treatments-xml/data/80/3F/87/803F87FDC02FFFCAFD89A96FFB6B9A10.xml

<document id="AC1CEBABFD44F9C1AC3833977BC5A080" ID-CLB-Dataset="21840" ID-DOI="10.5852/ejt.2012.13" ID-GBIF-Dataset="9691a997-b6e4-4d47-9082-bbcba7347a3e" ID-ISSN="2118-9773" ID-Zenodo-Dep="3857775" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1590516738498" checkinUser="carolina" docAuthor="Ryvkin, Alexandr B." docDate="2012" docId="803F87FDC02FFFCAFD89A96FFB6B9A10" docLanguage="en" docName="ejt-2012-13.pdf.imf" docOrigin="European Journal of Taxonomy 13" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Stenus canaliculatus Gyllenhal 1827" docType="treatment" docVersion="10" lastPageNumber="8" masterDocId="7C06FF85C02CFFCDFFA1AD48FFD59832" masterDocTitle="New species and records of Stenus (Nestus) of the canaliculatus group, with the erection of a new species group (Insecta: Coleoptera: Staphylinidae: Steninae)" masterLastPageNumber="62" masterPageNumber="1" pageNumber="4" updateTime="1698841419189" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="8B2A1AFFB338CF258D1DB7E857FDB1D0">New species and records of Stenus (Nestus) of the canaliculatus group, with the erection of a new species group (Insecta: Coleoptera: Staphylinidae: Steninae)</mods:title>
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<treatment id="803F87FDC02FFFCAFD89A96FFB6B9A10" ID-DOI="http://doi.org/10.5281/zenodo.3858638" ID-GBIF-Taxon="164302148" ID-Zenodo-Dep="3858638" LSID="urn:lsid:plazi:treatment:803F87FDC02FFFCAFD89A96FFB6B9A10" httpUri="http://treatment.plazi.org/id/803F87FDC02FFFCAFD89A96FFB6B9A10" lastPageId="7" lastPageNumber="8" pageId="3" pageNumber="4">
<subSubSection id="408C6560C02FFFCEFD89A96FFBDE9C73" box="[552,1035,1063,1090]" pageId="3" pageNumber="4" type="nomenclature">
<paragraph id="082936EBC02FFFCEFD89A96FFBDE9C73" blockId="3.[552,1035,1063,1090]" box="[552,1035,1063,1090]" pageId="3" pageNumber="4">
<heading id="53618187C02FFFCEFD89A96FFBDE9C73" box="[552,1035,1063,1090]" centered="true" fontSize="11" level="2" pageId="3" pageNumber="4" reason="2">
<emphasis id="3AE2EAF9C02FFFCEFD89A96FFBDE9C73" bold="true" box="[552,1035,1063,1090]" pageId="3" pageNumber="4">
Redefinition of the 
<taxonomicName id="CF964D68C02FFFCEFCB6A96FFC6F9C70" ID-CoL="5297K" authorityName="Gyllenhal" authorityYear="1827" box="[791,954,1063,1090]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="canaliculatus">
<emphasis id="3AE2EAF9C02FFFCEFCB6A96FFC6F9C70" bold="true" box="[791,954,1063,1090]" italics="true" pageId="3" pageNumber="4">canaliculatus</emphasis>
</taxonomicName>
group
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="408C6560C02FFFC9FF1CA920FEB79DD0" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4" type="diagnosis">
<paragraph id="082936EBC02FFFCEFF1CA920FEE39CB0" blockId="3.[189,310,1128,1154]" box="[189,310,1128,1154]" pageId="3" pageNumber="4">
<heading id="53618187C02FFFCEFF1CA920FEE39CB0" bold="true" box="[189,310,1128,1154]" fontSize="11" level="3" pageId="3" pageNumber="4" reason="3">
<emphasis id="3AE2EAF9C02FFFCEFF1CA920FEE39CB0" bold="true" box="[189,310,1128,1154]" pageId="3" pageNumber="4">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="082936EBC02FFFCEFF1CA9DFFC2F9C80" blockId="3.[189,1018,1175,1202]" box="[189,1018,1175,1202]" pageId="3" pageNumber="4">
Body size moderate; body length of the known species: 
<quantity id="CF6E9B0EC02FFFCEFCEFA9DFFC239C80" box="[846,1014,1175,1202]" metricMagnitude="-3" metricUnit="m" metricValue="3.75" metricValueMax="4.7" metricValueMin="2.8" pageId="3" pageNumber="4" unit="mm" value="3.75" valueMax="4.7" valueMin="2.8">2.8 to 4.7 mm</quantity>
.
</paragraph>
<paragraph id="082936EBC02FFFCEFF1CA991FD599DF5" blockId="3.[189,1398,1241,1479]" pageId="3" pageNumber="4">Head fairly small to fairly broad in comparison to elytra and pronotum. Upper surface between eyes feebly evenly convex, nearly flat, to slightly concave as a whole; with longitudinal elevations and impressions nearly absent to well developed. Antennae rather short to long, with the club segments more or less elongated. Internal tooth of each mandible placed a little before the middle, directed only slightly dorsally of the main plane. Maxillar palpi with yellow and not shortened basal segment. A considerable reduction of the adhesion capture apparatus may be supposed for all the members of the group (for more details see the Remarks section below).</paragraph>
<paragraph id="082936EBC02FFFCEFF1CA8A6FEBD9E18" blockId="3.[189,1398,1517,1578]" pageId="3" pageNumber="4">Pronotum with long and sharp longitudinal median furrow more or less deep, less often very feeble, stroke-shaped.</paragraph>
<paragraph id="082936EBC02FFFCEFF1CAB1AFEA49EBD" blockId="3.[189,1398,1618,1679]" pageId="3" pageNumber="4">Elytra nearly rectangular, large, with short rounded humeri, to shortened, slightly trapezoid with humeri less developed.</paragraph>
<paragraph id="082936EBC02FFFCEFF1CABFFFAF89EC6" blockId="3.[189,1399,1719,1781]" pageId="3" pageNumber="4">Legs moderately long; the segment 4 of tarsi without emargination; segment 1 of metatarsi about the same length as to a bit longer than segment 5 but distinctly shorter than segments 2 to 4 together.</paragraph>
<paragraph id="082936EBC02FFFCEFF1CAA53FCE99F4E" blockId="3.[189,1399,1819,1916]" pageId="3" pageNumber="4">Abdomen moderately convex, with paratergites evident on abdominal segments 3 to 7; four anterior visible tergites each with four short but evident longitudinal keels at basal part; posterior margin of the tergite 7 with well-developed fine membranous fringe.</paragraph>
<paragraph id="082936EBC02FFFCEFF1CAAEBFC319F8C" blockId="3.[189,996,1955,1982]" box="[189,996,1955,1982]" pageId="3" pageNumber="4">Puncturation of pronotum and elytra diverse, but always non-rugose.</paragraph>
<paragraph id="082936EBC028FFC9FF1CAC43FCBD9917" blockId="4.[189,872,267,294]" box="[189,872,267,294]" pageId="4" pageNumber="5">Ground sculpture cellular or net-shaped to entirely absent.</paragraph>
<paragraph id="082936EBC028FFC9FF1CAC05FC5F9955" blockId="4.[189,906,333,359]" box="[189,906,333,359]" pageId="4" pageNumber="5">Pubescence moderate, contiguous to fairly long, outstanding.</paragraph>
<paragraph id="082936EBC028FFC9FF1CACC7FD8E9AD7" blockId="4.[189,1399,398,742]" pageId="4" pageNumber="5">MALE. Legs without specific features; posterior margin of abdominal sternites 6–7 shallowly emarginated to about straight;abdominal sternite 8with a broad but not deep, rounded or angularly rounded emargination of posterior margin; abdominal sternite 9 with large posterolateral teeth not incurved inwards; ventral depressions of sternites 6–8, if present, flat and feeble, never flanked by sharp keels. Aedeagus relatively large; median lobe archetypically lanceolate, with apical sclerotized part to a variable extent angular to angularly rounded; endophallus with paired medial bands (optionally, an unpaired bar may also be distinct), small but visible lateral parts, and H-shaped expulsion clasp producing lateral portions fairly broad; basal tube of diverse structure, more or less sclerotized. Parameres cylindrical, with apical broadening, if present, very feeble, rather vague; apico-internal setiferous surface differentiated poorly to moderately, setae fairly uniform.</paragraph>
<paragraph id="082936EBC028FFC9FF1CAE45FAEA9B78" blockId="4.[189,1398,780,842]" pageId="4" pageNumber="5">FEMALE. Posterior margin of abdominal sternite 8 broadly rounded to angularly rounded; each valvifer with posterolateral tooth directed backwards. Spermatheca sclerotized to a greater or lesser extent.</paragraph>
<paragraph id="082936EBC028FFC9FF1CAE3AFEB79DD0" blockId="4.[189,1399,881,1506]" pageId="4" pageNumber="5">
In the structure of abdomen, the male genitalia, the abdominal sternite 9 of both males and females (valvifera), the group under consideration is most closely related to the 
<taxonomicName id="CF964D68C028FFC9FB8CAEDCFBA69B9C" authorityName="Ljungh" authorityYear="1810" box="[1069,1139,916,942]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="boops">
<emphasis id="3AE2EAF9C028FFC9FB8CAEDCFBA69B9C" box="[1069,1139,916,942]" italics="true" pageId="4" pageNumber="5">boops</emphasis>
</taxonomicName>
and 
<emphasis id="3AE2EAF9C028FFC9FB12AEDCFACE9B9C" box="[1203,1307,916,942]" italics="true" pageId="4" pageNumber="5">palposus</emphasis>
groups of the same subgenus 
<taxonomicName id="CF964D68C028FFC9FE69AEF0FDC39BE3" authorityName="Rey" authorityYear="1884" box="[456,534,952,977]" class="Insecta" family="Staphylinidae" genus="Nestus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="3AE2EAF9C028FFC9FE69AEF0FDC39BE3" box="[456,534,952,977]" italics="true" pageId="4" pageNumber="5">Nestus</emphasis>
</taxonomicName>
. It differs from these by the structure of the mandibles, by the adhesion capture apparatus more or less shortened, by the pronotum with long and sharp, line-shaped, longitudinal median furrow, by the evidently narrower aedeagus, by the endophallus with the expulsion clasp’s transversal bridge adjoining to the lateral pieces near their middle, by the abdominal sternite 9 with posterolateral teeth not being curved inwards, by the male legs without peculiar features (not all the species of the named groups have such features well developed); from the 
<emphasis id="3AE2EAF9C028FFC9FBEDA92FFB619CB3" box="[1100,1204,1127,1153]" italics="true" pageId="4" pageNumber="5">palposus</emphasis>
group it can be easily distinguished by the yellow basal segment of the maxillar palpi. The differences from 
<taxonomicName id="CF964D68C028FFC9FA89A9C3FE2A9CFA" authority="Marsham, 1802" authorityName="Marsham" authorityYear="1802" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="42" phylum="Arthropoda" rank="species" species="melanopus" subGenus="Nestus">
<emphasis id="3AE2EAF9C028FFC9FA89A9C3FAE29C97" box="[1320,1335,1163,1189]" italics="true" pageId="4" pageNumber="5">S</emphasis>
. (
<emphasis id="3AE2EAF9C028FFC9FAF3A9C4FAB39C97" box="[1362,1382,1164,1189]" italics="true" pageId="4" pageNumber="5">N</emphasis>
.) 
<emphasis id="3AE2EAF9C028FFC9FF1CA9E6FEE99CFA" box="[189,316,1198,1224]" italics="true" pageId="4" pageNumber="5">melanopus</emphasis>
<bibRefCitation id="6C074B1AC028FFC9FEE5A9E6FE2A9CFA" author="Marsham" box="[324,511,1198,1224]" firstAuthor="Marsham" pageId="4" pageNumber="5" refId="ref42860" refString="Marsham T. 1802. Entomologia britannica, sistens insecta, Britanniae indigena, secundum methodum Linnaeanum disposita. Tomus I. Coleoptera: I-XXXI + 1 - 549. Wilks &amp; Taylor, Londini." type="book" year="1802">Marsham, 1802</bibRefCitation>
</taxonomicName>
, similar externally to some species of the 
<taxonomicName id="CF964D68C028FFC9FC50A9E6FB5A9CFA" authorityName="Gyllenhal" authorityYear="1827" box="[1009,1167,1198,1224]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="canaliculatus">
<emphasis id="3AE2EAF9C028FFC9FC50A9E6FB5A9CFA" box="[1009,1167,1198,1224]" italics="true" pageId="4" pageNumber="5">canaliculatus</emphasis>
</taxonomicName>
group, are given in the diagnosis of the 
<taxonomicName id="CF964D68C028FFC9FE10A999FDE59CD9" authorityName="Marsham" authorityYear="1802" box="[433,560,1233,1259]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="melanopus">
<emphasis id="3AE2EAF9C028FFC9FE10A999FDE59CD9" box="[433,560,1233,1259]" italics="true" pageId="4" pageNumber="5">melanopus</emphasis>
</taxonomicName>
group and in the key (see below). In many characters of the structure of abdomen and male genitalia, the 
<taxonomicName id="CF964D68C028FFC9FDD9A9BCFCC39D3C" authorityName="Gyllenhal" authorityYear="1827" box="[632,790,1268,1294]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="canaliculatus">
<emphasis id="3AE2EAF9C028FFC9FDD9A9BCFCC39D3C" box="[632,790,1268,1294]" italics="true" pageId="4" pageNumber="5">canaliculatus</emphasis>
</taxonomicName>
group also resembles the aggregate including the large and diverse 
<emphasis id="3AE2EAF9C028FFC9FE2FA85FFCCD9D03" box="[398,792,1303,1330]" italics="true" pageId="4" pageNumber="5">atratulus -fuscipes -cautus -crassus</emphasis>
complex and some isolated American members, as 
<emphasis id="3AE2EAF9C028FFC9FF1CA873FF199D67" box="[189,204,1339,1365]" italics="true" pageId="4" pageNumber="5">S</emphasis>
. (
<emphasis id="3AE2EAF9C028FFC9FF43A874FF239D67" box="[226,246,1340,1365]" italics="true" pageId="4" pageNumber="5">N</emphasis>
.) 
<emphasis id="3AE2EAF9C028FFC9FEACA873FE549D67" box="[269,385,1339,1365]" italics="true" pageId="4" pageNumber="5">mendosus</emphasis>
<bibRefCitation id="6C074B1AC028FFC9FE27A873FDC49D67" author="Puthz V." box="[390,529,1339,1365]" pageId="4" pageNumber="5" pagination="74 - 121" refId="ref43744" refString="Puthz V. 1971 a. Kritische Faunistik der bisher aus Mitteleuropa bekannten Stenus - Arten nebst systematischen Bemerkungen und Neubeschreibungen (Coleoptera, Staphylinidae). 80. Beitrag zur Kenntnis der Steninen. Entomologische Blatter fur Biologie und Systematik der Kafer 67 (2): 74 - 121." type="journal article" year="1971">Puthz, 1971</bibRefCitation>
(
<bibRefCitation id="6C074B1AC028FFC9FDBFA873FDBF9D64" author="Puthz V." box="[542,618,1339,1366]" pageId="4" pageNumber="5" pagination="20 - 21" refId="ref43903" refString="Puthz V. 1971 d. 104. Beitrag zur Kenntnis der Steninen. Mitteilungen der Deutschen Entomologischen Gesellschaft 30 (2): 20 - 21." type="journal article" year="1971">1971d</bibRefCitation>
) and 
<emphasis id="3AE2EAF9C028FFC9FD09A873FD629D67" box="[680,695,1339,1365]" italics="true" pageId="4" pageNumber="5">S</emphasis>
. (
<emphasis id="3AE2EAF9C028FFC9FD6FA874FD379D67" box="[718,738,1340,1365]" italics="true" pageId="4" pageNumber="5">N</emphasis>
.) 
<emphasis id="3AE2EAF9C028FFC9FD59A873FC899D67" box="[760,860,1339,1365]" italics="true" pageId="4" pageNumber="5">sordidus</emphasis>
<bibRefCitation id="6C074B1AC028FFC9FCC0A873FC3B9D67" author="Puthz" box="[865,1006,1339,1365]" firstAuthor="Puthz" pageId="4" pageNumber="5" pagination="132 - 164" refId="ref44399" refString="Puthz V. 1988. Revision der nearktischen Steninenfauna 3 (Coleoptera, Staphylinidae). Neue Arten und Unterarten aus Nordamerika. Entomologische Blatter fur Biologie und Systematik der Kafer 84 (3): 132 - 164." type="journal article" year="1988">Puthz, 1988</bibRefCitation>
, but phylogenetic relations within the named aggregate remain poorly understood; therefore a thorough analysis seems to be premature. In external morphology, the 
<taxonomicName id="CF964D68C028FFC9FDADA8C9FD7F9DA9" authorityName="Gyllenhal" authorityYear="1827" box="[524,682,1409,1435]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="canaliculatus">
<emphasis id="3AE2EAF9C028FFC9FDADA8C9FD7F9DA9" box="[524,682,1409,1435]" italics="true" pageId="4" pageNumber="5">canaliculatus</emphasis>
</taxonomicName>
group can easily be distinguished from all the species of the mentioned complex by the presence of the long and sharp, line-shaped, longitudinal median furrow of the pronotum.
</paragraph>
</subSubSection>
<subSubSection id="408C6560C028FFCBFF1CAB41FE9A9E5D" lastPageId="6" lastPageNumber="7" pageId="4" pageNumber="5" type="discussion">
<paragraph id="082936EBC028FFC9FF1CAB41FEB89E11" blockId="4.[189,365,1545,1571]" box="[189,365,1545,1571]" pageId="4" pageNumber="5">
<heading id="53618187C028FFC9FF1CAB41FEB89E11" bold="true" box="[189,365,1545,1571]" fontSize="11" level="3" pageId="4" pageNumber="5" reason="3">
<emphasis id="3AE2EAF9C028FFC9FF1CAB41FEB89E11" bold="true" box="[189,365,1545,1571]" pageId="4" pageNumber="5">Taxa included</emphasis>
</heading>
</paragraph>
<paragraph id="082936EBC028FFC9FF1CAB70FB109F59" blockId="4.[189,1399,1591,1900]" pageId="4" pageNumber="5">
<taxonomicName id="CF964D68C028FFC9FF1CAB70FEB49E63" authority="Ryvkin, 2012" authorityName="Ryvkin" authorityYear="2012" box="[189,353,1591,1618]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="1" phylum="Arthropoda" rank="species" species="alopex" status="sp. nov." subGenus="Nestus">
<emphasis id="3AE2EAF9C028FFC9FF1CAB70FF199E60" box="[189,204,1592,1618]" italics="true" pageId="4" pageNumber="5">S</emphasis>
. (
<emphasis id="3AE2EAF9C028FFC9FF47AB70FF2F9E63" box="[230,250,1592,1617]" italics="true" pageId="4" pageNumber="5">N</emphasis>
.) 
<emphasis id="3AE2EAF9C028FFC9FEB2AB7FFEB49E63" box="[275,353,1591,1617]" italics="true" pageId="4" pageNumber="5">alopex</emphasis>
</taxonomicName>
<taxonomicNameLabel id="21D15782C028FFC9FECBAB70FE1C9E60" box="[362,457,1592,1618]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
, 
<emphasis id="3AE2EAF9C028FFC9FE77AB70FE309E60" box="[470,485,1592,1618]" italics="true" pageId="4" pageNumber="5">S</emphasis>
. 
<taxonomicName id="CF964D68C028FFC9FE54AB70FCC69E60" authority="Puthz, 1972" authorityName="Puthz" authorityYear="1972" box="[501,787,1591,1618]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="brivioi" subGenus="Nestus">
(
<emphasis id="3AE2EAF9C028FFC9FE5FAB70FDC79E63" box="[510,530,1592,1617]" italics="true" pageId="4" pageNumber="5">N</emphasis>
.) 
<emphasis id="3AE2EAF9C028FFC9FD8DAB7FFDAF9E63" box="[556,634,1591,1617]" italics="true" pageId="4" pageNumber="5">brivioi</emphasis>
<bibRefCitation id="6C074B1AC028FFC9FD22AB7FFCC69E60" author="Puthz V." box="[643,787,1591,1618]" pageId="4" pageNumber="5" pagination="239 - 292" refId="ref43931" refString="Puthz V. 1972 a. Zur Staphylinidenfauna des Balkans: die bisher aus Jugoslawien und angrenzenden Landern bekannten Steninen (Coleoptera. Staphylinidae). 63. [83!] Beitrag zur Kenntnis der Steninen. Wissenschafliche Mitteilungen des Bosnisch-herzegovinischen Landesmuseums 1 (C): 239 - 292." type="book chapter" year="1972">Puthz, 1972</bibRefCitation>
</taxonomicName>
, 
<emphasis id="3AE2EAF9C028FFC9FC80AB70FCE59E60" box="[801,816,1592,1618]" italics="true" pageId="4" pageNumber="5">S</emphasis>
. (
<emphasis id="3AE2EAF9C028FFC9FCE8AB70FC889E63" box="[841,861,1592,1617]" italics="true" pageId="4" pageNumber="5">N</emphasis>
.) 
<emphasis id="3AE2EAF9C028FFC9FCD6AB7FFBC09E63" box="[887,1045,1591,1617]" italics="true" pageId="4" pageNumber="5">canaliculatus</emphasis>
Gyllenhal, 
<taxonomicName id="CF964D68C028FFC9FB00AB70FA999E63" authority="Ryvkin, 2012" authorityName="Ryvkin" authorityYear="2012" box="[1185,1356,1591,1618]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="33" phylum="Arthropoda" rank="species" species="canalis" status="sp. nov." subGenus="Nestus">
<emphasis id="3AE2EAF9C028FFC9FB00AB70FB659E60" box="[1185,1200,1592,1618]" italics="true" pageId="4" pageNumber="5">S</emphasis>
. (
<emphasis id="3AE2EAF9C028FFC9FB6BAB70FB0B9E63" box="[1226,1246,1592,1617]" italics="true" pageId="4" pageNumber="5">N</emphasis>
.) 
<emphasis id="3AE2EAF9C028FFC9FB56AB7FFA999E63" box="[1271,1356,1591,1617]" italics="true" pageId="4" pageNumber="5">canalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="21D15782C028FFC9FAF4AB70FF249E47" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
, 1827, 
<taxonomicName id="CF964D68C028FFC9FEEDAB13FDD19E47" authority="Ryvkin, 2012" authorityName="Ryvkin" authorityYear="2012" box="[332,516,1627,1653]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="33" phylum="Arthropoda" rank="species" species="canosus" status="sp. nov." subGenus="Nestus">
<emphasis id="3AE2EAF9C028FFC9FEEDAB13FE8E9E47" box="[332,347,1627,1653]" italics="true" pageId="4" pageNumber="5">S</emphasis>
. (
<emphasis id="3AE2EAF9C028FFC9FED7AB14FE5F9E47" box="[374,394,1628,1653]" italics="true" pageId="4" pageNumber="5">N</emphasis>
.) 
<emphasis id="3AE2EAF9C028FFC9FE04AB14FDD19E47" box="[421,516,1628,1653]" italics="true" pageId="4" pageNumber="5">canosus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="21D15782C028FFC9FDAFAB13FDBB9E47" box="[526,622,1627,1653]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
, 
<taxonomicName id="CF964D68C028FFC9FDDCAB13FC6E9E47" authority="Puthz, 1972" authorityName="Puthz" authorityYear="1972" box="[637,955,1627,1653]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="33" phylum="Arthropoda" rank="species" species="caseyi" subGenus="Nestus">
<emphasis id="3AE2EAF9C028FFC9FDDCAB13FD599E47" box="[637,652,1627,1653]" italics="true" pageId="4" pageNumber="5">S</emphasis>
. (
<emphasis id="3AE2EAF9C028FFC9FD06AB14FD6E9E47" box="[679,699,1628,1653]" italics="true" pageId="4" pageNumber="5">N</emphasis>
.) 
<emphasis id="3AE2EAF9C028FFC9FD77AB14FCCA9E47" box="[726,799,1628,1653]" italics="true" pageId="4" pageNumber="5">caseyi</emphasis>
<bibRefCitation id="6C074B1AC028FFC9FC88AB13FC6E9E47" author="Puthz V." box="[809,955,1627,1653]" pageId="4" pageNumber="5" pagination="239 - 292" refId="ref43931" refString="Puthz V. 1972 a. Zur Staphylinidenfauna des Balkans: die bisher aus Jugoslawien und angrenzenden Landern bekannten Steninen (Coleoptera. Staphylinidae). 63. [83!] Beitrag zur Kenntnis der Steninen. Wissenschafliche Mitteilungen des Bosnisch-herzegovinischen Landesmuseums 1 (C): 239 - 292." type="book chapter" year="1972">Puthz, 1972</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="CF964D68C028FFC9FC6BAB13FAA79E47" authority="J. Sahlberg, 1876" authorityName="J. Sahlberg" authorityYear="1876" box="[970,1394,1627,1653]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="14" phylum="Arthropoda" rank="species" species="confusus" subGenus="Nestus">
<emphasis id="3AE2EAF9C028FFC9FC6BAB13FC0C9E47" box="[970,985,1627,1653]" italics="true" pageId="4" pageNumber="5">S</emphasis>
. (
<emphasis id="3AE2EAF9C028FFC9FC55AB14FBDD9E47" box="[1012,1032,1628,1653]" italics="true" pageId="4" pageNumber="5">N</emphasis>
.) 
<emphasis id="3AE2EAF9C028FFC9FB82AB13FB5F9E47" box="[1059,1162,1627,1653]" italics="true" pageId="4" pageNumber="5">confusus</emphasis>
J. 
<bibRefCitation id="6C074B1AC028FFC9FB1AAB13FAA79E47" author="Sahlberg" box="[1211,1394,1627,1653]" firstAuthor="Sahlberg" pageId="4" pageNumber="5" pagination="1 - 248" refId="ref45221" refString="Sahlberg J. R. 1876. Enumeratio Coleopterorum Brachelytrorum Fenniae. Systematisk forteckning ofver de inom Finlands naturalhistoriska omrade hittils funna Coleoptera Brachelytra jemte uppgift om arternas utbredning och &lt;...&gt;. I. Staphylinidae. Acta Societatis pro Fauna et Flora Fennica 1: 1 - 248." type="journal article" year="1876">Sahlberg, 1876</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="CF964D68C028FFC9FF1CAB36FE899EAA" authority="Ryvkin, 2012" authorityName="Ryvkin" authorityYear="2012" box="[189,348,1662,1688]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="33" phylum="Arthropoda" rank="species" species="delitor" status="sp. nov." subGenus="Nestus">
<emphasis id="3AE2EAF9C028FFC9FF1CAB36FF199EAA" box="[189,204,1662,1688]" italics="true" pageId="4" pageNumber="5">S</emphasis>
. (
<emphasis id="3AE2EAF9C028FFC9FF42AB37FF229EAA" box="[227,247,1663,1688]" italics="true" pageId="4" pageNumber="5">N</emphasis>
.) 
<emphasis id="3AE2EAF9C028FFC9FEAFAB36FE899EAA" box="[270,348,1662,1688]" italics="true" pageId="4" pageNumber="5">delitor</emphasis>
</taxonomicName>
<taxonomicNameLabel id="21D15782C028FFC9FEC3AB36FE6B9EAA" box="[354,446,1662,1688]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
, 
<emphasis id="3AE2EAF9C028FFC9FE68AB36FE0D9EAA" box="[457,472,1662,1688]" italics="true" pageId="4" pageNumber="5">S</emphasis>
. 
<taxonomicName id="CF964D68C028FFC9FE44AB36FCDC9EAA" authority="Casey, 1884" authorityName="Casey" authorityYear="1884" box="[485,777,1662,1688]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="dolosus" subGenus="Nestus">
(
<emphasis id="3AE2EAF9C028FFC9FE4FAB37FDD79EAA" box="[494,514,1663,1688]" italics="true" pageId="4" pageNumber="5">N</emphasis>
.) 
<emphasis id="3AE2EAF9C028FFC9FDB8AB36FDA69EAA" box="[537,627,1662,1688]" italics="true" pageId="4" pageNumber="5">dolosus</emphasis>
<bibRefCitation id="6C074B1AC028FFC9FDD8AB36FCDC9EAA" author="Casey" box="[633,777,1662,1688]" firstAuthor="Casey" pageId="4" pageNumber="5" refId="ref40664" refString="Casey Th. L. 1884. Revision of the Stenini of America north of Mexico. Insects of the family Staphylinidae, order Coleoptera. Collins Printing House, Philadelphia. 206 pp. http: // dx. doi. org / 10.5962 / bhl. title. 9208" type="book" year="1884">Casey, 1884</bibRefCitation>
</taxonomicName>
, 
<emphasis id="3AE2EAF9C028FFC9FCB5AB36FCF69EAA" box="[788,803,1662,1688]" italics="true" pageId="4" pageNumber="5">S</emphasis>
. 
<taxonomicName id="CF964D68C028FFC9FC90AB36FBAB9EAA" authority="Puthz, 1973" authorityName="Puthz" authorityYear="1973" box="[817,1150,1662,1688]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="geminorum" subGenus="Nestus">
(
<emphasis id="3AE2EAF9C028FFC9FC9BAB37FC9B9EAA" box="[826,846,1663,1688]" italics="true" pageId="4" pageNumber="5">N</emphasis>
.) 
<emphasis id="3AE2EAF9C028FFC9FCC4AB37FC3E9EAA" box="[869,1003,1663,1688]" italics="true" pageId="4" pageNumber="5">geminorum</emphasis>
<bibRefCitation id="6C074B1AC028FFC9FC50AB36FBAB9EAA" author="Puthz V." box="[1009,1150,1662,1688]" pageId="4" pageNumber="5" pagination="23 - 31" refId="ref44130" refString="Puthz V. 1973 a. Die von Claudius Rey beschreibenen Stenus - Arten (Coleoptera, Staphylinidae). 120. Beitrag zur Kenntnis der Steninen. Nouvelle Revue d'Entomologie 3 (1): 23 - 31." type="journal article" year="1973">Puthz, 1973</bibRefCitation>
</taxonomicName>
* (USA: Washington; Canada: BC, see 
<bibRefCitation id="6C074B1AC028FFC9FE34ABE9FDA49E89" author="Puthz" box="[405,625,1697,1723]" firstAuthor="Puthz" page="208" pageId="4" pageNumber="5" pagination="189 - 209" refId="ref44170" refString="Puthz V. 1973 b. Revision der nearktischen Steninenfauna (I). Neue nordamericanische Stenus - (s. str. + Nestus) Arten. 133. Beitrag zur Kenntnis der Steninen. Entomologische Blatter fur Biologie und Systematik der Kafer 69 (3): 189 - 209." type="journal article" year="1973" yearSuffix="b">Puthz 1973b: 208</bibRefCitation>
), 
<emphasis id="3AE2EAF9C028FFC9FD2BABEAFD4C9E8E" box="[650,665,1698,1724]" italics="true" pageId="4" pageNumber="5">S</emphasis>
. 
<taxonomicName id="CF964D68C028FFC9FD0DABE9FC369E89" authority="Sharp, 1887" authorityName="Sharp" authorityYear="1887" box="[684,995,1697,1724]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="idoneus" subGenus="Nestus">
(
<emphasis id="3AE2EAF9C028FFC9FD17ABEAFD1F9E89" box="[694,714,1698,1723]" italics="true" pageId="4" pageNumber="5">N</emphasis>
.) 
<emphasis id="3AE2EAF9C028FFC9FD46ABE9FC979E89" box="[743,834,1697,1723]" italics="true" pageId="4" pageNumber="5">idoneus</emphasis>
<bibRefCitation id="6C074B1AC028FFC9FCEFABE9FC369E89" author="Sharp" box="[846,995,1697,1723]" firstAuthor="Sharp" pageId="4" pageNumber="5" pagination="673 - 824" refId="ref45684" refString="Sharp D. 1887. Staphylinidae. In: Biologia Centrali-Americana. Insecta. Coleoptera. 1 (2): 673 - 824. Taylor &amp; Francis, London." type="journal article" year="1887">Sharp, 1887</bibRefCitation>
</taxonomicName>
* (Mexico, see 
<bibRefCitation id="6C074B1AC028FFC9FB02ABE9FAA79E89" author="Sharp" box="[1187,1394,1697,1723]" firstAuthor="Sharp" page="799" pageId="4" pageNumber="5" pagination="673 - 824" refId="ref45684" refString="Sharp D. 1887. Staphylinidae. In: Biologia Centrali-Americana. Insecta. Coleoptera. 1 (2): 673 - 824. Taylor &amp; Francis, London." type="journal article" year="1887">Sharp 1887: 799</bibRefCitation>
; 
<bibRefCitation id="6C074B1AC028FFC9FF1CAB8CFEA49EEC" author="Puthz" box="[189,369,1732,1758]" firstAuthor="Puthz" page="11" pageId="4" pageNumber="5" pagination="1 - 13" refId="ref43597" refString="Puthz V. 1968. Neue und alte Mittelamerikanische Steninen (Coleoptera. Staphylinidae). 40. Beitrag zur Kenntnis der Steninen. Annotationes Zoologicae et Botanicae 48 (1967): 1 - 13." type="journal article" year="1968">Puthz 1968: 11</bibRefCitation>
&lt;as 
<emphasis id="3AE2EAF9C028FFC9FE0CAB8CFDD79EEC" box="[429,514,1732,1758]" italics="true" pageId="4" pageNumber="5">laniger</emphasis>
<bibRefCitation id="6C074B1AC028FFC9FDAAAB8CFD499EED" author="Puthz" box="[523,668,1732,1759]" firstAuthor="Puthz" pageId="4" pageNumber="5" pagination="1 - 13" refId="ref43597" refString="Puthz V. 1968. Neue und alte Mittelamerikanische Steninen (Coleoptera. Staphylinidae). 40. Beitrag zur Kenntnis der Steninen. Annotationes Zoologicae et Botanicae 48 (1967): 1 - 13." type="journal article" year="1968">Puthz, 1968</bibRefCitation>
&gt;), 
<taxonomicName id="CF964D68C028FFC9FD62AB8DFBC19EEC" authority="Puthz, 1972" authorityName="Puthz" authorityYear="1972" box="[707,1044,1732,1759]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="14" phylum="Arthropoda" rank="species" species="illotulus" subGenus="Nestus">
<emphasis id="3AE2EAF9C028FFC9FD62AB8DFD079EED" box="[707,722,1733,1759]" italics="true" pageId="4" pageNumber="5">S</emphasis>
. (
<emphasis id="3AE2EAF9C028FFC9FD4DAB8DFCD59EEC" box="[748,768,1733,1758]" italics="true" pageId="4" pageNumber="5">N</emphasis>
.) 
<emphasis id="3AE2EAF9C028FFC9FCBBAB8CFCAF9EEC" box="[794,890,1732,1758]" italics="true" pageId="4" pageNumber="5">illotulus</emphasis>
<bibRefCitation id="6C074B1AC028FFC9FC25AB8CFBC19EEC" author="Puthz V." box="[900,1044,1732,1758]" pageId="4" pageNumber="5" pagination="239 - 292" refId="ref43931" refString="Puthz V. 1972 a. Zur Staphylinidenfauna des Balkans: die bisher aus Jugoslawien und angrenzenden Landern bekannten Steninen (Coleoptera. Staphylinidae). 63. [83!] Beitrag zur Kenntnis der Steninen. Wissenschafliche Mitteilungen des Bosnisch-herzegovinischen Landesmuseums 1 (C): 239 - 292." type="book chapter" year="1972">Puthz, 1972</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="CF964D68C028FFC9FB83AB8DFAA79EEC" authority="Ryvkin, 1987" authorityName="Ryvkin" authorityYear="1987" box="[1058,1394,1732,1759]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="42" phylum="Arthropoda" rank="species" species="illusor" subGenus="Nestus">
<emphasis id="3AE2EAF9C028FFC9FB83AB8DFBE49EED" box="[1058,1073,1733,1759]" italics="true" pageId="4" pageNumber="5">S</emphasis>
. (
<emphasis id="3AE2EAF9C028FFC9FBEAAB8DFB8A9EEC" box="[1099,1119,1733,1758]" italics="true" pageId="4" pageNumber="5">N</emphasis>
.) 
<emphasis id="3AE2EAF9C028FFC9FBD8AB8CFB139EEC" box="[1145,1222,1732,1758]" italics="true" pageId="4" pageNumber="5">illusor</emphasis>
<bibRefCitation id="6C074B1AC028FFC9FB6EAB8CFAA79EEC" author="Ryvkin" box="[1231,1394,1732,1758]" firstAuthor="Ryvkin" pageId="4" pageNumber="5" pagination="149 - 161" refId="ref44999" refString="Ryvkin A. B. 1987. New species of Stenus Latreille 1796 from the Far East of the Soviet Union (Insecta: Coleoptera: Staphylinidae). Senckenbergiana biologica 68 (1 / 3): 149 - 161." type="journal article" year="1987">Ryvkin, 1987</bibRefCitation>
</taxonomicName>
, 
<emphasis id="3AE2EAF9C028FFC9FF1CABA0FF199F30" box="[189,204,1768,1794]" italics="true" pageId="4" pageNumber="5">S</emphasis>
. 
<taxonomicName id="CF964D68C028FFC9FF7AABA0FDE59F33" authority="Puthz, 2006" authorityName="Puthz" authorityYear="2006" box="[219,560,1767,1794]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="immigratus" subGenus="Nestus">
(
<emphasis id="3AE2EAF9C028FFC9FF44ABA0FF2C9F33" box="[229,249,1768,1793]" italics="true" pageId="4" pageNumber="5">N</emphasis>
.) 
<emphasis id="3AE2EAF9C028FFC9FEB3ABA0FE4D9F33" box="[274,408,1768,1793]" italics="true" pageId="4" pageNumber="5">immigratus</emphasis>
<bibRefCitation id="6C074B1AC028FFC9FE01ABAFFDE59F33" author="Puthz" box="[416,560,1767,1793]" firstAuthor="Puthz" pageId="4" pageNumber="5" pagination="171 - 196" refId="ref44527" refString="Puthz V. 2006. Ein Dutzend neuer palaarktischer Stenus - Arten (Coleoptera, Staphylinidae). 290. Beitrag zur Kenntnis der Steninen. Entomologische Blatter fur Biologie und Systematik der Kafer 101 (2005) (2 - 3): 171 - 196." type="journal article" year="2006">Puthz, 2006</bibRefCitation>
</taxonomicName>
* (China: Taiwan: Ilan Hsien: Shen Mi Lake, 
<quantity id="CF6E9B0EC028FFC9FBEAABA0FB739F30" box="[1099,1190,1768,1794]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="4" pageNumber="5" unit="m" value="1100.0">1100 m</quantity>
, see 
<bibRefCitation id="6C074B1AC028FFC9FB40ABAFFF399F17" author="Puthz" firstAuthor="Puthz" page="187" pageId="4" pageNumber="5" pagination="171 - 196" refId="ref44527" refString="Puthz V. 2006. Ein Dutzend neuer palaarktischer Stenus - Arten (Coleoptera, Staphylinidae). 290. Beitrag zur Kenntnis der Steninen. Entomologische Blatter fur Biologie und Systematik der Kafer 101 (2005) (2 - 3): 171 - 196." type="journal article" year="2006">Puthz, 2006: 187</bibRefCitation>
), 
<taxonomicName id="CF964D68C028FFC9FEA0AA43FD839F17" authority="Erichson, 1840" authorityName="Erichson" authorityYear="1840" box="[257,598,1803,1829]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="9" phylum="Arthropoda" rank="species" species="labilis" subGenus="Nestus">
<emphasis id="3AE2EAF9C028FFC9FEA0AA43FEC59F17" box="[257,272,1803,1829]" italics="true" pageId="4" pageNumber="5">S</emphasis>
. (
<emphasis id="3AE2EAF9C028FFC9FE86AA44FEEE9F17" box="[295,315,1804,1829]" italics="true" pageId="4" pageNumber="5">N</emphasis>
.) 
<emphasis id="3AE2EAF9C028FFC9FEF3AA43FE4E9F17" box="[338,411,1803,1829]" italics="true" pageId="4" pageNumber="5">labilis</emphasis>
<bibRefCitation id="6C074B1AC028FFC9FE03AA43FD839F17" author="Erichson" box="[418,598,1803,1829]" firstAuthor="Erichson" pageId="4" pageNumber="5" refId="ref40960" refString="Erichson W. F. 1840. Genera et species staphylinorum insectorum coleopterorum familiae: 401 - 954. F. H. Morin, Berlin. http: // dx. doi. org / 10.5962 / bhl. title. 59644" type="book" year="1840">Erichson, 1840</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="CF964D68C028FFC9FDC0AA43FBD59F17" authority="J. Sahlberg, 1880" authorityName="J. Sahlberg" authorityYear="1880" box="[609,1024,1803,1829]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="42" phylum="Arthropoda" rank="species" species="latipennis" subGenus="Nestus">
<emphasis id="3AE2EAF9C028FFC9FDC0AA43FDA59F17" box="[609,624,1803,1829]" italics="true" pageId="4" pageNumber="5">S</emphasis>
. (
<emphasis id="3AE2EAF9C028FFC9FD26AA44FD4E9F17" box="[647,667,1804,1829]" italics="true" pageId="4" pageNumber="5">N</emphasis>
.) 
<emphasis id="3AE2EAF9C028FFC9FD12AA43FCF29F17" box="[691,807,1803,1829]" italics="true" pageId="4" pageNumber="5">latipennis</emphasis>
J. 
<bibRefCitation id="6C074B1AC028FFC9FCECAA43FBD59F17" author="Sahlberg" box="[845,1024,1803,1829]" firstAuthor="Sahlberg" pageId="4" pageNumber="5" pagination="1 - 115" refId="ref45272" refString="Sahlberg J. R. 1880. Bidrag till Nordvestra Sibiriens Insektfauna. Coleoptera. Insamlade under Expeditionerna till Obi och Jenissej 1876 och 1877. I. Cicindelidae, Carabidae, Dytiscidae, Hydrophilidae, Gyrinidae, [...] Staphylinidae och Micropepeplidae. Kongliga Svenska Vetenskaps- Akademiens Handlingar 17 (4): 1 - 115." type="journal article" year="1880">Sahlberg, 1880</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="CF964D68C028FFC9FBAAAA43FAA79F17" authority="Casey, 1884" authorityName="Casey" authorityYear="1884" box="[1035,1394,1803,1829]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="1" phylum="Arthropoda" rank="species" species="milleporus" subGenus="Nestus">
<emphasis id="3AE2EAF9C028FFC9FBAAAA43FBCF9F17" box="[1035,1050,1803,1829]" italics="true" pageId="4" pageNumber="5">S</emphasis>
. (
<emphasis id="3AE2EAF9C028FFC9FB90AA44FB909F17" box="[1073,1093,1804,1829]" italics="true" pageId="4" pageNumber="5">N</emphasis>
.) 
<emphasis id="3AE2EAF9C028FFC9FBFCAA43FB0F9F17" box="[1117,1242,1803,1829]" italics="true" pageId="4" pageNumber="5">milleporus</emphasis>
<bibRefCitation id="6C074B1AC028FFC9FB40AA43FAA79F17" author="Casey" box="[1249,1394,1803,1829]" firstAuthor="Casey" pageId="4" pageNumber="5" refId="ref40664" refString="Casey Th. L. 1884. Revision of the Stenini of America north of Mexico. Insects of the family Staphylinidae, order Coleoptera. Collins Printing House, Philadelphia. 206 pp. http: // dx. doi. org / 10.5962 / bhl. title. 9208" type="book" year="1884">Casey, 1884</bibRefCitation>
</taxonomicName>
, 
<emphasis id="3AE2EAF9C028FFC9FF1CAA66FF199F7A" box="[189,204,1838,1864]" italics="true" pageId="4" pageNumber="5">S</emphasis>
. 
<taxonomicName id="CF964D68C028FFC9FF7BAA66FDC49F7A" authority="Stephens, 1833" authorityName="Stephens" authorityYear="1833" box="[218,529,1838,1864]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="nitens" subGenus="Nestus">
(
<emphasis id="3AE2EAF9C028FFC9FF45AA67FF2D9F7A" box="[228,248,1839,1864]" italics="true" pageId="4" pageNumber="5">N</emphasis>
.) 
<emphasis id="3AE2EAF9C028FFC9FEAEAA67FE809F7A" box="[271,341,1839,1864]" italics="true" pageId="4" pageNumber="5">nitens</emphasis>
<bibRefCitation id="6C074B1AC028FFC9FEFDAA66FDC49F7A" author="Stephens" box="[348,529,1838,1864]" firstAuthor="Stephens" pageId="4" pageNumber="5" refId="ref46202" refString="Stephens J. F. 1833. Illustrations of British Entomology; or a synopsis of indigenous insects: containing their generic and specific distinctions; with an account of their metamorphoses, times of appearance, localities, food, and economy, as far as practicable. Mandibulata. Vol. V: 241 - 304. Baldwin and Cradock, London. http: // dx. doi. org / 10.5962 / bhl. title. 8133" type="book" year="1833">Stephens, 1833</bibRefCitation>
</taxonomicName>
, 
<emphasis id="3AE2EAF9C028FFC9FDBCAA66FDF99F7A" box="[541,556,1838,1864]" italics="true" pageId="4" pageNumber="5">S</emphasis>
. 
<taxonomicName id="CF964D68C028FFC9FD9BAA66FCB69F7A" authority="Ryvkin, 1987" authorityName="Ryvkin" authorityYear="1987" box="[570,867,1838,1864]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="raddei" subGenus="Nestus">
(
<emphasis id="3AE2EAF9C028FFC9FDE5AA67FD8D9F7A" box="[580,600,1839,1864]" italics="true" pageId="4" pageNumber="5">N</emphasis>
.) 
<emphasis id="3AE2EAF9C028FFC9FDCEAA66FD699F7A" box="[623,700,1838,1864]" italics="true" pageId="4" pageNumber="5">raddei</emphasis>
<bibRefCitation id="6C074B1AC028FFC9FD62AA66FCB69F7A" author="Ryvkin" box="[707,867,1838,1864]" firstAuthor="Ryvkin" pageId="4" pageNumber="5" pagination="149 - 161" refId="ref44999" refString="Ryvkin A. B. 1987. New species of Stenus Latreille 1796 from the Far East of the Soviet Union (Insecta: Coleoptera: Staphylinidae). Senckenbergiana biologica 68 (1 / 3): 149 - 161." type="journal article" year="1987">Ryvkin, 1987</bibRefCitation>
</taxonomicName>
, 
<emphasis id="3AE2EAF9C028FFC9FCCEAA66FCAB9F7A" box="[879,894,1838,1864]" italics="true" pageId="4" pageNumber="5">S</emphasis>
. 
<taxonomicName id="CF964D68C028FFC9FC2DAA66FB7E9F7A" authority="Puthz, 1987" authorityName="Puthz" authorityYear="1987" box="[908,1195,1838,1864]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="shogun" subGenus="Nestus">
(
<emphasis id="3AE2EAF9C028FFC9FC37AA67FC7F9F7A" box="[918,938,1839,1864]" italics="true" pageId="4" pageNumber="5">N</emphasis>
.) 
<emphasis id="3AE2EAF9C028FFC9FC60AA66FBC39F7A" box="[961,1046,1838,1864]" italics="true" pageId="4" pageNumber="5">shogun</emphasis>
<bibRefCitation id="6C074B1AC028FFC9FBBCAA66FB7E9F7A" author="Puthz" box="[1053,1195,1838,1864]" firstAuthor="Puthz" pageId="4" pageNumber="5" pagination="45 - 48" refId="ref44353" refString="Puthz V. 1987. Ein neuer Stenus (Col. Staphylinidae) aus Japan. 208. Beitrag zur Kenntnis der Steninen. Mitteilungen des Internationalen Entomologischen Vereins, Frankfurt a. M. 11 (2 / 3): 45 - 48." type="journal article" year="1987">Puthz, 1987</bibRefCitation>
</taxonomicName>
* (Japan: Nishigo Uzan, see 
<bibRefCitation id="6C074B1AC028FFC9FE97AA19FE399F5E" author="Puthz" box="[310,492,1873,1900]" firstAuthor="Puthz" page="45" pageId="4" pageNumber="5" pagination="45 - 48" refId="ref44353" refString="Puthz V. 1987. Ein neuer Stenus (Col. Staphylinidae) aus Japan. 208. Beitrag zur Kenntnis der Steninen. Mitteilungen des Internationalen Entomologischen Vereins, Frankfurt a. M. 11 (2 / 3): 45 - 48." type="journal article" year="1987">Puthz 1987: 45</bibRefCitation>
), 
<taxonomicName id="CF964D68C028FFC9FDA1AA1AFCB19F59" authority="Casey, 1884" authorityName="Casey" authorityYear="1884" box="[512,868,1873,1900]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="42" phylum="Arthropoda" rank="species" species="sphaerops" subGenus="Nestus">
<emphasis id="3AE2EAF9C028FFC9FDA1AA1AFDDA9F5E" box="[512,527,1874,1900]" italics="true" pageId="4" pageNumber="5">S</emphasis>
. (
<emphasis id="3AE2EAF9C028FFC9FD89AA1AFDE99F59" box="[552,572,1874,1899]" italics="true" pageId="4" pageNumber="5">N</emphasis>
.) 
<emphasis id="3AE2EAF9C028FFC9FDF5AA19FD199F59" box="[596,716,1873,1899]" italics="true" pageId="4" pageNumber="5">sphaerops</emphasis>
<bibRefCitation id="6C074B1AC028FFC9FD73AA19FCB19F59" author="Casey" box="[722,868,1873,1899]" firstAuthor="Casey" pageId="4" pageNumber="5" refId="ref40664" refString="Casey Th. L. 1884. Revision of the Stenini of America north of Mexico. Insects of the family Staphylinidae, order Coleoptera. Collins Printing House, Philadelphia. 206 pp. http: // dx. doi. org / 10.5962 / bhl. title. 9208" type="book" year="1884">Casey, 1884</bibRefCitation>
</taxonomicName>
, 
<emphasis id="3AE2EAF9C028FFC9FCD1AA1AFCAA9F5E" box="[880,895,1874,1900]" italics="true" pageId="4" pageNumber="5">S</emphasis>
. 
<taxonomicName id="CF964D68C028FFC9FC2FAA19FB149F59" authority="Casey, 1884" authorityName="Casey" authorityYear="1884" box="[910,1217,1873,1900]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="vinnulus" subGenus="Nestus">
(
<emphasis id="3AE2EAF9C028FFC9FC36AA1AFC7E9F59" box="[919,939,1874,1899]" italics="true" pageId="4" pageNumber="5">N</emphasis>
.) 
<emphasis id="3AE2EAF9C028FFC9FC62AA19FBF39F59" box="[963,1062,1873,1899]" italics="true" pageId="4" pageNumber="5">vinnulus</emphasis>
<bibRefCitation id="6C074B1AC028FFC9FB8FAA19FB149F59" author="Casey" box="[1070,1217,1873,1899]" firstAuthor="Casey" pageId="4" pageNumber="5" refId="ref40664" refString="Casey Th. L. 1884. Revision of the Stenini of America north of Mexico. Insects of the family Staphylinidae, order Coleoptera. Collins Printing House, Philadelphia. 206 pp. http: // dx. doi. org / 10.5962 / bhl. title. 9208" type="book" year="1884">Casey, 1884</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="082936EBC029FFC8FF1CAC43FCCB9917" blockId="5.[189,798,267,293]" box="[189,798,267,293]" pageId="5" pageNumber="6">
<heading id="53618187C029FFC8FF1CAC43FCCB9917" bold="true" box="[189,798,267,293]" fontSize="11" level="3" pageId="5" pageNumber="6" reason="3">
<emphasis id="3AE2EAF9C029FFC8FF1CAC43FCCB9917" bold="true" box="[189,798,267,293]" pageId="5" pageNumber="6">Notes on comparative morphology and evolution</emphasis>
</heading>
</paragraph>
<paragraph id="082936EBC029FFC8FF1CAC72FE6899A8" blockId="5.[189,1399,313,410]" pageId="5" pageNumber="6">
As most other polytypic groups in 
<taxonomicName id="CF964D68C029FFC8FDF0AC71FD6C9961" authorityName="W.S.MacLeay" authorityYear="1825" box="[593,697,313,339]" class="Insecta" family="Staphylinidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="subFamily" subFamily="Steninae">Steninae</taxonomicName>
, the 
<taxonomicName id="CF964D68C029FFC8FD4EAC71FC589961" authorityName="Gyllenhal" authorityYear="1827" box="[751,909,313,339]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="canaliculatus">
<emphasis id="3AE2EAF9C029FFC8FD4EAC71FC589961" box="[751,909,313,339]" italics="true" pageId="5" pageNumber="6">canaliculatus</emphasis>
</taxonomicName>
group reveals diverse evolutionary trends quite evident at a comparison between species of the same lineage. A concise review of the mentioned trends is given below.
</paragraph>
<paragraph id="082936EBC029FFC8FF1CAC89FD629AE0" blockId="5.[189,1399,449,722]" pageId="5" pageNumber="6">
A. Upper surface of head between eyes tends either to become flattened, at most slightly convex (
<taxonomicName id="CF964D68C029FFC8FF66ACADFEEC99CC" authorityName="Puthz" authorityYear="1987" box="[199,313,484,511]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="shogun">
<emphasis id="3AE2EAF9C029FFC8FF66ACADFF0399CD" box="[199,214,485,511]" italics="true" pageId="5" pageNumber="6">S</emphasis>
. 
<emphasis id="3AE2EAF9C029FFC8FF45ACACFEEC99CC" box="[228,313,484,510]" italics="true" pageId="5" pageNumber="6">shogun</emphasis>
</taxonomicName>
, 
<taxonomicName id="CF964D68C029FFC8FEE6ACADFE7C99CC" authorityName="Stephens" authorityYear="1833" box="[327,425,485,511]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="nitens">
<emphasis id="3AE2EAF9C029FFC8FEE6ACADFE8399CD" box="[327,342,485,511]" italics="true" pageId="5" pageNumber="6">S</emphasis>
. 
<emphasis id="3AE2EAF9C029FFC8FEC2ACADFE7C99CC" box="[355,425,485,510]" italics="true" pageId="5" pageNumber="6">nitens</emphasis>
</taxonomicName>
, 
<taxonomicName id="CF964D68C029FFC8FE19ACADFDE099CC" authorityName="Puthz" authorityYear="1972" box="[440,565,484,511]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="illotulus">
<emphasis id="3AE2EAF9C029FFC8FE19ACADFE1299CD" box="[440,455,485,511]" italics="true" pageId="5" pageNumber="6">S</emphasis>
. 
<emphasis id="3AE2EAF9C029FFC8FE74ACACFDE099CC" box="[469,565,484,510]" italics="true" pageId="5" pageNumber="6">illotulus</emphasis>
</taxonomicName>
, 
<taxonomicName id="CF964D68C029FFC8FDE2ACADFD7899CC" authorityName="Ryvkin" authorityYear="1987" box="[579,685,484,511]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="raddei">
<emphasis id="3AE2EAF9C029FFC8FDE2ACADFD8799CD" box="[579,594,485,511]" italics="true" pageId="5" pageNumber="6">S</emphasis>
. 
<emphasis id="3AE2EAF9C029FFC8FDC1ACACFD7899CC" box="[608,685,484,510]" italics="true" pageId="5" pageNumber="6">raddei</emphasis>
</taxonomicName>
, 
<taxonomicName id="CF964D68C029FFC8FD1AACADFCF599CC" authorityName="Puthz" authorityYear="1972" box="[699,800,485,511]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="caseyi">
<emphasis id="3AE2EAF9C029FFC8FD1AACADFD1F99CD" box="[699,714,485,511]" italics="true" pageId="5" pageNumber="6">S</emphasis>
. 
<emphasis id="3AE2EAF9C029FFC8FD76ACADFCF599CC" box="[727,800,485,510]" italics="true" pageId="5" pageNumber="6">caseyi</emphasis>
</taxonomicName>
, 
<taxonomicName id="CF964D68C029FFC8FC8EACADFC3F99CC" authorityName="Gyllenhal" authorityYear="1827" box="[815,1002,484,511]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="canaliculatus">
<emphasis id="3AE2EAF9C029FFC8FC8EACADFCEB99CD" box="[815,830,485,511]" italics="true" pageId="5" pageNumber="6">S</emphasis>
. 
<emphasis id="3AE2EAF9C029FFC8FCEDACACFC3F99CC" box="[844,1002,484,510]" italics="true" pageId="5" pageNumber="6">canaliculatus</emphasis>
</taxonomicName>
, 
<taxonomicName id="CF964D68C029FFC8FC59ACADFBBF99CC" authorityName="Ryvkin" authorityYear="2012" box="[1016,1130,484,511]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="canalis">
<emphasis id="3AE2EAF9C029FFC8FC59ACADFBD299CD" box="[1016,1031,485,511]" italics="true" pageId="5" pageNumber="6">S</emphasis>
. 
<emphasis id="3AE2EAF9C029FFC8FBB4ACACFBBF99CC" box="[1045,1130,484,510]" italics="true" pageId="5" pageNumber="6">canalis</emphasis>
</taxonomicName>
, 
<taxonomicName id="CF964D68C029FFC8FBD9ACADFB2199CC" authorityName="Ryvkin" authorityYear="2012" box="[1144,1268,485,511]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="canosus">
<emphasis id="3AE2EAF9C029FFC8FBD9ACADFB5299CD" box="[1144,1159,485,511]" italics="true" pageId="5" pageNumber="6">S</emphasis>
. 
<emphasis id="3AE2EAF9C029FFC8FB34ACADFB2199CC" box="[1173,1268,485,510]" italics="true" pageId="5" pageNumber="6">canosus</emphasis>
</taxonomicName>
, 
<taxonomicName id="CF964D68C029FFC8FAA0ACADFAB999CC" authorityName="Ryvkin" authorityYear="2012" box="[1281,1388,484,511]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="alopex">
<emphasis id="3AE2EAF9C029FFC8FAA0ACADFAC599CD" box="[1281,1296,485,511]" italics="true" pageId="5" pageNumber="6">S</emphasis>
. 
<emphasis id="3AE2EAF9C029FFC8FABFACACFAB999CC" box="[1310,1388,484,510]" italics="true" pageId="5" pageNumber="6">alopex</emphasis>
</taxonomicName>
) or to produce distinct longitudinal impressions with median elevation in between (
<taxonomicName id="CF964D68C029FFC8FB23AF40FB3D9A10" authorityName="Erichson" authorityYear="1840" box="[1154,1256,520,546]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="labilis">
<emphasis id="3AE2EAF9C029FFC8FB23AF40FB449A10" box="[1154,1169,520,546]" italics="true" pageId="5" pageNumber="6">S</emphasis>
. 
<emphasis id="3AE2EAF9C029FFC8FB3EAF40FB3D9A10" box="[1183,1256,520,546]" italics="true" pageId="5" pageNumber="6">labilis</emphasis>
</taxonomicName>
, 
<taxonomicName id="CF964D68C029FFC8FB56AF40FABA9A10" authorityName="Sharp" authorityYear="1887" box="[1271,1391,520,546]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="idoneus">
<emphasis id="3AE2EAF9C029FFC8FB56AF40FAD39A10" box="[1271,1286,520,546]" italics="true" pageId="5" pageNumber="6">S</emphasis>
. 
<emphasis id="3AE2EAF9C029FFC8FAB5AF40FABA9A10" box="[1300,1391,520,546]" italics="true" pageId="5" pageNumber="6">idoneus</emphasis>
</taxonomicName>
, 
<taxonomicName id="CF964D68C029FFC8FF1CAF63FE959A77" authorityName="Casey" authorityYear="1884" box="[189,320,555,581]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="vinnulus">
<emphasis id="3AE2EAF9C029FFC8FF1CAF63FF199A77" box="[189,204,555,581]" italics="true" pageId="5" pageNumber="6">S</emphasis>
. 
<emphasis id="3AE2EAF9C029FFC8FF7CAF63FE959A77" box="[221,320,555,581]" italics="true" pageId="5" pageNumber="6">vinnulus</emphasis>
</taxonomicName>
, 
<taxonomicName id="CF964D68C029FFC8FEF3AF63FE149A77" authorityName="Puthz" authorityYear="1972" box="[338,449,555,581]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="brivioi">
<emphasis id="3AE2EAF9C029FFC8FEF3AF63FEB49A77" box="[338,353,555,581]" italics="true" pageId="5" pageNumber="6">S</emphasis>
. 
<emphasis id="3AE2EAF9C029FFC8FED2AF63FE149A77" box="[371,449,555,581]" italics="true" pageId="5" pageNumber="6">brivioi</emphasis>
</taxonomicName>
, 
<taxonomicName id="CF964D68C029FFC8FE72AF63FD8F9A77" authorityName="J. Sahlberg" authorityYear="1876" box="[467,602,555,581]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="confusus">
<emphasis id="3AE2EAF9C029FFC8FE72AF63FE379A77" box="[467,482,555,581]" italics="true" pageId="5" pageNumber="6">S</emphasis>
. 
<emphasis id="3AE2EAF9C029FFC8FE52AF63FD8F9A77" box="[499,602,555,581]" italics="true" pageId="5" pageNumber="6">confusus</emphasis>
</taxonomicName>
, 
<taxonomicName id="CF964D68C029FFC8FDCDAF63FD0F9A77" authorityName="Ryvkin" authorityYear="2012" box="[620,730,555,581]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="delitor">
<emphasis id="3AE2EAF9C029FFC8FDCDAF63FDAE9A77" box="[620,635,555,581]" italics="true" pageId="5" pageNumber="6">S</emphasis>
. 
<emphasis id="3AE2EAF9C029FFC8FD2DAF63FD0F9A77" box="[652,730,555,581]" italics="true" pageId="5" pageNumber="6">delitor</emphasis>
</taxonomicName>
, 
<taxonomicName id="CF964D68C029FFC8FD4DAF63FC8F9A77" authorityName="Ryvkin" authorityYear="1987" box="[748,858,555,581]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="illusor">
<emphasis id="3AE2EAF9C029FFC8FD4DAF63FD2E9A77" box="[748,763,555,581]" italics="true" pageId="5" pageNumber="6">S</emphasis>
. 
<emphasis id="3AE2EAF9C029FFC8FCACAF63FC8F9A77" box="[781,858,555,581]" italics="true" pageId="5" pageNumber="6">illusor</emphasis>
</taxonomicName>
, 
<taxonomicName id="CF964D68C029FFC8FCCAAF63FBD59A77" authorityName="J. Sahlberg" authorityYear="1880" box="[875,1024,555,581]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="latipennis">
<emphasis id="3AE2EAF9C029FFC8FCCAAF63FCAF9A77" box="[875,890,555,581]" italics="true" pageId="5" pageNumber="6">S</emphasis>
. 
<emphasis id="3AE2EAF9C029FFC8FC2DAF63FBD59A77" box="[908,1024,555,581]" italics="true" pageId="5" pageNumber="6">latipennis</emphasis>
</taxonomicName>
, 
<taxonomicName id="CF964D68C029FFC8FBB0AF63FB629A77" authorityName="Puthz" authorityYear="2006" box="[1041,1207,555,581]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="immigratus">
<emphasis id="3AE2EAF9C029FFC8FBB0AF63FBF59A77" box="[1041,1056,555,581]" italics="true" pageId="5" pageNumber="6">S</emphasis>
. 
<emphasis id="3AE2EAF9C029FFC8FB90AF64FB629A77" box="[1073,1207,556,581]" italics="true" pageId="5" pageNumber="6">immigratus</emphasis>
</taxonomicName>
, 
<taxonomicName id="CF964D68C029FFC8FB68AF63FABA9A77" authorityName="Puthz" authorityYear="1973" box="[1225,1391,555,581]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="geminorum">
<emphasis id="3AE2EAF9C029FFC8FB68AF63FB0D9A77" box="[1225,1240,555,581]" italics="true" pageId="5" pageNumber="6">S</emphasis>
. 
<emphasis id="3AE2EAF9C029FFC8FB48AF64FABA9A77" box="[1257,1391,556,581]" italics="true" pageId="5" pageNumber="6">geminorum</emphasis>
</taxonomicName>
, 
<taxonomicName id="CF964D68C029FFC8FF1CAF06FE849A5A" authorityName="Casey" authorityYear="1884" box="[189,337,590,617]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="sphaerops">
<emphasis id="3AE2EAF9C029FFC8FF1CAF06FF199A5A" box="[189,204,590,616]" italics="true" pageId="5" pageNumber="6">S</emphasis>
. 
<emphasis id="3AE2EAF9C029FFC8FF78AF06FE849A5A" box="[217,337,590,616]" italics="true" pageId="5" pageNumber="6">sphaerops</emphasis>
</taxonomicName>
, 
<taxonomicName id="CF964D68C029FFC8FEFCAF06FE069A5A" authorityName="Casey" authorityYear="1884" box="[349,467,590,617]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="dolosus">
<emphasis id="3AE2EAF9C029FFC8FEFCAF06FEB99A5A" box="[349,364,590,616]" italics="true" pageId="5" pageNumber="6">S</emphasis>
. 
<emphasis id="3AE2EAF9C029FFC8FED8AF06FE069A5A" box="[377,467,590,616]" italics="true" pageId="5" pageNumber="6">dolosus</emphasis>
</taxonomicName>
, 
<taxonomicName id="CF964D68C029FFC8FE41AF06FDAC9A5A" authorityName="Casey" authorityYear="1884" box="[480,633,590,617]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="milleporus">
<emphasis id="3AE2EAF9C029FFC8FE41AF06FE3A9A5A" box="[480,495,590,616]" italics="true" pageId="5" pageNumber="6">S</emphasis>
. 
<emphasis id="3AE2EAF9C029FFC8FE5DAF06FDAC9A5A" box="[508,633,590,616]" italics="true" pageId="5" pageNumber="6">milleporus</emphasis>
</taxonomicName>
). The latter tendency seems to arise and develop independently, at least within the complexes 
<emphasis id="3AE2EAF9C029FFC8FE55AF39FB8F9AB9" box="[500,1114,625,651]" italics="true" pageId="5" pageNumber="6">labilis-confusus-delitor-illusor-latipennis-sphaerops</emphasis>
and 
<emphasis id="3AE2EAF9C029FFC8FB2FAF39FABB9AB9" box="[1166,1390,625,652]" italics="true" pageId="5" pageNumber="6">
<taxonomicName id="CF964D68C029FFC8FB2FAF39FB3D9AB9" authorityName="Casey" authorityYear="1884" box="[1166,1256,625,651]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="dolosus">dolosus</taxonomicName>
- 
<taxonomicName id="CF964D68C029FFC8FB50AF39FABB9AB9" authorityName="Casey" authorityYear="1884" box="[1265,1390,625,651]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="milleporus">milleporus</taxonomicName>
</emphasis>
, in spite of the fact that the latter is closely related in the shape of the male genitalia to the ‘ 
<taxonomicName id="CF964D68C029FFC8FB79AFDCFAA39A9C" authorityName="Gyllenhal" authorityYear="1827" box="[1240,1398,660,686]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" sensu="stricto" species="canaliculatus">
<emphasis id="3AE2EAF9C029FFC8FB79AFDCFAA39A9C" box="[1240,1398,660,686]" italics="true" pageId="5" pageNumber="6">canaliculatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="21D15782C029FFC8FF1CAFF0FF209AE0" box="[189,245,696,722]" pageId="5" pageNumber="6" sensu="stricto">s.str.</taxonomicNameLabel>
’ complex characterised by a flat front.
</paragraph>
<paragraph id="082936EBC029FFC8FF1CAFB2FAA29BF6" blockId="5.[189,1399,761,964]" pageId="5" pageNumber="6">
B. For three species studied by him (
<taxonomicName id="CF964D68C029FFC8FDDEAFB2FCE89B21" authorityName="Gyllenhal" authorityYear="1827" box="[639,829,761,788]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="canaliculatus">
<emphasis id="3AE2EAF9C029FFC8FDDEAFB2FD5B9B26" box="[639,654,762,788]" italics="true" pageId="5" pageNumber="6">S</emphasis>
. 
<emphasis id="3AE2EAF9C029FFC8FD3EAFB1FCE89B21" box="[671,829,761,787]" italics="true" pageId="5" pageNumber="6">canaliculatus</emphasis>
</taxonomicName>
, 
<taxonomicName id="CF964D68C029FFC8FCEFAFB2FC619B21" authorityName="Stephens" authorityYear="1833" box="[846,948,762,788]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="nitens">
<emphasis id="3AE2EAF9C029FFC8FCEFAFB2FC889B26" box="[846,861,762,788]" italics="true" pageId="5" pageNumber="6">S</emphasis>
. 
<emphasis id="3AE2EAF9C029FFC8FCCFAFB2FC619B21" box="[878,948,762,787]" italics="true" pageId="5" pageNumber="6">nitens</emphasis>
</taxonomicName>
, 
<taxonomicName id="CF964D68C029FFC8FC64AFB2FBFB9B21" authorityName="Erichson" authorityYear="1840" box="[965,1070,761,788]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="labilis">
<emphasis id="3AE2EAF9C029FFC8FC64AFB2FC019B26" box="[965,980,762,788]" italics="true" pageId="5" pageNumber="6">S</emphasis>
. 
<emphasis id="3AE2EAF9C029FFC8FC44AFB1FBFB9B21" box="[997,1070,761,787]" italics="true" pageId="5" pageNumber="6">labilis</emphasis>
</taxonomicName>
), 
<bibRefCitation id="6C074B1AC029FFC8FBEBAFB2FB029B26" author="Betz" box="[1098,1239,761,788]" firstAuthor="Betz" pageId="5" pageNumber="6" pagination="15 - 34" refId="ref40460" refString="Betz O. 1996. Function and evolution of the adhesion-capture apparatus of Stenus species (Coleoptera, Staphylinidae). Zoomorphology 116: 15 - 34. http: // dx. doi. org / 10.1007 / BF 02526926" type="journal article" year="1996">Betz (1996)</bibRefCitation>
has shown a considerable reduction of the adhesion capture apparatus. It is expressed both in the degeneration of sticky cushions of paraglossae (decreasing of their surface and number of adhesive setae etc.) and in a shortening of the labium. No data on labium length have been provided by the named author but, as it follows from the text, 
<taxonomicName id="CF964D68C029FFC8FDA7AECEFD779B92" authorityName="Casey" authorityYear="1884" box="[518,674,902,928]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="milleporus">
<emphasis id="3AE2EAF9C029FFC8FDA7AECEFDC09B92" box="[518,533,902,928]" italics="true" pageId="5" pageNumber="6">S</emphasis>
. 
<emphasis id="3AE2EAF9C029FFC8FD84AECEFD779B92" box="[549,674,902,928]" italics="true" pageId="5" pageNumber="6">milleporus</emphasis>
</taxonomicName>
(as 
<taxonomicName id="CF964D68C029FFC8FD79AECEFC099B92" authority="Casey, 1884" authorityName="Casey" authorityYear="1884" box="[728,988,902,928]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="sectilifer">
<emphasis id="3AE2EAF9C029FFC8FD79AECEFCEA9B92" box="[728,831,902,928]" italics="true" pageId="5" pageNumber="6">sectilifer</emphasis>
<bibRefCitation id="6C074B1AC029FFC8FCE9AECEFC099B92" author="Casey" box="[840,988,902,928]" firstAuthor="Casey" pageId="5" pageNumber="6" refId="ref40664" refString="Casey Th. L. 1884. Revision of the Stenini of America north of Mexico. Insects of the family Staphylinidae, order Coleoptera. Collins Printing House, Philadelphia. 206 pp. http: // dx. doi. org / 10.5962 / bhl. title. 9208" type="book" year="1884">Casey, 1884</bibRefCitation>
</taxonomicName>
), 
<taxonomicName id="CF964D68C029FFC8FC52AECEFB899B92" authorityName="Puthz" authorityYear="1972" box="[1011,1116,902,928]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="caseyi">
<emphasis id="3AE2EAF9C029FFC8FC52AECEFBD79B92" box="[1011,1026,902,928]" italics="true" pageId="5" pageNumber="6">S</emphasis>
. 
<emphasis id="3AE2EAF9C029FFC8FBB2AECFFB899B92" box="[1043,1116,903,928]" italics="true" pageId="5" pageNumber="6">caseyi</emphasis>
</taxonomicName>
, 
<taxonomicName id="CF964D68C029FFC8FBCCAECEFB3A9B92" authorityName="Casey" authorityYear="1884" box="[1133,1263,902,928]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="vinnulus">
<emphasis id="3AE2EAF9C029FFC8FBCCAECEFBA99B92" box="[1133,1148,902,928]" italics="true" pageId="5" pageNumber="6">S</emphasis>
. 
<emphasis id="3AE2EAF9C029FFC8FB2DAECEFB3A9B92" box="[1164,1263,902,928]" italics="true" pageId="5" pageNumber="6">vinnulus</emphasis>
</taxonomicName>
, as well as 
<taxonomicName id="CF964D68C029FFC8FF1CAEE2FE8C9BF1" authorityName="Marsham" authorityYear="1802" box="[189,345,937,964]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="melanopus">
<emphasis id="3AE2EAF9C029FFC8FF1CAEE2FF199BF6" box="[189,204,938,964]" italics="true" pageId="5" pageNumber="6">S</emphasis>
. 
<emphasis id="3AE2EAF9C029FFC8FF7BAEE1FE8C9BF1" box="[218,345,937,963]" italics="true" pageId="5" pageNumber="6">melanopus</emphasis>
</taxonomicName>
, which is regarded as a member of the same group, have also been studied in this respect.
</paragraph>
<paragraph id="082936EBC029FFC8FF1CAEA3FD009DC3" blockId="5.[189,1399,1002,1521]" pageId="5" pageNumber="6">
I have measured the relative length of the prementum (eulabium after 
<bibRefCitation id="6C074B1AC029FFC8FC5EAEA2FB6E9C37" author="Weinreich" box="[1023,1211,1002,1029]" firstAuthor="Weinreich" pageId="5" pageNumber="6" pagination="162 - 210" refId="ref46824" refString="Weinreich E. 1968. Uber den Klebfangapparat der Imagines von Stenus Latr. (Coleopt., Staphylinidae) mit einem Beitrag zur Kenntnis der Jugendstadien dieser Gattung. Zeitschrift fur Morphologie der Tiere 62: 162 - 210." type="journal article" year="1968">Weinreich 1968</bibRefCitation>
) for six species of the 
<taxonomicName id="CF964D68C029FFC8FEAFA946FE799C1A" authorityName="Gyllenhal" authorityYear="1827" box="[270,428,1038,1064]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="canaliculatus">
<emphasis id="3AE2EAF9C029FFC8FEAFA946FE799C1A" box="[270,428,1038,1064]" italics="true" pageId="5" pageNumber="6">canaliculatus</emphasis>
</taxonomicName>
group: 
<taxonomicName id="CF964D68C029FFC8FDACA946FD1E9C1A" authorityName="Gyllenhal" authorityYear="1827" box="[525,715,1038,1064]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="canaliculatus">
<emphasis id="3AE2EAF9C029FFC8FDACA946FDC99C1A" box="[525,540,1038,1064]" italics="true" pageId="5" pageNumber="6">S</emphasis>
. 
<emphasis id="3AE2EAF9C029FFC8FD8CA946FD1E9C1A" box="[557,715,1038,1064]" italics="true" pageId="5" pageNumber="6">canaliculatus</emphasis>
</taxonomicName>
(I: 17, 0.53; II: 16, 0.53), 
<taxonomicName id="CF964D68C029FFC8FBB5A946FBAE9C1A" authorityName="Stephens" authorityYear="1833" box="[1044,1147,1038,1064]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="nitens">
<emphasis id="3AE2EAF9C029FFC8FBB5A946FBF69C1A" box="[1044,1059,1038,1064]" italics="true" pageId="5" pageNumber="6">S</emphasis>
. 
<emphasis id="3AE2EAF9C029FFC8FB94A947FBAE9C1A" box="[1077,1147,1039,1064]" italics="true" pageId="5" pageNumber="6">nitens</emphasis>
</taxonomicName>
(33, 0.87), 
<taxonomicName id="CF964D68C029FFC8FAACA946FAA39C1A" authorityName="Erichson" authorityYear="1840" box="[1293,1398,1038,1064]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="labilis">
<emphasis id="3AE2EAF9C029FFC8FAACA946FAC99C1A" box="[1293,1308,1038,1064]" italics="true" pageId="5" pageNumber="6">S</emphasis>
. 
<emphasis id="3AE2EAF9C029FFC8FA8CA946FAA39C1A" box="[1325,1398,1038,1064]" italics="true" pageId="5" pageNumber="6">labilis</emphasis>
</taxonomicName>
(37, 1.12), 
<taxonomicName id="CF964D68C029FFC8FEE1A979FE6A9C79" authorityName="Puthz" authorityYear="1972" box="[320,447,1073,1099]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="illotulus">
<emphasis id="3AE2EAF9C029FFC8FEE1A979FE9A9C79" box="[320,335,1073,1099]" italics="true" pageId="5" pageNumber="6">S</emphasis>
. 
<emphasis id="3AE2EAF9C029FFC8FEFEA979FE6A9C79" box="[351,447,1073,1099]" italics="true" pageId="5" pageNumber="6">illotulus</emphasis>
</taxonomicName>
(26, 0.84), 
<taxonomicName id="CF964D68C029FFC8FDEAA979FD639C79" authorityName="Ryvkin" authorityYear="1987" box="[587,694,1073,1099]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="raddei">
<emphasis id="3AE2EAF9C029FFC8FDEAA979FD8F9C79" box="[587,602,1073,1099]" italics="true" pageId="5" pageNumber="6">S</emphasis>
. 
<emphasis id="3AE2EAF9C029FFC8FDC8A979FD639C79" box="[617,694,1073,1099]" italics="true" pageId="5" pageNumber="6">raddei</emphasis>
</taxonomicName>
(35, 1.09), 
<taxonomicName id="CF964D68C029FFC8FCE3A979FC019C79" authorityName="J. Sahlberg" authorityYear="1880" box="[834,980,1073,1099]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="latipennis">
<emphasis id="3AE2EAF9C029FFC8FCE3A979FC849C79" box="[834,849,1073,1099]" italics="true" pageId="5" pageNumber="6">S</emphasis>
. 
<emphasis id="3AE2EAF9C029FFC8FCC1A979FC019C79" box="[864,980,1073,1099]" italics="true" pageId="5" pageNumber="6">latipennis</emphasis>
</taxonomicName>
(31, 0.79); and for three species of other groups of the subgenus 
<taxonomicName id="CF964D68C029FFC8FDB8A91DFD3E9C5C" authority="Rey, 1884" authorityName="Rey" authorityYear="1884" box="[537,747,1108,1135]" class="Insecta" family="Staphylinidae" genus="Nestus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="3AE2EAF9C029FFC8FDB8A91DFDB29C5C" box="[537,615,1109,1134]" italics="true" pageId="5" pageNumber="6">Nestus</emphasis>
<bibRefCitation id="6C074B1AC029FFC8FDD1A91DFD3E9C5C" author="Rey" box="[624,747,1108,1135]" firstAuthor="Rey" pageId="5" pageNumber="6" pagination="153 - 415" refId="ref44858" refString="Rey Cl. 1884. Tribu des Brevipennes. Deuxieme groupe: Micropeplides. Troisieme groupe: Stenides. Annales de la Societe Linneenne de Lyon (n. ser.) 30: 153 - 415." type="journal article" year="1884">Rey, 1884</bibRefCitation>
</taxonomicName>
: 
<taxonomicName id="CF964D68C029FFC8FD56A91DFBCA9C5C" authority="Erichson, 1840" authorityName="Erichson" authorityYear="1840" box="[759,1055,1108,1135]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="ruralis">
<emphasis id="3AE2EAF9C029FFC8FD56A91DFCD39C5D" box="[759,774,1109,1135]" italics="true" pageId="5" pageNumber="6">S</emphasis>
. 
<emphasis id="3AE2EAF9C029FFC8FCB4A91CFCB09C5C" box="[789,869,1108,1134]" italics="true" pageId="5" pageNumber="6">ruralis</emphasis>
<bibRefCitation id="6C074B1AC029FFC8FCCDA91CFBCA9C5C" author="Erichson" box="[876,1055,1108,1134]" firstAuthor="Erichson" pageId="5" pageNumber="6" refId="ref40960" refString="Erichson W. F. 1840. Genera et species staphylinorum insectorum coleopterorum familiae: 401 - 954. F. H. Morin, Berlin. http: // dx. doi. org / 10.5962 / bhl. title. 59644" type="book" year="1840">Erichson, 1840</bibRefCitation>
</taxonomicName>
(47, 1.52) from the 
<emphasis id="3AE2EAF9C029FFC8FAAFA91CFAA39C5C" box="[1294,1398,1108,1134]" italics="true" pageId="5" pageNumber="6">palposus</emphasis>
group, 
<taxonomicName id="CF964D68C029FFC8FEB5A930FDB59CA3" authority="Gravenhorst, 1806" authorityName="Gravenhorst" authorityYear="1806" box="[276,608,1143,1170]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="argus">
<emphasis id="3AE2EAF9C029FFC8FEB5A930FEF69CA0" box="[276,291,1144,1170]" italics="true" pageId="5" pageNumber="6">S</emphasis>
. 
<emphasis id="3AE2EAF9C029FFC8FE94A930FEA29CA3" box="[309,375,1144,1169]" italics="true" pageId="5" pageNumber="6">argus</emphasis>
Gravenhorst, 1806
</taxonomicName>
(47, 1.74) from the 
<emphasis id="3AE2EAF9C029FFC8FCFEA93FFC6B9CA3" box="[863,958,1143,1169]" italics="true" pageId="5" pageNumber="6">fuscipes</emphasis>
group, and 
<taxonomicName id="CF964D68C029FFC8FBF4A930FF2D9C87" authority="Marsham, 1802" authorityName="Marsham" authorityYear="1802" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="melanopus">
<emphasis id="3AE2EAF9C029FFC8FBF4A930FBB19CA0" box="[1109,1124,1144,1170]" italics="true" pageId="5" pageNumber="6">S</emphasis>
. 
<emphasis id="3AE2EAF9C029FFC8FBD7A93FFB209CA3" box="[1142,1269,1143,1169]" italics="true" pageId="5" pageNumber="6">melanopus</emphasis>
<bibRefCitation id="6C074B1AC029FFC8FAA1A93FFF2D9C87" author="Marsham" firstAuthor="Marsham" pageId="5" pageNumber="6" refId="ref42860" refString="Marsham T. 1802. Entomologia britannica, sistens insecta, Britanniae indigena, secundum methodum Linnaeanum disposita. Tomus I. Coleoptera: I-XXXI + 1 - 549. Wilks &amp; Taylor, Londini." type="book" year="1802">Marsham, 1802</bibRefCitation>
</taxonomicName>
(41, 1.46) presumably representing a monotypic group. In each case above, the first number in brackets means length of prementum whereas the second one is a rounded ratio between the former and the length of the head (
<specimenCount id="1E90FD62C029FFC8FDA3A9A9FD679CC9" box="[514,690,1249,1275]" pageId="5" pageNumber="6" type="generic">two specimens</specimenCount>
have been measured for 
<taxonomicName id="CF964D68C029FFC8FC7CA9A9FB4C9CC9" authorityName="Gyllenhal" authorityYear="1827" box="[989,1177,1249,1275]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="canaliculatus">
<emphasis id="3AE2EAF9C029FFC8FC7CA9A9FC399CC9" box="[989,1004,1249,1275]" italics="true" pageId="5" pageNumber="6">S</emphasis>
. 
<emphasis id="3AE2EAF9C029FFC8FC5AA9A9FB4C9CC9" box="[1019,1177,1249,1275]" italics="true" pageId="5" pageNumber="6">canaliculatus</emphasis>
</taxonomicName>
). One can see that the data split the examined species into three clusters: a) 
<taxonomicName id="CF964D68C029FFC8FCC7A84DFBF79D2C" authorityName="Gyllenhal" authorityYear="1827" box="[870,1058,1284,1311]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="canaliculatus">
<emphasis id="3AE2EAF9C029FFC8FCC7A84DFCA09D2D" box="[870,885,1285,1311]" italics="true" pageId="5" pageNumber="6">S</emphasis>
. 
<emphasis id="3AE2EAF9C029FFC8FC25A84CFBF79D2C" box="[900,1058,1284,1310]" italics="true" pageId="5" pageNumber="6">canaliculatus</emphasis>
</taxonomicName>
; b) the remaining species of the 
<taxonomicName id="CF964D68C029FFC8FF4AA86FFE5C9D73" authorityName="Gyllenhal" authorityYear="1827" box="[235,393,1319,1345]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="canaliculatus">
<emphasis id="3AE2EAF9C029FFC8FF4AA86FFE5C9D73" box="[235,393,1319,1345]" italics="true" pageId="5" pageNumber="6">canaliculatus</emphasis>
</taxonomicName>
group; c) the species of other groups; both total and relative length of prementum increases from category a) to c). It can be supposed that the reduction of the eulabium is a characteristic feature of all the members of the 
<taxonomicName id="CF964D68C029FFC8FDEEA826FD389DBA" authorityName="Gyllenhal" authorityYear="1827" box="[591,749,1390,1416]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="canaliculatus">
<emphasis id="3AE2EAF9C029FFC8FDEEA826FD389DBA" box="[591,749,1390,1416]" italics="true" pageId="5" pageNumber="6">canaliculatus</emphasis>
</taxonomicName>
group, but it must be verified by a specific study. The shortening of the adhesion capture apparatus seems to correlate with some other modifications of the mouth parts, namely the more proximal position of the internal tooth of mandibles as well as its lesser deflection off the main plane of the mandible.
</paragraph>
<paragraph id="082936EBC029FFC8FF1CAB51FE129F1B" blockId="5.[189,1398,1561,1833]" pageId="5" pageNumber="6">
The small median notch of the labrum is a specific character of 
<taxonomicName id="CF964D68C029FFC8FC1AAB51FBAD9E01" authorityName="Gyllenhal" authorityYear="1827" box="[955,1144,1561,1587]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="canaliculatus">
<emphasis id="3AE2EAF9C029FFC8FC1AAB51FC1F9E01" box="[955,970,1561,1587]" italics="true" pageId="5" pageNumber="6">S</emphasis>
. 
<emphasis id="3AE2EAF9C029FFC8FC7BAB51FBAD9E01" box="[986,1144,1561,1587]" italics="true" pageId="5" pageNumber="6">canaliculatus</emphasis>
</taxonomicName>
and the most closely related Palaearctic species (
<taxonomicName id="CF964D68C029FFC8FDA3AB74FDA19E64" authorityName="Ryvkin" authorityYear="2012" box="[514,628,1596,1622]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="canalis">
<emphasis id="3AE2EAF9C029FFC8FDA3AB74FDC49E64" box="[514,529,1596,1622]" italics="true" pageId="5" pageNumber="6">S</emphasis>
. 
<emphasis id="3AE2EAF9C029FFC8FDBEAB74FDA19E64" box="[543,628,1596,1622]" italics="true" pageId="5" pageNumber="6">canalis</emphasis>
</taxonomicName>
, 
<taxonomicName id="CF964D68C029FFC8FD23AB74FD2A9E64" authorityName="Ryvkin" authorityYear="2012" box="[642,767,1596,1622]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="canosus">
<emphasis id="3AE2EAF9C029FFC8FD23AB74FD449E64" box="[642,657,1596,1622]" italics="true" pageId="5" pageNumber="6">S</emphasis>
. 
<emphasis id="3AE2EAF9C029FFC8FD01AB75FD2A9E64" box="[672,767,1597,1622]" italics="true" pageId="5" pageNumber="6">canosus</emphasis>
</taxonomicName>
, 
<taxonomicName id="CF964D68C029FFC8FCACAB74FCAD9E64" authorityName="Ryvkin" authorityYear="2012" box="[781,888,1596,1622]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="alopex">
<emphasis id="3AE2EAF9C029FFC8FCACAB74FCC99E64" box="[781,796,1596,1622]" italics="true" pageId="5" pageNumber="6">S</emphasis>
. 
<emphasis id="3AE2EAF9C029FFC8FC8BAB74FCAD9E64" box="[810,888,1596,1622]" italics="true" pageId="5" pageNumber="6">alopex</emphasis>
</taxonomicName>
). Though the functional explanation of this feature is obscure as yet, one can suppose it to correlate with the extreme reduction of the adhesion capture apparatus in this species complex. The Nearctic members of the same complex (
<taxonomicName id="CF964D68C029FFC8FB7BABCBFAA39EAE" authorityName="Casey" authorityYear="1884" box="[1242,1398,1666,1693]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="milleporus">
<emphasis id="3AE2EAF9C029FFC8FB7BABCBFB3C9EAF" box="[1242,1257,1667,1693]" italics="true" pageId="5" pageNumber="6">S</emphasis>
. 
<emphasis id="3AE2EAF9C029FFC8FB58ABCAFAA39EAE" box="[1273,1398,1666,1692]" italics="true" pageId="5" pageNumber="6">milleporus</emphasis>
</taxonomicName>
and 
<taxonomicName id="CF964D68C029FFC8FF53ABEEFEBB9EF2" authorityName="Casey" authorityYear="1884" box="[242,366,1702,1728]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="dolosus">
<emphasis id="3AE2EAF9C029FFC8FF53ABEEFED49EF2" box="[242,257,1702,1728]" italics="true" pageId="5" pageNumber="6">S</emphasis>
. 
<emphasis id="3AE2EAF9C029FFC8FEB5ABEEFEBB9EF2" box="[276,366,1702,1728]" italics="true" pageId="5" pageNumber="6">dolosus</emphasis>
</taxonomicName>
) have only a broad and shallow emargination on the anterior margin of the labrum; the margin is slightly sinuate at the middle in 
<taxonomicName id="CF964D68C029FFC8FD46AB81FC999ED1" authorityName="Stephens" authorityYear="1833" box="[743,844,1737,1763]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="nitens">
<emphasis id="3AE2EAF9C029FFC8FD46AB81FD239ED1" box="[743,758,1737,1763]" italics="true" pageId="5" pageNumber="6">S</emphasis>
. 
<emphasis id="3AE2EAF9C029FFC8FCA7AB82FC999ED1" box="[774,844,1738,1763]" italics="true" pageId="5" pageNumber="6">nitens</emphasis>
</taxonomicName>
, 
<taxonomicName id="CF964D68C029FFC8FCFCAB81FC099ED1" authorityName="Puthz" authorityYear="1972" box="[861,988,1737,1763]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="illotulus">
<emphasis id="3AE2EAF9C029FFC8FCFCAB81FCB99ED1" box="[861,876,1737,1763]" italics="true" pageId="5" pageNumber="6">S</emphasis>
. 
<emphasis id="3AE2EAF9C029FFC8FCDDAB81FC099ED1" box="[892,988,1737,1763]" italics="true" pageId="5" pageNumber="6">illotulus</emphasis>
</taxonomicName>
, 
<taxonomicName id="CF964D68C029FFC8FC4CAB81FB8C9ED1" authorityName="Ryvkin" authorityYear="1987" box="[1005,1113,1737,1763]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="raddei">
<emphasis id="3AE2EAF9C029FFC8FC4CAB81FC299ED1" box="[1005,1020,1737,1763]" italics="true" pageId="5" pageNumber="6">S</emphasis>
. 
<emphasis id="3AE2EAF9C029FFC8FBADAB81FB8C9ED1" box="[1036,1113,1737,1763]" italics="true" pageId="5" pageNumber="6">raddei</emphasis>
</taxonomicName>
, 
<taxonomicName id="CF964D68C029FFC8FBC8AB81FB049ED1" authorityName="Puthz" authorityYear="1972" box="[1129,1233,1737,1763]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="caseyi">
<emphasis id="3AE2EAF9C029FFC8FBC8AB81FBAD9ED1" box="[1129,1144,1737,1763]" italics="true" pageId="5" pageNumber="6">S</emphasis>
. 
<emphasis id="3AE2EAF9C029FFC8FB29AB82FB049ED1" box="[1160,1233,1738,1763]" italics="true" pageId="5" pageNumber="6">caseyi</emphasis>
</taxonomicName>
, 
<taxonomicName id="CF964D68C029FFC8FB43AB81FA9A9ED1" authorityName="Puthz" authorityYear="1972" box="[1250,1359,1737,1763]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="brivioi">
<emphasis id="3AE2EAF9C029FFC8FB43AB81FB249ED1" box="[1250,1265,1737,1763]" italics="true" pageId="5" pageNumber="6">S</emphasis>
. 
<emphasis id="3AE2EAF9C029FFC8FAA0AB81FA9A9ED1" box="[1281,1359,1737,1763]" italics="true" pageId="5" pageNumber="6">brivioi</emphasis>
</taxonomicName>
, 
<taxonomicName id="CF964D68C029FFC8FAC1AB81FEF59F34" authorityName="Casey" authorityYear="1884" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="vinnulus">
<emphasis id="3AE2EAF9C029FFC8FAC1AB81FABA9ED1" box="[1376,1391,1737,1763]" italics="true" pageId="5" pageNumber="6">S</emphasis>
. 
<emphasis id="3AE2EAF9C029FFC8FF1CABA4FEF59F34" box="[189,288,1772,1798]" italics="true" pageId="5" pageNumber="6">vinnulus</emphasis>
</taxonomicName>
; lastly, the species of the 
<emphasis id="3AE2EAF9C029FFC8FDC6ABA4FB189F34" box="[615,1229,1772,1798]" italics="true" pageId="5" pageNumber="6">labilis-confusus-delitor-illusor-latipennis-sphaerops</emphasis>
line have the margin nearly straight.
</paragraph>
<footnote id="6B8D2AE5C029FFC8FF1CAA19FEA9902E" pageId="5" pageNumber="6">
<paragraph id="082936EBC029FFC8FF1CAA19FEA9902E" blockId="5.[189,1399,1872,2040]" lastBlockId="5.[189,380,2048,2076]" pageId="5" pageNumber="6">
<bibRefCitation id="6C074B1AC029FFC8FF1CAA19FE9C9F59" author="Betz" box="[189,329,1872,1899]" firstAuthor="Betz" pageId="5" pageNumber="6" pagination="15 - 34" refId="ref40460" refString="Betz O. 1996. Function and evolution of the adhesion-capture apparatus of Stenus species (Coleoptera, Staphylinidae). Zoomorphology 116: 15 - 34. http: // dx. doi. org / 10.1007 / BF 02526926" type="journal article" year="1996">Betz (1996)</bibRefCitation>
supposed the reduction of the adhesion capture apparatus in the 
<taxonomicName id="CF964D68C029FFC8FBE8AA19FB329F59" authorityName="Gyllenhal" authorityYear="1827" box="[1097,1255,1873,1899]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="canaliculatus">
<emphasis id="3AE2EAF9C029FFC8FBE8AA19FB329F59" box="[1097,1255,1873,1899]" italics="true" pageId="5" pageNumber="6">canaliculatus</emphasis>
</taxonomicName>
group to be a result of miniaturisation, but this explanation does not stand up to any criticism. Indeed, the smallest specimens of the species within the group may be considered among the medium-sized representatives of the genus, while even much smaller members of both the 
<emphasis id="3AE2EAF9C029FFC8FC32AAF2FC3B9FE6" box="[915,1006,1978,2004]" italics="true" pageId="5" pageNumber="6">pusillus</emphasis>
and the 
<emphasis id="3AE2EAF9C029FFC8FBF6AAF2FB649FE6" box="[1111,1201,1978,2004]" italics="true" pageId="5" pageNumber="6">pumilio</emphasis>
groups have the adhesion capture apparatus well developed (
<emphasis id="3AE2EAF9C029FFC8FD71AA96FD0A9FCA" box="[720,735,2014,2040]" italics="true" pageId="5" pageNumber="6">S</emphasis>
. (
<emphasis id="3AE2EAF9C029FFC8FD58AA96FCD89FC5" box="[761,781,2014,2039]" italics="true" pageId="5" pageNumber="6">N</emphasis>
.) 
<emphasis id="3AE2EAF9C029FFC8FC86AA95FC579FC5" box="[807,898,2013,2039]" italics="true" pageId="5" pageNumber="6">pusillus</emphasis>
<bibRefCitation id="6C074B1AC029FFC8FC2DAA95FB949FCA" author="Stephens" box="[908,1089,2013,2040]" firstAuthor="Stephens" pageId="5" pageNumber="6" refId="ref46202" refString="Stephens J. F. 1833. Illustrations of British Entomology; or a synopsis of indigenous insects: containing their generic and specific distinctions; with an account of their metamorphoses, times of appearance, localities, food, and economy, as far as practicable. Mandibulata. Vol. V: 241 - 304. Baldwin and Cradock, London. http: // dx. doi. org / 10.5962 / bhl. title. 8133" type="book" year="1833">Stephens, 1833</bibRefCitation>
has been studied by 
<bibRefCitation id="6C074B1AC029FFC8FAE0AA96FEDE9029" author="Betz O." pageId="5" pageNumber="6" pagination="15 - 34" refId="ref40460" refString="Betz O. 1996. Function and evolution of the adhesion-capture apparatus of Stenus species (Coleoptera, Staphylinidae). Zoomorphology 116: 15 - 34. http: // dx. doi. org / 10.1007 / BF 02526926" type="journal article" year="1996">Betz (1996)</bibRefCitation>
himself).
</paragraph>
</footnote>
<paragraph id="082936EBC02AFFCBFF1CAC44FDA49A50" blockId="6.[189,1399,267,611]" pageId="6" pageNumber="7">
I believe that the modification of the mouth parts should be connected more likely to a particular suite of potential prey species. Unfortunately, Betz used only springtails and (to a lesser extent) winged aphids in his interesting prey-capture experiments, but those may not exhaust the list of possible prey objects for 
<taxonomicName id="CF964D68C02AFFCBFEEAAC3DFE6199BD" authorityName="W.S.MacLeay" authorityYear="1825" box="[331,436,373,399]" class="Insecta" family="Staphylinidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="subFamily" subFamily="Steninae">Steninae</taxonomicName>
. I did not experiment with species of the 
<taxonomicName id="CF964D68C02AFFCBFC64AC3DFBB699BD" authorityName="Gyllenhal" authorityYear="1827" box="[965,1123,373,399]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="canaliculatus">
<emphasis id="3AE2EAF9C02AFFCBFC64AC3DFBB699BD" box="[965,1123,373,399]" italics="true" pageId="6" pageNumber="7">canaliculatus</emphasis>
</taxonomicName>
group; however other 
<taxonomicName id="CF964D68C02AFFCBFF1CACD1FEDF9981" box="[189,266,409,435]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="3AE2EAF9C02AFFCBFF1CACD1FEDF9981" box="[189,266,409,435]" italics="true" pageId="6" pageNumber="7">Stenus</emphasis>
</taxonomicName>
species were capturing nymphs of 
<taxonomicName id="CF964D68C02AFFCBFD13ACD0FCDA9980" box="[690,783,408,434]" class="Insecta" family="Miridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="family">Miridae</taxonomicName>
(
<taxonomicName id="CF964D68C02AFFCBFC81ACD0FCAC9980" authorityName="Linnaeus" authorityYear="1758" box="[800,889,408,434]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="6" pageNumber="7" phylum="Arthropoda" rank="class">Insecta</taxonomicName>
: 
<taxonomicName id="CF964D68C02AFFCBFC27ACD0FBD29980" box="[902,1031,408,434]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="order">Hemiptera</taxonomicName>
: 
<taxonomicName id="CF964D68C02AFFCBFBB2ACD1FB759981" box="[1043,1184,409,435]" class="Magnoliopsida" higherTaxonomySource="GBIF" kingdom="Plantae" order="Heteroptera" pageId="6" pageNumber="7" phylum="Tracheophyta" rank="order">Heteroptera</taxonomicName>
) and 
<taxonomicName id="CF964D68C02AFFCBFB45ACD0FAA29980" authorityName="Latreille" authorityYear="1825" box="[1252,1399,408,434]" class="Insecta" family="Cicadellidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="family">Cicadellidae</taxonomicName>
(
<taxonomicName id="CF964D68C02AFFCBFF64ACF3FECB99E7" authorityName="Linnaeus" authorityYear="1758" box="[197,286,443,469]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="6" pageNumber="7" phylum="Arthropoda" rank="class">Insecta</taxonomicName>
: 
<taxonomicName id="CF964D68C02AFFCBFE89ACF3FE7C99E7" box="[296,425,443,469]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="order">Hemiptera</taxonomicName>
: 
<taxonomicName id="CF964D68C02AFFCBFE11ACF3FD5299E4" box="[432,647,443,470]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Auchenorrhyncha" pageId="6" pageNumber="7" phylum="Arthropoda" rank="order">Auchenorrhyncha</taxonomicName>
), juvenile spiders of the family 
<taxonomicName id="CF964D68C02AFFCBFC50ACF3FBAA99E7" authorityName="Blackwall" authorityYear="1859" box="[1009,1151,443,469]" class="Arachnida" family="Linyphiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="family">Linyphiidae</taxonomicName>
(
<taxonomicName id="CF964D68C02AFFCBFB2CACF3FADB99E7" box="[1165,1294,443,469]" class="Arachnida" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="6" pageNumber="7" phylum="Arthropoda" rank="class">Arachnida</taxonomicName>
: Aranei), small mites (
<taxonomicName id="CF964D68C02AFFCBFEF4AC96FE0399CA" box="[341,470,478,504]" class="Arachnida" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="6" pageNumber="7" phylum="Arthropoda" rank="class">Arachnida</taxonomicName>
: Acari), small pyralid caterpillars (
<taxonomicName id="CF964D68C02AFFCBFCD0AC97FC1F99CB" authorityName="Linnaeus" authorityYear="1758" box="[881,970,479,505]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="6" pageNumber="7" phylum="Arthropoda" rank="class">Insecta</taxonomicName>
: 
<taxonomicName id="CF964D68C02AFFCBFC76AC96FBBF99CA" authorityName="Linnaeus" authorityYear="1758" box="[983,1130,478,504]" class="Magnoliopsida" higherTaxonomySource="GBIF" kingdom="Plantae" order="Lepidoptera" pageId="6" pageNumber="7" phylum="Tracheophyta" rank="order">Lepidoptera</taxonomicName>
: 
<taxonomicName id="CF964D68C02AFFCBFBD4AC96FB3C99CB" box="[1141,1257,478,505]" class="Insecta" family="Pyralidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="family">Pyralidae</taxonomicName>
), small flies (Diptera), soil 
<taxonomicName id="CF964D68C02AFFCBFEC5AF4AFD609A2F" authority="(Oligochaeta)" baseAuthorityName="Oligochaeta" box="[356,693,514,541]" class="Magnoliopsida" family="Enchytraeidae" higherTaxonomySource="GBIF" kingdom="Plantae" order="Asterales" pageId="6" pageNumber="7" phylum="Tracheophyta" rank="family">Enchytraeidae (Oligochaeta)</taxonomicName>
, in addition to the aforementioned objects, in my unpublished experiments of 
<date id="7C28102BC02AFFCBFED6AF6DFE2A9A72" box="[375,511,549,576]" pageId="6" pageNumber="7" value="1985" valueMax="1986">1985–1986</date>
. I consider it necessary to undertake a more detailed study of prey for the species of the group being discussed.
</paragraph>
<paragraph id="082936EBC02AFFCBFF1CAFC2FE7D9B66" blockId="6.[189,1399,649,852]" pageId="6" pageNumber="7">
C. The tendency of legs to become longer has most likely taken place independently in two species complexes: 
<emphasis id="3AE2EAF9C02AFFCBFEEDAFE5FC8A9AF5" box="[332,863,685,711]" italics="true" pageId="6" pageNumber="7">delitor-illusor-latipennis-confusus-sphaerops</emphasis>
and 
<emphasis id="3AE2EAF9C02AFFCBFC3CAFE5FB769AF5" box="[925,1187,685,711]" italics="true" pageId="6" pageNumber="7">nitens-illotulus-raddei</emphasis>
(only the species with the 1 
<superScript id="FFE39BA3C02AFFCBFE9CAF87FE9D9AEF" attach="left" box="[317,328,719,733]" fontSize="6" pageId="6" pageNumber="7">st</superScript>
segment of the metatarsus longer than the 5 
<superScript id="FFE39BA3C02AFFCBFCDBAF86FC529AEE" attach="left" box="[890,903,718,732]" fontSize="6" pageId="6" pageNumber="7">th</superScript>
are listed). 
<taxonomicName id="CF964D68C02AFFCBFB82AF98FB369AD8" authorityName="Gyllenhal" authorityYear="1827" box="[1059,1251,720,746]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="canaliculatus">
<emphasis id="3AE2EAF9C02AFFCBFB82AF98FBE79AD8" box="[1059,1074,720,746]" italics="true" pageId="6" pageNumber="7">S</emphasis>
. 
<emphasis id="3AE2EAF9C02AFFCBFBE4AF98FB369AD8" box="[1093,1251,720,746]" italics="true" pageId="6" pageNumber="7">canaliculatus</emphasis>
</taxonomicName>
, 
<taxonomicName id="CF964D68C02AFFCBFB59AF98FABA9AD8" authorityName="Ryvkin" authorityYear="2012" box="[1272,1391,720,746]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="canalis">
<emphasis id="3AE2EAF9C02AFFCBFB59AF98FAD29AD8" box="[1272,1287,720,746]" italics="true" pageId="6" pageNumber="7">S</emphasis>
. 
<emphasis id="3AE2EAF9C02AFFCBFABBAF98FABA9AD8" box="[1306,1391,720,746]" italics="true" pageId="6" pageNumber="7">canalis</emphasis>
</taxonomicName>
, 
<taxonomicName id="CF964D68C02AFFCBFF1CAFBCFEEC9B3F" authorityName="Ryvkin" authorityYear="2012" box="[189,313,756,782]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="canosus">
<emphasis id="3AE2EAF9C02AFFCBFF1CAFBCFF199B3C" box="[189,204,756,782]" italics="true" pageId="6" pageNumber="7">S</emphasis>
. 
<emphasis id="3AE2EAF9C02AFFCBFF7BAFBCFEEC9B3F" box="[218,313,756,781]" italics="true" pageId="6" pageNumber="7">canosus</emphasis>
</taxonomicName>
, 
<taxonomicName id="CF964D68C02AFFCBFEE7AFBCFE649B3F" authorityName="Ryvkin" authorityYear="2012" box="[326,433,755,782]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="alopex">
<emphasis id="3AE2EAF9C02AFFCBFEE7AFBCFE809B3C" box="[326,341,756,782]" italics="true" pageId="6" pageNumber="7">S</emphasis>
. 
<emphasis id="3AE2EAF9C02AFFCBFEC2AFBBFE649B3F" box="[355,433,755,781]" italics="true" pageId="6" pageNumber="7">alopex</emphasis>
</taxonomicName>
, 
<taxonomicName id="CF964D68C02AFFCBFE1EAFBCFD8C9B3F" authorityName="Casey" authorityYear="1884" box="[447,601,755,782]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="milleporus">
<emphasis id="3AE2EAF9C02AFFCBFE1EAFBCFE1B9B3C" box="[447,462,756,782]" italics="true" pageId="6" pageNumber="7">S</emphasis>
. 
<emphasis id="3AE2EAF9C02AFFCBFE7DAFBBFD8C9B3F" box="[476,601,755,781]" italics="true" pageId="6" pageNumber="7">milleporus</emphasis>
</taxonomicName>
, 
<taxonomicName id="CF964D68C02AFFCBFDC9AFBCFD189B3F" authorityName="Puthz" authorityYear="1972" box="[616,717,756,782]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="caseyi">
<emphasis id="3AE2EAF9C02AFFCBFDC9AFBCFDA29B3C" box="[616,631,756,782]" italics="true" pageId="6" pageNumber="7">S</emphasis>
. 
<emphasis id="3AE2EAF9C02AFFCBFD25AFBCFD189B3F" box="[644,717,756,781]" italics="true" pageId="6" pageNumber="7">caseyi</emphasis>
</taxonomicName>
have metatarsi with the 1 
<superScript id="FFE39BA3C02AFFCBFC58AFBAFBD19B32" attach="left" box="[1017,1028,754,768]" fontSize="6" pageId="6" pageNumber="7">st</superScript>
segment about as long as 5 
<superScript id="FFE39BA3C02AFFCBFAE2AFB9FA859ACD" attach="none" box="[1347,1360,753,767]" fontSize="6" pageId="6" pageNumber="7">th</superScript>
; in 
<taxonomicName id="CF964D68C02AFFCBFF1CAE5FFEE09B02" authorityName="Casey" authorityYear="1884" box="[189,309,790,817]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="dolosus">
<emphasis id="3AE2EAF9C02AFFCBFF1CAE5FFF199B03" box="[189,204,791,817]" italics="true" pageId="6" pageNumber="7">S</emphasis>
. 
<emphasis id="3AE2EAF9C02AFFCBFF7AAE5EFEE09B02" box="[219,309,790,816]" italics="true" pageId="6" pageNumber="7">dolosus</emphasis>
</taxonomicName>
, 
<taxonomicName id="CF964D68C02AFFCBFEE5AE5FFE659B02" authorityName="Puthz" authorityYear="1972" box="[324,432,790,817]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="brivioi">
<emphasis id="3AE2EAF9C02AFFCBFEE5AE5FFE869B03" box="[324,339,791,817]" italics="true" pageId="6" pageNumber="7">S</emphasis>
. 
<emphasis id="3AE2EAF9C02AFFCBFEC3AE5EFE659B02" box="[354,432,790,816]" italics="true" pageId="6" pageNumber="7">brivioi</emphasis>
</taxonomicName>
, 
<taxonomicName id="CF964D68C02AFFCBFE61AE5FFD949B02" authorityName="Casey" authorityYear="1884" box="[448,577,790,817]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="vinnulus">
<emphasis id="3AE2EAF9C02AFFCBFE61AE5FFE1A9B03" box="[448,463,791,817]" italics="true" pageId="6" pageNumber="7">S</emphasis>
. 
<emphasis id="3AE2EAF9C02AFFCBFE7FAE5EFD949B02" box="[478,577,790,816]" italics="true" pageId="6" pageNumber="7">vinnulus</emphasis>
</taxonomicName>
, and 
<taxonomicName id="CF964D68C02AFFCBFD25AE5FFD3E9B02" authorityName="Erichson" authorityYear="1840" box="[644,747,790,817]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="labilis">
<emphasis id="3AE2EAF9C02AFFCBFD25AE5FFD469B03" box="[644,659,791,817]" italics="true" pageId="6" pageNumber="7">S</emphasis>
. 
<emphasis id="3AE2EAF9C02AFFCBFD03AE5EFD3E9B02" box="[674,747,790,816]" italics="true" pageId="6" pageNumber="7">labilis</emphasis>
</taxonomicName>
, metatarsi remain short (with the 1 
<superScript id="FFE39BA3C02AFFCBFB2DAE5DFB429B11" attach="left" box="[1164,1175,789,803]" fontSize="6" pageId="6" pageNumber="7">st</superScript>
segment evidently shorter than the 5 
<superScript id="FFE39BA3C02AFFCBFE28AE70FE439B74" attach="none" box="[393,406,824,838]" fontSize="6" pageId="6" pageNumber="7">th</superScript>
).
</paragraph>
<paragraph id="082936EBC02AFFCBFF1CAE34FD779CC7" blockId="6.[189,1399,891,1270]" pageId="6" pageNumber="7">
D. The plesiomorphic aedeagus with a lanceolate median lobe and unmodified cylindrical parameres (as in 
<taxonomicName id="CF964D68C02AFFCBFF7DAED7FE919B8A" authorityName="Erichson" authorityYear="1840" box="[220,324,926,953]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="labilis">
<emphasis id="3AE2EAF9C02AFFCBFF7DAED7FF3E9B8B" box="[220,235,927,953]" italics="true" pageId="6" pageNumber="7">S</emphasis>
. 
<emphasis id="3AE2EAF9C02AFFCBFF5AAED6FE919B8A" box="[251,324,926,952]" italics="true" pageId="6" pageNumber="7">labilis</emphasis>
</taxonomicName>
) can be transformed by different ways. The distal sclerotized part of the median lobe may be shortened, either angularly (as in 
<taxonomicName id="CF964D68C02AFFCBFDC2AE8AFD229BE9" authorityName="Casey" authorityYear="1884" box="[611,759,961,988]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="sphaerops">
<emphasis id="3AE2EAF9C02AFFCBFDC2AE8AFDA79BEE" box="[611,626,962,988]" italics="true" pageId="6" pageNumber="7">S</emphasis>
. 
<emphasis id="3AE2EAF9C02AFFCBFDDEAE89FD229BE9" box="[639,759,961,987]" italics="true" pageId="6" pageNumber="7">sphaerops</emphasis>
</taxonomicName>
) or rounded-angularly (in the 
<emphasis id="3AE2EAF9C02AFFCBFBF7AE89FAA29BE9" box="[1110,1399,961,987]" italics="true" pageId="6" pageNumber="7">delitor-illusor-latipennis</emphasis>
line); in only one case with a small apical denticle (in 
<taxonomicName id="CF964D68C02AFFCBFC9FAEADFC6A9BCD" authorityName="Casey" authorityYear="1884" box="[830,959,997,1023]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="vinnulus">
<emphasis id="3AE2EAF9C02AFFCBFC9FAEADFC989BCD" box="[830,845,997,1023]" italics="true" pageId="6" pageNumber="7">S</emphasis>
. 
<emphasis id="3AE2EAF9C02AFFCBFCFDAEADFC6A9BCD" box="[860,959,997,1023]" italics="true" pageId="6" pageNumber="7">vinnulus</emphasis>
</taxonomicName>
); this modification usually correlates to a lengthening of the whole aedeagus (see 
<figureCitation id="90AD2A6EC02AFFCBFD6CA940FCF49C10" box="[717,801,1032,1058]" captionStart="Fig" captionStartId="38.[189,232,1956,1982]" captionTargetBox="[177,1408,266,1913]" captionTargetId="figure@38.[161,1420,248,1934]" captionTargetPageId="38" captionText="Fig. 6. Aedeagus, ventral view. A. Stenus (Nestus) delitor sp. nov. (holotype). B. S. (N.) sphaerops Casey, 1884 (U.S.A.: New York: Albany Co.). – Scales = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3857787" httpUri="https://zenodo.org/record/3857787/files/figure.png" pageId="6" pageNumber="7">Figs. 6</figureCitation>
, 
<figureCitation id="90AD2A6EC02AFFCBFC8FA940FCE99C10" box="[814,828,1032,1058]" captionStart="Fig" captionStartId="41.[189,232,1803,1829]" captionTargetBox="[239,1339,1025,1762]" captionTargetId="figure@41.[206,1398,1001,1771]" captionTargetPageId="41" captionText="Fig. 7. ♂ copulative structures. A–I, K: Aedeagus, ventral view. J: Abdominal sternite 9. A. Stenus (Nestus) delitor sp. nov. (holotype). B. S. (N.) illusor Ryvkin, 1987 (Russia: Amur Area: Angelokit River). C–H: S. (N.) latipennis J. Sahlberg, 1880. C. (Russia: SW Taimyr: Ladannakh Lake). D. (Russia: Taimyr: Kotuy River). E. (Russia: Putorana: Ayan Lake). F–G. (Russia: Evenkia: Kruten’kiy Stream). H. (Russia: Magadan Area: ‘Aborigen’). I–J. S. (N.) sphaerops Casey, 1884 (U.S.A.: New York: Albany Co.). K. S. (N.) melanopus Marsham, 1802 (Ukraine: Odessa Area: Krinichnoye). – Scales = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3857789" httpUri="https://zenodo.org/record/3857789/files/figure.png" pageId="6" pageNumber="7">7</figureCitation>
A–I) or only the median lobe (as in 
<taxonomicName id="CF964D68C02AFFCBFB4BA940FABA9C10" authorityName="J. Sahlberg" authorityYear="1876" box="[1258,1391,1032,1058]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="confusus">
<emphasis id="3AE2EAF9C02AFFCBFB4BA940FB2C9C10" box="[1258,1273,1032,1058]" italics="true" pageId="6" pageNumber="7">S</emphasis>
. 
<emphasis id="3AE2EAF9C02AFFCBFAA9A940FABA9C10" box="[1288,1391,1032,1058]" italics="true" pageId="6" pageNumber="7">confusus</emphasis>
</taxonomicName>
, see 
<figureCitation id="90AD2A6EC02AFFCBFF4BA963FE919C77" box="[234,324,1067,1093]" captionStart="Fig" captionStartId="13.[189,232,1852,1878]" captionTargetBox="[244,1294,261,1809]" captionTargetId="figure@13.[219,1366,255,1824]" captionTargetPageId="13" captionText="Fig. 2. – A–B: Stenus (Nestus) raddei Ryvkin, 1987: ♂ (Russia: Magadan Area: Sibit-Tyhellakh). A. Aedeagus, ventral view. B. Apical part of aedeagus, ventral view. – C–D: S. (N.) illotulus Puthz, 1972: ♂ (Russia: Amur Area: Zeyskiy Nature Reserve). C. Aedeagus, ventral view. D. Apical part of aedeagus, ventral view. Right paramere removed. – E. S. (N.) confusus J. Sahlberg, 1876: ♂ (Russia: Kalinin Area: Putilovo). Aedeagus, ventral view. – Scales = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3857779" httpUri="https://zenodo.org/record/3857779/files/figure.png" pageId="6" pageNumber="7">Fig. 2E</figureCitation>
), although this correlation has not been observed in 
<taxonomicName id="CF964D68C02AFFCBFC0CA963FBCD9C77" authorityName="Ryvkin" authorityYear="1987" box="[941,1048,1067,1093]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="raddei">
<emphasis id="3AE2EAF9C02AFFCBFC0CA963FBCD9C77" box="[941,1048,1067,1093]" italics="true" pageId="6" pageNumber="7">S. raddei</emphasis>
</taxonomicName>
, belonging to another lineage (
<figureCitation id="90AD2A6EC02AFFCBFF67A906FE969C5B" box="[198,323,1102,1129]" captionStart="Fig" captionStartId="13.[189,232,1852,1878]" captionTargetBox="[244,1294,261,1809]" captionTargetId="figure@13.[219,1366,255,1824]" captionTargetPageId="13" captionText="Fig. 2. – A–B: Stenus (Nestus) raddei Ryvkin, 1987: ♂ (Russia: Magadan Area: Sibit-Tyhellakh). A. Aedeagus, ventral view. B. Apical part of aedeagus, ventral view. – C–D: S. (N.) illotulus Puthz, 1972: ♂ (Russia: Amur Area: Zeyskiy Nature Reserve). C. Aedeagus, ventral view. D. Apical part of aedeagus, ventral view. Right paramere removed. – E. S. (N.) confusus J. Sahlberg, 1876: ♂ (Russia: Kalinin Area: Putilovo). Aedeagus, ventral view. – Scales = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3857779" httpUri="https://zenodo.org/record/3857779/files/figure.png" pageId="6" pageNumber="7">Fig. 2A, B</figureCitation>
). The tendency of the median lobe to become narrower (as in 
<taxonomicName id="CF964D68C02AFFCBFBADA907FBA89C5A" authorityName="Puthz" authorityYear="1987" box="[1036,1149,1102,1129]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="shogun">
<emphasis id="3AE2EAF9C02AFFCBFBADA907FBA89C5A" box="[1036,1149,1102,1129]" italics="true" pageId="6" pageNumber="7">S. shogun</emphasis>
</taxonomicName>
, 
<taxonomicName id="CF964D68C02AFFCBFB2BA907FB209C5A" authorityName="Puthz" authorityYear="1972" box="[1162,1269,1102,1129]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="brivioi">
<emphasis id="3AE2EAF9C02AFFCBFB2BA907FB4C9C5B" box="[1162,1177,1103,1129]" italics="true" pageId="6" pageNumber="7">S</emphasis>
. 
<emphasis id="3AE2EAF9C02AFFCBFB06A906FB209C5A" box="[1191,1269,1102,1128]" italics="true" pageId="6" pageNumber="7">brivioi</emphasis>
</taxonomicName>
, 
<taxonomicName id="CF964D68C02AFFCBFAA3A907FAB99C5A" authorityName="Ryvkin" authorityYear="2012" box="[1282,1388,1102,1129]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="alopex">
<emphasis id="3AE2EAF9C02AFFCBFAA3A907FAB99C5A" box="[1282,1388,1102,1129]" italics="true" pageId="6" pageNumber="7">S. alopex</emphasis>
</taxonomicName>
) is another way of transformation, being peculiar to the lineages of 
<taxonomicName id="CF964D68C02AFFCBFC6CA93AFBC69CB9" authorityName="Stephens" authorityYear="1833" box="[973,1043,1138,1163]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="nitens">
<emphasis id="3AE2EAF9C02AFFCBFC6CA93AFBC69CB9" box="[973,1043,1138,1163]" italics="true" pageId="6" pageNumber="7">nitens</emphasis>
</taxonomicName>
and ‘ 
<taxonomicName id="CF964D68C02AFFCBFBF6A939FB209CB9" authorityName="Gyllenhal" authorityYear="1827" box="[1111,1269,1137,1163]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" sensu="stricto" species="canaliculatus">
<emphasis id="3AE2EAF9C02AFFCBFBF6A939FB209CB9" box="[1111,1269,1137,1163]" italics="true" pageId="6" pageNumber="7">canaliculatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="21D15782C02AFFCBFB5CA93AFAEC9CBE" box="[1277,1337,1138,1164]" pageId="6" pageNumber="7" sensu="stricto">s.str.</taxonomicNameLabel>
’; one can suppose that the aedeagus of Mexican 
<taxonomicName id="CF964D68C02AFFCBFD67A9DDFC949C9D" authorityName="Sharp" authorityYear="1887" box="[710,833,1173,1199]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="idoneus">
<emphasis id="3AE2EAF9C02AFFCBFD67A9DDFC949C9D" box="[710,833,1173,1199]" italics="true" pageId="6" pageNumber="7">S. idoneus</emphasis>
</taxonomicName>
has evolved in this way. All the modifications mentioned may be followed by forming a more or less developed medial impression or/and keel at the distal sclerotized part of the median lobe.
</paragraph>
<paragraph id="082936EBC02AFFCBFF1CA854FC829D4F" blockId="6.[189,1399,1308,1405]" pageId="6" pageNumber="7">The main trend in the evolution of the parameres is a differentiation of their setiferous surface. It is expressed more frequently in a flattening of the apicointernal side of each paramere or, on the contrary, in forming a one-sided swell which setae are attached to.</paragraph>
<paragraph id="082936EBC02AFFCBFF1CA8ECFE9A9E5D" blockId="6.[189,1398,1444,1647]" pageId="6" pageNumber="7">
Based on all the characters and tendencies briefly discussed above, we can subdivide the 
<taxonomicName id="CF964D68C02AFFCBFB79A8ECFAA39D8C" authorityName="Gyllenhal" authorityYear="1827" box="[1240,1398,1444,1470]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="canaliculatus">
<emphasis id="3AE2EAF9C02AFFCBFB79A8ECFAA39D8C" box="[1240,1398,1444,1470]" italics="true" pageId="6" pageNumber="7">canaliculatus</emphasis>
</taxonomicName>
group into four subgroups: the monotypic 
<taxonomicName id="CF964D68C02AFFCBFD59A88FFC949DD3" authorityName="Erichson" authorityYear="1840" box="[760,833,1479,1505]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="labilis">
<emphasis id="3AE2EAF9C02AFFCBFD59A88FFC949DD3" box="[760,833,1479,1505]" italics="true" pageId="6" pageNumber="7">labilis</emphasis>
</taxonomicName>
subgroup (
<taxonomicName id="CF964D68C02AFFCBFC41A880FB869DD3" authorityName="Erichson" authorityYear="1840" box="[992,1107,1479,1506]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="labilis">
<emphasis id="3AE2EAF9C02AFFCBFC41A880FC3A9DD0" box="[992,1007,1480,1506]" italics="true" pageId="6" pageNumber="7">S</emphasis>
. 
<emphasis id="3AE2EAF9C02AFFCBFBABA88FFB869DD3" box="[1034,1107,1479,1505]" italics="true" pageId="6" pageNumber="7">labilis</emphasis>
</taxonomicName>
), the 
<taxonomicName id="CF964D68C02AFFCBFB0EA880FB209DD3" authorityName="Stephens" authorityYear="1833" box="[1199,1269,1480,1505]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="nitens">
<emphasis id="3AE2EAF9C02AFFCBFB0EA880FB209DD3" box="[1199,1269,1480,1505]" italics="true" pageId="6" pageNumber="7">nitens</emphasis>
</taxonomicName>
subgroup (
<taxonomicName id="CF964D68C02AFFCBFF66A8A3FEBD9E37" authorityName="Puthz" authorityYear="2006" box="[199,360,1515,1541]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="immigratus">
<emphasis id="3AE2EAF9C02AFFCBFF66A8A3FF039E37" box="[199,214,1515,1541]" italics="true" pageId="6" pageNumber="7">S</emphasis>
. 
<emphasis id="3AE2EAF9C02AFFCBFF43A8A4FEBD9E37" box="[226,360,1516,1541]" italics="true" pageId="6" pageNumber="7">immigratus</emphasis>
</taxonomicName>
, 
<taxonomicName id="CF964D68C02AFFCBFED5A8A3FE0D9E37" authorityName="Stephens" authorityYear="1833" box="[372,472,1515,1541]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="nitens">
<emphasis id="3AE2EAF9C02AFFCBFED5A8A3FE569E37" box="[372,387,1515,1541]" italics="true" pageId="6" pageNumber="7">S</emphasis>
. 
<emphasis id="3AE2EAF9C02AFFCBFE2EA8A4FE0D9E37" box="[399,472,1516,1541]" italics="true" pageId="6" pageNumber="7">nitens</emphasis>
</taxonomicName>
<emphasis id="3AE2EAF9C02AFFCBFE79A8A4FE089E37" box="[472,477,1516,1541]" italics="true" pageId="6" pageNumber="7">,</emphasis>
<taxonomicName id="CF964D68C02AFFCBFE43A8A3FD9F9E37" authorityName="Ryvkin" authorityYear="1987" box="[482,586,1515,1541]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="raddei">
<emphasis id="3AE2EAF9C02AFFCBFE43A8A3FE249E37" box="[482,497,1515,1541]" italics="true" pageId="6" pageNumber="7">S</emphasis>
. 
<emphasis id="3AE2EAF9C02AFFCBFE5CA8A3FD9F9E37" box="[509,586,1515,1541]" italics="true" pageId="6" pageNumber="7">raddei</emphasis>
</taxonomicName>
, 
<taxonomicName id="CF964D68C02AFFCBFDF7A8A3FD019E37" authorityName="Puthz" authorityYear="1972" box="[598,724,1515,1541]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="illotulus">
<emphasis id="3AE2EAF9C02AFFCBFDF7A8A3FDB09E37" box="[598,613,1515,1541]" italics="true" pageId="6" pageNumber="7">S</emphasis>
. 
<emphasis id="3AE2EAF9C02AFFCBFDD0A8A3FD019E37" box="[625,724,1515,1541]" italics="true" pageId="6" pageNumber="7">illotulus</emphasis>
</taxonomicName>
<emphasis id="3AE2EAF9C02AFFCBFD75A8A3FC9B9E37" box="[724,846,1515,1541]" italics="true" pageId="6" pageNumber="7">
, 
<taxonomicName id="CF964D68C02AFFCBFD7FA8A3FC9B9E37" authorityName="Puthz" authorityYear="1987" box="[734,846,1515,1541]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="shogun">S. shogun</taxonomicName>
</emphasis>
, 
<taxonomicName id="CF964D68C02AFFCBFCFAA8A3FC6A9E37" authorityName="Puthz" authorityYear="1972" box="[859,959,1515,1541]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="caseyi">
<emphasis id="3AE2EAF9C02AFFCBFCFAA8A3FCBF9E37" box="[859,874,1515,1541]" italics="true" pageId="6" pageNumber="7">S</emphasis>
. 
<emphasis id="3AE2EAF9C02AFFCBFCD7A8A4FC6A9E37" box="[886,959,1516,1541]" italics="true" pageId="6" pageNumber="7">caseyi</emphasis>
</taxonomicName>
, 
<taxonomicName id="CF964D68C02AFFCBFC6DA8A3FBB89E37" authorityName="Puthz" authorityYear="1973" box="[972,1133,1515,1541]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="geminorum">
<emphasis id="3AE2EAF9C02AFFCBFC6DA8A3FC0E9E37" box="[972,987,1515,1541]" italics="true" pageId="6" pageNumber="7">S</emphasis>
. 
<emphasis id="3AE2EAF9C02AFFCBFC46A8A4FBB89E37" box="[999,1133,1516,1541]" italics="true" pageId="6" pageNumber="7">geminorum</emphasis>
</taxonomicName>
, 
<taxonomicName id="CF964D68C02AFFCBFBD8A8A3FB379E37" authorityName="Puthz" authorityYear="1972" box="[1145,1250,1515,1541]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="brivioi">
<emphasis id="3AE2EAF9C02AFFCBFBD8A8A3FB5D9E37" box="[1145,1160,1515,1541]" italics="true" pageId="6" pageNumber="7">S</emphasis>
. 
<emphasis id="3AE2EAF9C02AFFCBFB35A8A3FB379E37" box="[1172,1250,1515,1541]" italics="true" pageId="6" pageNumber="7">brivioi</emphasis>
</taxonomicName>
, 
<taxonomicName id="CF964D68C02AFFCBFB4EA8A3FAB09E37" authorityName="Sharp" authorityYear="1887" box="[1263,1381,1515,1541]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="idoneus">
<emphasis id="3AE2EAF9C02AFFCBFB4EA8A3FAB09E37" box="[1263,1381,1515,1541]" italics="true" pageId="6" pageNumber="7">S. idoneus</emphasis>
</taxonomicName>
), the 
<taxonomicName id="CF964D68C02AFFCBFF4DAB46FE5F9E1A" authorityName="Gyllenhal" authorityYear="1827" box="[236,394,1550,1576]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" sensu="stricto" species="canaliculatus">
<emphasis id="3AE2EAF9C02AFFCBFF4DAB46FE5F9E1A" box="[236,394,1550,1576]" italics="true" pageId="6" pageNumber="7">canaliculatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="21D15782C02AFFCBFE34AB46FE1E9E1A" box="[405,459,1550,1576]" pageId="6" pageNumber="7" sensu="stricto">s.str.</taxonomicNameLabel>
subgroup (
<taxonomicName id="CF964D68C02AFFCBFDF9AB46FD069E1A" authorityName="Casey" authorityYear="1884" box="[600,723,1550,1576]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="dolosus">
<emphasis id="3AE2EAF9C02AFFCBFDF9AB46FD069E1A" box="[600,723,1550,1576]" italics="true" pageId="6" pageNumber="7">S. dolosus</emphasis>
</taxonomicName>
, 
<taxonomicName id="CF964D68C02AFFCBFD44AB46FC569E1A" authorityName="Casey" authorityYear="1884" box="[741,899,1550,1576]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="milleporus">
<emphasis id="3AE2EAF9C02AFFCBFD44AB46FD219E1A" box="[741,756,1550,1576]" italics="true" pageId="6" pageNumber="7">S</emphasis>
. 
<emphasis id="3AE2EAF9C02AFFCBFCA7AB46FC569E1A" box="[774,899,1550,1576]" italics="true" pageId="6" pageNumber="7">milleporus</emphasis>
</taxonomicName>
, 
<taxonomicName id="CF964D68C02AFFCBFC37AB46FB809E1A" authorityName="Gyllenhal" authorityYear="1827" box="[918,1109,1550,1576]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="canaliculatus">
<emphasis id="3AE2EAF9C02AFFCBFC37AB46FC709E1A" box="[918,933,1550,1576]" italics="true" pageId="6" pageNumber="7">S</emphasis>
. 
<emphasis id="3AE2EAF9C02AFFCBFC16AB46FB809E1A" box="[951,1109,1550,1576]" italics="true" pageId="6" pageNumber="7">canaliculatus</emphasis>
</taxonomicName>
, 
<taxonomicName id="CF964D68C02AFFCBFBC6AB46FB089E1A" authorityName="Ryvkin" authorityYear="2012" box="[1127,1245,1550,1576]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="canalis">
<emphasis id="3AE2EAF9C02AFFCBFBC6AB46FBA39E1A" box="[1127,1142,1550,1576]" italics="true" pageId="6" pageNumber="7">S</emphasis>
. 
<emphasis id="3AE2EAF9C02AFFCBFB29AB46FB089E1A" box="[1160,1245,1550,1576]" italics="true" pageId="6" pageNumber="7">canalis</emphasis>
</taxonomicName>
, 
<taxonomicName id="CF964D68C02AFFCBFB51AB46FAA59E1A" authorityName="Ryvkin" authorityYear="2012" box="[1264,1392,1550,1576]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="canosus">
<emphasis id="3AE2EAF9C02AFFCBFB51AB46FB2A9E1A" box="[1264,1279,1550,1576]" italics="true" pageId="6" pageNumber="7">S</emphasis>
. 
<emphasis id="3AE2EAF9C02AFFCBFAB0AB47FAA59E1A" box="[1297,1392,1551,1576]" italics="true" pageId="6" pageNumber="7">canosus</emphasis>
</taxonomicName>
, 
<taxonomicName id="CF964D68C02AFFCBFF1CAB7AFEFE9E79" authorityName="Ryvkin" authorityYear="2012" box="[189,299,1585,1612]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="alopex">
<emphasis id="3AE2EAF9C02AFFCBFF1CAB7AFF199E7E" box="[189,204,1586,1612]" italics="true" pageId="6" pageNumber="7">S</emphasis>
. 
<emphasis id="3AE2EAF9C02AFFCBFF7CAB79FEFE9E79" box="[221,299,1585,1611]" italics="true" pageId="6" pageNumber="7">alopex</emphasis>
</taxonomicName>
), and the 
<taxonomicName id="CF964D68C02AFFCBFE08AB79FDC59E79" authorityName="J. Sahlberg" authorityYear="1876" box="[425,528,1585,1611]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="confusus">
<emphasis id="3AE2EAF9C02AFFCBFE08AB79FDC59E79" box="[425,528,1585,1611]" italics="true" pageId="6" pageNumber="7">confusus</emphasis>
</taxonomicName>
subgroup (
<taxonomicName id="CF964D68C02AFFCBFD3AAB7AFCF69E79" authorityName="J. Sahlberg" authorityYear="1876" box="[667,803,1585,1612]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="confusus">
<emphasis id="3AE2EAF9C02AFFCBFD3AAB7AFD7F9E7E" box="[667,682,1586,1612]" italics="true" pageId="6" pageNumber="7">S</emphasis>
. 
<emphasis id="3AE2EAF9C02AFFCBFD1DAB79FCF69E79" box="[700,803,1585,1611]" italics="true" pageId="6" pageNumber="7">confusus</emphasis>
</taxonomicName>
, 
<taxonomicName id="CF964D68C02AFFCBFC95AB7AFC779E79" authorityName="Ryvkin" authorityYear="2012" box="[820,930,1585,1612]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="delitor">
<emphasis id="3AE2EAF9C02AFFCBFC95AB7AFC779E79" box="[820,930,1585,1612]" italics="true" pageId="6" pageNumber="7">S. delitor</emphasis>
</taxonomicName>
, 
<taxonomicName id="CF964D68C02AFFCBFC15AB7AFBF49E79" authorityName="Ryvkin" authorityYear="1987" box="[948,1057,1585,1612]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="illusor">
<emphasis id="3AE2EAF9C02AFFCBFC15AB7AFBF49E79" box="[948,1057,1585,1612]" italics="true" pageId="6" pageNumber="7">S. illusor</emphasis>
</taxonomicName>
, 
<taxonomicName id="CF964D68C02AFFCBFB93AB7AFB139E79" authorityName="J. Sahlberg" authorityYear="1880" box="[1074,1222,1585,1612]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="latipennis">
<emphasis id="3AE2EAF9C02AFFCBFB93AB7AFB139E79" box="[1074,1222,1585,1612]" italics="true" pageId="6" pageNumber="7">S. latipennis</emphasis>
</taxonomicName>
, 
<taxonomicName id="CF964D68C02AFFCBFB76AB7AFAA59E79" authorityName="Casey" authorityYear="1884" box="[1239,1392,1585,1612]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="sphaerops">
<emphasis id="3AE2EAF9C02AFFCBFB76AB7AFB339E7E" box="[1239,1254,1586,1612]" italics="true" pageId="6" pageNumber="7">S</emphasis>
. 
<emphasis id="3AE2EAF9C02AFFCBFB59AB79FAA59E79" box="[1272,1392,1585,1611]" italics="true" pageId="6" pageNumber="7">sphaerops</emphasis>
</taxonomicName>
, 
<taxonomicName id="CF964D68C02AFFCBFF1CAB1DFEE89E5C" authorityName="Casey" authorityYear="1884" box="[189,317,1620,1647]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="vinnulus">
<emphasis id="3AE2EAF9C02AFFCBFF1CAB1DFF199E5D" box="[189,204,1621,1647]" italics="true" pageId="6" pageNumber="7">S</emphasis>
. 
<emphasis id="3AE2EAF9C02AFFCBFF7BAB1CFEE89E5C" box="[218,317,1620,1646]" italics="true" pageId="6" pageNumber="7">vinnulus</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="408C6560C02AFFCAFF1CABDEFB97997B" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7" type="biology_ecology">
<paragraph id="082936EBC02AFFCBFF1CABDEFEEA9E82" blockId="6.[189,319,1686,1712]" box="[189,319,1686,1712]" pageId="6" pageNumber="7">
<heading id="53618187C02AFFCBFF1CABDEFEEA9E82" bold="true" box="[189,319,1686,1712]" fontSize="11" level="3" pageId="6" pageNumber="7" reason="3">
<emphasis id="3AE2EAF9C02AFFCBFF1CABDEFEEA9E82" bold="true" box="[189,319,1686,1712]" pageId="6" pageNumber="7">Bionomics</emphasis>
</heading>
</paragraph>
<paragraph id="082936EBC02AFFCAFF1CAB8CFB97997B" blockId="6.[189,1399,1732,2005]" lastBlockId="7.[189,1399,267,329]" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">
The majority of the species for which enough data have been presented can be grouped into swamp inhabitants and alluvial ones. The former category, comprising the species dwelling in moss and litter of eutrophic and mesotrophic wet habitats, definitely includes 
<taxonomicName id="CF964D68C02AFFCBFC34AA43FC229F17" authorityName="Stephens" authorityYear="1833" box="[917,1015,1803,1829]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="nitens">
<emphasis id="3AE2EAF9C02AFFCBFC34AA43FC229F17" box="[917,1015,1803,1829]" italics="true" pageId="6" pageNumber="7">S. nitens</emphasis>
</taxonomicName>
, 
<taxonomicName id="CF964D68C02AFFCBFBA7AA43FBA59F17" authorityName="Ryvkin" authorityYear="1987" box="[1030,1136,1803,1829]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="raddei">
<emphasis id="3AE2EAF9C02AFFCBFBA7AA43FBA59F17" box="[1030,1136,1803,1829]" italics="true" pageId="6" pageNumber="7">S. raddei</emphasis>
</taxonomicName>
, 
<taxonomicName id="CF964D68C02AFFCBFBDFAA43FB2F9F17" authorityName="Puthz" authorityYear="1972" box="[1150,1274,1803,1829]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="illotulus">
<emphasis id="3AE2EAF9C02AFFCBFBDFAA43FB2F9F17" box="[1150,1274,1803,1829]" italics="true" pageId="6" pageNumber="7">S. illotulus</emphasis>
</taxonomicName>
, 
<taxonomicName id="CF964D68C02AFFCBFAA8AA43FABA9F17" authorityName="Puthz" authorityYear="1972" box="[1289,1391,1803,1829]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="caseyi">
<emphasis id="3AE2EAF9C02AFFCBFAA8AA43FABA9F17" box="[1289,1391,1803,1829]" italics="true" pageId="6" pageNumber="7">S. caseyi</emphasis>
</taxonomicName>
, 
<taxonomicName id="CF964D68C02AFFCBFF1CAA66FEF99F7A" authorityName="Puthz" authorityYear="1972" box="[189,300,1838,1864]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="brivioi">
<emphasis id="3AE2EAF9C02AFFCBFF1CAA66FEF99F7A" box="[189,300,1838,1864]" italics="true" pageId="6" pageNumber="7">S. brivioi</emphasis>
</taxonomicName>
and 
<taxonomicName id="CF964D68C02AFFCBFECCAA66FDD39F7A" authorityName="Casey" authorityYear="1884" box="[365,518,1838,1864]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="sphaerops">
<emphasis id="3AE2EAF9C02AFFCBFECCAA66FDD39F7A" box="[365,518,1838,1864]" italics="true" pageId="6" pageNumber="7">S. sphaerops</emphasis>
</taxonomicName>
(?), whereas the latter category consists of the members inhabiting open alluvia, like shingle beds, sandy, clayey or silty banks, sometimes sparsely overgrown with grass but without any more or less evident peat-moss or litter layer: 
<taxonomicName id="CF964D68C02AFFCBFCC3AA3DFC1D9FBC" authorityName="Erichson" authorityYear="1840" box="[866,968,1908,1935]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="labilis">
<emphasis id="3AE2EAF9C02AFFCBFCC3AA3DFC1D9FBC" box="[866,968,1908,1935]" italics="true" pageId="6" pageNumber="7">S. labilis</emphasis>
</taxonomicName>
, 
<taxonomicName id="CF964D68C02AFFCBFC77AA3DFB959FBC" authorityName="Ryvkin" authorityYear="2012" box="[982,1088,1908,1935]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="delitor">
<emphasis id="3AE2EAF9C02AFFCBFC77AA3DFB959FBC" box="[982,1088,1908,1935]" italics="true" pageId="6" pageNumber="7">S. delitor</emphasis>
</taxonomicName>
, 
<taxonomicName id="CF964D68C02AFFCBFBEFAA3DFB0A9FBC" authorityName="J. Sahlberg" authorityYear="1880" box="[1102,1247,1908,1935]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="latipennis">
<emphasis id="3AE2EAF9C02AFFCBFBEFAA3DFB0A9FBC" box="[1102,1247,1908,1935]" italics="true" pageId="6" pageNumber="7">S. latipennis</emphasis>
</taxonomicName>
, 
<taxonomicName id="CF964D68C02AFFCBFB4DAA3DFABA9FBC" authorityName="J. Sahlberg" authorityYear="1876" box="[1260,1391,1908,1935]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="confusus">
<emphasis id="3AE2EAF9C02AFFCBFB4DAA3DFABA9FBC" box="[1260,1391,1908,1935]" italics="true" pageId="6" pageNumber="7">S. confusus</emphasis>
</taxonomicName>
, 
<taxonomicName id="CF964D68C02AFFCBFF1CAAD0FEE09F83" authorityName="Casey" authorityYear="1884" box="[189,309,1943,1970]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="dolosus">
<emphasis id="3AE2EAF9C02AFFCBFF1CAAD0FEE09F83" box="[189,309,1943,1970]" italics="true" pageId="6" pageNumber="7">S. dolosus</emphasis>
</taxonomicName>
, 
<taxonomicName id="CF964D68C02AFFCBFEE2AAD0FE2A9F83" authorityName="Gyllenhal" authorityYear="1827" box="[323,511,1943,1970]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="canaliculatus">
<emphasis id="3AE2EAF9C02AFFCBFEE2AAD0FE2A9F83" box="[323,511,1943,1970]" italics="true" pageId="6" pageNumber="7">S. canaliculatus</emphasis>
</taxonomicName>
, 
<taxonomicName id="CF964D68C02AFFCBFDAFAAD0FD5F9F83" authorityName="Ryvkin" authorityYear="2012" box="[526,650,1944,1970]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="canosus">
<emphasis id="3AE2EAF9C02AFFCBFDAFAAD0FD5F9F83" box="[526,650,1944,1970]" italics="true" pageId="6" pageNumber="7">S. canosus</emphasis>
</taxonomicName>
, 
<taxonomicName id="CF964D68C02AFFCBFD38AAD0FCDE9F83" authorityName="Ryvkin" authorityYear="2012" box="[665,779,1943,1970]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="canalis">
<emphasis id="3AE2EAF9C02AFFCBFD38AAD0FCDE9F83" box="[665,779,1943,1970]" italics="true" pageId="6" pageNumber="7">S. canalis</emphasis>
</taxonomicName>
. There are also some species (
<taxonomicName id="CF964D68C02AFFCBFBD2AAD0FB0A9F83" authorityName="Ryvkin" authorityYear="2012" box="[1139,1247,1943,1970]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="alopex">
<emphasis id="3AE2EAF9C02AFFCBFBD2AAD0FB0A9F83" box="[1139,1247,1943,1970]" italics="true" pageId="6" pageNumber="7">S. alopex</emphasis>
</taxonomicName>
, 
<taxonomicName id="CF964D68C02AFFCBFB4FAAD0FABA9F83" authorityName="Casey" authorityYear="1884" box="[1262,1391,1943,1970]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="vinnulus">
<emphasis id="3AE2EAF9C02AFFCBFB4FAAD0FABA9F83" box="[1262,1391,1943,1970]" italics="true" pageId="6" pageNumber="7">S. vinnulus</emphasis>
</taxonomicName>
, and supposedly 
<taxonomicName id="CF964D68C02AFFCBFEDBAAF3FE319FE7" authorityName="Ryvkin" authorityYear="1987" box="[378,484,1979,2005]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="illusor">
<emphasis id="3AE2EAF9C02AFFCBFEDBAAF3FE319FE7" box="[378,484,1979,2005]" italics="true" pageId="6" pageNumber="7">S. illusor</emphasis>
</taxonomicName>
) living in biotopes of both categories. The peculiar swamp species, excluding 
<taxonomicName id="CF964D68C02BFFCAFF1CAC44FE869917" authorityName="Casey" authorityYear="1884" box="[189,339,267,294]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="sphaerops">
<emphasis id="3AE2EAF9C02BFFCAFF1CAC44FE869917" box="[189,339,267,294]" italics="true" pageId="7" pageNumber="8">S. sphaerops</emphasis>
</taxonomicName>
for which too little data are available, are represented only by the 
<taxonomicName id="CF964D68C02BFFCAFBC5AC44FB7F9917" authorityName="Stephens" authorityYear="1833" box="[1124,1194,268,293]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="nitens">
<emphasis id="3AE2EAF9C02BFFCAFBC5AC44FB7F9917" box="[1124,1194,268,293]" italics="true" pageId="7" pageNumber="8">nitens</emphasis>
</taxonomicName>
subgroup, which is supposed to be closest to the common ancestor of the 
<taxonomicName id="CF964D68C02BFFCAFCF1AC67FC3B997B" authorityName="Gyllenhal" authorityYear="1827" box="[848,1006,303,329]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="canaliculatus">
<emphasis id="3AE2EAF9C02BFFCAFCF1AC67FC3B997B" box="[848,1006,303,329]" italics="true" pageId="7" pageNumber="8">canaliculatus</emphasis>
</taxonomicName>
group.
</paragraph>
</subSubSection>
<subSubSection id="408C6560C02BFFCAFF1CAC38FB6B9A10" pageId="7" pageNumber="8" type="distribution">
<paragraph id="082936EBC02BFFCAFF1CAC38FE6899B8" blockId="7.[189,445,368,394]" box="[189,445,368,394]" pageId="7" pageNumber="8">
<heading id="53618187C02BFFCAFF1CAC38FE6899B8" bold="true" box="[189,445,368,394]" fontSize="11" level="3" pageId="7" pageNumber="8" reason="3">
<emphasis id="3AE2EAF9C02BFFCAFF1CAC38FE6899B8" bold="true" box="[189,445,368,394]" pageId="7" pageNumber="8">General distribution</emphasis>
</heading>
</paragraph>
<paragraph id="082936EBC02BFFCAFF1CACD6FB6B9A10" blockId="7.[189,1399,414,547]" pageId="7" pageNumber="8">
Palaearctic except subtropical regions (spotty-relict in the southernmost temperate territories); the whole Nearctic; the Caribbean Islands. Many species of the group are represented in the northernmost territories of the Holarctic.Thus, five of the fifteen Palaearctic species have been recorded for the Taimyr Peninsula and Putorana 
<collectingRegion id="CA52F809C02BFFCAFE79AF40FD9D9A10" box="[472,584,520,546]" country="United Kingdom" name="Highland" pageId="7" pageNumber="8">Highland</collectingRegion>
, which is more than 1/4 of the regional stenine fauna.
</paragraph>
</subSubSection>
</treatment>
</document>