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<document id="A1A56BAB46498461B762A6E576DCF7B6" ID-CLB-Dataset="299644" ID-DOI="10.3897/zse.100.117859" ID-GBIF-Dataset="f17d3dc6-151e-4b2c-bfe6-1e3978daa701" ID-ZooBank="15CA9ED4-C24C-4AD7-81AC-F699FA953FE8" ID-publisher-id="117859" URI-arpha="35315993-8E70-5007-A8AE-277D7EFC9889" URI-zoobank="http://zoobank.org/15CA9ED4-C24C-4AD7-81AC-F699FA953FE8" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" XM.treatmentCitations_approvedBy="admin" XM.treatments_approvedBy="admin" article-type="research-article" checkinTime="1720542344403" checkinUser="pensoft" docAuthor="Yuan, Zi-Ming, Jiang, Wei &amp; Sha, Zhong-Li" docDate="2024" docId="79F6468F6F07551DAA48492439C6E203" docLanguage="en" docName="Zoosystematics and Evolution 100(3): 965-987" docOrigin="Zoosystematics and Evolution 100 (3)" docSource="https://zse.pensoft.net/article/117859/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Gothus teemo Yuan, Jiang &amp; Sha, 2024, sp. nov." docType="treatment" docUuid="9A4FA138-D3F0-4FC0-8687-844B52C5BA15" docUuidSource="ZooBank" docVersion="5" dtd-version="3.0" lastPageNumber="987" masterDocId="353159938E705007A8AE277D7EFC9889" masterDocTitle="Morphological and molecular evidence for Gothus teemo gen. et sp. nov., a new xanthid crab (Crustacea, Brachyura, Xanthoidea) from coral reefs in the South China Sea, with a review of the taxonomy of Actaeodes consobrinus (A. Milne-Edwards, 1867)" masterLastPageNumber="987" masterPageNumber="965" pageNumber="965" updateTime="1720578290841" updateUser="ExternalLinkService">
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<mods:title id="8D8A50B0C568A0CF26BFD0EC45ED0065">Morphological and molecular evidence for Gothus teemo gen. et sp. nov., a new xanthid crab (Crustacea, Brachyura, Xanthoidea) from coral reefs in the South China Sea, with a review of the taxonomy of Actaeodes consobrinus (A. Milne-Edwards, 1867)</mods:title>
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<mods:namePart id="7526440F76CEFB4A4C059578596B2A20">Yuan, Zi-Ming</mods:namePart>
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<mods:affiliation id="03BEF6A1852C36746C1B31481973B46D">Department of Marine Organism Taxonomy and Phylogeny, Institute of Oceanology, Chinese Academy of Sciences, Qingdao 266071, China &amp; Laoshan Laboratory, Qingdao 266237, China &amp; Shandong Province Key Laboratory of Experimental Marine Biology, Institute of Oceanology, Chinese Academy of Sciences, Qingdao, China</mods:affiliation>
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<mods:namePart id="623699D15304645295CB648FFF7E92E8">Jiang, Wei</mods:namePart>
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<mods:affiliation id="6552C5BB4564B2126F85487F9DEA09F2">Department of Marine Organism Taxonomy and Phylogeny, Institute of Oceanology, Chinese Academy of Sciences, Qingdao 266071, China &amp; Laoshan Laboratory, Qingdao 266237, China</mods:affiliation>
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<mods:namePart id="B7ABE3FFEEE8985F62F29390A90BCD9E">Sha, Zhong-Li</mods:namePart>
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<mods:affiliation id="91BEFE26939C2116C97AA7D0A365ED95">Department of Marine Organism Taxonomy and Phylogeny, Institute of Oceanology, Chinese Academy of Sciences, Qingdao 266071, China &amp; Laoshan Laboratory, Qingdao 266237, China</mods:affiliation>
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<mods:date id="B18D886EA39E29EE3277EF008F6C8840">2024</mods:date>
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<treatment id="79F6468F6F07551DAA48492439C6E203" ID-DOI="http://doi.org/10.5281/zenodo.12701267" ID-GBIF-Taxon="235025295" ID-Zenodo-Dep="12701267" ID-arpha="79F6468F-6F07-551D-AA48-492439C6E203" ID-zoobank="https://zoobank.org/9A4FA138-D3F0-4FC0-8687-844B52C5BA15" LSID="urn:lsid:zoobank.org:act:9A4FA138-D3F0-4FC0-8687-844B52C5BA15" LSID-ZBK="urn:lsid:zoobank.org:act:9A4FA138-D3F0-4FC0-8687-844B52C5BA15" httpUri="http://treatment.plazi.org/id/79F6468F6F07551DAA48492439C6E203">
<subSubSection id="3B1B10E1ACF9B975E7095C89CF5A9494" type="nomenclature">
<paragraph id="0083659A78BA8F92EC213F2921A76EDA">
<taxonomicName id="E6F894BA992044D23A753E956C9A30E8" authorityName="Yuan &amp; Jiang &amp; Sha" authorityYear="2024" class="Malacostraca" family="Xanthidae" genus="Gothus" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="teemo" status="sp. nov.">
<emphasis id="E603FEA98A76769F9E0CD4EDB92F706C" italics="true">Gothus teemo</emphasis>
</taxonomicName>
<taxonomicNameLabel id="534A76156579B75C561ED54261FB4A86" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
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<paragraph id="3BBA3E8B60EC05C2230DD2ABAA50DFF1">
<figureCitation id="78F5434FF1634EAF4EE72EAD9BA98132" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Gothus teemo sp. nov. A C. Holotype, male, CW 3.7 mm, CL 2.6 mm, MBM 287027; D F. Paratype, CW 3.2 mm, CL 2.2 mm, MBM 287026; A, D. Dorsal view; B, E. Frontal view; C, F. Carapace. Scale bar: 1 mm." figureDoi="10.3897/zse.100.117859.figure1" httpUri="https://binary.pensoft.net/fig/1088094">Figs 1</figureCitation>
,
<figureCitation id="2B17FAB63C4437951F798A0F7744180C" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Gothus teemo sp. nov. A, C, E. Holotype, male, CW 3.7 mm, CL 2.6 mm, MBM 287027; B, D, F, G. Paratype, female, CW 3.2 mm, CL 2.2 mm, MBM 287026; A. Male chelipeds; B. Female chelipeds; C. Male thoracic sternites; D. Female thoracic sternites; E. Male pleon; F. Female pleon; G. Female vulva. Scale bar: 1 mm (A F); 0.2 mm (G)." figureDoi="10.3897/zse.100.117859.figure2" httpUri="https://binary.pensoft.net/fig/1088095">2</figureCitation>
,
<figureCitation id="F2CDFE7633DCFC3F33EC0178D9F70F75" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Gothus teemo sp. nov. A C, F, G L. Holotype, male, CW 3.7 mm, CL 2.6 mm, MBM 287027; D, E. Paratype, female, CW 3.2 mm, CL 2.2 mm, MBM 287026; A. Carapace; B. Maxilliped 3; C. Cheliped; D. Pereopod 5; E. Female pleon; F. Male pleon; G. right G 1, ventral view; H. Same, dorsal view; I. Right G 1, distal part, ventral view; J. Same, dorsal view; K. Right G 2, ventral view; L. Same, dorsal view. Scale bar: 1 mm (A, C, D); 0.5 mm (B, E, F, G, H); 0.1 mm (I L)." figureDoi="10.3897/zse.100.117859.figure3" httpUri="https://binary.pensoft.net/fig/1088096">3</figureCitation>
,
<figureCitation id="69C5C53BE9FCECE17B025D7CB6AED011" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Gothus teemo sp. nov. A, B. Holotype, male, CW 3.7 mm, CL 2.6 mm, MBM 287027; C E. Paratypes, 2 juvenile, CW 1.82.2 mm, CL 1.31.5 mm, MBM 287023; A D. Live coloration; E. Habitat and substrate conditions. Scale bar: 1 mm." figureDoi="10.3897/zse.100.117859.figure4" httpUri="https://binary.pensoft.net/fig/1088097">4</figureCitation>
,
<figureCitation id="5D901640834D78A5DADE1DCBFD95E533" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Gothus teemo sp. nov., paratype, female, CW 3.2 mm, CL 2.2 mm, MBM 287026, artistic illustration, displaying live coloration. Drawn by Fei Gao." figureDoi="10.3897/zse.100.117859.figure5" httpUri="https://binary.pensoft.net/fig/1088098">5</figureCitation>
</paragraph>
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<subSubSection id="SECID0E3ACI" type="type material">
<paragraph id="BED8E3E24340A9612D26D76F7531E72F">
<heading id="D94011904E847DFC7BE26BB2F9B32095" reason="title">Type material.</heading>
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<emphasis id="BFE1E35C2C3BCE77DF425FD8409D7BC7" bold="true" italics="true">
<typeStatus id="3674E96D9EED80E55D401C38B8FC02DA" type="holotype">Holotype</typeStatus>
.
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;
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;
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<collectorName id="8CEEC4A5FF4593D9223E35A33C264058">Ziming Yuan</collectorName>
coll.; 2404189149;
<specimenCode id="0398CCD27ABD9599AA00C1D5EC6CE831" collectionCode="MBM" country="0" httpUri="http://grbio.org/cool/fc7c-p25f" name="San Jose State University, Museum of Birds and Mammals" type="">MBM 287027</specimenCode>
.
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</paragraph>
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<typeStatus id="D4062C2E857CEC1D7E92B57D148BF6D1" type="paratype">Paratypes</typeStatus>
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.
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;
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<quantity id="32AB49D164087B3B0AC179261B6895AB" metricMagnitude="-3" metricUnit="m" metricValue="3.2" unit="mm" value="3.2">3.2 mm</quantity>
,
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<quantity id="B93A8CCD5D26F9FA51980603FCC8E254" metricMagnitude="-3" metricUnit="m" metricValue="2.2" unit="mm" value="2.2">2.2 mm</quantity>
;
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;
<collectorName id="F19DC4AD003750A8FC56B6A62C2746F4">Aiyang Wang</collectorName>
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<collectorName id="E5812540C1A2AB27CB63E9E5ED3DACAE">Bingqin Liu</collectorName>
coll.; 2304278379;
<specimenCode id="CD8146020B1C69401DE82EDD4841B730" collectionCode="MBM" country="0" httpUri="http://grbio.org/cool/fc7c-p25f" name="San Jose State University, Museum of Birds and Mammals" type="">MBM 287026</specimenCode>
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<materialsCitation id="6FBEACABEF2F8A930F904A777BE256E3" ID-GBIF-Occurrence="4904612313" collectingDate="2023-04-27" collectionCode="MBM" collectorName="Aiyang Wang &amp; Bingqin Liu" country="China" latitude="9.883653" location="Meiji Reef" longLatPrecision="1" longitude="115.560394" municipality="Nansha Islands" specimenCode="MBM 287024" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
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(decalcified);
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coll.; 2304278461;
<specimenCode id="C380EC0D16B4F3B1ADCB53FF1CBE6D4A" collectionCode="MBM" country="0" httpUri="http://grbio.org/cool/fc7c-p25f" name="San Jose State University, Museum of Birds and Mammals" type="">MBM 287024</specimenCode>
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<specimenCount id="9AA7DDEF894C16D81B9AE829DC295066" count="1" type="male">1 male</specimenCount>
(partially crushed);
<abbrev id="ABBRID0E5CCI" xlink_title="maximum carapace width">CW</abbrev>
<quantity id="EEB73BDCE6D0BC113E1471747A56C867" metricMagnitude="-3" metricUnit="m" metricValue="4.0" unit="mm" value="4.0">4 mm</quantity>
,
<abbrev id="ABBRID0ECDCI" xlink_title="median carapace length">CL</abbrev>
<quantity id="444D0606255710FEF064035A0289CD95" metricMagnitude="-3" metricUnit="m" metricValue="2.7" unit="mm" value="2.7">2.7 mm</quantity>
;
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,
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;
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<geoCoordinate id="739D28F811DDC53483A2C67F610F96C8" degrees="9" direction="north" minutes="53" orientation="latitude" precision="1" seconds="1.15" value="9.883653">9 ° 53 ' 1.15 &quot; N</geoCoordinate>
,
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;
<date id="26A9E7EDB4506160454F5AECFEE010E9" value="2023-04-27">
<collectingDate id="0C64ADCC288E3B91D89B3D3E0C59793D" value="2023-04-27">27 Apr. 2023</collectingDate>
</date>
;
<collectorName id="A8D019DD4323A158CB4960AC0DA7CB9B">Aiyang Wang</collectorName>
,
<collectorName id="A972E98BB35931E5232A65460C0B29F4">Bingqin Liu</collectorName>
coll.; 2304278486;
<specimenCode id="339435B4760C286936BC4B6CE27C0DFB" collectionCode="MBM" country="0" httpUri="http://grbio.org/cool/fc7c-p25f" name="San Jose State University, Museum of Birds and Mammals" type="">MBM 287025</specimenCode>
</materialsCitation>
<materialsCitation id="99AA99A4096DC3EF15BA149437D955F2" ID-GBIF-Occurrence="4904612303" collectingDate="2022-05-06" collectionCode="MBM" collectorName="Ziming Yuan &amp; Yuli Sun &amp; Shaobo Ma" country="China" elevation="3" latitude="9.907152" location="Meiji Reef" longLatPrecision="1" longitude="115.49706" municipality="Nansha Islands" specimenCode="MBM 287023" specimenCount="2" specimenCount-juvenile="2" typeStatus="paratype">
<specimenCount id="F3B106D64EEA6458F9935303CC4DEF0A" count="2" type="juvenile">2 juveniles</specimenCount>
;
<abbrev id="ABBRID0EODCI" xlink_title="maximum carapace width">CW</abbrev>
<quantity id="32457CFD4BB0D42CA91FF7F26687B0EB" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="2.2" metricValueMin="1.8" unit="mm" value="2.0" valueMax="2.2" valueMin="1.8">1.82.2 mm</quantity>
,
<abbrev id="ABBRID0ESDCI" xlink_title="median carapace length">CL</abbrev>
<quantity id="41508B70BDCCC16D0025AF23DA12E413" metricMagnitude="-3" metricUnit="m" metricValue="1.4" metricValueMax="1.5" metricValueMin="1.3" unit="mm" value="1.4" valueMax="1.5" valueMin="1.3">1.31.5 mm</quantity>
;
<location id="8D026B0438A8921A4951A1C40771D7C3" LSID="urn:lsid:plazi:treatment:79F6468F6F07551DAA48492439C6E203:8D026B0438A8921A4951A1C40771D7C3" country="China" latitude="9.907152" longLatPrecision="1" longitude="115.49706" municipality="Nansha Islands" name="Meiji Reef">Meiji Reef</location>
,
<collectingMunicipality id="6A1B6A374246A6DA2A5F4BFE5384AF02">Nansha Islands</collectingMunicipality>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EZDCI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[115.497067,9.907153]}">
<geoCoordinate id="83926F44FEB1125E4EDB4AA1EA3528E4" degrees="9" direction="north" minutes="54" orientation="latitude" precision="1" seconds="25.75" value="9.907152">9 ° 54 ' 25.75 &quot; N</geoCoordinate>
,
<geoCoordinate id="425660934ED2A377116DF5BDF131FD54" degrees="115" direction="east" minutes="29" orientation="longitude" precision="1" seconds="49.44" value="115.49706">115 ° 29 ' 49.44 &quot; E</geoCoordinate>
</named-content>
;
<quantity id="139C917412223DECA658F7BE5665B9E8" metricMagnitude="0" metricUnit="m" metricValue="3.0" unit="m" value="3.0">
<elevation id="C49FAA2548F6EDCCFF57A64B99CA8CEB" metricMagnitude="0" metricUnit="m" metricValue="3.0" unit="m" value="3.0">3 m</elevation>
</quantity>
;
<date id="5CB1C0F8F5FE05691276CDA7A20CAFD2" value="2022-05-06">
<collectingDate id="A6671C6F37E42AF6EE2F13930C858DA6" value="2022-05-06">6 May 2022</collectingDate>
</date>
;
<collectorName id="0F2BF1D453DD22EA5DAB3952A39B9FA5">Ziming Yuan</collectorName>
,
<collectorName id="E1BB7C6BF26BB3294E8B9D5BE9C5F0B1">Yuli Sun</collectorName>
,
<collectorName id="4E50BC402DA91BA51DD63F86C565974A">Shaobo Ma</collectorName>
coll.;
<specimenCode id="5389C9DCF9C987C7C144F9317B23BC28">NS-MJ- 2022-1287</specimenCode>
;
<specimenCode id="FDFFD220181AF49F57659073AE025497" collectionCode="MBM" country="0" httpUri="http://grbio.org/cool/fc7c-p25f" name="San Jose State University, Museum of Birds and Mammals" type="">MBM 287023</specimenCode>
</materialsCitation>
<materialsCitation id="0B87BFB8CDF8D7821F3E5F804E75DEEF" ID-GBIF-Occurrence="4904612356" collectingDate="2024-04-11" collectionCode="MBM" collectorName="Ziming Yuan" country="China" elevation="5" latitude="9.8919" location="Meiji Reef" longLatPrecision="1" longitude="115.57279" municipality="Nansha Islands" specimenCode="MBM 287022" specimenCount="1" specimenCount-juvenile="1" typeStatus="paratype">
<specimenCount id="270C554E5A65CE1588F4D5E9E7D36E4A" count="1" type="juvenile">1 juvenile</specimenCount>
;
<abbrev id="ABBRID0E5DCI" xlink_title="maximum carapace width">CW</abbrev>
<quantity id="8C9A809EFEA96364AF4F419029BDD4D5" metricMagnitude="-3" metricUnit="m" metricValue="2.0" unit="mm" value="2.0">2 mm</quantity>
,
<abbrev id="ABBRID0ECECI" xlink_title="median carapace length">CL</abbrev>
<quantity id="2D07F075D288646D80E9D7E8B1EB68E9" metricMagnitude="-3" metricUnit="m" metricValue="1.3" unit="mm" value="1.3">1.3 mm</quantity>
;
<location id="FD93DB2EEF107F185AA790FA0943EE7D" LSID="urn:lsid:plazi:treatment:79F6468F6F07551DAA48492439C6E203:FD93DB2EEF107F185AA790FA0943EE7D" country="China" latitude="9.8919" longLatPrecision="1" longitude="115.57279" municipality="Nansha Islands" name="Meiji Reef">Meiji Reef</location>
,
<collectingMunicipality id="704C07EC61E3870003786FBAF45EE14F">Nansha Islands</collectingMunicipality>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EJECI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[115.572792,9.891900]}">
<geoCoordinate id="CAADC176E4E8F3E035C573C18810DDDD" degrees="9" direction="north" minutes="53" orientation="latitude" precision="1" seconds="30.84" value="9.8919">9 ° 53 ' 30.84 &quot; N</geoCoordinate>
,
<geoCoordinate id="06D439815F9BD25895FD75F121200CA8" degrees="115" direction="east" minutes="34" orientation="longitude" precision="1" seconds="22.05" value="115.57279">115 ° 34 ' 22.05 &quot; E</geoCoordinate>
</named-content>
;
<quantity id="499EFCE2AFF9488653E5A8DA43C3A617" metricMagnitude="0" metricUnit="m" metricValue="5.0" unit="m" value="5.0">
<elevation id="4419CBFDC3532FE6025A6B1A2A627C23" metricMagnitude="0" metricUnit="m" metricValue="5.0" unit="m" value="5.0">5 m</elevation>
</quantity>
;
<date id="13A725A93F9E6005AC293EC728EFB448" value="2024-04-11">
<collectingDate id="EA54809F2F7488DADF5CD76FE97012F6" value="2024-04-11">11 Apr. 2024</collectingDate>
</date>
;
<collectorName id="C31AD5FBA315F0AD727CE874903491D0">Ziming Yuan</collectorName>
coll.; 2404188048;
<specimenCode id="37D839C053DB7A532BC781E28C312445" collectionCode="MBM" country="0" httpUri="http://grbio.org/cool/fc7c-p25f" name="San Jose State University, Museum of Birds and Mammals" type="">MBM 287022</specimenCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EOECI" type="description">
<paragraph id="153A6D020D12881BCF4B0752FD2F8F0F">
<heading id="E3AA2DCBB3D9C276BAFCF20128B5E440" reason="title">Description.</heading>
</paragraph>
<paragraph id="E2C301093FCE0FC6BE3D6BFB6F81DF66">
Carapace (Figs
<figureCitation id="75A04C00616ABEAB3B710395C167E454" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Gothus teemo sp. nov. A C. Holotype, male, CW 3.7 mm, CL 2.6 mm, MBM 287027; D F. Paratype, CW 3.2 mm, CL 2.2 mm, MBM 287026; A, D. Dorsal view; B, E. Frontal view; C, F. Carapace. Scale bar: 1 mm." figureDoi="10.3897/zse.100.117859.figure1" httpUri="https://binary.pensoft.net/fig/1088094">1</figureCitation>
,
<figureCitation id="39C5CE387EF2504849261AF9362B9AE4" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Gothus teemo sp. nov. A C, F, G L. Holotype, male, CW 3.7 mm, CL 2.6 mm, MBM 287027; D, E. Paratype, female, CW 3.2 mm, CL 2.2 mm, MBM 287026; A. Carapace; B. Maxilliped 3; C. Cheliped; D. Pereopod 5; E. Female pleon; F. Male pleon; G. right G 1, ventral view; H. Same, dorsal view; I. Right G 1, distal part, ventral view; J. Same, dorsal view; K. Right G 2, ventral view; L. Same, dorsal view. Scale bar: 1 mm (A, C, D); 0.5 mm (B, E, F, G, H); 0.1 mm (I L)." figureDoi="10.3897/zse.100.117859.figure3" httpUri="https://binary.pensoft.net/fig/1088096">3 A</figureCitation>
) broader than long,
<abbrev id="ABBRID0E3ECI" xlink_title="maximum carapace width">CW</abbrev>
about 1.5 times the
<abbrev id="ABBRID0EAFCI" xlink_title="median carapace length">CL</abbrev>
, dorsal surface bearing round granules, regions well defined, 1 M separated from 2 M by shallow transverse groove; 2 M indistinct divided, lateral lobe with elevated pointed tuberosity; 3 M distinct, undivided; 4 M indistinct; 1 L, 3 L, 4 L indistinct; 2 L, 5 l, 6 L distinct, with elevated sharp tuberosity each on 2 L, 5 L; 1 P distinct, 2 P indistinct, flat; 2 M, 5 L, and 6 L regions each with a tuft of setae; front wide, about 0.4 times
<abbrev id="ABBRID0EEFCI" xlink_title="maximum carapace width">CW</abbrev>
, not protruding, divided into two slightly triangular lobes by a V-shaped notch, a small pore visible from front introduced from median notch (Fig.
<figureCitation id="5A62F53C9153D0AAC52A4D1325CB9AA8" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Gothus teemo sp. nov. A C. Holotype, male, CW 3.7 mm, CL 2.6 mm, MBM 287027; D F. Paratype, CW 3.2 mm, CL 2.2 mm, MBM 287026; A, D. Dorsal view; B, E. Frontal view; C, F. Carapace. Scale bar: 1 mm." figureDoi="10.3897/zse.100.117859.figure1" httpUri="https://binary.pensoft.net/fig/1088094">1 B</figureCitation>
), frontal lobes and dorsal inner orbital angle separated by shallow depression; eyestalks densely granulated; area behind outer orbital angle slightly concave, not forming a subhepatic cavity; anterolateral margin starting from subhepatic region; first tooth nearly completely reduced; subsequent three teeth developed; second and third teeth nearly equal; fourth tooth smaller; carapace widest at second tooth; posterolateral margin almost straight; subhepatic region densely granulated; posterior margin nearly straight.
</paragraph>
<paragraph id="48BA2B67276CFE8018A38D3DD7DF8709">
Epistome (Fig.
<figureCitation id="8134A4B215EEB13015AAA357219D00C9" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Gothus teemo sp. nov. A C. Holotype, male, CW 3.7 mm, CL 2.6 mm, MBM 287027; D F. Paratype, CW 3.2 mm, CL 2.2 mm, MBM 287026; A, D. Dorsal view; B, E. Frontal view; C, F. Carapace. Scale bar: 1 mm." figureDoi="10.3897/zse.100.117859.figure1" httpUri="https://binary.pensoft.net/fig/1088094">1 B, E</figureCitation>
) central region with short median projection on posterior margin pronounced; lateral regions with undulating posterior margins; maxilliped 3 (Figs
<figureCitation id="BA0B5795F2C9B84F5578D8EE83D53E14" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Gothus teemo sp. nov. A C. Holotype, male, CW 3.7 mm, CL 2.6 mm, MBM 287027; D F. Paratype, CW 3.2 mm, CL 2.2 mm, MBM 287026; A, D. Dorsal view; B, E. Frontal view; C, F. Carapace. Scale bar: 1 mm." figureDoi="10.3897/zse.100.117859.figure1" httpUri="https://binary.pensoft.net/fig/1088094">1 B, E</figureCitation>
,
<figureCitation id="6CE72D5AD37A20399E19E79BD847D69F" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Gothus teemo sp. nov. A C, F, G L. Holotype, male, CW 3.7 mm, CL 2.6 mm, MBM 287027; D, E. Paratype, female, CW 3.2 mm, CL 2.2 mm, MBM 287026; A. Carapace; B. Maxilliped 3; C. Cheliped; D. Pereopod 5; E. Female pleon; F. Male pleon; G. right G 1, ventral view; H. Same, dorsal view; I. Right G 1, distal part, ventral view; J. Same, dorsal view; K. Right G 2, ventral view; L. Same, dorsal view. Scale bar: 1 mm (A, C, D); 0.5 mm (B, E, F, G, H); 0.1 mm (I L)." figureDoi="10.3897/zse.100.117859.figure3" httpUri="https://binary.pensoft.net/fig/1088096">3 B</figureCitation>
) granulated; ischium subrectangular; presenting submedian groove; merus subquadrate; anterior margin indented; external anteroexternal angle expanded. Antennule (Fig.
<figureCitation id="122CAD828CCB8902CA8B37B1F821D26E" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Gothus teemo sp. nov. A C. Holotype, male, CW 3.7 mm, CL 2.6 mm, MBM 287027; D F. Paratype, CW 3.2 mm, CL 2.2 mm, MBM 287026; A, D. Dorsal view; B, E. Frontal view; C, F. Carapace. Scale bar: 1 mm." figureDoi="10.3897/zse.100.117859.figure1" httpUri="https://binary.pensoft.net/fig/1088094">1 B, E</figureCitation>
) folding transversely; basal segment of antenna subrectangular; contacting ventral external frontal margin and ventral internal orbital angle; antennal flagellum filling orbital hiatus.
</paragraph>
<paragraph id="655067ACEB7A3FB9880D59D3C31A408F">
Chelipeds (Figs
<figureCitation id="CBAAAD6738A39C0CD7B8939D923AD783" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Gothus teemo sp. nov. A, C, E. Holotype, male, CW 3.7 mm, CL 2.6 mm, MBM 287027; B, D, F, G. Paratype, female, CW 3.2 mm, CL 2.2 mm, MBM 287026; A. Male chelipeds; B. Female chelipeds; C. Male thoracic sternites; D. Female thoracic sternites; E. Male pleon; F. Female pleon; G. Female vulva. Scale bar: 1 mm (A F); 0.2 mm (G)." figureDoi="10.3897/zse.100.117859.figure2" httpUri="https://binary.pensoft.net/fig/1088095">2 A, B</figureCitation>
,
<figureCitation id="80AC984DBB27D3FF410412134675055D" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Gothus teemo sp. nov. A C, F, G L. Holotype, male, CW 3.7 mm, CL 2.6 mm, MBM 287027; D, E. Paratype, female, CW 3.2 mm, CL 2.2 mm, MBM 287026; A. Carapace; B. Maxilliped 3; C. Cheliped; D. Pereopod 5; E. Female pleon; F. Male pleon; G. right G 1, ventral view; H. Same, dorsal view; I. Right G 1, distal part, ventral view; J. Same, dorsal view; K. Right G 2, ventral view; L. Same, dorsal view. Scale bar: 1 mm (A, C, D); 0.5 mm (B, E, F, G, H); 0.1 mm (I L)." figureDoi="10.3897/zse.100.117859.figure3" httpUri="https://binary.pensoft.net/fig/1088096">3 C</figureCitation>
) symmetrical; meri short; carpi robust, with square-shaped outward expansion, surface granulated, aggregated into nodules; palms dorsally with three protuberances, outer and dorsal surfaces densely granulated; fingers elongated, with triangular teeth, tips sharp, crossing at extremities when closed; dorsal surface of movable finger with three granular ridges; outer surface of immovable finger with two granular ridges.
</paragraph>
<paragraph id="2C4251BD2555E0AE10CA68C6937FD22E">
Ambulatory legs (Figs
<figureCitation id="817C2FC4A84F7F01504871C4E05D7184" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Gothus teemo sp. nov. A C. Holotype, male, CW 3.7 mm, CL 2.6 mm, MBM 287027; D F. Paratype, CW 3.2 mm, CL 2.2 mm, MBM 287026; A, D. Dorsal view; B, E. Frontal view; C, F. Carapace. Scale bar: 1 mm." figureDoi="10.3897/zse.100.117859.figure1" httpUri="https://binary.pensoft.net/fig/1088094">1 A, D</figureCitation>
,
<figureCitation id="12DC377AD4C19B83F442174870FCF30C" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Gothus teemo sp. nov. A C, F, G L. Holotype, male, CW 3.7 mm, CL 2.6 mm, MBM 287027; D, E. Paratype, female, CW 3.2 mm, CL 2.2 mm, MBM 287026; A. Carapace; B. Maxilliped 3; C. Cheliped; D. Pereopod 5; E. Female pleon; F. Male pleon; G. right G 1, ventral view; H. Same, dorsal view; I. Right G 1, distal part, ventral view; J. Same, dorsal view; K. Right G 2, ventral view; L. Same, dorsal view. Scale bar: 1 mm (A, C, D); 0.5 mm (B, E, F, G, H); 0.1 mm (I L)." figureDoi="10.3897/zse.100.117859.figure3" httpUri="https://binary.pensoft.net/fig/1088096">3 D</figureCitation>
) meri flattened, P 5 merus length about 3.6 times width of distal end, granulated dorsally and along anterior and posterior edges; carpi granulated dorsally and along anterior edge, dorsal surface with a grooved indentation near anterior edge; propodi granulated dorsally and along edges; dactyli elongated, margins armed with granules and setae, terminal end chitinous, sharp, slightly curved backward, dactylo-propodal lock very weak and inconspicuous.
</paragraph>
<paragraph id="5DF323E82E1B604A4194B4EB94079E6D">
Male thoracic sternum (Fig.
<figureCitation id="F2CC5C029FA3D4A2DFAB6F8671C5722F" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Gothus teemo sp. nov. A, C, E. Holotype, male, CW 3.7 mm, CL 2.6 mm, MBM 287027; B, D, F, G. Paratype, female, CW 3.2 mm, CL 2.2 mm, MBM 287026; A. Male chelipeds; B. Female chelipeds; C. Male thoracic sternites; D. Female thoracic sternites; E. Male pleon; F. Female pleon; G. Female vulva. Scale bar: 1 mm (A F); 0.2 mm (G)." figureDoi="10.3897/zse.100.117859.figure2" httpUri="https://binary.pensoft.net/fig/1088095">2 C</figureCitation>
) with sternites 1 and 2 completely fused, suture between sternites 2 and 3 straight, complete, sternites 3 and 4 mostly fused, suture between them visible only at margins, sternites 3 short, sternite 4 with central longitudinal groove, tubercle of sterno-pleonal lock (press-button mechanism) located on posterior of sternite 5. Male pleon (Figs
<figureCitation id="5F126AD12E919B07F7E8DA3E47F9D6D8" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Gothus teemo sp. nov. A, C, E. Holotype, male, CW 3.7 mm, CL 2.6 mm, MBM 287027; B, D, F, G. Paratype, female, CW 3.2 mm, CL 2.2 mm, MBM 287026; A. Male chelipeds; B. Female chelipeds; C. Male thoracic sternites; D. Female thoracic sternites; E. Male pleon; F. Female pleon; G. Female vulva. Scale bar: 1 mm (A F); 0.2 mm (G)." figureDoi="10.3897/zse.100.117859.figure2" httpUri="https://binary.pensoft.net/fig/1088095">2 C, E</figureCitation>
,
<figureCitation id="BA07865FA9E3EEC2A185DF423AF0C766" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Gothus teemo sp. nov. A C, F, G L. Holotype, male, CW 3.7 mm, CL 2.6 mm, MBM 287027; D, E. Paratype, female, CW 3.2 mm, CL 2.2 mm, MBM 287026; A. Carapace; B. Maxilliped 3; C. Cheliped; D. Pereopod 5; E. Female pleon; F. Male pleon; G. right G 1, ventral view; H. Same, dorsal view; I. Right G 1, distal part, ventral view; J. Same, dorsal view; K. Right G 2, ventral view; L. Same, dorsal view. Scale bar: 1 mm (A, C, D); 0.5 mm (B, E, F, G, H); 0.1 mm (I L)." figureDoi="10.3897/zse.100.117859.figure3" httpUri="https://binary.pensoft.net/fig/1088096">3 F</figureCitation>
) narrow; pleonites 3 to 5 completely fused, lateral margins of pleonite 6 slightly concave, telson long and broad, width slightly greater than length, truncated oval, basal margin wider than terminal margin of pleonite 6; paired tufts of setae present on margins of middle part and terminal margin of pleonites 35 and terminal margin of pleonite 6. Female pleon (Figs
<figureCitation id="EF463C02162EE922AE42285FFD002ACA" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Gothus teemo sp. nov. A, C, E. Holotype, male, CW 3.7 mm, CL 2.6 mm, MBM 287027; B, D, F, G. Paratype, female, CW 3.2 mm, CL 2.2 mm, MBM 287026; A. Male chelipeds; B. Female chelipeds; C. Male thoracic sternites; D. Female thoracic sternites; E. Male pleon; F. Female pleon; G. Female vulva. Scale bar: 1 mm (A F); 0.2 mm (G)." figureDoi="10.3897/zse.100.117859.figure2" httpUri="https://binary.pensoft.net/fig/1088095">2 D, F</figureCitation>
,
<figureCitation id="BD24E0EB5D98A32A51D2DF8C8947AD22" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Gothus teemo sp. nov. A C, F, G L. Holotype, male, CW 3.7 mm, CL 2.6 mm, MBM 287027; D, E. Paratype, female, CW 3.2 mm, CL 2.2 mm, MBM 287026; A. Carapace; B. Maxilliped 3; C. Cheliped; D. Pereopod 5; E. Female pleon; F. Male pleon; G. right G 1, ventral view; H. Same, dorsal view; I. Right G 1, distal part, ventral view; J. Same, dorsal view; K. Right G 2, ventral view; L. Same, dorsal view. Scale bar: 1 mm (A, C, D); 0.5 mm (B, E, F, G, H); 0.1 mm (I L)." figureDoi="10.3897/zse.100.117859.figure3" httpUri="https://binary.pensoft.net/fig/1088096">3 E</figureCitation>
) oval-shaped, margins of pleonite 6 slightly concave; telson triangular; vulva longitudinally ovate, located at upper of sternite 6, occupying anteromedial half, near sternites 5 / 6 (Fig.
<figureCitation id="5DEEA4C48B57372DF3F9581487BF1821" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Gothus teemo sp. nov. A, C, E. Holotype, male, CW 3.7 mm, CL 2.6 mm, MBM 287027; B, D, F, G. Paratype, female, CW 3.2 mm, CL 2.2 mm, MBM 287026; A. Male chelipeds; B. Female chelipeds; C. Male thoracic sternites; D. Female thoracic sternites; E. Male pleon; F. Female pleon; G. Female vulva. Scale bar: 1 mm (A F); 0.2 mm (G)." figureDoi="10.3897/zse.100.117859.figure2" httpUri="https://binary.pensoft.net/fig/1088095">2 G</figureCitation>
).
</paragraph>
<paragraph id="AD31A083FF4428089977F2FBFA6D7DAE">
<abbrev id="ABBRID0EOHCI" xlink_title="first gonopod of male">G 1</abbrev>
(Fig.
<figureCitation id="C3B0562568C849BC0442560D9C72F2E8" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Gothus teemo sp. nov. A C, F, G L. Holotype, male, CW 3.7 mm, CL 2.6 mm, MBM 287027; D, E. Paratype, female, CW 3.2 mm, CL 2.2 mm, MBM 287026; A. Carapace; B. Maxilliped 3; C. Cheliped; D. Pereopod 5; E. Female pleon; F. Male pleon; G. right G 1, ventral view; H. Same, dorsal view; I. Right G 1, distal part, ventral view; J. Same, dorsal view; K. Right G 2, ventral view; L. Same, dorsal view. Scale bar: 1 mm (A, C, D); 0.5 mm (B, E, F, G, H); 0.1 mm (I L)." figureDoi="10.3897/zse.100.117859.figure3" httpUri="https://binary.pensoft.net/fig/1088096">3 G, H, I, J</figureCitation>
) slender, distal lobe prominent but not excessively elongated, long setae on inner subdistal side, small spines on outer side.
<abbrev id="ABBRID0EWHCI" xlink_title="second gonopod of male">G 2</abbrev>
(Fig.
<figureCitation id="980300CFA6486957D6F157A642A562A8" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Gothus teemo sp. nov. A C, F, G L. Holotype, male, CW 3.7 mm, CL 2.6 mm, MBM 287027; D, E. Paratype, female, CW 3.2 mm, CL 2.2 mm, MBM 287026; A. Carapace; B. Maxilliped 3; C. Cheliped; D. Pereopod 5; E. Female pleon; F. Male pleon; G. right G 1, ventral view; H. Same, dorsal view; I. Right G 1, distal part, ventral view; J. Same, dorsal view; K. Right G 2, ventral view; L. Same, dorsal view. Scale bar: 1 mm (A, C, D); 0.5 mm (B, E, F, G, H); 0.1 mm (I L)." figureDoi="10.3897/zse.100.117859.figure3" httpUri="https://binary.pensoft.net/fig/1088096">3 K, L</figureCitation>
) short, distal lobe elongated, slightly curved upwards.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E5HCI" type="Live coloration">
<paragraph id="D015C5AA7A629C3EE3509F1A6A953B7D">
<heading id="6979BEC763E4EF36C7CE8CABB402EDD1" reason="title">Live coloration.</heading>
</paragraph>
<paragraph id="48DA380B53796A774C13F4C9F8109EFA">
Overall white to coral pink in coloration, carapace adorned with symmetrical black to brown stripes, ambulatory legs and chelipeds bearing stripes of similar coloration, anterior part of chelipeds carpus red (Figs
<figureCitation id="6E6F5A8969029E8B748086A148FD43A2" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Gothus teemo sp. nov. A, B. Holotype, male, CW 3.7 mm, CL 2.6 mm, MBM 287027; C E. Paratypes, 2 juvenile, CW 1.82.2 mm, CL 1.31.5 mm, MBM 287023; A D. Live coloration; E. Habitat and substrate conditions. Scale bar: 1 mm." figureDoi="10.3897/zse.100.117859.figure4" httpUri="https://binary.pensoft.net/fig/1088097">4</figureCitation>
,
<figureCitation id="81A5927510C3DAA7B075DB41ED00FBF6" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Gothus teemo sp. nov., paratype, female, CW 3.2 mm, CL 2.2 mm, MBM 287026, artistic illustration, displaying live coloration. Drawn by Fei Gao." figureDoi="10.3897/zse.100.117859.figure5" httpUri="https://binary.pensoft.net/fig/1088098">5</figureCitation>
). Fingers brownish-black, the coloration of immovable finger extending onto the palm along both the inner and outer surfaces in male (Fig.
<figureCitation id="73BCC1950EB4FCB2E9214B89FA3E36A4" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Gothus teemo sp. nov. A, C, E. Holotype, male, CW 3.7 mm, CL 2.6 mm, MBM 287027; B, D, F, G. Paratype, female, CW 3.2 mm, CL 2.2 mm, MBM 287026; A. Male chelipeds; B. Female chelipeds; C. Male thoracic sternites; D. Female thoracic sternites; E. Male pleon; F. Female pleon; G. Female vulva. Scale bar: 1 mm (A F); 0.2 mm (G)." figureDoi="10.3897/zse.100.117859.figure2" httpUri="https://binary.pensoft.net/fig/1088095">2 A, B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EQICI" type="etymology">
<paragraph id="9F5CC984CCC3D157DAEB3E8C200E341D">
<heading id="1A36C5485A94EE57836CB57FB89746A7" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="1B5B44262968D65F4437517338039C32">
The new species is named after Teemo, a character from the
<collectionCode id="4E69C21F1D62FFFA3E53259817A7DF1A">MOBA</collectionCode>
(Multiplayer Online Battle Arena) video game League of Legends. This character, modeled after a raccoon, has a fluffy, diminutive stature with a brown and white intermingled fur coat. This alludes to the new species small size, densely covered short setae, and brown-striped coloration.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EVICI" type="distribution">
<paragraph id="275D90E0EE597A9D32C29BF4888F4A66">
<heading id="108E3946F4FD27536FBFA95474A6BCB5" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="AD31FEB31934B204CC62F1A24066008D">
Currently known from the type locality at Triton Island, Xisha Islands (Paracel Islands), and Meiji Reef (Mischief Reef), Nansha Islands (
<collectingCountry id="7E4ABDE797ACF553E29F9F4177C3E424" name="Spratly Islands">Spratly Islands</collectingCountry>
), it inhabits crevices in shallow coral reefs.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E1ICI" type="remarks">
<paragraph id="894B6B6C401466FAD05E1F33DD197A8A">
<heading id="65E5AD2232846D9C459BBAE9C25AA5FA" reason="title">Remarks.</heading>
</paragraph>
<paragraph id="4795F7F60322F1715F7026AFEBF46065">
Apart from the members of the subfamily
<taxonomicName id="468C744446652A2698BC8A3CD778D7EA" authorityName="Alcock" authorityYear="1898" class="Malacostraca" family="Xanthidae" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="subFamily" subFamily="Euxanthinae">Euxanthinae</taxonomicName>
already compared in the remarks of
<taxonomicName id="C6F7B342AA61293F698A16636B61D7F3" authorityName="Yuan &amp; Jiang &amp; Sha" authorityYear="2024" class="Malacostraca" family="Xanthidae" genus="Gothus" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="0F6F26701EF255FE25400F668327F602" italics="true">Gothus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CADF2EA391EBCF07201EA894371D616D" rank="genus">gen. nov.</taxonomicNameLabel>
, the species is most similar to
<taxonomicName id="CED8B772ECDA482D4D06EA0C66BDEEF1" authority="(A. Milne-Edwards, 1873)" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1873" class="Malacostraca" family="Xanthidae" genus="Actaeodes" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="consobrinus">
<emphasis id="E2260FD39C3E60C97696D36D79999DDD" italics="true">Actaeodes consobrinus</emphasis>
(A.
<bibRefCitation id="71595F5FE576CDCC57D97844D55D3D42" DOI="10.5962/bhl.title.10644" author="Milne-Edwards A" journalOrPublisher="Journal des Museum Godeffroy" pagination="253-264" refId="B27" refString="Milne-Edwards A (1873) Description de Quelques Crustaces nouveaux ou peu connus provenant du musee de M. C. Godeffroy. Journal des Museum Godeffroy 1: 253 - 264. [pls. 12 - 13] https: // doi. org / 10.5962 / bhl. title. 10644" title="Description de Quelques Crustacés nouveaux ou peu connus provenant du musée de M. C. Godeffroy." volume="1" year="1873">Milne-Edwards, 1873</bibRefCitation>
)
</taxonomicName>
. They share similarities in the carapace outline, the shape of the male pleon, and even in the pattern of the live coloration. However,
<taxonomicName id="E782218F0D9D108BBC2435E5A289ABE8" authorityName="Yuan &amp; Jiang &amp; Sha" authorityYear="2024" class="Malacostraca" family="Xanthidae" genus="Gothus" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="teemo" status="sp. nov.">
<emphasis id="89C136E9863A2421E7E17DB0C38D888B" italics="true">G. teemo</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D88C229A340E93ACFD91B56B2E415237" rank="species">sp. nov.</taxonomicNameLabel>
can be differentiated from
<taxonomicName id="7CEF6D80837E40B21AB162C42E8323C6" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1873" class="Malacostraca" family="Xanthidae" genus="Actaeodes" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="consobrinus">
<emphasis id="D2543004F50C5E035E74D1489395D623" italics="true">A. consobrinus</emphasis>
</taxonomicName>
by the following features: the first tooth on the anterolateral margin of the carapace is completely reduced, almost invisible (Figs
<figureCitation id="3AB3988104F9E1C9FED3C0056DDB0669" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Gothus teemo sp. nov. A C. Holotype, male, CW 3.7 mm, CL 2.6 mm, MBM 287027; D F. Paratype, CW 3.2 mm, CL 2.2 mm, MBM 287026; A, D. Dorsal view; B, E. Frontal view; C, F. Carapace. Scale bar: 1 mm." figureDoi="10.3897/zse.100.117859.figure1" httpUri="https://binary.pensoft.net/fig/1088094">1 A, C</figureCitation>
,
<figureCitation id="9F03920996A525D4967D871CE889CC4F" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Gothus teemo sp. nov. A C, F, G L. Holotype, male, CW 3.7 mm, CL 2.6 mm, MBM 287027; D, E. Paratype, female, CW 3.2 mm, CL 2.2 mm, MBM 287026; A. Carapace; B. Maxilliped 3; C. Cheliped; D. Pereopod 5; E. Female pleon; F. Male pleon; G. right G 1, ventral view; H. Same, dorsal view; I. Right G 1, distal part, ventral view; J. Same, dorsal view; K. Right G 2, ventral view; L. Same, dorsal view. Scale bar: 1 mm (A, C, D); 0.5 mm (B, E, F, G, H); 0.1 mm (I L)." figureDoi="10.3897/zse.100.117859.figure3" httpUri="https://binary.pensoft.net/fig/1088096">3 A</figureCitation>
) (vs. the first tooth low but still visible in
<taxonomicName id="18265C0B8D2979000A2FCF4A24AA285C" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1873" class="Malacostraca" family="Xanthidae" genus="Actaeodes" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="consobrinus">
<emphasis id="068A0791CC45F031A01BB36B183A7640" italics="true">A. consobrinus</emphasis>
</taxonomicName>
; Figs
<figureCitation id="74AC23A0589D5A81CE75180EB2720F08" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Gothus consobrinus (A. Milne-Edwards, 1873), male, CW 6.7 mm, CL 4.4 mm, NS-YS- 2022-1336. A. Dorsal habitus; B. Carapace; C. Frontal view; D. Thoracic sternites and pleon; E. Chelipeds. Scale bar: 2 mm." figureDoi="10.3897/zse.100.117859.figure7" httpUri="https://binary.pensoft.net/fig/1088100">7 A, B</figureCitation>
,
<figureCitation id="E1A873C79BA69EDF47B6EF47812343DA" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Gothus consobrinus (A. Milne-Edwards, 1873), male, CW 6.7 mm, CL 4.4 mm, NS-YS- 2022-1336. A. Carapace; B. Maxilliped 3; C. Cheliped; D. Pereopod 5; E. Pleon; F. Left G 1, ventral view; G. Left G 1; H. Left G 2, ventral view. Scale bar: 2 mm (A); 0.5 mm (B, F, H); 1 mm (C, D, E); 0.1 mm (G)." figureDoi="10.3897/zse.100.117859.figure8" httpUri="https://binary.pensoft.net/fig/1088101">8 A</figureCitation>
,
<figureCitation id="8AAB150F82C965717F3829C4E9533A73" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Gothus consobrinus (A. Milne-Edwards, 1873). A C. Female, CW 10.5 mm, CL 6.8 mm, MNHN-IU- 2000-3885 (= MP-B 3885 S); D F. Female, CW 8.8 mm, CL 5.7 mm, MNHN-IU- 2024-3483; photographed by Sébastien Soubzmaigne; G I. Female, CW 4.5 mm, CL 2.9 mm, XS-QL- 2022-1014. Scale bar: 2 mm (A F); 1 mm (G I)." figureDoi="10.3897/zse.100.117859.figure9" httpUri="https://binary.pensoft.net/fig/1088102">9 A, B</figureCitation>
); carapace regions more pronounced, with tufts of setae (Figs
<figureCitation id="46AE2E57573FCF9B0DC6FF2E0682B3C2" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Gothus teemo sp. nov. A C. Holotype, male, CW 3.7 mm, CL 2.6 mm, MBM 287027; D F. Paratype, CW 3.2 mm, CL 2.2 mm, MBM 287026; A, D. Dorsal view; B, E. Frontal view; C, F. Carapace. Scale bar: 1 mm." figureDoi="10.3897/zse.100.117859.figure1" httpUri="https://binary.pensoft.net/fig/1088094">1 C</figureCitation>
,
<figureCitation id="B0D26AAA447EDAA78E7D3B9256BC0F91" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Gothus teemo sp. nov. A C, F, G L. Holotype, male, CW 3.7 mm, CL 2.6 mm, MBM 287027; D, E. Paratype, female, CW 3.2 mm, CL 2.2 mm, MBM 287026; A. Carapace; B. Maxilliped 3; C. Cheliped; D. Pereopod 5; E. Female pleon; F. Male pleon; G. right G 1, ventral view; H. Same, dorsal view; I. Right G 1, distal part, ventral view; J. Same, dorsal view; K. Right G 2, ventral view; L. Same, dorsal view. Scale bar: 1 mm (A, C, D); 0.5 mm (B, E, F, G, H); 0.1 mm (I L)." figureDoi="10.3897/zse.100.117859.figure3" httpUri="https://binary.pensoft.net/fig/1088096">3 A</figureCitation>
) (vs. carapace regions flatter, without tufts of setae in
<taxonomicName id="9EAAF7674AF0B2A60351A4B7E89739B9" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1873" class="Malacostraca" family="Xanthidae" genus="Actaeodes" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="consobrinus">
<emphasis id="FBDF5F2071524B5E78E45DC2AA1D0E1D" italics="true">A. consobrinus</emphasis>
</taxonomicName>
; Figs
<figureCitation id="BE60C507C4D40C62EDB5D099DBD1F7EC" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Gothus consobrinus (A. Milne-Edwards, 1873), male, CW 6.7 mm, CL 4.4 mm, NS-YS- 2022-1336. A. Dorsal habitus; B. Carapace; C. Frontal view; D. Thoracic sternites and pleon; E. Chelipeds. Scale bar: 2 mm." figureDoi="10.3897/zse.100.117859.figure7" httpUri="https://binary.pensoft.net/fig/1088100">7 A, B</figureCitation>
,
<figureCitation id="3DFD5F86DE04BA9B7C4899EEDE5DE1F0" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Gothus consobrinus (A. Milne-Edwards, 1873), male, CW 6.7 mm, CL 4.4 mm, NS-YS- 2022-1336. A. Carapace; B. Maxilliped 3; C. Cheliped; D. Pereopod 5; E. Pleon; F. Left G 1, ventral view; G. Left G 1; H. Left G 2, ventral view. Scale bar: 2 mm (A); 0.5 mm (B, F, H); 1 mm (C, D, E); 0.1 mm (G)." figureDoi="10.3897/zse.100.117859.figure8" httpUri="https://binary.pensoft.net/fig/1088101">8 A</figureCitation>
,
<figureCitation id="5F36AAF9381D64B459585A8961148704" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Gothus consobrinus (A. Milne-Edwards, 1873). A C. Female, CW 10.5 mm, CL 6.8 mm, MNHN-IU- 2000-3885 (= MP-B 3885 S); D F. Female, CW 8.8 mm, CL 5.7 mm, MNHN-IU- 2024-3483; photographed by Sébastien Soubzmaigne; G I. Female, CW 4.5 mm, CL 2.9 mm, XS-QL- 2022-1014. Scale bar: 2 mm (A F); 1 mm (G I)." figureDoi="10.3897/zse.100.117859.figure9" httpUri="https://binary.pensoft.net/fig/1088102">9 A, B</figureCitation>
); cheliped carpus with square-shaped outward expansion (Fig.
<figureCitation id="D06DDDD326825D148BED2A641B904A11" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Gothus teemo sp. nov. A C. Holotype, male, CW 3.7 mm, CL 2.6 mm, MBM 287027; D F. Paratype, CW 3.2 mm, CL 2.2 mm, MBM 287026; A, D. Dorsal view; B, E. Frontal view; C, F. Carapace. Scale bar: 1 mm." figureDoi="10.3897/zse.100.117859.figure1" httpUri="https://binary.pensoft.net/fig/1088094">1 A</figureCitation>
) (vs. cheliped carpus nearly spherical in
<taxonomicName id="186F19F46BFA182A0BC2B9890EEB00E4" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1873" class="Malacostraca" family="Xanthidae" genus="Actaeodes" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="consobrinus">
<emphasis id="5FCF629D1301C7D94D8BBF78AA5320E1" italics="true">A. consobrinus</emphasis>
</taxonomicName>
; Figs
<figureCitation id="FDBB65532B20F4A2C42628A6A5B0BB62" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Gothus consobrinus (A. Milne-Edwards, 1873), male, CW 6.7 mm, CL 4.4 mm, NS-YS- 2022-1336. A. Dorsal habitus; B. Carapace; C. Frontal view; D. Thoracic sternites and pleon; E. Chelipeds. Scale bar: 2 mm." figureDoi="10.3897/zse.100.117859.figure7" httpUri="https://binary.pensoft.net/fig/1088100">7 A</figureCitation>
,
<figureCitation id="665E6B9ACBAF35AE4F5A0A4A38342078" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Gothus consobrinus (A. Milne-Edwards, 1873), male, CW 6.7 mm, CL 4.4 mm, NS-YS- 2022-1336. A. Carapace; B. Maxilliped 3; C. Cheliped; D. Pereopod 5; E. Pleon; F. Left G 1, ventral view; G. Left G 1; H. Left G 2, ventral view. Scale bar: 2 mm (A); 0.5 mm (B, F, H); 1 mm (C, D, E); 0.1 mm (G)." figureDoi="10.3897/zse.100.117859.figure8" httpUri="https://binary.pensoft.net/fig/1088101">8 A, B</figureCitation>
); male pleon relatively broader, telson width slightly greater than length (Figs
<figureCitation id="844371CCC72052FD0074BCC6D9C38A64" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Gothus teemo sp. nov. A, C, E. Holotype, male, CW 3.7 mm, CL 2.6 mm, MBM 287027; B, D, F, G. Paratype, female, CW 3.2 mm, CL 2.2 mm, MBM 287026; A. Male chelipeds; B. Female chelipeds; C. Male thoracic sternites; D. Female thoracic sternites; E. Male pleon; F. Female pleon; G. Female vulva. Scale bar: 1 mm (A F); 0.2 mm (G)." figureDoi="10.3897/zse.100.117859.figure2" httpUri="https://binary.pensoft.net/fig/1088095">2 F</figureCitation>
,
<figureCitation id="DC5D9F182DA6B6DE413C51E2AC72221C" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Gothus teemo sp. nov. A C, F, G L. Holotype, male, CW 3.7 mm, CL 2.6 mm, MBM 287027; D, E. Paratype, female, CW 3.2 mm, CL 2.2 mm, MBM 287026; A. Carapace; B. Maxilliped 3; C. Cheliped; D. Pereopod 5; E. Female pleon; F. Male pleon; G. right G 1, ventral view; H. Same, dorsal view; I. Right G 1, distal part, ventral view; J. Same, dorsal view; K. Right G 2, ventral view; L. Same, dorsal view. Scale bar: 1 mm (A, C, D); 0.5 mm (B, E, F, G, H); 0.1 mm (I L)." figureDoi="10.3897/zse.100.117859.figure3" httpUri="https://binary.pensoft.net/fig/1088096">3 F</figureCitation>
) (vs. pleon very narrow, telson longer than wide in
<taxonomicName id="EEE7F723D7E5FC7F0069050F213980A4" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1873" class="Malacostraca" family="Xanthidae" genus="Actaeodes" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="consobrinus">
<emphasis id="D7AAB42CD16F73639BC5E3EBFC1841E5" italics="true">A. consobrinus</emphasis>
</taxonomicName>
; Figs
<figureCitation id="FA4B0FE0262CB29C32383510975AD18B" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Gothus consobrinus (A. Milne-Edwards, 1873), male, CW 6.7 mm, CL 4.4 mm, NS-YS- 2022-1336. A. Dorsal habitus; B. Carapace; C. Frontal view; D. Thoracic sternites and pleon; E. Chelipeds. Scale bar: 2 mm." figureDoi="10.3897/zse.100.117859.figure7" httpUri="https://binary.pensoft.net/fig/1088100">7 D</figureCitation>
,
<figureCitation id="511BB7CF8C9EE7D1E68526076E0F4281" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Gothus consobrinus (A. Milne-Edwards, 1873), male, CW 6.7 mm, CL 4.4 mm, NS-YS- 2022-1336. A. Carapace; B. Maxilliped 3; C. Cheliped; D. Pereopod 5; E. Pleon; F. Left G 1, ventral view; G. Left G 1; H. Left G 2, ventral view. Scale bar: 2 mm (A); 0.5 mm (B, F, H); 1 mm (C, D, E); 0.1 mm (G)." figureDoi="10.3897/zse.100.117859.figure8" httpUri="https://binary.pensoft.net/fig/1088101">8 E</figureCitation>
);
<abbrev id="ABBRID0E4NCI" xlink_title="first gonopod of male">G 1</abbrev>
distal lobe shorter (Fig.
<figureCitation id="134F2F3DAC632D459EEE8B6601C67593" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Gothus teemo sp. nov. A C, F, G L. Holotype, male, CW 3.7 mm, CL 2.6 mm, MBM 287027; D, E. Paratype, female, CW 3.2 mm, CL 2.2 mm, MBM 287026; A. Carapace; B. Maxilliped 3; C. Cheliped; D. Pereopod 5; E. Female pleon; F. Male pleon; G. right G 1, ventral view; H. Same, dorsal view; I. Right G 1, distal part, ventral view; J. Same, dorsal view; K. Right G 2, ventral view; L. Same, dorsal view. Scale bar: 1 mm (A, C, D); 0.5 mm (B, E, F, G, H); 0.1 mm (I L)." figureDoi="10.3897/zse.100.117859.figure3" httpUri="https://binary.pensoft.net/fig/1088096">3 G J</figureCitation>
) (vs.
<abbrev id="ABBRID0EFOCI" xlink_title="first gonopod of male">G 1</abbrev>
distal lobe significantly elongated in
<taxonomicName id="249EA131C67CC49A2BD9BE1AAF603BC3" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1873" class="Malacostraca" family="Xanthidae" genus="Actaeodes" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="consobrinus">
<emphasis id="B4B9E9599A4421BA2DE3D404AC32884D" italics="true">A. consobrinus</emphasis>
</taxonomicName>
; Fig.
<figureCitation id="9AADAF40D6E3DE8A26DAFF8719849DB0" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Gothus consobrinus (A. Milne-Edwards, 1873), male, CW 6.7 mm, CL 4.4 mm, NS-YS- 2022-1336. A. Carapace; B. Maxilliped 3; C. Cheliped; D. Pereopod 5; E. Pleon; F. Left G 1, ventral view; G. Left G 1; H. Left G 2, ventral view. Scale bar: 2 mm (A); 0.5 mm (B, F, H); 1 mm (C, D, E); 0.1 mm (G)." figureDoi="10.3897/zse.100.117859.figure8" httpUri="https://binary.pensoft.net/fig/1088101">8 F G</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>