treatments-xml/data/80/3F/87/803F87FDC037FFD0FDF4AFE6FAA29D89.xml
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<document id="9EF8D8A06B5977F7E7C33C91C118159E" ID-CLB-Dataset="21840" ID-DOI="10.5852/ejt.2012.13" ID-GBIF-Dataset="9691a997-b6e4-4d47-9082-bbcba7347a3e" ID-ISSN="2118-9773" ID-Zenodo-Dep="3857775" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1590516738498" checkinUser="carolina" docAuthor="Ryvkin, Alexandr B." docDate="2012" docId="803F87FDC037FFD0FDF4AFE6FAA29D89" docLanguage="en" docName="ejt-2012-13.pdf.imf" docOrigin="European Journal of Taxonomy 13" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Stenus (Nestus) canosus Ryvkin 2012, sp. nov." docType="treatment" docUuid="06D9027A-5EDA-4C66-B54C-B7E7F9FD283A" docUuidSource="ZooBank" docVersion="12" lastPageNumber="30" masterDocId="7C06FF85C02CFFCDFFA1AD48FFD59832" masterDocTitle="New species and records of Stenus (Nestus) of the canaliculatus group, with the erection of a new species group (Insecta: Coleoptera: Staphylinidae: Steninae)" masterLastPageNumber="62" masterPageNumber="1" pageNumber="28" updateTime="1698841419189" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="FB6E565F897B19C3140594A02504AB21">New species and records of Stenus (Nestus) of the canaliculatus group, with the erection of a new species group (Insecta: Coleoptera: Staphylinidae: Steninae)</mods:title>
</mods:titleInfo>
<mods:name id="4103FAFDA818BB95AAB5F6FD134A09FD" type="personal">
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<mods:roleTerm id="823E39D2EBEF3BB928DAC7AB10A1ABC7">Author</mods:roleTerm>
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<mods:namePart id="697853EB4EE001C9D2A7B3B0451ED5B4">Ryvkin, Alexandr B.</mods:namePart>
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<mods:title id="B4D4F2F7094E45D06411D3D9A4C1C933">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part id="BAF662C256545FB87C8940290EE95839">
<mods:date id="70A42AE7ECEF165562D938A903E64673">2012</mods:date>
<mods:detail id="8B3D26E357CE6AC7AFB6B6A2CFC56C3E" type="pubDate">
<mods:number id="5F5108F8FFB3EA50B3D724CA666D070E">2012-05-24</mods:number>
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<mods:number id="9BCB88C76026C4E9F490BF7EDE8DAC34">13</mods:number>
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<mods:classification id="3351FC27C81E0EF02FC2C0D90F9A0D70">journal article</mods:classification>
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<mods:identifier id="8C29FB92BCD201E9D96161F4BC1B803F" type="DOI">10.5852/ejt.2012.13</mods:identifier>
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</mods:mods>
<treatment id="803F87FDC037FFD0FDF4AFE6FAA29D89" ID-DOI="http://doi.org/10.5281/zenodo.3858646" ID-GBIF-Taxon="164302144" ID-Zenodo-Dep="3858646" LSID="urn:lsid:zoobank.org:act:06D9027A-5EDA-4C66-B54C-B7E7F9FD283A" httpUri="http://treatment.plazi.org/id/803F87FDC037FFD0FDF4AFE6FAA29D89" lastPageId="29" lastPageNumber="30" pageId="27" pageNumber="28">
<subSubSection id="408C6560C037FFD6FDF4AFE6FC0B9AFB" box="[597,990,686,713]" pageId="27" pageNumber="28" type="nomenclature">
<paragraph id="082936EBC037FFD6FDF4AFE6FC0B9AFB" blockId="27.[597,990,686,753]" box="[597,990,686,713]" pageId="27" pageNumber="28">
<heading id="53618187C037FFD6FDF4AFE6FC0B9AFB" box="[597,990,686,713]" centered="true" fontSize="11" level="2" pageId="27" pageNumber="28" reason="2">
<taxonomicName id="CF964D68C037FFD6FDF4AFE6FCA89AFB" ID-CoL="8Y9Z9" authority="Ryvkin, 2012" authorityName="Ryvkin" authorityYear="2012" box="[597,893,686,713]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="canosus" status="sp. nov." subGenus="Nestus">
<emphasis id="3AE2EAF9C037FFD6FDF4AFE6FCA89AFB" bold="true" box="[597,893,686,713]" pageId="27" pageNumber="28">
<emphasis id="3AE2EAF9C037FFD6FDF4AFE6FD729AFA" bold="true" box="[597,679,686,712]" italics="true" pageId="27" pageNumber="28">Stenus</emphasis>
(
<emphasis id="3AE2EAF9C037FFD6FD19AFE7FCDC9AFB" bold="true" box="[696,777,687,713]" italics="true" pageId="27" pageNumber="28">Nestus</emphasis>
)
<emphasis id="3AE2EAF9C037FFD6FCBAAFE7FCA89AFB" bold="true" box="[795,893,687,713]" italics="true" pageId="27" pageNumber="28">canosus</emphasis>
</emphasis>
</taxonomicName>
<taxonomicNameLabel id="21D15782C037FFD6FC25AFE7FC0B9AFB" box="[900,990,687,713]" pageId="27" pageNumber="28" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="408C6560C037FFD6FD08AF9BFC5E9AC2" box="[681,907,723,753]" pageId="27" pageNumber="28" type="description">
<paragraph id="082936EBC037FFD6FD08AF9BFC5E9AC2" blockId="27.[597,990,686,753]" box="[681,907,723,753]" pageId="27" pageNumber="28">
(
<figureCitation id="90AD2A6EC037FFD6FD13AF9BFC9E9AC2" box="[690,843,723,753]" captionStart="Fig" captionStartId="28.[189,232,1895,1921]" captionTargetBox="[228,1365,265,1852]" captionTargetId="figure@28.[217,1370,253,1873]" captionTargetPageId="28" captionText="Fig. 4. AB: Stenus (Nestus) canosus sp. nov.: ♂ (holotype). A. Aedeagus, ventral view. B. Apical part of aedeagus, ventral view. CD: S. (N.) canalis sp. nov.: ♂ (holotype). C. Aedeagus, ventral view. D. Apical part of aedeagus, ventral view. [Note that the difference in shape of the apical lobe compared to that in C is due to the level of magnification.] Scales = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3857783" httpUri="https://zenodo.org/record/3857783/files/figure.png" pageId="27" pageNumber="28">
Figs. 4A
<emphasis id="3AE2EAF9C037FFD6FC83AF90FCE19AC3" box="[802,820,728,753]" italics="true" pageId="27" pageNumber="28"></emphasis>
B
</figureCitation>
,
<figureCitation id="90AD2A6EC037FFD6FCF8AF9BFC549AC2" box="[857,897,723,752]" captionStart="Fig" captionStartId="32.[189,232,1851,1877]" captionTargetBox="[249,1336,263,1808]" captionTargetId="figure@32.[244,1344,255,1810]" captionTargetPageId="32" captionText="Fig. 5. ♂ copulative structures. AE: Abdominal sternite 9. A. Stenus (Nestus) latipennis J. Sahlberg, 1880 (Russia: Krasnoyarsk Territory: Mirnoye). B. S. (N.) illusor Ryvkin, 1987 (Russia: Magadan Area: 2 km W of Splavnaya). C. S. (N.) delitor sp. nov. (holotype). D. S. (N.) canosus sp. nov. (holotype). E. S. (N.) canalis sp. nov. (holotype). F. S. (N.) caseyi Puthz, 1972 (U.S.A.: Michigan: Berrien Co.). Aedeagus, ventral view. Scales = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3857785" httpUri="https://zenodo.org/record/3857785/files/figure.png" pageId="27" pageNumber="28">5D</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="408C6560C037FFD6FF1CAE5EFD019BDE" pageId="27" pageNumber="28" type="materials_examined">
<paragraph id="082936EBC037FFD6FF1CAE5EFE7F9B02" blockId="27.[189,426,790,816]" box="[189,426,790,816]" pageId="27" pageNumber="28">
<heading id="53618187C037FFD6FF1CAE5EFE7F9B02" bold="true" box="[189,426,790,816]" fontSize="11" level="3" pageId="27" pageNumber="28" reason="3">
<emphasis id="3AE2EAF9C037FFD6FF1CAE5EFE7F9B02" bold="true" box="[189,426,790,816]" pageId="27" pageNumber="28">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="082936EBC037FFD6FF1CAE0DFD019BDE" blockId="27.[189,1398,836,1004]" pageId="27" pageNumber="28">
<materialsCitation id="B8FE3CB6C037FFD6FF1CAE0DFD189BD9" ID-GBIF-Occurrence="3890233420" collectingLocationText="holotype." collectionCode="ZMMU" collectorName="A. B. Ryvkin" country="China" elevation="3200" location="Tien" municipality="Zizhiqu" pageId="27" pageNumber="3" specimenCount="2" specimenCount-male="1" stateProvince="Xinjiang Uygur" typeStatus="holotype">
<collectingCountry id="7081767BC037FFD6FF1CAE0DFECF9B6D" box="[189,282,837,863]" name="China" pageId="27" pageNumber="28" value="China">CHINA</collectingCountry>
:
<typeStatus id="D72D8849C037FFD6FE8CAE0CFE4A9B6C" box="[301,415,836,862]" pageId="27" pageNumber="28" type="holotype">Holotype</typeStatus>
:
<specimenCount id="1E90FD62C037FFD6FE0CAE0DFE109B6D" box="[429,453,837,863]" pageId="27" pageNumber="28" type="male"></specimenCount>
, NW
<collectingCountry id="7081767BC037FFD6FDBBAE0CFDB69B6C" box="[538,611,836,862]" name="China" pageId="27" pageNumber="28">China</collectingCountry>
,
<collectingRegion id="CA52F809C037FFD6FDD3AE0CFCFA9B6D" box="[626,815,836,863]" country="China" name="Xinjiang Uygur" pageId="27" pageNumber="28">Xinjiang Uygur</collectingRegion>
|
<collectingMunicipality id="E84DAC91C037FFD6FCEBAE0CFC7D9B6C" box="[842,936,836,862]" pageId="27" pageNumber="28">Zizhiqu</collectingMunicipality>
,
<location id="0D496030C037FFD6FC16AE0CFC399B6C" LSID="urn:lsid:plazi:treatment:803F87FDC037FFD0FDF4AFE6FAA29D89:0D496030C037FFD6FC16AE0CFC399B6C" box="[951,1004,836,862]" country="China" municipality="Zizhiqu" name="Tien" pageId="27" pageNumber="28" stateProvince="Xinjiang Uygur">Tien</location>
|
<collectingRegion id="CA52F809C037FFD6FBA9AE0CFEE39BB0" pageId="27" pageNumber="28">Shan, Southern slope of Narat Mt Ridge</collectingRegion>
: |
<location id="0D496030C037FFD6FEF7AE20FE3D9BB0" LSID="urn:lsid:plazi:treatment:803F87FDC037FFD0FDF4AFE6FAA29D89:0D496030C037FFD6FEF7AE20FE3D9BB0" box="[342,488,872,898]" country="China" municipality="Zizhiqu" name="Koktau Mts" pageId="27" pageNumber="28" stateProvince="Xinjiang Uygur">Koktau Mts</location>
, right confluent | of
<location id="0D496030C037FFD6FD48AE20FC519BB0" LSID="urn:lsid:plazi:treatment:803F87FDC037FFD0FDF4AFE6FAA29D89:0D496030C037FFD6FD48AE20FC519BB0" box="[745,900,872,898]" country="China" municipality="Zizhiqu" name="Koksu River" pageId="27" pageNumber="28" stateProvince="Xinjiang Uygur">Koksu River</location>
,
<quantity id="CF6E9B0EC037FFD6FC35AE20FC269BB0" box="[916,1011,872,898]" metricMagnitude="3" metricUnit="m" metricValue="3.2" pageId="27" pageNumber="28" unit="m" value="3200.0">
<elevation id="83BBD1D8C037FFD6FC35AE20FC269BB0" box="[916,1011,872,898]" metricMagnitude="3" metricUnit="m" metricValue="3.2" pageId="27" pageNumber="28" unit="m" value="3200.0">3200 m</elevation>
</quantity>
, stone-| sandy bank of rill with sedge tussocks and | small willows. 25.07.[20]01. [S.V.]
<location id="0D496030C037FFD6FCD7AEC2FC269BAA" LSID="urn:lsid:plazi:treatment:803F87FDC037FFD0FDF4AFE6FAA29D89:0D496030C037FFD6FCD7AEC2FC269BAA" box="[886,1011,906,933]" country="China" municipality="Zizhiqu" name="Saluk. No" pageId="27" pageNumber="28" stateProvince="Xinjiang Uygur">
Saluk.
<superScript id="FFE39BA3C037FFD6FC75AEC3FC269BAA" attach="right" box="[980,1011,906,933]" fontSize="6" pageId="27" pageNumber="28">No</superScript>
</location>
11 &lt;printed label in
<location id="0D496030C037FFD6FAA3AEC3FAB79B97" LSID="urn:lsid:plazi:treatment:803F87FDC037FFD0FDF4AFE6FAA29D89:0D496030C037FFD6FAA3AEC3FAB79B97" box="[1282,1378,907,933]" country="China" municipality="Zizhiqu" name="Russian" pageId="27" pageNumber="28" stateProvince="Xinjiang Uygur">Russian</location>
&gt;, HOLOTYPUS &lt;my standard printed red label&gt;,
<taxonomicName id="CF964D68C037FFD6FCBBAEE7FCB29BFB" box="[794,871,943,969]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="genus">
<emphasis id="3AE2EAF9C037FFD6FCBBAEE7FCB29BFB" box="[794,871,943,969]" italics="true" pageId="27" pageNumber="28">
<location id="0D496030C037FFD6FCBBAEE7FCB29BFB" LSID="urn:lsid:plazi:treatment:803F87FDC037FFD0FDF4AFE6FAA29D89:0D496030C037FFD6FCBBAEE7FCB29BFB" box="[794,871,943,969]" country="China" municipality="Zizhiqu" name="Stenus" pageId="27" pageNumber="28" stateProvince="Xinjiang Uygur">Stenus</location>
</emphasis>
</taxonomicName>
HT |
<taxonomicName id="CF964D68C037FFD6FC0AAEE7FBDF9BFA" authority="Ryvkin, 2012" authorityName="Ryvkin" authorityYear="2012" box="[939,1034,943,968]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="canosus" status="sp. nov.">
<emphasis id="3AE2EAF9C037FFD6FC0AAEE7FBDF9BFA" box="[939,1034,943,968]" italics="true" pageId="27" pageNumber="28">canosus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="21D15782C037FFD6FBB0AEE7FB859BFB" box="[1041,1104,943,969]" pageId="27" pageNumber="28" rank="species">
<specimenCount id="1E90FD62C037FFD6FBB0AEE7FBE69BFB" box="[1041,1075,943,969]" pageId="27" pageNumber="28" type="generic">sp.</specimenCount>
n.
</taxonomicNameLabel>
|
<collectorName id="A563533DC037FFD6FBC4AEE6FB249BFA" box="[1125,1265,942,968]" pageId="27" pageNumber="28">A.B.Ryvkin</collectorName>
det., 2010 &lt;my standard determinative label&gt; (
<collectionCode id="6E87AE2EC037FFD6FDC6AE99FD139BD9" box="[615,710,977,1003]" collectionName="ZMMU" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="27" pageNumber="28" type="Museum">ZMMU</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="408C6560C037FFD6FF1CA95BFE1B9CDA" pageId="27" pageNumber="28" type="diagnosis">
<paragraph id="082936EBC037FFD6FF1CA95BFE1C9C1F" blockId="27.[189,457,1043,1069]" box="[189,457,1043,1069]" pageId="27" pageNumber="28">
<heading id="53618187C037FFD6FF1CA95BFE1C9C1F" bold="true" box="[189,457,1043,1069]" fontSize="11" level="3" pageId="27" pageNumber="28" reason="3">
<emphasis id="3AE2EAF9C037FFD6FF1CA95BFE1C9C1F" bold="true" box="[189,457,1043,1069]" pageId="27" pageNumber="28">Differential diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="082936EBC037FFD6FF1CA909FE1B9CDA" blockId="27.[189,1399,1089,1256]" pageId="27" pageNumber="28">
This new species is externally very similar to
<taxonomicName id="CF964D68C037FFD6FD4BA90AFB7F9C69" authority="Gyllenhal, 1827" authorityName="Gyllenhal" authorityYear="1827" box="[746,1194,1089,1116]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="canaliculatus" subGenus="Nestus">
<emphasis id="3AE2EAF9C037FFD6FD4BA90AFCD59C6E" box="[746,768,1090,1116]" italics="true" pageId="27" pageNumber="28">S.</emphasis>
(
<emphasis id="3AE2EAF9C037FFD6FCB5A90AFCFA9C69" box="[788,815,1090,1115]" italics="true" pageId="27" pageNumber="28">N.</emphasis>
)
<emphasis id="3AE2EAF9C037FFD6FCE3A909FC359C69" box="[834,992,1089,1115]" italics="true" pageId="27" pageNumber="28">canaliculatus</emphasis>
<bibRefCitation id="6C074B1AC037FFD6FC4BA909FB7F9C69" author="Gyllenhal" box="[1002,1194,1089,1115]" firstAuthor="Gyllenhal" pageId="27" pageNumber="28" refId="ref41615" refString="Gyllenhal L. 1827. Insecta Suecica descripta. Classis I: Coleoptera sive Eleuterata. Vol. 1. ps. IV: i-viii + 1 - 762. Friederichum Fleischer, Lipsiae. http: // dx. doi. org / 10.5962 / bhl. title. 8767" type="book" year="1827">Gyllenhal, 1827</bibRefCitation>
</taxonomicName>
and differs from it by the character of the puncturation, which is much coarser, more sparse and irregular, by the less developed microsculpture, by the median furrow of pronotum, which is deeper and broader, and by the shape of the aedeagus. External differences from all the known species of the
<taxonomicName id="CF964D68C037FFD6FBFFA9E3FB299CF7" authorityName="Gyllenhal" authorityYear="1827" box="[1118,1276,1195,1221]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="canaliculatus">
<emphasis id="3AE2EAF9C037FFD6FBFFA9E3FB299CF7" box="[1118,1276,1195,1221]" italics="true" pageId="27" pageNumber="28">canaliculatus</emphasis>
</taxonomicName>
-group are given in the key below.
</paragraph>
</subSubSection>
<subSubSection id="408C6560C037FFD6FF1CA858FBD09D6B" box="[189,1029,1296,1369]" pageId="27" pageNumber="28" type="etymology">
<paragraph id="082936EBC037FFD6FF1CA858FE919D18" blockId="27.[189,324,1296,1322]" box="[189,324,1296,1322]" pageId="27" pageNumber="28">
<heading id="53618187C037FFD6FF1CA858FE919D18" bold="true" box="[189,324,1296,1322]" fontSize="11" level="3" pageId="27" pageNumber="28" reason="3">
<emphasis id="3AE2EAF9C037FFD6FF1CA858FE919D18" bold="true" box="[189,324,1296,1322]" pageId="27" pageNumber="28">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="082936EBC037FFD6FF1CA876FBD09D6B" blockId="27.[189,1029,1342,1369]" box="[189,1029,1342,1369]" pageId="27" pageNumber="28">
The name of this species is the Latin adjective “
<taxonomicName id="CF964D68C037FFD6FD51A877FC9A9D6A" authorityName="Ryvkin" authorityYear="2012" box="[752,847,1343,1368]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="canosus">
<emphasis id="3AE2EAF9C037FFD6FD51A877FC9A9D6A" box="[752,847,1343,1368]" italics="true" pageId="27" pageNumber="28">canosus</emphasis>
</taxonomicName>
” (grey-haired).
</paragraph>
</subSubSection>
<subSubSection id="408C6560C037FFD0FF1CA837FE4B9D36" lastPageId="29" lastPageNumber="30" pageId="27" pageNumber="28" type="description">
<paragraph id="082936EBC037FFD6FF1CA837FE9B9DAB" blockId="27.[189,334,1407,1433]" box="[189,334,1407,1433]" pageId="27" pageNumber="28">
<heading id="53618187C037FFD6FF1CA837FE9B9DAB" bold="true" box="[189,334,1407,1433]" fontSize="11" level="3" pageId="27" pageNumber="28" reason="3">
<emphasis id="3AE2EAF9C037FFD6FF1CA837FE9B9DAB" bold="true" box="[189,334,1407,1433]" pageId="27" pageNumber="28">Description</emphasis>
</heading>
</paragraph>
<paragraph id="082936EBC037FFD6FF1CA8E7FD6D9DFB" blockId="27.[189,696,1454,1481]" box="[189,696,1454,1481]" pageId="27" pageNumber="28">LENGTH. 4.0 mm (with abdomen extended).</paragraph>
<paragraph id="082936EBC037FFD6FF1CA8B8FCEA9E41" blockId="27.[189,1398,1519,1652]" pageId="27" pageNumber="28">COLORATION. Black, distinctly shining, with moderately long and dense silvery pubescence. Legs dark brown with knees somewhat darkened, brownish-black. Antennae dark brown, with segments 12 pitchy-black and club vaguely infuscate; 1st segment of maxillar palpi yellow, 2nd brown with yellow base, 3rd dark brown with very base yellowish.</paragraph>
<paragraph id="082936EBC037FFD6FF1CABD3FD7E9FC0" blockId="27.[189,1398,1690,2034]" pageId="27" pageNumber="28">HEAD. Distinctly broader than pronotum (50:44), somewhat narrower than elytra between humeri (50:53), much narrower than those in their broadest part (47:54). Front feebly evenly convex, with very vague and shallow lateral impressions distinguishable mainly between antennal tubercles and eyes. Puncturation moderately dense, somewhat irregular, evidently smaller and denser between antennal tubercles, partly nonrugosely confluent both there and at periocular areas, without distinct smooth spots and strips, but obviously sparser and greater in the middle; average diameter of punctures in the middle distinctly larger than the greatest cross-section of antennal segment 3. Frontal slope before antennal fossae rather flat. Anterior margin of labrum with small median notch. Antennae moderately long, scarcely reaching middle of pronotum. Length proportions of antennal segments 211 = 6:5:6:4:4:4:3:4:4.5:6; segments of club longer than broad (4:3, 4.5:4, 6:4).</paragraph>
<caption id="5CE96663C030FFD1FF1CAA2FFC6E9FD9" ID-DOI="http://doi.org/10.5281/zenodo.3857783" ID-Zenodo-Dep="3857783" httpUri="https://zenodo.org/record/3857783/files/figure.png" pageId="28" pageNumber="29" startId="28.[189,232,1895,1921]" targetBox="[228,1365,265,1852]" targetPageId="28">
<paragraph id="082936EBC030FFD1FF1CAA2FFC6E9FD9" blockId="28.[189,1399,1895,2027]" pageId="28" pageNumber="29">
<emphasis id="3AE2EAF9C030FFD1FF1CAA2FFE899FB3" bold="true" box="[189,348,1895,1922]" pageId="28" pageNumber="29">Fig. 4. AB</emphasis>
:
<taxonomicName id="CF964D68C030FFD1FECBAA20FD519FB3" authority="Ryvkin, 2012" authorityName="Ryvkin" authorityYear="2012" box="[362,644,1895,1922]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="canosus" status="sp. nov." subGenus="Nestus">
<emphasis id="3AE2EAF9C030FFD1FECBAA20FE629FB0" box="[362,439,1896,1922]" italics="true" pageId="28" pageNumber="29">Stenus</emphasis>
(
<emphasis id="3AE2EAF9C030FFD1FE66AA20FDC09FB3" box="[455,533,1896,1921]" italics="true" pageId="28" pageNumber="29">Nestus</emphasis>
)
<emphasis id="3AE2EAF9C030FFD1FD84AA20FD519FB3" box="[549,644,1896,1921]" italics="true" pageId="28" pageNumber="29">canosus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="21D15782C030FFD1FD2BAA20FD3D9FB0" box="[650,744,1896,1922]" pageId="28" pageNumber="29" rank="species">sp. nov.</taxonomicNameLabel>
: ♂ (holotype).
<emphasis id="3AE2EAF9C030FFD1FC37AA2FFC7E9FB3" bold="true" box="[918,939,1895,1921]" pageId="28" pageNumber="29">A</emphasis>
. Aedeagus, ventral view.
<emphasis id="3AE2EAF9C030FFD1FB74AA2FFB3C9FB3" bold="true" box="[1237,1257,1895,1921]" pageId="28" pageNumber="29">B</emphasis>
. Apical part of aedeagus, ventral view.
<emphasis id="3AE2EAF9C030FFD1FDACAAC2FD939F96" bold="true" box="[525,582,1930,1956]" pageId="28" pageNumber="29">CD</emphasis>
:
<taxonomicName id="CF964D68C030FFD1FDF4AAC3FD2B9F96" authority="Ryvkin, 2012" authorityName="Ryvkin" authorityYear="2012" box="[597,766,1930,1957]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="canalis" status="sp. nov." subGenus="Nestus">
<emphasis id="3AE2EAF9C030FFD1FDF4AAC3FDBE9F97" box="[597,619,1931,1957]" italics="true" pageId="28" pageNumber="29">S.</emphasis>
(
<emphasis id="3AE2EAF9C030FFD1FDDCAAC3FD4D9F96" box="[637,664,1931,1956]" italics="true" pageId="28" pageNumber="29">N.</emphasis>
)
<emphasis id="3AE2EAF9C030FFD1FD08AAC2FD2B9F96" box="[681,766,1930,1956]" italics="true" pageId="28" pageNumber="29">canalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="21D15782C030FFD1FCA4AAC3FCB19F97" box="[773,868,1931,1957]" pageId="28" pageNumber="29" rank="species">sp. nov.</taxonomicNameLabel>
: ♂ (holotype).
<emphasis id="3AE2EAF9C030FFD1FBB7AAC2FBFE9F96" bold="true" box="[1046,1067,1930,1956]" pageId="28" pageNumber="29">C</emphasis>
. Aedeagus, ventral view.
<emphasis id="3AE2EAF9C030FFD1FAFBAAC2FABA9F96" bold="true" box="[1370,1391,1930,1956]" pageId="28" pageNumber="29">D</emphasis>
. Apical part of aedeagus, ventral view. [Note that the difference in shape of the apical lobe compared to that in C is due to the level of magnification.] Scales = 0.1 mm.
</paragraph>
</caption>
<paragraph id="082936EBC031FFD0FF1CAC44FE4E9980" blockId="29.[189,1399,267,434]" pageId="29" pageNumber="30">PRONOTUM. A bit longer than broad (45:44), broadest near middle of length, narrowed feebly convexly towards anterior margin and concavely towards posterior one. Longitudinal median furrow deep and sharp over entire length except for very anterior piece. Laterobasal depressions very shallow, nearly vanishing. Puncturation deep and irregular, partly nonrugosely confluent; punctures distinctly larger than those of head.</paragraph>
<paragraph id="082936EBC031FFD0FF1CAC92FDCE9AB2" blockId="29.[189,1398,473,641]" pageId="29" pageNumber="30">ELYTRA. A bit broader than long (60:58), much longer than pronotum (58:45), at suture distinctly longer than the latter (48:45). Humeri anglarly rounded, lateral sides uniformly moderately divergent posteriorly (53:60). Humeral and sutural depressions feeble but evident in anterior half of disk. Puncturation deep, much more regular, evidently larger in diameter than that of head and pronotum, sometimes confluent, but not forming long furrows.</paragraph>
<paragraph id="082936EBC031FFD0FF1CAFE0FE0D9AD7" blockId="29.[189,1398,680,742]" pageId="29" pageNumber="30">LEGS. Fairly short; metatibia about 1/3 longer than metatarsus (40:29); segment 1 of metatarsi about as long as segment 5 (8:8).</paragraph>
<paragraph id="082936EBC031FFD0FF1CAE45FD7E9B86" blockId="29.[189,1399,780,948]" pageId="29" pageNumber="30">ABDOMEN. Moderately convex, with well developed paratergites and two pairs of keels at bases of anterior visible tergites. Lateral sides feebly uniformly convergent. Posterior margin of tergite 7 with fine light fringe. Puncturation of tergites evidently smaller than that of head and pronotum; on anterior visible tergites much denser and smaller laterally, medioposteriorly with some smooth interstices distinctly wider than average diameter of punctures.</paragraph>
<paragraph id="082936EBC031FFD0FF1CAE93FAA29C0E" blockId="29.[189,1399,987,1084]" pageId="29" pageNumber="30">MICROSCULPTURE. Very fine mesh-like ground sculpture visible between punctures, mainly on elytra and abdominal tergites 89, as well as on lateral parts of head, pronotum and anterior abdominal tergites, median parts of abdominal tergites quite smooth or slightly irregularly netted only on edges of punctures.</paragraph>
<paragraph id="082936EBC031FFD0FF1CA92BFC5A9CF1" blockId="29.[189,1398,1122,1220]" pageId="29" pageNumber="30">
MALE. Meso- and metatibiae without specific characters; 8th abdominal sternite without conspicuous impressions and keels, its posterior margin with very broad and not deep angularly-rounded emargination; abdominal sternite 9 as in
<figureCitation id="90AD2A6EC031FFD0FE53A9E1FD9B9CF6" box="[498,590,1193,1220]" captionStart="Fig" captionStartId="32.[189,232,1851,1877]" captionTargetBox="[249,1336,263,1808]" captionTargetId="figure@32.[244,1344,255,1810]" captionTargetPageId="32" captionText="Fig. 5. ♂ copulative structures. AE: Abdominal sternite 9. A. Stenus (Nestus) latipennis J. Sahlberg, 1880 (Russia: Krasnoyarsk Territory: Mirnoye). B. S. (N.) illusor Ryvkin, 1987 (Russia: Magadan Area: 2 km W of Splavnaya). C. S. (N.) delitor sp. nov. (holotype). D. S. (N.) canosus sp. nov. (holotype). E. S. (N.) canalis sp. nov. (holotype). F. S. (N.) caseyi Puthz, 1972 (U.S.A.: Michigan: Berrien Co.). Aedeagus, ventral view. Scales = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3857785" httpUri="https://zenodo.org/record/3857785/files/figure.png" pageId="29" pageNumber="30">Fig. 5D</figureCitation>
; aedeagus as in
<figureCitation id="90AD2A6EC031FFD0FCADA9E1FC5A9CF1" box="[780,911,1193,1219]" captionStart="Fig" captionStartId="28.[189,232,1895,1921]" captionTargetBox="[228,1365,265,1852]" captionTargetId="figure@28.[217,1370,253,1873]" captionTargetPageId="28" captionText="Fig. 4. AB: Stenus (Nestus) canosus sp. nov.: ♂ (holotype). A. Aedeagus, ventral view. B. Apical part of aedeagus, ventral view. CD: S. (N.) canalis sp. nov.: ♂ (holotype). C. Aedeagus, ventral view. D. Apical part of aedeagus, ventral view. [Note that the difference in shape of the apical lobe compared to that in C is due to the level of magnification.] Scales = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3857783" httpUri="https://zenodo.org/record/3857783/files/figure.png" pageId="29" pageNumber="30">Fig. 4A, B.</figureCitation>
</paragraph>
<paragraph id="082936EBC031FFD0FF1CA9A3FE4B9D36" blockId="29.[189,414,1258,1285]" box="[189,414,1258,1285]" pageId="29" pageNumber="30">FEMALE. Unknown.</paragraph>
</subSubSection>
<subSubSection id="408C6560C031FFD0FF1CA864FAA29D89" pageId="29" pageNumber="30" type="discussion">
<paragraph id="082936EBC031FFD0FF1CA864FEFA9D74" blockId="29.[189,303,1324,1350]" box="[189,303,1324,1350]" pageId="29" pageNumber="30">
<heading id="53618187C031FFD0FF1CA864FEFA9D74" bold="true" box="[189,303,1324,1350]" fontSize="11" level="3" pageId="29" pageNumber="30" reason="3">
<emphasis id="3AE2EAF9C031FFD0FF1CA864FEFA9D74" bold="true" box="[189,303,1324,1350]" pageId="29" pageNumber="30">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="082936EBC031FFD0FF1CA813FAA29D89" blockId="29.[189,1399,1370,1467]" pageId="29" pageNumber="30">
No records of
<taxonomicName id="CF964D68C031FFD0FECAA813FDB59D46" authorityName="Gyllenhal" authorityYear="1827" box="[363,608,1370,1397]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="canaliculatus" subGenus="Nestus">
<emphasis id="3AE2EAF9C031FFD0FECAA813FEAF9D47" box="[363,378,1371,1397]" italics="true" pageId="29" pageNumber="30">S</emphasis>
. (
<emphasis id="3AE2EAF9C031FFD0FE35A813FE7A9D46" box="[404,431,1371,1396]" italics="true" pageId="29" pageNumber="30">N.</emphasis>
)
<emphasis id="3AE2EAF9C031FFD0FE63A812FDB59D46" box="[450,608,1370,1396]" italics="true" pageId="29" pageNumber="30">canaliculatus</emphasis>
</taxonomicName>
were previously known from China or from adjacent countries of Central Asia. The capture of
<taxonomicName id="CF964D68C031FFD0FDB4A836FD1C9DA5" authority="Ryvkin, 2012" authorityName="Ryvkin" authorityYear="2012" box="[533,713,1406,1432]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="canosus" status="sp. nov." subGenus="Nestus">
<emphasis id="3AE2EAF9C031FFD0FDB4A836FDFE9DAA" box="[533,555,1406,1432]" italics="true" pageId="29" pageNumber="30">S.</emphasis>
(
<emphasis id="3AE2EAF9C031FFD0FD9CA836FD8C9DA5" box="[573,601,1406,1431]" italics="true" pageId="29" pageNumber="30">N.</emphasis>
)
<emphasis id="3AE2EAF9C031FFD0FDCBA836FD1C9DA5" box="[618,713,1406,1431]" italics="true" pageId="29" pageNumber="30">canosus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="21D15782C031FFD0FD70A836FCFA9DAA" box="[721,815,1406,1432]" pageId="29" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
, which is closely related to
<taxonomicName id="CF964D68C031FFD0FBDDA836FABA9DAA" authorityName="Gyllenhal" authorityYear="1827" box="[1148,1391,1406,1432]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="canaliculatus" subGenus="Nestus">
<emphasis id="3AE2EAF9C031FFD0FBDDA836FB5E9DAA" box="[1148,1163,1406,1432]" italics="true" pageId="29" pageNumber="30">S</emphasis>
. (
<emphasis id="3AE2EAF9C031FFD0FB05A836FB6A9DA5" box="[1188,1215,1406,1431]" italics="true" pageId="29" pageNumber="30">N.</emphasis>
)
<emphasis id="3AE2EAF9C031FFD0FB70A836FABA9DAA" box="[1233,1391,1406,1432]" italics="true" pageId="29" pageNumber="30">canaliculatus</emphasis>
</taxonomicName>
, proves that their common ancestor was recently widely distributed throughout these spacious territories.
</paragraph>
</subSubSection>
</treatment>
</document>