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<document id="41A2558648FE5C4CF84E82E709E4B68D" ID-CLB-Dataset="29524" ID-DOI="10.11646/zootaxa.4446.1.4" ID-GBIF-Dataset="d35cedde-4442-4a14-be2d-d5213a12ab29" ID-ISSN="1175-5326" ID-Zenodo-Dep="1437384" ID-ZooBank="E5ABBD4C-C643-4091-9AD4-D9D4028A218A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1538121156330" checkinUser="plazi" docAuthor="Ban, Teruaki" docDate="2018" docId="9B474B53FFE73D60FF01B6FEFDCB5391" docLanguage="en" docName="zootaxa.4446.1.4.pdf" docOrigin="Zootaxa 4446 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Sadoletus subpellucidus Ban 2018, sp. nov." docType="treatment" docVersion="8" lastPageNumber="66" masterDocId="677E332BFFEE3D6FFF96B264FFD05011" masterDocTitle="The lygaeoid bug family Heterogastridae (Hemiptera: Heteroptera: Lygaeoidea) from Laos and Thailand, with descriptions of three new species" masterLastPageNumber="67" masterPageNumber="51" pageNumber="60" updateTime="1698555679930" updateUser="ExternalLinkService">
<mods:mods id="5DB29902076CF7C1EA545BC61D6EF49D" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="AC727E493E957DFB4E749338D5C56922">
<mods:title id="1A68EFBAAD9E8806DD6FCC27A641963C">The lygaeoid bug family Heterogastridae (Hemiptera: Heteroptera: Lygaeoidea) from Laos and Thailand, with descriptions of three new species</mods:title>
</mods:titleInfo>
<mods:name id="97F91CD4BD5C30C1C6A8AFEF08776084" type="personal">
<mods:role id="9EE2EE6BAA95161F28B96B25863A38E6">
<mods:roleTerm id="B1171784486EE83F9A7D458C52141480">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="42FB7F6E380338E72B1FF9EFE0E45A35">Ban, Teruaki</mods:namePart>
</mods:name>
<mods:typeOfResource id="161AF405575759FFC6FD88F5F0638DB5">text</mods:typeOfResource>
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<mods:title id="1F75D3472DF07A45AB029E66181B8D6B">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="11F3C795FB153FFC83393C31C09A4853">
<mods:date id="A30A8521E571F101E14ADA4B2EA9026E">2018</mods:date>
<mods:detail id="1E9F22F0627564A290ABAD6B541A6EAC" type="pubDate">
<mods:number id="93556D21B4EFDB842B8272FF0E09B8A8">2018-07-16</mods:number>
</mods:detail>
<mods:detail id="013E4E29ED1875690C6625B2A3AF22FD" type="volume">
<mods:number id="D1C7E35C4846597A0F405139D50A7CB7">4446</mods:number>
</mods:detail>
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<mods:number id="57F83E04ED87D35DD23DE29AA1D0E616">1</mods:number>
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<mods:start id="87969C32B747D7E957EC71AA3EC5AE3F">51</mods:start>
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<mods:classification id="94C72C0900767D91CCC6CB873669480A">journal article</mods:classification>
<mods:identifier id="F36DE5A7B594158B6C9D855A2E929256" type="CLB-Dataset">29524</mods:identifier>
<mods:identifier id="90028D49DED35102F43DE433AAE45052" type="DOI">10.11646/zootaxa.4446.1.4</mods:identifier>
<mods:identifier id="1A0BFD0C21863889A871FCA363F9ED60" type="GBIF-Dataset">d35cedde-4442-4a14-be2d-d5213a12ab29</mods:identifier>
<mods:identifier id="D45C313C50ED19992E766A848E81B0DD" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="5B6693F0F11A88B595B6370D1AA219E7" type="Zenodo-Dep">1437384</mods:identifier>
<mods:identifier id="DF43D90B34BAFCEE8AA77CBC19ED8E89" type="ZooBank">E5ABBD4C-C643-4091-9AD4-D9D4028A218A</mods:identifier>
</mods:mods>
<treatment id="9B474B53FFE73D60FF01B6FEFDCB5391" ID-DOI="http://doi.org/10.5281/zenodo.3503042" ID-GBIF-Taxon="148403382" ID-Zenodo-Dep="3503042" LSID="urn:lsid:plazi:treatment:9B474B53FFE73D60FF01B6FEFDCB5391" httpUri="http://treatment.plazi.org/id/9B474B53FFE73D60FF01B6FEFDCB5391" lastPageId="15" lastPageNumber="66" pageId="9" pageNumber="60">
<subSubSection id="5BF4A9CEFFE73D66FF01B6FEFE0654C7" pageId="9" pageNumber="60" type="nomenclature">
<paragraph id="1351FA45FFE73D66FF01B6FEFDCE54A4" blockId="9.[151,542,1178,1238]" box="[151,542,1178,1205]" pageId="9" pageNumber="60">
<heading id="48194D29FFE73D66FF01B6FEFDCE54A4" bold="true" box="[151,542,1178,1205]" fontSize="11" level="1" pageId="9" pageNumber="60" reason="1">
<emphasis id="219A2657FFE73D66FF01B6FEFDCE54A4" bold="true" box="[151,542,1178,1205]" pageId="9" pageNumber="60">
<taxonomicName id="D4EE81C6FFE73D66FF01B6FEFE6954A5" ID-CoL="8QP6D" authority="Ban, 2018" authorityName="Ban" authorityYear="2018" box="[151,441,1178,1204]" class="Insecta" family="Heterogastridae" genus="Sadoletus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="60" phylum="Arthropoda" rank="species" species="subpellucidus" status="sp. nov.">
<emphasis id="219A2657FFE73D66FF01B6FEFE6954A5" bold="true" box="[151,441,1178,1204]" italics="true" pageId="9" pageNumber="60">Sadoletus subpellucidus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3AA99B2CFFE73D66FE57B6FFFDCE54A4" box="[449,542,1179,1205]" pageId="9" pageNumber="60" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="1351FA45FFE73D66FF01B6D9FE0654C7" blockId="9.[151,542,1178,1238]" box="[151,470,1213,1238]" pageId="9" pageNumber="60">
(
<figureCitation id="8BD5E6C0FFE73D66FF09B6D9FEF754C7" box="[159,295,1213,1238]" captionStart="FIGURES 10–13" captionStartId="10.[151,264,1970,1993]" captionTargetBox="[256,1363,244,1888]" captionTargetId="figure@10.[216,1374,193,1949]" captionTargetPageId="10" captionText="FIGURES 10–13. Habitus of Heterogastridae species from Laos and Thailand. 10–11, S. ryukyuensis, male (10), female (11); 12–13, Sadoletus subpellucidus sp. nov., holotype, male (12), female (13). Scales: 1.0 mm." httpUri="https://zenodo.org/record/1437390/files/figure.png" pageId="9" pageNumber="60">Figs. 12–13</figureCitation>
,
<figureCitation id="8BD5E6C0FFE73D66FEA4B6D9FE8054C7" box="[306,336,1213,1238]" captionStart="FIGURES 14–16" captionStartId="11.[151,264,768,791]" captionTargetBox="[199,1371,188,745]" captionTargetId="figure@11.[181,1405,183,761]" captionTargetPageId="11" captionText="FIGURES 14–16. Females of each new Sadoletus species, ventral view. 14, Sadoletus laosensis sp. nov.; 15, S. melasmus sp. nov.; 16, S. subpellucidus sp. nov. Scales: 1.0 mm." httpUri="https://zenodo.org/record/1437392/files/figure.png" pageId="9" pageNumber="60">16</figureCitation>
,
<figureCitation id="8BD5E6C0FFE73D66FECBB6DAFEAB54C7" box="[349,379,1214,1238]" captionStart="FIGURES 17–19" captionStartId="11.[151,264,1221,1244]" captionTargetBox="[173,1370,892,1151]" captionTargetId="figure@11.[151,1436,861,1218]" captionTargetPageId="11" captionText="FIGURES 17–19. Abdominal sternite VI and ovipositer of each new Sadoletus species, ventral view. 17, Sadoletus laosensis sp. nov.; 18, S. melasmus sp. nov.; 19, S. subpellucidus sp. nov. Scales: 1.0 mm." httpUri="https://zenodo.org/record/1437394/files/figure.png" pageId="9" pageNumber="60">19</figureCitation>
,
<figureCitation id="8BD5E6C0FFE73D66FE10B6DAFE1E54C7" box="[390,462,1214,1238]" captionStart="FIGURES 30–34" captionStartId="14.[151,264,1933,1956]" captionTargetBox="[188,1367,238,1845]" captionTargetId="figure@14.[151,1436,238,1885]" captionTargetPageId="14" captionText="FIGURES 30–34. Sadoletus subpellucidus sp. nov. 30, Pygophore, dorsal view (setae omitted from left half); 31, left paramere, dorsal view; 32–33, phallus (endosoma not everted), ventral (32) and lateral (33) views; 34, spermatheca. Scales: 0.1 mm." httpUri="https://zenodo.org/record/1437402/files/figure.png" pageId="9" pageNumber="60">30–34</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="5BF4A9CEFFE73D66FF01B761FCE1559B" pageId="9" pageNumber="60" type="materials_examined">
<paragraph id="1351FA45FFE73D66FF01B761FCE1559B" blockId="9.[151,1437,1285,2030]" pageId="9" pageNumber="60">
<materialsCitation id="A386F018FFE73D66FF01B761FE2E5553" ID-GBIF-Occurrence="1913858567" collectingDate="2010-02-19" country="Laos" elevation="195" latitude="18.216667" location="Teruaki Ban" longLatPrecision="1276" longitude="102.46667" municipality="Wat Par Ba Phong Khong" pageId="9" pageNumber="60" specimenCount="1" specimenCount-male="1" stateProvince="Vientiane" typeStatus="holotype">
<emphasis id="219A2657FFE73D66FF01B761FE84550F" bold="true" box="[151,340,1285,1310]" pageId="9" pageNumber="60">
<typeStatus id="CC5544E7FFE73D66FF01B761FF02550F" box="[151,210,1285,1310]" pageId="9" pageNumber="60">Type</typeStatus>
Materials.
</emphasis>
<typeStatus id="CC5544E7FFE73D66FECAB761FE19550F" box="[348,457,1285,1310]" pageId="9" pageNumber="60" type="holotype">Holotype</typeStatus>
:
<specimenCount id="05E831CCFFE73D66FE42B762FE3B550C" box="[468,491,1286,1309]" pageId="9" pageNumber="60" type="male">♂</specimenCount>
,
<emphasis id="219A2657FFE73D66FE61B761FD9D550F" bold="true" box="[503,589,1285,1310]" pageId="9" pageNumber="60">
<collectingCountry id="6BF9BAD5FFE73D66FE61B761FD98550F" box="[503,584,1285,1310]" name="Laos" pageId="9" pageNumber="60">LAOS</collectingCountry>
:
</emphasis>
<geoCoordinate id="76DA9C82FFE73D66FDC3B762FD63550F" box="[597,691,1286,1310]" direction="north" orientation="latitude" pageId="9" pageNumber="60" precision="925" value="18.216667">18°13'N</geoCoordinate>
,
<geoCoordinate id="76DA9C82FFE73D66FD56B762FCFA550F" box="[704,810,1286,1310]" direction="east" orientation="longitude" pageId="9" pageNumber="60" precision="925" value="102.46667">102°28'E</geoCoordinate>
, alt.
<elevation id="98C31D76FFE73D66FCF4B761FC7C550C" box="[866,940,1285,1310]" metricMagnitude="2" metricUnit="m" metricValue="1.95" pageId="9" pageNumber="60" unit="m" value="195.0">
<quantity id="D41657A0FFE73D66FCF4B761FC7C550C" box="[866,940,1285,1310]" metricMagnitude="2" metricUnit="m" metricValue="1.95" pageId="9" pageNumber="60" unit="m" value="195.0">195 m</quantity>
</elevation>
,
<collectingMunicipality id="F335603FFFE73D66FC2FB762FB30550F" box="[953,1248,1285,1310]" pageId="9" pageNumber="60">Wat Par Ba Phong Khong</collectingMunicipality>
,
<collectingRegion id="D12A34A7FFE73D66FB7AB761FA87550F" box="[1260,1367,1285,1310]" country="Laos" name="Viangchan" pageId="9" pageNumber="60">Vientiane</collectingRegion>
Dist.,
<collectingDate id="7714256DFFE73D66FF01B74EFEDE5553" box="[151,270,1322,1346]" pageId="9" pageNumber="60" value="2010-02-19">
<date id="6750DC85FFE73D66FF01B74EFEDE5553" box="[151,270,1322,1346]" pageId="9" pageNumber="60" value="2010-02-19">19.II.2010</date>
</collectingDate>
,
<location id="1631AC9EFFE73D66FE8DB74DFE765550" LSID="urn:lsid:plazi:treatment:9B474B53FFE73D60FF01B6FEFDCB5391:1631AC9EFFE73D66FE8DB74DFE765550" box="[283,422,1321,1346]" country="Laos" latitude="18.216667" longLatPrecision="1276" longitude="102.46667" municipality="Wat Par Ba Phong Khong" name="Teruaki Ban" pageId="9" pageNumber="60" stateProvince="Vientiane">Teruaki Ban</location>
(TUA)
</materialsCitation>
.
<materialsCitation id="A386F018FFE73D66FD9DB74EFCE1559B" ID-GBIF-Occurrence="1913858571" collectingDate="2010-02-15" collectingDateMax="2010-02-19" collectingDateMin="2010-02-15" collectorName="Same &amp; Ban Nakha &amp; Mika Watanabe &amp; J. Kantoh" country="Laos" county="Vientiane Dist." elevation="124" latitude="15.25" location="Teruaki Ban" longLatPrecision="1285" longitude="106.73333" municipality="Champasak Prov." pageId="9" pageNumber="60" specimenCount="16" specimenCount-female="7" specimenCount-male="9" stateProvince="Sekong" typeStatus="paratype">
<typeStatus id="CC5544E7FFE73D66FD9DB74EFDAD5553" box="[523,637,1322,1346]" pageId="9" pageNumber="60" type="paratype">Paratypes</typeStatus>
:
<emphasis id="219A2657FFE73D66FD1EB74DFD0D5553" bold="true" box="[648,733,1321,1346]" pageId="9" pageNumber="60">
<collectingCountry id="6BF9BAD5FFE73D66FD1EB74DFD085553" box="[648,728,1321,1346]" name="Laos" pageId="9" pageNumber="60">LAOS</collectingCountry>
:
</emphasis>
<specimenCount id="05E831CCFFE73D66FD70B74EFCC25550" box="[742,786,1322,1345]" pageId="9" pageNumber="60" type="male">7 ♂</specimenCount>
<specimenCount id="05E831CCFFE73D66FC8DB74DFC965550" box="[795,838,1321,1346]" pageId="9" pageNumber="60" type="female">5 ♀</specimenCount>
,
<collectorName id="BE1B9F93FFE73D66FCC0B74EFC445553" box="[854,916,1322,1346]" pageId="9" pageNumber="60">Same</collectorName>
date as
<typeStatus id="CC5544E7FFE73D66FC62B74DFB8A5553" box="[1012,1114,1321,1346]" pageId="9" pageNumber="60" type="holotype">holotype</typeStatus>
;
<specimenCount id="05E831CCFFE73D66FBF1B74EFB455550" box="[1127,1173,1322,1346]" pageId="9" pageNumber="60" type="male">1 ♂</specimenCount>
,
<collectorName id="BE1B9F93FFE73D66FB34B74EFAF45553" box="[1186,1316,1321,1346]" pageId="9" pageNumber="60">Ban Nakha</collectorName>
,
<collectingCounty id="FA3082C9FFE73D66FAA7B74DFF1F5577" pageId="9" pageNumber="60">Vientiane Dist.</collectingCounty>
,
<collectingDate id="7714256DFFE73D66FF4DB72AFE895577" box="[219,345,1358,1382]" pageId="9" pageNumber="60" value="2010-02-19">
<date id="6750DC85FFE73D66FF4DB72AFE895577" box="[219,345,1358,1382]" pageId="9" pageNumber="60" value="2010-02-19">19.II. 2010</date>
</collectingDate>
,
<collectorName id="BE1B9F93FFE73D66FEF3B729FDC55577" box="[357,533,1357,1382]" pageId="9" pageNumber="60">Mika Watanabe</collectorName>
&amp;
<collectorName id="BE1B9F93FFE73D66FDAFB72AFD745577" box="[569,676,1357,1382]" pageId="9" pageNumber="60">J. Kantoh</collectorName>
(TUA);
<specimenCount id="05E831CCFFE73D66FC91B72AFCE15574" box="[775,817,1358,1382]" pageId="9" pageNumber="60" type="male">1 ♂</specimenCount>
<specimenCount id="05E831CCFFE73D66FCAEB72AFCB25574" box="[824,866,1358,1382]" pageId="9" pageNumber="60" type="female">2 ♀</specimenCount>
,
<geoCoordinate id="76DA9C82FFE73D66FCE7B729FC1F5577" box="[881,975,1357,1382]" direction="north" orientation="latitude" pageId="9" pageNumber="60" precision="925" value="15.25">15°15'N</geoCoordinate>
,
<geoCoordinate id="76DA9C82FFE73D66FC4DB729FB955577" box="[987,1093,1357,1382]" direction="east" orientation="longitude" pageId="9" pageNumber="60" precision="925" value="106.73333">106°44'E</geoCoordinate>
, alt.
<elevation id="98C31D76FFE73D66FBEDB72AFB145574" box="[1147,1220,1358,1382]" metricMagnitude="2" metricUnit="m" metricValue="1.24" pageId="9" pageNumber="60" unit="m" value="124.0">
<quantity id="D41657A0FFE73D66FBEDB72AFB145574" box="[1147,1220,1358,1382]" metricMagnitude="2" metricUnit="m" metricValue="1.24" pageId="9" pageNumber="60" unit="m" value="124.0">124 m</quantity>
</elevation>
,
<quantity id="D41657A0FFE73D66FB46B729FAC05577" box="[1232,1296,1357,1382]" metricMagnitude="4" metricUnit="m" metricValue="1.3" pageId="9" pageNumber="60" unit="km" value="13.0">13km</quantity>
South From
<collectingRegion id="D12A34A7FFE73D66FF01B715FF3E559B" box="[151,238,1393,1418]" country="Laos" name="Xekong" pageId="9" pageNumber="60">Sekong</collectingRegion>
,
<collectingMunicipality id="F335603FFFE73D66FF6FB715FE105598" box="[249,448,1393,1418]" pageId="9" pageNumber="60">Champasak Prov.</collectingMunicipality>
,
<collectingDate id="7714256DFFE73D66FE5DB715FD92559B" box="[459,578,1393,1418]" pageId="9" pageNumber="60" value="2010-02-15">
<date id="6750DC85FFE73D66FE5DB715FD92559B" box="[459,578,1393,1418]" pageId="9" pageNumber="60" value="2010-02-15">15.II.2010</date>
</collectingDate>
,
<location id="1631AC9EFFE73D66FDDBB715FD065598" LSID="urn:lsid:plazi:treatment:9B474B53FFE73D60FF01B6FEFDCB5391:1631AC9EFFE73D66FDDBB715FD065598" box="[589,726,1393,1418]" country="Laos" county="Vientiane Dist." latitude="15.25" longLatPrecision="1285" longitude="106.73333" municipality="Champasak Prov." name="Teruaki Ban" pageId="9" pageNumber="60" stateProvince="Sekong">Teruaki Ban</location>
(TUA).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="5BF4A9CEFFE73D66FF51B7F1FF1356BB" pageId="9" pageNumber="60" type="diagnosis">
<paragraph id="1351FA45FFE73D66FF51B7F1FF1356BB" blockId="9.[151,1437,1285,2030]" pageId="9" pageNumber="60">
<emphasis id="219A2657FFE73D66FF51B7F1FE9155BF" bold="true" box="[199,321,1429,1454]" pageId="9" pageNumber="60">Diagnosis.</emphasis>
Recognized among congeners of
<taxonomicName id="D4EE81C6FFE73D66FD5BB7F1FCEA55BC" authorityName="Distant" authorityYear="1903" box="[717,826,1429,1453]" class="Insecta" family="Heterogastridae" genus="Sadoletus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="60" phylum="Arthropoda" rank="genus">
<emphasis id="219A2657FFE73D66FD5BB7F1FCEA55BC" box="[717,826,1429,1453]" italics="true" pageId="9" pageNumber="60">Sadoletus</emphasis>
</taxonomicName>
by a combination of the following characters: head, anterior lobe of pronotum and basal half of scutellum reddish brown (
<figureCitation id="8BD5E6C0FFE73D66FC22B7DDFBEE55C3" box="[948,1086,1465,1490]" captionStart="FIGURES 10–13" captionStartId="10.[151,264,1970,1993]" captionTargetBox="[256,1363,244,1888]" captionTargetId="figure@10.[216,1374,193,1949]" captionTargetPageId="10" captionText="FIGURES 10–13. Habitus of Heterogastridae species from Laos and Thailand. 10–11, S. ryukyuensis, male (10), female (11); 12–13, Sadoletus subpellucidus sp. nov., holotype, male (12), female (13). Scales: 1.0 mm." httpUri="https://zenodo.org/record/1437390/files/figure.png" pageId="9" pageNumber="60">Figs. 12, 13</figureCitation>
); antennal segment
<date id="6750DC85FFE73D66FAB5B7DEFA8755C3" box="[1315,1367,1466,1490]" pageId="9" pageNumber="60">IV 3</date>
times longer than segment I; labium attaining metacoxae; posterior lobe of pronotum and legs pale yellow (
<figureCitation id="8BD5E6C0FFE73D66FA90B7B9FA5D55E7" box="[1286,1421,1501,1526]" captionStart="FIGURES 10–13" captionStartId="10.[151,264,1970,1993]" captionTargetBox="[256,1363,244,1888]" captionTargetId="figure@10.[216,1374,193,1949]" captionTargetPageId="10" captionText="FIGURES 10–13. Habitus of Heterogastridae species from Laos and Thailand. 10–11, S. ryukyuensis, male (10), female (11); 12–13, Sadoletus subpellucidus sp. nov., holotype, male (12), female (13). Scales: 1.0 mm." httpUri="https://zenodo.org/record/1437390/files/figure.png" pageId="9" pageNumber="60">Figs. 12, 13</figureCitation>
); femora and tibiae with pale brown annulations; hemelytra attaining apex of abdomen; corium subhyaline (
<figureCitation id="8BD5E6C0FFE73D66FAACB465FF65562F" captionStart="FIGURES 10–13" captionStartId="10.[151,264,1970,1993]" captionTargetBox="[256,1363,244,1888]" captionTargetId="figure@10.[216,1374,193,1949]" captionTargetPageId="10" captionText="FIGURES 10–13. Habitus of Heterogastridae species from Laos and Thailand. 10–11, S. ryukyuensis, male (10), female (11); 12–13, Sadoletus subpellucidus sp. nov., holotype, male (12), female (13). Scales: 1.0 mm." httpUri="https://zenodo.org/record/1437390/files/figure.png" pageId="9" pageNumber="60">Figs. 12, 13</figureCitation>
), with pale brown shallow punctures; hemelytral membrane hyaline; posterior margin of sternite VI weakly convex at middle (
<figureCitation id="8BD5E6C0FFE73D66FEF0B42DFE695673" box="[358,441,1609,1634]" captionStart="FIGURES 14–16" captionStartId="11.[151,264,768,791]" captionTargetBox="[199,1371,188,745]" captionTargetId="figure@11.[181,1405,183,761]" captionTargetPageId="11" captionText="FIGURES 14–16. Females of each new Sadoletus species, ventral view. 14, Sadoletus laosensis sp. nov.; 15, S. melasmus sp. nov.; 16, S. subpellucidus sp. nov. Scales: 1.0 mm." httpUri="https://zenodo.org/record/1437392/files/figure.png" pageId="9" pageNumber="60">Fig. 16</figureCitation>
); dorsal sinus of posterior aperture of pygophore without collar (
<figureCitation id="8BD5E6C0FFE73D66FB1CB42DFB0E5673" box="[1162,1246,1609,1634]" captionStart="FIGURES 30–34" captionStartId="14.[151,264,1933,1956]" captionTargetBox="[188,1367,238,1845]" captionTargetId="figure@14.[151,1436,238,1885]" captionTargetPageId="14" captionText="FIGURES 30–34. Sadoletus subpellucidus sp. nov. 30, Pygophore, dorsal view (setae omitted from left half); 31, left paramere, dorsal view; 32–33, phallus (endosoma not everted), ventral (32) and lateral (33) views; 34, spermatheca. Scales: 0.1 mm." httpUri="https://zenodo.org/record/1437402/files/figure.png" pageId="9" pageNumber="60">Fig. 30</figureCitation>
); cuplike sclerite fused to posterior margin of pygophore and arising in the middle of it (
<figureCitation id="8BD5E6C0FFE73D66FC50B409FBCC5697" box="[966,1052,1645,1670]" captionStart="FIGURES 30–34" captionStartId="14.[151,264,1933,1956]" captionTargetBox="[188,1367,238,1845]" captionTargetId="figure@14.[151,1436,238,1885]" captionTargetPageId="14" captionText="FIGURES 30–34. Sadoletus subpellucidus sp. nov. 30, Pygophore, dorsal view (setae omitted from left half); 31, left paramere, dorsal view; 32–33, phallus (endosoma not everted), ventral (32) and lateral (33) views; 34, spermatheca. Scales: 0.1 mm." httpUri="https://zenodo.org/record/1437402/files/figure.png" pageId="9" pageNumber="60">Fig. 30</figureCitation>
); paramere laminate at apex (
<figureCitation id="8BD5E6C0FFE73D66FAF9B409FF6656BB" captionStart="FIGURES 30–34" captionStartId="14.[151,264,1933,1956]" captionTargetBox="[188,1367,238,1845]" captionTargetId="figure@14.[151,1436,238,1885]" captionTargetPageId="14" captionText="FIGURES 30–34. Sadoletus subpellucidus sp. nov. 30, Pygophore, dorsal view (setae omitted from left half); 31, left paramere, dorsal view; 32–33, phallus (endosoma not everted), ventral (32) and lateral (33) views; 34, spermatheca. Scales: 0.1 mm." httpUri="https://zenodo.org/record/1437402/files/figure.png" pageId="9" pageNumber="60">Fig. 31</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="5BF4A9CEFFE73D60FF51B4D1FA4351BC" lastPageId="15" lastPageNumber="66" pageId="9" pageNumber="60" type="description">
<paragraph id="1351FA45FFE73D66FF51B4D1FB8957FF" blockId="9.[151,1437,1285,2030]" pageId="9" pageNumber="60">
<emphasis id="219A2657FFE73D66FF51B4D1FE8956DF" bold="true" box="[199,345,1717,1742]" pageId="9" pageNumber="60">Description.</emphasis>
Male (
<typeStatus id="CC5544E7FFE73D66FE3EB4D1FDC156DF" box="[424,529,1717,1742]" pageId="9" pageNumber="60" type="holotype">holotype</typeStatus>
): Short decumbent setae on body silvery. Punctures on pronotum, scutellum, and corium brown. Head (
<figureCitation id="8BD5E6C0FFE73D66FE07B4BDFE3456E3" box="[401,484,1753,1778]" captionStart="FIGURES 10–13" captionStartId="10.[151,264,1970,1993]" captionTargetBox="[256,1363,244,1888]" captionTargetId="figure@10.[216,1374,193,1949]" captionTargetPageId="10" captionText="FIGURES 10–13. Habitus of Heterogastridae species from Laos and Thailand. 10–11, S. ryukyuensis, male (10), female (11); 12–13, Sadoletus subpellucidus sp. nov., holotype, male (12), female (13). Scales: 1.0 mm." httpUri="https://zenodo.org/record/1437390/files/figure.png" pageId="9" pageNumber="60">Fig. 12</figureCitation>
) reddish brown. Clypeus pale yellow. Antennae brownish yellow. Labium (
<figureCitation id="8BD5E6C0FFE73D66FAD7B4BDFA4456E3" box="[1345,1428,1753,1778]" captionStart="FIGURES 14–16" captionStartId="11.[151,264,768,791]" captionTargetBox="[199,1371,188,745]" captionTargetId="figure@11.[181,1405,183,761]" captionTargetPageId="11" captionText="FIGURES 14–16. Females of each new Sadoletus species, ventral view. 14, Sadoletus laosensis sp. nov.; 15, S. melasmus sp. nov.; 16, S. subpellucidus sp. nov. Scales: 1.0 mm." httpUri="https://zenodo.org/record/1437392/files/figure.png" pageId="9" pageNumber="60">Fig. 16</figureCitation>
) pale yellow, apex of segment IV dark brown. Pronotum (
<figureCitation id="8BD5E6C0FFE73D66FC82B499FCB55707" box="[788,869,1789,1814]" captionStart="FIGURES 10–13" captionStartId="10.[151,264,1970,1993]" captionTargetBox="[256,1363,244,1888]" captionTargetId="figure@10.[216,1374,193,1949]" captionTargetPageId="10" captionText="FIGURES 10–13. Habitus of Heterogastridae species from Laos and Thailand. 10–11, S. ryukyuensis, male (10), female (11); 12–13, Sadoletus subpellucidus sp. nov., holotype, male (12), female (13). Scales: 1.0 mm." httpUri="https://zenodo.org/record/1437390/files/figure.png" pageId="9" pageNumber="60">Fig. 12</figureCitation>
) reddish brown in anterior lobe and pale yellow in posterior lobe, with 2 dark brown markings arranged transversely near posterior margin. Venter of mesothorax and metathorax reddish brown; peritreme brownish yellow. Scutellum (
<figureCitation id="8BD5E6C0FFE73D66FC27B521FBD8574F" box="[945,1032,1861,1886]" captionStart="FIGURES 10–13" captionStartId="10.[151,264,1970,1993]" captionTargetBox="[256,1363,244,1888]" captionTargetId="figure@10.[216,1374,193,1949]" captionTargetPageId="10" captionText="FIGURES 10–13. Habitus of Heterogastridae species from Laos and Thailand. 10–11, S. ryukyuensis, male (10), female (11); 12–13, Sadoletus subpellucidus sp. nov., holotype, male (12), female (13). Scales: 1.0 mm." httpUri="https://zenodo.org/record/1437390/files/figure.png" pageId="9" pageNumber="60">Fig. 12</figureCitation>
) reddish brown in basal half and brownish in remaining part. Apical half of scutellum with pale longitudinal stripe along meson. Corium (
<figureCitation id="8BD5E6C0FFE73D66FAD7B50DFA445793" box="[1345,1428,1897,1922]" captionStart="FIGURES 10–13" captionStartId="10.[151,264,1970,1993]" captionTargetBox="[256,1363,244,1888]" captionTargetId="figure@10.[216,1374,193,1949]" captionTargetPageId="10" captionText="FIGURES 10–13. Habitus of Heterogastridae species from Laos and Thailand. 10–11, S. ryukyuensis, male (10), female (11); 12–13, Sadoletus subpellucidus sp. nov., holotype, male (12), female (13). Scales: 1.0 mm." httpUri="https://zenodo.org/record/1437390/files/figure.png" pageId="9" pageNumber="60">Fig. 12</figureCitation>
) and clavus subhyaline; hemelytral membrane hyaline. Coxae and trochanter pale yellow; femora and tibiae pale yellow with pale brown annulations; tarsi pale yellow, apex of tarsomere III dark brown. Abdomen brown; each laterotergite pale brown anteriorly and dark brown posteriorly; genital capsule brown.
</paragraph>
<caption id="4791AACDFFE43D65FF01B5D6FBE357F6" httpUri="https://zenodo.org/record/1437390/files/figure.png" pageId="10" pageNumber="61" startId="10.[151,264,1970,1993]" targetBox="[256,1363,244,1888]" targetPageId="10">
<paragraph id="1351FA45FFE43D65FF01B5D6FBE357F6" blockId="10.[151,1436,1970,2023]" pageId="10" pageNumber="61">
<emphasis id="219A2657FFE43D65FF01B5D6FE8457D9" bold="true" box="[151,340,1970,1993]" pageId="10" pageNumber="61">FIGURES 10–13.</emphasis>
Habitus of
<taxonomicName id="D4EE81C6FFE43D65FE5AB5D7FDBC57D8" authorityName="Stal" authorityYear="1872" box="[460,620,1971,1993]" class="Insecta" family="Heterogastridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="61" phylum="Arthropoda" rank="family">Heterogastridae</taxonomicName>
species from Laos and Thailand. 10–11,
<taxonomicName id="D4EE81C6FFE43D65FB84B5D0FB7357D8" authorityName="Ban &amp; Ishikawa" authorityYear="2013" box="[1042,1187,1971,1993]" class="Insecta" family="Heterogastridae" genus="Sadoletus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="61" phylum="Arthropoda" rank="species" species="ryukyuensis">
<emphasis id="219A2657FFE43D65FB84B5D0FB7357D8" box="[1042,1187,1971,1993]" italics="true" pageId="10" pageNumber="61">S. ryukyuensis</emphasis>
</taxonomicName>
, male (10), female (11); 12–13,
<taxonomicName id="D4EE81C6FFE43D65FF75B5B5FE0757F6" authority="Ban, 2018" authorityName="Ban" authorityYear="2018" box="[227,471,2001,2023]" class="Insecta" family="Heterogastridae" genus="Sadoletus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="61" phylum="Arthropoda" rank="species" species="subpellucidus" status="sp. nov.">
<emphasis id="219A2657FFE43D65FF75B5B5FE0757F6" box="[227,471,2001,2023]" italics="true" pageId="10" pageNumber="61">Sadoletus subpellucidus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3AA99B2CFFE43D65FE4BB5B6FDFD57F6" box="[477,557,2002,2023]" pageId="10" pageNumber="61" rank="species">
<emphasis id="219A2657FFE43D65FE4BB5B6FDFD57F6" bold="true" box="[477,557,2002,2023]" pageId="10" pageNumber="61">sp. nov.</emphasis>
</taxonomicNameLabel>
, holotype, male (12), female (13). Scales: 1.0 mm.
</paragraph>
</caption>
<caption id="4791AACDFFE53D64FF01B164FD4B5324" httpUri="https://zenodo.org/record/1437392/files/figure.png" pageId="11" pageNumber="62" startId="11.[151,264,768,791]" targetBox="[199,1371,188,745]" targetPageId="11">
<paragraph id="1351FA45FFE53D64FF01B164FD4B5324" blockId="11.[151,1436,768,821]" pageId="11" pageNumber="62">
<emphasis id="219A2657FFE53D64FF01B164FE845307" bold="true" box="[151,340,768,791]" pageId="11" pageNumber="62">FIGURES 14–16.</emphasis>
Females of each new
<taxonomicName id="D4EE81C6FFE53D64FDA1B165FD4A5306" authorityName="Distant" authorityYear="1903" box="[567,666,769,791]" class="Insecta" family="Heterogastridae" genus="Sadoletus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="62" phylum="Arthropoda" rank="genus">
<emphasis id="219A2657FFE53D64FDA1B165FD4A5306" box="[567,666,769,791]" italics="true" pageId="11" pageNumber="62">Sadoletus</emphasis>
</taxonomicName>
species, ventral view. 14,
<taxonomicName id="D4EE81C6FFE53D64FC31B165FBBD5306" authority="Ban, 2018" authorityName="Ban" authorityYear="2018" box="[935,1133,769,791]" class="Insecta" family="Heterogastridae" genus="Sadoletus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="62" phylum="Arthropoda" rank="species" species="laosensis" status="sp. nov.">
<emphasis id="219A2657FFE53D64FC31B165FBBD5306" box="[935,1133,769,791]" italics="true" pageId="11" pageNumber="62">Sadoletus laosensis</emphasis>
</taxonomicName>
<emphasis id="219A2657FFE53D64FBE2B165FB145307" bold="true" box="[1140,1220,769,790]" pageId="11" pageNumber="62">
<taxonomicNameLabel id="3AA99B2CFFE53D64FBE2B165FB145307" box="[1140,1220,769,790]" pageId="11" pageNumber="62" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; 15,
<taxonomicName id="D4EE81C6FFE53D64FB6FB166FAA75306" authority="Ban, 2018" authorityName="Ban" authorityYear="2018" box="[1273,1399,769,791]" class="Insecta" family="Heterogastridae" genus="Sadoletus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="62" phylum="Arthropoda" rank="species" species="melasmus" status="sp. nov.">
<emphasis id="219A2657FFE53D64FB6FB166FAA75306" box="[1273,1399,769,791]" italics="true" pageId="11" pageNumber="62">S. melasmus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3AA99B2CFFE53D64FAEBB165FF135325" pageId="11" pageNumber="62" rank="species">
<emphasis id="219A2657FFE53D64FAEBB165FF135325" bold="true" pageId="11" pageNumber="62">sp. nov.</emphasis>
</taxonomicNameLabel>
; 16,
<taxonomicName id="D4EE81C6FFE53D64FF60B144FE4B5324" authority="Ban, 2018" authorityName="Ban" authorityYear="2018" box="[246,411,799,821]" class="Insecta" family="Heterogastridae" genus="Sadoletus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="62" phylum="Arthropoda" rank="species" species="subpellucidus" status="sp. nov.">
<emphasis id="219A2657FFE53D64FF60B144FE4B5324" box="[246,411,799,821]" italics="true" pageId="11" pageNumber="62">S. subpellucidus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3AA99B2CFFE53D64FE37B17BFE225325" box="[417,498,799,820]" pageId="11" pageNumber="62" rank="species">
<emphasis id="219A2657FFE53D64FE37B17BFE225325" bold="true" box="[417,498,799,820]" pageId="11" pageNumber="62">sp. nov.</emphasis>
</taxonomicNameLabel>
Scales: 1.0 mm.
</paragraph>
</caption>
<caption id="4791AACDFFE53D64FF01B6A1FC1754EA" httpUri="https://zenodo.org/record/1437394/files/figure.png" pageId="11" pageNumber="62" startId="11.[151,264,1221,1244]" targetBox="[173,1370,892,1151]" targetPageId="11">
<paragraph id="1351FA45FFE53D64FF01B6A1FC1754EA" blockId="11.[151,1436,1221,1275]" pageId="11" pageNumber="62">
<emphasis id="219A2657FFE53D64FF01B6A1FE8454CA" bold="true" box="[151,340,1221,1244]" pageId="11" pageNumber="62">FIGURES 17–19.</emphasis>
Abdominal sternite VI and ovipositer of each new
<taxonomicName id="D4EE81C6FFE53D64FCF2B6A2FC1754CD" authorityName="Distant" authorityYear="1903" box="[868,967,1222,1244]" class="Insecta" family="Heterogastridae" genus="Sadoletus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="62" phylum="Arthropoda" rank="genus">
<emphasis id="219A2657FFE53D64FCF2B6A2FC1754CD" box="[868,967,1222,1244]" italics="true" pageId="11" pageNumber="62">Sadoletus</emphasis>
</taxonomicName>
species, ventral view. 17,
<taxonomicName id="D4EE81C6FFE53D64FB43B6A2FA4C54CD" authority="Ban, 2018" authorityName="Ban" authorityYear="2018" box="[1237,1436,1222,1244]" class="Insecta" family="Heterogastridae" genus="Sadoletus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="62" phylum="Arthropoda" rank="species" species="laosensis" status="sp. nov.">
<emphasis id="219A2657FFE53D64FB43B6A2FA4C54CD" box="[1237,1436,1222,1244]" italics="true" pageId="11" pageNumber="62">Sadoletus laosensis</emphasis>
</taxonomicName>
<emphasis id="219A2657FFE53D64FF01B681FF3754EB" bold="true" box="[151,231,1253,1274]" pageId="11" pageNumber="62">
<taxonomicNameLabel id="3AA99B2CFFE53D64FF01B681FF3754EB" box="[151,231,1253,1274]" pageId="11" pageNumber="62" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; 18,
<taxonomicName id="D4EE81C6FFE53D64FE8DB682FE4854EA" authority="Ban, 2018" authorityName="Ban" authorityYear="2018" box="[283,408,1253,1275]" class="Insecta" family="Heterogastridae" genus="Sadoletus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="62" phylum="Arthropoda" rank="species" species="melasmus" status="sp. nov.">
<emphasis id="219A2657FFE53D64FE8DB682FE4854EA" box="[283,408,1253,1275]" italics="true" pageId="11" pageNumber="62">S. melasmus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3AA99B2CFFE53D64FE08B681FE3F54EB" box="[414,495,1253,1274]" pageId="11" pageNumber="62" rank="species">
<emphasis id="219A2657FFE53D64FE08B681FE3F54EB" bold="true" box="[414,495,1253,1274]" pageId="11" pageNumber="62">sp. nov.</emphasis>
</taxonomicNameLabel>
; 19,
<taxonomicName id="D4EE81C6FFE53D64FDB5B682FD1754EA" authority="Ban, 2018" authorityName="Ban" authorityYear="2018" box="[547,711,1253,1275]" class="Insecta" family="Heterogastridae" genus="Sadoletus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="62" phylum="Arthropoda" rank="species" species="subpellucidus" status="sp. nov.">
<emphasis id="219A2657FFE53D64FDB5B682FD1754EA" box="[547,711,1253,1275]" italics="true" pageId="11" pageNumber="62">S. subpellucidus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3AA99B2CFFE53D64FD5BB681FCCE54EB" box="[717,798,1253,1274]" pageId="11" pageNumber="62" rank="species">
<emphasis id="219A2657FFE53D64FD5BB681FCCE54EB" bold="true" box="[717,798,1253,1274]" pageId="11" pageNumber="62">sp. nov.</emphasis>
</taxonomicNameLabel>
Scales: 1.0 mm.
</paragraph>
</caption>
<paragraph id="1351FA45FFE53D64FF51B746FBBD55DA" blockId="11.[151,1437,1314,2023]" pageId="11" pageNumber="62">
Body length (
<figureCitation id="8BD5E6C0FFE53D64FEC9B746FE61552A" box="[351,433,1314,1339]" captionStart="FIGURES 10–13" captionStartId="10.[151,264,1970,1993]" captionTargetBox="[256,1363,244,1888]" captionTargetId="figure@10.[216,1374,193,1949]" captionTargetPageId="10" captionText="FIGURES 10–13. Habitus of Heterogastridae species from Laos and Thailand. 10–11, S. ryukyuensis, male (10), female (11); 12–13, Sadoletus subpellucidus sp. nov., holotype, male (12), female (13). Scales: 1.0 mm." httpUri="https://zenodo.org/record/1437390/files/figure.png" pageId="11" pageNumber="62">Fig. 12</figureCitation>
) 3 times width across humeri, covered with pubescence. Head (
<figureCitation id="8BD5E6C0FFE53D64FBECB746FB1B552A" box="[1146,1227,1314,1339]" captionStart="FIGURES 10–13" captionStartId="10.[151,264,1970,1993]" captionTargetBox="[256,1363,244,1888]" captionTargetId="figure@10.[216,1374,193,1949]" captionTargetPageId="10" captionText="FIGURES 10–13. Habitus of Heterogastridae species from Laos and Thailand. 10–11, S. ryukyuensis, male (10), female (11); 12–13, Sadoletus subpellucidus sp. nov., holotype, male (12), female (13). Scales: 1.0 mm." httpUri="https://zenodo.org/record/1437390/files/figure.png" pageId="11" pageNumber="62">Fig. 12</figureCitation>
) 1.7 times as wide as its length, covered with both rough and fine punctures; tylus tumid upwards; clypeus weakly projecting anteriad; juga with decumbent setae. Eyes occupying nearly half area of head in lateral view, not reaching anterior margin of pronotum. Antennae covered with suberect setae; proportional lengths of segments I to IV 1.0: 2.6: 2.2: 3.0. Labium attaining metacoxae; proportional lengths of segments I to IV 1.3: 1.9: 1.3: 1.0.
</paragraph>
<paragraph id="1351FA45FFE53D64FF51B7B3FCB456D7" blockId="11.[151,1437,1314,2023]" pageId="11" pageNumber="62">
Pronotum (
<figureCitation id="8BD5E6C0FFE53D64FED3B7B2FE4655FE" box="[325,406,1494,1519]" captionStart="FIGURES 10–13" captionStartId="10.[151,264,1970,1993]" captionTargetBox="[256,1363,244,1888]" captionTargetId="figure@10.[216,1374,193,1949]" captionTargetPageId="10" captionText="FIGURES 10–13. Habitus of Heterogastridae species from Laos and Thailand. 10–11, S. ryukyuensis, male (10), female (11); 12–13, Sadoletus subpellucidus sp. nov., holotype, male (12), female (13). Scales: 1.0 mm." httpUri="https://zenodo.org/record/1437390/files/figure.png" pageId="11" pageNumber="62">Fig. 12</figureCitation>
) a little more than 1.4 times as wide as its length, nearly trapezoidal, lacking collar, densely covered with punctures, and with lateral margin of anterior lobe weakly serrate; posterior lobe 1.3 times as long as anterior lobe, slightly more convex than anterior lobe; posterior margin weakly curved to posterior. Venter of prothorax, mesepisterna and metepisterna covered with rough punctures. Scutellum (
<figureCitation id="8BD5E6C0FFE53D64FBFEB426FB6C564A" box="[1128,1212,1602,1627]" captionStart="FIGURES 10–13" captionStartId="10.[151,264,1970,1993]" captionTargetBox="[256,1363,244,1888]" captionTargetId="figure@10.[216,1374,193,1949]" captionTargetPageId="10" captionText="FIGURES 10–13. Habitus of Heterogastridae species from Laos and Thailand. 10–11, S. ryukyuensis, male (10), female (11); 12–13, Sadoletus subpellucidus sp. nov., holotype, male (12), female (13). Scales: 1.0 mm." httpUri="https://zenodo.org/record/1437390/files/figure.png" pageId="11" pageNumber="62">Fig. 12</figureCitation>
) triangular, weakly tumid transversely in basal half, entirely covered with punctures. Hemelytra (
<figureCitation id="8BD5E6C0FFE53D64FB92B402FB87566E" box="[1028,1111,1638,1663]" captionStart="FIGURES 10–13" captionStartId="10.[151,264,1970,1993]" captionTargetBox="[256,1363,244,1888]" captionTargetId="figure@10.[216,1374,193,1949]" captionTargetPageId="10" captionText="FIGURES 10–13. Habitus of Heterogastridae species from Laos and Thailand. 10–11, S. ryukyuensis, male (10), female (11); 12–13, Sadoletus subpellucidus sp. nov., holotype, male (12), female (13). Scales: 1.0 mm." httpUri="https://zenodo.org/record/1437390/files/figure.png" pageId="11" pageNumber="62">Fig. 12</figureCitation>
) attaining apex of abdomen; apical margin of corium as long as claval suture. Profemur moderately expanded, armed with a spine ventrally at about apical third; tibiae and tarsi covered with long erect setae.
</paragraph>
<paragraph id="1351FA45FFE53D64FF51B4B6FF3E571E" blockId="11.[151,1437,1314,2023]" pageId="11" pageNumber="62">Abdomen wider than hemelytral width; sternum covered with fine punctures; posterior margin of sternite VI sinuate.</paragraph>
<paragraph id="1351FA45FFE53D64FF51B57EFC7857F6" blockId="11.[151,1437,1314,2023]" pageId="11" pageNumber="62">
Genital structures of
<typeStatus id="CC5544E7FFE53D64FE2AB57FFDFC5722" box="[444,556,1819,1843]" pageId="11" pageNumber="62" type="paratype">paratypes</typeStatus>
: pygophore (
<figureCitation id="8BD5E6C0FFE53D64FD52B57EFCC95722" box="[708,793,1818,1843]" captionStart="FIGURES 30–34" captionStartId="14.[151,264,1933,1956]" captionTargetBox="[188,1367,238,1845]" captionTargetId="figure@14.[151,1436,238,1885]" captionTargetPageId="14" captionText="FIGURES 30–34. Sadoletus subpellucidus sp. nov. 30, Pygophore, dorsal view (setae omitted from left half); 31, left paramere, dorsal view; 32–33, phallus (endosoma not everted), ventral (32) and lateral (33) views; 34, spermatheca. Scales: 0.1 mm." httpUri="https://zenodo.org/record/1437402/files/figure.png" pageId="11" pageNumber="62">Fig. 30</figureCitation>
) nearly spherical in dorsal view, covered with suberect setae; maximum width of dorsal sinus of posterior aperture more than one-third as wide at middle as pygophore, without collar along anterior margin; cuplike sclerite fused to posterior margin of pygophore and arising in the middle of it; antero-lateral margin of cuplike sclerite acute. Paramere (
<figureCitation id="8BD5E6C0FFE53D64FC23B5E2FBD7578E" box="[949,1031,1926,1951]" captionStart="FIGURES 30–34" captionStartId="14.[151,264,1933,1956]" captionTargetBox="[188,1367,238,1845]" captionTargetId="figure@14.[151,1436,238,1885]" captionTargetPageId="14" captionText="FIGURES 30–34. Sadoletus subpellucidus sp. nov. 30, Pygophore, dorsal view (setae omitted from left half); 31, left paramere, dorsal view; 32–33, phallus (endosoma not everted), ventral (32) and lateral (33) views; 34, spermatheca. Scales: 0.1 mm." httpUri="https://zenodo.org/record/1437402/files/figure.png" pageId="11" pageNumber="62">Fig. 31</figureCitation>
) twisted at basal third, laminated at apex, covered with several erect setae, sensory processes absent. Phallosoma (
<figureCitation id="8BD5E6C0FFE53D64FB89B5CEFB7B57D2" box="[1055,1195,1962,1987]" captionStart="FIGURES 30–34" captionStartId="14.[151,264,1933,1956]" captionTargetBox="[188,1367,238,1845]" captionTargetId="figure@14.[151,1436,238,1885]" captionTargetPageId="14" captionText="FIGURES 30–34. Sadoletus subpellucidus sp. nov. 30, Pygophore, dorsal view (setae omitted from left half); 31, left paramere, dorsal view; 32–33, phallus (endosoma not everted), ventral (32) and lateral (33) views; 34, spermatheca. Scales: 0.1 mm." httpUri="https://zenodo.org/record/1437402/files/figure.png" pageId="11" pageNumber="62">Figs. 32, 33</figureCitation>
) twice as long as its width in ventral view, roundly concave and wrinkled at apical margin.
</paragraph>
<caption id="4791AACDFFE23D63FF01B520FABB576B" httpUri="https://zenodo.org/record/1437396/files/figure.png" pageId="12" pageNumber="63" startId="12.[151,264,1860,1883]" targetBox="[173,1319,245,1801]" targetPageId="12">
<paragraph id="1351FA45FFE23D63FF01B520FABB576B" blockId="12.[151,1436,1860,1914]" pageId="12" pageNumber="63">
<emphasis id="219A2657FFE23D63FF01B520FE86574B" bold="true" box="[151,342,1860,1883]" pageId="12" pageNumber="63">FIGURES 20–24.</emphasis>
<taxonomicName id="D4EE81C6FFE23D63FECBB521FDF5574A" authority="Ban, 2018" authorityName="Ban" authorityYear="2018" box="[349,549,1861,1883]" class="Insecta" family="Heterogastridae" genus="Sadoletus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="63" phylum="Arthropoda" rank="species" species="laosensis" status="sp. nov.">
<emphasis id="219A2657FFE23D63FECBB521FDF5574A" box="[349,549,1861,1883]" italics="true" pageId="12" pageNumber="63">Sadoletus laosensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3AA99B2CFFE23D63FDBBB521FDAF574B" box="[557,639,1861,1882]" pageId="12" pageNumber="63" rank="species">
<emphasis id="219A2657FFE23D63FDBBB521FDAF574B" bold="true" box="[557,639,1861,1882]" pageId="12" pageNumber="63">sp. nov.</emphasis>
</taxonomicNameLabel>
20, Pygophore, dorsal view (setae omitted from left half); 21, left paramere, dorsal view; 22–23, phallus (endosoma not everted), ventral (22) and lateral (23) views; 24, spermatheca. Scales: 0.1 mm.
</paragraph>
</caption>
<paragraph id="1351FA45FFE23D62FF51B5C5FDAA50E9" blockId="12.[151,1436,1953,2013]" lastBlockId="13.[151,1436,151,248]" lastPageId="13" lastPageNumber="64" pageId="12" pageNumber="63">
Female (
<typeStatus id="CC5544E7FFE23D63FEBBB5C5FE7057A8" box="[301,416,1953,1977]" pageId="12" pageNumber="63" type="paratype">paratypes</typeStatus>
): resembles male in general appearance. Body (
<figureCitation id="8BD5E6C0FFE23D63FC42B5C5FBF957A8" box="[980,1065,1953,1978]" captionStart="FIGURES 10–13" captionStartId="10.[151,264,1970,1993]" captionTargetBox="[256,1363,244,1888]" captionTargetId="figure@10.[216,1374,193,1949]" captionTargetPageId="10" captionText="FIGURES 10–13. Habitus of Heterogastridae species from Laos and Thailand. 10–11, S. ryukyuensis, male (10), female (11); 12–13, Sadoletus subpellucidus sp. nov., holotype, male (12), female (13). Scales: 1.0 mm." httpUri="https://zenodo.org/record/1437390/files/figure.png" pageId="12" pageNumber="63">Fig. 13</figureCitation>
) evidently wider; antennae and abdominal color paler than male; ovipositor (
<figureCitation id="8BD5E6C0FFE23D63FD01B5A0FD3957CC" box="[663,745,1988,2013]" captionStart="FIGURES 17–19" captionStartId="11.[151,264,1221,1244]" captionTargetBox="[173,1370,892,1151]" captionTargetId="figure@11.[151,1436,861,1218]" captionTargetPageId="11" captionText="FIGURES 17–19. Abdominal sternite VI and ovipositer of each new Sadoletus species, ventral view. 17, Sadoletus laosensis sp. nov.; 18, S. melasmus sp. nov.; 19, S. subpellucidus sp. nov. Scales: 1.0 mm." httpUri="https://zenodo.org/record/1437394/files/figure.png" pageId="12" pageNumber="63">Fig. 19</figureCitation>
) straight, reaching sternite VI, approximate oviposital length
<quantity id="D41657A0FFE33D62FF01B2F3FF2550BE" box="[151,245,151,176]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="13" pageNumber="64" unit="mm" value="0.5">0.5 mm</quantity>
; the two hemiventrites of sternite VII are short therefore posterior margins of sternites III–V almost straight, sternite VI weakly sinuate (
<figureCitation id="8BD5E6C0FFE33D62FDABB2D8FD4350C5" box="[573,659,188,213]" captionStart="FIGURES 17–19" captionStartId="11.[151,264,1221,1244]" captionTargetBox="[173,1370,892,1151]" captionTargetId="figure@11.[151,1436,861,1218]" captionTargetPageId="11" captionText="FIGURES 17–19. Abdominal sternite VI and ovipositer of each new Sadoletus species, ventral view. 17, Sadoletus laosensis sp. nov.; 18, S. melasmus sp. nov.; 19, S. subpellucidus sp. nov. Scales: 1.0 mm." httpUri="https://zenodo.org/record/1437394/files/figure.png" pageId="13" pageNumber="64">Fig. 19</figureCitation>
); spermatheca (
<figureCitation id="8BD5E6C0FFE33D62FCDCB2D8FC4E50C5" box="[842,926,188,213]" captionStart="FIGURES 30–34" captionStartId="14.[151,264,1933,1956]" captionTargetBox="[188,1367,238,1845]" captionTargetId="figure@14.[151,1436,238,1885]" captionTargetPageId="14" captionText="FIGURES 30–34. Sadoletus subpellucidus sp. nov. 30, Pygophore, dorsal view (setae omitted from left half); 31, left paramere, dorsal view; 32–33, phallus (endosoma not everted), ventral (32) and lateral (33) views; 34, spermatheca. Scales: 0.1 mm." httpUri="https://zenodo.org/record/1437402/files/figure.png" pageId="13" pageNumber="64">Fig. 34</figureCitation>
) rounded at apex, as is typical in
<taxonomicName id="D4EE81C6FFE33D62FAB9B2D8FA4C50C5" authorityName="Distant" authorityYear="1903" box="[1327,1436,188,212]" class="Insecta" family="Heterogastridae" genus="Sadoletus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="64" phylum="Arthropoda" rank="genus">
<emphasis id="219A2657FFE33D62FAB9B2D8FA4C50C5" box="[1327,1436,188,212]" italics="true" pageId="13" pageNumber="64">Sadoletus</emphasis>
</taxonomicName>
species (as in
<bibRefCitation id="777F87B4FFE33D62FEA5B2BBFE0850E9" box="[307,472,223,248]" pageId="13" pageNumber="64" refId="ref11559" refString="Scudder, G. G. E. (1962) New Heterogastrinae (Hemiptera) with a key to the genera of the world. Opuscula Entomologica, 27, 117 - 127." type="journal article">Scudder, 1962</bibRefCitation>
: 121,
<figureCitation id="8BD5E6C0FFE33D62FD8DB2BBFDBD50E9" box="[539,621,223,248]" captionStart="FIGURES 14–16" captionStartId="11.[151,264,768,791]" captionTargetBox="[199,1371,188,745]" captionTargetId="figure@11.[181,1405,183,761]" captionTargetPageId="11" captionText="FIGURES 14–16. Females of each new Sadoletus species, ventral view. 14, Sadoletus laosensis sp. nov.; 15, S. melasmus sp. nov.; 16, S. subpellucidus sp. nov. Scales: 1.0 mm." httpUri="https://zenodo.org/record/1437392/files/figure.png" pageId="13" pageNumber="64">Fig. 14</figureCitation>
).
</paragraph>
<caption id="4791AACDFFE33D62FF01B5CDFABB57CF" httpUri="https://zenodo.org/record/1437398/files/figure.png" pageId="13" pageNumber="64" startId="13.[151,264,1961,1984]" targetBox="[152,1431,355,1907]" targetPageId="13">
<paragraph id="1351FA45FFE33D62FF01B5CDFABB57CF" blockId="13.[151,1436,1961,2014]" pageId="13" pageNumber="64">
<emphasis id="219A2657FFE33D62FF01B5CDFE8557AF" bold="true" box="[151,341,1961,1984]" pageId="13" pageNumber="64">FIGURES 25–29.</emphasis>
<taxonomicName id="D4EE81C6FFE33D62FECAB5CDFDFA57AE" authority="Ban, 2018" authorityName="Ban" authorityYear="2018" box="[348,554,1961,1983]" class="Insecta" family="Heterogastridae" genus="Sadoletus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="64" phylum="Arthropoda" rank="species" species="melasmus" status="sp. nov.">
<emphasis id="219A2657FFE33D62FECAB5CDFDFA57AE" box="[348,554,1961,1983]" italics="true" pageId="13" pageNumber="64">Sadoletus melasmus</emphasis>
</taxonomicName>
<emphasis id="219A2657FFE33D62FDA4B5CEFD5357AE" bold="true" box="[562,643,1962,1983]" pageId="13" pageNumber="64">
<taxonomicNameLabel id="3AA99B2CFFE33D62FDA4B5CEFD5357AE" box="[562,643,1962,1983]" pageId="13" pageNumber="64" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
25, Pygophore, dorsal view (setae omitted from left half); 26, left paramere, dorsal view; 27–28, phallus (endosoma not everted), ventral (27) and lateral (28) views; 29, spermatheca. Scales: 0.1 mm.
</paragraph>
</caption>
<caption id="4791AACDFFE03D61FF01B5E9FF1657F3" httpUri="https://zenodo.org/record/1437402/files/figure.png" pageId="14" pageNumber="65" startId="14.[151,264,1933,1956]" targetBox="[188,1367,238,1845]" targetPageId="14">
<paragraph id="1351FA45FFE03D61FF01B5E9FF1657F3" blockId="14.[151,1436,1933,2018]" pageId="14" pageNumber="65">
<emphasis id="219A2657FFE03D61FF01B5E9FE8957B2" bold="true" box="[151,345,1933,1956]" pageId="14" pageNumber="65">FIGURES 30–34.</emphasis>
<taxonomicName id="D4EE81C6FFE03D61FEF3B5EAFD8E57B5" authority="Ban, 2018" authorityName="Ban" authorityYear="2018" box="[357,606,1934,1956]" class="Insecta" family="Heterogastridae" genus="Sadoletus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="65" phylum="Arthropoda" rank="species" species="subpellucidus" status="sp. nov.">
<emphasis id="219A2657FFE03D61FEF3B5EAFD8E57B5" box="[357,606,1934,1956]" italics="true" pageId="14" pageNumber="65">Sadoletus subpellucidus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3AA99B2CFFE03D61FDFDB5EBFD1157B5" box="[619,705,1935,1956]" pageId="14" pageNumber="65" rank="species">
<emphasis id="219A2657FFE03D61FDFDB5EBFD1157B5" bold="true" box="[619,705,1935,1956]" pageId="14" pageNumber="65">sp. nov.</emphasis>
</taxonomicNameLabel>
30, Pygophore, dorsal view (setae omitted from left half); 31, left paramere, dorsal view; 32–33, phallus (endosoma not everted), ventral (32) and lateral (33) views; 34, spermatheca. Scales: 0.1 mm.
</paragraph>
</caption>
<paragraph id="1351FA45FFE13D60FF51B2F3FA4351BC" blockId="15.[151,1437,151,896]" pageId="15" pageNumber="66">
<emphasis id="219A2657FFE13D60FF51B2F3FEA650A1" bold="true" box="[199,374,151,176]" pageId="15" pageNumber="66">Measurements</emphasis>
[
<specimenCount id="05E831CCFFE13D60FE1EB2FCFE4F50BE" box="[392,415,152,175]" pageId="15" pageNumber="66" type="male">♂</specimenCount>
(n=10) /
<specimenCount id="05E831CCFFE13D60FD84B2FCFDF750BE" box="[530,551,152,175]" pageId="15" pageNumber="66" type="female">♀</specimenCount>
(n=7), value for
<typeStatus id="CC5544E7FFE13D60FD62B2F3FC8650A1" box="[756,854,151,176]" pageId="15" pageNumber="66" type="holotype">holotype</typeStatus>
male in parentheses]. Body length 4.2–4.5 (4.4) / 4.75–5.2; body height between coxae and dorsum 0.95–1.15 (1.15) / 1.18–1.3; head length 0.65–0.75 (0.75) / 0.75– 0.9, head width across eyes 1.15–1.25 (1.25) / 1.2–1.35; length of antennal segments I–IV respectively 0.25–0.31 (0.25) / 0.25–0.35, 0.63–0.7 (0.65) / 0.63–0.73, 0.5–0.6 (0.55) / 0.53–0.63, 0.7–0.75 (0.75) / 0.7–0.78; length of rostral segments I–IV respectively 0.5–0.63 (0.5) / 0.55–0.7, 0.65–0.7 (0.7) / 0.69–0.75, 0.45–0.58 (0.5) / 0.45– 0.55, 0.35–0.4 (0.38) / 0.35–0.43; mesal pronotal length 0.9–1.1 (1.05) / 1.05–1.25; posterior pronotal width 1.35– 1.5 (1.5) / 1.25–1.75; mesal scutellal length 0.75–0.85 (0.8) / 0.8–1.0; basal scutellal width 0.75–0.88 (0.85) / 0.88– 1.0; hemelytral length 2.75–2.95 (2.9) / 2.9 5–3.4; maximum width across hemelytra 1.38–1.45 (1.45) / 1.55–1.75.
</paragraph>
</subSubSection>
<subSubSection id="5BF4A9CEFFE13D60FF51B3D3FCE851C1" box="[199,824,439,464]" pageId="15" pageNumber="66" type="distribution">
<paragraph id="1351FA45FFE13D60FF51B3D3FCE851C1" blockId="15.[151,1437,151,896]" box="[199,824,439,464]" pageId="15" pageNumber="66">
<emphasis id="219A2657FFE13D60FF51B3D3FEB051C1" bold="true" box="[199,352,439,464]" pageId="15" pageNumber="66">Distribution.</emphasis>
<collectingCountry id="6BF9BAD5FFE13D60FEF1B3DCFE4D51DE" box="[359,413,440,463]" name="Laos" pageId="15" pageNumber="66">Laos</collectingCountry>
(
<collectingRegion id="D12A34A7FFE13D60FE3AB3D3FDC951C1" box="[428,537,439,464]" country="Laos" name="Viangchan" pageId="15" pageNumber="66">Vientiane</collectingRegion>
Dist.,
<collectingRegion id="D12A34A7FFE13D60FDF2B3D3FD3751C1" box="[612,743,439,464]" country="Laos" name="Champasak" pageId="15" pageNumber="66">Champasak</collectingRegion>
Prov.).
</paragraph>
</subSubSection>
<subSubSection id="5BF4A9CEFFE13D60FF51B3B8FD9A5209" pageId="15" pageNumber="66" type="etymology">
<paragraph id="1351FA45FFE13D60FF51B3B8FD9A5209" blockId="15.[151,1437,151,896]" pageId="15" pageNumber="66">
<emphasis id="219A2657FFE13D60FF51B3B8FE9E51E4" bold="true" box="[199,334,476,501]" pageId="15" pageNumber="66">Etymology.</emphasis>
The species epithet is the Latin adjective “
<taxonomicName id="D4EE81C6FFE13D60FCD4B3B8FC0C51E5" authorityName="Ban" authorityYear="2018" box="[834,988,476,500]" class="Insecta" family="Heterogastridae" genus="Sadoletus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="66" phylum="Arthropoda" rank="species" species="subpellucidus">
<emphasis id="219A2657FFE13D60FCD4B3B8FC0C51E5" box="[834,988,476,500]" italics="true" pageId="15" pageNumber="66">subpellucidus</emphasis>
</taxonomicName>
” (meaning subtransparent) referring to characteristic coloration of the corium.
</paragraph>
</subSubSection>
<subSubSection id="5BF4A9CEFFE13D60FF51B040FDCB5391" pageId="15" pageNumber="66" type="discussion">
<paragraph id="1351FA45FFE13D60FF51B040FB4A5329" blockId="15.[151,1437,151,896]" pageId="15" pageNumber="66">
<emphasis id="219A2657FFE13D60FF51B040FEEB522C" bold="true" box="[199,315,548,573]" pageId="15" pageNumber="66">Remarks.</emphasis>
This new species is most similar in general appearance to
<taxonomicName id="D4EE81C6FFE13D60FC59B040FB6B522C" authority="Distant" authorityName="Distant" box="[975,1211,548,573]" class="Insecta" family="Heterogastridae" genus="Sadoletus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="66" phylum="Arthropoda" rank="species" species="pallescens">
<emphasis id="219A2657FFE13D60FC59B040FBB0522D" box="[975,1120,548,572]" italics="true" pageId="15" pageNumber="66">S. pallescens</emphasis>
Distant
</taxonomicName>
,
<taxonomicName id="D4EE81C6FFE13D60FB50B040FA4C522C" authority="Bergroth" authorityName="Bergroth" box="[1222,1436,548,573]" class="Insecta" family="Heterogastridae" genus="Sadoletus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="66" phylum="Arthropoda" rank="species" species="valdezi">
<emphasis id="219A2657FFE13D60FB50B040FAE1522D" box="[1222,1329,548,572]" italics="true" pageId="15" pageNumber="66">S. valdezi</emphasis>
Bergroth
</taxonomicName>
and
<taxonomicName id="D4EE81C6FFE13D60FF50B02CFE5F5271" authority="Distant" authorityName="Distant" box="[198,399,583,608]" class="Insecta" family="Heterogastridae" genus="Sadoletus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="66" phylum="Arthropoda" rank="species" species="validus">
<emphasis id="219A2657FFE13D60FF50B02CFEE45271" box="[198,308,584,608]" italics="true" pageId="15" pageNumber="66">S. validus</emphasis>
Distant
</taxonomicName>
, but it can be distinguished from them by a combination of the following characters: labium attaining to metacoxae (in
<taxonomicName id="D4EE81C6FFE13D60FE5EB008FDE55295" box="[456,565,620,644]" class="Insecta" family="Heterogastridae" genus="Sadoletus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="66" phylum="Arthropoda" rank="species" species="valdezi">
<emphasis id="219A2657FFE13D60FE5EB008FDE55295" box="[456,565,620,644]" italics="true" pageId="15" pageNumber="66">S. valdezi</emphasis>
</taxonomicName>
labium attaining to mesocoxae); hemelytra not surpassing apex of abdomen (vs. hemelytra surpassing apex of abdomen); corium subhyaline (vs. corium white); femora with pale brown annulations (vs. femora without annulation); posterior margin of sternite VI of female convex at middle (vs. concave at middle); dorsal sinus of posterior aperture of pygophore without collar (in
<taxonomicName id="D4EE81C6FFE13D60FBE0B0BCFB3352E1" box="[1142,1251,728,752]" class="Insecta" family="Heterogastridae" genus="Sadoletus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="66" phylum="Arthropoda" rank="species" species="valdezi">
<emphasis id="219A2657FFE13D60FBE0B0BCFB3352E1" box="[1142,1251,728,752]" italics="true" pageId="15" pageNumber="66">S. valdezi</emphasis>
</taxonomicName>
, dorsal sinus of posterior aperture of pygophore with collar); apex of paramere laminate (in
<taxonomicName id="D4EE81C6FFE13D60FC6FB098FBB55305" box="[1017,1125,764,788]" class="Insecta" family="Heterogastridae" genus="Sadoletus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="66" phylum="Arthropoda" rank="species" species="valdezi">
<emphasis id="219A2657FFE13D60FC6FB098FBB55305" box="[1017,1125,764,788]" italics="true" pageId="15" pageNumber="66">S. valdezi</emphasis>
</taxonomicName>
apex of paramere peaked); sensory processes of paramere absent (in
<taxonomicName id="D4EE81C6FFE13D60FDFEB144FD055329" box="[616,725,800,824]" class="Insecta" family="Heterogastridae" genus="Sadoletus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="66" phylum="Arthropoda" rank="species" species="validus">
<emphasis id="219A2657FFE13D60FDFEB144FD055329" box="[616,725,800,824]" italics="true" pageId="15" pageNumber="66">S. validus</emphasis>
</taxonomicName>
sensory processes of paramere present).
</paragraph>
<paragraph id="1351FA45FFE13D60FF51B120FDCB5391" blockId="15.[151,1437,151,896]" pageId="15" pageNumber="66">
<emphasis id="219A2657FFE13D60FF51B120FE5A534C" bold="true" box="[199,394,836,861]" pageId="15" pageNumber="66">Biological notes.</emphasis>
The specimens were collected from fruiting trees of
<taxonomicName id="D4EE81C6FFE13D60FC77B121FBCF534D" box="[993,1055,837,860]" class="Magnoliopsida" family="Moraceae" genus="Ficus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Rosales" pageId="15" pageNumber="66" phylum="Tracheophyta" rank="genus">
<emphasis id="219A2657FFE13D60FC77B121FBCF534D" box="[993,1055,837,860]" italics="true" pageId="15" pageNumber="66">Ficus</emphasis>
</taxonomicName>
sp. in comparatively open forests adjacent to farmland and a temple.
</paragraph>
</subSubSection>
</treatment>
</document>
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