treatments-xml/data/6C/7F/0C/6C7F0C4EFFC25C6E38DE461CC4E9FE51.xml

<document id="6B22DB68DC978CE53DC99645E9152045" ID-CLB-Dataset="31990" ID-DOI="10.11646/zootaxa.4324.1.3" ID-GBIF-Dataset="cb90f22f-94a2-4606-9c3e-0b497389dae7" ID-ISSN="1175-5326" ID-Zenodo-Dep="997051" ID-ZooBank="77728C1D-430E-4B84-98Be-462E2A1F3Aaf" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1506406729148" checkinUser="plazi" docAuthor="Marin, Ivan" docDate="2017" docId="6C7F0C4EFFC25C6E38DE461CC4E9FE51" docLanguage="en" docName="zootaxa.4324.1.3.pdf" docOrigin="Zootaxa 4324 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Athanas alpheusophilus Marin 2017" docType="treatment" docVersion="8" masterDocId="90467436FFC35C6438494423C170FFBC" masterDocTitle="Athanas alpheusophilus sp. nov. (Decapoda: Alpheidae) - a new Alpheus - associated shrimp from the Russian coast of the Sea of Japan" updateTime="1698468981414" updateUser="ExternalLinkService">
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<mods:title id="16B6A8535F243A37E2B22CBEDFAA937B">Athanas alpheusophilus sp. nov. (Decapoda: Alpheidae) - a new Alpheus - associated shrimp from the Russian coast of the Sea of Japan</mods:title>
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<mods:namePart id="205DE87621D55AA69CC15F8F3BE355DB">Marin, Ivan</mods:namePart>
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<mods:part id="4DA59B7D624913BC9D1F0EA69D0C48A9">
<mods:date id="29DB3132D33450B1D8D08B4684FA796D">2017</mods:date>
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<mods:number id="90D6196202B94AC911ACBF5D02869721">2017-09-26</mods:number>
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<mods:identifier id="762CF3843C480A6899E0271E87482D6C" type="DOI">10.11646/zootaxa.4324.1.3</mods:identifier>
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<treatment id="6C7F0C4EFFC25C6E38DE461CC4E9FE51" ID-DOI="http://doi.org/10.5281/zenodo.6046834" ID-GBIF-Taxon="134840113" ID-Zenodo-Dep="6046834" LSID="urn:lsid:plazi:treatment:6C7F0C4EFFC25C6E38DE461CC4E9FE51" httpUri="http://treatment.plazi.org/id/6C7F0C4EFFC25C6E38DE461CC4E9FE51" lastPageId="10" pageId="1">
<subSubSection id="ACCCEED3FFC25C6538DE461CC062FDC7" pageId="1" type="nomenclature">
<paragraph id="E469BD58FFC25C6538DE461CC350FDE6" blockId="1.[151,544,575,635]" box="[151,544,575,602]" pageId="1">
<heading id="BF210A34FFC25C6538DE461CC350FDE6" bold="true" box="[151,544,575,602]" fontSize="11" level="1" pageId="1" reason="1">
<emphasis id="D6A2614AFFC25C6538DE461CC350FDE6" bold="true" box="[151,544,575,602]" pageId="1">
<taxonomicName id="23D6C6DBFFC25C6538DE461CC0CBFDE5" ID-CoL="85MBK" authority="Marin, 2017" authorityName="Marin" authorityYear="2017" box="[151,443,575,601]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="1" phylum="Arthropoda" rank="species" species="alpheusophilus" status="sp. nov.">
<emphasis id="D6A2614AFFC25C6538DE461CC0CBFDE5" bold="true" box="[151,443,575,601]" italics="true" pageId="1">Athanas alpheusophilus</emphasis>
</taxonomicName>
sp. nov.
</emphasis>
</heading>
</paragraph>
<paragraph id="E469BD58FFC25C6538DE4641C062FDC7" blockId="1.[151,544,575,635]" box="[151,274,610,635]" pageId="1">
(
<figureCitation id="7CEDA1DDFFC25C6538D64641C079FDC7" box="[159,265,610,635]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStart-5="FIGURE 6" captionStartId-0="1.[151,250,1956,1978]" captionStartId-1="3.[151,250,1954,1976]" captionStartId-2="4.[151,250,1952,1974]" captionStartId-3="5.[151,250,1983,2005]" captionStartId-4="6.[151,250,1931,1953]" captionStartId-5="7.[151,250,1924,1946]" captionTargetBox-0="[169,1422,953,1927]" captionTargetBox-1="[166,1411,202,1909]" captionTargetBox-2="[168,1430,207,1912]" captionTargetBox-3="[169,1414,218,1948]" captionTargetBox-4="[208,1378,220,1888]" captionTargetBox-5="[151,1436,193,1902]" captionTargetId-0="figure@1.[151,1436,937,1932]" captionTargetId-1="figure@3.[159,1419,193,1928]" captionTargetId-2="figure@4.[151,1436,193,1931]" captionTargetId-3="figure@5.[161,1425,193,1962]" captionTargetId-4="figure@6.[195,1391,211,1899]" captionTargetId-5="figure@7.[151,1436,193,1902]" captionTargetPageId-0="1" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="5" captionTargetPageId-4="6" captionTargetPageId-5="7" captionText-0="FIGURE 1. Athanas alpheusophillus sp. nov., Vitjaz, Posjeta Bay, the Sea of Japan, general view of paratype female (a) and holotype male (ZMMU Ma 5839) (b)." captionText-1="FIGURE 2. Athanas alpheusophillus sp. nov., Vitjaz, Posjeta Bay, the Sea of Japan, paratypes (a, d, e – g — ovigerous female; b, c — male): a, b — front of carapace, dorsal view; c, d — front of carapace, lateral view; e — distal abdominal segments and telson, lateral view; f — telson; g — exopod of uropods." captionText-2="FIGURE 3. Athanas alpheusophillus sp. nov., Vitjaz, Posjeta Bay, the Sea of Japan, paratypes (a, c, d — ovigerous female; b, e — male): a, b — antennula; c, e — pereiopods I (chelipeds); d — distal propodus and chela of pereiopod I." captionText-3="FIGURE 4. Athanas alpheusophillus sp. nov., Vitjaz, Posjeta Bay, the Sea of Japan, paratypes (a – c — male; d, e — female): a, b — minor pereiopod I; c — distal propodus and fingers of pereiopod I; d — pereiopod II; e — pereiopod III." captionText-4="FIGURE 5. Athanas alpheusophillus sp. nov., Vitjaz, Posjeta Bay, the Sea of Japan, paratype male, major pereiopod I: a — general view of pereiopod I; b — distal segments of pereiopod I; c, d — distal propodus and fingers of pereiopod I, different views." captionText-5="FIGURE 6. Athanas alpheusophillus sp. nov., Vitjaz, Posjeta Bay, the Sea of Japan, alive coloration of female paratype (a, b) and male holotype (ZMMU Ma 5839) (c)." httpUri-0="https://zenodo.org/record/997053/files/figure.png" httpUri-1="https://zenodo.org/record/997055/files/figure.png" httpUri-2="https://zenodo.org/record/997057/files/figure.png" httpUri-3="https://zenodo.org/record/997059/files/figure.png" httpUri-4="https://zenodo.org/record/997061/files/figure.png" httpUri-5="https://zenodo.org/record/997063/files/figure.png" pageId="1">Figs. 1–6</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="ACCCEED3FFC25C6538DE4689C25CFD7C" box="[151,812,682,704]" pageId="1" type="description">
<paragraph id="E469BD58FFC25C6538DE4689C25CFD7C" blockId="1.[151,812,682,704]" box="[151,812,682,704]" pageId="1">
<taxonomicName id="23D6C6DBFFC25C6538DE4689C0A0FD7C" authority="Kubo, 1936" authorityName="Kubo" authorityYear="1936" box="[151,464,682,704]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="1" phylum="Arthropoda" rank="species" species="japonicus">
<emphasis id="D6A2614AFFC25C6538DE4689C023FD7C" box="[151,339,682,704]" italics="true" pageId="1">Athanas japonicus</emphasis>
Kubo, 1936
</taxonomicName>
.—Marin, 2013: 61, Fig. 24–3a, b.
</paragraph>
</subSubSection>
<subSubSection id="ACCCEED3FFC25C6538DE46CEC18DFCCE" pageId="1" type="materials_examined">
<paragraph id="E469BD58FFC25C6538DE46CEC18DFCCE" blockId="1.[151,1436,749,882]" pageId="1">
<emphasis id="D6A2614AFFC25C6538DE46CEC00BFCBA" bold="true" box="[151,379,749,774]" pageId="1">Material examined.</emphasis>
Holotype: male (cl 
<quantity id="232E10BDFFC25C653A1246CEC3C2FCB9" box="[603,690,749,774]" metricMagnitude="-3" metricUnit="m" metricValue="5.6" pageId="1" unit="mm" value="5.6">5.6 mm</quantity>
, tl 11.0 mm), ZMMU Ma 5839, Russia, Sea of Japan, Posjeta Bay, Vitjaz, 42°36′7″N 131°10′43″E, depth 
<quantity id="232E10BDFFC25C653A0F4731C3E0FC95" box="[582,656,786,810]" metricMagnitude="0" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="1" unit="m" value="1.5" valueMax="2.0" valueMin="1.0">1–2 m</quantity>
, 
<taxonomicName id="23D6C6DBFFC25C653AD44731C380FC95" authorityName="Linnaeus" authorityYear="1753" box="[669,752,786,809]" class="Liliopsida" family="Zosteraceae" genus="Zostera" kingdom="Plantae" order="Alismatales" pageId="1" phylum="Tracheophyta" rank="genus">
<emphasis id="D6A2614AFFC25C653AD44731C380FC95" box="[669,752,786,809]" italics="true" pageId="1">Zostera</emphasis>
</taxonomicName>
beds, yabby pump, from burrows of 
<taxonomicName id="23D6C6DBFFC25C653CDE4732C4E5FC95" box="[1175,1429,785,809]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="1" phylum="Arthropoda" rank="species" species="breviscristatus">
<emphasis id="D6A2614AFFC25C653CDE4732C4E5FC95" box="[1175,1429,785,809]" italics="true" pageId="1">Alpheus breviscristatus</emphasis>
</taxonomicName>
, coll. I. Marin, 
<date id="90689B98FFC25C6539774715C0AAFCF2" box="[318,474,821,846]" pageId="1" value="2015-06-14">14 June 2015</date>
. Paratypes: 12 ovigerous females and 7 males), LEMMI, same collection data as for holotype.
</paragraph>
</subSubSection>
<caption id="B0A9EDD0FFC25C6538DE4387C368F865" httpUri="https://zenodo.org/record/997053/files/figure.png" pageId="1" targetBox="[169,1422,953,1927]" targetPageId="1">
<paragraph id="E469BD58FFC25C6538DE4387C368F865" blockId="1.[151,1435,1956,2009]" pageId="1">
<emphasis id="D6A2614AFFC25C6538DE4387C305F806" bold="true" box="[151,629,1956,1979]" pageId="1">
FIGURE 1. 
<taxonomicName id="23D6C6DBFFC25C6539534387C36DF807" authority="Marin, 2017" authorityName="Marin" authorityYear="2017" box="[282,541,1956,1979]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="1" phylum="Arthropoda" rank="species" species="alpheusophillus" status="sp. nov.">
<emphasis id="D6A2614AFFC25C6539534387C36DF807" bold="true" box="[282,541,1956,1979]" italics="true" pageId="1">Athanas alpheusophillus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CD91DC31FFC25C653A6D4386C305F806" box="[548,629,1957,1978]" pageId="1" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, Vitjaz, Posjeta Bay, the Sea of Japan, general view of paratype female (
<emphasis id="D6A2614AFFC25C653D154386C418F806" box="[1372,1384,1957,1978]" italics="true" pageId="1">a</emphasis>
) and holotype male (ZMMU Ma 5839) (
<emphasis id="D6A2614AFFC25C6539B543E0C379F865" box="[508,521,1987,2009]" italics="true" pageId="1">b</emphasis>
).
</paragraph>
</caption>
<subSubSection id="ACCCEED3FFC15C6C388E44B4C36AFE60" lastPageId="8" pageId="2" type="description">
<paragraph id="E469BD58FFC15C66388E44B4C098FEEB" blockId="2.[151,1437,151,2010]" pageId="2">
<emphasis id="D6A2614AFFC15C66388E44B4C029FF0C" bold="true" box="[199,345,151,176]" pageId="2">Description.</emphasis>
Small-sized shrimp with moderately slender, subcylindrical body (
<figureCitation id="7CEDA1DDFFC15C663C2544B4C5C2FF0C" box="[1132,1202,151,176]" captionStart="FIGURE 1" captionStartId="1.[151,250,1956,1978]" captionTargetBox="[169,1422,953,1927]" captionTargetId="figure@1.[151,1436,937,1932]" captionTargetPageId="1" captionText="FIGURE 1. Athanas alpheusophillus sp. nov., Vitjaz, Posjeta Bay, the Sea of Japan, general view of paratype female (a) and holotype male (ZMMU Ma 5839) (b)." httpUri="https://zenodo.org/record/997053/files/figure.png" pageId="2">Fig. 1</figureCitation>
). Carapace smooth, sparsely covered with simple erect setae (
<figureCitation id="7CEDA1DDFFC15C663AC8449AC3B7FF6D" box="[641,711,185,210]" captionStart="FIGURE 1" captionStartId="1.[151,250,1956,1978]" captionTargetBox="[169,1422,953,1927]" captionTargetId="figure@1.[151,1436,937,1932]" captionTargetPageId="1" captionText="FIGURE 1. Athanas alpheusophillus sp. nov., Vitjaz, Posjeta Bay, the Sea of Japan, general view of paratype female (a) and holotype male (ZMMU Ma 5839) (b)." httpUri="https://zenodo.org/record/997053/files/figure.png" pageId="2">Fig. 1</figureCitation>
). Rostrum long and slender, almost reaching distal margin of second antennular article (
<figureCitation id="7CEDA1DDFFC15C66398044F9C34DFF4F" box="[457,573,218,243]" captionStart="FIGURE 2" captionStartId="3.[151,250,1954,1976]" captionTargetBox="[166,1411,202,1909]" captionTargetId="figure@3.[159,1419,193,1928]" captionTargetPageId="3" captionText="FIGURE 2. Athanas alpheusophillus sp. nov., Vitjaz, Posjeta Bay, the Sea of Japan, paratypes (a, d, e – g — ovigerous female; b, c — male): a, b — front of carapace, dorsal view; c, d — front of carapace, lateral view; e — distal abdominal segments and telson, lateral view; f — telson; g — exopod of uropods." httpUri="https://zenodo.org/record/997055/files/figure.png" pageId="2">
Fig. 2 
<emphasis id="D6A2614AFFC15C663A4644F8C34DFF4F" box="[527,573,219,243]" italics="true" pageId="2">a, b</emphasis>
</figureCitation>
), distally acute, without dorsal or ventral ridges, with well-marked lateral ridges (
<figureCitation id="7CEDA1DDFFC15C6638A244DFC027FEA8" box="[235,343,252,277]" captionStart="FIGURE 2" captionStartId="3.[151,250,1954,1976]" captionTargetBox="[166,1411,202,1909]" captionTargetId="figure@3.[159,1419,193,1928]" captionTargetPageId="3" captionText="FIGURE 2. Athanas alpheusophillus sp. nov., Vitjaz, Posjeta Bay, the Sea of Japan, paratypes (a, d, e – g — ovigerous female; b, c — male): a, b — front of carapace, dorsal view; c, d — front of carapace, lateral view; e — distal abdominal segments and telson, lateral view; f — telson; g — exopod of uropods." httpUri="https://zenodo.org/record/997055/files/figure.png" pageId="2">
Fig. 2 
<emphasis id="D6A2614AFFC15C66396744DEC027FEA8" box="[302,343,252,276]" italics="true" pageId="2">c, d</emphasis>
</figureCitation>
). Extra-corneal and infra-corneal teeth well developed, sharp, extra-corneal slightly smaller than infra-corneal (
<figureCitation id="7CEDA1DDFFC15C663971453EC0D4FE89" box="[312,420,285,310]" captionStart="FIGURE 2" captionStartId="3.[151,250,1954,1976]" captionTargetBox="[166,1411,202,1909]" captionTargetId="figure@3.[159,1419,193,1928]" captionTargetPageId="3" captionText="FIGURE 2. Athanas alpheusophillus sp. nov., Vitjaz, Posjeta Bay, the Sea of Japan, paratypes (a, d, e – g — ovigerous female; b, c — male): a, b — front of carapace, dorsal view; c, d — front of carapace, lateral view; e — distal abdominal segments and telson, lateral view; f — telson; g — exopod of uropods." httpUri="https://zenodo.org/record/997055/files/figure.png" pageId="2">
Fig. 2 
<emphasis id="D6A2614AFFC15C663932453DC0D4FE89" box="[379,420,285,309]" italics="true" pageId="2">c, d</emphasis>
</figureCitation>
). Pterygostomial angle feebly marked, sometimes bluntly protruding (
<figureCitation id="7CEDA1DDFFC15C663C89453EC45DFE89" box="[1216,1325,285,310]" captionStart="FIGURE 2" captionStartId="3.[151,250,1954,1976]" captionTargetBox="[166,1411,202,1909]" captionTargetId="figure@3.[159,1419,193,1928]" captionTargetPageId="3" captionText="FIGURE 2. Athanas alpheusophillus sp. nov., Vitjaz, Posjeta Bay, the Sea of Japan, paratypes (a, d, e – g — ovigerous female; b, c — male): a, b — front of carapace, dorsal view; c, d — front of carapace, lateral view; e — distal abdominal segments and telson, lateral view; f — telson; g — exopod of uropods." httpUri="https://zenodo.org/record/997055/files/figure.png" pageId="2">
Fig. 2 
<emphasis id="D6A2614AFFC15C663D4A453DC45DFE89" box="[1283,1325,285,309]" italics="true" pageId="2">c, d</emphasis>
</figureCitation>
). Cardiac notch well developed (
<figureCitation id="7CEDA1DDFFC15C6639DF451DC0A7FEEB" box="[406,471,318,343]" captionStart="FIGURE 1" captionStartId="1.[151,250,1956,1978]" captionTargetBox="[169,1422,953,1927]" captionTargetId="figure@1.[151,1436,937,1932]" captionTargetPageId="1" captionText="FIGURE 1. Athanas alpheusophillus sp. nov., Vitjaz, Posjeta Bay, the Sea of Japan, general view of paratype female (a) and holotype male (ZMMU Ma 5839) (b)." httpUri="https://zenodo.org/record/997053/files/figure.png" pageId="2">Fig. 1</figureCitation>
).
</paragraph>
<paragraph id="E469BD58FFC15C66388E457CC2D9FE42" blockId="2.[151,1437,151,2010]" pageId="2">
Pleomeres I–IV with ventrally rounded pleura, pleura of pleonite V with blunt posteroventral angle (
<figureCitation id="7CEDA1DDFFC15C663D0E457CC4FAFEC4" box="[1351,1418,351,376]" captionStart="FIGURE 1" captionStartId="1.[151,250,1956,1978]" captionTargetBox="[169,1422,953,1927]" captionTargetId="figure@1.[151,1436,937,1932]" captionTargetPageId="1" captionText="FIGURE 1. Athanas alpheusophillus sp. nov., Vitjaz, Posjeta Bay, the Sea of Japan, general view of paratype female (a) and holotype male (ZMMU Ma 5839) (b)." httpUri="https://zenodo.org/record/997053/files/figure.png" pageId="2">Fig. 1</figureCitation>
); pleonite VI with large subtriangular articulated plate and bluntly produced posterolateral angle (
<figureCitation id="7CEDA1DDFFC15C663CAE45A2C448FE25" box="[1255,1336,385,410]" captionStart="FIGURE 2" captionStartId="3.[151,250,1954,1976]" captionTargetBox="[166,1411,202,1909]" captionTargetId="figure@3.[159,1419,193,1928]" captionTargetPageId="3" captionText="FIGURE 2. Athanas alpheusophillus sp. nov., Vitjaz, Posjeta Bay, the Sea of Japan, paratypes (a, d, e – g — ovigerous female; b, c — male): a, b — front of carapace, dorsal view; c, d — front of carapace, lateral view; e — distal abdominal segments and telson, lateral view; f — telson; g — exopod of uropods." httpUri="https://zenodo.org/record/997055/files/figure.png" pageId="2">
Fig. 2 
<emphasis id="D6A2614AFFC15C663D6545A1C448FE25" box="[1324,1336,386,409]" italics="true" pageId="2">e</emphasis>
</figureCitation>
). Telson (
<figureCitation id="7CEDA1DDFFC15C6638D64581C19CFE07" box="[159,236,418,443]" captionStart="FIGURE 2" captionStartId="3.[151,250,1954,1976]" captionTargetBox="[166,1411,202,1909]" captionTargetId="figure@3.[159,1419,193,1928]" captionTargetPageId="3" captionText="FIGURE 2. Athanas alpheusophillus sp. nov., Vitjaz, Posjeta Bay, the Sea of Japan, paratypes (a, d, e – g — ovigerous female; b, c — male): a, b — front of carapace, dorsal view; c, d — front of carapace, lateral view; e — distal abdominal segments and telson, lateral view; f — telson; g — exopod of uropods." httpUri="https://zenodo.org/record/997055/files/figure.png" pageId="2">
Fig. 2 
<emphasis id="D6A2614AFFC15C6638AD4580C19CFE07" box="[228,236,419,443]" italics="true" pageId="2">f</emphasis>
</figureCitation>
) subrectangular, slightly tapering distally; dorsal surface with two pairs of stout spines, situated at about 1/2 and 1/4 of telson length, respectively; posterior margin rounded, posterolateral angle with 
<emphasis id="D6A2614AFFC15C663D6845E7C45FFE61" bold="true" box="[1313,1327,452,477]" pageId="2">2</emphasis>
pairs of submarginal spines, the lateral spine is 
<emphasis id="D6A2614AFFC15C663A0445C6C3D7FE42" bold="true" box="[589,679,485,510]" pageId="2">about 3</emphasis>
times shorter (
<figureCitation id="7CEDA1DDFFC15C663B0645C6C2E9FE41" box="[847,921,485,510]" captionStart="FIGURE 2" captionStartId="3.[151,250,1954,1976]" captionTargetBox="[166,1411,202,1909]" captionTargetId="figure@3.[159,1419,193,1928]" captionTargetPageId="3" captionText="FIGURE 2. Athanas alpheusophillus sp. nov., Vitjaz, Posjeta Bay, the Sea of Japan, paratypes (a, d, e – g — ovigerous female; b, c — male): a, b — front of carapace, dorsal view; c, d — front of carapace, lateral view; e — distal abdominal segments and telson, lateral view; f — telson; g — exopod of uropods." httpUri="https://zenodo.org/record/997055/files/figure.png" pageId="2">
Fig. 2 
<emphasis id="D6A2614AFFC15C663BD845C6C2E9FE41" box="[913,921,485,509]" italics="true" pageId="2">f</emphasis>
</figureCitation>
).
</paragraph>
<paragraph id="E469BD58FFC15C66388E4624C282FDFC" blockId="2.[151,1437,151,2010]" pageId="2">
Eyes only partly concealed by carapace, most-posterior portion invisible in dorsal and lateral views; cornea well developed and pigmented, occupying most of eye peduncle (
<figureCitation id="7CEDA1DDFFC15C663B3F4604C292FDFC" box="[886,994,551,576]" captionStart="FIGURE 2" captionStartId="3.[151,250,1954,1976]" captionTargetBox="[166,1411,202,1909]" captionTargetId="figure@3.[159,1419,193,1928]" captionTargetPageId="3" captionText="FIGURE 2. Athanas alpheusophillus sp. nov., Vitjaz, Posjeta Bay, the Sea of Japan, paratypes (a, d, e – g — ovigerous female; b, c — male): a, b — front of carapace, dorsal view; c, d — front of carapace, lateral view; e — distal abdominal segments and telson, lateral view; f — telson; g — exopod of uropods." httpUri="https://zenodo.org/record/997055/files/figure.png" pageId="2">
Fig. 
<emphasis id="D6A2614AFFC15C663BE3460BC292FDFC" box="[938,994,552,576]" italics="true" pageId="2">2a–d</emphasis>
</figureCitation>
).
</paragraph>
<paragraph id="E469BD58FFC15C66388E466AC577FD5B" blockId="2.[151,1437,151,2010]" pageId="2">
Antennula (
<figureCitation id="7CEDA1DDFFC15C663904466AC0B7FDDD" box="[333,455,585,610]" captionStart="FIGURE 2" captionStartId="3.[151,250,1954,1976]" captionTargetBox="[166,1411,202,1909]" captionTargetId="figure@3.[159,1419,193,1928]" captionTargetPageId="3" captionText="FIGURE 2. Athanas alpheusophillus sp. nov., Vitjaz, Posjeta Bay, the Sea of Japan, paratypes (a, d, e – g — ovigerous female; b, c — male): a, b — front of carapace, dorsal view; c, d — front of carapace, lateral view; e — distal abdominal segments and telson, lateral view; f — telson; g — exopod of uropods." httpUri="https://zenodo.org/record/997055/files/figure.png" pageId="2">
Figs. 2 
<emphasis id="D6A2614AFFC15C6639D54669C0B7FDDD" box="[412,455,585,609]" italics="true" pageId="2">a, b</emphasis>
</figureCitation>
, 
<figureCitation id="7CEDA1DDFFC15C66399E466AC361FDDD" box="[471,529,585,609]" captionStart="FIGURE 3" captionStartId="4.[151,250,1952,1974]" captionTargetBox="[168,1430,207,1912]" captionTargetId="figure@4.[151,1436,193,1931]" captionTargetPageId="4" captionText="FIGURE 3. Athanas alpheusophillus sp. nov., Vitjaz, Posjeta Bay, the Sea of Japan, paratypes (a, c, d — ovigerous female; b, e — male): a, b — antennula; c, e — pereiopods I (chelipeds); d — distal propodus and chela of pereiopod I." httpUri="https://zenodo.org/record/997057/files/figure.png" pageId="2">
3 
<emphasis id="D6A2614AFFC15C6639AC4669C361FDDD" box="[485,529,585,609]" italics="true" pageId="2">a, b</emphasis>
</figureCitation>
) with antennular peduncle relatively slender; basal segment about twice longer than wide, with strong tooth on mesioventral margin, stylocerite relatively slender, acute distally, overreaching distal margin of basal antennular segment, but not reaching mid-length of second segment of antennular peduncle; latter about 0.8 times as long as basal segment, slightly longer than distal segment, about 1.5 times as long as wide in dorsal view; distal segment about 1.5 times longer than wide in dorsal view.
</paragraph>
<paragraph id="E469BD58FFC15C66388E46D3C232FCD0" blockId="2.[151,1437,151,2010]" pageId="2">
Antenna (
<figureCitation id="7CEDA1DDFFC15C66397E46CCC0D7FCB4" box="[311,423,751,776]" captionStart="FIGURE 2" captionStartId="3.[151,250,1954,1976]" captionTargetBox="[166,1411,202,1909]" captionTargetId="figure@3.[159,1419,193,1928]" captionTargetPageId="3" captionText="FIGURE 2. Athanas alpheusophillus sp. nov., Vitjaz, Posjeta Bay, the Sea of Japan, paratypes (a, d, e – g — ovigerous female; b, c — male): a, b — front of carapace, dorsal view; c, d — front of carapace, lateral view; e — distal abdominal segments and telson, lateral view; f — telson; g — exopod of uropods." httpUri="https://zenodo.org/record/997055/files/figure.png" pageId="2">
Fig. 2 
<emphasis id="D6A2614AFFC15C66393246D2C0D7FCB4" box="[379,423,752,776]" italics="true" pageId="2">a, b</emphasis>
</figureCitation>
) with basicerite fairly stout, bearing strong, sharp ventrolateral tooth overreaching distal margin of the segment (
<figureCitation id="7CEDA1DDFFC15C6639ED4732C360FC95" box="[420,528,785,810]" captionStart="FIGURE 2" captionStartId="3.[151,250,1954,1976]" captionTargetBox="[166,1411,202,1909]" captionTargetId="figure@3.[159,1419,193,1928]" captionTargetPageId="3" captionText="FIGURE 2. Athanas alpheusophillus sp. nov., Vitjaz, Posjeta Bay, the Sea of Japan, paratypes (a, d, e – g — ovigerous female; b, c — male): a, b — front of carapace, dorsal view; c, d — front of carapace, lateral view; e — distal abdominal segments and telson, lateral view; f — telson; g — exopod of uropods." httpUri="https://zenodo.org/record/997055/files/figure.png" pageId="2">
Fig. 2 
<emphasis id="D6A2614AFFC15C6639AE4731C360FC95" box="[487,528,785,809]" italics="true" pageId="2">c, d</emphasis>
</figureCitation>
); scaphocerite subrectangular, reaching distal margin of antennular segment and distal margin of carpocerite; distolateral tooth strong, acute; distolateral tooth, convex, sloping posteriorly overreaching distal margin of scaphocerite blade (
<figureCitation id="7CEDA1DDFFC15C663A8E4777C242FCD0" box="[711,818,852,877]" captionStart="FIGURE 2" captionStartId="3.[151,250,1954,1976]" captionTargetBox="[166,1411,202,1909]" captionTargetId="figure@3.[159,1419,193,1928]" captionTargetPageId="3" captionText="FIGURE 2. Athanas alpheusophillus sp. nov., Vitjaz, Posjeta Bay, the Sea of Japan, paratypes (a, d, e – g — ovigerous female; b, c — male): a, b — front of carapace, dorsal view; c, d — front of carapace, lateral view; e — distal abdominal segments and telson, lateral view; f — telson; g — exopod of uropods." httpUri="https://zenodo.org/record/997055/files/figure.png" pageId="2">
Fig. 2 
<emphasis id="D6A2614AFFC15C663B404776C242FCD0" box="[777,818,852,876]" italics="true" pageId="2">c, d</emphasis>
</figureCitation>
).
</paragraph>
<paragraph id="E469BD58FFC15C66388E4756C41AFC4E" blockId="2.[151,1437,151,2010]" pageId="2">Mouthparts typical for the genus. Maxilliped III slender, pediform; coxa with ear-shaped, distally subacute lateral plate; arthrobranch absent; antepenultimate segment about 6 times as long as wide, with a long slender subdistal spinule (or spine-like seta) on distoventral margin; penultimate segment about twice longer than wide; ultimate segment about 4.5 times as long as wide, tip furnished with long strong setae, without marked spines.</paragraph>
<paragraph id="E469BD58FFC15C66388E47D9C415F9D7" blockId="2.[151,1437,151,2010]" pageId="2">
Pereiopods I (chelipeds) differ in males and females. Chelipeds relatively small, subequal and symmetrical (
<figureCitation id="7CEDA1DDFFC15C6638D6403FC18FFB88" box="[159,255,1052,1077]" captionStart="FIGURE 1" captionStartId="1.[151,250,1956,1978]" captionTargetBox="[169,1422,953,1927]" captionTargetId="figure@1.[151,1436,937,1932]" captionTargetPageId="1" captionText="FIGURE 1. Athanas alpheusophillus sp. nov., Vitjaz, Posjeta Bay, the Sea of Japan, general view of paratype female (a) and holotype male (ZMMU Ma 5839) (b)." httpUri="https://zenodo.org/record/997053/files/figure.png" pageId="2">
Figs. 1 
<emphasis id="D6A2614AFFC15C6638B8403EC18FFB88" box="[241,255,1053,1076]" italics="true" pageId="2">a</emphasis>
</figureCitation>
, 
<figureCitation id="7CEDA1DDFFC15C66395B403FC05BFB88" box="[274,299,1052,1076]" captionStart="FIGURE 3" captionStartId="4.[151,250,1952,1974]" captionTargetBox="[168,1430,207,1912]" captionTargetId="figure@4.[151,1436,193,1931]" captionTargetPageId="4" captionText="FIGURE 3. Athanas alpheusophillus sp. nov., Vitjaz, Posjeta Bay, the Sea of Japan, paratypes (a, c, d — ovigerous female; b, e — male): a, b — antennula; c, e — pereiopods I (chelipeds); d — distal propodus and chela of pereiopod I." httpUri="https://zenodo.org/record/997057/files/figure.png" pageId="2">
3 
<emphasis id="D6A2614AFFC15C663956403EC05BFB88" box="[287,299,1053,1076]" italics="true" pageId="2">c</emphasis>
</figureCitation>
) in females, with coxa and basis unarmed, coxa with epipod; basis with minuscule rudimentary exopod; ischium about 4 times as long as wide, with unarmed margin; merus relatively slender, with straight margin, about 7 times as long as wide, slightly longer than ischium and carpus; carpus relatively short, slender, distally widening, with rows of long simple setae along distodorsal margin; palm (
<figureCitation id="7CEDA1DDFFC15C663C7D405CC5F5FB24" box="[1076,1157,1151,1176]" captionStart="FIGURE 3" captionStartId="4.[151,250,1952,1974]" captionTargetBox="[168,1430,207,1912]" captionTargetId="figure@4.[151,1436,193,1931]" captionTargetPageId="4" captionText="FIGURE 3. Athanas alpheusophillus sp. nov., Vitjaz, Posjeta Bay, the Sea of Japan, paratypes (a, c, d — ovigerous female; b, e — male): a, b — antennula; c, e — pereiopods I (chelipeds); d — distal propodus and chela of pereiopod I." httpUri="https://zenodo.org/record/997057/files/figure.png" pageId="2">
Fig. 3 
<emphasis id="D6A2614AFFC15C663C3E40A3C5F5FB24" box="[1143,1157,1152,1176]" italics="true" pageId="2">d</emphasis>
</figureCitation>
) smooth, subcylindrical, with straight lateral margins, about twice longer than wide; fingers slender, subcylindrical, straight, about 4 times longer than wide, with straight cutting margins. Chelipeds greatly enlarged, subequal in size, but slightly asymmetrical in shape (
<figureCitation id="7CEDA1DDFFC15C6639E140C7C089FB40" box="[424,505,1252,1277]" captionStart="FIGURE 3" captionStartId="4.[151,250,1952,1974]" captionTargetBox="[168,1430,207,1912]" captionTargetId="figure@4.[151,1436,193,1931]" captionTargetPageId="4" captionText="FIGURE 3. Athanas alpheusophillus sp. nov., Vitjaz, Posjeta Bay, the Sea of Japan, paratypes (a, c, d — ovigerous female; b, e — male): a, b — antennula; c, e — pereiopods I (chelipeds); d — distal propodus and chela of pereiopod I." httpUri="https://zenodo.org/record/997057/files/figure.png" pageId="2">
Fig. 3 
<emphasis id="D6A2614AFFC15C6639A440C6C089FB40" box="[493,505,1253,1276]" italics="true" pageId="2">e</emphasis>
</figureCitation>
) in males, carried folded with ventral portion of palm fitting in ventrally deeply excavated merus in life (
<figureCitation id="7CEDA1DDFFC15C6639E24126C08BFAA1" box="[427,507,1285,1310]" captionStart="FIGURE 1" captionStartId="1.[151,250,1956,1978]" captionTargetBox="[169,1422,953,1927]" captionTargetId="figure@1.[151,1436,937,1932]" captionTargetPageId="1" captionText="FIGURE 1. Athanas alpheusophillus sp. nov., Vitjaz, Posjeta Bay, the Sea of Japan, general view of paratype female (a) and holotype male (ZMMU Ma 5839) (b)." httpUri="https://zenodo.org/record/997053/files/figure.png" pageId="2">
Fig. 1 
<emphasis id="D6A2614AFFC15C6639A44126C08BFAA1" box="[493,507,1285,1309]" italics="true" pageId="2">b</emphasis>
</figureCitation>
). Major cheliped (
<figureCitation id="7CEDA1DDFFC15C663A854126C27EFAA2" box="[716,782,1285,1310]" captionStart="FIGURE 5" captionStartId="6.[151,250,1931,1953]" captionTargetBox="[208,1378,220,1888]" captionTargetId="figure@6.[195,1391,211,1899]" captionTargetPageId="6" captionText="FIGURE 5. Athanas alpheusophillus sp. nov., Vitjaz, Posjeta Bay, the Sea of Japan, paratype male, major pereiopod I: a — general view of pereiopod I; b — distal segments of pereiopod I; c, d — distal propodus and fingers of pereiopod I, different views." httpUri="https://zenodo.org/record/997061/files/figure.png" pageId="2">Fig. 5</figureCitation>
) with stout coxa and short basis, both unarmed; coxa with epipod; basis with minuscule rudimentary exopod; ischium about 2.5 times longer than wide, with single long simple spine in distodorsal angle; merus significantly broadened medially, ventral surface excavated, excavated lobe armed with several blunt small teeth along distal margin; carpus relatively slender, vase-shaped, distally widening, about 3 times longer than maximum width; palm smooth, subcylindrical, medially inflated distinctly flattened, with moderately convex dorsal and strongly convex ventral margins, without lateral teeth; polex (fixed finger) (
<figureCitation id="7CEDA1DDFFC15C6638BA41EEC02EFA59" box="[243,350,1485,1510]" captionStart="FIGURE 5" captionStartId="6.[151,250,1931,1953]" captionTargetBox="[208,1378,220,1888]" captionTargetId="figure@6.[195,1391,211,1899]" captionTargetPageId="6" captionText="FIGURE 5. Athanas alpheusophillus sp. nov., Vitjaz, Posjeta Bay, the Sea of Japan, paratype male, major pereiopod I: a — general view of pereiopod I; b — distal segments of pereiopod I; c, d — distal propodus and fingers of pereiopod I, different views." httpUri="https://zenodo.org/record/997061/files/figure.png" pageId="2">
Fig. 5 
<emphasis id="D6A2614AFFC15C66397F41EEC02EFA59" box="[310,350,1485,1509]" italics="true" pageId="2">b, c</emphasis>
</figureCitation>
) subcylindrical, tapering and curved in distal part, with sharp tip, cutting margin with numerous small teeth, unarmed with dense rows of setae; dactylus (
<figureCitation id="7CEDA1DDFFC15C663B5641CDC2FBF9BA" box="[799,907,1518,1543]" captionStart="FIGURE 5" captionStartId="6.[151,250,1931,1953]" captionTargetBox="[208,1378,220,1888]" captionTargetId="figure@6.[195,1391,211,1899]" captionTargetPageId="6" captionText="FIGURE 5. Athanas alpheusophillus sp. nov., Vitjaz, Posjeta Bay, the Sea of Japan, paratype male, major pereiopod I: a — general view of pereiopod I; b — distal segments of pereiopod I; c, d — distal propodus and fingers of pereiopod I, different views." httpUri="https://zenodo.org/record/997061/files/figure.png" pageId="2">
Fig. 5 
<emphasis id="D6A2614AFFC15C663B2B41CCC2FBF9BA" box="[866,907,1518,1542]" italics="true" pageId="2">c, d</emphasis>
</figureCitation>
) equal in size to polex, tapering and curved in distal part, with sharp tip, cutting margin with numerous small teeth, with well-marked large dorsal crest, outer lateral part of finger furnished with dense row of long simple setae, forming dense lateroventrally directed setal brush. Minor cheliped (
<figureCitation id="7CEDA1DDFFC15C6639D74271C087F9D6" box="[414,503,1618,1643]" captionStart="FIGURE 3" captionStartId="4.[151,250,1952,1974]" captionTargetBox="[168,1430,207,1912]" captionTargetId="figure@4.[151,1436,193,1931]" captionTargetPageId="4" captionText="FIGURE 3. Athanas alpheusophillus sp. nov., Vitjaz, Posjeta Bay, the Sea of Japan, paratypes (a, c, d — ovigerous female; b, e — male): a, b — antennula; c, e — pereiopods I (chelipeds); d — distal propodus and chela of pereiopod I." httpUri="https://zenodo.org/record/997057/files/figure.png" pageId="2">
Figs. 3 
<emphasis id="D6A2614AFFC15C6639A24270C087F9D6" box="[491,503,1619,1642]" italics="true" pageId="2">e</emphasis>
</figureCitation>
, 
<figureCitation id="7CEDA1DDFFC15C663A4F4270C34CF9D6" box="[518,572,1619,1643]" captionStart="FIGURE 4" captionStartId="5.[151,250,1983,2005]" captionTargetBox="[169,1414,218,1948]" captionTargetId="figure@5.[161,1425,193,1962]" captionTargetPageId="5" captionText="FIGURE 4. Athanas alpheusophillus sp. nov., Vitjaz, Posjeta Bay, the Sea of Japan, paratypes (a – c — male; d, e — female): a, b — minor pereiopod I; c — distal propodus and fingers of pereiopod I; d — pereiopod II; e — pereiopod III." httpUri="https://zenodo.org/record/997059/files/figure.png" pageId="2">
4 
<emphasis id="D6A2614AFFC15C663A5D4270C34CF9D6" box="[532,572,1619,1642]" italics="true" pageId="2">a–c</emphasis>
</figureCitation>
) morphologically similar and only slightly smaller than major cheliped.
</paragraph>
<paragraph id="E469BD58FFC15C66388E4257C4F2F94C" blockId="2.[151,1437,151,2010]" pageId="2">
Pereiopod II (
<figureCitation id="7CEDA1DDFFC15C66392B4257C0C3F930" box="[354,435,1652,1677]" captionStart="FIGURE 4" captionStartId="5.[151,250,1983,2005]" captionTargetBox="[169,1414,218,1948]" captionTargetId="figure@5.[161,1425,193,1962]" captionTargetPageId="5" captionText="FIGURE 4. Athanas alpheusophillus sp. nov., Vitjaz, Posjeta Bay, the Sea of Japan, paratypes (a – c — male; d, e — female): a, b — minor pereiopod I; c — distal propodus and fingers of pereiopod I; d — pereiopod II; e — pereiopod III." httpUri="https://zenodo.org/record/997059/files/figure.png" pageId="2">
Fig. 4 
<emphasis id="D6A2614AFFC15C6639EC4257C0C3F930" box="[421,435,1652,1676]" italics="true" pageId="2">d</emphasis>
</figureCitation>
) relatively slender; coxa and basis short, coxal with small epipod; basis without exopod; ischium unarmed, about 6 times as long as wide, about 1.5 times shorter than merus; merus unarmed, with straight margins, about 6 times as long as wide; carpus with 
<emphasis id="D6A2614AFFC15C663AB24295C279F973" bold="true" box="[763,777,1718,1743]" pageId="2">5</emphasis>
segments with ratio approximately equal to (proximal to distal) 4:1:1:1:2; chela simple, slender, slightly shorter that fifth carpal segment; fingers simple, as long as palm.
</paragraph>
<paragraph id="E469BD58FFC15C66388E42DAC58CF866" blockId="2.[151,1437,151,2010]" pageId="2">
Ambulatory pereiopods (III–V) moderately slender. Pereiopod III (
<figureCitation id="7CEDA1DDFFC15C663BFA42DAC571F8AD" box="[947,1025,1785,1810]" captionStart="FIGURE 4" captionStartId="5.[151,250,1983,2005]" captionTargetBox="[169,1414,218,1948]" captionTargetId="figure@5.[161,1425,193,1962]" captionTargetPageId="5" captionText="FIGURE 4. Athanas alpheusophillus sp. nov., Vitjaz, Posjeta Bay, the Sea of Japan, paratypes (a – c — male; d, e — female): a, b — minor pereiopod I; c — distal propodus and fingers of pereiopod I; d — pereiopod II; e — pereiopod III." httpUri="https://zenodo.org/record/997059/files/figure.png" pageId="2">
Fig. 4 
<emphasis id="D6A2614AFFC15C663BBC42D9C571F8AD" box="[1013,1025,1786,1809]" italics="true" pageId="2">e</emphasis>
</figureCitation>
) with short coxal segment and small epipod; basis short, as long as wide; ischium about 3 times as long as wide, with 2 small ventrolateral spines; merus unarmed, about 5.5 times as long as wide, unarmed; carpus about 4 times as long as maximal width, widening distally, unarmed; propodus about 6 times as long as wide, with 5 minute ventral spines and one pair of distoventral spinules; dactylus relatively slender, simple, curved distally, about half of propodus length. Pereiopods IV and V similar to pereiopod III, but more slender; propodus of pereiopod V armed with 2 minute spines in proximal third of length and several rows of short setae distally (propodal cleaning brush); dactylus simple, slender, slightly curved.
</paragraph>
<caption id="B0A9EDD0FFC05C6738DE4381C3C9F84A" httpUri="https://zenodo.org/record/997055/files/figure.png" pageId="3" targetBox="[166,1411,202,1909]" targetPageId="3">
<paragraph id="E469BD58FFC05C6738DE4381C3C9F84A" blockId="3.[151,1436,1954,2038]" pageId="3">
<emphasis id="D6A2614AFFC05C6738DE4381C305F804" bold="true" box="[151,629,1954,1977]" pageId="3">
FIGURE 2. 
<taxonomicName id="23D6C6DBFFC05C6739534381C36EF805" authority="Marin, 2017" authorityName="Marin" authorityYear="2017" box="[282,542,1954,1977]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="3" phylum="Arthropoda" rank="species" species="alpheusophillus" status="sp. nov.">
<emphasis id="D6A2614AFFC05C6739534381C36EF805" bold="true" box="[282,542,1954,1977]" italics="true" pageId="3">Athanas alpheusophillus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CD91DC31FFC05C673A6D4380C305F804" box="[548,629,1955,1976]" pageId="3" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, Vitjaz, Posjeta Bay, the Sea of Japan, paratypes (
<emphasis id="D6A2614AFFC05C673C3D4380C5BBF804" box="[1140,1227,1954,1976]" italics="true" pageId="3">a, d, e–g</emphasis>
—ovigerous female; 
<emphasis id="D6A2614AFFC05C6738DE43E2C1CEF86B" box="[151,190,1985,2007]" italics="true" pageId="3">b, c</emphasis>
—male): 
<emphasis id="D6A2614AFFC05C67396843E1C039F86B" box="[289,329,1985,2007]" italics="true" pageId="3">a, b</emphasis>
—front of carapace, dorsal view; 
<emphasis id="D6A2614AFFC05C673AE543E1C3A4F86B" box="[684,724,1985,2007]" italics="true" pageId="3">c, d</emphasis>
—front of carapace, lateral view; 
<emphasis id="D6A2614AFFC05C673C7E43E1C532F86B" box="[1079,1090,1986,2007]" italics="true" pageId="3">e</emphasis>
—distal abdominal segments and telson, lateral view; 
<emphasis id="D6A2614AFFC05C67392D43C3C01BF84A" box="[356,363,2016,2038]" italics="true" pageId="3">f</emphasis>
—telson; 
<emphasis id="D6A2614AFFC05C67398743C2C0AAF84A" box="[462,474,2017,2038]" italics="true" pageId="3">g</emphasis>
—exopod of uropods.
</paragraph>
</caption>
<caption id="B0A9EDD0FFC75C6038DE4383C5C4F869" httpUri="https://zenodo.org/record/997057/files/figure.png" pageId="4" targetBox="[168,1430,207,1912]" targetPageId="4">
<paragraph id="E469BD58FFC75C6038DE4383C5C4F869" blockId="4.[151,1435,1952,2005]" pageId="4">
<emphasis id="D6A2614AFFC75C6038DE4383C07DF809" bold="true" box="[151,269,1952,1974]" pageId="4">FIGURE 3</emphasis>
. 
<emphasis id="D6A2614AFFC75C6039534383C305F80A" bold="true" box="[282,629,1952,1975]" pageId="4">
<taxonomicName id="23D6C6DBFFC75C6039534383C36EF80B" authority="Marin, 2017" authorityName="Marin" authorityYear="2017" box="[282,542,1952,1975]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="4" phylum="Arthropoda" rank="species" species="alpheusophillus" status="sp. nov.">
<emphasis id="D6A2614AFFC75C6039534383C36EF80B" bold="true" box="[282,542,1952,1975]" italics="true" pageId="4">Athanas alpheusophillus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CD91DC31FFC75C603A6D4382C305F80A" box="[548,629,1953,1974]" pageId="4" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, Vitjaz, Posjeta Bay, the Sea of Japan, paratypes (
<emphasis id="D6A2614AFFC75C603C3D4381C5E8F80B" box="[1140,1176,1954,1975]" italics="true" pageId="4">a, c</emphasis>
, 
<emphasis id="D6A2614AFFC75C603CEF4383C5C2F80A" box="[1190,1202,1952,1974]" italics="true" pageId="4">d</emphasis>
—ovigerous female; 
<emphasis id="D6A2614AFFC75C603DC04383C1D2F869" italics="true" pageId="4">b, e</emphasis>
—male): 
<emphasis id="D6A2614AFFC75C60394B43E3C059F869" box="[258,297,1983,2005]" italics="true" pageId="4">a, b</emphasis>
—antennula; 
<emphasis id="D6A2614AFFC75C6039F843E3C0A4F869" box="[433,468,1984,2005]" italics="true" pageId="4">c, e</emphasis>
—pereiopods I (chelipeds); 
<emphasis id="D6A2614AFFC75C603AA6439CC38BF869" box="[751,763,1983,2005]" italics="true" pageId="4">d</emphasis>
—distal propodus and chela of pereiopod I.
</paragraph>
</caption>
<caption id="B0A9EDD0FFC65C6138DE439CC5CCF84F" httpUri="https://zenodo.org/record/997059/files/figure.png" pageId="5" targetBox="[169,1414,218,1948]" targetPageId="5">
<paragraph id="E469BD58FFC65C6138DE439CC5CCF84F" blockId="5.[151,1435,1982,2036]" pageId="5">
<emphasis id="D6A2614AFFC65C6138DE439CC07DF868" bold="true" box="[151,269,1983,2005]" pageId="5">FIGURE 4</emphasis>
. 
<emphasis id="D6A2614AFFC65C613953439DC305F868" bold="true" box="[282,629,1982,2005]" pageId="5">
<taxonomicName id="23D6C6DBFFC65C613953439DC36DF869" authority="Marin, 2017" authorityName="Marin" authorityYear="2017" box="[282,541,1982,2005]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="5" phylum="Arthropoda" rank="species" species="alpheusophillus" status="sp. nov.">
<emphasis id="D6A2614AFFC65C613953439DC36DF869" bold="true" box="[282,541,1982,2005]" italics="true" pageId="5">Athanas alpheusophillus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CD91DC31FFC65C613A6D439CC305F868" box="[548,629,1983,2004]" pageId="5" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, Vitjaz, Posjeta Bay, the Sea of Japan, paratypes (
<emphasis id="D6A2614AFFC65C613C3A43E3C5E7F869" box="[1139,1175,1984,2005]" italics="true" pageId="5">a–c</emphasis>
—male; 
<emphasis id="D6A2614AFFC65C613CB9439CC464F869" box="[1264,1300,1983,2005]" italics="true" pageId="5">d, e</emphasis>
—female): 
<emphasis id="D6A2614AFFC65C613DC043E3C1D4F84F" italics="true" pageId="5">a, b</emphasis>
—minor pereiopod I; 
<emphasis id="D6A2614AFFC65C6139C843FCC0FCF848" box="[385,396,2015,2036]" italics="true" pageId="5">c</emphasis>
—distal propodus and fingers of pereiopod I; 
<emphasis id="D6A2614AFFC65C613B1243FEC218F84F" box="[859,872,2013,2035]" italics="true" pageId="5">d</emphasis>
—pereiopod II; 
<emphasis id="D6A2614AFFC65C613C4243FCC566F848" box="[1035,1046,2015,2036]" italics="true" pageId="5">e</emphasis>
—pereiopod III.
</paragraph>
</caption>
<caption id="B0A9EDD0FFC55C6238DE43A8C1A9F862" httpUri="https://zenodo.org/record/997061/files/figure.png" pageId="6" targetBox="[208,1378,220,1888]" targetPageId="6">
<paragraph id="E469BD58FFC55C6238DE43A8C1A9F862" blockId="6.[151,1436,1930,2014]" pageId="6">
<emphasis id="D6A2614AFFC55C6238DE43A8C07EF81C" bold="true" box="[151,270,1931,1953]" pageId="6">FIGURE 5</emphasis>
. 
<emphasis id="D6A2614AFFC55C62395543A9C30AF81C" bold="true" box="[284,634,1930,1953]" pageId="6">
<taxonomicName id="23D6C6DBFFC55C62395543A9C350F81D" authority="Marin, 2017" authorityName="Marin" authorityYear="2017" box="[284,544,1930,1953]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="6" phylum="Arthropoda" rank="species" species="alpheusophillus" status="sp. nov.">
<emphasis id="D6A2614AFFC55C62395543A9C350F81D" bold="true" box="[284,544,1930,1953]" italics="true" pageId="6">Athanas alpheusophillus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CD91DC31FFC55C623A6143A8C30AF81C" box="[552,634,1931,1952]" pageId="6" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, Vitjaz, Posjeta Bay, the Sea of Japan, paratype male, major pereiopod I: 
<emphasis id="D6A2614AFFC55C623D3F43AFC4F3F81D" box="[1398,1411,1932,1953]" italics="true" pageId="6">a</emphasis>
— general view of pereiopod I; 
<emphasis id="D6A2614AFFC55C623983438AC0A7F803" box="[458,471,1961,1983]" italics="true" pageId="6">b</emphasis>
—distal segments of pereiopod I; 
<emphasis id="D6A2614AFFC55C623B754389C214F803" box="[828,868,1961,1983]" italics="true" pageId="6">c, d</emphasis>
—distal propodus and fingers of pereiopod I, different views.
</paragraph>
</caption>
<caption id="B0A9EDD0FFC45C6338DE43A7C332F805" httpUri="https://zenodo.org/record/997063/files/figure.png" pageId="7" targetBox="[151,1436,193,1902]" targetPageId="7">
<paragraph id="E469BD58FFC45C6338DE43A7C332F805" blockId="7.[151,1436,1923,1977]" pageId="7">
<emphasis id="D6A2614AFFC45C6338DE43A7C303F825" bold="true" box="[151,627,1923,1946]" pageId="7">
FIGURE 6. 
<taxonomicName id="23D6C6DBFFC45C63395043A0C36CF826" authority="Marin, 2017" authorityName="Marin" authorityYear="2017" box="[281,540,1923,1946]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="7" phylum="Arthropoda" rank="species" species="alpheusophillus" status="sp. nov.">
<emphasis id="D6A2614AFFC45C63395043A0C36CF826" bold="true" box="[281,540,1923,1946]" italics="true" pageId="7">Athanas alpheusophillus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CD91DC31FFC45C633A6B43A7C303F825" box="[546,627,1924,1945]" pageId="7" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, Vitjaz, Posjeta Bay, the Sea of Japan, alive coloration of female paratype (
<emphasis id="D6A2614AFFC45C633D2443A6C4E4F826" box="[1389,1428,1924,1946]" italics="true" pageId="7">a, b</emphasis>
) and male holotype (ZMMU Ma 5839) (
<emphasis id="D6A2614AFFC45C633A6E4387C342F805" box="[551,562,1956,1977]" italics="true" pageId="7">c</emphasis>
).
</paragraph>
</caption>
<paragraph id="E469BD58FFCB5C6C388E44B4C025FF6E" blockId="8.[151,1437,151,2022]" pageId="8">
Uropod with exopod armed with fairly stout distolateral spine (
<figureCitation id="7CEDA1DDFFCB5C6C3BE744B4C572FF0C" box="[942,1026,151,176]" captionStart="FIGURE 2" captionStartId="3.[151,250,1954,1976]" captionTargetBox="[166,1411,202,1909]" captionTargetId="figure@3.[159,1419,193,1928]" captionTargetPageId="3" captionText="FIGURE 2. Athanas alpheusophillus sp. nov., Vitjaz, Posjeta Bay, the Sea of Japan, paratypes (a, d, e – g — ovigerous female; b, c — male): a, b — front of carapace, dorsal view; c, d — front of carapace, lateral view; e — distal abdominal segments and telson, lateral view; f — telson; g — exopod of uropods." httpUri="https://zenodo.org/record/997055/files/figure.png" pageId="8">
Fig. 
<quantity id="232E10BDFFCB5C6C3BAF44BBC572FF0C" box="[998,1026,152,176]" metricMagnitude="-3" metricUnit="kg" metricValue="2.0" pageId="8" unit="g" value="2.0">
2 
<emphasis id="D6A2614AFFCB5C6C3BBD44BAC572FF0C" box="[1012,1026,153,176]" italics="true" pageId="8">g</emphasis>
</quantity>
</figureCitation>
); diaeresis almost straight, without specific features.
</paragraph>
<paragraph id="E469BD58FFCB5C6C388E44F9C5B8FEA8" blockId="8.[151,1437,151,2022]" pageId="8">Gill/exopod formula: 5 pleurobranchs (PI–V); 0 arthrobranchs; 0 podobranchs; 4 mastigobranchs (MxpIII, PI– III); 5 setobranchs (PI–IV); 3 well-developed exopods (MxpI–III) + 1 rudimentary exopod (PI).</paragraph>
<paragraph id="E469BD58FFCB5C6C388E453EC251FEC4" blockId="8.[151,1437,151,2022]" pageId="8">
<emphasis id="D6A2614AFFCB5C6C388E453EC050FE8A" bold="true" box="[199,288,285,310]" pageId="8">Colour.</emphasis>
Body and appendages deep red; white dorsal line stretching from rostral tip through mid-dorsal area of carapace to distal margin of pleomere VI; distal parts of uropodal exopod white; some small isolated white spots present on lateral surface of carapace and pleurae (
<figureCitation id="7CEDA1DDFFCB5C6C3A86457CC263FEC4" box="[719,787,351,376]" captionStart="FIGURE 7" captionStartId="9.[151,250,1571,1593]" captionTargetBox="[151,1436,628,1535]" captionTargetId="figure@9.[151,1436,628,1550]" captionTargetPageId="9" captionText="FIGURE 7. The map of records of representatives of the genus Athanas Leach, 1814 in the Sea of Japan and adjacent area. Empty sign indicates the type locality of the species." httpUri="https://zenodo.org/record/997065/files/figure.png" pageId="8">Fig. 7</figureCitation>
).
</paragraph>
<paragraph id="E469BD58FFCB5C6C388E45A2C36AFE60" blockId="8.[151,1437,151,2022]" pageId="8">
<emphasis id="D6A2614AFFCB5C6C388E45A2C30DFE26" bold="true" box="[199,637,384,410]" pageId="8">
Size. 
<taxonomicName id="23D6C6DBFFCB5C6C394C45A3C36CFE25" authority="Marin, 2017" authorityName="Marin" authorityYear="2017" box="[261,540,384,409]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="8" phylum="Arthropoda" rank="species" species="alpheusophilus" status="sp. nov.">
<emphasis id="D6A2614AFFCB5C6C394C45A3C36CFE25" bold="true" box="[261,540,384,409]" italics="true" pageId="8">Athanas alpheusophilus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CD91DC31FFCB5C6C3A6A45A2C30DFE26" box="[547,637,385,410]" pageId="8" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is a relatively large species within the genus. Carapace length reaching 6.0 mm and total length about 
<quantity id="232E10BDFFCB5C6C39A64580C34EFE06" box="[495,574,419,443]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="8" unit="mm" value="12.0">12 mm</quantity>
in males, while females are slightly larger with carapace length reaching 7.0 mm and total length about 
<quantity id="232E10BDFFCB5C6C398D45E7C366FE60" box="[452,534,452,476]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" pageId="8" unit="mm" value="14.0">14 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="ACCCEED3FFCB5C6C388E45C6C016FD0E" pageId="8" type="biology_ecology">
<paragraph id="E469BD58FFCB5C6C388E45C6C016FD0E" blockId="8.[151,1437,151,2022]" pageId="8">
<emphasis id="D6A2614AFFCB5C6C388E45C6C05BFE42" bold="true" box="[199,299,485,510]" pageId="8">Ecology.</emphasis>
All described specimens were collected from burrows of large snapping shrimp 
<taxonomicName id="23D6C6DBFFCB5C6C3D0B45C6C040FD9D" authorityName="De Haan" authorityYear="1844" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="8" phylum="Arthropoda" rank="species" species="brevicristatus">
<emphasis id="D6A2614AFFCB5C6C3D0B45C6C040FD9D" italics="true" pageId="8">Alpheus brevicristatus</emphasis>
</taxonomicName>
built at depths of 
<quantity id="232E10BDFFCB5C6C39B4462AC334FD9D" box="[509,580,521,545]" metricMagnitude="0" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="8" unit="m" value="1.5" valueMax="2.0" valueMin="1.0">1–2 m</quantity>
into muddy sand among 
<taxonomicName id="23D6C6DBFFCB5C6C3B2A4629C2C9FD9D" authorityName="Linnaeus" authorityYear="1753" box="[867,953,522,545]" class="Liliopsida" family="Zosteraceae" genus="Zostera" kingdom="Plantae" order="Alismatales" pageId="8" phylum="Tracheophyta" rank="genus">
<emphasis id="D6A2614AFFCB5C6C3B2A4629C2C9FD9D" box="[867,953,522,545]" italics="true" pageId="8">Zostera</emphasis>
</taxonomicName>
beds. The species appears to be associated with 
<taxonomicName id="23D6C6DBFFCB5C6C389D460DC0FFFDF9" authorityName="De Haan" authorityYear="1844" box="[212,399,557,581]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="8" phylum="Arthropoda" rank="species" species="brevicristatus">
<emphasis id="D6A2614AFFCB5C6C389D460DC0FFFDF9" box="[212,399,557,581]" italics="true" pageId="8">A. brevicristatus</emphasis>
</taxonomicName>
as it has never been found outside of the host burrows at the 
<typeStatus id="3B6D03FAFFCB5C6C3C3A460DC5D3FDFA" box="[1139,1187,558,582]" pageId="8">type</typeStatus>
locality or any other investigated areas along Russian coast of the Sea of 
<collectingCountry id="9CC1FDC8FFCB5C6C3B454671C23FFDD5" box="[780,847,594,617]" name="Japan" pageId="8">Japan</collectingCountry>
. Within the burrows of 
<taxonomicName id="23D6C6DBFFCB5C6C3C394671C45DFDD5" authorityName="De Haan" authorityYear="1844" box="[1136,1325,593,617]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="8" phylum="Arthropoda" rank="species" species="brevicristatus">
<emphasis id="D6A2614AFFCB5C6C3C394671C45DFDD5" box="[1136,1325,593,617]" italics="true" pageId="8">A. brevicristatus</emphasis>
</taxonomicName>
, 
<taxonomicName id="23D6C6DBFFCB5C6C3D094672C031FD31" authority="Marin, 2017" authorityName="Marin" authorityYear="2017" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="8" phylum="Arthropoda" rank="species" species="alpheusophilus" status="sp. nov.">
<emphasis id="D6A2614AFFCB5C6C3D094672C031FD31" italics="true" pageId="8">Athanas alpheusophilus</emphasis>
</taxonomicName>
<emphasis id="D6A2614AFFCB5C6C390E4656C0D0FD32" bold="true" box="[327,416,629,654]" pageId="8">
<taxonomicNameLabel id="CD91DC31FFCB5C6C390E4656C0D0FD32" box="[327,416,629,654]" pageId="8" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
lives in small groups composed of several ovigerous and non-ovigerous females and one or two mature males.
</paragraph>
</subSubSection>
<subSubSection id="ACCCEED3FFCB5C6C388E469EC43FFD46" pageId="8" type="etymology">
<paragraph id="E469BD58FFCB5C6C388E469EC43FFD46" blockId="8.[151,1437,151,2022]" pageId="8">
<emphasis id="D6A2614AFFCB5C6C388E469EC03EFD6A" bold="true" box="[199,334,701,726]" pageId="8">Etymology.</emphasis>
The specific name refers to the association with larger host alpheid shrimp of the genus 
<taxonomicName id="23D6C6DBFFCB5C6C3D0B469EC037FD46" authority="Fabricius, 1798" authorityName="Fabricius" authorityYear="1798" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="8" phylum="Arthropoda" rank="genus">
<emphasis id="D6A2614AFFCB5C6C3D0B469EC4ECFD69" box="[1346,1436,701,725]" italics="true" pageId="8">Alpheus</emphasis>
Fabricius, 1798
</taxonomicName>
, which burrows are inhabited by the new species; from 
<taxonomicName id="23D6C6DBFFCB5C6C3BF146C2C562FD45" authorityName="Fabricius" authorityYear="1798" box="[952,1042,737,761]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="8" phylum="Arthropoda" rank="genus">
<emphasis id="D6A2614AFFCB5C6C3BF146C2C562FD45" box="[952,1042,737,761]" italics="true" pageId="8">Alpheus</emphasis>
</taxonomicName>
+ 
<emphasis id="D6A2614AFFCB5C6C3C7946C2C505FD45" box="[1072,1141,737,761]" italics="true" pageId="8">philus</emphasis>
(Greek for 
<emphasis id="D6A2614AFFCB5C6C3CB146C2C44EFD45" box="[1272,1342,737,761]" italics="true" pageId="8">loving</emphasis>
).
</paragraph>
</subSubSection>
<subSubSection id="ACCCEED3FFCB5C6C388E4726C58BFA86" pageId="8" type="distribution">
<paragraph id="E469BD58FFCB5C6C388E4726C5EFFCDA" blockId="8.[151,1437,151,2022]" pageId="8">
<emphasis id="D6A2614AFFCB5C6C388E4726C010FCA2" bold="true" box="[199,352,773,798]" pageId="8">Distribution.</emphasis>
Presently known only from the 
<typeStatus id="3B6D03FAFFCB5C6C3A9D4725C274FCA2" box="[724,772,774,798]" pageId="8">type</typeStatus>
locality in Vityaz Bay (
<geoCoordinate id="81E2DB9FFFCB5C6C3C504726C5E1FCA2" box="[1049,1169,773,798]" direction="north" orientation="latitude" pageId="8" precision="15" value="42.601944">42°36′7″N</geoCoordinate>
<geoCoordinate id="81E2DB9FFFCB5C6C3CD34726C45EFCA2" box="[1178,1326,773,798]" direction="east" orientation="longitude" pageId="8" precision="15" value="131.17862">131°10′43″E</geoCoordinate>
), Russian coast of the Sea of 
<collectingCountry id="9CC1FDC8FFCB5C6C39264709C0C2FCFD" box="[367,434,810,833]" name="Japan" pageId="8">Japan</collectingCountry>
. The species probably belongs to cold current living fauna of the Sea of 
<collectingCountry id="9CC1FDC8FFCB5C6C3CAA4709C453FCFD" box="[1251,1315,810,833]" name="Japan" pageId="8">Japan</collectingCountry>
inhabiting northern mainland coast influenced by Liman cold current from the Sea of Okhotsk (
<figureCitation id="7CEDA1DDFFCB5C6C3C04476EC5E1FCDA" box="[1101,1169,845,870]" captionStart="FIGURE 7" captionStartId="9.[151,250,1571,1593]" captionTargetBox="[151,1436,628,1535]" captionTargetId="figure@9.[151,1436,628,1550]" captionTargetPageId="9" captionText="FIGURE 7. The map of records of representatives of the genus Athanas Leach, 1814 in the Sea of Japan and adjacent area. Empty sign indicates the type locality of the species." httpUri="https://zenodo.org/record/997065/files/figure.png" pageId="8">Fig. 7</figureCitation>
).
</paragraph>
<paragraph id="E469BD58FFCB5C6C388E4752C288FBDD" blockId="8.[151,1437,151,2022]" pageId="8">
<emphasis id="D6A2614AFFCB5C6C388E4752C237FC36" bold="true" box="[199,839,881,906]" pageId="8">
Taxonomic remarks. 
<taxonomicName id="23D6C6DBFFCB5C6C39814752C392FC36" authority="Marin, 2017" authorityName="Marin" authorityYear="2017" box="[456,738,881,906]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="8" phylum="Arthropoda" rank="species" species="alpheusophilus" status="sp. nov.">
<emphasis id="D6A2614AFFCB5C6C39814752C392FC36" bold="true" box="[456,738,881,906]" italics="true" pageId="8">Athanas alpheusophilus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CD91DC31FFCB5C6C3AA24752C237FC36" box="[747,839,881,906]" pageId="8" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
belongs to increasingly heterogeneous and possibly non-monophyletic “ 
<taxonomicName id="23D6C6DBFFCB5C6C393447B6C32DFC11" authorityName="Otmann" authorityYear="1894" box="[381,605,917,941]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="8" phylum="Arthropoda" rank="species" species="dimorphus">
<emphasis id="D6A2614AFFCB5C6C393447B6C32DFC11" box="[381,605,917,941]" italics="true" pageId="8">Athanas dimorphus</emphasis>
</taxonomicName>
” species group (cf. Anker &amp; Jeng 2007) characterized mainly by the general shape of enlarged chelipeds (pereiopods I), especially in males, including their ability to be carried flexed, with chela fitting in merus specially excavated for this purpose. The new species shows more close similarities with the representatives of “ 
<taxonomicName id="23D6C6DBFFCB5C6C39F94022C3F7FBA5" authorityName="Kubo" authorityYear="1936" box="[432,647,1025,1049]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="8" phylum="Arthropoda" rank="species" species="japonicus">
<emphasis id="D6A2614AFFCB5C6C39F94022C3F7FBA5" box="[432,647,1025,1049]" italics="true" pageId="8">Athanas japonicus</emphasis>
</taxonomicName>
” species group described and presently known only from 
<collectingCountry id="9CC1FDC8FFCB5C6C3D1C4021C4E8FBA5" box="[1365,1432,1026,1049]" name="Japan" pageId="8">Japan</collectingCountry>
, including 
<taxonomicName id="23D6C6DBFFCB5C6C39594006C3FAFB82" authority="Yokoya, 1936" authorityName="Yokoya" authorityYear="1936" baseAuthorityName="Yokoya" baseAuthorityYear="1936" box="[272,650,1061,1086]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="8" phylum="Arthropoda" rank="species" species="ohsimai">
<emphasis id="D6A2614AFFCB5C6C39594006C0A4FB81" box="[272,468,1061,1085]" italics="true" pageId="8">Athanas ohsimai</emphasis>
Yokoya, 1936
</taxonomicName>
, 
<taxonomicName id="23D6C6DBFFCB5C6C3AD44006C564FB82" authority="Kubo, 1940" authorityName="Kubo" authorityYear="1940" box="[669,1044,1061,1086]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="8" phylum="Arthropoda" rank="species" species="japonicus">
<emphasis id="D6A2614AFFCB5C6C3AD44006C205FB81" box="[669,885,1061,1085]" italics="true" pageId="8">Athanas japonicus</emphasis>
<bibRefCitation id="8047C0A9FFCB5C6C3BCC4006C564FB82" author="Kubo" box="[901,1044,1061,1086]" pageId="8" refString="Kubo, I. (1940) Notes on the Japanese shrimps of the genus Athanas with a description of one new species. Annotationes Zoologicae Japonenses, 19, 99 - 106." type="journal article" year="1940">Kubo, 1940</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="23D6C6DBFFCB5C6C3C614006C4ECFB82" authority="Kubo, 1940" authorityName="Kubo" authorityYear="1940" box="[1064,1436,1061,1086]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="8" phylum="Arthropoda" rank="species" species="lamellifer">
<emphasis id="D6A2614AFFCB5C6C3C614006C470FB81" box="[1064,1280,1061,1085]" italics="true" pageId="8">Athanas lamellifer</emphasis>
<bibRefCitation id="8047C0A9FFCB5C6C3D464006C4ECFB82" author="Kubo" box="[1295,1436,1061,1086]" pageId="8" refString="Kubo, I. (1940) Notes on the Japanese shrimps of the genus Athanas with a description of one new species. Annotationes Zoologicae Japonenses, 19, 99 - 106." type="journal article" year="1940">Kubo, 1940</bibRefCitation>
</taxonomicName>
(considered as junior synonym of 
<taxonomicName id="23D6C6DBFFCB5C6C3A5D4069C3D1FBDD" authorityName="Kubo" authorityYear="1940" box="[532,673,1098,1121]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="8" phylum="Arthropoda" rank="species" species="japonicus">
<emphasis id="D6A2614AFFCB5C6C3A5D4069C3D1FBDD" box="[532,673,1098,1121]" italics="true" pageId="8">A. japonicus</emphasis>
</taxonomicName>
(after 
<bibRefCitation id="8047C0A9FFCB5C6C3AA2406AC2AFFBDE" author="Miya" box="[747,991,1097,1122]" pageId="8" refString="Miya, Y. &amp; Miyake, S. (1968) Revision of the genus Athanas of Japan and the Ryukyu Islands, with description of a new species (Crustacea, Decapoda, Alpheidae). Publications from the Amakusa Marine Biology Laboratory, 1, 129 - 162." type="journal article" year="1968">Miya &amp; Miyake 1968</bibRefCitation>
)).
</paragraph>
<paragraph id="E469BD58FFCB5C6C388E404DC58BFA86" blockId="8.[151,1437,151,2022]" pageId="8">
At the same time, the new species can be clearly separated from 
<taxonomicName id="23D6C6DBFFCB5C6C3BFF404DC541FB39" authorityName="Yokoya" authorityYear="1936" baseAuthorityName="Yokoya" baseAuthorityYear="1936" box="[950,1073,1133,1157]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="8" phylum="Arthropoda" rank="species" species="ohsimai">
<emphasis id="D6A2614AFFCB5C6C3BFF404DC541FB39" box="[950,1073,1133,1157]" italics="true" pageId="8">A. ohsimai</emphasis>
</taxonomicName>
by longer rostrum reaching the distal antennular segment (vs. reaching the midlength of second antennular segment in 
<taxonomicName id="23D6C6DBFFCB5C6C3C2C40B1C4E8FB16" authority="(Yokoya 1936))" authorityName="Yokoya" authorityYear="1936" baseAuthorityName="Yokoya" baseAuthorityYear="1936" box="[1125,1432,1169,1194]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="8" phylum="Arthropoda" rank="species" species="ohsimai">
<emphasis id="D6A2614AFFCB5C6C3C2C40B1C5ADFB15" box="[1125,1245,1169,1193]" italics="true" pageId="8">A. ohsimai</emphasis>
(Yokoya 1936))
</taxonomicName>
, different proportions in pereiopod I with longer carpus both in males and females (carpus of pereiopod I about half of propodal length in males (
<figureCitation id="7CEDA1DDFFCB5C6C399240FAC344FB4D" box="[475,564,1241,1266]" captionStart="FIGURE 3" captionStartId="4.[151,250,1952,1974]" captionTargetBox="[168,1430,207,1912]" captionTargetId="figure@4.[151,1436,193,1931]" captionTargetPageId="4" captionText="FIGURE 3. Athanas alpheusophillus sp. nov., Vitjaz, Posjeta Bay, the Sea of Japan, paratypes (a, c, d — ovigerous female; b, e — male): a, b — antennula; c, e — pereiopods I (chelipeds); d — distal propodus and chela of pereiopod I." httpUri="https://zenodo.org/record/997057/files/figure.png" pageId="8">
Figs. 3 
<emphasis id="D6A2614AFFCB5C6C3A6140F9C344FB4D" box="[552,564,1242,1265]" italics="true" pageId="8">e</emphasis>
</figureCitation>
, 
<figureCitation id="7CEDA1DDFFCB5C6C3A0A40F9C30BFB4D" box="[579,635,1241,1266]" captionStart="FIGURE 4" captionStartId="5.[151,250,1983,2005]" captionTargetBox="[169,1414,218,1948]" captionTargetId="figure@5.[161,1425,193,1962]" captionTargetPageId="5" captionText="FIGURE 4. Athanas alpheusophillus sp. nov., Vitjaz, Posjeta Bay, the Sea of Japan, paratypes (a – c — male; d, e — female): a, b — minor pereiopod I; c — distal propodus and fingers of pereiopod I; d — pereiopod II; e — pereiopod III." httpUri="https://zenodo.org/record/997059/files/figure.png" pageId="8">
4 
<emphasis id="D6A2614AFFCB5C6C3A1840F9C30BFB4D" box="[593,635,1241,1265]" italics="true" pageId="8">a, b</emphasis>
</figureCitation>
, 
<figureCitation id="7CEDA1DDFFCB5C6C3AC040FAC3B1FB4D" box="[649,705,1241,1266]" captionStart="FIGURE 5" captionStartId="6.[151,250,1931,1953]" captionTargetBox="[208,1378,220,1888]" captionTargetId="figure@6.[195,1391,211,1899]" captionTargetPageId="6" captionText="FIGURE 5. Athanas alpheusophillus sp. nov., Vitjaz, Posjeta Bay, the Sea of Japan, paratype male, major pereiopod I: a — general view of pereiopod I; b — distal segments of pereiopod I; c, d — distal propodus and fingers of pereiopod I, different views." httpUri="https://zenodo.org/record/997061/files/figure.png" pageId="8">
5 
<emphasis id="D6A2614AFFCB5C6C3ADE40F9C3B1FB4D" box="[663,705,1241,1265]" italics="true" pageId="8">a, b</emphasis>
</figureCitation>
) and equal to chela length in females (
<figureCitation id="7CEDA1DDFFCB5C6C3C3C40FAC5B3FB4D" box="[1141,1219,1241,1266]" captionStart="FIGURE 3" captionStartId="4.[151,250,1952,1974]" captionTargetBox="[168,1430,207,1912]" captionTargetId="figure@4.[151,1436,193,1931]" captionTargetPageId="4" captionText="FIGURE 3. Athanas alpheusophillus sp. nov., Vitjaz, Posjeta Bay, the Sea of Japan, paratypes (a, c, d — ovigerous female; b, e — male): a, b — antennula; c, e — pereiopods I (chelipeds); d — distal propodus and chela of pereiopod I." httpUri="https://zenodo.org/record/997057/files/figure.png" pageId="8">
Fig. 3 
<emphasis id="D6A2614AFFCB5C6C3CFE40F9C5B3FB4D" box="[1207,1219,1242,1265]" italics="true" pageId="8">c</emphasis>
</figureCitation>
)) (vs. carpus about 1/3 of propodal length in males and markedly shorter than chela in females of 
<taxonomicName id="23D6C6DBFFCB5C6C3C5840DDC5FAFAA9" authorityName="Yokoya" authorityYear="1936" baseAuthorityName="Yokoya" baseAuthorityYear="1936" box="[1041,1162,1277,1301]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="8" phylum="Arthropoda" rank="species" species="ohsimai">
<emphasis id="D6A2614AFFCB5C6C3C5840DDC5FAFAA9" box="[1041,1162,1277,1301]" italics="true" pageId="8">A. ohsimai</emphasis>
</taxonomicName>
(Yokoya 1936: 
<figureCitation id="7CEDA1DDFFCB5C6C3D0940DEC1D9FA86" captionStart="FIGURE 1" captionStartId="1.[151,250,1956,1978]" captionTargetBox="[169,1422,953,1927]" captionTargetId="figure@1.[151,1436,937,1932]" captionTargetPageId="1" captionText="FIGURE 1. Athanas alpheusophillus sp. nov., Vitjaz, Posjeta Bay, the Sea of Japan, general view of paratype female (a) and holotype male (ZMMU Ma 5839) (b)." httpUri="https://zenodo.org/record/997053/files/figure.png" pageId="8">Fig. 1B, L</figureCitation>
) and different form of dactylus with marked dorsal crest (
<figureCitation id="7CEDA1DDFFCB5C6C3B7B4102C2FBFA85" box="[818,907,1313,1338]" captionStart="FIGURE 3" captionStartId="4.[151,250,1952,1974]" captionTargetBox="[168,1430,207,1912]" captionTargetId="figure@4.[151,1436,193,1931]" captionTargetPageId="4" captionText="FIGURE 3. Athanas alpheusophillus sp. nov., Vitjaz, Posjeta Bay, the Sea of Japan, paratypes (a, c, d — ovigerous female; b, e — male): a, b — antennula; c, e — pereiopods I (chelipeds); d — distal propodus and chela of pereiopod I." httpUri="https://zenodo.org/record/997057/files/figure.png" pageId="8">
Figs. 3 
<emphasis id="D6A2614AFFCB5C6C3B364101C2FBFA85" box="[895,907,1314,1337]" italics="true" pageId="8">e</emphasis>
</figureCitation>
, 
<figureCitation id="7CEDA1DDFFCB5C6C3BD34101C2A0FA85" box="[922,976,1314,1338]" captionStart="FIGURE 4" captionStartId="5.[151,250,1983,2005]" captionTargetBox="[169,1414,218,1948]" captionTargetId="figure@5.[161,1425,193,1962]" captionTargetPageId="5" captionText="FIGURE 4. Athanas alpheusophillus sp. nov., Vitjaz, Posjeta Bay, the Sea of Japan, paratypes (a – c — male; d, e — female): a, b — minor pereiopod I; c — distal propodus and fingers of pereiopod I; d — pereiopod II; e — pereiopod III." httpUri="https://zenodo.org/record/997059/files/figure.png" pageId="8">
4 
<emphasis id="D6A2614AFFCB5C6C3BEE4101C2A0FA85" box="[935,976,1314,1337]" italics="true" pageId="8">a–c</emphasis>
</figureCitation>
, 
<figureCitation id="7CEDA1DDFFCB5C6C3B974102C566FA85" box="[990,1046,1313,1338]" captionStart="FIGURE 5" captionStartId="6.[151,250,1931,1953]" captionTargetBox="[208,1378,220,1888]" captionTargetId="figure@6.[195,1391,211,1899]" captionTargetPageId="6" captionText="FIGURE 5. Athanas alpheusophillus sp. nov., Vitjaz, Posjeta Bay, the Sea of Japan, paratype male, major pereiopod I: a — general view of pereiopod I; b — distal segments of pereiopod I; c, d — distal propodus and fingers of pereiopod I, different views." httpUri="https://zenodo.org/record/997061/files/figure.png" pageId="8">
5 
<emphasis id="D6A2614AFFCB5C6C3BA54102C566FA85" box="[1004,1046,1313,1337]" italics="true" pageId="8">a–d</emphasis>
</figureCitation>
) in the new species.
</paragraph>
</subSubSection>
<subSubSection id="ACCCEED3FFCB5C6E388E4165C019FF4F" lastPageId="10" pageId="8" type="reference_group">
<paragraph id="E469BD58FFCB5C6C388E4165C1ACF9E6" blockId="8.[151,1437,151,2022]" pageId="8">
From 
<taxonomicName id="23D6C6DBFFCB5C6C39594165C0D1FAE1" ID-CoL="85MC9" authorityName="Kubo" authorityYear="1940" box="[272,417,1349,1373]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="8" phylum="Arthropoda" rank="species" species="lamellifer">
<emphasis id="D6A2614AFFCB5C6C39594165C0D1FAE1" box="[272,417,1349,1373]" italics="true" pageId="8">A. lamellifer</emphasis>
</taxonomicName>
the new species can be clearly separated by shorter antennal stylocerite not reaching midlength of second antennular segment (vs. overreaching distal margin of second antennular segment in 
<taxonomicName id="23D6C6DBFFCB5C6C3DCD4149C074FA19" ID-CoL="85MC9" authorityName="Kubo" authorityYear="1940" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="8" phylum="Arthropoda" rank="species" species="lamellifer">
<emphasis id="D6A2614AFFCB5C6C3DCD4149C074FA19" italics="true" pageId="8">A. lamellifer</emphasis>
</taxonomicName>
<bibRefCitation id="8047C0A9FFCB5C6C394241AEC37BFA1A" author="Kubo" box="[267,523,1421,1446]" pageId="8" refString="Kubo, I. (1940) Notes on the Japanese shrimps of the genus Athanas with a description of one new species. Annotationes Zoologicae Japonenses, 19, 99 - 106." type="journal article" year="1940">
(Kubo 1940: 
<figureCitation id="7CEDA1DDFFCB5C6C39EB41AEC088FA1A" box="[418,504,1421,1446]" captionStart="FIGURE 4" captionStartId="5.[151,250,1983,2005]" captionTargetBox="[169,1414,218,1948]" captionTargetId="figure@5.[161,1425,193,1962]" captionTargetPageId="5" captionText="FIGURE 4. Athanas alpheusophillus sp. nov., Vitjaz, Posjeta Bay, the Sea of Japan, paratypes (a – c — male; d, e — female): a, b — minor pereiopod I; c — distal propodus and fingers of pereiopod I; d — pereiopod II; e — pereiopod III." httpUri="https://zenodo.org/record/997059/files/figure.png" pageId="8">Fig. 4B</figureCitation>
))
</bibRefCitation>
, equal in size and similar in shape pereiopods I in males (vs. unequal in size and dissimilar pereiopod I in males of 
<taxonomicName id="23D6C6DBFFCB5C6C3A5D4191C3D0FA75" ID-CoL="85MC9" authorityName="Kubo" authorityYear="1940" box="[532,672,1457,1481]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="8" phylum="Arthropoda" rank="species" species="lamellifer">
<emphasis id="D6A2614AFFCB5C6C3A5D4191C3D0FA75" box="[532,672,1457,1481]" italics="true" pageId="8">A. lamellifer</emphasis>
</taxonomicName>
<bibRefCitation id="8047C0A9FFCB5C6C3AEE4192C2BDFA76" author="Kubo" box="[679,973,1457,1482]" pageId="8" refString="Kubo, I. (1940) Notes on the Japanese shrimps of the genus Athanas with a description of one new species. Annotationes Zoologicae Japonenses, 19, 99 - 106." type="journal article" year="1940">
(Kubo 1940: 
<figureCitation id="7CEDA1DDFFCB5C6C3B724192C2E3FA76" box="[827,915,1457,1482]" captionStart="FIGURE 4" captionStartId="5.[151,250,1983,2005]" captionTargetBox="[169,1414,218,1948]" captionTargetId="figure@5.[161,1425,193,1962]" captionTargetPageId="5" captionText="FIGURE 4. Athanas alpheusophillus sp. nov., Vitjaz, Posjeta Bay, the Sea of Japan, paratypes (a – c — male; d, e — female): a, b — minor pereiopod I; c — distal propodus and fingers of pereiopod I; d — pereiopod II; e — pereiopod III." httpUri="https://zenodo.org/record/997059/files/figure.png" pageId="8">Fig. 4D</figureCitation>
, E’))
</bibRefCitation>
, different proportions in pereiopod I with longer carpus both in males and females (
<figureCitation id="7CEDA1DDFFCB5C6C3A3F41F6C3A0FA51" box="[630,720,1493,1518]" captionStart="FIGURE 3" captionStartId="4.[151,250,1952,1974]" captionTargetBox="[168,1430,207,1912]" captionTargetId="figure@4.[151,1436,193,1931]" captionTargetPageId="4" captionText="FIGURE 3. Athanas alpheusophillus sp. nov., Vitjaz, Posjeta Bay, the Sea of Japan, paratypes (a, c, d — ovigerous female; b, e — male): a, b — antennula; c, e — pereiopods I (chelipeds); d — distal propodus and chela of pereiopod I." httpUri="https://zenodo.org/record/997057/files/figure.png" pageId="8">
Figs. 3 
<emphasis id="D6A2614AFFCB5C6C3A8D41F5C3A0FA51" box="[708,720,1494,1517]" italics="true" pageId="8">e</emphasis>
</figureCitation>
, 
<figureCitation id="7CEDA1DDFFCB5C6C3AA841F5C26AFA51" box="[737,794,1493,1518]" captionStart="FIGURE 4" captionStartId="5.[151,250,1983,2005]" captionTargetBox="[169,1414,218,1948]" captionTargetId="figure@5.[161,1425,193,1962]" captionTargetPageId="5" captionText="FIGURE 4. Athanas alpheusophillus sp. nov., Vitjaz, Posjeta Bay, the Sea of Japan, paratypes (a – c — male; d, e — female): a, b — minor pereiopod I; c — distal propodus and fingers of pereiopod I; d — pereiopod II; e — pereiopod III." httpUri="https://zenodo.org/record/997059/files/figure.png" pageId="8">
4 
<emphasis id="D6A2614AFFCB5C6C3AA641F5C26AFA51" box="[751,794,1493,1517]" italics="true" pageId="8">a, b</emphasis>
</figureCitation>
, 
<figureCitation id="7CEDA1DDFFCB5C6C3B6341F6C214FA51" box="[810,868,1493,1518]" captionStart="FIGURE 5" captionStartId="6.[151,250,1931,1953]" captionTargetBox="[208,1378,220,1888]" captionTargetId="figure@6.[195,1391,211,1899]" captionTargetPageId="6" captionText="FIGURE 5. Athanas alpheusophillus sp. nov., Vitjaz, Posjeta Bay, the Sea of Japan, paratype male, major pereiopod I: a — general view of pereiopod I; b — distal segments of pereiopod I; c, d — distal propodus and fingers of pereiopod I, different views." httpUri="https://zenodo.org/record/997061/files/figure.png" pageId="8">
5 
<emphasis id="D6A2614AFFCB5C6C3B7141F5C214FA51" box="[824,868,1493,1517]" italics="true" pageId="8">a, b</emphasis>
</figureCitation>
) (vs. carpus of pereiopod I about 1/3 of propodal length in males of 
<taxonomicName id="23D6C6DBFFCB5C6C392241D9C089F9AD" ID-CoL="85MC9" authorityName="Kubo" authorityYear="1940" box="[363,505,1529,1553]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="8" phylum="Arthropoda" rank="species" species="lamellifer">
<emphasis id="D6A2614AFFCB5C6C392241D9C089F9AD" box="[363,505,1529,1553]" italics="true" pageId="8">A. lamellifer</emphasis>
</taxonomicName>
<bibRefCitation id="8047C0A9FFCB5C6C3A4941DAC251F9AE" author="Kubo" box="[512,801,1529,1554]" pageId="8" refString="Kubo, I. (1940) Notes on the Japanese shrimps of the genus Athanas with a description of one new species. Annotationes Zoologicae Japonenses, 19, 99 - 106." type="journal article" year="1940">
(Kubo 1940: 
<figureCitation id="7CEDA1DDFFCB5C6C3AD141DAC27FF9AE" box="[664,783,1529,1554]" captionStart="FIGURE 4" captionStartId="5.[151,250,1983,2005]" captionTargetBox="[169,1414,218,1948]" captionTargetId="figure@5.[161,1425,193,1962]" captionTargetPageId="5" captionText="FIGURE 4. Athanas alpheusophillus sp. nov., Vitjaz, Posjeta Bay, the Sea of Japan, paratypes (a – c — male; d, e — female): a, b — minor pereiopod I; c — distal propodus and fingers of pereiopod I; d — pereiopod II; e — pereiopod III." httpUri="https://zenodo.org/record/997059/files/figure.png" pageId="8">Fig. 4D, E</figureCitation>
))
</bibRefCitation>
and different form of dactylus with marked dorsal crest (
<figureCitation id="7CEDA1DDFFCB5C6C38D6423EC18BF989" box="[159,251,1565,1590]" captionStart="FIGURE 3" captionStartId="4.[151,250,1952,1974]" captionTargetBox="[168,1430,207,1912]" captionTargetId="figure@4.[151,1436,193,1931]" captionTargetPageId="4" captionText="FIGURE 3. Athanas alpheusophillus sp. nov., Vitjaz, Posjeta Bay, the Sea of Japan, paratypes (a, c, d — ovigerous female; b, e — male): a, b — antennula; c, e — pereiopods I (chelipeds); d — distal propodus and chela of pereiopod I." httpUri="https://zenodo.org/record/997057/files/figure.png" pageId="8">
Figs. 3 
<emphasis id="D6A2614AFFCB5C6C38A6423DC18BF989" box="[239,251,1566,1589]" italics="true" pageId="8">e</emphasis>
</figureCitation>
, 
<figureCitation id="7CEDA1DDFFCB5C6C3944423DC033F989" box="[269,323,1566,1590]" captionStart="FIGURE 4" captionStartId="5.[151,250,1983,2005]" captionTargetBox="[169,1414,218,1948]" captionTargetId="figure@5.[161,1425,193,1962]" captionTargetPageId="5" captionText="FIGURE 4. Athanas alpheusophillus sp. nov., Vitjaz, Posjeta Bay, the Sea of Japan, paratypes (a – c — male; d, e — female): a, b — minor pereiopod I; c — distal propodus and fingers of pereiopod I; d — pereiopod II; e — pereiopod III." httpUri="https://zenodo.org/record/997059/files/figure.png" pageId="8">
4 
<emphasis id="D6A2614AFFCB5C6C3952423DC033F989" box="[283,323,1566,1589]" italics="true" pageId="8">a–c</emphasis>
</figureCitation>
, 
<figureCitation id="7CEDA1DDFFCB5C6C391D423EC0FCF989" box="[340,396,1565,1590]" captionStart="FIGURE 5" captionStartId="6.[151,250,1931,1953]" captionTargetBox="[208,1378,220,1888]" captionTargetId="figure@6.[195,1391,211,1899]" captionTargetPageId="6" captionText="FIGURE 5. Athanas alpheusophillus sp. nov., Vitjaz, Posjeta Bay, the Sea of Japan, paratype male, major pereiopod I: a — general view of pereiopod I; b — distal segments of pereiopod I; c, d — distal propodus and fingers of pereiopod I, different views." httpUri="https://zenodo.org/record/997061/files/figure.png" pageId="8">
5 
<emphasis id="D6A2614AFFCB5C6C392A423EC0FCF989" box="[355,396,1565,1589]" italics="true" pageId="8">a–d</emphasis>
</figureCitation>
) (vs. dactylus of major pereiopod I without distal crest in 
<taxonomicName id="23D6C6DBFFCB5C6C3C72423DC5BAF989" ID-CoL="85MC9" authorityName="Kubo" authorityYear="1940" box="[1083,1226,1565,1589]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="8" phylum="Arthropoda" rank="species" species="lamellifer">
<emphasis id="D6A2614AFFCB5C6C3C72423DC5BAF989" box="[1083,1226,1565,1589]" italics="true" pageId="8">A. lamellifer</emphasis>
</taxonomicName>
<bibRefCitation id="8047C0A9FFCB5C6C3C9D423EC1A8F9E6" author="Kubo" pageId="8" refString="Kubo, I. (1940) Notes on the Japanese shrimps of the genus Athanas with a description of one new species. Annotationes Zoologicae Japonenses, 19, 99 - 106." type="journal article" year="1940">
(Kubo 1940: 
<figureCitation id="7CEDA1DDFFCB5C6C3D39423EC1D7F9E6" captionStart="FIGURE 4" captionStartId="5.[151,250,1983,2005]" captionTargetBox="[169,1414,218,1948]" captionTargetId="figure@5.[161,1425,193,1962]" captionTargetPageId="5" captionText="FIGURE 4. Athanas alpheusophillus sp. nov., Vitjaz, Posjeta Bay, the Sea of Japan, paratypes (a – c — male; d, e — female): a, b — minor pereiopod I; c — distal propodus and fingers of pereiopod I; d — pereiopod II; e — pereiopod III." httpUri="https://zenodo.org/record/997059/files/figure.png" pageId="8">Fig. 4</figureCitation>
D’))
</bibRefCitation>
.
</paragraph>
<paragraph id="E469BD58FFCB5C6D388E4245C0ADFD81" blockId="8.[151,1437,151,2022]" lastBlockId="9.[151,1437,151,573]" lastPageId="9" pageId="8">
From 
<taxonomicName id="23D6C6DBFFCB5C6C39454245C356F9C2" ID-CoL="85MC7" authority="Kubo, 1936" authorityName="Kubo" authorityYear="1936" box="[268,550,1637,1662]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="8" phylum="Arthropoda" rank="species" species="japonicus">
<emphasis id="D6A2614AFFCB5C6C39454245C0E9F9C1" box="[268,409,1638,1661]" italics="true" pageId="8">A. japonicus</emphasis>
Kubo, 1936
</taxonomicName>
the new species can be clearly separated by different form and proportions of pereiopods I both in males and females as well as different body coloration. The new species possess large slightly unequal in size but similar in shape pereiopods I in males (
<figureCitation id="7CEDA1DDFFCB5C6C3B64428EC20EF979" box="[813,894,1709,1734]" captionStart="FIGURE 1" captionStartId="1.[151,250,1956,1978]" captionTargetBox="[169,1422,953,1927]" captionTargetId="figure@1.[151,1436,937,1932]" captionTargetPageId="1" captionText="FIGURE 1. Athanas alpheusophillus sp. nov., Vitjaz, Posjeta Bay, the Sea of Japan, general view of paratype female (a) and holotype male (ZMMU Ma 5839) (b)." httpUri="https://zenodo.org/record/997053/files/figure.png" pageId="8">
Fig. 1 
<emphasis id="D6A2614AFFCB5C6C3B39428EC20EF979" box="[880,894,1709,1733]" italics="true" pageId="8">b</emphasis>
</figureCitation>
) and small similar pereiopods I in females (
<figureCitation id="7CEDA1DDFFCB5C6C3D26428EC1C3F955" captionStart="FIGURE 1" captionStartId="1.[151,250,1956,1978]" captionTargetBox="[169,1422,953,1927]" captionTargetId="figure@1.[151,1436,937,1932]" captionTargetPageId="1" captionText="FIGURE 1. Athanas alpheusophillus sp. nov., Vitjaz, Posjeta Bay, the Sea of Japan, general view of paratype female (a) and holotype male (ZMMU Ma 5839) (b)." httpUri="https://zenodo.org/record/997053/files/figure.png" pageId="8">
Fig. 1 
<emphasis id="D6A2614AFFCB5C6C38EC42F1C1C3F955" box="[165,179,1746,1769]" italics="true" pageId="8">a</emphasis>
</figureCitation>
) while it was indicated in the original description of 
<taxonomicName id="23D6C6DBFFCB5C6C3B5742F1C539F956" ID-CoL="85MC7" authority="(Kubo 1936)" authorityName="Kubo" authorityYear="1936" baseAuthorityName="Kubo" baseAuthorityYear="1936" box="[798,1097,1745,1770]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="8" phylum="Arthropoda" rank="species" species="japonicus">
<emphasis id="D6A2614AFFCB5C6C3B5742F1C2DDF955" box="[798,941,1746,1769]" italics="true" pageId="8">A. japonicus</emphasis>
(Kubo 1936)
</taxonomicName>
that “ 
<emphasis id="D6A2614AFFCB5C6C3CDB42F2C5D9F88D" italics="true" pageId="8">first pair of pereiopods robust as compared with the other legs, asymmetrical in size, usually larger in the right side than the left in both sexes, in male they are stouter than in female and slightly outreach the antennal scale</emphasis>
”. Such unequal and dissimilar pereiopods I in females were also described in 
<bibRefCitation id="8047C0A9FFCB5C6C3B53431DC55BF8EA" author="Mya" box="[794,1067,1853,1878]" pageId="8" refString="Miya, Y. &amp; Miyake, S. (1968) Revision of the genus Athanas of Japan and the Ryukyu Islands, with description of a new species (Crustacea, Decapoda, Alpheidae). Publications from the Amakusa Marine Biology Laboratory, 1, 129 - 162." type="journal article" year="1968">Mya and Miyake (1968)</bibRefCitation>
as “Pattern II A–1” and “Pattern B” (see 
<bibRefCitation id="8047C0A9FFCB5C6C38B94341C0AEF8C6" author="Mya" box="[240,478,1889,1914]" pageId="8" refString="Miya, Y. &amp; Miyake, S. (1968) Revision of the genus Athanas of Japan and the Ryukyu Islands, with description of a new species (Crustacea, Decapoda, Alpheidae). Publications from the Amakusa Marine Biology Laboratory, 1, 129 - 162." type="journal article" year="1968">Mya &amp; Miyake 1968</bibRefCitation>
: 
<figureCitation id="7CEDA1DDFFCB5C6C39A14342C32DF8C6" box="[488,605,1889,1914]" captionStart="FIGURE 5" captionStartId="6.[151,250,1931,1953]" captionTargetBox="[208,1378,220,1888]" captionTargetId="figure@6.[195,1391,211,1899]" captionTargetPageId="6" captionText="FIGURE 5. Athanas alpheusophillus sp. nov., Vitjaz, Posjeta Bay, the Sea of Japan, paratype male, major pereiopod I: a — general view of pereiopod I; b — distal segments of pereiopod I; c, d — distal propodus and fingers of pereiopod I, different views." httpUri="https://zenodo.org/record/997061/files/figure.png" pageId="8">Fig. 5B, C</figureCitation>
), while pereiopods I characteristic for females of 
<emphasis id="D6A2614AFFCB5C6C3CCC4342C180F822" bold="true" pageId="8">
<taxonomicName id="23D6C6DBFFCB5C6C3CCC4342C4ECF8C6" ID-CoL="85MBK" authority="Marin, 2017" authorityName="Marin" authorityYear="2017" box="[1157,1436,1889,1914]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="8" phylum="Arthropoda" rank="species" species="alpheusophilus" status="sp. nov.">
<emphasis id="D6A2614AFFCB5C6C3CCC4342C4ECF8C6" bold="true" box="[1157,1436,1889,1914]" italics="true" pageId="8">Athanas alpheusophilus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CD91DC31FFCB5C6C38DE43A6C180F822" box="[151,240,1925,1950]" pageId="8" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
were probably indicated as “Pattern A” (
<bibRefCitation id="8047C0A9FFCB5C6C3AF743A5C564F822" author="Mya" box="[702,1044,1925,1950]" pageId="8" refString="Miya, Y. &amp; Miyake, S. (1968) Revision of the genus Athanas of Japan and the Ryukyu Islands, with description of a new species (Crustacea, Decapoda, Alpheidae). Publications from the Amakusa Marine Biology Laboratory, 1, 129 - 162." type="journal article" year="1968">
Mya &amp; Miyake 1968: 
<figureCitation id="7CEDA1DDFFCB5C6C3BF243A6C564F822" box="[955,1044,1925,1950]" captionStart="FIGURE 5" captionStartId="6.[151,250,1931,1953]" captionTargetBox="[208,1378,220,1888]" captionTargetId="figure@6.[195,1391,211,1899]" captionTargetPageId="6" captionText="FIGURE 5. Athanas alpheusophillus sp. nov., Vitjaz, Posjeta Bay, the Sea of Japan, paratype male, major pereiopod I: a — general view of pereiopod I; b — distal segments of pereiopod I; c, d — distal propodus and fingers of pereiopod I, different views." httpUri="https://zenodo.org/record/997061/files/figure.png" pageId="8">Fig. 5A</figureCitation>
</bibRefCitation>
); dactylus of pereiopods I in males is less than half of propodus while in Kubo’s description (1936: Pl. XIII: C, I) it significantly larger than half of propodal length. Such large dactylus is also characteristic for 
<taxonomicName id="23D6C6DBFFCB5C6C3B2D43EEC52EF859" ID-CoL="85MC7" baseAuthorityName="Kubo" baseAuthorityYear="1936" box="[868,1118,1997,2022]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="8" phylum="Arthropoda" rank="species" species="japonicus">
<emphasis id="D6A2614AFFCB5C6C3B2D43EEC2B0F859" box="[868,960,1997,2021]" italics="true" pageId="8">Athanas</emphasis>
cf. 
<emphasis id="D6A2614AFFCB5C6C3BB843EDC52EF859" box="[1009,1118,1998,2021]" italics="true" pageId="8">japonicus</emphasis>
</taxonomicName>
described by 
<bibRefCitation id="8047C0A9FFCB5C6D3D4A43EEC058FF0C" author="Anker" lastPageId="9" pageId="8" refString="Anker, A. (2003) Alpheid shrimps from the mangroves and mudflats of Singapore. Part I. Genera Salmoneus, Athanas and Potamalpheops, with the description of two new species (Crustacea: Decapoda: Caridea). The Raffles Bulletin of Zoology, 51, 283 - 314." type="journal article" year="2003">Anker (2003: Figs. 14–17)</bibRefCitation>
and 
<bibRefCitation id="8047C0A9FFCA5C6D392144BBC381FF0C" author="Mya" box="[360,753,151,176]" pageId="9" refString="Miya, Y. &amp; Miyake, S. (1968) Revision of the genus Athanas of Japan and the Ryukyu Islands, with description of a new species (Crustacea, Decapoda, Alpheidae). Publications from the Amakusa Marine Biology Laboratory, 1, 129 - 162." type="journal article" year="1968">
Mya and Miyake (1968: 
<figureCitation id="7CEDA1DDFFCA5C6D3AC644B4C398FF0C" box="[655,744,151,176]" captionStart="FIGURE 5" captionStartId="6.[151,250,1931,1953]" captionTargetBox="[208,1378,220,1888]" captionTargetId="figure@6.[195,1391,211,1899]" captionTargetPageId="6" captionText="FIGURE 5. Athanas alpheusophillus sp. nov., Vitjaz, Posjeta Bay, the Sea of Japan, paratype male, major pereiopod I: a — general view of pereiopod I; b — distal segments of pereiopod I; c, d — distal propodus and fingers of pereiopod I, different views." httpUri="https://zenodo.org/record/997061/files/figure.png" pageId="9">Fig. 5F</figureCitation>
)
</bibRefCitation>
. Dactylus of both male pereiopods I in males of 
<emphasis id="D6A2614AFFCA5C6D3D7244B4C0D7FF69" bold="true" pageId="9">
<taxonomicName id="23D6C6DBFFCA5C6D3D7244B4C037FF68" ID-CoL="85MBK" authority="Marin, 2017" authorityName="Marin" authorityYear="2017" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="9" phylum="Arthropoda" rank="species" species="alpheusophilus" status="sp. nov.">
<emphasis id="D6A2614AFFCA5C6D3D7244B4C037FF68" bold="true" italics="true" pageId="9">Athanas alpheusophilus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CD91DC31FFCA5C6D3904449FC0D7FF69" box="[333,423,188,213]" pageId="9" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is armed with marked dorsal crest (
<figureCitation id="7CEDA1DDFFCA5C6D3B73449FC2F9FF68" box="[826,905,188,213]" captionStart="FIGURE 3" captionStartId="4.[151,250,1952,1974]" captionTargetBox="[168,1430,207,1912]" captionTargetId="figure@4.[151,1436,193,1931]" captionTargetPageId="4" captionText="FIGURE 3. Athanas alpheusophillus sp. nov., Vitjaz, Posjeta Bay, the Sea of Japan, paratypes (a, c, d — ovigerous female; b, e — male): a, b — antennula; c, e — pereiopods I (chelipeds); d — distal propodus and chela of pereiopod I." httpUri="https://zenodo.org/record/997057/files/figure.png" pageId="9">
Fig. 3 
<emphasis id="D6A2614AFFCA5C6D3B34449EC2F9FF68" box="[893,905,189,212]" italics="true" pageId="9">e</emphasis>
</figureCitation>
, 
<figureCitation id="7CEDA1DDFFCA5C6D3BDE449FC2BEFF68" box="[919,974,188,212]" captionStart="FIGURE 4" captionStartId="5.[151,250,1983,2005]" captionTargetBox="[169,1414,218,1948]" captionTargetId="figure@5.[161,1425,193,1962]" captionTargetPageId="5" captionText="FIGURE 4. Athanas alpheusophillus sp. nov., Vitjaz, Posjeta Bay, the Sea of Japan, paratypes (a – c — male; d, e — female): a, b — minor pereiopod I; c — distal propodus and fingers of pereiopod I; d — pereiopod II; e — pereiopod III." httpUri="https://zenodo.org/record/997059/files/figure.png" pageId="9">
4 
<emphasis id="D6A2614AFFCA5C6D3BEC449EC2BEFF68" box="[933,974,189,212]" italics="true" pageId="9">a–c</emphasis>
</figureCitation>
, 
<figureCitation id="7CEDA1DDFFCA5C6D3B95449FC563FF68" box="[988,1043,188,213]" captionStart="FIGURE 5" captionStartId="6.[151,250,1931,1953]" captionTargetBox="[208,1378,220,1888]" captionTargetId="figure@6.[195,1391,211,1899]" captionTargetPageId="6" captionText="FIGURE 5. Athanas alpheusophillus sp. nov., Vitjaz, Posjeta Bay, the Sea of Japan, paratype male, major pereiopod I: a — general view of pereiopod I; b — distal segments of pereiopod I; c, d — distal propodus and fingers of pereiopod I, different views." httpUri="https://zenodo.org/record/997061/files/figure.png" pageId="9">
5 
<emphasis id="D6A2614AFFCA5C6D3BA3449FC563FF68" box="[1002,1043,188,212]" italics="true" pageId="9">a–d</emphasis>
</figureCitation>
) also referred by 
<bibRefCitation id="8047C0A9FFCA5C6D3C95449EC191FF44" author="Mya" pageId="9" refString="Miya, Y. &amp; Miyake, S. (1968) Revision of the genus Athanas of Japan and the Ryukyu Islands, with description of a new species (Crustacea, Decapoda, Alpheidae). Publications from the Amakusa Marine Biology Laboratory, 1, 129 - 162." type="journal article" year="1968">Mya and Miyake (1968)</bibRefCitation>
as “Pattern IV B” (
<bibRefCitation id="8047C0A9FFCA5C6D39F044C3C27BFF44" author="Mya" box="[441,779,223,248]" pageId="9" refString="Miya, Y. &amp; Miyake, S. (1968) Revision of the genus Athanas of Japan and the Ryukyu Islands, with description of a new species (Crustacea, Decapoda, Alpheidae). Publications from the Amakusa Marine Biology Laboratory, 1, 129 - 162." type="journal article" year="1968">
Mya &amp; Miyake 1968: 
<figureCitation id="7CEDA1DDFFCA5C6D3AFA44FCC27BFF44" box="[691,779,223,248]" captionStart="FIGURE 5" captionStartId="6.[151,250,1931,1953]" captionTargetBox="[208,1378,220,1888]" captionTargetId="figure@6.[195,1391,211,1899]" captionTargetPageId="6" captionText="FIGURE 5. Athanas alpheusophillus sp. nov., Vitjaz, Posjeta Bay, the Sea of Japan, paratype male, major pereiopod I: a — general view of pereiopod I; b — distal segments of pereiopod I; c, d — distal propodus and fingers of pereiopod I, different views." httpUri="https://zenodo.org/record/997061/files/figure.png" pageId="9">Fig. 5G</figureCitation>
</bibRefCitation>
). At the same time, such significant feature is missing both in the description and figures given by Kubo (1936: pl. XIII: C, H, I) for 
<taxonomicName id="23D6C6DBFFCA5C6D3BBD4527C5F3FEA0" ID-CoL="85MC7" baseAuthorityName="Kubo" baseAuthorityYear="1936" box="[1012,1155,260,284]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="9" phylum="Arthropoda" rank="species" species="japonicus">
<emphasis id="D6A2614AFFCA5C6D3BBD4527C5F3FEA0" box="[1012,1155,260,284]" italics="true" pageId="9">A. japonicus</emphasis>
</taxonomicName>
. Pattern of pereiopods I similar to the original description of 
<taxonomicName id="23D6C6DBFFCA5C6D3A7A450BC3B0FE83" ID-CoL="85MC7" baseAuthorityName="Kubo" baseAuthorityYear="1936" box="[563,704,296,319]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="9" phylum="Arthropoda" rank="species" species="japonicus">
<emphasis id="D6A2614AFFCA5C6D3A7A450BC3B0FE83" box="[563,704,296,319]" italics="true" pageId="9">A. japonicus</emphasis>
</taxonomicName>
(Kubo, 1936: Pl. XIII C, H, I) is probably presented by 
<bibRefCitation id="8047C0A9FFCA5C6D3D70450BC0E8FED8" author="Mya" pageId="9" refString="Miya, Y. &amp; Miyake, S. (1968) Revision of the genus Athanas of Japan and the Ryukyu Islands, with description of a new species (Crustacea, Decapoda, Alpheidae). Publications from the Amakusa Marine Biology Laboratory, 1, 129 - 162." type="journal article" year="1968">Mya and Miyake (1968: Fig. F)</bibRefCitation>
as “Pattern IV A”. Thus, it is possible to conclude that combination of small equal-andsimilar pereiopods I in females and large unequal-and-similar pereiopods I armed with large dorsal dactylar crest in males characteristic for 
<emphasis id="D6A2614AFFCA5C6D39FA45B0C24BFE11" bold="true" box="[435,827,403,429]" pageId="9">
<taxonomicName id="23D6C6DBFFCA5C6D39FA45B0C3A0FE10" ID-CoL="85MBK" authority="Marin, 2017" authorityName="Marin" authorityYear="2017" box="[435,720,403,428]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="9" phylum="Arthropoda" rank="species" species="alpheusophilus" status="sp. nov.">
<emphasis id="D6A2614AFFCA5C6D39FA45B0C3A0FE10" bold="true" box="[435,720,403,428]" italics="true" pageId="9">Athanas alpheusophilus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CD91DC31FFCA5C6D3A9545B7C24BFE11" box="[732,827,404,429]" pageId="9" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
show a unique pattern within “ 
<taxonomicName id="23D6C6DBFFCA5C6D3CF345B7C4FFFE10" ID-CoL="85MC7" baseAuthorityName="Kubo" baseAuthorityYear="1936" box="[1210,1423,404,428]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="9" phylum="Arthropoda" rank="species" species="japonicus">
<emphasis id="D6A2614AFFCA5C6D3CF345B7C4FFFE10" box="[1210,1423,404,428]" italics="true" pageId="9">Athanas japonicus</emphasis>
</taxonomicName>
” species complex. One more feature allowing separating the new species and 
<taxonomicName id="23D6C6DBFFCA5C6D3C4C459BC5E3FE73" ID-CoL="85MC7" baseAuthorityName="Kubo" baseAuthorityYear="1936" box="[1029,1171,440,463]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="9" phylum="Arthropoda" rank="species" species="japonicus">
<emphasis id="D6A2614AFFCA5C6D3C4C459BC5E3FE73" box="[1029,1171,440,463]" italics="true" pageId="9">A. japonicus</emphasis>
</taxonomicName>
is the body coloration. Kubo (1936) mentioned that body coloration of 
<taxonomicName id="23D6C6DBFFCA5C6D3AF145FFC235FE48" ID-CoL="85MC7" baseAuthorityName="Kubo" baseAuthorityYear="1936" box="[696,837,476,500]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="9" phylum="Arthropoda" rank="species" species="japonicus">
<emphasis id="D6A2614AFFCA5C6D3AF145FFC235FE48" box="[696,837,476,500]" italics="true" pageId="9">A. japonicus</emphasis>
</taxonomicName>
is “ 
<emphasis id="D6A2614AFFCA5C6D3B3D45FFC291FE48" box="[884,993,476,500]" italics="true" pageId="9">deep blue</emphasis>
” (Kubo 1936: 46) that is characteristic for many 
<taxonomicName id="23D6C6DBFFCA5C6D39594623C01CFDA4" ID-CoL="84LMQ" authorityName="Leach" authorityYear="1814" box="[272,364,512,536]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="9" phylum="Arthropoda" rank="genus">
<emphasis id="D6A2614AFFCA5C6D39594623C01CFDA4" box="[272,364,512,536]" italics="true" pageId="9">Athanas</emphasis>
</taxonomicName>
species (e.g. 
<bibRefCitation id="8047C0A9FFCA5C6D3A5F45DCC3D5FDA4" author="Anker" box="[534,677,511,536]" pageId="9" refString="Anker, A. (2003) Alpheid shrimps from the mangroves and mudflats of Singapore. Part I. Genera Salmoneus, Athanas and Potamalpheops, with the description of two new species (Crustacea: Decapoda: Caridea). The Raffles Bulletin of Zoology, 51, 283 - 314." type="journal article" year="2003">Anker 2003</bibRefCitation>
), while body coloration of 
<emphasis id="D6A2614AFFCA5C6D3BBB45DCC40CFDA4" bold="true" box="[1010,1404,511,536]" pageId="9">
<taxonomicName id="23D6C6DBFFCA5C6D3BBB45DCC47FFDA4" ID-CoL="85MBK" authority="Marin, 2017" authorityName="Marin" authorityYear="2017" box="[1010,1295,511,536]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="9" phylum="Arthropoda" rank="species" species="alpheusophilus" status="sp. nov.">
<emphasis id="D6A2614AFFCA5C6D3BBB45DCC47FFDA4" bold="true" box="[1010,1295,511,536]" italics="true" pageId="9">Athanas alpheusophilus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CD91DC31FFCA5C6D3D5545DCC40CFDA4" box="[1308,1404,511,536]" pageId="9" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is generally deeply red (
<figureCitation id="7CEDA1DDFFCA5C6D39C24607C0BFFD81" box="[395,463,548,573]" captionStart="FIGURE 6" captionStartId="7.[151,250,1924,1946]" captionTargetBox="[151,1436,193,1902]" captionTargetId="figure@7.[151,1436,193,1902]" captionTargetPageId="7" captionText="FIGURE 6. Athanas alpheusophillus sp. nov., Vitjaz, Posjeta Bay, the Sea of Japan, alive coloration of female paratype (a, b) and male holotype (ZMMU Ma 5839) (c)." httpUri="https://zenodo.org/record/997063/files/figure.png" pageId="9">Fig. 6</figureCitation>
).
</paragraph>
<caption id="B0A9EDD0FFCA5C6D38DE4200C3DCF9E4" httpUri="https://zenodo.org/record/997065/files/figure.png" pageId="9" targetBox="[151,1436,628,1535]" targetPageId="9">
<paragraph id="E469BD58FFCA5C6D38DE4200C3DCF9E4" blockId="9.[151,1437,1571,1624]" pageId="9">
<emphasis id="D6A2614AFFCA5C6D38DE4200C07EF985" bold="true" box="[151,270,1571,1593]" pageId="9">FIGURE 7</emphasis>
. The map of records of representatives of the genus 
<taxonomicName id="23D6C6DBFFCA5C6D3B7A4200C57DF985" ID-CoL="84LMQ" authority="Leach, 1814" authorityName="Leach" authorityYear="1814" box="[819,1037,1571,1593]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="9" phylum="Arthropoda" rank="genus">
<emphasis id="D6A2614AFFCA5C6D3B7A4200C2F6F985" box="[819,902,1571,1593]" italics="true" pageId="9">Athanas</emphasis>
Leach, 1814
</taxonomicName>
in the Sea of Japan and adjacent area. Empty sign indicates the type locality of the species.
</paragraph>
</caption>
<paragraph id="E469BD58FFCA5C6E388E42BCC019FF4F" blockId="9.[151,1436,1693,2019]" lastBlockId="10.[151,1436,151,493]" lastPageId="10" pageId="9">
It is also worth noting that proportions of pereiopods I in males of 
<emphasis id="D6A2614AFFCA5C6D3BE342BEC451F90B" bold="true" box="[938,1313,1693,1719]" pageId="9">
<taxonomicName id="23D6C6DBFFCA5C6D3BE342BEC5B0F90A" ID-CoL="85MBK" authority="Marin, 2017" authorityName="Marin" authorityYear="2017" box="[938,1216,1693,1718]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="9" phylum="Arthropoda" rank="species" species="alpheusophilus" status="sp. nov.">
<emphasis id="D6A2614AFFCA5C6D3BE342BEC5B0F90A" bold="true" box="[938,1216,1693,1718]" italics="true" pageId="9">Athanas alpheusophilus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CD91DC31FFCA5C6D3C8E42BDC451F90B" box="[1223,1313,1694,1719]" pageId="9" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
are mostly similar to 
<taxonomicName id="23D6C6DBFFCA5C6D394242E3C0E9F96B" ID-CoL="85MC7" baseAuthorityName="Kubo" baseAuthorityYear="1936" box="[267,409,1728,1751]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="9" phylum="Arthropoda" rank="species" species="japonicus">
<emphasis id="D6A2614AFFCA5C6D394242E3C0E9F96B" box="[267,409,1728,1751]" italics="true" pageId="9">A. japonicus</emphasis>
</taxonomicName>
reported by Kubo (1936) and clearly differs from 
<taxonomicName id="23D6C6DBFFCA5C6D3B9242E3C524F96B" ID-CoL="85MCM" authorityName="Yokoya" authorityYear="1936" baseAuthorityName="Yokoya" baseAuthorityYear="1936" box="[987,1108,1727,1751]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="9" phylum="Arthropoda" rank="species" species="ohsimai">
<emphasis id="D6A2614AFFCA5C6D3B9242E3C524F96B" box="[987,1108,1727,1751]" italics="true" pageId="9">A. ohsimai</emphasis>
</taxonomicName>
described by Yokoya (1936: 
<figureCitation id="7CEDA1DDFFCA5C6D38DE42C3C07CF945" box="[151,268,1760,1785]" captionStart="FIGURE 1" captionStartId="1.[151,250,1956,1978]" captionTargetBox="[169,1422,953,1927]" captionTargetId="figure@1.[151,1436,937,1932]" captionTargetPageId="1" captionText="FIGURE 1. Athanas alpheusophillus sp. nov., Vitjaz, Posjeta Bay, the Sea of Japan, general view of paratype female (a) and holotype male (ZMMU Ma 5839) (b)." httpUri="https://zenodo.org/record/997053/files/figure.png" pageId="9">Fig. 1B, L</figureCitation>
), 
<taxonomicName id="23D6C6DBFFCA5C6D396942C2C0DCF945" ID-CoL="85MC9" authorityName="Kubo" authorityYear="1940" box="[288,428,1761,1785]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="9" phylum="Arthropoda" rank="species" species="lamellifer">
<emphasis id="D6A2614AFFCA5C6D396942C2C0DCF945" box="[288,428,1761,1785]" italics="true" pageId="9">A. lamellifer</emphasis>
</taxonomicName>
(after 
<bibRefCitation id="8047C0A9FFCA5C6D39BC42C3C306F945" author="Kubo" box="[501,630,1760,1785]" pageId="9" refString="Kubo, I. (1940) Notes on the Japanese shrimps of the genus Athanas with a description of one new species. Annotationes Zoologicae Japonenses, 19, 99 - 106." type="journal article" year="1940">Kubo 1940</bibRefCitation>
: 
<figureCitation id="7CEDA1DDFFCA5C6D3AC842C3C386F945" box="[641,758,1760,1785]" captionStart="FIGURE 4" captionStartId="5.[151,250,1983,2005]" captionTargetBox="[169,1414,218,1948]" captionTargetId="figure@5.[161,1425,193,1962]" captionTargetPageId="5" captionText="FIGURE 4. Athanas alpheusophillus sp. nov., Vitjaz, Posjeta Bay, the Sea of Japan, paratypes (a – c — male; d, e — female): a, b — minor pereiopod I; c — distal propodus and fingers of pereiopod I; d — pereiopod II; e — pereiopod III." httpUri="https://zenodo.org/record/997059/files/figure.png" pageId="9">Fig. 4D, E</figureCitation>
) and 
<taxonomicName id="23D6C6DBFFCA5C6D3B7D42C2C51AF945" ID-CoL="85MC7" authority="Kubo" authorityName="Kubo" authorityYear="1936" box="[820,1130,1760,1785]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="9" phylum="Arthropoda" rank="species" species="japonicus">
<emphasis id="D6A2614AFFCA5C6D3B7D42C2C2E0F945" box="[820,912,1761,1785]" italics="true" pageId="9">Athanas</emphasis>
cf. 
<emphasis id="D6A2614AFFCA5C6D3BF042C2C556F944" box="[953,1062,1761,1784]" italics="true" pageId="9">japonicus</emphasis>
Kubo
</taxonomicName>
described by 
<bibRefCitation id="8047C0A9FFCA5C6D3D4F42C3C058F8A6" author="Anker" pageId="9" refString="Anker, A. (2003) Alpheid shrimps from the mangroves and mudflats of Singapore. Part I. Genera Salmoneus, Athanas and Potamalpheops, with the description of two new species (Crustacea: Decapoda: Caridea). The Raffles Bulletin of Zoology, 51, 283 - 314." type="journal article" year="2003">Anker (2003: Figs. 14–17)</bibRefCitation>
. Both 
<taxonomicName id="23D6C6DBFFCA5C6D39384321C099F8A6" ID-CoL="85MCM" authorityName="Yokoya" authorityYear="1936" baseAuthorityName="Yokoya" baseAuthorityYear="1936" box="[369,489,1794,1818]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="9" phylum="Arthropoda" rank="species" species="ohsimai">
<emphasis id="D6A2614AFFCA5C6D39384321C099F8A6" box="[369,489,1794,1818]" italics="true" pageId="9">A. ohsimai</emphasis>
</taxonomicName>
and 
<taxonomicName id="23D6C6DBFFCA5C6D3A694321C3DCF8A6" ID-CoL="85MC9" authorityName="Kubo" authorityYear="1940" box="[544,684,1794,1818]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="9" phylum="Arthropoda" rank="species" species="lamellifer">
<emphasis id="D6A2614AFFCA5C6D3A694321C3DCF8A6" box="[544,684,1794,1818]" italics="true" pageId="9">A. lamellifer</emphasis>
</taxonomicName>
can be easily separated from 
<emphasis id="D6A2614AFFCA5C6D3BB24322C404F8A7" bold="true" box="[1019,1396,1793,1819]" pageId="9">
<taxonomicName id="23D6C6DBFFCA5C6D3BB24322C463F8A6" ID-CoL="85MBK" authority="Marin, 2017" authorityName="Marin" authorityYear="2017" box="[1019,1299,1793,1818]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="9" phylum="Arthropoda" rank="species" species="alpheusophilus" status="sp. nov.">
<emphasis id="D6A2614AFFCA5C6D3BB24322C463F8A6" bold="true" box="[1019,1299,1793,1818]" italics="true" pageId="9">Athanas alpheusophilus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CD91DC31FFCA5C6D3D534321C404F8A7" box="[1306,1396,1794,1819]" pageId="9" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(as well as from 
<taxonomicName id="23D6C6DBFFCA5C6D39614307C0C4F887" ID-CoL="85MC7" authorityName="Kubo" authorityYear="1936" box="[296,436,1828,1851]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="9" phylum="Arthropoda" rank="species" species="japonicus">
<emphasis id="D6A2614AFFCA5C6D39614307C0C4F887" box="[296,436,1828,1851]" italics="true" pageId="9">A. japonicus</emphasis>
</taxonomicName>
too) by significantly shorter carpus of pereiopods I in males: carpus in 
<taxonomicName id="23D6C6DBFFCA5C6D3C844307C429F887" ID-CoL="85MC7" authorityName="Kubo" authorityYear="1936" box="[1229,1369,1828,1851]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="9" phylum="Arthropoda" rank="species" species="japonicus">
<emphasis id="D6A2614AFFCA5C6D3C844307C429F887" box="[1229,1369,1828,1851]" italics="true" pageId="9">A. japonicus</emphasis>
</taxonomicName>
(after Kubo 1936) and 
<emphasis id="D6A2614AFFCA5C6D39104367C3A6F8E1" bold="true" box="[345,726,1860,1885]" pageId="9">
<taxonomicName id="23D6C6DBFFCA5C6D39104367C302F8E1" ID-CoL="85MBK" authority="Marin, 2017" authorityName="Marin" authorityYear="2017" box="[345,626,1860,1885]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="9" phylum="Arthropoda" rank="species" species="alpheusophilus" status="sp. nov.">
<emphasis id="D6A2614AFFCA5C6D39104367C302F8E1" bold="true" box="[345,626,1860,1885]" italics="true" pageId="9">Athanas alpheusophilus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CD91DC31FFCA5C6D3A334367C3A6F8E1" box="[634,726,1860,1885]" pageId="9" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="7CEDA1DDFFCA5C6D3AAF4367C247F8E1" box="[742,823,1860,1885]" captionStart="FIGURE 3" captionStartId="4.[151,250,1952,1974]" captionTargetBox="[168,1430,207,1912]" captionTargetId="figure@4.[151,1436,193,1931]" captionTargetPageId="4" captionText="FIGURE 3. Athanas alpheusophillus sp. nov., Vitjaz, Posjeta Bay, the Sea of Japan, paratypes (a, c, d — ovigerous female; b, e — male): a, b — antennula; c, e — pereiopods I (chelipeds); d — distal propodus and chela of pereiopod I." httpUri="https://zenodo.org/record/997057/files/figure.png" pageId="9">
Fig. 3 
<emphasis id="D6A2614AFFCA5C6D3B624365C247F8E1" box="[811,823,1862,1885]" italics="true" pageId="9">e</emphasis>
</figureCitation>
) accounting for at least half of propodal length while carpus is much shorter, not exceeding 1/3 of propodal length in 
<taxonomicName id="23D6C6DBFFCA5C6D3BD94345C5FCF8C2" ID-CoL="85MC7" authorityName="Kubo" authorityYear="1936" box="[912,1164,1894,1919]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="9" phylum="Arthropoda" rank="species" species="japonicus">
<emphasis id="D6A2614AFFCA5C6D3BD94345C29CF8C2" box="[912,1004,1894,1918]" italics="true" pageId="9">Athanas</emphasis>
cf. 
<emphasis id="D6A2614AFFCA5C6D3C564344C5FCF8C2" box="[1055,1164,1895,1918]" italics="true" pageId="9">japonicus</emphasis>
</taxonomicName>
(after 
<bibRefCitation id="8047C0A9FFCA5C6D3C964345C41CF8C3" author="Anker" box="[1247,1388,1894,1919]" pageId="9" refString="Anker, A. (2003) Alpheid shrimps from the mangroves and mudflats of Singapore. Part I. Genera Salmoneus, Athanas and Potamalpheops, with the description of two new species (Crustacea: Decapoda: Caridea). The Raffles Bulletin of Zoology, 51, 283 - 314." type="journal article" year="2003">Anker 2003</bibRefCitation>
), 
<taxonomicName id="23D6C6DBFFCA5C6D3DCD4345C074F823" ID-CoL="85MC9" authorityName="Kubo" authorityYear="1940" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="9" phylum="Arthropoda" rank="species" species="lamellifer">
<emphasis id="D6A2614AFFCA5C6D3DCD4345C074F823" italics="true" pageId="9">A. lamellifer</emphasis>
</taxonomicName>
(after 
<bibRefCitation id="8047C0A9FFCA5C6D391A43A4C0A3F81C" author="Kubo" box="[339,467,1927,1952]" pageId="9" refString="Kubo, I. (1940) Notes on the Japanese shrimps of the genus Athanas with a description of one new species. Annotationes Zoologicae Japonenses, 19, 99 - 106." type="journal article" year="1940">Kubo 1940</bibRefCitation>
) and 
<taxonomicName id="23D6C6DBFFCA5C6D3A5C43ABC3FFF823" ID-CoL="85MCM" authorityName="Yokoya" authorityYear="1936" baseAuthorityName="Yokoya" baseAuthorityYear="1936" box="[533,655,1927,1951]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="9" phylum="Arthropoda" rank="species" species="ohsimai">
<emphasis id="D6A2614AFFCA5C6D3A5C43ABC3FFF823" box="[533,655,1927,1951]" italics="true" pageId="9">A. ohsimai</emphasis>
</taxonomicName>
(after Yokoya 1936). Male pereiopods I with long carpus were also described by 
<bibRefCitation id="8047C0A9FFCA5C6D397D438AC33FF87D" author="Mya" box="[308,591,1960,1985]" pageId="9" refString="Miya, Y. &amp; Miyake, S. (1968) Revision of the genus Athanas of Japan and the Ryukyu Islands, with description of a new species (Crustacea, Decapoda, Alpheidae). Publications from the Amakusa Marine Biology Laboratory, 1, 129 - 162." type="journal article" year="1968">Mya and Miyake (1968)</bibRefCitation>
as “Pattern III” (
<bibRefCitation id="8047C0A9FFCA5C6D3B57438AC504F87D" author="Mya" box="[798,1140,1960,1985]" pageId="9" refString="Miya, Y. &amp; Miyake, S. (1968) Revision of the genus Athanas of Japan and the Ryukyu Islands, with description of a new species (Crustacea, Decapoda, Alpheidae). Publications from the Amakusa Marine Biology Laboratory, 1, 129 - 162." type="journal article" year="1968">Mya &amp; Miyake 1968: Fig. E</bibRefCitation>
). Thus, 
<bibRefCitation id="8047C0A9FFCA5C6D3C9C438AC191F85F" author="Mya" pageId="9" refString="Miya, Y. &amp; Miyake, S. (1968) Revision of the genus Athanas of Japan and the Ryukyu Islands, with description of a new species (Crustacea, Decapoda, Alpheidae). Publications from the Amakusa Marine Biology Laboratory, 1, 129 - 162." type="journal article" year="1968">Mya and Miyake (1968)</bibRefCitation>
synonymized 
<taxonomicName id="23D6C6DBFFCA5C6D39C243E8C36BF85E" ID-CoL="85MC9" authorityName="Kubo" authorityYear="1940" box="[395,539,1994,2018]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="9" phylum="Arthropoda" rank="species" species="lamellifer">
<emphasis id="D6A2614AFFCA5C6D39C243E8C36BF85E" box="[395,539,1994,2018]" italics="true" pageId="9">A. lamellifer</emphasis>
</taxonomicName>
and 
<taxonomicName id="23D6C6DBFFCA5C6D3A1143E8C397F85E" ID-CoL="85MC7" authorityName="Kubo" authorityYear="1936" box="[600,743,1995,2018]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="9" phylum="Arthropoda" rank="species" species="japonicus">
<emphasis id="D6A2614AFFCA5C6D3A1143E8C397F85E" box="[600,743,1995,2018]" italics="true" pageId="9">A. japonicus</emphasis>
</taxonomicName>
using this feature incorrectly and 
<taxonomicName id="23D6C6DBFFCA5C6E3C3043E9C1BFFF0C" ID-CoL="85MC9" authority="Kubo, 1940" authorityName="Kubo" authorityYear="1940" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" lastPageId="10" order="Decapoda" pageId="9" phylum="Arthropoda" rank="species" species="lamellifer">
<emphasis id="D6A2614AFFCA5C6D3C3043E9C43CF85E" box="[1145,1356,1994,2018]" italics="true" pageId="9">Athanas lamellifer</emphasis>
<bibRefCitation id="8047C0A9FFCA5C6E3D1F43E9C1BFFF0C" author="Kubo" lastPageId="10" pageId="9" refString="Kubo, I. (1940) Notes on the Japanese shrimps of the genus Athanas with a description of one new species. Annotationes Zoologicae Japonenses, 19, 99 - 106." type="journal article" year="1940">Kubo, 1940</bibRefCitation>
</taxonomicName>
could be considered as valid species until more careful and complete revision will be accomplished. Besides, some previous records of 
<taxonomicName id="23D6C6DBFFC95C6E398B449AC3E4FF6D" ID-CoL="85MC7" authorityName="Kubo" authorityYear="1936" box="[450,660,185,209]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="10" phylum="Arthropoda" rank="species" species="japonicus">
<emphasis id="D6A2614AFFC95C6E398B449AC3E4FF6D" box="[450,660,185,209]" italics="true" pageId="10">Athanas japonicus</emphasis>
</taxonomicName>
from Indo-West Pacific (e.g. 
<bibRefCitation id="8047C0A9FFC95C6E3BA74499C58DFF6D" author="Banner" box="[1006,1277,185,209]" pageId="10" refString="Banner, D. M. &amp; Banner, A. H. (1973) The alpheid shrimp of Australia. Part I. The lower genera. Records of the Australian Museum, 28 (15), 291 - 382. https: // doi. org / 10.3853 / j. 0067 - 1975.28.1973.407" type="journal article" year="1973">Banner &amp; Banner 1973</bibRefCitation>
; 
<bibRefCitation id="8047C0A9FFC95C6E3D42449AC4E5FF6D" author="Anker" box="[1291,1429,185,210]" pageId="10" refString="Anker, A. (2003) Alpheid shrimps from the mangroves and mudflats of Singapore. Part I. Genera Salmoneus, Athanas and Potamalpheops, with the description of two new species (Crustacea: Decapoda: Caridea). The Raffles Bulletin of Zoology, 51, 283 - 314." type="journal article" year="2003">Anker 2003</bibRefCitation>
) need to be revised.
</paragraph>
</subSubSection>
<subSubSection id="ACCCEED3FFC95C6E388E44DFC4E9FE51" pageId="10" type="description">
<paragraph id="E469BD58FFC95C6E388E44DFC4E9FE51" blockId="10.[151,1436,151,493]" pageId="10">
The only 
<taxonomicName id="23D6C6DBFFC95C6E397E44DFC0E3FEA8" authorityName="Leach" authorityYear="1814" box="[311,403,252,276]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="10" phylum="Arthropoda" rank="genus">
<emphasis id="D6A2614AFFC95C6E397E44DFC0E3FEA8" box="[311,403,252,276]" italics="true" pageId="10">Athanas</emphasis>
</taxonomicName>
species recorded north from the latitude of the description of the new species is 
<taxonomicName id="23D6C6DBFFC95C6E3D0944DFC0A2FE84" authority="Hayashi, 2002" authorityName="Hayashi" authorityYear="2002" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="10" phylum="Arthropoda" rank="species" species="squillophilus">
<emphasis id="D6A2614AFFC95C6E3D0944DFC058FE84" italics="true" pageId="10">Athanas squillophilus</emphasis>
<bibRefCitation id="8047C0A9FFC95C6E3967453CC0A2FE84" author="Hayashi" box="[302,466,287,312]" pageId="10" refString="Hayashi, K. - I. (2002) A new species of the genus Athanas (Decapoda, Caridea, Alpheidae) living in the burrows of a mantis shrimp. Crustaceana, 75, 395 - 403. https: // doi. org / 10.1163 / 156854002760095462" type="journal article" year="2002">Hayashi, 2002</bibRefCitation>
</taxonomicName>
, which can be clearly separated from the new species by longer rostrum reaching distal antennular segment (vs. reaching midlength of second antennular segment in A. 
<taxonomicName id="23D6C6DBFFC95C6E3C6A4567C5C4FEE0" authorityName="Hayashi" authorityYear="2002" box="[1059,1204,324,348]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="10" phylum="Arthropoda" rank="species" species="squillophilus">
<emphasis id="D6A2614AFFC95C6E3C6A4567C5C4FEE0" box="[1059,1204,324,348]" italics="true" pageId="10">squillophilus</emphasis>
</taxonomicName>
<bibRefCitation id="8047C0A9FFC95C6E3CF54567C1BBFE3C" author="Hayashi" pageId="10" refString="Hayashi, K. - I. (2002) A new species of the genus Athanas (Decapoda, Caridea, Alpheidae) living in the burrows of a mantis shrimp. Crustaceana, 75, 395 - 403. https: // doi. org / 10.1163 / 156854002760095462" type="journal article" year="2002">
(Hayashi 2002: 
<figureCitation id="7CEDA1DDFFC95C6E3D394567C1C9FE3C" captionStart="FIGURE 2" captionStartId="3.[151,250,1954,1976]" captionTargetBox="[166,1411,202,1909]" captionTargetId="figure@3.[159,1419,193,1928]" captionTargetPageId="3" captionText="FIGURE 2. Athanas alpheusophillus sp. nov., Vitjaz, Posjeta Bay, the Sea of Japan, paratypes (a, d, e – g — ovigerous female; b, c — male): a, b — front of carapace, dorsal view; c, d — front of carapace, lateral view; e — distal abdominal segments and telson, lateral view; f — telson; g — exopod of uropods." httpUri="https://zenodo.org/record/997055/files/figure.png" pageId="10">Fig. 2A</figureCitation>
))
</bibRefCitation>
as well as shorter stylocerite, different position of dorsal spines on telson and morphology of chelipeds (pereiopods I) both in males and females (see 
<bibRefCitation id="8047C0A9FFC95C6E3AF745AFC213FE18" author="Hayashi" box="[702,867,396,421]" pageId="10" refString="Hayashi, K. - I. (2002) A new species of the genus Athanas (Decapoda, Caridea, Alpheidae) living in the burrows of a mantis shrimp. Crustaceana, 75, 395 - 403. https: // doi. org / 10.1163 / 156854002760095462" type="journal article" year="2002">Hayashi 2002</bibRefCitation>
: 
<figureCitation id="7CEDA1DDFFC95C6E3B3845AFC2C8FE18" box="[881,952,396,421]" captionStart="FIGURE 3" captionStartId="4.[151,250,1952,1974]" captionTargetBox="[168,1430,207,1912]" captionTargetId="figure@4.[151,1436,193,1931]" captionTargetPageId="4" captionText="FIGURE 3. Athanas alpheusophillus sp. nov., Vitjaz, Posjeta Bay, the Sea of Japan, paratypes (a, c, d — ovigerous female; b, e — male): a, b — antennula; c, e — pereiopods I (chelipeds); d — distal propodus and chela of pereiopod I." httpUri="https://zenodo.org/record/997057/files/figure.png" pageId="10">Fig. 3</figureCitation>
). Moreover, carpus of pereiopod II in 
<taxonomicName id="23D6C6DBFFC95C6E3DCD45AFC058FE74" authorityName="Hayashi" authorityYear="2002" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="10" phylum="Arthropoda" rank="species" species="squillophilus">
<emphasis id="D6A2614AFFC95C6E3DCD45AFC058FE74" italics="true" pageId="10">A. squillophilus</emphasis>
</taxonomicName>
consist of 6 segments (
<bibRefCitation id="8047C0A9FFC95C6E3A7F458CC24DFE74" author="Hayashi" box="[566,829,431,456]" pageId="10" refString="Hayashi, K. - I. (2002) A new species of the genus Athanas (Decapoda, Caridea, Alpheidae) living in the burrows of a mantis shrimp. Crustaceana, 75, 395 - 403. https: // doi. org / 10.1163 / 156854002760095462" type="journal article" year="2002">
Hayashi 2002: 
<figureCitation id="7CEDA1DDFFC95C6E3AAC458CC24DFE74" box="[741,829,431,456]" captionStart="FIGURE 3" captionStartId="4.[151,250,1952,1974]" captionTargetBox="[168,1430,207,1912]" captionTargetId="figure@4.[151,1436,193,1931]" captionTargetPageId="4" captionText="FIGURE 3. Athanas alpheusophillus sp. nov., Vitjaz, Posjeta Bay, the Sea of Japan, paratypes (a, c, d — ovigerous female; b, e — male): a, b — antennula; c, e — pereiopods I (chelipeds); d — distal propodus and chela of pereiopod I." httpUri="https://zenodo.org/record/997057/files/figure.png" pageId="10">Fig. 3D</figureCitation>
</bibRefCitation>
) while carpus of pereiopod II consists of 5 segments (
<figureCitation id="7CEDA1DDFFC95C6E38D645F7C19FFE50" box="[159,239,468,493]" captionStart="FIGURE 4" captionStartId="5.[151,250,1983,2005]" captionTargetBox="[169,1414,218,1948]" captionTargetId="figure@5.[161,1425,193,1962]" captionTargetPageId="5" captionText="FIGURE 4. Athanas alpheusophillus sp. nov., Vitjaz, Posjeta Bay, the Sea of Japan, paratypes (a – c — male; d, e — female): a, b — minor pereiopod I; c — distal propodus and fingers of pereiopod I; d — pereiopod II; e — pereiopod III." httpUri="https://zenodo.org/record/997059/files/figure.png" pageId="10">
Fig. 4 
<emphasis id="D6A2614AFFC95C6E38A845F7C19FFE50" box="[225,239,468,492]" italics="true" pageId="10">d</emphasis>
</figureCitation>
) in the new species. By these features, 
<taxonomicName id="23D6C6DBFFC95C6E3AE045F7C229FE50" authorityName="Hayashi" authorityYear="2002" box="[681,857,468,492]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="10" phylum="Arthropoda" rank="species" species="squillophilus">
<emphasis id="D6A2614AFFC95C6E3AE045F7C229FE50" box="[681,857,468,492]" italics="true" pageId="10">A. squillophilus</emphasis>
</taxonomicName>
seems not to be closely relative to the new species.
</paragraph>
</subSubSection>
</treatment>
</document>