216 lines
22 KiB
XML
216 lines
22 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.544.6074" ID-GBIF-Dataset="0e5e47a0-f8f3-4039-b0e1-c159fef0b9f6" ID-PMC="PMC4714384" ID-Pensoft-Pub="1313-2970-544-1" ID-PubMed="26798289" ID-ZBK="C5978BD0145B40F8ACDEB27371B7B9A4" ModsDocAuthor="" ModsDocDate="2015" ModsDocID="1313-2970-544-1" ModsDocOrigin="ZooKeys 544" ModsDocTitle="The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation" checkinTime="1451243640953" checkinUser="pensoft" docAuthor="Liebherr, James K." docDate="2015" docId="A82125154158892C12BAC6FD998202AC" docLanguage="en" docName="ZooKeys 544: 1-407" docOrigin="ZooKeys 544" docSource="http://dx.doi.org/10.3897/zookeys.544.6074" docTitle="Mecyclothorax cognatus Sharp" docType="treatment" docVersion="4" lastPageNumber="56" masterDocId="FF92FFC2FF9FFFC7191EFFB8FF91AF5D" masterDocTitle="The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation" masterLastPageNumber="407" masterPageNumber="1" pageNumber="54" updateTime="1668162300918" updateUser="ExternalLinkService">
|
||
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo>
|
||
<mods:title>The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Liebherr, James K.</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource>text</mods:typeOfResource>
|
||
<mods:relatedItem type="host">
|
||
<mods:titleInfo>
|
||
<mods:title>ZooKeys</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part>
|
||
<mods:date>2015</mods:date>
|
||
<mods:detail type="volume">
|
||
<mods:number>544</mods:number>
|
||
</mods:detail>
|
||
<mods:extent unit="page">
|
||
<mods:start>1</mods:start>
|
||
<mods:end>407</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:location>
|
||
<mods:url>http://dx.doi.org/10.3897/zookeys.544.6074</mods:url>
|
||
</mods:location>
|
||
<mods:classification>journal article</mods:classification>
|
||
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.544.6074</mods:identifier>
|
||
<mods:identifier type="Pensoft-Pub">1313-2970-544-1</mods:identifier>
|
||
<mods:identifier type="ZBK">C5978BD0145B40F8ACDEB27371B7B9A4</mods:identifier>
|
||
<mods:identifier type="ZooBank">C5978BD0145B40F8ACDEB27371B7B9A4</mods:identifier>
|
||
</mods:mods>
|
||
<treatment ID-GBIF-Taxon="127853570" LSID="urn:lsid:plazi:treatment:A82125154158892C12BAC6FD998202AC" httpUri="http://treatment.plazi.org/id/A82125154158892C12BAC6FD998202AC" lastPageId="55" lastPageNumber="56" pageId="53" pageNumber="54">
|
||
<subSubSection pageId="53" pageNumber="54" type="multiple">
|
||
<paragraph pageId="53" pageNumber="54">
|
||
<pageBreakToken pageId="53" pageNumber="54" start="start">Taxon</pageBreakToken>
|
||
classification Animalia Coleoptera Carabidae
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="53" pageNumber="54" type="nomenclature">
|
||
<paragraph pageId="53" pageNumber="54">
|
||
(018)
|
||
<taxonomicName authority="Sharp" class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax cognatus" order="Coleoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="cognatus">Mecyclothorax cognatus Sharp</taxonomicName>
|
||
Figs 28C, 29
|
||
<normalizedToken originalValue="E–H">E-H</normalizedToken>
|
||
, 30B, 31B, 32
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="53" pageNumber="54" type="reference_group">
|
||
<paragraph pageId="53" pageNumber="54">
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax cognatus" order="Coleoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="cognatus">Mecyclothorax cognatus</taxonomicName>
|
||
<bibRefCitation author="Sharp, D" editor="Sharp, D" journalOrPublisher="The University Press, Cambridge" pageId="310" pageNumber="311" pagination="175 - 292" title="Coleoptera II. Caraboidea." volumeTitle="Fauna Hawaiiensis" year="1903">Sharp 1903</bibRefCitation>
|
||
: 255;
|
||
<bibRefCitation author="Britton, EB" journalOrPublisher="Bishop Museum" pageId="307" pageNumber="308" pagination="107 - 166" title="A revision of the Hawaiian species of Mecyclothorax (Coleoptera: Carabidae). Occasional Papers of the Bernice P." volume="19" year="1948 b">Britton 1948b</bibRefCitation>
|
||
: 165.
|
||
</paragraph>
|
||
<paragraph pageId="53" pageNumber="54">
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Atelothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Atelothorax optatus" order="Coleoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="optatus">Atelothorax optatus</taxonomicName>
|
||
<bibRefCitation author="Sharp, D" editor="Sharp, D" journalOrPublisher="The University Press, Cambridge" pageId="310" pageNumber="311" pagination="175 - 292" title="Coleoptera II. Caraboidea." volumeTitle="Fauna Hawaiiensis" year="1903">Sharp 1903</bibRefCitation>
|
||
: 269;
|
||
<bibRefCitation author="Britton, EB" journalOrPublisher="Bishop Museum" pageId="307" pageNumber="308" pagination="107 - 166" title="A revision of the Hawaiian species of Mecyclothorax (Coleoptera: Carabidae). Occasional Papers of the Bernice P." volume="19" year="1948 b">Britton 1948b</bibRefCitation>
|
||
: 165 (synonymy).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="54" lastPageNumber="55" pageId="53" pageNumber="54" type="diagnosis">
|
||
<paragraph pageId="53" pageNumber="54">Diagnosis.</paragraph>
|
||
<paragraph lastPageId="54" lastPageNumber="55" pageId="53" pageNumber="54">
|
||
Among the mid-sized species in this group-standardized body length 4.7-5.7 mm-this species exhibits the broadest body in both pronotal and elytral dimensions. The pronotum is transverse, MPW/PL = 1.26-1.33, and basally broad, MPW/BPW = 1.31-1.42. This species is broadly sympatric in the Waikamoi area with
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax consanguineus" order="Coleoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="consanguineus">Mecyclothorax consanguineus</taxonomicName>
|
||
, the species most similar in appearance and thus likely to cause confusion.
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax cognatus" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="cognatus">
|
||
<pageBreakToken pageId="54" pageNumber="55" start="start">Mecyclothorax</pageBreakToken>
|
||
cognatus
|
||
</taxonomicName>
|
||
can be distinguished by the broad elytra (Fig. 28C), with broad humeri, MEW/HuW = 1.83-1.94 versus MEW/HuW = 1.91-2.04 for
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax consanguineus" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="consanguineus">Mecyclothorax consanguineus</taxonomicName>
|
||
(Fig. 33B). The pronotum of
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax cognatus" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="cognatus">Mecyclothorax cognatus</taxonomicName>
|
||
also exhibits more broadly explanate lateral margins just before the hind angles, that area broadly extended from the deep, smooth laterobasal depressions.
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax consanguineus" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="consanguineus">Mecyclothorax consanguineus</taxonomicName>
|
||
conversely exhibits less explanate lateral margins at that position. If a male is available, the aedeagal tip is absolutely diagnostic even if extended only slightly from the specimen. Those of
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax cognatus" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="cognatus">Mecyclothorax cognatus</taxonomicName>
|
||
males have an angulate apex with subangulate tip (Fig. 29
|
||
<normalizedToken originalValue="E–G">E-G</normalizedToken>
|
||
), whereas those of
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax consanguineus" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="consanguineus">Mecyclothorax consanguineus</taxonomicName>
|
||
males have an elongate apex with a mucronate tip (Fig. 34
|
||
<normalizedToken originalValue="A–I">A-I</normalizedToken>
|
||
).
|
||
<bibRefCitation author="Sharp, D" editor="Sharp, D" journalOrPublisher="The University Press, Cambridge" pageId="310" pageNumber="311" pagination="175 - 292" title="Coleoptera II. Caraboidea." volumeTitle="Fauna Hawaiiensis" year="1903">Sharp (1903)</bibRefCitation>
|
||
described
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Atelothrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Atelothrus optatus" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="optatus">Atelothrus optatus</taxonomicName>
|
||
based his interpretation that the lateral pronotal setae were missing in the unique type specimen. The left lateral seta is indeed absent, but the articulatory socket for the right lateral seta is evident, resulting in the species setal formula being scored here as 2 2(1) 2 2.
|
||
</paragraph>
|
||
<caption pageId="54" pageNumber="55">
|
||
<paragraph pageId="54" pageNumber="55">
|
||
Figure 33.
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax robustus" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="robustus">Mecyclothorax robustus</taxonomicName>
|
||
group species, dorsal habitus view. A
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax anchisteus" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="anchisteus">Mecyclothorax anchisteus</taxonomicName>
|
||
(Kīpahulu, 1960 m) B
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax consanguineus" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="consanguineus">Mecyclothorax consanguineus</taxonomicName>
|
||
(Ukulele Camp Pipeline, 1510 m) C
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax aeneus" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="aeneus">Mecyclothorax aeneus</taxonomicName>
|
||
(Honomanu, 1820-1850 m) D
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax antaeus" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="antaeus">Mecyclothorax antaeus</taxonomicName>
|
||
(ESE Kuiki, 2145 m).
|
||
</paragraph>
|
||
</caption>
|
||
<caption pageId="54" pageNumber="55">
|
||
<paragraph pageId="54" pageNumber="55">
|
||
Figure 34. Male aedeagus,
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax robustus" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="robustus">Mecyclothorax robustus</taxonomicName>
|
||
group species (for abbreviations see Table 2, p. 23).
|
||
<normalizedToken originalValue="A–I">A-I</normalizedToken>
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax consanguineus" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="consanguineus">Mecyclothorax consanguineus</taxonomicName>
|
||
. A Right view, sac everted (Ukulele Camp Pipeline, 1534 m)
|
||
<normalizedToken originalValue="B–C">B-C</normalizedToken>
|
||
Right and ventral views (Ukulele Camp Pipeline, 1534 m) D Right view (Honomanu, 1750 m). E Right view (ESE Kuiki, 2164 m) F Right view (ESE Kuiki, 2135 m) G Right view (Paliku, 1900 m)
|
||
<normalizedToken originalValue="H–I">H-I</normalizedToken>
|
||
Right view (Kīpahulu, 1960 m)
|
||
<normalizedToken originalValue="J–L">J-L</normalizedToken>
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax antaeus" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="antaeus">Mecyclothorax antaeus</taxonomicName>
|
||
J Right view, sac everted (ESE Kuiki, 2145 m)
|
||
<normalizedToken originalValue="K–L">K-L</normalizedToken>
|
||
Right and left views (ESE Kuiki, 2105 m) M
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax cymindicus" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="cymindicus">Mecyclothorax cymindicus</taxonomicName>
|
||
, right view (near Ukulele Camp, 1210-1365 m) N
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax cymindulus" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="cymindulus">Mecyclothorax cymindulus</taxonomicName>
|
||
, right view, sac everted (Kīpahulu, 1960 m).
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection pageId="54" pageNumber="55" type="identification">
|
||
<paragraph pageId="54" pageNumber="55">Identification</paragraph>
|
||
<paragraph pageId="54" pageNumber="55">
|
||
(n = 5). The eyes are moderately developed-ocular ratio = 1.41-1.46, ocular lobe ratio = 0.75-0.81-with the ocular lobe smoothly joined to the gena. The pronotal front angles are broadly protruded, subangulate externally, with the anteriorly broad pronotal lateral marginal depression narrowed to the position of the lateral seta, and then broadened toward the back of the pronotum. The elytral striae are present across the width of the elytra, depth of striae 6 and 7 subequal to slightly shallower than striae 1-5 and 8. The discal elytral intervals are only slightly convex, with very fine punctures in the associated striae. The metepisternum bears ~16 punctures in 2-3 rows, about twice as many punctures as seen in
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax consanguineus" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="consanguineus">Mecyclothorax consanguineus</taxonomicName>
|
||
. Cuticular microsculpture is essentially identical to that observed in
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax consanguineus" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="consanguineus">Mecyclothorax consanguineus</taxonomicName>
|
||
: 1, vertex with transverse mesh, sculpticell breadth 2
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
length; 2, pronotal disc with transverse mesh, sculpticell breadth 2
|
||
<normalizedToken originalValue="–3×">-3x</normalizedToken>
|
||
length; 3, pronotal base with distinct isodiametric and transverse sculpticells; 4, elytral disc with transverse mesh, sculpticell breadth 3
|
||
<normalizedToken originalValue="–4×">-4x</normalizedToken>
|
||
length; and 5, elytral apex with transverse mesh, sculpticell breadth 3
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
length, to transverse lines.
|
||
</paragraph>
|
||
<paragraph pageId="54" pageNumber="55">
|
||
Male genitalia (n = 6). Aedeagal median lobe gracile with broad, trapezoidal apex, distance from parameral articulation to tip 4.1
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
median breadth (Fig. 29G), apex with flat ventral and apical faces, the tip angulate; median lobe nearly straight in ventral view, the right margin concave, and left margin incurved before the apparently rounded tip (Fig. 29H); internal sac broad, parallel sided, with moderate dorsal microtrichial patch and smaller ventral microtrichical patch that is near base of sac (Fig. 29
|
||
<normalizedToken originalValue="E–F">E-F</normalizedToken>
|
||
); flagellar plate well sclerotized, length 0.42
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
distance from parameral articulation to tip.
|
||
</paragraph>
|
||
<paragraph pageId="54" pageNumber="55">
|
||
Female reproductive tract (n = 1). Bursa copulatrix columnar with rounded apex, length 1.32 mm, breadth 0.46 mm, base as broad as vagina (Fig. 30B); bursal walls translucent with thick wrinkles; gonocoxite 1 with 4 apical fringe setae, 6-7 smaller setae on medial surface (Fig. 31B); gonocoxite 2 narrowly subtriangular with subacuminate apex, base narrowly extended laterally, 2 lateral ensiform setae with apical seta broader, apical nematiform setae on medioventral surface at 0.73
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
gonocoxite length.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="55" lastPageNumber="56" pageId="54" pageNumber="55" type="types">
|
||
<paragraph pageId="54" pageNumber="55">Types.</paragraph>
|
||
<paragraph lastPageId="55" lastPageNumber="56" pageId="54" pageNumber="55">
|
||
For
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax cognatus" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="cognatus">Mecyclothorax cognatus</taxonomicName>
|
||
Sharp holotype female (BMNH) labeled:
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax cognatus" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="cognatus">Mecyclothorax cognatus</taxonomicName>
|
||
Type D.S. Haleakala Perkins 111 // Type // Hawaiian Is. Perkins 1904-336. // Haleakala Maui 5000 ft. Perkins IV 1894 //
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Atelothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Atelothorax optatus" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="optatus">Atelothorax optatus</taxonomicName>
|
||
Sharp compared with type E.B.B. // HOLOTYPE
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax cognatus" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="cognatus">Mecyclothorax cognatus</taxonomicName>
|
||
Sharp J.K. Lieb
|
||
<pageBreakToken pageId="55" pageNumber="56" start="start">herr</pageBreakToken>
|
||
1998 (black-margined red label). For
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Atelothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Atelothorax optatus" order="Coleoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="optatus">Atelothorax optatus</taxonomicName>
|
||
Sharp holotype male (BPBM) platen mounted and labeled:
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Atelothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Atelothorax" order="Coleoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="genus">Atelothorax</taxonomicName>
|
||
/
|
||
<taxonomicName lsidName="optatus" pageId="55" pageNumber="56" rank="species" species="optatus">optatus</taxonomicName>
|
||
/ Type / D.S. / Haleakala / 1902 (written on obverse of mounting card) //
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax" order="Coleoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="genus">Mecyclothorax</taxonomicName>
|
||
/
|
||
<taxonomicName lsidName="cognatus" pageId="55" pageNumber="56" rank="species" species="cognatus">cognatus</taxonomicName>
|
||
Sharp / Compared / with type E.B.B. // HSPA # / 1960 // HOLOTYPE /
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Atelothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Atelothorax" order="Coleoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="genus">Atelothorax</taxonomicName>
|
||
/
|
||
<taxonomicName lsidName="optatus" pageId="55" pageNumber="56" rank="species" species="optatus">optatus</taxonomicName>
|
||
/ Sharp / J.K. Liebherr 1998 (black-margined red label).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="55" pageNumber="56" type="distribution and habitat">
|
||
<paragraph pageId="55" pageNumber="56">Distribution and habitat.</paragraph>
|
||
<paragraph pageId="55" pageNumber="56">
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax cognatus" order="Coleoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="cognatus">Mecyclothorax cognatus</taxonomicName>
|
||
is restricted to forests in the Waikamoi region (Fig. 32) from 1200-1850 m elevation. The only recorded Perkins lot (No. 111) was collected
|
||
<normalizedToken originalValue="iv–">iv-</normalizedToken>
|
||
1894 near Ukulele Camp, with modern collections centered on the Ukulele Pipeline Koa-
|
||
<normalizedToken originalValue="‘Ōhi‘a">'Ōhi'a</normalizedToken>
|
||
Mesic Forest at ~1500 m elevation northeast of the Ukulele Camp site. Beetles occur on and under bark of
|
||
<taxonomicName lsidName="koa" pageId="55" pageNumber="56" rank="species" species="koa">koa</taxonomicName>
|
||
, in moss on trunks of
|
||
<normalizedToken originalValue="‘ōhi‘a">'ōhi'a</normalizedToken>
|
||
, and on the ground in the leaf litter or under logs. Several have been collected in yellow-pan traps set on the ground.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |