treatments-xml/data/79/B0/F8/79B0F851A71455EAB277F5A36D237D1B.xml

<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.67.6703" ID-PMC="PMC4972245" ID-Pensoft-Pub="1314-2003-67-1" ID-Pensoft-UUID="D846EB3F7746FFFE4A469751FFEF3B22" ID-PubMed="27536193" ID-Zenodo-Dep="133270" ModsDocID="1314-2003-67-1" checkinTime="1469054247116" checkinUser="pensoft" docAuthor="Kriebel, Ricardo" docDate="2016" docId="79B0F851A71455EAB277F5A36D237D1B" docLanguage="en" docName="PhytoKeys 67: 1-326" docOrigin="PhytoKeys 67" docPubDate="2016-07-20" docSource="http://dx.doi.org/10.3897/phytokeys.67.6703" docTitle="Conostegia papillopetala Kriebel 2016, comb. nov." docType="treatment" docVersion="3" id="D846EB3F7746FFFE4A469751FFEF3B22" lastPageNumber="205" masterDocId="D846EB3F7746FFFE4A469751FFEF3B22" masterDocTitle="A Monograph of Conostegia (Melastomataceae, Miconieae)" masterLastPageNumber="326" masterPageNumber="1" pageNumber="202" updateTime="1668141747722" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>A Monograph of Conostegia (Melastomataceae, Miconieae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Kriebel, Ricardo</mods:namePart>
<mods:affiliation>Department of Botany, University of Wisconsin-Madison, 430 Lincoln Drive Madison, Wisconsin 53706, USA</mods:affiliation>
<mods:nameIdentifier type="email">kriebelr@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>PhytoKeys</mods:title>
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<mods:part>
<mods:date>2016</mods:date>
<mods:detail type="pubDate">
<mods:number>2016-07-20</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>67</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>326</mods:end>
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<mods:url>http://dx.doi.org/10.3897/phytokeys.67.6703</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/phytokeys.67.6703</mods:identifier>
<mods:identifier type="Pensoft-Pub">1314-2003-67-1</mods:identifier>
<mods:identifier type="Pensoft-UUID">D846EB3F7746FFFE4A469751FFEF3B22</mods:identifier>
<mods:identifier type="Zenodo-Dep">133270</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="182226968" LSID="urn:lsid:plazi:treatment:79B0F851A71455EAB277F5A36D237D1B" httpUri="http://treatment.plazi.org/id/79B0F851A71455EAB277F5A36D237D1B" lastPageId="204" lastPageNumber="205" pageId="201" pageNumber="202">
<subSubSection pageId="201" pageNumber="202" type="nomenclature">
<paragraph pageId="201" pageNumber="202">
<taxonomicName LSID="79B0F851-A714-55EA-B277-F5A36D237D1B" authority="(Kriebel &amp; Almeda) Kriebel" authorityName="Kriebel" authorityYear="2016" baseAuthorityName="Kriebel &amp; Almeda" baseAuthorityYear="2016" class="Magnoliopsida" family="Melastomataceae" genus="Conostegia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Conostegia papillopetala" order="Myrtales" pageId="201" pageNumber="202" phylum="Tracheophyta" rank="species" species="papillopetala" status="comb. nov.">
<pageBreakToken pageId="201" pageNumber="202" start="start">Conostegia</pageBreakToken>
papillopetala (Kriebel &amp; Almeda) Kriebel
</taxonomicName>
<taxonomicNameLabel pageId="201" pageNumber="202">comb. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 180" captionStartId="F180" captionText="Figure 180. Conostegia papillopetala. A Stem apex B Branch C Inflorescence D Infructescence E Flower bud with closed hyaline calyx and apiculum F Pickled flower G Longitudinal cut of flower with style removed H Petal I Stamen. Photos of specimen vouchered R. Kriebel and J. Burke 5718." figureDoi="10.3897/phytokeys.67.6703.figure180" httpUri="https://binary.pensoft.net/fig/96538" pageId="201" pageNumber="202">Fig. 180</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="202" pageNumber="203" type="reference_group">
<paragraph pageId="202" pageNumber="203">
<taxonomicName authorityName="Kriebel" authorityYear="2016" baseAuthorityName="Kriebel &amp; Almeda" baseAuthorityYear="2016" class="Magnoliopsida" family="Melastomataceae" genus="Conostegia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Conostegia papillopetala" order="Myrtales" pageId="202" pageNumber="203" phylum="Tracheophyta" rank="species" species="papillopetala" status="comb. nov.">
<pageBreakToken pageId="202" pageNumber="203" start="start">Conostegia</pageBreakToken>
papillopetala
</taxonomicName>
(Kriebel &amp; Almeda) Kriebel. Basionym: 
<taxonomicName authorityName="Kriebel" authorityYear="2016" class="Magnoliopsida" family="Melastomataceae" genus="Miconia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Miconia papillopetala" order="Myrtales" pageId="202" pageNumber="203" phylum="Tracheophyta" rank="species" species="papillopetala" status="comb. nov.">Miconia papillopetala</taxonomicName>
, Phytotaxa 134 (1): 32. 2013. Type: Panama. Veraguas: Parque Nacional Santa 
<normalizedToken originalValue="Fé">Fe</normalizedToken>
. Sendero a la cima del Cerro Mariposa, 960 m, N 08.50412, W 081.11999, 16 September 2011, R. Kriebel &amp; J. Burke 5718 (holotype: NY!, isotypes: INB!, PMA!).
</paragraph>
</subSubSection>
<subSubSection lastPageId="204" lastPageNumber="205" pageId="202" pageNumber="203" type="description">
<paragraph pageId="202" pageNumber="203">Description.</paragraph>
<paragraph lastPageId="204" lastPageNumber="205" pageId="202" pageNumber="203">
Shrubs 1-2.5 m tall with young stems copiously covered with pinoid hairs intermixed with asperous-headed hairs that are both golden-orange in color, nodal line not evident and concealed by the copious indument. Petioles 0.1-1 cm. 
<pageBreakToken pageId="203" pageNumber="204" start="start">Leaves</pageBreakToken>
subisophyllous to anisophyllous; blades 3.5-16 
<normalizedToken originalValue="×">x</normalizedToken>
1.5-8.5 cm, 3-5-plinerved, diverging from the midvein 0.5-3 cm above the blade base usually asymmetrically, elliptic, base obtuse to rounded and sometimes oblique, apex acuminate, the mar 
<pageBreakToken pageId="204" pageNumber="205" start="start">gin</pageBreakToken>
denticulate, adaxially glabrous except for short and long pinoid hairs on the main veins towards the base, somewhat bullate and dark green when alive, abaxially densely pubescent on tertiary and higher order veins with pinoid golden-orange hairs. Inflorescences terminal, lax dichasia branched from the base, (4-)7-10 cm long, copiously covered with golden-orange pinoid hairs intermixed with asperous-headed hairs; bracts to 8 mm long, linear oblong; bracteoles 0.5-1 mm long, lanceolate, less pubescent than the rest of inflorescence rachis, drying pinkish. Pedicels ca. 0.5 mm. Hypanthia campanulate 1.25-2 
<normalizedToken originalValue="×">x</normalizedToken>
1-1.25 mm, densely covered with asperous-headed that appear somewhat stellate. Flowers 5-merous. Calyx fused in bud, shortly apiculate and less pubescent than the hypanthium, rupturing at anthesis into irregular, broadly rounded hyaline lobes 0.25-0.75 mm long and 0.5-0.75 mm wide at the base, the exterior calyx teeth 0.25-0.5 mm long, linear oblong, the calyx tube 0.25-0.5 mm long. Petals 1.25-2 
<normalizedToken originalValue="×">x</normalizedToken>
2.5-3 mm, ovate, pink, papillose abaxially, reflexed at anthesis, emarginate. Stamens 10, 3-3.5 mm long, radially arranged around the style; filaments 1.5-2 mm long, geniculate near the apex, translucent white; anthers 1.25-1.75 
<normalizedToken originalValue="×">x</normalizedToken>
0.4-0.6 mm, linear-oblong, somewhat laterally compressed, cream yellow, pores 0.1-0.15 mm, truncate to somewhat ventrally inclined. Ovaries 5-locular, half inferior, the apex elevated into a low papillose collar. Styles 4.5-4.75 mm long, slightly curved, distance between the anther apex and the stigma 1-1.5 mm; stigmas truncate to capitellate, 0.4 mm wide. Berries pink when immature and turning purple at maturity, 3.3-4.5 
<normalizedToken originalValue="×">x</normalizedToken>
3.5-4.5 mm; seeds ovoid and angled, 0.4-0.5 
<normalizedToken originalValue="×">x</normalizedToken>
0.2-0.3 mm, orange-brown, lateral symmetrical plane ovate to triangular, the highest point toward the chalazal side, antiraphal symmetrical plane ovate-triangular and inconspicuously verruculose on the angles, raphal zone narrowly triangular and extending the length of the seed, expanded into an appendage that covers about 30% of the seed length.
</paragraph>
<caption ID-Zenodo-Dep="974017" doi="10.3897/phytokeys.67.6703.figure180" httpUri="https://binary.pensoft.net/fig/96538" pageId="204" pageNumber="205" start="Figure 180" startId="F180">
<paragraph pageId="204" pageNumber="205">
<emphasis bold="true" pageId="204" pageNumber="205">Figure 180.</emphasis>
<taxonomicName authorityName="Kriebel" authorityYear="2016" baseAuthorityName="Kriebel &amp; Almeda" baseAuthorityYear="2016" class="Magnoliopsida" family="Melastomataceae" genus="Conostegia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Conostegia papillopetala" order="Myrtales" pageId="204" pageNumber="205" phylum="Tracheophyta" rank="species" species="papillopetala">
<emphasis italics="true" pageId="204" pageNumber="205">Conostegia papillopetala</emphasis>
</taxonomicName>
. 
<emphasis bold="true" pageId="204" pageNumber="205">A</emphasis>
Stem apex 
<emphasis bold="true" pageId="204" pageNumber="205">B</emphasis>
Branch 
<emphasis bold="true" pageId="204" pageNumber="205">C</emphasis>
Inflorescence 
<emphasis bold="true" pageId="204" pageNumber="205">D</emphasis>
Infructescence 
<emphasis bold="true" pageId="204" pageNumber="205">E</emphasis>
Flower bud with closed hyaline calyx and apiculum 
<emphasis bold="true" pageId="204" pageNumber="205">F</emphasis>
Pickled flower 
<emphasis bold="true" pageId="204" pageNumber="205">G</emphasis>
Longitudinal cut of flower with style removed 
<emphasis bold="true" pageId="204" pageNumber="205">H</emphasis>
Petal 
<emphasis bold="true" pageId="204" pageNumber="205">I</emphasis>
Stamen. Photos of specimen vouchered 
<emphasis italics="true" pageId="204" pageNumber="205">R. Kriebel and J. Burke 5718</emphasis>
.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="204" pageNumber="205" type="distribution">
<paragraph pageId="204" pageNumber="205">Distribution</paragraph>
<paragraph pageId="204" pageNumber="205">
(Fig. 
<figureCitation captionStart="Figure 181" captionStartId="F181" captionText="Figure 181. Distribution of Conostegia papillopetala." figureDoi="10.3897/phytokeys.67.6703.figure181" httpUri="https://binary.pensoft.net/fig/96539" pageId="204" pageNumber="205">181</figureCitation>
). Endemic to cloud forests in the provinces of 
<normalizedToken originalValue="Coclé">Cocle</normalizedToken>
and Veraguas, 
<normalizedToken originalValue="Panamá">Panama</normalizedToken>
, at 750-1400 m elevation.
</paragraph>
<caption ID-Zenodo-Dep="974019" doi="10.3897/phytokeys.67.6703.figure181" httpUri="https://binary.pensoft.net/fig/96539" pageId="204" pageNumber="205" start="Figure 181" startId="F181">
<paragraph pageId="204" pageNumber="205">
<emphasis bold="true" pageId="204" pageNumber="205">Figure 181.</emphasis>
Distribution of 
<taxonomicName authorityName="Kriebel" authorityYear="2016" baseAuthorityName="Kriebel &amp; Almeda" baseAuthorityYear="2016" class="Magnoliopsida" family="Melastomataceae" genus="Conostegia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Conostegia papillopetala" order="Myrtales" pageId="204" pageNumber="205" phylum="Tracheophyta" rank="species" species="papillopetala">
<emphasis italics="true" pageId="204" pageNumber="205">Conostegia papillopetala</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph pageId="204" pageNumber="205">
<taxonomicName authorityName="Kriebel" authorityYear="2016" baseAuthorityName="Kriebel &amp; Almeda" baseAuthorityYear="2016" class="Magnoliopsida" family="Melastomataceae" genus="Conostegia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Conostegia papillopetala" order="Myrtales" pageId="204" pageNumber="205" phylum="Tracheophyta" rank="species" species="papillopetala">
<emphasis italics="true" pageId="204" pageNumber="205">Conostegia papillopetala</emphasis>
</taxonomicName>
can be recognized by its indument of long asperous headed hairs, short petiolate, somewhat bullate, narrow and plinerved leaves, small, herkogamous flowers with pink papillose petals and seeds with tubercles on the edges. It is similar and closely related to 
<taxonomicName authorityName="Kriebel" authorityYear="2016" baseAuthorityName="Almeda &amp; Umana" baseAuthorityYear="2016" class="Magnoliopsida" family="Melastomataceae" genus="Conostegia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Conostegia friedmaniorum" order="Myrtales" pageId="204" pageNumber="205" phylum="Tracheophyta" rank="species" species="friedmaniorum">
<emphasis italics="true" pageId="204" pageNumber="205">Conostegia friedmaniorum</emphasis>
</taxonomicName>
, 
<taxonomicName authorityName="Kriebel" authorityYear="2016" baseAuthorityName="Kriebel &amp; Almeda" baseAuthorityYear="2016" class="Magnoliopsida" family="Melastomataceae" genus="Conostegia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Conostegia galdamesiae" order="Myrtales" pageId="204" pageNumber="205" phylum="Tracheophyta" rank="species" species="galdamesiae">
<emphasis italics="true" pageId="204" pageNumber="205">Conostegia galdamesiae</emphasis>
</taxonomicName>
, and 
<taxonomicName authorityName="Kriebel" authorityYear="2016" baseAuthorityName="Umana &amp; Almeda" baseAuthorityYear="2016" class="Magnoliopsida" family="Melastomataceae" genus="Conostegia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Conostegia pendula" order="Myrtales" pageId="204" pageNumber="205" phylum="Tracheophyta" rank="species" species="pendula">
<emphasis italics="true" pageId="204" pageNumber="205">Conostegia pendula</emphasis>
</taxonomicName>
. See the discussion section under these species for difference with 
<taxonomicName authorityName="Kriebel" authorityYear="2016" baseAuthorityName="Kriebel &amp; Almeda" baseAuthorityYear="2016" class="Magnoliopsida" family="Melastomataceae" genus="Conostegia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Conostegia papillopetala" order="Myrtales" pageId="204" pageNumber="205" phylum="Tracheophyta" rank="species" species="papillopetala">
<emphasis italics="true" pageId="204" pageNumber="205">Conostegia papillopetala</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection pageId="204" pageNumber="205" type="materials_examined">
<paragraph pageId="204" pageNumber="205">Specimens examined.</paragraph>
<paragraph pageId="204" pageNumber="205">
<materialsCitation collectingDate="2016-01-01" collectingDateMax="2016-12-31" collectingDateMin="2016-01-01" collectorName="Folsom, Near, Hammel" country="PANAMA" county="Atlantic" elevation="900" location="Croat" municipality="Atlantic" specimenCount="1" stateProvince="Cocle">
<emphasis bold="true" pageId="204" pageNumber="205">
<collectingCountry name="Panama">PANAMA</collectingCountry>
. 
<normalizedToken originalValue="Coclé">
<collectingRegion country="Panama" name="Cocle">Cocle</collectingRegion>
</normalizedToken>
</emphasis>
: 
<collectingCounty>Atlantic</collectingCounty>
slope near the continental divide along lumbering road 
<locationDeviation location="El Cope">
N of El 
<normalizedToken originalValue="Copé">Cope</normalizedToken>
</locationDeviation>
, 
<locationDeviation location="El Cope">
<quantity metricMagnitude="3" metricUnit="m" metricValue="9.4" unit="km" value="9.4">9.4 km</quantity>
above El 
<normalizedToken originalValue="Copé">Cope</normalizedToken>
</locationDeviation>
, Croat 44624 (CAS, PMA); 
<collectingMunicipality>Atlantic</collectingMunicipality>
slope near the continental divide along lumbering road 
<locationDeviation location="El Cope">
N of El 
<normalizedToken originalValue="Copé">Cope</normalizedToken>
</locationDeviation>
, 
<quantity metricMagnitude="3" metricUnit="m" metricValue="2.2" unit="km" value="2.2">2.2 km</quantity>
N of lumber sawmill, 
<location LSID="urn:lsid:plazi:treatment:79B0F851A71455EAB277F5A36D237D1B:9BC6A7AB7BBF2F529B8894C5EC2426D6" country="PANAMA" county="Atlantic" municipality="Atlantic" name="Croat" stateProvince="Cocle">Croat</location>
44668 (CAS, MO, PMA); 
<location LSID="urn:lsid:plazi:treatment:79B0F851A71455EAB277F5A36D237D1B:B06538F34A3DCC55044617DC9EA8367A" country="PANAMA" county="Atlantic" municipality="Atlantic" name="Lumber" stateProvince="Cocle">Lumber</location>
camp at 
<location LSID="urn:lsid:plazi:treatment:79B0F851A71455EAB277F5A36D237D1B:C6635640C06BDEC5079F01EC28A2A882" country="PANAMA" county="Atlantic" municipality="Atlantic" name="Alto Calvario" stateProvince="Cocle">Alto Calvario</location>
, 
<locationDeviation location="El Cope">
<quantity metricMagnitude="3" metricUnit="m" metricValue="7.0" unit="km" value="7.0">7 km</quantity>
N of El 
<normalizedToken originalValue="Copé">Cope</normalizedToken>
</locationDeviation>
, 
<quantity metricMagnitude="2" metricUnit="m" metricValue="9.0" unit="m" value="900.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="9.0" unit="m" value="900.0">900 m</elevation>
</quantity>
, 
<collectorName>Folsom</collectorName>
1276 (CAS, INB, MEXU, MO, PMA); 
<collectorName>Near</collectorName>
continental divide along lumbering road, 
<quantity metricMagnitude="3" metricUnit="m" metricValue="2.3" unit="km" value="2.3">2.3 km</quantity>
beyond sawmill above 
<location LSID="urn:lsid:plazi:treatment:79B0F851A71455EAB277F5A36D237D1B:48B23A9DF6C7CFCE05ACED370CF5A4A9" country="PANAMA" county="Atlantic" municipality="Atlantic" name="El Cope" stateProvince="Cocle">
El 
<normalizedToken originalValue="Copé">Cope</normalizedToken>
</location>
, 
<collectorName>Hammel</collectorName>
990, 1054 (CAS, MO, PMA)
</materialsCitation>
. 
<materialsCitation collectingDate="2016-01-01" collectingDateMax="2016-12-31" collectingDateMin="2016-01-01" collectorName="de Tute, de Piedra" country="Panama" county="Cordillera de Tute" location="Altos de Piedra" municipality="Escuela Agricola Altos de Piedra" specimenCount="1" stateProvince="Veraguas">
<emphasis bold="true" pageId="204" pageNumber="205">
<collectingRegion country="Panama" name="Veraguas">Veraguas</collectingRegion>
</emphasis>
: Trail on ridge summit of Cerro Tute, 
<collectingCounty>
Cordillera 
<collectorName>de Tute</collectorName>
</collectingCounty>
, 
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.0" unit="km" value="1.0">1 km</quantity>
past 
<collectingMunicipality>Escuela Agricola Altos de Piedra</collectingMunicipality>
, W of Santa Fe, Knapp and Sytsma 2654 (BM, CAS, MO, PMA); Santa Fe, 
<location LSID="urn:lsid:plazi:treatment:79B0F851A71455EAB277F5A36D237D1B:EF7146D1358735B69E381BEB573002B0" country="Panama" county="Cordillera de Tute" municipality="Escuela Agricola Altos de Piedra" name="Altos de Piedra" stateProvince="Veraguas">
Altos 
<collectorName>de Piedra</collectorName>
</location>
, trail leading up Cerro Mariposa (=Cerro Arizona)
</materialsCitation>
about 
<quantity metricMagnitude="3" metricUnit="m" metricValue="2.0" unit="km" value="2.0">2 km</quantity>
past the Escuela Agricola, ca. 
<quantity metricMagnitude="3" metricUnit="m" metricValue="3.0" unit="km" value="3.0">3 km</quantity>
from summit, Penneys and Blanco 1707 (CAS, FLAS, NY, PMA, 
<collectingCountry name="United States of America">US</collectingCountry>
).
</paragraph>
</subSubSection>
</treatment>
</document>