282 lines
33 KiB
XML
282 lines
33 KiB
XML
<document id="0E18867FD0D2F8276FDF49BE814F60E0" ID-CLB-Dataset="42241" ID-DOI="10.11646/zootaxa.3914.4.8" ID-GBIF-Dataset="f5485b2b-12a6-40f9-92ae-fd227d61981e" ID-ISSN="1175-5326" ID-Zenodo-Dep="234115" ID-ZooBank="32058514-10EC-4579-813C-091734A0321C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460963296775" checkinUser="plazi" docAuthor="Hotopp, Kenneth P." docDate="2015" docId="9B3087978A43FF81FF7DD78CFE637D65" docLanguage="en" docName="zt03914p494.pdf" docOrigin="Zootaxa 3914 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Triodopsis juxtidens subsp. robinae Hotopp, 2015, subsp. nov." docType="treatment" docVersion="8" lastPageNumber="493" masterDocId="6709FFEF8A43FF82FFEAD167FF9A7B79" masterDocTitle="A new Triodopsis juxtidens subspecies (Gastropoda: Pulmonata) from West Virginia, U. S. A." masterLastPageNumber="494" masterPageNumber="490" pageNumber="490" updateTime="1698587409130" updateUser="ExternalLinkService">
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<mods:titleInfo id="7119082724BC03270F8F937C9C9F7F07">
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<mods:title id="64C5D0DDFCF7CCB728935AEC4D59B110">A new Triodopsis juxtidens subspecies (Gastropoda: Pulmonata) from West Virginia, U. S. A.</mods:title>
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<mods:name id="FBF289305458C9A926CAC9862F2C8E62" type="personal">
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<mods:namePart id="BDDD0DEBF1BF0BF15FAD313CF86139C9">Hotopp, Kenneth P.</mods:namePart>
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<mods:typeOfResource id="1FB762B0FBC5D287933171E1529FC72D">text</mods:typeOfResource>
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<mods:title id="A1FC28959C01CC830095A3137B9996A1">Zootaxa</mods:title>
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<mods:part id="CF30FF0A121655EFA6B3D92751DD5526">
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<mods:date id="F8A0D4FE7787BF575E26801E22C5BEA8">2015</mods:date>
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<mods:number id="F6F1FC222F9EFA94086F7056D227C206">3914</mods:number>
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<mods:number id="2B06799D7164DCE7F36135F317D15B40">4</mods:number>
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<mods:identifier id="897BA71B7D9FBA145416DE03A08B47EC" type="DOI">10.11646/zootaxa.3914.4.8</mods:identifier>
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<mods:identifier id="C7A46CCA69A89BCE585BB74EE343CD58" type="GBIF-Dataset">f5485b2b-12a6-40f9-92ae-fd227d61981e</mods:identifier>
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<treatment id="9B3087978A43FF81FF7DD78CFE637D65" ID-DOI="http://doi.org/10.5281/zenodo.6120022" ID-GBIF-Taxon="119593244" ID-Zenodo-Dep="6120022" LSID="urn:lsid:plazi:treatment:9B3087978A43FF81FF7DD78CFE637D65" httpUri="http://treatment.plazi.org/id/9B3087978A43FF81FF7DD78CFE637D65" lastPageId="3" lastPageNumber="493" pageId="0" pageNumber="490">
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<subSubSection id="5B83650A8A43FF82FF7DD78CFD1A7C7C" box="[151,640,1771,1797]" pageId="0" pageNumber="490" type="nomenclature">
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<paragraph id="132636818A43FF82FF7DD78CFD1A7C7C" blockId="0.[151,640,1771,1797]" box="[151,640,1771,1797]" pageId="0" pageNumber="490">
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<heading id="486E81ED8A43FF82FF7DD78CFD1A7C7C" bold="true" box="[151,640,1771,1797]" fontSize="11" level="1" pageId="0" pageNumber="490" reason="1">
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<emphasis id="21EDEA938A43FF82FF7DD78CFD1A7C7C" bold="true" box="[151,640,1771,1797]" pageId="0" pageNumber="490">
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<taxonomicName id="D4994D028A43FF82FF7DD78CFE757C7C" ID-CoL="7V3NC" box="[151,495,1771,1797]" class="Gastropoda" family="Polygyridae" genus="Triodopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="0" pageNumber="490" phylum="Mollusca" rank="subSpecies" species="juxtidens" status="subsp. nov." subSpecies="robinae">
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<emphasis id="21EDEA938A43FF82FF7DD78CFE757C7C" bold="true" box="[151,495,1771,1797]" italics="true" pageId="0" pageNumber="490">Triodopsis juxtidens robinae</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="3ADE57E88A43FF82FE1CD78CFD1A7C7C" box="[502,640,1771,1797]" pageId="0" pageNumber="490" rank="subSpecies">subsp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="5B83650A8A43FF81FF7DD657FBE979A0" lastPageId="3" lastPageNumber="493" pageId="0" pageNumber="490" type="materials_examined">
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<paragraph id="132636818A43FF82FF7DD657FBF57C10" blockId="0.[151,1436,1840,2036]" pageId="0" pageNumber="490">
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<emphasis id="21EDEA938A43FF82FF7DD657FEAA7C3E" bold="true" box="[151,304,1840,1863]" pageId="0" pageNumber="490">
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<typeStatus id="CC2288238A43FF82FF7DD657FF557C3E" box="[151,207,1840,1863]" pageId="0" pageNumber="490">Type</typeStatus>
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locality.
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</emphasis>
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Brush Creek Preserve, The Nature Conservancy, Mercer County, West Virginia,
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<collectingCountry id="6B8E76118A43FF82FB43D657FB6F7C3E" box="[1193,1269,1840,1863]" name="United States of America" pageId="0" pageNumber="490">U.S.A.</collectingCountry>
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; on and in leaf litter on steep forested slopes of Brush Creek above its confluence with the Bluestone River.
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</paragraph>
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<paragraph id="132636818A43FF82FF2DD612FD9C7C8D" blockId="0.[151,1436,1840,2036]" pageId="0" pageNumber="490">
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<emphasis id="21EDEA938A43FF82FF2DD612FEB37CF5" box="[199,297,1909,1932]" italics="true" pageId="0" pageNumber="490">
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<typeStatus id="CC2288238A43FF82FF2DD612FEB37CF5" box="[199,297,1909,1932]" pageId="0" pageNumber="490" type="holotype">Holotype</typeStatus>
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</emphasis>
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.
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<accessionNumber id="0CCAAB628A43FF82FEDDD612FE287CF5" box="[311,434,1909,1932]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/CM103371" pageId="0" pageNumber="490" type="EnaNcbi">CM103371</accessionNumber>
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,
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<collectingCountry id="6B8E76118A43FF82FE55D612FD9B7CF5" box="[447,513,1909,1932]" name="United States of America" pageId="0" pageNumber="490">U.S.A.</collectingCountry>
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West Virginia, Mercer County, Brush Creek Preserve, Timothy A. Pearce,
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<date id="672710418A43FF82FADAD612FF4A7CD6" pageId="0" pageNumber="490" value="2007-10-01">1 October 2007</date>
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. Shell (
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<figureCitation id="8BA22A048A43FF82FEC9D6FFFE1A7CD6" box="[291,384,1944,1967]" captionStart="FIGURE 1" captionStartId="1.[151,244,1108,1130]" captionTargetBox="[340,1246,178,1085]" captionTargetId="figure@1.[340,1247,178,1085]" captionTargetPageId="1" captionText="FIGURE 1. Shell of Triodopsis j. robinae subsp. nov. holotype from Brush Creek, Mercer Co., West Virginia (CM 103371). Photographed by Charles F. Sturm." httpUri="https://zenodo.org/record/234116/files/figure.png" pageId="0" pageNumber="490">Figure 1</figureCitation>
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) on a steep rocky slope with a mixed eastern hemlock (
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<taxonomicName id="D4994D028A43FF82FC1BD6FFFB357CD6" box="[1009,1199,1944,1967]" class="Pinopsida" family="Pinaceae" genus="Tsuga" kingdom="Plantae" order="Pinales" pageId="0" pageNumber="490" phylum="Tracheophyta" rank="species" species="canadensis">
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<emphasis id="21EDEA938A43FF82FC1BD6FFFB357CD6" box="[1009,1199,1944,1967]" italics="true" pageId="0" pageNumber="490">Tsuga canadensis</emphasis>
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</taxonomicName>
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L.(Carr.))-hardwood forest canopy, on the east side of Brush Creek above its confluence with the Bluestone River (within
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<quantity id="D4619B648A43FF82FAEAD6DDFAAA7CA8" box="[1280,1328,1978,2001]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="0" pageNumber="490" unit="m" value="10.0">10m</quantity>
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of UTM, NAD83, 17S, 0 494625, 4147911).
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</paragraph>
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<caption id="47E666098A42FF83FF7DD533FE727FFE" httpUri="https://zenodo.org/record/234116/files/figure.png" pageId="1" pageNumber="491" targetBox="[340,1246,178,1085]" targetPageId="1">
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<paragraph id="132636818A42FF83FF7DD533FE727FFE" blockId="1.[151,1437,1108,1159]" pageId="1" pageNumber="491">
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<emphasis id="21EDEA938A42FF83FF7DD533FE887F13" bold="true" box="[151,274,1108,1130]" pageId="1" pageNumber="491">FIGURE 1.</emphasis>
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Shell of
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<taxonomicName id="D4994D028A42FF83FE90D532FDD47F13" box="[378,590,1109,1130]" class="Gastropoda" family="Polygyridae" genus="Triodopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="1" pageNumber="491" phylum="Mollusca" rank="subSpecies" species="juxtidens" status="subsp. nov." subSpecies="robinae">
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<emphasis id="21EDEA938A42FF83FE90D532FDD47F13" box="[378,590,1109,1130]" italics="true" pageId="1" pageNumber="491">Triodopsis j. robinae</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="3ADE57E88A42FF83FDB3D532FD517F13" box="[601,715,1109,1130]" pageId="1" pageNumber="491" rank="subSpecies">subsp. nov.</taxonomicNameLabel>
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holotype from Brush Creek, Mercer Co., West Virginia (
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<accessionNumber id="0CCAAB628A42FF83FAF5D532FA0B7F13" box="[1311,1425,1109,1130]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/CM103371" pageId="1" pageNumber="491" type="EnaNcbi">CM103371</accessionNumber>
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). Photographed by Charles F. Sturm.
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</paragraph>
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</caption>
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<caption id="47E666098A42FF83FF7DD6DBFEA57C96" httpUri="https://zenodo.org/record/234117/files/figure.png" pageId="1" pageNumber="491" targetBox="[321,1265,1210,1956]" targetPageId="1">
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<paragraph id="132636818A42FF83FF7DD6DBFEA57C96" blockId="1.[151,1436,1980,2031]" pageId="1" pageNumber="491">
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<emphasis id="21EDEA938A42FF83FF7DD6DBFE957CAB" bold="true" box="[151,271,1980,2002]" pageId="1" pageNumber="491">FIGURE 2.</emphasis>
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View of the aperture and basal denticle of
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<taxonomicName id="D4994D028A42FF83FD28D6DBFC157CA8" box="[706,911,1980,2001]" class="Gastropoda" family="Polygyridae" genus="Triodopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="1" pageNumber="491" phylum="Mollusca" rank="subSpecies" species="juxtidens" status="subsp. nov." subSpecies="robinae">
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<emphasis id="21EDEA938A42FF83FD28D6DBFC157CA8" box="[706,911,1980,2001]" italics="true" pageId="1" pageNumber="491">Triodopsis j. robinae</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="3ADE57E88A42FF83FC72D6DBFB9C7CA8" box="[920,1030,1980,2001]" pageId="1" pageNumber="491" rank="subSpecies">subsp. nov.</taxonomicNameLabel>
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holotype (
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<accessionNumber id="0CCAAB628A42FF83FB9BD6DBFB797CA8" box="[1137,1251,1980,2001]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/CM103371" pageId="1" pageNumber="491" type="EnaNcbi">CM103371</accessionNumber>
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). Photographed by Charles F. Sturm.
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</paragraph>
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</caption>
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<caption id="47E666098A41FF80FF7DD78AFC047C94" httpUri="https://zenodo.org/record/234118/files/figure.png" pageId="2" pageNumber="492" targetBox="[465,1112,194,1742]" targetPageId="2">
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<paragraph id="132636818A41FF80FF7DD78AFC047C94" blockId="2.[151,1436,1773,2029]" pageId="2" pageNumber="492">
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<emphasis id="21EDEA938A41FF80FF7DD78AFE967C7A" bold="true" box="[151,268,1773,1795]" pageId="2" pageNumber="492">FIGURE 3.</emphasis>
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Comparison of seven
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<taxonomicName id="D4994D028A41FF80FE0ED78AFDD37C7B" box="[484,585,1773,1794]" class="Gastropoda" family="Polygyridae" genus="Triodopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="492" phylum="Mollusca" rank="genus">
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<emphasis id="21EDEA938A41FF80FE0ED78AFDD37C7B" box="[484,585,1773,1794]" italics="true" pageId="2" pageNumber="492">Triodopsis</emphasis>
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</taxonomicName>
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species shells (left) to a series of six
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<taxonomicName id="D4994D028A41FF80FC5BD78AFBE27C7B" box="[945,1144,1773,1794]" class="Gastropoda" family="Polygyridae" genus="Triodopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="492" phylum="Mollusca" rank="subSpecies" species="juxtidens" status="subsp. nov." subSpecies="robinae">
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<emphasis id="21EDEA938A41FF80FC5BD78AFBE27C7B" box="[945,1144,1773,1794]" italics="true" pageId="2" pageNumber="492">Triodopsis j. robinae</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="3ADE57E88A41FF80FB95D78AFB717C7B" box="[1151,1259,1773,1794]" pageId="2" pageNumber="492" rank="subSpecies">subsp. nov.</taxonomicNameLabel>
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shells (right). The
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<taxonomicName id="D4994D028A41FF80FF7DD66CFF667C59" box="[151,252,1803,1824]" class="Gastropoda" family="Polygyridae" genus="Triodopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="492" phylum="Mollusca" rank="genus">
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<emphasis id="21EDEA938A41FF80FF7DD66CFF667C59" box="[151,252,1803,1824]" italics="true" pageId="2" pageNumber="492">Triodopsis</emphasis>
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</taxonomicName>
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species representatives have similar shells to the new subspecies or are geographically near. To the left, from top to bottom, are: a,
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<taxonomicName id="D4994D028A41FF80FF3CD64FFE3A7C44" authority="Durham" authorityName="Durham" box="[214,416,1832,1853]" class="Gastropoda" family="Polygyridae" genus="Triodopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="492" phylum="Mollusca" rank="species" species="juxtidens">
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<emphasis id="21EDEA938A41FF80FF3CD64FFED97C44" box="[214,323,1832,1853]" italics="true" pageId="2" pageNumber="492">T. juxtidens</emphasis>
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, Durham
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</taxonomicName>
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, NC, 11 August 2008 (W. Frank, personal collection); b,
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<taxonomicName id="D4994D028A41FF80FC54D64FFBB07C44" box="[958,1066,1832,1853]" class="Gastropoda" family="Polygyridae" genus="Triodopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="492" phylum="Mollusca" rank="species" species="juxtidens">
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<emphasis id="21EDEA938A41FF80FC54D64FFBB07C44" box="[958,1066,1832,1853]" italics="true" pageId="2" pageNumber="492">T. juxtidens</emphasis>
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</taxonomicName>
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also from Durham Co., NC (H.G. Lee, personal collection); c,
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<taxonomicName id="D4994D028A41FF80FE92D622FE497C23" box="[376,467,1861,1882]" class="Gastropoda" family="Polygyridae" genus="Triodopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="492" phylum="Mollusca" rank="species" species="rugosa">
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<emphasis id="21EDEA938A41FF80FE92D622FE497C23" box="[376,467,1861,1882]" italics="true" pageId="2" pageNumber="492">T. rugosa</emphasis>
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</taxonomicName>
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from Logan Co., WV (FMNH264690); d,
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<taxonomicName id="D4994D028A41FF80FC9AD622FC7F7C23" box="[880,997,1861,1882]" class="Gastropoda" family="Polygyridae" genus="Triodopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="492" phylum="Mollusca" rank="species" species="anteridon">
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<emphasis id="21EDEA938A41FF80FC9AD622FC7F7C23" box="[880,997,1861,1882]" italics="true" pageId="2" pageNumber="492">T. anteridon</emphasis>
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</taxonomicName>
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from Wyoming Co., WV (FMNH264768); e,
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<taxonomicName id="D4994D028A41FF80FF7DD604FF7E7C01" box="[151,228,1891,1912]" class="Gastropoda" family="Polygyridae" genus="Triodopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="492" phylum="Mollusca" rank="species" species="picea">
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<emphasis id="21EDEA938A41FF80FF7DD604FF7E7C01" box="[151,228,1891,1912]" italics="true" pageId="2" pageNumber="492">T. picea</emphasis>
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</taxonomicName>
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from Pocahontas Co., WV (FMNH266426); f,
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<taxonomicName id="D4994D028A41FF80FD50D604FCBB7C01" box="[698,801,1891,1912]" class="Gastropoda" family="Polygyridae" genus="Triodopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="492" phylum="Mollusca" rank="species" species="pendula">
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<emphasis id="21EDEA938A41FF80FD50D604FCBB7C01" box="[698,801,1891,1912]" italics="true" pageId="2" pageNumber="492">T. pendula</emphasis>
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</taxonomicName>
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from Iredell Co., NC (FMNH266352), and g,
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<taxonomicName id="D4994D028A41FF80FB1AD604FAFF7C01" box="[1264,1381,1891,1912]" class="Gastropoda" family="Polygyridae" genus="Triodopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="492" phylum="Mollusca" rank="species" species="discoidea">
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<emphasis id="21EDEA938A41FF80FB1AD604FAFF7C01" box="[1264,1381,1891,1912]" italics="true" pageId="2" pageNumber="492">T. discoidea</emphasis>
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</taxonomicName>
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from Crawford Co., IN (H.G. Lee, personal collection). To the right are paratype h,
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<taxonomicName id="D4994D028A41FF80FC4AD6E7FBF67CEC" box="[928,1132,1920,1941]" class="Gastropoda" family="Polygyridae" genus="Triodopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="492" phylum="Mollusca" rank="subSpecies" species="juxtidens" status="subsp. nov." subSpecies="robinae">
|
||
<emphasis id="21EDEA938A41FF80FC4AD6E7FBF67CEC" box="[928,1132,1920,1941]" italics="true" pageId="2" pageNumber="492">Triodopsis j. robinae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="3ADE57E88A41FF80FB9ED6E7FB797CEC" box="[1140,1251,1920,1941]" pageId="2" pageNumber="492" rank="subSpecies">subsp. nov.</taxonomicNameLabel>
|
||
from Brush Creek (CM97548); paratypes i, j, Brush Creek (CM97549; two shells); k, the holotype from Brush Creek (
|
||
<accessionNumber id="0CCAAB628A41FF80FBBFD6FAFB5D7CCB" box="[1109,1223,1949,1970]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/CM103371" pageId="2" pageNumber="492" type="EnaNcbi">CM103371</accessionNumber>
|
||
); paratype l, Pipestem State Park (
|
||
<accessionNumber id="0CCAAB628A41FF80FEE2D6DCFEE07CA9" box="[264,378,1979,2000]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/CM141101" pageId="2" pageNumber="492" type="EnaNcbi">CM141101</accessionNumber>
|
||
); and paratype m, Brush Creek (CM85754). Adapted from Frank and Lee (2014), with permission, and with photography of
|
||
<taxonomicName id="D4994D028A41FF80FEDAD6BFFE6C7C94" box="[304,502,2008,2029]" class="Gastropoda" family="Polygyridae" genus="Triodopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="492" phylum="Mollusca" rank="subSpecies" species="juxtidens" status="subsp. nov." subSpecies="robinae">
|
||
<emphasis id="21EDEA938A41FF80FEDAD6BFFE6C7C94" box="[304,502,2008,2029]" italics="true" pageId="2" pageNumber="492">Triodopsis j. robinae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="3ADE57E88A41FF80FE17D6BFFDF37C94" box="[509,617,2008,2029]" pageId="2" pageNumber="492" rank="subSpecies">subsp. nov.</taxonomicNameLabel>
|
||
specimens by Charles F. Sturm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="132636818A40FF81FF2DD1FFFEF57A9E" blockId="3.[151,1436,152,1564]" pageId="3" pageNumber="493">
|
||
<emphasis id="21EDEA938A40FF81FF2DD1FFFEDA7BD6" bold="true" box="[199,320,152,175]" pageId="3" pageNumber="493">
|
||
<typeStatus id="CC2288238A40FF81FF2DD1FFFEA17BD6" box="[199,315,152,175]" pageId="3" pageNumber="493" type="paratype">Paratypes</typeStatus>
|
||
.
|
||
</emphasis>
|
||
CM97548, five shells from the west side of Brush Creek close to its confluence with the Bluestone River, Kenneth P. Hotopp and Brett Freedman,
|
||
<date id="672710418A40FF81FD7ED1DDFCAC7BA8" box="[660,822,186,209]" pageId="3" pageNumber="493" value="2005-08-03">3 August 2005</date>
|
||
; CM97549, five shells from mesic forest among cliffs on the west side of Brush Creek near the
|
||
<typeStatus id="CC2288238A40FF81FDC5D1BAFDC77B8D" box="[559,605,221,244]" pageId="3" pageNumber="493">type</typeStatus>
|
||
locality, Kenneth P. Hotopp and Brett Freedman,
|
||
<date id="672710418A40FF81FB99D1BAFA8B7B8D" box="[1139,1297,221,244]" pageId="3" pageNumber="493" value="2005-08-03">3 August 2005</date>
|
||
(within
|
||
<quantity id="D4619B648A40FF81FA86D1BAFA017B8D" box="[1388,1435,221,244]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="493" unit="m" value="50.0">50m</quantity>
|
||
of UTM, NAD83, 17S, 0 494627, 4147741);
|
||
<accessionNumber id="0CCAAB628A40FF81FD64D067FC937A6E" box="[654,777,256,279]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/CM141101" pageId="3" pageNumber="493" type="EnaNcbi">
|
||
CM
|
||
<date id="672710418A40FF81FD53D067FC937A6E" box="[697,777,256,279]" pageId="3" pageNumber="493" value="01-11-14">141101</date>
|
||
</accessionNumber>
|
||
, five shells from hardwood forest near a hillside spring at Pipestem State Park, Kenneth P. Hotopp and Kenneth R. Hotopp (Sr.),
|
||
<date id="672710418A40FF81FC55D045FB027A40" box="[959,1176,290,313]" pageId="3" pageNumber="493" value="2008-09-19">19 September 2008</date>
|
||
(within
|
||
<quantity id="D4619B648A40FF81FB17D045FAB67A40" box="[1277,1324,290,313]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="493" unit="m" value="50.0">50m</quantity>
|
||
of UTM, NAD83, 17S, 0 500316, 4152917);
|
||
<accessionNumber id="0CCAAB628A40FF81FDF1D022FD0C7A25" box="[539,662,325,348]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/CM141102" pageId="3" pageNumber="493" type="EnaNcbi">
|
||
CM
|
||
<date id="672710418A40FF81FDAFD022FD0C7A25" box="[581,662,325,348]" pageId="3" pageNumber="493" value="02-11-14">141102</date>
|
||
</accessionNumber>
|
||
, five shells from mixed forest upon sandstone talus at Pipestem State Park, Kenneth P. Hotopp and Kenneth R. Hotopp (Sr.),
|
||
<date id="672710418A40FF81FD0BD00FFC2A7A06" box="[737,944,360,383]" pageId="3" pageNumber="493" value="2008-09-19">19 September 2008</date>
|
||
(
|
||
<figureCitation id="8BA22A048A40FF81FC57D00FFB837A06" box="[957,1049,360,383]" captionStart="FIGURE 3" captionStartId="2.[151,244,1773,1795]" captionTargetBox="[465,1112,194,1742]" captionTargetId="figure@2.[465,1113,194,1743]" captionTargetPageId="2" captionText="FIGURE 3. Comparison of seven Triodopsis species shells (left) to a series of six Triodopsis j. robinae subsp. nov. shells (right). The Triodopsis species representatives have similar shells to the new subspecies or are geographically near. To the left, from top to bottom, are: a, T. juxtidens, Durham, NC, 11 August 2008 (W. Frank, personal collection); b, T. juxtidens also from Durham Co., NC (H. G. Lee, personal collection); c, T. rugosa from Logan Co., WV (FMNH 264690); d, T. anteridon from Wyoming Co., WV (FMNH 264768); e, T. picea from Pocahontas Co., WV (FMNH 266426); f, T. pendula from Iredell Co., NC (FMNH 266352), and g, T. discoidea from Crawford Co., IN (H. G. Lee, personal collection). To the right are paratype h, Triodopsis j. robinae subsp. nov. from Brush Creek (CM 97548); paratypes i, j, Brush Creek (CM 97549; two shells); k, the holotype from Brush Creek (CM 103371); paratype l, Pipestem State Park (CM 141101); and paratype m, Brush Creek (CM 85754). Adapted from Frank and Lee (2014), with permission, and with photography of Triodopsis j. robinae subsp. nov. specimens by Charles F. Sturm." httpUri="https://zenodo.org/record/234118/files/figure.png" pageId="3" pageNumber="493">Figure 3</figureCitation>
|
||
; within
|
||
<quantity id="D4619B648A40FF81FB9AD00FFB057A06" box="[1136,1183,360,383]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="493" unit="m" value="50.0">50m</quantity>
|
||
of UTM, NAD83, 17S, 0 498866, 4154255); CM85754, one live specimen on a steep, wooded slope at the foot of a cliff above a ravine, east side of Brush Creek, Timothy A. Pearce and Alice W. Doolittle,
|
||
<date id="672710418A40FF81FCA9D0CAFC6E7ABD" box="[835,1012,429,452]" pageId="3" pageNumber="493" value="2007-10-01">1 October 2007</date>
|
||
, (within
|
||
<quantity id="D4619B648A40FF81FBB6D0CAFB117ABD" box="[1116,1163,429,452]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="493" unit="m" value="10.0">10m</quantity>
|
||
of UTM, NAD83, 17S, 0 494866, 4147454).
|
||
</paragraph>
|
||
<paragraph id="132636818A40FF81FF2DD095FBE979A0" blockId="3.[151,1436,152,1564]" pageId="3" pageNumber="493">
|
||
<emphasis id="21EDEA938A40FF81FF2DD095FE367970" bold="true" box="[199,428,498,521]" pageId="3" pageNumber="493">Additional material.</emphasis>
|
||
Daniel C. Dourson, personal collection, 16 shells from cliffline at Brush Creek, Jeffrey J. Hajenga,
|
||
<date id="672710418A40FF81FF11D372FEFB7955" box="[251,353,533,556]" pageId="3" pageNumber="493" value="2012-03">12 March</date>
|
||
, 2013; Daniel C. Dourson, personal collection, 16 shells from forest at Brush Creek, Jeffrey J. Hajenga, undated; Daniel C. Dourson, personal collection, 5 shells from forest at Sand Branch, Jeffrey J. Hajenga,
|
||
<date id="672710418A40FF81FB0AD35FFACC7936" box="[1248,1366,568,591]" pageId="3" pageNumber="493" value="2013-10">13 October</date>
|
||
, 2013;
|
||
<accessionNumber id="0CCAAB628A40FF81FF7DD33DFE897908" box="[151,275,602,625]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/CM103371" pageId="3" pageNumber="493" type="EnaNcbi">CM103371</accessionNumber>
|
||
, one immature shell from mixed forest, southeast side of junction of Brush Creek with Bluestone River, Timothy A. Pearce,
|
||
<date id="672710418A40FF81FE9FD31AFD8579ED" box="[373,543,637,660]" pageId="3" pageNumber="493" value="2007-10-01">1 October 2007</date>
|
||
(within
|
||
<quantity id="D4619B648A40FF81FD94D31AFD3479ED" box="[638,686,637,660]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="493" unit="m" value="10.0">10m</quantity>
|
||
of UTM, WGS84, 17S, 0 494625, 4147911); CM85754, five broken shells from a steep, wooded slope at the foot of a cliff above a ravine, east side of Brush Creek, Timothy A. Pearce and Alice W. Doolittle,
|
||
<date id="672710418A40FF81FE8DD3A5FD9779A0" box="[359,525,706,729]" pageId="3" pageNumber="493" value="2007-10-01">1 October 2007</date>
|
||
(within
|
||
<quantity id="D4619B648A40FF81FD8DD3A5FD0C79A0" box="[615,662,706,729]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="493" unit="m" value="10.0">10m</quantity>
|
||
of UTM, NAD83, 17S, 0 494866, 4147454).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="5B83650A8A40FF81FF2DD382FB807858" pageId="3" pageNumber="493" type="etymology">
|
||
<paragraph id="132636818A40FF81FF2DD382FB807858" blockId="3.[151,1436,152,1564]" pageId="3" pageNumber="493">
|
||
<emphasis id="21EDEA938A40FF81FF2DD382FEDD7985" bold="true" box="[199,327,741,764]" pageId="3" pageNumber="493">Etymology.</emphasis>
|
||
The subspecific name
|
||
<emphasis id="21EDEA938A40FF81FDA3D382FD017985" box="[585,667,741,764]" italics="true" pageId="3" pageNumber="493">robinae</emphasis>
|
||
is in honor of my wonderful wife, Mary “Robin” Robertson Gorrell, originally of Clarksburg, West Virginia, on the year of our 25th wedding anniversary.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="5B83650A8A40FF81FF2DD24AFEDE781E" pageId="3" pageNumber="493" type="distribution">
|
||
<paragraph id="132636818A40FF81FF2DD24AFEDE781E" blockId="3.[151,1436,152,1564]" pageId="3" pageNumber="493">
|
||
<emphasis id="21EDEA938A40FF81FF2DD24AFEC3783D" bold="true" box="[199,345,813,836]" pageId="3" pageNumber="493">Distribution.</emphasis>
|
||
Forested slopes of the Bluestone River Valley and tributaries in Mercer and Summers Counties, West Virginia,
|
||
<collectingCountry id="6B8E76118A40FF81FF11D237FEDE781E" box="[251,324,848,871]" name="United States of America" pageId="3" pageNumber="493">U.S.A.</collectingCountry>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="5B83650A8A40FF81FF2DD215FE637D65" pageId="3" pageNumber="493" type="description">
|
||
<paragraph id="132636818A40FF81FF2DD215FE637D65" blockId="3.[151,1436,152,1564]" pageId="3" pageNumber="493">
|
||
<emphasis id="21EDEA938A40FF81FF2DD215FEC878F0" bold="true" box="[199,338,882,905]" pageId="3" pageNumber="493">Description.</emphasis>
|
||
Shell (Figures 1,2)
|
||
<quantity id="D4619B648A40FF81FDC5D215FDE078F0" box="[559,634,882,905]" metricMagnitude="-3" metricUnit="m" metricValue="7.7" pageId="3" pageNumber="493" unit="mm" value="7.7">7.7mm</quantity>
|
||
tall (
|
||
<typeStatus id="CC2288238A40FF81FD5CD215FC8478F0" box="[694,798,882,905]" pageId="3" pageNumber="493" type="paratype">paratypes</typeStatus>
|
||
7.0–9.0mm; measured from apex to lowest point on shell with axis vertical) x
|
||
<quantity id="D4619B648A40FF81FE6FD2F2FE4778D5" box="[389,477,917,940]" metricMagnitude="-2" metricUnit="m" metricValue="1.47" pageId="3" pageNumber="493" unit="mm" value="14.7">14.7mm</quantity>
|
||
wide (
|
||
<typeStatus id="CC2288238A40FF81FDC5D2F2FD0D78D5" box="[559,663,917,940]" pageId="3" pageNumber="493" type="paratype">paratypes</typeStatus>
|
||
<quantity id="D4619B648A40FF81FD48D2F2FCA378D5" box="[674,825,917,940]" metricMagnitude="-2" metricUnit="m" metricValue="1.535" metricValueMax="1.76" metricValueMin="1.31" pageId="3" pageNumber="493" unit="mm" value="15.35" valueMax="17.6" valueMin="13.1">13.1–17.6mm</quantity>
|
||
; measured including the peristome), height:width 0.49 (
|
||
<typeStatus id="CC2288238A40FF81FF74D2DFFE9C78B6" box="[158,262,952,975]" pageId="3" pageNumber="493" type="paratype">paratypes</typeStatus>
|
||
0.45–0.58); umbilicus open,
|
||
<quantity id="D4619B648A40FF81FDAFD2DFFD1578B6" box="[581,655,952,975]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="3" pageNumber="493" unit="mm" value="3.2">3.2mm</quantity>
|
||
wide (
|
||
<typeStatus id="CC2288238A40FF81FD31D2DFFCD978B6" box="[731,835,952,975]" pageId="3" pageNumber="493" type="paratype">paratypes</typeStatus>
|
||
<quantity id="D4619B648A40FF81FCA1D2DFFC5278B6" box="[843,968,952,975]" metricMagnitude="-3" metricUnit="m" metricValue="3.25" metricValueMax="4.2" metricValueMin="2.3" pageId="3" pageNumber="493" unit="mm" value="3.25" valueMax="4.2" valueMin="2.3">2.3–4.2mm</quantity>
|
||
; measured at the whorl suture immediately behind the peristome to the suture on the farthest side of the umbilicus), umbilicus:width 0.20 (
|
||
<typeStatus id="CC2288238A40FF81FB5AD2BDFA827888" box="[1200,1304,986,1009]" pageId="3" pageNumber="493" type="paratype">paratypes</typeStatus>
|
||
0.17–0.24); whorls 5.2 (4.75–5.5; counted along a tangent starting parallel with the beginning of the apical suture, following Pilsbry [1939]); color light brown to reddish-brown. Protoconch and juvenile whorls with fine growth wrinkles, subsequent whorls with growth wrinkles and fine, regular ribs, approximately 3.5 ribs/mm (
|
||
<typeStatus id="CC2288238A40FF81FBF3D525FB1B7F20" box="[1049,1153,1090,1113]" pageId="3" pageNumber="493" type="paratype">paratypes</typeStatus>
|
||
3.5–4.5) on final quarter whorl; sculpture becoming minutely papillose, primarily at the suture and final whorl; final whorl inflated near the end, the suture rising upon the wall of the penultimate whorl, then abruptly descending and well-constricted behind the aperture. Aperture widely reflected, white,
|
||
<quantity id="D4619B648A40FF81FD96D5CDFD5D7FB8" box="[636,711,1194,1217]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="3" pageNumber="493" unit="mm" value="7.5">7.5mm</quantity>
|
||
tall (outside distance, measured on plane of aperture;
|
||
<typeStatus id="CC2288238A40FF81FADFD5CDFA017FB8" box="[1333,1435,1194,1217]" pageId="3" pageNumber="493" type="paratype">paratypes</typeStatus>
|
||
<quantity id="D4619B648A40FF81FF7DD5AAFE887F9D" box="[151,274,1229,1252]" metricMagnitude="-3" metricUnit="m" metricValue="6.85" metricValueMax="7.8" metricValueMin="5.9" pageId="3" pageNumber="493" unit="mm" value="6.85" valueMax="7.8" valueMin="5.9">5.9–7.8mm</quantity>
|
||
) x
|
||
<quantity id="D4619B648A40FF81FED9D5AAFEE47F9D" box="[307,382,1229,1252]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" pageId="3" pageNumber="493" unit="mm" value="8.5">8.5mm</quantity>
|
||
wide (outside distance, measured from columellar edge of basal peristome to farthest palatal limit;
|
||
<typeStatus id="CC2288238A40FF81FF7DD597FF677E7E" box="[151,253,1264,1287]" pageId="3" pageNumber="493" type="paratype">paratypes</typeStatus>
|
||
<quantity id="D4619B648A40FF81FEEFD597FE197E7E" box="[261,387,1264,1287]" metricMagnitude="-3" metricUnit="m" metricValue="7.949999999999999" metricValueMax="9.1" metricValueMin="6.8" pageId="3" pageNumber="493" unit="mm" value="7.949999999999999" valueMax="9.1" valueMin="6.8">6.8–9.1mm</quantity>
|
||
), broadly triangular (viewed in the plane of the aperture), with the triangle’s apex usually to the right of the point at which the peristome meets the body whorl, and widest below an imaginary horizontal midline; peristome slightly-to-moderately dished and projecting forward at the top edge, with parietal, basal and palatal lamellae, often nearly straight along its basal edge; a light parietal callus at the aperture. Parietal lamella narrow, slightly to strongly curved toward the palatal denticle, its umbilical end sometimes continuous with the basal peristome; palatal denticle variable, usually wide at the distal end but sometimes pointed, and pointing inward behind parietal lamella; interdenticular sinus usually a defined semicircle; basal denticle deeper than wide, set perpendicular to the aperture upon a horizontal buttress (
|
||
<figureCitation id="8BA22A048A40FF81FE91D485FE427E80" box="[379,472,1506,1529]" captionStart="FIGURE 2" captionStartId="1.[151,244,1980,2002]" captionTargetBox="[321,1265,1210,1956]" captionTargetId="figure@1.[321,1265,1210,1956]" captionTargetPageId="1" captionText="FIGURE 2. View of the aperture and basal denticle of Triodopsis j. robinae subsp. nov. holotype (CM 103371). Photographed by Charles F. Sturm." httpUri="https://zenodo.org/record/234117/files/figure.png" pageId="3" pageNumber="493">Figure 2</figureCitation>
|
||
), the buttress usually taller to the palatal side of the tooth. Body of animal gray with dorsal surface of upper tentacles darker.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |