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<document id="A8AAAEBA7EC2F4CC20AEB55718F9617B" ID-DOI="10.1649/611" ID-ISSN="1938-4394" ID-Zenodo-Dep="10104359" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630168753109" checkinUser="tatiana" docAuthor="Sokolov, Igor M., Carlton, Christopher &amp; Cornell, James F." docDate="2004" docId="845C87831250FE115EA1FEAAFC63F9C2" docLanguage="en" docName="611.pdf.imf" docOrigin="The Coleopterists Bulletin 58 (2)" docSource="http://dx.doi.org/10.1649/611" docStyle="DocumentStyle:07711681861DB98305E87147CAB6DAD5.4:TheColeopBull.2003-2009.journal_article.0cover" docStyleId="07711681861DB98305E87147CAB6DAD5" docStyleName="TheColeopBull.2003-2009.journal_article.0cover" docStyleVersion="4" docTitle="Anillinus fortis" docType="treatment" docVersion="4" lastPageNumber="201" masterDocId="7865FFFB1240FE015F43FFFDFFABFFE0" masterDocTitle="Review of Anillinus, with Descriptions of 17 New Species and a Key to Soil and Litter Species (Coleoptera: Carabidae: Trechinae: Bembidiini)" masterLastPageNumber="233" masterPageNumber="185" pageNumber="201" updateTime="1699629423378" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="C45227A1656B50A8415E31213B9C3187">Review of Anillinus, with Descriptions of 17 New Species and a Key to Soil and Litter Species (Coleoptera: Carabidae: Trechinae: Bembidiini)</mods:title>
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<mods:namePart id="AF558769B187E9C26D557E7B14D3402D">Sokolov, Igor M.</mods:namePart>
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<mods:namePart id="8A346A5A203C35CBFD0D54EAB32CCA4A">Carlton, Christopher</mods:namePart>
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<mods:namePart id="78A358297490F8AEF004B3930D096C0C">Cornell, James F.</mods:namePart>
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<mods:title id="9288D49F0CE6B82D30A4F87D2D100E27">The Coleopterists Bulletin</mods:title>
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<mods:date id="6C6491E5B63BBCBB669949A3605CE1DE">2004</mods:date>
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<mods:identifier id="EE347076E54671761656725B4F54E790" type="DOI">10.1649/611</mods:identifier>
<mods:identifier id="03FE4973480724DEB7B8B51CE7D77E99" type="ISSN">1938-4394</mods:identifier>
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<treatment id="845C87831250FE115EA1FEAAFC63F9C2" ID-DOI="http://doi.org/10.5281/zenodo.5398715" ID-Zenodo-Dep="5398715" LSID="urn:lsid:plazi:treatment:845C87831250FE115EA1FEAAFC63F9C2" httpUri="http://treatment.plazi.org/id/845C87831250FE115EA1FEAAFC63F9C2" lastPageNumber="201" pageId="16" pageNumber="201">
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<taxonomicName id="CBF54D161250FE115EA1FEAAFD12FE8C" ID-CoL="5V28F" authority="(Horn)" baseAuthorityName="Horn" box="[482,697,343,364]" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="201" phylum="Arthropoda" rank="species" species="fortis">
<emphasis id="3E81EA871250FE115EA1FEAAFDDAFE8C" box="[482,625,343,364]" italics="true" pageId="16" pageNumber="201">Anillinus fortis</emphasis>
(Horn)
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</heading>
</paragraph>
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<subSubSection id="44EF651E1250FE115ED1FE8FFCA1FE67" box="[402,778,370,391]" pageId="16" pageNumber="201" type="description">
<paragraph id="0C4A36951250FE115ED1FE8FFCA1FE67" blockId="16.[402,778,343,391]" box="[402,778,370,391]" pageId="16" pageNumber="201">
(
<figureCitation id="94CE2A101250FE115EDAFE8FFE5CFE67" box="[409,503,370,391]" captionStart="Figs" captionStartId="5.[169,208,1067,1086]" captionTargetBox="[151,974,194,1045]" captionTargetId="figure-44@5.[151,975,194,1045]" captionTargetPageId="5" captionText="Figs. 1–5. Anillinus spp. 1) Anillinus fortis (Horn), dorsal aspect, scale ¼ 1.0 mm. 2–7) Dorsal aspects of heads, showing microsculpture, scale ¼ 0.2 mm. 2) Anillinus magazinensis n. sp.; 3) A. fortis (Horn); 4) A. robisoni n. sp.; 5) A. erwini n. sp." figureDoi="http://doi.org/10.5281/zenodo.10104361" httpUri="https://zenodo.org/record/10104361/files/figure.png" pageId="16" pageNumber="201">Figs. 1, 3</figureCitation>
,
<figureCitation id="94CE2A101250FE115D47FE8FFDB9FE67" box="[516,530,370,391]" captionStart="Figs" captionStartId="6.[197,236,941,960]" captionTargetBox="[177,1000,194,920]" captionTargetId="figure-348@6.[177,1001,194,920]" captionTargetPageId="6" captionText="Figs. 6–9. Pronota of Anillinus spp., scale ¼ 0.3 mm. 6) Anillinus erwini n. sp.; 7) A. fortis" figureDoi="http://doi.org/10.5281/zenodo.10104363" httpUri="https://zenodo.org/record/10104363/files/figure.png" pageId="16" pageNumber="201">7</figureCitation>
,
<figureCitation id="94CE2A101250FE115D5DFE8FFD9CFE67" box="[542,567,370,391]" captionStart="Figs" captionStartId="7.[169,208,1334,1353]" captionText="Figs. 10–13. Anillinus spp., right metafemora, setae omitted. 10) A. elongatus Jeannel;" pageId="16" pageNumber="201">11</figureCitation>
,
<figureCitation id="94CE2A101250FE115D07FE8FFDF5FE67" box="[580,606,370,391]" captionStart="Figs" captionStartId="7.[169,208,1404,1423]" captionTargetBox="[160,966,196,1313]" captionTargetId="figure-1@7.[159,966,194,1313]" captionTargetPageId="7" captionText="Figs. 14–17. Anillinus spp., right metatibiae, setae and spurs omitted. 14) A. elongatus Jeannel; 15) A. lescheni n. sp.; 16) A. fortis (Horn); 17) A. stephani n. sp." figureDoi="http://doi.org/10.5281/zenodo.10104366" httpUri="https://zenodo.org/record/10104366/files/figure.png" pageId="16" pageNumber="201">16</figureCitation>
,
<figureCitation id="94CE2A101250FE115D29FE8FFD2FFE67" box="[618,644,370,391]" captionStart="Figs" captionStartId="8.[197,236,1197,1216]" captionTargetBox="[180,1001,196,1175]" captionTargetId="figure-191@8.[178,1002,194,1176]" captionTargetPageId="8" captionText="Figs. 18–24. Aedeagi of Anillinus spp., right lateral aspects, scale ¼ 0.1 mm. 18) A. elongatus Jeannel; 19) A. fortis (Horn); 20) A. steevesi Barr; 21) A. balli n. sp.; 22) A. barri n. sp.; 23) A. cornelli n. sp.; 24) A. daggyi n. sp." figureDoi="http://doi.org/10.5281/zenodo.10104368" httpUri="https://zenodo.org/record/10104368/files/figure.png" pageId="16" pageNumber="201">19</figureCitation>
, Map fig. 39)
</paragraph>
</subSubSection>
<subSubSection id="44EF651E1250FE115FECFE67FC1CFDF9" pageId="16" pageNumber="201" type="reference_group">
<paragraph id="0C4A36951250FE115FECFE67FC7AFE29" blockId="16.[175,1004,410,537]" pageId="16" pageNumber="201">
<treatmentCitationGroup id="2CE511BB1250FE115FECFE67FC7AFE29" pageId="16" pageNumber="201">
<treatmentCitation id="8D5410841250FE115FECFE67FE7CFE4F" author="Horn" box="[175,471,410,431]" page="127" pageId="16" pageNumber="201" year="1868">
<taxonomicName id="CBF54D161250FE115FECFE67FE7CFE4F" ID-CoL="5V28W" authority="Horn, 1868: 127" authorityName="Horn" authorityPageNumber="127" authorityYear="1868" box="[175,471,410,431]" class="Insecta" family="Carabidae" genus="Anillus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="201" phylum="Arthropoda" rank="species" species="fortis">
<emphasis id="3E81EA871250FE115FECFE67FE84FE4F" box="[175,303,410,431]" italics="true" pageId="16" pageNumber="201">Anillus fortis</emphasis>
<materialsCitation id="BC9D3CC81250FE115E7AFE67FE7CFE4F" box="[313,471,410,431]" county="Eastern" location="Philadelphia" municipality="Deposited Academy of Natural Sciences" pageId="16" pageNumber="201" specimenCount="1" stateProvince="Tennessee">
<bibRefCitation id="68644B641250FE115E7AFE67FE7CFE4F" author="Horn" box="[313,471,410,431]" pageId="16" pageNumber="201" pagination="123 - 128" refId="ref32763" refString="Horn, G. H. 1868. Catalogue of Coleoptera from south Western Virginia. Transactions of the American Entomological Society 2 (1868 - 1869): 123 - 128." type="journal article" year="1868">Horn, 1868:127</bibRefCitation>
</materialsCitation>
</taxonomicName>
</treatmentCitation>
.
<typeStatus id="D34E88371250FE115EA7FE67FDBEFE4F" box="[484,533,410,431]" pageId="16" pageNumber="201">Type</typeStatus>
(sex unknown) from ‘‘mountainous regions of
<collectingCounty id="E52B4E191250FE115FACFE49FE9DFE29" box="[239,310,436,457]" pageId="16" pageNumber="201">Eastern</collectingCounty>
<collectingRegion id="CE31F8771250FE115E7DFE49FE01FE29" box="[318,426,436,457]" country="United States of America" name="Tennessee" pageId="16" pageNumber="201">Tennessee</collectingRegion>
.’’
<collectingMunicipality id="EC2EACEF1250FE115EFDFE49FCE3FE29" box="[446,840,436,457]" pageId="16" pageNumber="201">Deposited Academy of Natural Sciences</collectingMunicipality>
,
<location id="092A604E1250FE115C17FE49FC66FE29" LSID="urn:lsid:plazi:treatment:845C87831250FE115EA1FEAAFC63F9C2:092A604E1250FE115C17FE49FC66FE29" box="[852,973,436,457]" county="Eastern" municipality="Deposited Academy of Natural Sciences" name="Philadelphia" pageId="16" pageNumber="201" stateProvince="Tennessee">Philadelphia</location>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="0C4A36951250FE115FECFE32FDEFFE04" blockId="16.[175,1004,410,537]" box="[175,580,463,484]" pageId="16" pageNumber="201">
<treatmentCitationGroup id="2CE511BB1250FE115FECFE32FDEFFE04" box="[175,580,463,484]" pageId="16" pageNumber="201">
<taxonomicName id="CBF54D161250FE115FECFE32FE22FE04" ID-CoL="5V28F" authority="(Horn)" baseAuthorityName="Horn" box="[175,393,463,484]" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="201" phylum="Arthropoda" rank="species" species="fortis">
<emphasis id="3E81EA871250FE115FECFE32FE94FE04" box="[175,319,463,484]" italics="true" pageId="16" pageNumber="201">Anillinus fortis</emphasis>
(Horn)
</taxonomicName>
,
<treatmentCitation id="8D5410841250FE115ED6FE32FDEBFE04" author="Jeannel" box="[405,576,463,484]" page="75" pageId="16" pageNumber="201" year="1963">
<bibRefCitation id="68644B641250FE115ED6FE32FDEBFE04" author="Jeannel" box="[405,576,463,484]" pageId="16" pageNumber="201" pagination="33 - 204" refId="ref32919" refString="Jeannel, R. 1963 a. Monographie des '' Anillini' ', Bembidiides endoges [Coleoptera Trechidae]. Memoires du Museum National d'Histoire Naturelle, Serie A, Zoologie 28 (2): 33 - 204." type="journal article" year="1963">
Jeannel 1963
<emphasis id="3E81EA871250FE115D50FE32FDB4FE04" box="[531,543,463,484]" italics="true" pageId="16" pageNumber="201">a</emphasis>
:75
</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="0C4A36951250FE115FECFE14FE74FE1E" blockId="16.[175,1004,410,537]" box="[175,479,489,510]" pageId="16" pageNumber="201">
<treatmentCitationGroup id="2CE511BB1250FE115FECFE14FE74FE1E" box="[175,479,489,510]" pageId="16" pageNumber="201">
<treatmentCitation id="8D5410841250FE115FECFE14FE70FE1E" author="Barr" box="[175,475,489,510]" page="241" pageId="16" pageNumber="201" year="1995">
<taxonomicName id="CBF54D161250FE115FECFE14FE70FE1E" ID-CoL="5V28F" authority=", Barr 1995: 241" authorityName=", Barr" authorityPageNumber="241" authorityYear="1995" box="[175,475,489,510]" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="201" phylum="Arthropoda" rank="species" species="fortis">
<emphasis id="3E81EA871250FE115FECFE14FE94FE1E" box="[175,319,489,510]" italics="true" pageId="16" pageNumber="201">Anillinus fortis</emphasis>
,
<bibRefCitation id="68644B641250FE115E0FFE14FE70FE1E" author="Barr" box="[332,475,489,510]" pageId="16" pageNumber="201" pagination="239 - 248" refId="ref31942" refString="Barr, T. C. 1995. Notes on some anillines (Coleoptera, Carabidae, Bembidiinae) from southeastern United States, with description of a new genus and two new species. Special Bulletin Japanese Society of Coleopterology, Tokyo 4: 239 - 248." type="journal article" year="1995">Barr 1995:241</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="0C4A36951250FE115FECFDF9FC1CFDF9" blockId="16.[175,1004,410,537]" box="[175,951,516,537]" pageId="16" pageNumber="201">
<treatmentCitationGroup id="2CE511BB1250FE115FECFDF9FC1CFDF9" box="[175,951,516,537]" pageId="16" pageNumber="201">
<treatmentCitation id="8D5410841250FE115FECFDF9FDC4FDF9" author="Casey" box="[175,623,516,537]" page="168" pageId="16" pageNumber="201" year="1918">
<taxonomicName id="CBF54D161250FE115FECFDF9FDC4FDF9" authority="Casey, 1918: 168" authorityName="Casey" authorityPageNumber="168" authorityYear="1918" box="[175,623,516,537]" class="Insecta" family="Carabidae" genus="Anillus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="201" phylum="Arthropoda" rank="species" species="carolinae" subGenus="Anillinus">
<emphasis id="3E81EA871250FE115FECFDF9FF59FDF9" box="[175,242,516,537]" italics="true" pageId="16" pageNumber="201">Anillus</emphasis>
(
<emphasis id="3E81EA871250FE115E41FDF9FEFCFDF9" box="[258,343,516,537]" italics="true" pageId="16" pageNumber="201">Anillinus</emphasis>
)
<emphasis id="3E81EA871250FE115E24FDF9FE69FDF9" box="[359,450,516,537]" italics="true" pageId="16" pageNumber="201">carolinae</emphasis>
<bibRefCitation id="68644B641250FE115E89FDF9FDC4FDF9" author="Casey" box="[458,623,516,537]" pageId="16" pageNumber="201" refId="ref32208" refString="Casey, T. L. 1918. Memoirs on the Coleoptera. VIII. The New Era Printing Company, Lancaster PA. 427 pp." type="book" year="1918">Casey, 1918:168</bibRefCitation>
</taxonomicName>
</treatmentCitation>
. Synonymized by
<treatmentCitation id="8D5410841250FE115C66FDF9FC18FDF9" author="Barr" box="[805,947,516,537]" page="241" pageId="16" pageNumber="201" year="1995">
<bibRefCitation id="68644B641250FE115C66FDF9FC18FDF9" author="Barr" box="[805,947,516,537]" pageId="16" pageNumber="201" pagination="239 - 248" refId="ref31942" refString="Barr, T. C. 1995. Notes on some anillines (Coleoptera, Carabidae, Bembidiinae) from southeastern United States, with description of a new genus and two new species. Special Bulletin Japanese Society of Coleopterology, Tokyo 4: 239 - 248." type="journal article" year="1995">Barr 1995:241</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="44EF651E1250FE115F84FDD1FC46FD96" pageId="16" pageNumber="201" type="materials_examined">
<paragraph id="0C4A36951250FE115F84FDD1FC46FD96" blockId="16.[175,1005,556,630]" pageId="16" pageNumber="201">
<typeStatus id="D34E88371250FE115F84FDD1FEB4FDA1" box="[199,287,556,577]" pageId="16" pageNumber="201" type="holotype">Holotype</typeStatus>
not examined. Our interpretation of the species is based on examination of the cotype of
<taxonomicName id="CBF54D161250FE115E77FDBBFE00FDBB" authorityName="Casey" authorityYear="1918" box="[308,427,582,603]" class="Insecta" family="Carabidae" genus="Anillus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="201" phylum="Arthropoda" rank="species" species="carolinae">
<emphasis id="3E81EA871250FE115E77FDBBFE00FDBB" box="[308,427,582,603]" italics="true" pageId="16" pageNumber="201">A. carolinae</emphasis>
</taxonomicName>
(female bearing labels: ‘‘Black Mts. N.C. IX.’’ and ‘‘fortis, compared with type’’). We are in agreement with
<bibRefCitation id="68644B641250FE115DC9FD9CFCA1FD96" author="Barr" box="[650,778,609,630]" pageId="16" pageNumber="201" pagination="239 - 248" refId="ref31942" refString="Barr, T. C. 1995. Notes on some anillines (Coleoptera, Carabidae, Bembidiinae) from southeastern United States, with description of a new genus and two new species. Special Bulletin Japanese Society of Coleopterology, Tokyo 4: 239 - 248." type="journal article" year="1995">Barr’s (1995)</bibRefCitation>
concept of this species.
</paragraph>
</subSubSection>
<subSubSection id="44EF651E1250FE115F86FD77FD56FBFC" pageId="16" pageNumber="201" type="description">
<paragraph id="0C4A36951250FE115F86FD77FD1CFD1D" blockId="16.[175,1005,650,1052]" pageId="16" pageNumber="201">
<emphasis id="3E81EA871250FE115F86FD77FE98FD7D" bold="true" box="[197,307,650,669]" pageId="16" pageNumber="201">Description.</emphasis>
Medium to large-sized for the genus (ABL range,
<quantity id="CB0D9B701250FE115DA5FD76FCCFFD7E" box="[742,868,651,670]" metricMagnitude="-3" metricUnit="m" metricValue="1.89" metricValueMax="2.08" metricValueMin="1.7" pageId="16" pageNumber="201" unit="mm" value="1.89" valueMax="2.08" valueMin="1.7">1.70–2.08 mm</quantity>
; mean,
<geoCoordinate id="69C150521250FE115CE6FD76FC70FD7E" box="[933,987,651,670]" degrees="1.96" direction="north" orientation="latitude" pageId="16" pageNumber="201" precision="555" value="1.96">1.96 n</geoCoordinate>
<emphasis id="3E81EA871250FE115C9DFD77FC47FD7D" bold="true" box="[990,1004,650,669]" pageId="16" pageNumber="201">¼</emphasis>
10). Habitus (
<figureCitation id="94CE2A101250FE115E6AFD5FFEF7FD55" box="[297,348,674,693]" captionStart="Figs" captionStartId="5.[169,208,1067,1086]" captionTargetBox="[151,974,194,1045]" captionTargetId="figure-44@5.[151,975,194,1045]" captionTargetPageId="5" captionText="Figs. 1–5. Anillinus spp. 1) Anillinus fortis (Horn), dorsal aspect, scale ¼ 1.0 mm. 2–7) Dorsal aspects of heads, showing microsculpture, scale ¼ 0.2 mm. 2) Anillinus magazinensis n. sp.; 3) A. fortis (Horn); 4) A. robisoni n. sp.; 5) A. erwini n. sp." figureDoi="http://doi.org/10.5281/zenodo.10104361" httpUri="https://zenodo.org/record/10104361/files/figure.png" pageId="16" pageNumber="201">Fig. 1</figureCitation>
) markedly convex, ovoid, head of moderate size (WH/WPm, 0.73), elytra relatively wide (WPm/WE, 0.78). Color of body various, from brunneous to brunneorufous, appendages testaceous. Pronotal microsculpture fine but distinct across entire disc. Microsculpture on head absent on two paralateral patches at vertex (
<figureCitation id="94CE2A101250FE115D35FD17FD00FD1D" box="[630,683,746,765]" captionStart="Figs" captionStartId="5.[169,208,1067,1086]" captionTargetBox="[151,974,194,1045]" captionTargetId="figure-44@5.[151,975,194,1045]" captionTargetPageId="5" captionText="Figs. 1–5. Anillinus spp. 1) Anillinus fortis (Horn), dorsal aspect, scale ¼ 1.0 mm. 2–7) Dorsal aspects of heads, showing microsculpture, scale ¼ 0.2 mm. 2) Anillinus magazinensis n. sp.; 3) A. fortis (Horn); 4) A. robisoni n. sp.; 5) A. erwini n. sp." figureDoi="http://doi.org/10.5281/zenodo.10104361" httpUri="https://zenodo.org/record/10104361/files/figure.png" pageId="16" pageNumber="201">Fig. 3</figureCitation>
).
</paragraph>
<paragraph id="0C4A36951250FE115F86FCFFFD27FCBD" blockId="16.[175,1005,650,1052]" pageId="16" pageNumber="201">
Pronotum moderately convex and transverse (WPm/LP, 1.30), with margins slightly constricted and rectilinear convergent posteriad (WPm/WPp, 1.24) (
<figureCitation id="94CE2A101250FE115DD0FCE7FD63FCCD" box="[659,712,794,813]" captionStart="Figs" captionStartId="6.[197,236,941,960]" captionTargetBox="[177,1000,194,920]" captionTargetId="figure-348@6.[177,1001,194,920]" captionTargetPageId="6" captionText="Figs. 6–9. Pronota of Anillinus spp., scale ¼ 0.3 mm. 6) Anillinus erwini n. sp.; 7) A. fortis" figureDoi="http://doi.org/10.5281/zenodo.10104363" httpUri="https://zenodo.org/record/10104363/files/figure.png" pageId="16" pageNumber="201">Fig. 7</figureCitation>
). Anterior angles evident, slightly prominent. Posterior angles slightly obtuse (105–1158). Width between posterior angles much greater than between anterior angles (WPa/WPp, 0.92).
</paragraph>
<paragraph id="0C4A36951250FE115F86FC9FFE7FFC45" blockId="16.[175,1005,650,1052]" pageId="16" pageNumber="201">Elytra markedly convex, slightly depressed along suture, length normal for genus (LE/ABL, 0.57), with traces of 1–3 (occasionally 4) interneurs. Humeri moderately prominent, slightly rounded. Vestiture of elytra short.</paragraph>
<paragraph id="0C4A36951250FE115F86FC54FD56FC5C" blockId="16.[175,1005,650,1052]" box="[197,765,937,956]" pageId="16" pageNumber="201">
Males with metafemora (
<figureCitation id="94CE2A101250FE115EDFFC54FE71FC5C" box="[412,474,937,956]" captionStart="Figs" captionStartId="7.[169,208,1334,1353]" captionText="Figs. 10–13. Anillinus spp., right metafemora, setae omitted. 10) A. elongatus Jeannel;" pageId="16" pageNumber="201">Fig. 11</figureCitation>
) and metatibiae (
<figureCitation id="94CE2A101250FE115D31FC54FD1AFC5C" box="[626,689,937,956]" captionStart="Figs" captionStartId="7.[169,208,1404,1423]" captionTargetBox="[160,966,196,1313]" captionTargetId="figure-1@7.[159,966,194,1313]" captionTargetPageId="7" captionText="Figs. 14–17. Anillinus spp., right metatibiae, setae and spurs omitted. 14) A. elongatus Jeannel; 15) A. lescheni n. sp.; 16) A. fortis (Horn); 17) A. stephani n. sp." figureDoi="http://doi.org/10.5281/zenodo.10104366" httpUri="https://zenodo.org/record/10104366/files/figure.png" pageId="16" pageNumber="201">Fig. 16</figureCitation>
) simple.
</paragraph>
<paragraph id="0C4A36951250FE115F86FC3CFD56FBFC" blockId="16.[175,1005,650,1052]" pageId="16" pageNumber="201">
Median lobe (
<figureCitation id="94CE2A101250FE115E7EFC3CFED7FC34" box="[317,380,961,980]" captionStart="Figs" captionStartId="8.[197,236,1197,1216]" captionTargetBox="[180,1001,196,1175]" captionTargetId="figure-191@8.[178,1002,194,1176]" captionTargetPageId="8" captionText="Figs. 18–24. Aedeagi of Anillinus spp., right lateral aspects, scale ¼ 0.1 mm. 18) A. elongatus Jeannel; 19) A. fortis (Horn); 20) A. steevesi Barr; 21) A. balli n. sp.; 22) A. barri n. sp.; 23) A. cornelli n. sp.; 24) A. daggyi n. sp." figureDoi="http://doi.org/10.5281/zenodo.10104368" httpUri="https://zenodo.org/record/10104368/files/figure.png" pageId="16" pageNumber="201">Fig. 19</figureCitation>
) evenly arcuate, with small protrusion at dorsal margin of apex. Internal sac with copulatory pieces of spirally-curved dorsal sclerites, in form of a blade-like structure with a peculiar prominence near base; ventral sclerite of internal sac absent; small oval cluster of spines often visible at base of copulatory sclerite under high magnification.
</paragraph>
</subSubSection>
<subSubSection id="44EF651E1250FE115F84FBD2FC80FB29" pageId="16" pageNumber="201" type="materials_examined">
<paragraph id="0C4A36951250FE115F84FBD2FC80FB29" blockId="16.[175,1005,1071,1570]" pageId="16" pageNumber="201">
<emphasis id="3E81EA871250FE115F84FBD2FE24FBA4" bold="true" box="[199,399,1071,1092]" pageId="16" pageNumber="201">Material Examined</emphasis>
(39).
<materialsCitation id="BC9D3CC81250FE115E8EFBD2FD5DFBA4" box="[461,758,1071,1092]" collectionCode="NC" location="Twelve" pageId="16" pageNumber="201" specimenCount="1">
<location id="092A604E1250FE115E8EFBD2FDBFFBA4" LSID="urn:lsid:plazi:treatment:845C87831250FE115EA1FEAAFC63F9C2:092A604E1250FE115E8EFBD2FDBFFBA4" box="[461,532,1071,1092]" name="Twelve" pageId="16" pageNumber="201">Twelve</location>
specimens labeled /
<collectionCode id="6AE4AE501250FE115D97FBD2FD5DFBA4" box="[724,758,1071,1092]" pageId="16" pageNumber="201">NC</collectionCode>
</materialsCitation>
,
<materialsCitation id="BC9D3CC81250FE115C43FBD2FDFCFBBF" collectingDate="1996-08-17" collectionCode="JFC" collectorName="J. F. Cornell" county="Avery Co." location="Globe" pageId="16" pageNumber="201" specimenCount="3">
<collectingCounty id="E52B4E191250FE115C43FBD2FCCDFBA4" box="[768,870,1071,1092]" pageId="16" pageNumber="201">Avery Co.</collectingCounty>
, nr.
<location id="092A604E1250FE115CCDFBD2FC61FBA4" LSID="urn:lsid:plazi:treatment:845C87831250FE115EA1FEAAFC63F9C2:092A604E1250FE115CCDFBD2FC61FBA4" box="[910,970,1071,1092]" county="Avery Co." name="Globe" pageId="16" pageNumber="201">Globe</location>
,
<date id="784B10551250FE115C97FBD2FEBAFBBF" pageId="16" pageNumber="201" value="1996-08-17">
<collectingDate id="680FE9BD1250FE115C97FBD2FEBAFBBF" pageId="16" pageNumber="201" value="1996-08-17">17 Aug 1996</collectingDate>
</date>
,
<collectorName id="A10053431250FE115E58FBB7FE3AFBBF" box="[283,401,1098,1119]" pageId="16" pageNumber="201">J. F. Cornell</collectorName>
/ (
<collectionCode id="6AE4AE501250FE115EE5FBB7FE7AFBBF" box="[422,465,1098,1119]" pageId="16" pageNumber="201">JFC</collectionCode>
);
<specimenCount id="1AF3FD1C1250FE115EA1FBB7FDFCFBBF" box="[482,599,1098,1119]" pageId="16" pageNumber="201" type="generic">3 specimens</specimenCount>
</materialsCitation>
labeled /
<collectionCode id="6AE4AE501250FE115DECFBB7FD7AFBBF" box="[687,721,1098,1119]" pageId="16" pageNumber="201">NC</collectionCode>
,
<materialsCitation id="BC9D3CC81250FE115D9FFBB7FD2AFB99" collectingDate="1996-08-17" collectionCode="JFC" collectorName="J. F. Cornell" county="Caldwell Co." location="Edgemont" pageId="16" pageNumber="201" specimenCount="17">
<collectingCounty id="E52B4E191250FE115D9FFBB7FCF7FBBF" box="[732,860,1098,1119]" pageId="16" pageNumber="201">Caldwell Co.</collectingCounty>
,
<location id="092A604E1250FE115C25FBB7FC61FBBF" LSID="urn:lsid:plazi:treatment:845C87831250FE115EA1FEAAFC63F9C2:092A604E1250FE115C25FBB7FC61FBBF" box="[870,970,1098,1119]" county="Caldwell Co." name="Edgemont" pageId="16" pageNumber="201">Edgemont</location>
,
<date id="784B10551250FE115C97FBB7FEBEFB99" pageId="16" pageNumber="201" value="1996-08-17">
<collectingDate id="680FE9BD1250FE115C97FBB7FEBEFB99" pageId="16" pageNumber="201" value="1996-08-17">17 Aug 1996</collectingDate>
</date>
,
<collectorName id="A10053431250FE115E67FB99FE09FB99" box="[292,418,1124,1145]" pageId="16" pageNumber="201">J. F. Cornell</collectorName>
/ (
<collectionCode id="6AE4AE501250FE115EF8FB99FE4DFB99" box="[443,486,1124,1145]" pageId="16" pageNumber="201">JFC</collectionCode>
);
<specimenCount id="1AF3FD1C1250FE115EBFFB99FD2AFB99" box="[508,641,1124,1145]" pageId="16" pageNumber="201" type="generic">17 specimens</specimenCount>
</materialsCitation>
<materialsCitation id="BC9D3CC81250FE115DCFFB99FCEEFB99" box="[652,837,1124,1145]" collectionCode="NC" country="United States of America" location="USA" pageId="16" pageNumber="201" specimenCount="1">
labeled /
<collectingCountry id="74E276051250FE115DA1FB99FCBFFB99" box="[738,788,1124,1145]" name="United States of America" pageId="16" pageNumber="201">USA</collectingCountry>
:
<collectionCode id="6AE4AE501250FE115C61FB99FCEEFB99" box="[802,837,1124,1145]" pageId="16" pageNumber="201">NC</collectionCode>
</materialsCitation>
,
<materialsCitation id="BC9D3CC81250FE115C17FB99FD95FB74" collectionCode="R" collectorName="Deerlick Gap Overlook" country="United States of America" county="McDowell Co." location="McDowell Co." pageId="16" pageNumber="201" specimenCount="1">
<collectingCounty id="E52B4E191250FE115C17FB99FC42FB99" box="[852,1001,1124,1145]" pageId="16" pageNumber="201">McDowell Co.</collectingCounty>
,
<collectorName id="A10053431250FE115FECFB82FE33FB74" box="[175,408,1151,1172]" pageId="16" pageNumber="201">Deerlick Gap Overlook</collectorName>
, elv. 3,500, B.
<collectionCode id="6AE4AE501250FE115D6FFB82FD95FB74" box="[556,574,1151,1172]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="16" pageNumber="201">R</collectionCode>
</materialsCitation>
.
<materialsCitation id="BC9D3CC81250FE115D02FB82FC18FB74" box="[577,947,1151,1172]" collectionCode="VI" country="United States of America" location="Pkw." pageId="16" pageNumber="201" specimenCount="1">
<location id="092A604E1250FE115D02FB82FDDDFB74" LSID="urn:lsid:plazi:treatment:845C87831250FE115EA1FEAAFC63F9C2:092A604E1250FE115D02FB82FDDDFB74" box="[577,630,1151,1172]" country="United States of America" name="Pkw." pageId="16" pageNumber="201">Pkw.</location>
<quantity id="CB0D9B701250FE115D3CFB82FD6BFB74" box="[639,704,1151,1172]" metricMagnitude="3" metricUnit="m" metricValue="3.5405568" pageId="16" pageNumber="201" unit="mi" value="2.2">2.2 mi</quantity>
SW intersec. 221,
<date id="784B10551250FE115CC5FB82FC18FB74" box="[902,947,1151,1172]" pageId="16" pageNumber="201" value="1991-06-02">
2.
<collectionCode id="6AE4AE501250FE115CD5FB82FC18FB74" box="[918,947,1151,1172]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="16" pageNumber="201">VI</collectionCode>
</date>
</materialsCitation>
<materialsCitation id="BC9D3CC81250FE115CF1FB82FDA3FB4E" collectionCode="LSAM" collectorName="Carlton" country="United States of America" pageId="16" pageNumber="201" specimenCount="7">
.1991, C.E.
<collectorName id="A10053431250FE115F97FB64FE8BFB4E" box="[212,288,1177,1198]" pageId="16" pageNumber="201">Carlton</collectorName>
/ (
<collectionCode id="6AE4AE501250FE115E74FB64FED5FB4E" box="[311,382,1177,1198]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33726" lsid="urn:lsid:biocol.org:col:33726" name="Louisiana State Arthropod Museum" pageId="16" pageNumber="201">LSAM</collectionCode>
);
<specimenCount id="1AF3FD1C1250FE115ED2FB64FDA3FB4E" box="[401,520,1177,1198]" pageId="16" pageNumber="201" type="generic">7 specimens</specimenCount>
</materialsCitation>
labeled /
<collectionCode id="6AE4AE501250FE115D26FB64FD2CFB4E" box="[613,647,1177,1198]" pageId="16" pageNumber="201">NC</collectionCode>
,
<materialsCitation id="BC9D3CC81250FE115DD7FB64FEB9FB29" collectionCode="MP" collectorName="Green Knob" country="United States of America" county="McDowell Co." location="McDowell Co." pageId="16" pageNumber="201" specimenCode="MP 350.4" specimenCount="1">
<collectingCounty id="E52B4E191250FE115DD7FB64FC8CFB4E" box="[660,807,1177,1198]" pageId="16" pageNumber="201">McDowell Co.</collectingCounty>
,
<collectorName id="A10053431250FE115C70FB64FC07FB4E" box="[819,940,1177,1198]" pageId="16" pageNumber="201">Green Knob</collectorName>
, BRP.
<specimenCode id="5C539EEE1250FE115FECFB49FEB9FB29" box="[175,274,1204,1225]" collectionCode="MP" country="0" name="Mohonk Preserve, Inc." pageId="16" pageNumber="201">MP 350.4</specimenCode>
</materialsCitation>
,
<materialsCitation id="BC9D3CC81250FE115E5EFB49FC8CFB29" box="[285,807,1204,1225]" collectingDate="1991-08-07" collectionCode="JFC" collectorName="Cornell" country="United States of America" pageId="16" pageNumber="201" specimenCount="1">
<date id="784B10551250FE115E5EFB49FE3FFB29" box="[285,404,1204,1225]" pageId="16" pageNumber="201" value="1991-08-07">
<collectingDate id="680FE9BD1250FE115E5EFB49FE3FFB29" box="[285,404,1204,1225]" pageId="16" pageNumber="201" value="1991-08-07">7 Aug 1991</collectingDate>
</date>
, J. F.
<collectorName id="A10053431250FE115E91FB49FDB0FB29" box="[466,539,1204,1225]" pageId="16" pageNumber="201">Cornell</collectorName>
, rhododendron litter/ (
<collectionCode id="6AE4AE501250FE115DB6FB49FC8BFB29" box="[757,800,1204,1225]" pageId="16" pageNumber="201">JFC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="44EF651E1250FE115F84FB33FD95FAF9" pageId="16" pageNumber="201" type="distribution">
<paragraph id="0C4A36951250FE115F84FB33FC0BFB1E" blockId="16.[175,1005,1071,1570]" pageId="16" pageNumber="201">
<emphasis id="3E81EA871250FE115F84FB33FEE1FB03" bold="true" box="[199,330,1230,1251]" pageId="16" pageNumber="201">Distribution.</emphasis>
Known from north of the French Broad River in
<collectingRegion id="CE31F8771250FE115C6AFB32FC13FB04" box="[809,952,1231,1252]" country="United States of America" name="North Carolina" pageId="16" pageNumber="201">North Carolina</collectingRegion>
(
<figureCitation id="94CE2A101250FE115C85FB32FF63FB1E" captionStart="Fig" captionStartId="17.[169,200,845,864]" captionText="Fig. 39. Locality records for Anillinus spp." pageId="16" pageNumber="201">Fig. 39</figureCitation>
) and presumably eastern
<collectingRegion id="CE31F8771250FE115E81FB14FD83FB1E" box="[450,552,1257,1278]" country="United States of America" name="Tennessee" pageId="16" pageNumber="201">Tennessee</collectingRegion>
.
<collectingRegion id="CE31F8771250FE115D70FB14FD3DFB1E" box="[563,662,1257,1278]" country="United States of America" name="Tennessee" pageId="16" pageNumber="201">Tennessee</collectingRegion>
localities not corroborated.
</paragraph>
<paragraph id="0C4A36951250FE115F84FAFEFD95FAF9" blockId="16.[175,1005,1071,1570]" box="[199,574,1283,1305]" pageId="16" pageNumber="201">
<emphasis id="3E81EA871250FE115F84FAFEFEB7FAF8" bold="true" box="[199,284,1283,1304]" pageId="16" pageNumber="201">Habitat.</emphasis>
Deep, moist hardwood litter.
</paragraph>
</subSubSection>
<subSubSection id="44EF651E1250FE115F84FAE3FC63F9C2" pageId="16" pageNumber="201" type="diagnosis">
<paragraph id="0C4A36951250FE115F84FAE3FC63F9C2" blockId="16.[175,1005,1071,1570]" pageId="16" pageNumber="201">
<emphasis id="3E81EA871250FE115F84FAE3FE18FAD3" bold="true" box="[199,435,1310,1331]" pageId="16" pageNumber="201">Differential Diagnosis.</emphasis>
<taxonomicName id="CBF54D161250FE115EFCFAE3FDFFFAD3" authorityName=", Barr" authorityYear="1995" box="[447,596,1310,1331]" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="201" phylum="Arthropoda" rank="species" species="fortis">
<emphasis id="3E81EA871250FE115EFCFAE3FDFFFAD3" box="[447,596,1310,1331]" italics="true" pageId="16" pageNumber="201">Anillinus fortis</emphasis>
</taxonomicName>
is ovoid in form with patches lacking microsculpture on the lateral surfaces of the vertex (
<tableCitation id="4177032E1250FE115DE9FAC4FD5CFAAE" box="[682,759,1337,1358]" captionStart="Table 2" captionStartId="43.[167,219,627,646]" captionText="Table 2. Grouping of Anillinus species in accordance with their microsculpture patterns." pageId="16" pageNumber="201">Table 2</tableCitation>
, species group II of litter species). The median lobe is diagnostic. Based on similarities in the structure of the median lobe and the distribution,
<taxonomicName id="CBF54D161250FE115D50FA93FDCCFA63" authorityName=", Barr" authorityYear="1995" box="[531,615,1390,1411]" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="201" phylum="Arthropoda" rank="species" species="fortis">
<emphasis id="3E81EA871250FE115D50FA93FDCCFA63" box="[531,615,1390,1411]" italics="true" pageId="16" pageNumber="201">A. fortis</emphasis>
</taxonomicName>
may be the allopatric sister-taxon to
<taxonomicName id="CBF54D161250FE115FECFA74FEA5FA7E" box="[175,270,1417,1438]" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="201" phylum="Arthropoda" rank="species" species="loweae">
<emphasis id="3E81EA871250FE115FECFA74FEA5FA7E" box="[175,270,1417,1438]" italics="true" pageId="16" pageNumber="201">A. loweae</emphasis>
</taxonomicName>
, from which it differs in having a smaller apical process of the median lobe (larger in
<taxonomicName id="CBF54D161250FE115E4EFA5EFEC0FA58" box="[269,363,1443,1464]" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="201" phylum="Arthropoda" rank="species" species="loweae">
<emphasis id="3E81EA871250FE115E4EFA5EFEC0FA58" box="[269,363,1443,1464]" italics="true" pageId="16" pageNumber="201">A. loweae</emphasis>
</taxonomicName>
) and in the absence of a group of closed spines associated with the copulatory sclerites.
<taxonomicName id="CBF54D161250FE115E35FA43FDAFFA33" authorityName=", Barr" authorityYear="1995" box="[374,516,1470,1491]" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="201" phylum="Arthropoda" rank="species" species="fortis">
<emphasis id="3E81EA871250FE115E35FA43FDAFFA33" box="[374,516,1470,1491]" italics="true" pageId="16" pageNumber="201">Anillinus fortis</emphasis>
</taxonomicName>
is sympatric with at least two species of
<taxonomicName id="CBF54D161250FE115CD2FA43FC4DFA33" authorityName="Casey" authorityYear="1918" box="[913,998,1470,1491]" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="201" phylum="Arthropoda" rank="genus">
<emphasis id="3E81EA871250FE115CD2FA43FC4DFA33" box="[913,998,1470,1491]" italics="true" pageId="16" pageNumber="201">Anillinus</emphasis>
</taxonomicName>
, described herein as
<taxonomicName id="CBF54D161250FE115E36FA25FE64FA0D" box="[373,463,1496,1517]" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="201" phylum="Arthropoda" rank="species" species="erwini">
<emphasis id="3E81EA871250FE115E36FA25FE64FA0D" box="[373,463,1496,1517]" italics="true" pageId="16" pageNumber="201">A. erwini</emphasis>
</taxonomicName>
and
<taxonomicName id="CBF54D161250FE115D47FA25FDC8FA0D" box="[516,611,1496,1517]" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="201" phylum="Arthropoda" rank="species" species="daggyi">
<emphasis id="3E81EA871250FE115D47FA25FDC8FA0D" box="[516,611,1496,1517]" italics="true" pageId="16" pageNumber="201">A. daggyi</emphasis>
</taxonomicName>
. From the first species it differs by the presence of distinct microsculpture on the disc of the pronotum and from the second by its substantially larger size. All three may be separated using aedeagal characters.
</paragraph>
</subSubSection>
</treatment>
</document>
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