId="17" pageNumber="346" precision="925" value="116.11667">116°07’E</geoCoordinate>
</materialsCitation>
, 
<materialsCitation id="97F1017EFF8DFF9813F6FEE5D9934C88" ID-GBIF-Occurrence="2423596360" box="[151,404,369,395]" collectingDate="2009-08-11" collectionCode="KAUM" country="Malaysia" pageId="17" pageNumber="346" specimenCount="1" stateProvince="Sabah">
<collectionCode id="418893E6FF8DFF9813F6FEE5D8E84C88" box="[151,239,369,395]" country="Japan" httpUri="http://grbio.org/cool/r700-376n" name="Kagoshima University Museum" pageId="17" pageNumber="346">KAUM</collectionCode>
, 
<date id="53272DE3FF8DFF98139CFEE5D9934C88" box="[253,404,369,395]" pageId="17" pageNumber="346" value="2009-08-11">
<collectingDate id="4363D40BFF8DFF98139CFEE5D9934C88" box="[253,404,369,395]" pageId="17" pageNumber="346" value="2009-08-11">11 Aug 2009</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation id="97F1017EFF8DFF9812C3FEE5DB904C88" ID-GBIF-Occurrence="2423596358" box="[418,919,369,395]" collectionCode="KAUM" collectorName="Gulf" country="Thailand" location="Thailand" pageId="17" pageNumber="346" specimenCount="1">
<collectionCode id="418893E6FF8DFF9812C3FEE5D9FF4C88" box="[418,504,369,395]" country="Japan" httpUri="http://grbio.org/cool/r700-376n" name="Kagoshima University Museum" pageId="17" pageNumber="346">KAUM</collectionCode>
I. 
<quantity id="E061A6C6FF8DFF98116FFEE5DAC04C88" box="[526,711,369,395]" metricMagnitude="4" metricUnit="m" metricValue="2.3091114" pageId="17" pageNumber="346" unit="mm" value="2.3091114E7">23091, 114 mm</quantity>
, 
<collectorName id="8A6C6EF5FF8DFF9811B4FEE5DB0F4C88" box="[725,776,369,395]" pageId="17" pageNumber="346">Gulf</collectorName>
of 
<collectingCountry id="5F8E4BB3FF8DFF981050FEE5DB904C88" box="[817,919,369,395]" name="Thailand" pageId="17" pageNumber="346">Thailand</collectingCountry>
</materialsCitation>
, 
<materialsCitation id="97F1017EFF8DFF9810C4FEE5DC9F4C88" ID-GBIF-Occurrence="2423596354" box="[933,1176,369,395]" collectingDate="2009-09-06" collectionCode="KAUM" country="Thailand" pageId="17" pageNumber="346" specimenCount="1">
<collectionCode id="418893E6FF8DFF9810C4FEE5DBFA4C88" box="[933,1021,369,395]" country="Japan" httpUri="http://grbio.org/cool/r700-376n" name="Kagoshima University Museum" pageId="17" pageNumber="346">KAUM</collectionCode>
, 
<date id="53272DE3FF8DFF98176AFEE5DC9F4C88" box="[1035,1176,369,395]" pageId="17" pageNumber="346" value="2009-09-06">
<collectingDate id="4363D40BFF8DFF98176AFEE5DC9F4C88" box="[1035,1176,369,395]" pageId="17" pageNumber="346" value="2009-09-06">6 Sept 2009</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation id="97F1017EFF8DFF9817C7FEE5D98A4CAC" ID-GBIF-Occurrence="2423596367" collectionCode="KAUM" collectorName="Gulf" country="Thailand" location="Thailand" pageId="17" pageNumber="346" specimenCount="1">
<collectionCode id="418893E6FF8DFF9817C7FEE5DCFB4C88" box="[1190,1276,369,395]" country="Japan" httpUri="http://grbio.org/cool/r700-376n" name="Kagoshima University Museum" pageId="17" pageNumber="346">KAUM</collectionCode>
I.23282, 
<quantity id="E061A6C6FF8DFF98160AFEE5D8C24CAC" metricMagnitude="-2" metricUnit="m" metricValue="8.69" pageId="17" pageNumber="346" unit="mm" value="86.9">86.9 mm</quantity>
, 
<collectorName id="8A6C6EF5FF8DFF9813B1FE01D9044CAC" box="[208,259,405,431]" pageId="17" pageNumber="346">Gulf</collectorName>
of 
<collectingCountry id="5F8E4BB3FF8DFF981246FE01D98A4CAC" box="[295,397,405,431]" name="Thailand" pageId="17" pageNumber="346">Thailand</collectingCountry>
</materialsCitation>
, 
<materialsCitation id="97F1017EFF8DFF9812F6FE01DA884CAC" ID-GBIF-Occurrence="2423596365" box="[407,655,405,431]" collectingDate="2009-09-10" collectionCode="KAUM" country="Thailand" pageId="17" pageNumber="346" specimenCount="1">
<collectionCode id="418893E6FF8DFF9812F6FE01D9E84CAC" box="[407,495,405,431]" country="Japan" httpUri="http://grbio.org/cool/r700-376n" name="Kagoshima University Museum" pageId="17" pageNumber="346">KAUM</collectionCode>
, 
<date id="53272DE3FF8DFF98129BFE01DA884CAC" box="[506,655,405,431]" pageId="17" pageNumber="346" value="2009-09-10">
<collectingDate id="4363D40BFF8DFF98129BFE01DA884CAC" box="[506,655,405,431]" pageId="17" pageNumber="346" value="2009-09-10">10 Sept 2009</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation id="97F1017EFF8DFF9811FBFE01D90D4CD0" ID-GBIF-Occurrence="2423596362" collectingDate="1990-11-16" collectionCode="QM" collectorName="D. Tuma" country="Australia" location="North West Shelf" pageId="17" pageNumber="346" specimenCount="1">
<collectionCode id="418893E6FF8DFF9811FBFE01DAC04CAC" box="[666,711,405,431]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="17" pageNumber="346" type="Museum">QM</collectionCode>
I. 
<quantity id="E061A6C6FF8DFF9811BAFE01DB884CAC" box="[731,911,404,431]" metricMagnitude="4" metricUnit="m" metricValue="2.7488305" pageId="17" pageNumber="346" unit="mm" value="2.7488305E7">27488, 305 mm</quantity>
, 
<location id="22465DF8FF8DFF9810FBFE01DC5A4CAC" LSID="urn:lsid:plazi:treatment:AF30BA35FF9FFF9813F6FA65DA124FCC:22465DF8FF8DFF9810FBFE01DC5A4CAC" box="[922,1117,405,431]" country="Australia" name="North West Shelf" pageId="17" pageNumber="346">North West Shelf</location>
, WA, 
<collectingCountry id="5F8E4BB3FF8DFF9817FFFE01DD0F4CAC" box="[1182,1288,405,431]" name="Australia" pageId="17" pageNumber="346">Australia</collectingCountry>
, 
<collectorName id="8A6C6EF5FF8DFF981673FE01DD714CAC" box="[1298,1398,405,431]" pageId="17" pageNumber="346">D. Tuma</collectorName>
, 
<date id="53272DE3FF8DFF9816E1FE01D90D4CD0" pageId="17" pageNumber="346" value="1990-11-16">
<collectingDate id="4363D40BFF8DFF9816E1FE01D90D4CD0" pageId="17" pageNumber="346" value="1990-11-16">16 Nov 1990</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation id="97F1017EFF8DFF981277FE2DDCF74CD0" ID-GBIF-Occurrence="2423596370" box="[278,1264,441,467]" collectionCode="QM" country="Australia" location="Groote Eylandt" municipality="Gulf of Carpentaria" pageId="17" pageNumber="346" specimenCount="1">
<collectionCode id="418893E6FF8DFF981277FE2DD9444CD0" box="[278,323,441,467]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="17" pageNumber="346" type="Museum">QM</collectionCode>
I. 
<quantity id="E061A6C6FF8DFF981236FE2DDA094CD0" box="[343,526,441,467]" metricMagnitude="4" metricUnit="m" metricValue="3.4961127999999997" pageId="17" pageNumber="346" unit="mm" value="3.4961128E7">34961, 128 mm</quantity>
, east of 
<location id="22465DF8FF8DFF98110AFE2DDB1D4CD0" LSID="urn:lsid:plazi:treatment:AF30BA35FF9FFF9813F6FA65DA124FCC:22465DF8FF8DFF98110AFE2DDB1D4CD0" box="[619,794,441,467]" country="Australia" municipality="Gulf of Carpentaria" name="Groote Eylandt" pageId="17" pageNumber="346">Groote Eylandt</location>
, 
<collectingMunicipality id="C7429159FF8DFF981044FE2DDC024CD0" box="[805,1029,441,467]" pageId="17" pageNumber="346">Gulf of Carpentaria</collectingMunicipality>
, NT, 
<collectingCountry id="5F8E4BB3FF8DFF98175EFE2DDCAE4CD0" box="[1087,1193,441,467]" name="Australia" pageId="17" pageNumber="346">Australia</collectingCountry>
, trawl
</materialsCitation>
, 
<materialsCitation id="97F1017EFF8DFF98179DFE2DD8D44CF4" ID-GBIF-Occurrence="2423596369" collectingDate="2003-02-09" collectionCode="CSIRO" country="Australia" pageId="17" pageNumber="346" specimenCount="1">
<collectionCode id="418893E6FF8DFF98179DFE2DDD564CD0" box="[1276,1361,441,467]" country="Australia" httpUri="http://grbio.org/cool/0k2v-y6wz" name="Australian National Fish Collection" pageId="17" pageNumber="346" type="Museum">CSIRO</collectionCode>
, 
<date id="53272DE3FF8DFF98163CFE2DD8D44CF4" pageId="17" pageNumber="346" value="2003-02-09">
<collectingDate id="4363D40BFF8DFF98163CFE2DD8D44CF4" pageId="17" pageNumber="346" value="2003-02-09">9 Feb 2003</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
<paragraph id="27260B23FF8DFF9813A6FD95DB094F84" blockId="17.[151,1437,152,719]" pageId="17" pageNumber="346">
<taxonomicName id="E09970A0FF8DFF9813A6FD95D9E04F18" box="[199,487,513,539]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="346" phylum="Chordata" rank="species" species="magniscuttis">
<emphasis id="15EDD731FF8DFF9813A6FD95D9E04F18" box="[199,487,513,539]" italics="true" pageId="17" pageNumber="346">Epinephelus magniscuttis</emphasis>
</taxonomicName>
<materialsCitation id="97F1017EFF8DFF98128FFD95D9A44F3C" ID-GBIF-Occurrence="2423596359" collectingDate="1979-02-04" collectionCode="WAM" collectorName="Timor Sea, NT &amp; Cliff" country="Australia" elevation="200" latitude="-13.883333" location="Australia" longLatPrecision="1289" longitude="129.41667" pageId="17" pageNumber="346" specimenCount="1">
<collectionCode id="418893E6FF8DFF98128FFD95DA354F18" box="[494,562,513,539]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="17" pageNumber="346" type="Museum">WAM</collectionCode>
P.26610–001, 
<quantity id="E061A6C6FF8DFF9811BBFD94DB3D4F18" box="[730,826,512,539]" metricMagnitude="-1" metricUnit="m" metricValue="5.58" pageId="17" pageNumber="346" unit="mm" value="558.0">558 mm</quantity>
, 
<collectorName id="8A6C6EF5FF8DFF981027FD95DBF74F18" box="[838,1008,513,539]" pageId="17" pageNumber="346">Timor Sea, NT</collectorName>
, 
<collectingCountry id="5F8E4BB3FF8DFF98109BFD95DC634F18" box="[1018,1124,513,539]" name="Australia" pageId="17" pageNumber="346">Australia</collectingCountry>
, 
<geoCoordinate id="42AD6DE4FF8DFF981711FD94DCCB4F18" box="[1136,1228,512,539]" degrees="13" direction="south" minutes="53" orientation="latitude" pageId="17" pageNumber="346" precision="925" value="-13.883333">13°53’S</geoCoordinate>
<geoCoordinate id="42AD6DE4FF8DFF9817B2FD94DD454F18" box="[1235,1346,512,539]" degrees="129" direction="east" minutes="25" orientation="longitude" pageId="17" pageNumber="346" precision="925" value="129.41667">129°25’E</geoCoordinate>
, 
<quantity id="E061A6C6FF8DFF98162FFD95DD904F18" box="[1358,1431,513,539]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="17" pageNumber="346" unit="m" value="200.0">
<elevation id="ACB4EC10FF8DFF98162FFD95DD904F18" box="[1358,1431,513,539]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="17" pageNumber="346" unit="m" value="200.0">200 m</elevation>
</quantity>
, trawl, 
<collectorName id="8A6C6EF5FF8DFF9813BEFDB1D9114F3C" box="[223,278,549,575]" pageId="17" pageNumber="346">Cliff</collectorName>
, 
<date id="53272DE3FF8DFF981243FDB1D9A44F3C" box="[290,419,549,575]" pageId="17" pageNumber="346" value="1979-02-04">
<collectingDate id="4363D40BFF8DFF981243FDB1D9A44F3C" box="[290,419,549,575]" pageId="17" pageNumber="346" value="1979-02-04">4 Feb 1979</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation id="97F1017EFF8DFF9812CFFDB1DA764F60" ID-GBIF-Occurrence="2423596356" collectingDate="1992-06" collectionCode="WAM" collectorName="S. Kramer" country="Australia" elevation="290" latitude="-14.066667" location="Ribbon Reefs" longLatPrecision="1289" longitude="145.68333" municipality="Qld" pageId="17" pageNumber="346" specimenCount="1">
<collectionCode id="418893E6FF8DFF9812CFFDB1D9F54F3C" box="[430,498,549,575]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="17" pageNumber="346" type="Museum">WAM</collectionCode>
P.30510–007, 
<quantity id="E061A6C6FF8DFF9811FBFDB1DAFE4F3C" box="[666,761,549,575]" metricMagnitude="-1" metricUnit="m" metricValue="4.1899999999999995" pageId="17" pageNumber="346" unit="mm" value="419.0">419 mm</quantity>
, off 
<location id="22465DF8FF8DFF98104DFDB1DBCE4F3C" LSID="urn:lsid:plazi:treatment:AF30BA35FF9FFF9813F6FA65DA124FCC:22465DF8FF8DFF98104DFDB1DBCE4F3C" box="[812,969,549,575]" country="Australia" latitude="-14.066667" longLatPrecision="1289" longitude="145.68333" municipality="Qld" name="Ribbon Reefs" pageId="17" pageNumber="346">Ribbon Reefs</location>
, 
<collectingMunicipality id="C7429159FF8DFF9810B5FDB1DC064F3C" box="[980,1025,549,575]" pageId="17" pageNumber="346">Qld</collectingMunicipality>
, 
<collectingCountry id="5F8E4BB3FF8DFF98176AFDB1DC724F3C" box="[1035,1141,549,575]" name="Australia" pageId="17" pageNumber="346">Australia</collectingCountry>
, 
<geoCoordinate id="42AD6DE4FF8DFF98171EFDB1DCDC4F3C" box="[1151,1243,549,575]" degrees="14" direction="south" minutes="04" orientation="latitude" pageId="17" pageNumber="346" precision="925" value="-14.066667">14°04’S</geoCoordinate>
<geoCoordinate id="42AD6DE4FF8DFF981782FDB0DD564F3C" box="[1251,1361,548,575]" degrees="145" direction="east" minutes="41" orientation="longitude" pageId="17" pageNumber="346" precision="925" value="145.68333">145°41’E</geoCoordinate>
, handline, 
<quantity id="E061A6C6FF8DFF9813B1FDDDD91D4F60" box="[208,282,585,611]" metricMagnitude="2" metricUnit="m" metricValue="2.9" pageId="17" pageNumber="346" unit="m" value="290.0">
<elevation id="ACB4EC10FF8DFF9813B1FDDDD91D4F60" box="[208,282,585,611]" metricMagnitude="2" metricUnit="m" metricValue="2.9" pageId="17" pageNumber="346" unit="m" value="290.0">290 m</elevation>
</quantity>
, 
<collectorName id="8A6C6EF5FF8DFF981246FDDDD99C4F60" box="[295,411,585,611]" pageId="17" pageNumber="346">S. Kramer</collectorName>
&amp; party, 
<date id="53272DE3FF8DFF981166FDDDDA764F60" box="[519,625,585,611]" pageId="17" pageNumber="346" value="1992-06">
<collectingDate id="4363D40BFF8DFF981166FDDDDA764F60" box="[519,625,585,611]" pageId="17" pageNumber="346" value="1992-06">Jun 1992</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation id="97F1017EFF8DFF98111CFDDDDB0D4F84" ID-GBIF-Occurrence="2423596357" collectingDate="2018-03-20" collectionCode="WAM" collectorName="A. Heslewood" country="Australia" elevation="300" latitude="-34.316666" location="Augusta" longLatPrecision="1200" longitude="115.166664" pageId="17" pageNumber="346" specimenCount="1">
<collectionCode id="418893E6FF8DFF98111CFDDDDAC64F60" box="[637,705,585,611]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="17" pageNumber="346" type="Museum">WAM</collectionCode>
P.34888–001, 
<quantity id="E061A6C6FF8DFF98100BFDDCDBCD4F60" box="[874,970,584,611]" metricMagnitude="-1" metricUnit="m" metricValue="5.77" pageId="17" pageNumber="346" unit="mm" value="577.0">577 mm</quantity>
, SW of 
<location id="22465DF8FF8DFF981747FDDDDC814F60" LSID="urn:lsid:plazi:treatment:AF30BA35FF9FFF9813F6FA65DA124FCC:22465DF8FF8DFF981747FDDDDC814F60" box="[1062,1158,585,611]" country="Australia" latitude="-34.316666" longLatPrecision="1200" longitude="115.166664" name="Augusta" pageId="17" pageNumber="346">Augusta</location>
, WA, 
<collectingCountry id="5F8E4BB3FF8DFF9817ABFDDDDD334F60" box="[1226,1332,585,611]" name="Australia" pageId="17" pageNumber="346">Australia</collectingCountry>
, 
<geoCoordinate id="42AD6DE4FF8DFF981621FDDDDD9A4F60" box="[1344,1437,585,611]" degrees="34" direction="south" minutes="19" orientation="latitude" pageId="17" pageNumber="346" precision="925" value="-34.316666">34°19’S</geoCoordinate>
<geoCoordinate id="42AD6DE4FF8DFF9813F6FDF8D9034F84" box="[151,260,620,647]" degrees="115" direction="east" minutes="10" orientation="longitude" pageId="17" pageNumber="346" precision="925" value="115.166664">115°10’E</geoCoordinate>
, dropline 
<quantity id="E061A6C6FF8DFF981215FDF9D9BA4F84" box="[372,445,621,647]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="17" pageNumber="346" unit="m" value="300.0">
<elevation id="ACB4EC10FF8DFF981215FDF9D9BA4F84" box="[372,445,621,647]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="17" pageNumber="346" unit="m" value="300.0">300 m</elevation>
</quantity>
, 
<collectorName id="8A6C6EF5FF8DFF9812A6FDF9DA6C4F84" box="[455,619,621,647]" pageId="17" pageNumber="346">A. Heslewood</collectorName>
, 
<date id="53272DE3FF8DFF981117FDF9DB0D4F84" box="[630,778,621,647]" pageId="17" pageNumber="346" value="2018-03-20">
<collectingDate id="4363D40BFF8DFF981117FDF9DB0D4F84" box="[630,778,621,647]" pageId="17" pageNumber="346" value="2018-03-20">20 Mar 2018</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
<paragraph id="27260B23FF8DFF9813A6FD05DA124FCC" blockId="17.[151,1437,152,719]" pageId="17" pageNumber="346">
<taxonomicName id="E09970A0FF8DFF9813A6FD05D9E04FA8" baseAuthorityName="Temminck &amp; Schlegel" baseAuthorityYear="1842" box="[199,487,657,683]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="346" phylum="Chordata" rank="species" species="poecilonotus">
<emphasis id="15EDD731FF8DFF9813A6FD05D9E04FA8" box="[199,487,657,683]" italics="true" pageId="17" pageNumber="346">Epinephelus poecilonotus</emphasis>
</taxonomicName>
, 
<materialsCitation id="97F1017EFF8DFF981292FD05DA164FCC" ID-GBIF-Occurrence="2423596355" collectingDate="1997-06-13" collectionCode="WAM" collectorName="Scott Reef, WA &amp; S. Newman" country="Australia" elevation="162" latitude="-14.033334" location="Australia" longLatPrecision="1289" longitude="121.76667" pageId="17" pageNumber="346" specimenCount="1">
<collectionCode id="418893E6FF8DFF981292FD05DA304FA8" box="[499,567,657,683]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="17" pageNumber="346" type="Museum">WAM</collectionCode>
P.31329–001, 
<quantity id="E061A6C6FF8DFF9811BDFD04DB3D4FA8" box="[732,826,656,683]" metricMagnitude="-1" metricUnit="m" metricValue="4.85" pageId="17" pageNumber="346" unit="mm" value="485.0">485 mm</quantity>
, off 
<collectorName id="8A6C6EF5FF8DFF98100AFD05DC1C4FA8" box="[875,1051,657,683]" pageId="17" pageNumber="346">Scott Reef, WA</collectorName>
, 
<collectingCountry id="5F8E4BB3FF8DFF981745FD05DC894FA8" box="[1060,1166,657,683]" name="Australia" pageId="17" pageNumber="346">Australia</collectingCountry>
, 
<geoCoordinate id="42AD6DE4FF8DFF9817F6FD05DCF44FA8" box="[1175,1267,657,683]" degrees="14" direction="south" minutes="02" orientation="latitude" pageId="17" pageNumber="346" precision="925" value="-14.033334">14°02’S</geoCoordinate>
<geoCoordinate id="42AD6DE4FF8DFF981798FD05DD6F4FA8" box="[1273,1384,657,683]" degrees="121" direction="east" minutes="46" orientation="longitude" pageId="17" pageNumber="346" precision="925" value="121.76667">121°46’E</geoCoordinate>
, 
<quantity id="E061A6C6FF8DFF981613FD05D8A84FCC" metricMagnitude="2" metricUnit="m" metricValue="1.6199999999999999" pageId="17" pageNumber="346" unit="m" value="162.0">
<elevation id="ACB4EC10FF8DFF981613FD05D8A84FCC" metricMagnitude="2" metricUnit="m" metricValue="1.6199999999999999" pageId="17" pageNumber="346" unit="m" value="162.0">162 m</elevation>
</quantity>
, trap, 
<collectorName id="8A6C6EF5FF8DFF981394FD21D97D4FCC" box="[245,378,693,719]" pageId="17" pageNumber="346">S. Newman</collectorName>
, 
<date id="53272DE3FF8DFF9812E4FD21DA164FCC" box="[389,529,693,719]" pageId="17" pageNumber="346" value="1997-06-13">
<collectingDate id="4363D40BFF8DFF9812E4FD21DA164FCC" box="[389,529,693,719]" pageId="17" pageNumber="346" value="1997-06-13">13 Jun 1997</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>