640 lines
54 KiB
XML
640 lines
54 KiB
XML
<document id="CA8C28F517A59A5975AE9AC78D416BAE" ID-DOI="10.3897/zookeys.1223.138220" ID-ZooBank="3557A991-63DF-42D8-B8B1-F310CD52FD00" ID-publisher-id="138220" URI-arpha="7352E8C9-65CA-546F-BE46-EF55E52C51A6" URI-zoobank="http://zoobank.org/3557A991-63DF-42D8-B8B1-F310CD52FD00" article-type="research-article" checkinTime="1736187739607" checkinUser="pensoft" docAuthor="Hendrich, Lars & Brancucci, Michel" docDate="2025" docId="A88F9289A45B52BEB5326BFDB80550D2" docLanguage="en" docName="ZooKeys 1223: 87-147" docOrigin="ZooKeys 1223" docSource="https://zookeys.pensoft.net/article/138220/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Sandracottus bizonatus Regimbart Hendrich & Brancucci 1899" docType="treatment" docVersion="2" dtd-version="3.0" lastPageNumber="147" masterDocId="7352E8C965CA546FBE46EF55E52C51A6" masterDocTitle="Revision of the Oriental and Australasian diving beetle genus Sandracottus Sharp, 1882 (Coleoptera, Dytiscidae, Dytiscinae)" masterLastPageNumber="147" masterPageNumber="87" pageNumber="87" updateTime="1736188416300" updateUser="ExternalLinkService">
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<mods:mods id="49D3D28B971152D5CBAFEC9EB4976E8A" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="24085383FD3664C6C1AEEFAAB5E76901">
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<mods:title id="65CB67D65828666C6055B2AF39FBA6CA">Revision of the Oriental and Australasian diving beetle genus Sandracottus Sharp, 1882 (Coleoptera, Dytiscidae, Dytiscinae)</mods:title>
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<mods:name id="E739658F7885777FD55057DAD08A608F" type="personal">
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<mods:namePart id="48CF3D8CDB451B1EA774ADEDC7E0F175">Hendrich, Lars</mods:namePart>
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<mods:affiliation id="0F24A5721C8F8EDA4C42C2F7102D6ED1">SNSB - Zoologische Staatssammlung München, Münchhausenstraße 21, D – 81247 München, Germany</mods:affiliation>
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<mods:namePart id="ABE0D815ECBFD66BAEC7CDC5CDAD7572">Brancucci, Michel</mods:namePart>
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<mods:affiliation id="2A74FFD8487956AAFB13426CF53E8630">Naturhistorisches Museum Basel, Basel, Switzerland</mods:affiliation>
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<mods:typeOfResource id="0BF352FB1EE4914CFB18F151DBBCF202">text</mods:typeOfResource>
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<mods:title id="F85EA71DAA640B5761219799FCD108C8">ZooKeys</mods:title>
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<mods:part id="93AB477E46E8D1B0478CF7C4FEE50FA1">
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<mods:date id="D5FBF50E2E408AFE3AAF0E52B44A9246">2025</mods:date>
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<mods:number id="60C604497AEA11AD2C984FE2A99FC739">2025-01-06</mods:number>
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<mods:detail id="3691C189E51CCA8429F7C6A7A3F7BA28" type="volume">
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<mods:number id="01FAF30B06E8515C8DE2893E8112A51F">1223</mods:number>
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<mods:start id="5ABCD1CBE68D4E2DE4E1E13E8153D0E4">87</mods:start>
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<mods:end id="CE2112099EF2E2958B13317EB350B212">147</mods:end>
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<mods:classification id="5CC16DF47456B72D78C9148FED69D056">journal article</mods:classification>
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<mods:identifier id="35875E102F07E7600338B949DFF0B7CE" type="DOI">10.3897/zookeys.1223.138220</mods:identifier>
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<mods:identifier id="1BFB7B4C03E91ED9A8862790EF83CCB3" type="ZooBank">3557A991-63DF-42D8-B8B1-F310CD52FD00</mods:identifier>
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<treatment id="A88F9289A45B52BEB5326BFDB80550D2" ID-DOI="http://doi.org/10.5281/zenodo.14606319" ID-Zenodo-Dep="14606319" ID-arpha="A88F9289-A45B-52BE-B532-6BFDB80550D2" LSID="urn:lsid:plazi:treatment:A88F9289A45B52BEB5326BFDB80550D2" httpUri="http://treatment.plazi.org/id/A88F9289A45B52BEB5326BFDB80550D2">
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<subSubSection id="6A66CE9E0394809B38C49E095E168EF0" type="nomenclature">
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<paragraph id="32A443BAAA064EF6B3A9F07CDC8E0E68">
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<taxonomicName id="FA080BF9442818D673A72125F3433DEE" ID-CoL="6XHLD" authority="Regimbart, 1899" authorityName="Regimbart Hendrich & Brancucci" authorityYear="1899" class="Insecta" family="Dytiscidae" genus="Sandracottus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="bizonatus">
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<emphasis id="3498E371DA55D30B1F0F247092FC0536" italics="true">Sandracottus bizonatus</emphasis>
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<bibRefCitation id="F10D53A8DC6D891CC9EBEE38D84B80E0" DOI="10.1080/21686351.1899.12279737" author="Régimbart M" journalOrPublisher="Annales de la Société Entomologique de France" pagination="186-367" refId="B42" refString="Regimbart M (1899) Revision des Dytiscidae de la region Indo-Sino-Malaise. Annales de la Societe Entomologique de France 68: 186 - 367. https: // doi. org / 10.1080 / 21686351.1899. 12279737" title="Révision des Dytiscidae de la région Indo-Sino-Malaise." volume="68" year="1899">Régimbart, 1899</bibRefCitation>
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</taxonomicName>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="B254E6062C9FA0119F3CC27E03C2F4B3" type="description">
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<paragraph id="18E3F52B0B1D4BB16EC382AB826CE9E5">
|
||
<figureCitation id="EE6C99B9AE9D7FFC61AFF32CAD22BFBD" captionStart="Figures 1–3" captionStartId="F1" captionText="Figures 1–3. Habitus of 1 Sandracottus bakewellii bakewellii (Northern Territory, Australia) 2 S. bizonatus (Borneo, Sabah, Kinabalu) 3 S. chevrolati (Sumba Island, Indonesia)." figureDoi="10.3897/zookeys.1223.138220.figures1-3" httpUri="https://binary.pensoft.net/fig/1214408">Figs 2</figureCitation>
|
||
,
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||
<figureCitation id="43793575AFC1BBA2932C47AA442A2FD6" captionStart="Figures 12–15" captionStartId="F4" captionText="Figures 12–15. Median lobe of aedeagus in ventral view (a), and right paramere in lateral view (b) 12 Sandracottus bakewellii bakewellii 13 S. bizonatus 14 S. chevrolati and 15 S. dejeanii." figureDoi="10.3897/zookeys.1223.138220.figures12-15" httpUri="https://binary.pensoft.net/fig/1214411">13</figureCitation>
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||
,
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||
<figureCitation id="D733CFB07D5BBAE254A254A569E97CBE" captionStart="Figure 24" captionStartId="F8" captionText="Figure 24. Distribution of Sandracottus bizonatus [red squares] and S. maculatus [black dots] in Southeast Asia." figureDoi="10.3897/zookeys.1223.138220.figure24" httpUri="https://binary.pensoft.net/fig/1214415">24</figureCitation>
|
||
,
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||
<figureCitation id="78245072D1EEB9E51D155AF97ED7622C" captionStart="Figures 46–49" captionStartId="F18" captionText="Figures 46–49. Colouration and habitus of 46 Sandracottus bakewellii bakewellii (Australia, Northern Queensland, Atherton Tableland, Mareeba) 47 S. bakewellii guttatus (Australia, Northern Territory, Ormiston Gorge) 48 S. bizonatus (Borneo, Sabah, Keningau) 49 S. chevrolati (Indonesia, Sumba Island, Waingapu)." figureDoi="10.3897/zookeys.1223.138220.figures46-49" httpUri="https://binary.pensoft.net/fig/1214425">48</figureCitation>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="4204E964AF899469CFEADDBD48B722E9" type="reference_group">
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<paragraph id="D12BBB85517AE55D17EF150A5A585CF0">
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||
<treatmentCitationGroup id="24465EA1FBCECE46BBA0644469CFF746">
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||
<treatmentCitation id="9A4268B7581D47F386154FB4B321ED37">
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||
<taxonomicName id="9BE652F42A584614B6B54167E8948C09" ID-CoL="6XHLD" authority="Regimbart, 1899 a: 336" authorityName="Regimbart Hendrich & Brancucci" authorityPageNumber="336" authorityYear="1899" class="Insecta" family="Dytiscidae" genus="Sandracottus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="bizonatus">
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||
<emphasis id="13FCD653D2C90DE6279938B082B05376" italics="true">Sandracottus bizonatus</emphasis>
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||
<comment id="02EF7204C080A1D886FCAD6BE8EEE414">
|
||
<bibRefCitation id="7B40071AD2E91988BBC727039EE52900" DOI="10.1080/21686351.1899.12279737" author="Régimbart M" journalOrPublisher="Annales de la Société Entomologique de France" pagination="186-367" refId="B42" refString="Regimbart M (1899) Revision des Dytiscidae de la region Indo-Sino-Malaise. Annales de la Societe Entomologique de France 68: 186 - 367. https: // doi. org / 10.1080 / 21686351.1899. 12279737" title="Révision des Dytiscidae de la région Indo-Sino-Malaise." volume="68" year="1899">Régimbart, 1899</bibRefCitation>
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||
a: 336 (
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||
<typeStatus id="40C1B10792C51A623D0B8F9E7B42760E">type</typeStatus>
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||
locality
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||
<collectingCountry id="A982A65597FA5996BB91B1F8E2D4D064" name="Malaysia">Malaysia</collectingCountry>
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||
,
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||
<collectingRegion id="3B46D514997509A9F5553B82B2929BA2" country="Malaysia" name="Sabah">Sabah</collectingRegion>
|
||
);
|
||
<treatmentCitation id="8C89E948C79251367DDE8740D0184F6B" author="Zimmermann A" page="234" year="1920">
|
||
<bibRefCitation id="1668FF9D2ED78AA0325F9B1B5B1F0D93" author="Zimmermann A" firstAuthor="Zimmermann" journalOrPublisher="In: Junk W, Schenkling S (Eds) Berlin, Coleopterorum Catalogus 4, pars 71" page="234" refId="B60" refString="Zimmermann A (1920) Dytiscidae. In: Junk W, Schenkling S (Eds) Berlin, Coleopterorum Catalogus 4, pars 71, 326 pp." title="Dytiscidae." year="1920">Zimmermann 1920: 234</bibRefCitation>
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||
</treatmentCitation>
|
||
(cat.); Hájek and
|
||
<treatmentCitation id="A0041EA1F24F0E6529895A95EBC6ACB7" author="Nilsson A & Hájek J" page="91" year="2024">
|
||
<bibRefCitation id="847E66D003F64CB7549C2AC0CE197BBA" author="Nilsson A & Hájek J" refId="B38" refString="Nilsson A, Hajek J (2024) A world catalogue of the family Dytiscidae, or the diving beetles (Coleoptera, Adephaga). Version 1. I. 2024. Distributed as a PDF file via Internet, 323 pp. http: // www. waterbeetles. eu / documents / W _ CAT _ Dytiscidae _ 2024. pdf" url="http://www.waterbeetles.eu/documents/W_CAT_Dytiscidae_2024.pdf" year="2024">Nilsson 2024: 91</bibRefCitation>
|
||
</treatmentCitation>
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||
(cat.).
|
||
</comment>
|
||
</taxonomicName>
|
||
</treatmentCitation>
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0EXPBG" type="type material">
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||
<paragraph id="A17B4BD1E7878614AD97E8A9436707B6">
|
||
<heading id="C39056C1BD93958B858CD730816F62B3" reason="title">Type material.</heading>
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||
</paragraph>
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||
<paragraph id="849ACE63B461EBCB638ABC47C0AB6ABF">
|
||
<materialsCitation id="A528DAC5FC255C81EB19813DBD1878D5" collectionCode="MNHN" specimenCount="2" typeStatus="lectotype">
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||
<emphasis id="9E1D238CE6B38CD93CBDA0A8B0F7069F" bold="true" italics="true">
|
||
<typeStatus id="A04099E967ADF54CBBF3A7ECBAA5ADE8" type="lectotype">Lectotype</typeStatus>
|
||
</emphasis>
|
||
(herewith designated): Male, “ Borneo Sandakan Windrath ” [white printed label],
|
||
<taxonomicName id="58ECAF18CC7C8001AB7927C28566AFD5" authority="Reg." authorityName="Regimbart Hendrich & Brancucci" authorityYear="1899" class="Insecta" family="Dytiscidae" genus="Sandracottus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="bizonatus">
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||
“
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||
<emphasis id="858B3DF3EBF0EC88AA4F8328655FAF02" italics="true">bizonatus</emphasis>
|
||
Reg.
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||
</taxonomicName>
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||
” [handwritten label by Régimbart], “
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||
<typeStatus id="2AF191E9862294AEA9180DEA179DB0E3" type="lectotype">Lectotype</typeStatus>
|
||
<taxonomicName id="86DCBD4A842EE022A089D7C36B4615D4" authority="Regimbart Hendrich & Brancucci" authorityName="Regimbart Hendrich & Brancucci" authorityYear="1899" class="Insecta" family="Dytiscidae" genus="Sandracottus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="bizonatus">
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||
<emphasis id="3CE2DECE09694512CABDE327A38A9FFF" italics="true">Sandracottus bizonatus</emphasis>
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||
Régimbart Hendrich & Brancucci
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||
</taxonomicName>
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||
des. ” [red printed label] (
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||
<abbrev id="ABBRID0EVQBG" xlink_title="Muséum national d´Histoire naturelle, Paris">
|
||
<collectionCode id="6C6FE9DA7B59267511816E08F6763FD2" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" type="">MNHN</collectionCode>
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||
</abbrev>
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||
). Examined.
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||
</materialsCitation>
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||
</paragraph>
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||
<caption id="D795EABEF4F46293083E0C8BD0324277" ID-DOI="10.3897/zookeys.1223.138220.figures20-22" ID-arpha="72446F20-D1B7-53EA-A655-996B2DCCF488" httpUri="https://binary.pensoft.net/fig/1214413" startId="F6">
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||
<paragraph id="BC53747545DCB9F7AB340EC344377393">
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||
<label id="C0D76CB3B95926B9A561718D779AB36B">Figures 20–22.</label>
|
||
</paragraph>
|
||
<paragraph id="7493F6AC33E539AD6F4E65EBFE4627BD">
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||
Median lobe of aedeagus in ventral view (a), and right paramere in lateral view (b):
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||
<emphasis id="EDBA7F28D0F2C09DCDF2872F82643792" bold="true">20</emphasis>
|
||
<taxonomicName id="22B47A903576FC2ABB08DFE21AFDDCDE" authorityName="Wewalka & Vazirani" authorityYear="1975" class="Insecta" family="Dytiscidae" genus="Sandracottus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="jaechi">
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||
<emphasis id="985E7F6F8637A208AD62512D266210A0" italics="true">Sandracottus jaechi</emphasis>
|
||
</taxonomicName>
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||
<emphasis id="70A32489111D00E6C4A1570F1664C881" bold="true">21</emphasis>
|
||
<taxonomicName id="A6B58CAD060C6FD2FF672354578ABBA7" authorityName="Hendrich & Brancucci" baseAuthorityName="Wehncke" baseAuthorityYear="1876" class="Insecta" family="Dytiscidae" genus="Sandracottus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="maculatus">
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||
<emphasis id="9D42FB7C2A96D9D493E25FBEEE43222E" italics="true">S. maculatus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<emphasis id="D49C5A25ADE78A04DCEF6DFCF68FF5A7" bold="true">22</emphasis>
|
||
<taxonomicName id="2DA8877D5B62C75563DFA81D8F220EB1" authorityName="Sharp" authorityYear="1882" class="Insecta" family="Dytiscidae" genus="Sandracottus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="rotundus">
|
||
<emphasis id="BB6CEA13DEA403ECAEECB90287C80833" italics="true">S. rotundus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0ENSBG" type="material">
|
||
<paragraph id="313A766D61065F7B6D52D82D71FC5984">
|
||
<heading id="5153A237EA7F0E5201243FDFB9A29765" reason="title">Additional material.</heading>
|
||
</paragraph>
|
||
<paragraph id="91E186E210634E8AD6E53D68930E34DB">
|
||
<emphasis id="402FD1917692F6D6DE9435E4A3648CAA" bold="true">
|
||
(
|
||
<specimenCount id="07E06F16AB73435135929B8E8F5D1ADC" count="45" type="generic">45 specimens</specimenCount>
|
||
)
|
||
</emphasis>
|
||
:
|
||
<emphasis id="6C0882CBB5C03C414BAE74BC3C5C482F" bold="true">
|
||
<collectingCountry id="679115C5CE7A54E7A140EAD70232B0A1" name="Malaysia">Malaysia</collectingCountry>
|
||
.
|
||
</emphasis>
|
||
•
|
||
<materialsCitation id="E191030FD98E604A0B19B3A469E3E87F" collectingDate="1903-08-05" collectingDateMax="2006-05-04" collectingDateMin="1903-08-05" collectionCode="NHMUK, CAS, MNHN, RMNH, CGW, NMB, NMW, CLH, ZSM, SMTD" collectorName="E. Malaysia & Local & A. Skale Hof & N. Borneo & H. E. Andrewes Bequest. & N. Borneo Mt. Kina Balu & J. Waterstradt & H. M. Pendlbury Collection & H. Bolle Berlin & Bolm & J. Kodada & F. Ciampor & Samml. A. Zimmermann & Nordost Borneo Gebruder W. Muller Vermachtnis" country="Malaysia" county="Borneo" elevation="1500" location="Steven Chew" municipality="Ranau" specimenCount="38" stateProvince="Sabah">
|
||
<emphasis id="14BB96247AF1E2058694D5889E8D667D" bold="true">
|
||
<collectingRegion id="12145E5FEDA249563B8A99C3D5D09924" country="Malaysia" name="Sabah">Sabah</collectingRegion>
|
||
</emphasis>
|
||
:
|
||
<specimenCount id="F097D49167ABDAAB6153415BD6C243C6" count="1" type="generic">1 ex.</specimenCount>
|
||
, “
|
||
<collectingCounty id="8B0B2A433945D55255DF854F91494AE6">Borneo</collectingCounty>
|
||
,
|
||
<collectingRegion id="6A2C7B5C36793A66B072AA730F096964" country="Malaysia" name="Sabah">Sabah</collectingRegion>
|
||
<collectingMunicipality id="D310F76963A1882A7ABB923B1DC0F9FB">Ranau</collectingMunicipality>
|
||
,
|
||
<date id="2D7F3F4812131B19DFBAAA747CEFB0C0" value="2006-05-04">
|
||
<collectingDate id="33778AB7C53809B4B9F99A84C060C52E" value="2006-05-04">4. v. 2006</collectingDate>
|
||
</date>
|
||
<location id="16CEA4DEEDBA66784822107B8D15019E" LSID="urn:lsid:plazi:treatment:A88F9289A45B52BEB5326BFDB80550D2:16CEA4DEEDBA66784822107B8D15019E" country="Malaysia" county="Borneo" municipality="Ranau" name="Steven Chew" stateProvince="Sabah">Steven Chew</location>
|
||
<abbrev id="ABBRID0EZSBG" xlink_title="Natural History Museum, London">NHMUK</abbrev>
|
||
2006-36 ” (
|
||
<abbrev id="ABBRID0E4SBG" xlink_title="Natural History Museum, London">
|
||
<collectionCode id="0F0677A23562EDF8655B7FE3DAA426BC" country="United Kingdom" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
|
||
</abbrev>
|
||
);
|
||
<specimenCount id="CB8775FF4E88BF9C4FABE9BFD32D5379" count="1" type="generic">1 ex.</specimenCount>
|
||
, “
|
||
<collectorName id="AC39AB1E4738903535FD5746F3565561">E. Malaysia</collectorName>
|
||
,
|
||
<collectingRegion id="0C2282251754CE3C989383530BED2C8B" country="Malaysia" name="Sabah">Sabah</collectingRegion>
|
||
<location id="BB977581723C99E29F4E257EF94646A6" LSID="urn:lsid:plazi:treatment:A88F9289A45B52BEB5326BFDB80550D2:BB977581723C99E29F4E257EF94646A6" country="Malaysia" county="Borneo" municipality="Ranau" name="Borneo" stateProvince="Sabah">Borneo</location>
|
||
,
|
||
<location id="5DB6336CDFFDF3C36BBFE23F9ED20E6F" LSID="urn:lsid:plazi:treatment:A88F9289A45B52BEB5326BFDB80550D2:5DB6336CDFFDF3C36BBFE23F9ED20E6F" country="Malaysia" county="Borneo" municipality="Ranau" name="Mt. Trus March-April" stateProvince="Sabah">Mt. Trus March-April</location>
|
||
2010
|
||
<collectorName id="832C65818DF9A635458A32975A69F07E">Local</collectorName>
|
||
leg. ”, “ coll.
|
||
<collectorName id="DCEC87F92775586679AC0AC3EEC42D42">A. Skale Hof</collectorName>
|
||
/
|
||
<location id="8C00F636C2FD527ACBB830B3FAABD5FB" LSID="urn:lsid:plazi:treatment:A88F9289A45B52BEB5326BFDB80550D2:8C00F636C2FD527ACBB830B3FAABD5FB" country="Malaysia" county="Borneo" municipality="Ranau" name="Germany" stateProvince="Sabah">Germany</location>
|
||
” (
|
||
<abbrev id="ABBRID0EBTBG" xlink_title="Collection André Skale">
|
||
<collectionCode id="526A8144E9E0E27464DEFB8DC42B8AAB" country="USA" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" type="Museum">CAS</collectionCode>
|
||
</abbrev>
|
||
);
|
||
<specimenCount id="E9D6D4463D0EABAE1FF344F95E6B189B" count="5" type="generic">5 exs.</specimenCount>
|
||
, “
|
||
<location id="2278315CC65ABCF61D2F605CBA135BA2" LSID="urn:lsid:plazi:treatment:A88F9289A45B52BEB5326BFDB80550D2:2278315CC65ABCF61D2F605CBA135BA2" country="Malaysia" county="Borneo" municipality="Ranau" name="Nord Borneo Mt." stateProvince="Sabah">
|
||
Nord Borneo
|
||
<location id="9638E59B317F29EABC1F8D2D06D03A9E" LSID="urn:lsid:plazi:treatment:A88F9289A45B52BEB5326BFDB80550D2:9638E59B317F29EABC1F8D2D06D03A9E" country="Malaysia" county="Borneo" municipality="Ranau" name="Mt. Kina-Balu" stateProvince="Sabah">Mt. Kina-Balu</location>
|
||
</location>
|
||
<date id="BB167181320371260D3F253DD3A8D5E3" value="1903-08-05">
|
||
<collectingDate id="6B7A7BCBB02CBEC58C3A879551D281D8" value="1903-08-05">5. VIII. 1903</collectingDate>
|
||
</date>
|
||
<location id="4A516E4DBBB56E149F1BF7D7ABB9714C" LSID="urn:lsid:plazi:treatment:A88F9289A45B52BEB5326BFDB80550D2:4A516E4DBBB56E149F1BF7D7ABB9714C" country="Malaysia" county="Borneo" municipality="Ranau" name="John Waterstradth" stateProvince="Sabah">John Waterstradth</location>
|
||
” (
|
||
<abbrev id="ABBRID0EFTBG" xlink_title="Muséum national d´Histoire naturelle, Paris">
|
||
<collectionCode id="9AB624143B250E1DEB4D0E16ED1FDEF0" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" type="">MNHN</collectionCode>
|
||
</abbrev>
|
||
);
|
||
<specimenCount id="CCCFE62698E886DF42D10BBD9AE3F46C" count="2" type="generic">2 ex.</specimenCount>
|
||
, “
|
||
<location id="FDDE1369E110ECD6C9A2FD921E99CFCD" LSID="urn:lsid:plazi:treatment:A88F9289A45B52BEB5326BFDB80550D2:FDDE1369E110ECD6C9A2FD921E99CFCD" country="Malaysia" county="Borneo" municipality="Ranau" name="Kinabalu Borneo" stateProvince="Sabah">Kinabalu Borneo</location>
|
||
” (
|
||
<abbrev id="ABBRID0EJTBG" xlink_title="Muséum national d´Histoire naturelle, Paris">
|
||
<collectionCode id="6291BCEE507AB7209A0667024DF6F691" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" type="">MNHN</collectionCode>
|
||
</abbrev>
|
||
);
|
||
<specimenCount id="2046C72BB17AB434358ADBB155B6C275" count="1" type="generic">1 ex.</specimenCount>
|
||
, “
|
||
<collectorName id="A1A31B0B71B0AEFBA074E079CF4CD5B8">N. Borneo</collectorName>
|
||
,
|
||
<location id="38D8C61E822B49858FFDFC156DF01756" LSID="urn:lsid:plazi:treatment:A88F9289A45B52BEB5326BFDB80550D2:38D8C61E822B49858FFDFC156DF01756" country="Malaysia" county="Borneo" municipality="Ranau" name="Kinabalu" stateProvince="Sabah">Kinabalu</location>
|
||
” (
|
||
<abbrev id="ABBRID0ENTBG" xlink_title="Muséum national d´Histoire naturelle, Paris">
|
||
<collectionCode id="EE2A881B97344B793B33334469D42150" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" type="">MNHN</collectionCode>
|
||
</abbrev>
|
||
);
|
||
<specimenCount id="33FAA0D1CD8048A0C187EFEBD1D437CF" count="8" type="generic">8 exs.</specimenCount>
|
||
, “ v. d.
|
||
<location id="38E914122C70D71ECB149CE9918BFC1C" LSID="urn:lsid:plazi:treatment:A88F9289A45B52BEB5326BFDB80550D2:38E914122C70D71ECB149CE9918BFC1C" country="Malaysia" county="Borneo" municipality="Ranau" name="Does de Beye" stateProvince="Sabah">Does de Beye</location>
|
||
,
|
||
<collectingRegion id="402E01085073CC04114518B1E2FE01FD" country="Malaysia" name="Melaka">Malakka</collectingRegion>
|
||
” (
|
||
<abbrev id="ABBRID0ERTBG" xlink_title="Nationaal Natuurhistorisch Museum">
|
||
<collectionCode id="EA9900FB6CD8E49A5DF86ECDDB14EA6E" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" type="Museum">RMNH</collectionCode>
|
||
</abbrev>
|
||
);
|
||
<specimenCount id="99BF245FCF5BDF1108AE52987E906338" count="1" type="generic">1 ex.</specimenCount>
|
||
, “ Borneo,
|
||
<collectorName id="55D0B15B227751859D079FE32FEA181B">H. E. Andrewes Bequest.</collectorName>
|
||
B. M. 1922-221 ” (
|
||
<abbrev id="ABBRID0EVTBG" xlink_title="Natural History Museum, London">
|
||
<collectionCode id="29B2E45F15018D6269D963A5CD601852" country="United Kingdom" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
|
||
</abbrev>
|
||
);
|
||
<specimenCount id="7EA7932CE63ABC80A34DE066AC630B5D" count="1" type="generic">1 ex.</specimenCount>
|
||
, “
|
||
<location id="0FE6620363F712F2C52AB191B5DE2E57" LSID="urn:lsid:plazi:treatment:A88F9289A45B52BEB5326BFDB80550D2:0FE6620363F712F2C52AB191B5DE2E57" country="Malaysia" county="Borneo" municipality="Ranau" name="Kina Balu" stateProvince="Sabah">Kina Balu</location>
|
||
” (
|
||
<abbrev id="ABBRID0EZTBG" xlink_title="Muséum national d´Histoire naturelle, Paris">
|
||
<collectionCode id="2760DDDA7E6780B60450CF41F0D919F4" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" type="">MNHN</collectionCode>
|
||
</abbrev>
|
||
);
|
||
<specimenCount id="E3B2E608307F2F5BA558F12D5492B884" count="2" type="generic">2 exs.</specimenCount>
|
||
, ”
|
||
<collectorName id="6BE0ADA764E64CF08ABF331E81352D82">
|
||
N.
|
||
<location id="0803F9F8BBC4877C9CFC07544CE71180" LSID="urn:lsid:plazi:treatment:A88F9289A45B52BEB5326BFDB80550D2:0803F9F8BBC4877C9CFC07544CE71180" country="Malaysia" county="Borneo" municipality="Ranau" name="Borneo Mt." stateProvince="Sabah">
|
||
Borneo
|
||
<location id="44BB54F0591087E4F2A36C1FFD2CA86E" LSID="urn:lsid:plazi:treatment:A88F9289A45B52BEB5326BFDB80550D2:44BB54F0591087E4F2A36C1FFD2CA86E" country="Malaysia" county="Borneo" municipality="Ranau" name="Mt. Kina Balu" stateProvince="Sabah">Mt. Kina Balu</location>
|
||
</location>
|
||
</collectorName>
|
||
<date id="A94B5C0C43595E025096133037FC5246" value="1903-08-05">
|
||
<collectingDate id="445C08B41C4EAB8B3E831AF923B9DAE1" value="1903-08-05">5. VIII. 1903</collectingDate>
|
||
</date>
|
||
<collectorName id="10B4503F9C0577C2FA266FA502789C4F">J. Waterstradt</collectorName>
|
||
leg. ” (
|
||
<abbrev id="ABBRID0E4TBG" xlink_title="Muséum national d´Histoire naturelle, Paris">
|
||
<collectionCode id="99B4092222D64F33D5F9ED390AA186B7" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" type="">MNHN</collectionCode>
|
||
</abbrev>
|
||
);
|
||
<specimenCount id="E3506774D18C9122BDF0E70587355951" count="1" type="generic">1 ex.</specimenCount>
|
||
, “
|
||
<location id="9D205E0861000A1F31B6A698AC2B3B2B" LSID="urn:lsid:plazi:treatment:A88F9289A45B52BEB5326BFDB80550D2:9D205E0861000A1F31B6A698AC2B3B2B" country="Malaysia" county="Borneo" municipality="Ranau" name="Borneo" stateProvince="Sabah">Borneo</location>
|
||
,
|
||
<collectingRegion id="794BC0A74870E9177E509D1E0B6242D8" country="Malaysia" name="Sabah">Sabah</collectingRegion>
|
||
<location id="BD90293A04A4A450061F2A82F7CAA4AA" LSID="urn:lsid:plazi:treatment:A88F9289A45B52BEB5326BFDB80550D2:BD90293A04A4A450061F2A82F7CAA4AA" country="Malaysia" county="Borneo" municipality="Ranau" name="Mt. Kinabalu" stateProvince="Sabah">Mt. Kinabalu</location>
|
||
3000 f,
|
||
<date id="777E3C676F589906ADDCA732208E2D3B" value="1929-04-21">
|
||
<collectingDate id="56B0B39340C08C981B8B6965044DEED4" value="1929-04-21">21. IV. 1929</collectingDate>
|
||
</date>
|
||
ex
|
||
<collectorName id="3B501F1BAB5ED0CE35737B907D63F35A">H. M. Pendlbury Collection</collectorName>
|
||
” (
|
||
<abbrev id="ABBRID0EBUBG" xlink_title="Collection Günther Wewalka">
|
||
<collectionCode id="8594A261D96C453AEF6CD7B5E319A3BE">CGW</collectionCode>
|
||
</abbrev>
|
||
);
|
||
<specimenCount id="B9A92FD65227B70B40EA8B08983C74C7" count="1" type="generic">1 ex.</specimenCount>
|
||
, “
|
||
<location id="4E4FD7178775107DD62282A6CA8A6A1B" LSID="urn:lsid:plazi:treatment:A88F9289A45B52BEB5326BFDB80550D2:4E4FD7178775107DD62282A6CA8A6A1B" country="Malaysia" county="Borneo" municipality="Ranau" name="Borneo Kinabalu" stateProvince="Sabah">Borneo Kinabalu</location>
|
||
<quantity id="E3E58442886719D13CFB51D625DF1153" metricMagnitude="3" metricUnit="m" metricValue="1.5" unit="m" value="1500.0">
|
||
<elevation id="7C9A63012CD08DC2972603A7977ECC0B" metricMagnitude="3" metricUnit="m" metricValue="1.5" unit="m" value="1500.0">1500 m</elevation>
|
||
</quantity>
|
||
<collectorName id="99CD806E470EB97C8D217F721F8327C8">H. Bolle Berlin</collectorName>
|
||
SW 11 ” (
|
||
<abbrev id="ABBRID0EFUBG" xlink_title="Muséum national d´Histoire naturelle, Paris">
|
||
<collectionCode id="41C3B7E1158B4789ED59D0F75F3D04AA" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" type="">MNHN</collectionCode>
|
||
</abbrev>
|
||
);
|
||
<specimenCount id="FCE38B0FB9CA36BFCAD717150F3DC6FF" count="1" type="generic">1 ex.</specimenCount>
|
||
, “ Borneo,
|
||
<collectingRegion id="B3E6CDE8A2CCA935C22F15494409C9E1" country="Malaysia" name="Sabah">Sabah</collectingRegion>
|
||
, Tibow,
|
||
<locationDeviation id="F1776257C03CE5618858142D16FFEA0F" location="Sapulut">
|
||
<quantity id="A1B8634090E7860C5847D71172014645" metricMagnitude="4" metricUnit="m" metricValue="4.5" unit="km" value="45.0">45 km</quantity>
|
||
NE of Sapulut
|
||
</locationDeviation>
|
||
,
|
||
<quantity id="0F3D594763CD4E1490B174839A40121C" metricMagnitude="2" metricUnit="m" metricValue="7.5" metricValueMax="9.0" metricValueMin="6.0" unit="m" value="750.0" valueMax="900.0" valueMin="600.0">
|
||
<elevation id="A31EBAF070867BFCB8D0D40D2E595794" metricMagnitude="2" metricUnit="m" metricValue="7.5" metricValueMax="9.0" metricValueMin="6.0" unit="m" value="750.0" valueMax="900.0" valueMin="600.0">600-900 m</elevation>
|
||
</quantity>
|
||
,
|
||
<date id="38B5D3B6636F1611DACDE7DF03F05ED3" value="2000-04-07" valueMax="2000-04-15" valueMin="2000-04-07">
|
||
<collectingDate id="F8E5D02A29A824D787B2C0EAAE59CECE" value="2000-04-07" valueMax="2000-04-15" valueMin="2000-04-07">7. - 15. IV. 2000</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="C982C3DBC31308A2DE31AA7AF6B6CFA5">Bolm</collectorName>
|
||
leg. ” (
|
||
<abbrev id="ABBRID0EJUBG" xlink_title="Naturhistorisches Museum Basel">
|
||
<collectionCode id="467C9F4065F3F12F628675A4E0DD4FD9" country="Switzerland" httpUri="http://grbio.org/cool/8wbz-jkc3" name="Naturhistorishes Museum" type="">NMB</collectionCode>
|
||
</abbrev>
|
||
);
|
||
<specimenCount id="0C32076186AC61A11E0B2D50D8797A5A" count="1" type="generic">1 ex.</specimenCount>
|
||
, “ Borneo,
|
||
<collectingRegion id="0CD987C6574D9D98B1036D4F96BFB2FC" country="Malaysia" name="Sabah">Sabah</collectingRegion>
|
||
, Kampung Pisang, Pisang env., tributary of
|
||
<location id="DADABA9F94F1377095F84FFBCA3231F3" LSID="urn:lsid:plazi:treatment:A88F9289A45B52BEB5326BFDB80550D2:DADABA9F94F1377095F84FFBCA3231F3" country="Malaysia" county="Borneo" municipality="Ranau" name="Kuamut river" stateProvince="Sabah">Kuamut river</location>
|
||
,
|
||
<date id="7A3736297FDFAB0A1DA290CE118E95F0" value="1998-06-29">
|
||
<collectingDate id="33081F760D8EFC083F309E585ABE6CCB" value="1998-06-29">29. VI. 1998</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="19AFF824B19C7295871C22E3E9500FFF">J. Kodada</collectorName>
|
||
&
|
||
<collectorName id="3BC54AC71FB82D7945247B1CC6C631A3">F. Ciampor</collectorName>
|
||
leg. ” (
|
||
<abbrev id="ABBRID0ENUBG" xlink_title="Naturhistorisches Museum Wien">
|
||
<collectionCode id="DB532F589C6E934E742A9F0C7B83510D" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" type="">NMW</collectionCode>
|
||
</abbrev>
|
||
);
|
||
<specimenCount id="5329FD14793863074F2EF6A8D61DF35C" count="1" type="generic">1 ex.</specimenCount>
|
||
, “
|
||
<collectingRegion id="746A8946F22D4EF327894BC093C907BE" country="Malaysia" name="Sabah">Sabah</collectingRegion>
|
||
, Borneo env. Keningau
|
||
<date id="4E864CB513A6BEC7139B6C1B88C03999" value="1993-05">
|
||
<collectingDate id="1A99201A3518DB65D67BCF4035D793B0" value="1993-05">V. 1993</collectingDate>
|
||
</date>
|
||
” (
|
||
<abbrev id="ABBRID0ERUBG" xlink_title="Collection Lars Hendrich">
|
||
<collectionCode id="F7540F99F0979E50CADF26B3C58903E6">CLH</collectionCode>
|
||
</abbrev>
|
||
);
|
||
<specimenCount id="D0462044199D4E4F262F7EC826879541" count="2" type="generic">2 exs.</specimenCount>
|
||
, “ Nord Borneo ”, “
|
||
<collectorName id="2FBFBC82855C87150888764273421A0E">Samml. A. Zimmermann</collectorName>
|
||
” (
|
||
<abbrev id="ABBRID0EVUBG" xlink_title="Zoologische Staatssammlung, München">
|
||
<collectionCode id="45CF2FB80158B5B3F0BBB1957691662D" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" type="Museum">ZSM</collectionCode>
|
||
</abbrev>
|
||
);
|
||
<specimenCount id="093B5FFFD5E9D087793BD0ECAC0F7AFE" count="2" type="generic">2 exs.</specimenCount>
|
||
, “ Kinabalu Nord Borneo ”, “
|
||
<collectorName id="3B0B62EC0D6B3201BE5B3C45448B7AD1">Samml. A. Zimmermann</collectorName>
|
||
” (
|
||
<abbrev id="ABBRID0EZUBG" xlink_title="Zoologische Staatssammlung, München">
|
||
<collectionCode id="6234DB70098BC170D1E4434ACA4B7284" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" type="Museum">ZSM</collectionCode>
|
||
</abbrev>
|
||
);
|
||
<specimenCount id="BF5BF78ED41DA9AFCDB669178C29D8C7" count="5" type="generic">5 exs.</specimenCount>
|
||
, “ Kinabalu Borneo
|
||
<quantity id="1098E2BD862EF0F47F0123B26766CF0F" metricMagnitude="3" metricUnit="m" metricValue="1.5" unit="m" value="1500.0">
|
||
<elevation id="3C0433DC45D3C9A9E61BDE17D2FF9820" metricMagnitude="3" metricUnit="m" metricValue="1.5" unit="m" value="1500.0">1500 m</elevation>
|
||
</quantity>
|
||
”, “
|
||
<collectorName id="3D803A8F23C4E68BD3BE3133B9E81E92">Samml. A. Zimmermann</collectorName>
|
||
” (
|
||
<abbrev id="ABBRID0E4UBG" xlink_title="Zoologische Staatssammlung, München">
|
||
<collectionCode id="AC1D17403732BCB3CB9943543A864003" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" type="Museum">ZSM</collectionCode>
|
||
</abbrev>
|
||
);
|
||
<specimenCount id="79D0C02552DE774875A426BCE66E7023" count="1" type="generic">1 ex.</specimenCount>
|
||
, “
|
||
<collectorName id="D85C87923E2E4121CBD8429267FE13B3">Nordost Borneo Gebrüder W. Müller Vermächtnis</collectorName>
|
||
1909 ” (
|
||
<abbrev id="ABBRID0EBVBG" xlink_title="Staatliches Museum für Tierkunde">
|
||
<collectionCode id="65DDD31D4392211C6DB77CF7DAB632C7">SMTD</collectionCode>
|
||
</abbrev>
|
||
);
|
||
<specimenCount id="19624B9B6AE363883FC2E470F092BAC0" count="1" type="generic">1 ex.</specimenCount>
|
||
, without locality label, “
|
||
<taxonomicName id="BE42EBC2DEABF1485547C58AED25EAE7" authority="Gschwendtner" authorityName="Gschwendtner" class="Insecta" family="Dytiscidae" genus="Sandracottus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="bizonatus">
|
||
<emphasis id="5B6EC3D41800DF3BB5F581DEFD83B893" italics="true">Sandracottus bizonatus</emphasis>
|
||
Gschwendtner
|
||
</taxonomicName>
|
||
det. ” (
|
||
<abbrev id="ABBRID0EQVBG" xlink_title="Staatliches Museum für Tierkunde">
|
||
<collectionCode id="395D8B9D980AD88B193463A0329435B9">SMTD</collectionCode>
|
||
</abbrev>
|
||
)
|
||
</materialsCitation>
|
||
. •
|
||
<materialsCitation id="B659A3975B0AB04B0B585A05FD34C5DC" collectingDate="1994-03-03" collectingDateMax="2003-06-25" collectingDateMin="1994-03-03" collectionCode="NMW, CJS" collectorName="J. Horak & J. Stastny" country="Malaysia" county="Kapit distr." location="Sut river" municipality="Rumah Ugap" specimenCount="7" stateProvince="Sarawak">
|
||
<emphasis id="BF552078901296748322482627FF2FBE" bold="true">
|
||
<collectingCountry id="883078C664579F80C8B42C4EEADD4515" name="Malaysia">Sarawak</collectingCountry>
|
||
</emphasis>
|
||
:
|
||
<specimenCount id="153054524295BE42488C9FB2A7B4A122" count="1" type="generic">1 ex.</specimenCount>
|
||
, “
|
||
<collectingRegion id="522A7FE59F7F7BEA4FA37D2E82302ACC" country="Malaysia" name="Sarawak">Sarawak</collectingRegion>
|
||
,
|
||
<collectingCounty id="7EF79EB9958B02626C8DDE0A6967C24F">Kapit distr.</collectingCounty>
|
||
,
|
||
<collectingMunicipality id="06B4EC7CE172C6E3D6D00FC56D885A21">Rumah Ugap</collectingMunicipality>
|
||
vill.,
|
||
<location id="926C6817ACAF332D6C54C8A1068F4054" LSID="urn:lsid:plazi:treatment:A88F9289A45B52BEB5326BFDB80550D2:926C6817ACAF332D6C54C8A1068F4054" country="Malaysia" county="Kapit distr." municipality="Rumah Ugap" name="Sut river" stateProvince="Sarawak">Sut river</location>
|
||
,
|
||
<date id="B71BFE5FF395ED45EA46D2B88E999A8D" value="1994-03-03" valueMax="1994-03-09" valueMin="1994-03-03">
|
||
<collectingDate id="CD34DBED718629962A05F8330415D890" value="1994-03-03" valueMax="1994-03-09" valueMin="1994-03-03">3. - 9. III. 1994</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="A3800262553A3C97486AA2FB98FAC197">J. Horák</collectorName>
|
||
leg. ” (
|
||
<abbrev id="ABBRID0EWVBG" xlink_title="Naturhistorisches Museum Wien">
|
||
<collectionCode id="B3894D35404D18FF0F6E3B1B1AC26E3A" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" type="">NMW</collectionCode>
|
||
</abbrev>
|
||
);
|
||
<specimenCount id="A3D05B1DA0511D4424357AAF55374A70" count="4" type="generic">4 exs.</specimenCount>
|
||
, “
|
||
<collectingRegion id="7ABF22F91411A7C5891746D48DD0D6E9" country="Malaysia" name="Sarawak">Sarawak</collectingRegion>
|
||
, Bario env., Pa Ukat,
|
||
<date id="41A076118BDCF735368C9A52A212180B" value="2003-06-24">
|
||
<collectingDate id="B123B9C819D91C730F55833C761FC6C9" value="2003-06-24">24. VI. 2003</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="C8697B74F61D19C31728FAB21EC53DD8">J. Šťastný</collectorName>
|
||
lgt. ” (
|
||
<abbrev id="ABBRID0E1VBG" xlink_title="Collection Jaroslav Šťastný">
|
||
<collectionCode id="61C17D4BFF28ED069FDB2F69356E7BF8">CJS</collectionCode>
|
||
</abbrev>
|
||
);
|
||
<specimenCount id="5A3212F5D9BBE7108D4987A9E7A9F15A" count="2" type="generic">2 exs.</specimenCount>
|
||
, “
|
||
<collectingRegion id="DC58CA9CD2251A425729643B7E9846A7" country="Malaysia" name="Sarawak">Sarawak</collectingRegion>
|
||
, Kelabit, Bario env.
|
||
<date id="01DCF7273E98B482BD11B76592C56470" value="2003-06-21" valueMax="2003-06-25" valueMin="2003-06-21">
|
||
<collectingDate id="F7AA96FF73789D867DF1192D47D3DEF8" value="2003-06-21" valueMax="2003-06-25" valueMin="2003-06-21">21. - 25. VI. 2003</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="D47D09998CAB2927F287EEA37220960F">J. Šťastný</collectorName>
|
||
lgt. ” (
|
||
<abbrev id="ABBRID0E5VBG" xlink_title="Collection Jaroslav Šťastný">
|
||
<collectionCode id="E3DF1DFA4C66105284831AF46748B976">CJS</collectionCode>
|
||
</abbrev>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0ECWBG" type="redescription">
|
||
<paragraph id="417A7ECED8019BA4A89C870E9DDE58E9">
|
||
<heading id="A5140C180A4CB0D946D64CC213CB8FFF" reason="title">Redescription.</heading>
|
||
</paragraph>
|
||
<paragraph id="67ED9E1267DD22BD7C81B924CAD767B5">
|
||
Body broad oval, shiny, reddish brown with broad black markings on elytra (Figs
|
||
<figureCitation id="EDD260821DBC9FE08E1753CC9FE41BD8" captionStart="Figures 1–3" captionStartId="F1" captionText="Figures 1–3. Habitus of 1 Sandracottus bakewellii bakewellii (Northern Territory, Australia) 2 S. bizonatus (Borneo, Sabah, Kinabalu) 3 S. chevrolati (Sumba Island, Indonesia)." figureDoi="10.3897/zookeys.1223.138220.figures1-3" httpUri="https://binary.pensoft.net/fig/1214408">2</figureCitation>
|
||
,
|
||
<figureCitation id="DA4B1D736A34F40B44F3CE4A9E211B36" captionStart="Figures 46–49" captionStartId="F18" captionText="Figures 46–49. Colouration and habitus of 46 Sandracottus bakewellii bakewellii (Australia, Northern Queensland, Atherton Tableland, Mareeba) 47 S. bakewellii guttatus (Australia, Northern Territory, Ormiston Gorge) 48 S. bizonatus (Borneo, Sabah, Keningau) 49 S. chevrolati (Indonesia, Sumba Island, Waingapu)." figureDoi="10.3897/zookeys.1223.138220.figures46-49" httpUri="https://binary.pensoft.net/fig/1214425">48</figureCitation>
|
||
). Ventral side completely dark brown to black, legs testaceous, hind legs somewhat darker.
|
||
</paragraph>
|
||
<paragraph id="0647F4D790B840542A0CE57E387A3DF6">Head ferrugineus with posterior part broadly black, shiny. Black band protruding forwards to frons. Surface almost smooth consisting of dense and very numerous punctures of different sizes and of larger, much sparser ones, particularly numerous on frons. Clypeal grooves, punctures alongside eyes and transverse depressions beside eyes distinctly impressed, punctures large and coalescent. Antennae ferrugineus; antennomeres slender, fifth 4 × as long as broad.</paragraph>
|
||
<paragraph id="6CA1F94969ACD2F47DCE547AD4B33581">
|
||
Pronotum ferrugineus with large median black marking reaching from posterior to anterior margins (Figs
|
||
<figureCitation id="747C9DB4B57A15FF0AA374582B78850C" captionStart="Figures 1–3" captionStartId="F1" captionText="Figures 1–3. Habitus of 1 Sandracottus bakewellii bakewellii (Northern Territory, Australia) 2 S. bizonatus (Borneo, Sabah, Kinabalu) 3 S. chevrolati (Sumba Island, Indonesia)." figureDoi="10.3897/zookeys.1223.138220.figures1-3" httpUri="https://binary.pensoft.net/fig/1214408">2</figureCitation>
|
||
,
|
||
<figureCitation id="307F3048D9DA4546EE01D42D28E6308D" captionStart="Figures 46–49" captionStartId="F18" captionText="Figures 46–49. Colouration and habitus of 46 Sandracottus bakewellii bakewellii (Australia, Northern Queensland, Atherton Tableland, Mareeba) 47 S. bakewellii guttatus (Australia, Northern Territory, Ormiston Gorge) 48 S. bizonatus (Borneo, Sabah, Keningau) 49 S. chevrolati (Indonesia, Sumba Island, Waingapu)." figureDoi="10.3897/zookeys.1223.138220.figures46-49" httpUri="https://binary.pensoft.net/fig/1214425">48</figureCitation>
|
||
). Surface shagreened, with dense punctation; punctures medium-sized mixed with smaller ones. Anterior and lateral puncture rows dense and coalescent, punctures becoming sparse towards middle and lacking in very middle of anterior margin. Posterior row of punctures with coarse and coalescent punctures in middle of each side, distinctly smaller and spaced on disc.
|
||
</paragraph>
|
||
<paragraph id="51B1652D2D8AD1F3AA84CA6EB79C92A0">
|
||
Elytra ferrugineus brown with black and broad markings consisting of three transverse bands (Figs
|
||
<figureCitation id="62066923421820F6E8AB63DF2FE2B4AB" captionStart="Figures 1–3" captionStartId="F1" captionText="Figures 1–3. Habitus of 1 Sandracottus bakewellii bakewellii (Northern Territory, Australia) 2 S. bizonatus (Borneo, Sabah, Kinabalu) 3 S. chevrolati (Sumba Island, Indonesia)." figureDoi="10.3897/zookeys.1223.138220.figures1-3" httpUri="https://binary.pensoft.net/fig/1214408">2</figureCitation>
|
||
,
|
||
<figureCitation id="A6D7FA8B183729D663B9C0AC982CDA60" captionStart="Figures 46–49" captionStartId="F18" captionText="Figures 46–49. Colouration and habitus of 46 Sandracottus bakewellii bakewellii (Australia, Northern Queensland, Atherton Tableland, Mareeba) 47 S. bakewellii guttatus (Australia, Northern Territory, Ormiston Gorge) 48 S. bizonatus (Borneo, Sabah, Keningau) 49 S. chevrolati (Indonesia, Sumba Island, Waingapu)." figureDoi="10.3897/zookeys.1223.138220.figures46-49" httpUri="https://binary.pensoft.net/fig/1214425">48</figureCitation>
|
||
); an antemedian one, a postmedian one and a preapical one. Epipleura ferrugineus brown. Surface of elytra very slightly and superficially shagreened and with double punctation, a smaller and dense punctation as well as a larger one much more sparsely distributed. Row of punctures with groups of medium-sized punctures mostly grouped in five or six punctures, groups closer together on discal row.
|
||
</paragraph>
|
||
<caption id="C149A0EC817BA8C57F72D9685AFE3C8E" ID-DOI="10.3897/zookeys.1223.138220.figure23" ID-arpha="A3B52DAC-89C1-589D-A7E1-6271F83DFC2B" httpUri="https://binary.pensoft.net/fig/1214414" startId="F7">
|
||
<paragraph id="D65B601BDDF7EBD670CE7559F57E981A">
|
||
<label id="C85B3C672ED998997CC4BD304F7340BF">Figure 23.</label>
|
||
</paragraph>
|
||
<paragraph id="DC49C1CF7DA0BA77A38ABE8F5ABFD60E">
|
||
Distribution of
|
||
<taxonomicName id="C03B3DEFDD85E276B642033230C4D4EB" baseAuthorityName="Clark" baseAuthorityYear="1864" class="Insecta" family="Dytiscidae" genus="Sandracottus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="subSpecies" species="bakewellii" subSpecies="bakewellii">
|
||
<emphasis id="0B1366B8269308167E1689CBF4386280" italics="true">Sandracottus bakewellii bakewellii</emphasis>
|
||
</taxonomicName>
|
||
[black dots] and
|
||
<taxonomicName id="3A8CD6943C1AB8C6A3D780D3CA544BE0" authorityName="Sharp" authorityYear="1882" baseAuthorityName="Sharp Det. C. Watts" baseAuthorityYear="1882" class="Insecta" family="Dytiscidae" genus="Sandracottus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="subSpecies" species="bakewellii" subSpecies="guttatus">
|
||
<emphasis id="FBF7A29F599CCA8F9F224510B50F81B3" italics="true">S. bakewellii guttatus</emphasis>
|
||
</taxonomicName>
|
||
[red squares] in Australia.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="3EAE54B3FDDEBCD3D06FDC8421C717AD">Ventral side dark brown. Fore and mid legs particularly testaceous, hind legs ferrugineus brown to dark brown. Prosternal process short and broad, 1.5 × longer than broad, flattened finely but distinctly sculptured; posterior margin broadly rounded. Whole surface very superficially shagreened and finely punctured. Metatibia with sparse medium-sized punctures on outer half. Ventrites II – VI very superficially shagreened, slightly and longitudinally wrinkled on lateral parts, on whole surface densely covered with very small punctures and larger and sparser ones. Posterior margins rounded, bordered with some large and coalescent punctures in middle of each side.</paragraph>
|
||
<paragraph id="92F2E0BAD030D52A53E211108224B34C">
|
||
Measurements:
|
||
<abbrev id="ABBRID0ETYBG" xlink_title="total length">TL</abbrev>
|
||
= 12.5–13.0 mm,
|
||
<abbrev id="ABBRID0EXYBG" xlink_title="total length minus head length">TL-h</abbrev>
|
||
=
|
||
<quantity id="1D1349882516DDDA2DD9B3C6D29E004C" metricMagnitude="-2" metricUnit="m" metricValue="1.18" metricValueMax="1.22" metricValueMin="1.14" unit="mm" value="11.8" valueMax="12.2" valueMin="11.4">11.4–12.2 mm</quantity>
|
||
,
|
||
<abbrev id="ABBRID0E2YBG" xlink_title="maximum width">TW</abbrev>
|
||
= 8.5–9.0 mm.
|
||
</paragraph>
|
||
<paragraph id="DDD5C403BAF9A4B5FE917D88336A90D1">
|
||
<specimenCount id="AAEC2241BF4C26F4AF73F991790C4F05" count="1" type="male">♂</specimenCount>
|
||
. Protarsomeres I – III strongly enlarged with three larger suckers and numerous smaller one. Mesotarsomeres I – III with two rows of small suckers. Median lobe of aedeagus, in ventral view, broad, parallel-sided up to apex where it is slightly broadened and broadly rounded (Fig.
|
||
<figureCitation id="BA220EDA23C46324AE2D3369F3ED5A5D" captionStart="Figures 12–15" captionStartId="F4" captionText="Figures 12–15. Median lobe of aedeagus in ventral view (a), and right paramere in lateral view (b) 12 Sandracottus bakewellii bakewellii 13 S. bizonatus 14 S. chevrolati and 15 S. dejeanii." figureDoi="10.3897/zookeys.1223.138220.figures12-15" httpUri="https://binary.pensoft.net/fig/1214411">13 a</figureCitation>
|
||
). Parameres broad, same length as median lobe, and pointed at apex (Fig.
|
||
<figureCitation id="88AA9136DBD1878C6728FB4F158B3030" captionStart="Figures 12–15" captionStartId="F4" captionText="Figures 12–15. Median lobe of aedeagus in ventral view (a), and right paramere in lateral view (b) 12 Sandracottus bakewellii bakewellii 13 S. bizonatus 14 S. chevrolati and 15 S. dejeanii." figureDoi="10.3897/zookeys.1223.138220.figures12-15" httpUri="https://binary.pensoft.net/fig/1214411">13 b</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="3B06806B457A9E0A5B0B68549A33FFE6">
|
||
<specimenCount id="43C848C84A4E5AFC501D33BC837C3FC0" count="1" type="female">♀</specimenCount>
|
||
. Similar to male, tarsi not enlarged. Microsculpture on ventrite VI as in male.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0EKZBG" type="Differential diagnosis">
|
||
<paragraph id="DE67B3C40958418AE5507553863557B4">
|
||
<heading id="42C9F21DCB2D3D3FBAE4E757A7CF3D7C" reason="title">Differential diagnosis.</heading>
|
||
</paragraph>
|
||
<paragraph id="05CF316AB8E50BCDB748B63649B9D37F">
|
||
The combination of dorsal colour pattern (Figs
|
||
<figureCitation id="8954347FB776C9779BE512C29D823D87" captionStart="Figures 1–3" captionStartId="F1" captionText="Figures 1–3. Habitus of 1 Sandracottus bakewellii bakewellii (Northern Territory, Australia) 2 S. bizonatus (Borneo, Sabah, Kinabalu) 3 S. chevrolati (Sumba Island, Indonesia)." figureDoi="10.3897/zookeys.1223.138220.figures1-3" httpUri="https://binary.pensoft.net/fig/1214408">2</figureCitation>
|
||
,
|
||
<figureCitation id="E8F8A7ECA5377DAC00481C67162D6AEF" captionStart="Figures 46–49" captionStartId="F18" captionText="Figures 46–49. Colouration and habitus of 46 Sandracottus bakewellii bakewellii (Australia, Northern Queensland, Atherton Tableland, Mareeba) 47 S. bakewellii guttatus (Australia, Northern Territory, Ormiston Gorge) 48 S. bizonatus (Borneo, Sabah, Keningau) 49 S. chevrolati (Indonesia, Sumba Island, Waingapu)." figureDoi="10.3897/zookeys.1223.138220.figures46-49" httpUri="https://binary.pensoft.net/fig/1214425">48</figureCitation>
|
||
) and shape of median lobe of aedeagus and parameres (Fig.
|
||
<figureCitation id="A8A8510BB3637F6321E427890B4B5A9B" captionStart="Figures 12–15" captionStartId="F4" captionText="Figures 12–15. Median lobe of aedeagus in ventral view (a), and right paramere in lateral view (b) 12 Sandracottus bakewellii bakewellii 13 S. bizonatus 14 S. chevrolati and 15 S. dejeanii." figureDoi="10.3897/zookeys.1223.138220.figures12-15" httpUri="https://binary.pensoft.net/fig/1214411">13 a, b</figureCitation>
|
||
) separates
|
||
<taxonomicName id="595B7693D50D2E80F1F7C6F583F1EA4D" class="Insecta" family="Dytiscidae" genus="Sandracottus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="bizonatus">
|
||
<emphasis id="413FDC3A08DDF87E34F1FFD693479D28" italics="true">S. bizonatus</emphasis>
|
||
</taxonomicName>
|
||
from all other species of the genus. The species is endemic to Borneo and co-occurs with
|
||
<taxonomicName id="FE1B99CC7325D62849BE6FDC530FF826" authorityName="Hendrich & Brancucci" baseAuthorityName="Wehncke" baseAuthorityYear="1876" class="Insecta" family="Dytiscidae" genus="Sandracottus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="maculatus">
|
||
<emphasis id="0527B4C1352E22BDC6682F82B3BE1D54" italics="true">S. maculatus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0ES1BG" type="distribution">
|
||
<paragraph id="3E004412496021C0C63B4953544FC720">
|
||
<heading id="22B71001D8111D705F68C5ECD03EB555" reason="title">Distribution.</heading>
|
||
</paragraph>
|
||
<paragraph id="84A1CCFD8C9DA1E67E5291BDEDEB2029">
|
||
<collectingCountry id="E4D87AEF9F6181706C71C63E413A6F3A" name="Malaysia">Malaysia</collectingCountry>
|
||
(
|
||
<collectingRegion id="64097D437B89B5ED5826A413B55F61C7" country="Malaysia" name="Sabah">Sabah</collectingRegion>
|
||
,
|
||
<collectingCountry id="A5EE6F69D86CC0A23ABBB502322E7990" name="Malaysia">Sarawak</collectingCountry>
|
||
) (Fig.
|
||
<figureCitation id="BCD8A7F67BCD539FC875857ED20ACC28" captionStart="Figure 24" captionStartId="F8" captionText="Figure 24. Distribution of Sandracottus bizonatus [red squares] and S. maculatus [black dots] in Southeast Asia." figureDoi="10.3897/zookeys.1223.138220.figure24" httpUri="https://binary.pensoft.net/fig/1214415">24</figureCitation>
|
||
). The
|
||
<specimenCount id="3E9BA62525C7AE2DED160A86295BA6FD" count="8" type="generic">eight specimens</specimenCount>
|
||
deposited in
|
||
<abbrev id="ABBRID0E31BG" xlink_title="Nationaal Natuurhistorisch Museum">RMNH</abbrev>
|
||
from
|
||
<collectingCountry id="14A10F5E70D0C67F4290F0CBAEBE2686" name="Malaysia">Malacca</collectingCountry>
|
||
are most probably mislabelled. According to a photo on a website from an insect dealer in
|
||
<collectingCountry id="D77958F0E67E57349661FFE325E4B8D7" name="Indonesia">Indonesia</collectingCountry>
|
||
, the species was also collected in
|
||
<collectingRegion id="7E50C23BA2A8DE958D67EF7E4F4DC27D" country="Indonesia" name="Kalimantan Utara">Kalimantan</collectingRegion>
|
||
(
|
||
<ext-link id="1723C73DDF7DC30890C8E099B58B2C01" ext-link-type="uri" xlink_href="https://www.giradis-insect.com/" xlink_type="simple">https://www.giradis-insect.com/</ext-link>
|
||
<date id="7B1AC87D745D1CC10E49F91952E4182E" value="2024-09-27">27.9.2024</date>
|
||
). Specimens were collected between 300 and
|
||
<quantity id="33E0A0A92E4C8B2F5E0FC0E2FECC76ED" metricMagnitude="0" metricUnit="m" metricValue="1.5" unit="m" value="1.5">1.500 m</quantity>
|
||
a. s. l.
|
||
</paragraph>
|
||
<caption id="513FA509DD090BA3E1D38E6F10CB2F40" ID-DOI="10.3897/zookeys.1223.138220.figure24" ID-arpha="13B10FC3-0A0A-50F3-9081-31C7B2DDA3D8" httpUri="https://binary.pensoft.net/fig/1214415" startId="F8">
|
||
<paragraph id="A1E88B48B02A3AD01AF3E69692A43671">
|
||
<label id="3AB9AA31750F12FC0BF77D0E5625A244">Figure 24.</label>
|
||
</paragraph>
|
||
<paragraph id="F944F612733432066BBC61D0D02FF420">
|
||
Distribution of
|
||
<taxonomicName id="21DC040A36A2DF4E0AD2E095C59C8763" class="Insecta" family="Dytiscidae" genus="Sandracottus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="bizonatus">
|
||
<emphasis id="6B7FCDF72D50EFA4B1D1E1A0341AE64D" italics="true">Sandracottus bizonatus</emphasis>
|
||
</taxonomicName>
|
||
[red squares] and
|
||
<taxonomicName id="FED3687EA2617DE46B9E1AF8FF9C648A" authorityName="Hendrich & Brancucci" baseAuthorityName="Wehncke" baseAuthorityYear="1876" class="Insecta" family="Dytiscidae" genus="Sandracottus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="maculatus">
|
||
<emphasis id="AADB0ACC8A8F6E9A0BB52FA34AABB19C" italics="true">S. maculatus</emphasis>
|
||
</taxonomicName>
|
||
[black dots] in Southeast Asia.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0EI3BG" type="habitat">
|
||
<paragraph id="15F41690B853B675673F2ABC4FFE6F05">
|
||
<heading id="5C25E45C1C33844220E494AD85BC1F9E" reason="title">Habitat.</heading>
|
||
</paragraph>
|
||
<paragraph id="1D2777F4CB82CD6CF5CE8859A7128B3D">
|
||
Muddy and shaded forest pools, rich in rotten leaves and more permanent side pools of larger forest streams in primary rainforests, partly shaded and enriched with rotten leaves and twigs (J. Kodada and J. Šťastný, pers. comm.
|
||
<date id="89B26B493CAFB7BF397F206504399342" value="2013-03">March 2013</date>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0EN3BG" type="Conservation">
|
||
<paragraph id="5FA68816CD9E5B6FA2E46CE45C1B3DFE">
|
||
<heading id="FA2DF8D73797BDA743C182C22E863901" reason="title">Conservation.</heading>
|
||
</paragraph>
|
||
<paragraph id="3FA952EE88C05946055BF02A5FDA8983">
|
||
A rare and highly endemic species of Borneo, probably associated with the declining primary lowland and hilly rainforests on the island (see Southeast Asian species
|
||
<taxonomicName id="90A2864EE67140C21A8A9022958109AA" authorityName="Heller" authorityYear="1934" class="Insecta" family="Dytiscidae" genus="Sandracottus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="femoralis">
|
||
<emphasis id="8382C4971335336B490FC40B563FF391" italics="true">S. femoralis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="15F992D277148D2FE951B7816E234AFF" authorityName="Hendrich" baseAuthorityName="Wehncke" baseAuthorityYear="1876" class="Insecta" family="Dytiscidae" genus="Sandracottus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="insignis">
|
||
<emphasis id="22E153AC165E7DA794ED101BC1509EA5" italics="true">S. insignis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="A9394D3059A3BEE2C1EDCD25820709AA" authorityName="Hendrich & Brancucci" baseAuthorityName="Wehncke" baseAuthorityYear="1876" class="Insecta" family="Dytiscidae" genus="Sandracottus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="maculatus">
|
||
<emphasis id="168775F2447560011C6252916CA22C52" italics="true">S. maculatus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="39829F565637EB06E326A6CF0BF100E0" authorityName="Sharp" authorityYear="1882" class="Insecta" family="Dytiscidae" genus="Sandracottus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="rotundus">
|
||
<emphasis id="86A6D037990757D65379FC7A92BD9166" italics="true">S. rotundus</emphasis>
|
||
</taxonomicName>
|
||
). Most records are from the end of the 19
|
||
<superScript id="C1ED03BFFF947B15C99AC045EB4177F4">th</superScript>
|
||
and the beginning of the 20
|
||
<superScript id="46EC1ECA866CB1990DFC21A99736CCDF">th</superScript>
|
||
century. According to present knowledge it is an endangered species. It is recommended to be listed in the next IUCN red list.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |