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<document id="5C3CBA4DA793B4B4D80C1DA681A74AC1" ENCODING="UTF-8" ID-GBIF-Dataset="de6121e1-d370-43ff-8a55-54eff752a6f8" ModsDocID="z00462p001" checkinTime="1247045511124" checkinUser="thomas" docAuthor="James L. Van Tassell &amp; Carole C. Baldwin" docDate="2004" docId="9A95908552F8602F793A5435E98F2229" docLanguage="en" docName="2004_VanTassell_Baldwin_gg1.xml" docOrigin="Zootaxa 462" docSource="http://www.zoobank.org/urn:lsid:zoobank.org:pub:73EC2E7B-E100-4C34-9485-3BBC53531905" docTitle="Akko Birdsong and Robins" docType="treatment" docVersion="15" lastPageNumber="2" masterDocId="802F49E080BEEDBE85A511D3CC247C7C" masterDocTitle="A review of the gobiid genus Akko (Teleostei: Gobiidae) with description of a new species." masterLastPageNumber="15" masterPageNumber="1" pageNumber="1" updateTime="1701313966396" updateUser="plazi">
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<mods:title id="6CA5F16D921FE9D0D8FA1C0F4EF1A5D5">A review of the gobiid genus Akko (Teleostei: Gobiidae) with description of a new species.</mods:title>
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<mods:namePart id="904ECC17F877BE4D80E7120D43AC7301">James L. Van Tassell</mods:namePart>
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<mods:namePart id="0C95EA1BDD9360BA4B3DA805D84EDA45">Carole C. Baldwin</mods:namePart>
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<treatment id="9A95908552F8602F793A5435E98F2229" ID-DOI="http://doi.org/10.5281/zenodo.6270520" ID-GBIF-Taxon="100121584" ID-Zenodo-Dep="6270520" LSID="urn:lsid:plazi:treatment:9A95908552F8602F793A5435E98F2229" httpUri="http://treatment.plazi.org/id/9A95908552F8602F793A5435E98F2229" lastPageNumber="2" pageNumber="1">
<subSubSection id="C38342807179E0D46B561039E37A2E43" type="nomenclature">
<paragraph id="9462B1A6BBFD389AEC6835F8B0206C75">
[[
<taxonomicName id="4EA9B38105E0A3E9506180FBDA556087" ID-CoL="RB2" ID-ENA="203341" LSID-ZBK="urn:lsid:zoobank.org:act:1654F3F7-4BF7-44EE-BD21-80A538552441" family="Gobiidae" genus="Akko" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Akko Birdsong &amp; Robins 1995:676" order="Perciformes" pageNumber="1" phylum="Chordata" rank="genus">Akko Birdsong and Robins</taxonomicName>
]]
</paragraph>
</subSubSection>
<subSubSection id="608C7E5031743903795B3DB802CD5561" type="discussion">
<paragraph id="73CC2D10BA5DF0FADBFD1DD87250B6D0" pageNumber="1">Introduction</paragraph>
<paragraph id="9BC3D1B6452F42B07A5CAF02767B23A0" lastPageNumber="2" pageNumber="1">
<taxonomicName id="1E5216A183CAF07162B3E8C28848DDE5" LSID-ZBK="urn:lsid:zoobank.org:act:1D669904-3B1B-4EB8-99A5-493BE7889CCA" family="Gobiidae" genus="Amblyopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Amblyopus brevis Günther 1864:151" order="Perciformes" pageNumber="1" phylum="Chordata" rank="species" species="brevis">Amblyopus brevis</taxonomicName>
was described by
<normalizedToken id="67346A0AD8A492811C4220577578C188" originalValue="Günther">Guenther</normalizedToken>
(1864) from a single specimen collected along the Pacific coast of Panama. Two additional specimens were later obtained from stomach contents of a
<taxonomicName id="783624A5E27820A3370FD5E0A9085E2D" LSID-ZBK="urn:lsid:zoobank.org:act:24D43392-272C-4DCA-9755-A65C5ED99DBE" family="Centropomidae" genus="Centropomus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Centropomus Lacépède 1802:248" order="Perciformes" pageNumber="1" phylum="Chordata" rank="genus">Centropomus</taxonomicName>
and added to the collection at the British Museum.
<normalizedToken id="026ECE914CFCD84023C57DBEC2A3F6E1" originalValue="Günther">Guenther</normalizedToken>
(1869) placed
<taxonomicName id="6EEAC971855F35F0E781AFA4466A1DBE" LSID-ZBK="urn:lsid:zoobank.org:act:1D669904-3B1B-4EB8-99A5-493BE7889CCA" family="Gobiidae" genus="Amblyopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Amblyopus brevis Günther 1864:151" order="Perciformes" pageNumber="1" phylum="Chordata" rank="species" species="brevis">A. brevis</taxonomicName>
in the subgenus
<taxonomicName id="BFEDF0AACB90A367B4B76C3EFDF7AC61" LSID-ZBK="urn:lsid:zoobank.org:act:475E2464-87A8-40E3-9A7A-F76987320994" family="Gobiidae" genus="Tyntlastes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Tyntlastes Amblyopus Günther 1862:194" order="Perciformes" pageNumber="1" phylum="Chordata" rank="genus">Tyntlastes</taxonomicName>
based on its dentition (teeth in a single series). Jordan and Eigenmann (1886[1887]) elevated
<taxonomicName id="73AB00C55941BAC5B20A6A89E3A31A56" LSID-ZBK="urn:lsid:zoobank.org:act:475E2464-87A8-40E3-9A7A-F76987320994" family="Gobiidae" genus="Tyntlastes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Tyntlastes Amblyopus Günther 1862:194" order="Perciformes" pageNumber="1" phylum="Chordata" rank="genus">Tyntlastes</taxonomicName>
to a genus and assigned to it the same species that comprised
<normalizedToken id="D18ED6EB97B3404C78359DC6B1147258" originalValue="Günther">Guenther</normalizedToken>
’s subgenus (
<taxonomicName id="4AF7D9969D4133790FEE65156A106782" LSID-ZBK="urn:lsid:zoobank.org:act:1D669904-3B1B-4EB8-99A5-493BE7889CCA" family="Gobiidae" genus="Amblyopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Amblyopus brevis Günther 1864:151" order="Perciformes" pageNumber="1" phylum="Chordata" rank="species" species="brevis">A. brevis</taxonomicName>
and
<taxonomicName id="D88A1B596BBB988178D615B27C8D4B57" LSID-ZBK="urn:lsid:zoobank.org:act:C4D25825-CD53-45D4-BD20-1FF262826F9F" family="Gobiidae" genus="Amblyopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Amblyopus sagitta Günther 1862:193 [6]" order="Perciformes" pageNumber="1" phylum="Chordata" rank="species" species="sagitta">A. sagitta</taxonomicName>
). When Palmer (1952) reviewed the genus
<taxonomicName id="7CFA08CDA872DB320265D0D84F0DE5BD" LSID-ZBK="urn:lsid:zoobank.org:act:BA04C1EE-19AC-4F48-B98A-B4A18304A427" family="Gobiidae" genus="Gobioides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Gobioides Lacépède 1800:576" order="Perciformes" pageNumber="1" phylum="Chordata" rank="genus">Gobioides</taxonomicName>
, he created two subgenera:
<taxonomicName id="565EB2ADC71BEC38FFE1D649563A6FCD" LSID-ZBK="urn:lsid:zoobank.org:act:BA04C1EE-19AC-4F48-B98A-B4A18304A427" family="Gobiidae" genus="Gobioides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Gobioides Lacépède 1800:576" order="Perciformes" pageNumber="1" phylum="Chordata" rank="genus">Gobioides</taxonomicName>
, with 25-26 vertebrae and 14-16 anal-fin rays, and
<taxonomicName id="E7030507E06879C6A050C4730AC53292" LSID-ZBK="urn:lsid:zoobank.org:act:475E2464-87A8-40E3-9A7A-F76987320994" family="Gobiidae" genus="Tyntlastes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Tyntlastes Amblyopus Günther 1862:194" order="Perciformes" pageNumber="1" phylum="Chordata" rank="genus">Tyntlastes</taxonomicName>
, with 31 vertebrae and 19 anal-fin rays. He placed
<taxonomicName id="7EC77C9BAD6FE4A4F37A7743588C8AA8" LSID-ZBK="urn:lsid:zoobank.org:act:1D669904-3B1B-4EB8-99A5-493BE7889CCA" family="Gobiidae" genus="Amblyopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Amblyopus brevis Günther 1864:151" order="Perciformes" pageNumber="1" phylum="Chordata" rank="species" species="brevis">A. brevis</taxonomicName>
in
<taxonomicName id="EDF62B5B60CDD29E64E57695600351BF" LSID-ZBK="urn:lsid:zoobank.org:act:BA04C1EE-19AC-4F48-B98A-B4A18304A427" family="Gobiidae" genus="Gobioides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Gobioides Lacépède 1800:576" order="Perciformes" pageNumber="1" phylum="Chordata" rank="genus">Gobioides</taxonomicName>
based on its vertebral count of 26 (an incorrect count) and anal-fin count of 15.
</paragraph>
<paragraph id="EAAC1D0D56927F5B35DA2549A2DEEEFF" pageNumber="2">
Murdy (1998) reviewed
<taxonomicName id="D1BB6D17D3F12C96BF692201F5058F57" LSID-ZBK="urn:lsid:zoobank.org:act:BA04C1EE-19AC-4F48-B98A-B4A18304A427" family="Gobiidae" genus="Gobioides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Gobioides Lacépède 1800:576" order="Perciformes" pageNumber="2" phylum="Chordata" rank="genus">Gobioides</taxonomicName>
and noted that
<taxonomicName id="AC149E674FAF52B8857A8A78CAC646AB" LSID-ZBK="urn:lsid:zoobank.org:act:1D669904-3B1B-4EB8-99A5-493BE7889CCA" family="Gobiidae" genus="Amblyopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Amblyopus brevis Günther 1864:151" order="Perciformes" pageNumber="2" phylum="Chordata" rank="species" species="brevis">
Amblyopus brevis
<normalizedToken id="A1B50687FC5D72E063DA128DBD501AE4" originalValue="Günther">Guenther</normalizedToken>
</taxonomicName>
does not belong in
<taxonomicName id="43195CCE8D50AB3E5E4D1FE25E6B39EC" LSID-ZBK="urn:lsid:zoobank.org:act:BA04C1EE-19AC-4F48-B98A-B4A18304A427" family="Gobiidae" genus="Gobioides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Gobioides Lacépède 1800:576" order="Perciformes" pageNumber="2" phylum="Chordata" rank="genus">Gobioides</taxonomicName>
based on its dorsal pterygiophore pattern of 3-221110 and a vertebral count of 11+16 (not 26 as given by Palmer [1952]). Those characters would place it in the “
<taxonomicName id="5230BEB66F636EE7DCA85B29B47DDB87" family="Gobiidae" genus="Gobiosoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="2" phylum="Chordata" rank="genus">Gobiosoma</taxonomicName>
” or “
<taxonomicName id="F3D7BA60EE7601F85BEA6810F7CF900F" LSID-ZBK="urn:lsid:zoobank.org:act:44046A17-7637-4AF9-9121-F993672528FB" family="Gobiidae" genus="Microgobius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Microgobius Bleeker 1931:101" order="Perciformes" pageNumber="2" phylum="Chordata" rank="genus">Microgobius</taxonomicName>
” groups of the Gobiosomatini and thus part of the Gobiinae (sensu Pezold, 1993).
</paragraph>
<paragraph id="72D4302D00B59B30DD61B4D86A11FD1D" pageNumber="2">
Birdsong and Robins (1995) erected
<taxonomicName id="FD5E1C82B0A44BDD9CC59284612B490A" LSID-ZBK="urn:lsid:zoobank.org:act:1654F3F7-4BF7-44EE-BD21-80A538552441" family="Gobiidae" genus="Akko" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Akko Birdsong &amp; Robins 1995:676" order="Perciformes" pageNumber="2" phylum="Chordata" rank="genus">Akko</taxonomicName>
for a new species,
<taxonomicName id="06199AB65EEE0F3BD66706BBDE4AB65F" LSID-ZBK="urn:lsid:zoobank.org:act:40FD9887-81A7-4035-B4B2-30CB36945D18" family="Gobiidae" genus="Akko" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Akko dionaea Birdsong &amp; Robins 1995:676" order="Perciformes" pageNumber="2" phylum="Chordata" rank="species" species="dionaea">A. dionaea [1]</taxonomicName>
, from off Brazil.
<taxonomicName id="31A7B21754F7A230A65A083E4A383021" LSID-ZBK="urn:lsid:zoobank.org:act:40FD9887-81A7-4035-B4B2-30CB36945D18" family="Gobiidae" genus="Akko" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Akko dionaea Birdsong &amp; Robins 1995:676" order="Perciformes" pageNumber="2" phylum="Chordata" rank="species" species="dionaea">Akko dionaea</taxonomicName>
has a dorsal pterygiophore pattern of 3-221110, 11+16 vertebrae, and a distinctive sensory papillae pattern in which horizontal row b is elongate, extending anteriorly to a vertical through the middle of the eye; dorsal row n is elongate, the left and right elements approaching or meeting at the dorsal midline; and rows x1 and x2 are continuous. Birdsong and Robins (1995) placed the genus in the Gobiosomatini but did not comment further on its relationships within the tribe because of the large number of osteological autapomorphies it possesses.
</paragraph>
<paragraph id="F9B3BE216130046949A2170F2FF3C806" pageNumber="2">
Fish collections made during a Smithsonian Institution cruise aboard the R/V Uracca to the Darien province of Panama in 2000 resulted in several specimens initially identified as
<taxonomicName id="6E8482CB7634C3E44A6761370E896789" family="Gobiidae" genus="Gobioides" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="2" phylum="Chordata" rank="species" species="brevis">Gobioides brevis</taxonomicName>
. A Smithsonian expedition to El Salvador in 2002 resulted in 144 additional specimens. This species has a dorsal pterygiophore pattern of 3-221110, 11 precaudal and 16 caudal vertebrae, and the distinct papillae pattern of and most of the osteological autapomorphies present in
<taxonomicName id="C00E3E0385A973D8602EE52E9B3834E9" LSID-ZBK="urn:lsid:zoobank.org:act:40FD9887-81A7-4035-B4B2-30CB36945D18" family="Gobiidae" genus="Akko" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Akko dionaea Birdsong &amp; Robins 1995:676" order="Perciformes" pageNumber="2" phylum="Chordata" rank="species" species="dionaea">Akko dionaea</taxonomicName>
. Several specimens were sent to the British Museum where Anthony Gill confirmed that our specimens match those of the holotype of
<taxonomicName id="7792474C0FDDEFFE0D1642EB0CD56D06" LSID-ZBK="urn:lsid:zoobank.org:act:1D669904-3B1B-4EB8-99A5-493BE7889CCA" family="Gobiidae" genus="Amblyopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Amblyopus brevis Günther 1864:151" order="Perciformes" pageNumber="2" phylum="Chordata" rank="species" species="brevis">Amblyopus brevis</taxonomicName>
. We therefore take the opportunity to redescribe
<taxonomicName id="8AB60E7A21314D4941132C944973B325" LSID-ZBK="urn:lsid:zoobank.org:act:1D669904-3B1B-4EB8-99A5-493BE7889CCA" family="Gobiidae" genus="Amblyopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Amblyopus brevis Günther 1864:151" order="Perciformes" pageNumber="2" phylum="Chordata" rank="species" species="brevis">Amblyopus brevis</taxonomicName>
based on the new material and place it in the genus
<taxonomicName id="B12B8A96B64198F58C35A7D60F501793" LSID-ZBK="urn:lsid:zoobank.org:act:1654F3F7-4BF7-44EE-BD21-80A538552441" family="Gobiidae" genus="Akko" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Akko Birdsong &amp; Robins 1995:676" order="Perciformes" pageNumber="2" phylum="Chordata" rank="genus">Akko</taxonomicName>
, thus extending the range of
<taxonomicName id="A5CF69EBBFD4C703A8F090DE72045638" LSID-ZBK="urn:lsid:zoobank.org:act:1654F3F7-4BF7-44EE-BD21-80A538552441" family="Gobiidae" genus="Akko" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Akko Birdsong &amp; Robins 1995:676" order="Perciformes" pageNumber="2" phylum="Chordata" rank="genus">Akko</taxonomicName>
from the Atlantic to the eastern Pacific Ocean.
</paragraph>
<paragraph id="14FC91015002D98EAFDF64713CE52BBD" pageNumber="2">
In addition, a previously undescribed gobiid species obtained during the El Salvador expedition appears to represent a new species of
<taxonomicName id="7484CFCB366546B042B4F2CF4C1EF22A" LSID-ZBK="urn:lsid:zoobank.org:act:1654F3F7-4BF7-44EE-BD21-80A538552441" family="Gobiidae" genus="Akko" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Akko Birdsong &amp; Robins 1995:676" order="Perciformes" pageNumber="2" phylum="Chordata" rank="genus">Akko</taxonomicName>
. This species, which is larger than
<taxonomicName id="F3C69B625DE24FE1294E1A564BEC368E" LSID-ZBK="urn:lsid:zoobank.org:act:1D669904-3B1B-4EB8-99A5-493BE7889CCA" family="Gobiidae" genus="Amblyopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Amblyopus brevis Günther 1864:151" order="Perciformes" pageNumber="2" phylum="Chordata" rank="species" species="brevis">A. brevis</taxonomicName>
, found in shallower waters (7-9 meters as opposed to 15-25 meters for
<taxonomicName id="8E7D8C59E5156344414C6E4F519CB7A9" LSID-ZBK="urn:lsid:zoobank.org:act:1D669904-3B1B-4EB8-99A5-493BE7889CCA" family="Gobiidae" genus="Amblyopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Amblyopus brevis Günther 1864:151" order="Perciformes" pageNumber="2" phylum="Chordata" rank="species" species="brevis">A. brevis</taxonomicName>
), and separable from
<taxonomicName id="C9B1C396AA627076B1CE077CCCB69FA3" LSID-ZBK="urn:lsid:zoobank.org:act:1D669904-3B1B-4EB8-99A5-493BE7889CCA" family="Gobiidae" genus="Amblyopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Amblyopus brevis Günther 1864:151" order="Perciformes" pageNumber="2" phylum="Chordata" rank="species" species="brevis">A. brevis</taxonomicName>
on the basis of numbers of lateral-line scales, vertebrae, dorsal- and anal-fin rays, and pigment pattern, is described herein.
</paragraph>
<paragraph id="3BD81E399C7BB9A660DAA1ACECF6B0B4" pageNumber="2">[1] Birdsong and Robins (1995) spelled the name of their new species “dionea” in the abstract but “dionaea ” in all other places in the paper. The species was named after the plant genus Dionaea. We conclude that “dionea” is a misspelling.</paragraph>
</subSubSection>
</treatment>
</document>
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