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<document id="A30D5EF663806F2D2CC72C94F8226EC9" ID-CLB-Dataset="25190" ID-DOI="10.11646/zootaxa.4687.1.1" ID-GBIF-Dataset="de00ca6c-2843-4ae1-a39c-a29603947612" ID-ISSN="1175-5326" ID-Zenodo-Dep="3508098" ID-ZooBank="CE73264D-C234-4B82-A634-CAD6254C5957" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1571388220410" checkinUser="plazi" docAuthor="Ho, - Z." docDate="2019" docId="6C3DA91C51BE18DEFF0EFAD1EE801A14" docLanguage="en" docName="zootaxa.4687.1.1.pdf.imf" docOrigin="Zootaxa 4687 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Pygoluciola nitescens Ho 2019, comb. nov." docType="treatment" docVersion="19" lastPageNumber="137" masterDocId="9004D164513B1856FF99FF9FEF381C47" masterDocTitle="The Luciolinae of S. E. Asia and the Australopacific region: a revisionary checklist (Coleoptera: Lampyridae) including description of three new genera and 13 new species" masterLastPageNumber="174" masterPageNumber="1" pageNumber="134" updateTime="1698760145766" updateUser="ExternalLinkService">
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<mods:title id="18E903B799F4C4C6D06637E8B11D494D">The Luciolinae of S. E. Asia and the Australopacific region: a revisionary checklist (Coleoptera: Lampyridae) including description of three new genera and 13 new species</mods:title>
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<mods:namePart id="FF7B8B667146D52AA9E9BCA926A6A8A7">Ho, - Z.</mods:namePart>
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<mods:date id="2A8F53E78E054E6B322508DB41D99F04">2019</mods:date>
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<mods:number id="9E143717CDD63DBB25548F3CFCCC92E2">2019-10-18</mods:number>
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<treatment id="6C3DA91C51BE18DEFF0EFAD1EE801A14" ID-DOI="http://doi.org/10.5281/zenodo.4688941" ID-GBIF-Taxon="159757405" ID-Zenodo-Dep="4688941" LSID="urn:lsid:plazi:treatment:6C3DA91C51BE18DEFF0EFAD1EE801A14" httpUri="http://treatment.plazi.org/id/6C3DA91C51BE18DEFF0EFAD1EE801A14" lastPageId="136" lastPageNumber="137" pageId="133" pageNumber="134">
<subSubSection id="AC8E4B8151BE18D3FF0EFAD1EDD7192E" box="[151,751,1358,1385]" pageId="133" pageNumber="134" type="nomenclature">
<paragraph id="E42B180A51BE18D3FF0EFAD1EDD7192E" blockId="133.[151,751,1358,1421]" box="[151,751,1358,1385]" pageId="133" pageNumber="134">
<heading id="BF63AF6651BE18D3FF0EFAD1EDD7192E" bold="true" box="[151,751,1358,1385]" fontSize="11" level="1" pageId="133" pageNumber="134" reason="1">
<emphasis id="D6E0C41851BE18D3FF0EFAD1EDD7192E" bold="true" box="[151,751,1358,1385]" pageId="133" pageNumber="134">
<taxonomicName id="2394638951BE18D3FF0EFAD1ED59192E" ID-CoL="8NC5L" authority="(Olivier 1903 b)" authorityName="Ho" authorityYear="2019" baseAuthorityName="Olivier" baseAuthorityYear="1903" box="[151,609,1358,1385]" class="Insecta" family="Lampyridae" genus="Pygoluciola" kingdom="Animalia" order="Coleoptera" pageId="133" pageNumber="134" phylum="Arthropoda" rank="species" species="nitescens" status="comb. nov.">
<emphasis id="D6E0C41851BE18D3FF0EFAD1EEA5192E" bold="true" box="[151,413,1358,1385]" italics="true" pageId="133" pageNumber="134">Pygoluciola nitescens</emphasis>
(
<bibRefCitation id="800565FB51BE18D3FE35FAD0ED61192E" author="Olivier" box="[428,601,1359,1385]" firstAuthor="Olivier" pageId="133" pageNumber="134" pagination="19 - 20" refId="ref120510" refString="Olivier, E. (1903 b) Coleopteres Lampyrides captures ae Dardjilling par M. le Dr. Harmand. Bulletin de Museum d'histoire naturelle de Paris, 9, 19 - 20." type="journal article" year="1903" yearSuffix="b">Olivier 1903b</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="CDD3796351BE18D3FDF0FAD0EDD7192E" box="[617,751,1359,1385]" pageId="133" pageNumber="134" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="AC8E4B8151BE18D3FF0EFAECEE0A19CA" box="[151,306,1395,1421]" pageId="133" pageNumber="134" type="description">
<paragraph id="E42B180A51BE18D3FF0EFAECEE0A19CA" blockId="133.[151,751,1358,1421]" box="[151,306,1395,1421]" pageId="133" pageNumber="134">
<figureCitation id="7CAF048F51BE18D3FF0EFAECEE0A19CA" box="[151,306,1395,1421]" captionStart-0="FIGURES 422429" captionStart-1="FIGURES 430436" captionStartId-0="134.[151,264,1911,1936]" captionStartId-1="135.[151,264,1893,1918]" captionTargetBox-0="[213,1349,183,1892]" captionTargetBox-1="[218,1364,166,1867]" captionTargetId-0="figure@134.[213,1375,183,1892]" captionTargetId-1="figure@135.[218,1369,166,1873]" captionTargetPageId-0="134" captionTargetPageId-1="135" captionText-0="FIGURES 422429. Pygoluciola nitescens comb. nov. male. 422, 423 Borneo coll. Chaper (MNHN); 424429 Sarawak coll Polunin (ANIC). 422, 424 habitus dorsal; 423 habitus ventral; 425, 426 terminal abdomen: 425 ventral; 426 dorsal; 427429 aedeagal sheath: 427 dorsal; 428 ventral (outline of lobes arising from boomerang shaped piece at apex of sheath sternite arrowed); 429 left lateral." captionText-1="FIGURES 430436. Pygoluciola nitescens comb. nov. 430432 male (Sarawak coll. Polunin;ANIC); 433436 female (Ranau; NHML). 430432 aedeagus: 430 ventral; 431 dorsal; 432 left lateral; 433 habitus dorsal; 434 habitus ventral; 435 abdominal ventrite 8, ventral; 436 reproductive system, ovipositor to right, spermatophore digesting gland and spermatheca to left (arrow indicates area of bursa and possible thin bursa plates). Scale lines are 1 mm in 430, 435 and 2 mm in 433." figureDoi-0="http://doi.org/10.5281/zenodo.3508262" figureDoi-1="http://doi.org/10.5281/zenodo.3508264" httpUri-0="https://zenodo.org/record/3508262/files/figure.png" httpUri-1="https://zenodo.org/record/3508264/files/figure.png" pageId="133" pageNumber="134">Figs 422436</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="AC8E4B8151BE18D3FF0EFA28EC531988" box="[151,875,1462,1487]" pageId="133" pageNumber="134" type="reference_group">
<paragraph id="E42B180A51BE18D3FF0EFA28EC531988" blockId="133.[151,875,1462,1518]" box="[151,875,1462,1487]" pageId="133" pageNumber="134">
<treatmentCitationGroup id="C4843F2451BE18D3FF0EFA28EC531988" box="[151,875,1462,1487]" pageId="133" pageNumber="134">
<taxonomicName id="2394638951BE18D3FF0EFA28ED3D1988" ID-CoL="8MYPJ" authority="Olivier 1903 b: 19" authorityName="Olivier" authorityPageNumber="19" authorityYear="1903" box="[151,517,1462,1487]" class="Insecta" family="Lampyridae" genus="Luciola" kingdom="Animalia" order="Coleoptera" pageId="133" pageNumber="134" phylum="Arthropoda" rank="species" species="nitescens">
<emphasis id="D6E0C41851BE18D3FF0EFA28EE701988" box="[151,328,1463,1487]" italics="true" pageId="133" pageNumber="134">Luciola nitescens</emphasis>
<treatmentCitation id="65353E1B51BE18D3FED7FA29ED3D1988" author="Olivier, E." box="[334,517,1462,1487]" page="19" pageId="133" pageNumber="134" year="1903">
<bibRefCitation id="800565FB51BE18D3FED7FA29ED3D1988" author="Olivier" box="[334,517,1462,1487]" firstAuthor="Olivier" pageId="133" pageNumber="134" pagination="19 - 20" refId="ref120510" refString="Olivier, E. (1903 b) Coleopteres Lampyrides captures ae Dardjilling par M. le Dr. Harmand. Bulletin de Museum d'histoire naturelle de Paris, 9, 19 - 20." type="journal article" year="1903" yearSuffix="b">Olivier 1903b: 19</bibRefCitation>
</treatmentCitation>
</taxonomicName>
;
<treatmentCitation id="65353E1B51BE18D3FD89FA29ED411988" author="Olivier, E." box="[528,633,1462,1487]" page="206" pageId="133" pageNumber="134" year="1905">
<bibRefCitation id="800565FB51BE18D3FD89FA29ED411988" author="Olivier" box="[528,633,1462,1487]" firstAuthor="Olivier" pageId="133" pageNumber="134" pagination="206 - 209" refId="ref120545" refString="Olivier, E. (1905) Descriptions de Lampyrides nouveaux. Annales de la Societe Entomologique de Belgique, XLIX, 206 - 209." type="book chapter" year="1905">1905: 206</bibRefCitation>
</treatmentCitation>
.
<treatmentCitation id="65353E1B51BE18D3FD1BFA28EC5F1988" author="McDermott, F. A." box="[642,871,1462,1487]" page="110" pageId="133" pageNumber="134" year="1966">
<bibRefCitation id="800565FB51BE18D3FD1BFA28EC5F1988" author="McDermott" box="[642,871,1462,1487]" firstAuthor="McDermott" pageId="133" pageNumber="134" pagination="1 - 149" refId="ref119380" refString="McDermott, F. A. (1966) Lampyridae. In: Steel, W. O. (Ed.), Coleopterorum Catalogus Supplementa. Pars 9. Editio Secunda. W. Junk, S'Gravenhage, pp. 1 - 149." type="book chapter" year="1966">McDermott 1966: 110</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="AC8E4B8151BE18D3FF0EFA49ED651AE5" pageId="133" pageNumber="134" type="materials_examined">
<paragraph id="E42B180A51BE18D3FF0EFA49ED8219A9" blockId="133.[151,875,1462,1518]" box="[151,698,1493,1518]" pageId="133" pageNumber="134">
<materialsCitation id="54FC125751BE18D3FF0EFA49ED8F19A9" ID-GBIF-Occurrence="2427485573" box="[151,695,1493,1518]" country="India" location="Darjeeling" pageId="133" pageNumber="134" specimenCount="1" typeStatus="holotype">
<emphasis id="D6E0C41851BE18D3FF0EFA49EE1C19A9" bold="true" box="[151,292,1493,1518]" pageId="133" pageNumber="134">
<typeStatus id="3B2FA6A851BE18D3FF0EFA49EFF619A9" box="[151,206,1494,1518]" pageId="133" pageNumber="134">Type</typeStatus>
.
<collectingCountry id="9C83589A51BE18D3FF40FA4AEE1C19A9" box="[217,292,1493,1518]" name="India" pageId="133" pageNumber="134">INDIA</collectingCountry>
</emphasis>
.
<location id="E14B4ED151BE18D3FEA9FA4AEEA319A9" LSID="urn:lsid:plazi:treatment:6C3DA91C51BE18DEFF0EFAD1EE801A14:E14B4ED151BE18D3FEA9FA4AEEA319A9" box="[304,411,1493,1518]" country="India" name="Darjeeling" pageId="133" pageNumber="134">Darjeeling</location>
(as Dardjilling) not located
</materialsCitation>
.
</paragraph>
<paragraph id="E42B180A51BE18D3FF0EF984ED651AE5" blockId="133.[151,1437,1563,2023]" pageId="133" pageNumber="134">
<emphasis id="D6E0C41851BE18D3FF0EF984EEE11A71" bold="true" box="[151,473,1563,1590]" pageId="133" pageNumber="134">Other specimens examined</emphasis>
.
<materialsCitation id="54FC125751BE18D3FE7EF984ED621AE5" ID-GBIF-Occurrence="2427485756" collectingDate="2006-05-07" collectionCode="NHML, MNHN, ANIC" collectorName="S. Chew &amp; Borneo &amp; Chaper &amp; E. Olivier &amp; Leluan I. Polunin" country="Malaysia" location="Ranau" municipality="Borneo" pageId="133" pageNumber="134" specimenCount="1" stateProvince="Sabah">
<emphasis id="D6E0C41851BE18D3FE7EF984ED451A71" bold="true" box="[487,637,1563,1590]" pageId="133" pageNumber="134">
<collectingCountry id="9C83589A51BE18D3FE7EF984ED451A71" box="[487,637,1563,1590]" name="Malaysia" pageId="133" pageNumber="134">MALAYSIA</collectingCountry>
</emphasis>
as
<collectingMunicipality id="044F827051BE18D3FD3BF982EDC11A70" box="[674,761,1565,1591]" pageId="133" pageNumber="134">Borneo</collectingMunicipality>
:
<collectingRegion id="2650D6E851BE18D3FC9DF983EC701A71" box="[772,840,1564,1590]" country="Malaysia" name="Sabah" pageId="133" pageNumber="134">Sabah</collectingRegion>
<location id="E14B4ED151BE18D3FCD6F982ECAF1A70" LSID="urn:lsid:plazi:treatment:6C3DA91C51BE18DEFF0EFAD1EE801A14:E14B4ED151BE18D3FCD6F982ECAF1A70" box="[847,919,1565,1591]" country="Malaysia" municipality="Borneo" name="Ranau" pageId="133" pageNumber="134" stateProvince="Sabah">Ranau</location>
<date id="902A3ECA51BE18D3FC07F983ECC61A71" box="[926,1022,1564,1590]" pageId="133" pageNumber="134" value="2006-05-07">
<collectingDate id="806EC72251BE18D3FC07F983ECC61A71" box="[926,1022,1564,1590]" pageId="133" pageNumber="134" value="2006-05-07">7.v.2006</collectingDate>
</date>
<collectorName id="49617DDC51BE18D3FB9CF983EB5C1A71" box="[1029,1124,1564,1590]" pageId="133" pageNumber="134">S. Chew</collectorName>
female (
<collectionCode id="828580CF51BE18D3FB5FF983EA241A71" box="[1222,1308,1564,1590]" collectionName="Libya, Tripoli, Natural History Museum" country="Libya" httpUri="http://grbio.org/cool/49nc-28y0" name="Natural History Museum, Tripoli" pageId="133" pageNumber="134">NHML</collectionCode>
) (
<figureCitation id="7CAF048F51BE18D3FAABF983EFFE1A1D" captionStart="FIGURES 430436" captionStartId="135.[151,264,1893,1918]" captionTargetBox="[218,1364,166,1867]" captionTargetId="figure@135.[218,1369,166,1873]" captionTargetPageId="135" captionText="FIGURES 430436. Pygoluciola nitescens comb. nov. 430432 male (Sarawak coll. Polunin;ANIC); 433436 female (Ranau; NHML). 430432 aedeagus: 430 ventral; 431 dorsal; 432 left lateral; 433 habitus dorsal; 434 habitus ventral; 435 abdominal ventrite 8, ventral; 436 reproductive system, ovipositor to right, spermatophore digesting gland and spermatheca to left (arrow indicates area of bursa and possible thin bursa plates). Scale lines are 1 mm in 430, 435 and 2 mm in 433." figureDoi="http://doi.org/10.5281/zenodo.3508264" httpUri="https://zenodo.org/record/3508264/files/figure.png" pageId="133" pageNumber="134">Figs 433, 434</figureCitation>
).
<collectorName id="49617DDC51BE18D3FF43F9DEEE171A1C" box="[218,303,1601,1627]" pageId="133" pageNumber="134">Borneo</collectorName>
,
<collectorName id="49617DDC51BE18D3FEA5F9DFEEB51A1D" box="[316,397,1600,1626]" pageId="133" pageNumber="134">Chaper</collectorName>
1891, male, identified by
<collectorName id="49617DDC51BE18D3FD25F9DEEC141A1D" box="[700,812,1600,1627]" pageId="133" pageNumber="134">E. Olivier</collectorName>
(
<figureCitation id="7CAF048F51BE18D3FCA5F9DFECE31A1D" box="[828,987,1600,1626]" captionStart="FIGURES 422429" captionStartId="134.[151,264,1911,1936]" captionTargetBox="[213,1349,183,1892]" captionTargetId="figure@134.[213,1375,183,1892]" captionTargetPageId="134" captionText="FIGURES 422429. Pygoluciola nitescens comb. nov. male. 422, 423 Borneo coll. Chaper (MNHN); 424429 Sarawak coll Polunin (ANIC). 422, 424 habitus dorsal; 423 habitus ventral; 425, 426 terminal abdomen: 425 ventral; 426 dorsal; 427429 aedeagal sheath: 427 dorsal; 428 ventral (outline of lobes arising from boomerang shaped piece at apex of sheath sternite arrowed); 429 left lateral." figureDoi="http://doi.org/10.5281/zenodo.3508262" httpUri="https://zenodo.org/record/3508262/files/figure.png" pageId="133" pageNumber="134">Figs 422, 423</figureCitation>
) (
<collectionCode id="828580CF51BE18D3FC6DF9DFEB751A1D" box="[1012,1101,1600,1626]" collectionName="France, Paris, Museum National d'Histoire Naturelle" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="133" pageNumber="134">MNHN</collectionCode>
) (identification label reads 1. Handwritten
<taxonomicName id="2394638951BE18D3FEB4F9FBED5F1A39" authority="Ern Oliv." authorityName="Ern Oliv." box="[301,615,1636,1663]" class="Insecta" family="Lampyridae" genus="Luciola" kingdom="Animalia" order="Coleoptera" pageId="133" pageNumber="134" phylum="Arthropoda" rank="species" species="nitescens">Luciola nitescens Ern Oliv.</taxonomicName>
; 2. Printed Museum Paris Borneo Chaper 1891).
<collectingRegion id="2650D6E851BE18D3FB04F9FBEBC51A39" box="[1181,1277,1636,1662]" country="Malaysia" name="Sarawak" pageId="133" pageNumber="134">
<collectingCountry id="9C83589A51BE18D3FB04F9FBEBC51A39" box="[1181,1277,1636,1662]" name="Malaysia" pageId="133" pageNumber="134">Sarawak</collectingCountry>
</collectingRegion>
inland
<collectorName id="49617DDC51BE18D3FAD7F9FBEE3E1AE5" pageId="133" pageNumber="134">Leluan I. Polunin</collectorName>
male (
<collectionCode id="828580CF51BE18D3FECBF917EEA21AE5" box="[338,410,1672,1698]" collectionName="Australia, Australian Capital Territory, Canberra City, CSIRO, Australian National Insect Collection" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="133" pageNumber="134">ANIC</collectionCode>
) (
<figureCitation id="7CAF048F51BE18D3FE29F917ED691AE5" box="[432,593,1671,1698]" captionStart-0="FIGURES 422429" captionStart-1="FIGURES 430436" captionStartId-0="134.[151,264,1911,1936]" captionStartId-1="135.[151,264,1893,1918]" captionTargetBox-0="[213,1349,183,1892]" captionTargetBox-1="[218,1364,166,1867]" captionTargetId-0="figure@134.[213,1375,183,1892]" captionTargetId-1="figure@135.[218,1369,166,1873]" captionTargetPageId-0="134" captionTargetPageId-1="135" captionText-0="FIGURES 422429. Pygoluciola nitescens comb. nov. male. 422, 423 Borneo coll. Chaper (MNHN); 424429 Sarawak coll Polunin (ANIC). 422, 424 habitus dorsal; 423 habitus ventral; 425, 426 terminal abdomen: 425 ventral; 426 dorsal; 427429 aedeagal sheath: 427 dorsal; 428 ventral (outline of lobes arising from boomerang shaped piece at apex of sheath sternite arrowed); 429 left lateral." captionText-1="FIGURES 430436. Pygoluciola nitescens comb. nov. 430432 male (Sarawak coll. Polunin;ANIC); 433436 female (Ranau; NHML). 430432 aedeagus: 430 ventral; 431 dorsal; 432 left lateral; 433 habitus dorsal; 434 habitus ventral; 435 abdominal ventrite 8, ventral; 436 reproductive system, ovipositor to right, spermatophore digesting gland and spermatheca to left (arrow indicates area of bursa and possible thin bursa plates). Scale lines are 1 mm in 430, 435 and 2 mm in 433." figureDoi-0="http://doi.org/10.5281/zenodo.3508262" figureDoi-1="http://doi.org/10.5281/zenodo.3508264" httpUri-0="https://zenodo.org/record/3508262/files/figure.png" httpUri-1="https://zenodo.org/record/3508264/files/figure.png" pageId="133" pageNumber="134">Figs 425432</figureCitation>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="AC8E4B8151BE18D3FF5EF934EDA01B74" pageId="133" pageNumber="134" type="diagnosis">
<paragraph id="E42B180A51BE18D3FF5EF934EDA01B74" blockId="133.[151,1437,1563,2023]" pageId="133" pageNumber="134">
<emphasis id="D6E0C41851BE18D3FF5EF934EE021A81" bold="true" box="[199,314,1707,1734]" pageId="133" pageNumber="134">Diagnosis</emphasis>
. A large heavy bodied species (12.0 mm long) with orange pronotum and black elytra which may be narrowly orange marked across their bases; distinguished from the similarly coloured
<taxonomicName id="2394638951BE18D3FBCFF94EEBE91AAD" authorityName="Ho" authorityYear="2019" baseAuthorityName="Olivier" baseAuthorityYear="1905" box="[1110,1233,1744,1770]" class="Insecta" family="Lampyridae" genus="Pygoluciola" kingdom="Animalia" order="Coleoptera" pageId="133" pageNumber="134" phylum="Arthropoda" rank="species" species="calceata" status="comb. nov.">
<emphasis id="D6E0C41851BE18D3FBCFF94EEBE91AAD" box="[1110,1233,1744,1770]" italics="true" pageId="133" pageNumber="134">P. calceata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CDD3796351BE18D3FB4EF94FEA611AAC" box="[1239,1369,1744,1771]" pageId="133" pageNumber="134" rank="species">
<emphasis id="D6E0C41851BE18D3FB4EF94FEA6A1AAD" bold="true" box="[1239,1362,1744,1770]" pageId="133" pageNumber="134">comb. nov</emphasis>
.
</taxonomicNameLabel>
by its size (
<taxonomicName id="2394638951BE18D3FF4AF96BEE0A1B49" authorityName="Ho" authorityYear="2019" baseAuthorityName="Olivier" baseAuthorityYear="1905" box="[211,306,1780,1806]" class="Insecta" family="Lampyridae" genus="Pygoluciola" kingdom="Animalia" order="Coleoptera" pageId="133" pageNumber="134" phylum="Arthropoda" rank="species" species="calceata">
<emphasis id="D6E0C41851BE18D3FF4AF96BEE0A1B49" box="[211,306,1780,1806]" italics="true" pageId="133" pageNumber="134">calceata</emphasis>
</taxonomicName>
is
<quantity id="236CB5EF51BE18D3FECAF96BEEAC1B48" box="[339,404,1780,1807]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="133" pageNumber="134" unit="mm" value="9.0">9 mm</quantity>
long) and the outline of the MPP (in
<taxonomicName id="2394638951BE18D3FCA1F96AECA71B49" box="[824,927,1781,1806]" class="Insecta" family="Lampyridae" genus="Luciola" kingdom="Animalia" order="Coleoptera" pageId="133" pageNumber="134" phylum="Arthropoda" rank="species" species="nitescens">
<emphasis id="D6E0C41851BE18D3FCA1F96AECA71B49" box="[824,927,1781,1806]" italics="true" pageId="133" pageNumber="134">nitescens</emphasis>
</taxonomicName>
it is short and apically acute; in
<taxonomicName id="2394638951BE18D3FA94F96BEA541B49" authorityName="Ho" authorityYear="2019" baseAuthorityName="Olivier" baseAuthorityYear="1905" box="[1293,1388,1780,1806]" class="Insecta" family="Lampyridae" genus="Pygoluciola" kingdom="Animalia" order="Coleoptera" pageId="133" pageNumber="134" phylum="Arthropoda" rank="species" species="calceata">
<emphasis id="D6E0C41851BE18D3FA94F96BEA541B49" box="[1293,1388,1780,1806]" italics="true" pageId="133" pageNumber="134">calceata</emphasis>
</taxonomicName>
it is about as long as wide and apically truncated).
</paragraph>
</subSubSection>
<subSubSection id="AC8E4B8151BE18DEFF5EF8A4EC93188F" lastPageId="136" lastPageNumber="137" pageId="133" pageNumber="134" type="description">
<paragraph id="E42B180A51BE18DEFF5EF8A4EB931E88" blockId="133.[151,1437,1563,2023]" lastBlockId="136.[151,1437,153,1619]" lastPageId="136" lastPageNumber="137" pageId="133" pageNumber="134">
<emphasis id="D6E0C41851BE18D3FF5EF8A4EEF71B11" bold="true" box="[199,463,1851,1878]" pageId="133" pageNumber="134">Redescription of male</emphasis>
. 12.0 mm long. Colour (
<figureCitation id="7CAF048F51BE18D3FD7EF8A3ECB21B11" box="[743,906,1852,1878]" captionStart="FIGURES 422429" captionStartId="134.[151,264,1911,1936]" captionTargetBox="[213,1349,183,1892]" captionTargetId="figure@134.[213,1375,183,1892]" captionTargetPageId="134" captionText="FIGURES 422429. Pygoluciola nitescens comb. nov. male. 422, 423 Borneo coll. Chaper (MNHN); 424429 Sarawak coll Polunin (ANIC). 422, 424 habitus dorsal; 423 habitus ventral; 425, 426 terminal abdomen: 425 ventral; 426 dorsal; 427429 aedeagal sheath: 427 dorsal; 428 ventral (outline of lobes arising from boomerang shaped piece at apex of sheath sternite arrowed); 429 left lateral." figureDoi="http://doi.org/10.5281/zenodo.3508262" httpUri="https://zenodo.org/record/3508262/files/figure.png" pageId="133" pageNumber="134">Figs 422424</figureCitation>
): pronotum, MS and MN orange; elytra almost black, having humerus and narrow basal area orange in
<typeStatus id="3B2FA6A851BE18D3FD63F8FEEC161B3C" box="[762,814,1889,1915]" pageId="133" pageNumber="134">type</typeStatus>
; head between eyes, antennae and palpi very dark brown almost black; venter of pro and mesothorax yellowish, of metathorax slightly darker (venter of all thoracic segments light brown in
<typeStatus id="3B2FA6A851BE18D3FEA5F836EE491B84" box="[316,369,1961,1987]" pageId="133" pageNumber="134">type</typeStatus>
); legs yellowish (light brown in
<typeStatus id="3B2FA6A851BE18D3FD40F836EC331B84" box="[729,779,1961,1987]" pageId="133" pageNumber="134">type</typeStatus>
) with all tibiae and tarsi dark brown; basal abdominal ventrites yellowish;
<date id="902A3ECA51BE18D3FEC8F853EE4D1BA1" box="[337,373,1996,2022]" pageId="133" pageNumber="134">V6</date>
, 7 with whitish LO,
<date id="902A3ECA51BE18D3FDC0F852ED431BA0" box="[601,635,1997,2023]" pageId="133" pageNumber="134">V7</date>
LO retracted from all margins in Leluan male, retracted along posterior margin only in
<typeStatus id="3B2FA6A851B318DEFED9FF06EE4C1CF4" box="[320,372,153,179]" pageId="136" pageNumber="137">type</typeStatus>
; all abdominal tergites yellowish in Leluan male (colour not observed in
<typeStatus id="3B2FA6A851B318DEFB03FF06EBF71CF4" box="[1178,1231,153,179]" pageId="136" pageNumber="137">type</typeStatus>
). Pronotum: width subequal to (in
<typeStatus id="3B2FA6A851B318DEFED8FF22EE4D1C90" box="[321,373,189,215]" pageId="136" pageNumber="137">type</typeStatus>
), or slightly wider than humeral width; lateral margins strongly divergent and posterolateral corners rounded and projecting. Elytron: slightly convex sided; interstitial lines not well defined. Head: GHW
<date id="902A3ECA51B318DEFAA3FF7EEA641CBC" box="[1338,1372,225,251]" pageId="136" pageNumber="137">6 x</date>
SIW: antennal sockets very close but not contiguous. Mouthparts: apical labial palpomere laterally flattened of the form of a broad triangle with inner edge dentate (3 teeth). Antennae:&gt; GHW but not exceeding
<date id="902A3ECA51B318DEFBC5FEB6EB461D04" box="[1116,1150,297,323]" pageId="136" pageNumber="137">2 x</date>
GHW; all segments elongate (longer than
<date id="902A3ECA51B318DEFEC5FED2EE471D20" box="[348,383,333,359]" pageId="136" pageNumber="137">2 x</date>
width) slender. Abdomen (
<figureCitation id="7CAF048F51B318DEFD36FED2EC771D20" box="[687,847,332,359]" captionStart="FIGURES 422429" captionStartId="134.[151,264,1911,1936]" captionTargetBox="[213,1349,183,1892]" captionTargetId="figure@134.[213,1375,183,1892]" captionTargetPageId="134" captionText="FIGURES 422429. Pygoluciola nitescens comb. nov. male. 422, 423 Borneo coll. Chaper (MNHN); 424429 Sarawak coll Polunin (ANIC). 422, 424 habitus dorsal; 423 habitus ventral; 425, 426 terminal abdomen: 425 ventral; 426 dorsal; 427429 aedeagal sheath: 427 dorsal; 428 ventral (outline of lobes arising from boomerang shaped piece at apex of sheath sternite arrowed); 429 left lateral." figureDoi="http://doi.org/10.5281/zenodo.3508262" httpUri="https://zenodo.org/record/3508262/files/figure.png" pageId="136" pageNumber="137">Figs 423, 425</figureCitation>
): posterior margin of
<date id="902A3ECA51B318DEFBD9FED2EB5A1D20" box="[1088,1122,333,359]" pageId="136" pageNumber="137">V7</date>
without PLP and with short broadly rounded MPP; T8 with truncate posterior margin and subparallel sided, anterolateral prolongations long slender expanded vertically. Aedeagal sheath (
<figureCitation id="7CAF048F51B318DEFD35FE0AEC691DE8" box="[684,849,405,431]" captionStart="FIGURES 422429" captionStartId="134.[151,264,1911,1936]" captionTargetBox="[213,1349,183,1892]" captionTargetId="figure@134.[213,1375,183,1892]" captionTargetPageId="134" captionText="FIGURES 422429. Pygoluciola nitescens comb. nov. male. 422, 423 Borneo coll. Chaper (MNHN); 424429 Sarawak coll Polunin (ANIC). 422, 424 habitus dorsal; 423 habitus ventral; 425, 426 terminal abdomen: 425 ventral; 426 dorsal; 427429 aedeagal sheath: 427 dorsal; 428 ventral (outline of lobes arising from boomerang shaped piece at apex of sheath sternite arrowed); 429 left lateral." figureDoi="http://doi.org/10.5281/zenodo.3508262" httpUri="https://zenodo.org/record/3508262/files/figure.png" pageId="136" pageNumber="137">Figs 427429</figureCitation>
): tergite with anterior transverse darkened separate plate associated with but not clearly attached to the anterior margin of the sheath tergite; basal ¾ of sheath sternite with sclerotized margins very narrow, expanding in apical ¼ where it is terminated by broad paired rounded hairy lobes which are united across their bases by a darkened semicircular piece; dark transverse strip of cuticle (outline of lobes arrowed in
<figureCitation id="7CAF048F51B318DEFEEEFDBAEEE01E78" box="[375,472,549,575]" captionStart="FIGURES 422429" captionStartId="134.[151,264,1911,1936]" captionTargetBox="[213,1349,183,1892]" captionTargetId="figure@134.[213,1375,183,1892]" captionTargetPageId="134" captionText="FIGURES 422429. Pygoluciola nitescens comb. nov. male. 422, 423 Borneo coll. Chaper (MNHN); 424429 Sarawak coll Polunin (ANIC). 422, 424 habitus dorsal; 423 habitus ventral; 425, 426 terminal abdomen: 425 ventral; 426 dorsal; 427429 aedeagal sheath: 427 dorsal; 428 ventral (outline of lobes arising from boomerang shaped piece at apex of sheath sternite arrowed); 429 left lateral." figureDoi="http://doi.org/10.5281/zenodo.3508262" httpUri="https://zenodo.org/record/3508262/files/figure.png" pageId="136" pageNumber="137">Fig. 428</figureCitation>
). Aedeagus (
<figureCitation id="7CAF048F51B318DEFDFEFDBAEC321E78" box="[615,778,549,575]" captionStart="FIGURES 430436" captionStartId="135.[151,264,1893,1918]" captionTargetBox="[218,1364,166,1867]" captionTargetId="figure@135.[218,1369,166,1873]" captionTargetPageId="135" captionText="FIGURES 430436. Pygoluciola nitescens comb. nov. 430432 male (Sarawak coll. Polunin;ANIC); 433436 female (Ranau; NHML). 430432 aedeagus: 430 ventral; 431 dorsal; 432 left lateral; 433 habitus dorsal; 434 habitus ventral; 435 abdominal ventrite 8, ventral; 436 reproductive system, ovipositor to right, spermatophore digesting gland and spermatheca to left (arrow indicates area of bursa and possible thin bursa plates). Scale lines are 1 mm in 430, 435 and 2 mm in 433." figureDoi="http://doi.org/10.5281/zenodo.3508264" httpUri="https://zenodo.org/record/3508264/files/figure.png" pageId="136" pageNumber="137">Figs 430432</figureCitation>
): slightly more than twice as long as wide and evenly wide along its entire length; BP in two well cuticularised narrow parts; membranous apical section of LL with dense line of long hairs inside their lateral margins, and quite widely expanded in horizontal plane; ML tapering to a rounded narrow apex, not much shorter than LL; darker basal portion of LL with apical 1/3 separated, rest of LL fused at their bases; anterior margin of basal portion of LL asymmetrical, produced and rounded on its left.
</paragraph>
<caption id="B0EB488251BD18D0FF0EF8E8EEBF1BBB" ID-DOI="http://doi.org/10.5281/zenodo.3508262" ID-Zenodo-Dep="3508262" httpUri="https://zenodo.org/record/3508262/files/figure.png" pageId="134" pageNumber="135" startId="134.[151,264,1911,1936]" targetBox="[213,1349,183,1892]" targetPageId="134">
<paragraph id="E42B180A51BD18D0FF0EF8E8EEBF1BBB" blockId="134.[151,1437,1911,2044]" pageId="134" pageNumber="135">
<emphasis id="D6E0C41851BD18D0FF0EF8E8EE571BD7" bold="true" box="[151,367,1911,1936]" pageId="134" pageNumber="135">FIGURES 422429.</emphasis>
<taxonomicName id="2394638951BD18D0FEEFF8E7ED6C1BD7" authorityName="Ho" authorityYear="2019" baseAuthorityName="Olivier" baseAuthorityYear="1903" box="[374,596,1912,1936]" class="Insecta" family="Lampyridae" genus="Pygoluciola" kingdom="Animalia" order="Coleoptera" pageId="134" pageNumber="135" phylum="Arthropoda" rank="species" species="nitescens" status="comb. nov.">
<emphasis id="D6E0C41851BD18D0FEEFF8E7ED6C1BD7" box="[374,596,1912,1936]" italics="true" pageId="134" pageNumber="135">Pygoluciola nitescens</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CDD3796351BD18D0FDC2F8E7EDE91BC8" box="[603,721,1912,1936]" pageId="134" pageNumber="135" rank="species">
<emphasis id="D6E0C41851BD18D0FDC2F8E7EDF21BD7" bold="true" box="[603,714,1912,1936]" pageId="134" pageNumber="135">comb. nov</emphasis>
.
</taxonomicNameLabel>
male. 422, 423 Borneo coll. Chaper (MNHN); 424429 Sarawak coll Polunin (ANIC). 422, 424 habitus dorsal; 423 habitus ventral; 425, 426 terminal abdomen: 425 ventral; 426 dorsal; 427429 aedeagal sheath: 427 dorsal; 428 ventral (outline of lobes arising from boomerang shaped piece at apex of sheath sternite arrowed); 429 left lateral.
</paragraph>
</caption>
<caption id="B0EB488251BC18D1FF0EF8FAEBFD1BAE" ID-DOI="http://doi.org/10.5281/zenodo.3508264" ID-Zenodo-Dep="3508264" httpUri="https://zenodo.org/record/3508264/files/figure.png" pageId="135" pageNumber="136" startId="135.[151,264,1893,1918]" targetBox="[218,1364,166,1867]" targetPageId="135">
<paragraph id="E42B180A51BC18D1FF0EF8FAEBFD1BAE" blockId="135.[151,1437,1893,2026]" pageId="135" pageNumber="136">
<emphasis id="D6E0C41851BC18D1FF0EF8FAEE5E1B39" bold="true" box="[151,358,1893,1918]" pageId="135" pageNumber="136">FIGURES 430436</emphasis>
.
<taxonomicName id="2394638951BC18D1FEE8F8FAED761B3A" authorityName="Ho" authorityYear="2019" baseAuthorityName="Olivier" baseAuthorityYear="1903" box="[369,590,1893,1917]" class="Insecta" family="Lampyridae" genus="Pygoluciola" kingdom="Animalia" order="Coleoptera" pageId="135" pageNumber="136" phylum="Arthropoda" rank="species" species="nitescens" status="comb. nov.">
<emphasis id="D6E0C41851BC18D1FEE8F8FAED761B3A" box="[369,590,1893,1917]" italics="true" pageId="135" pageNumber="136">Pygoluciola nitescens</emphasis>
</taxonomicName>
<emphasis id="D6E0C41851BC18D1FDCAF8FAEDFD1B3A" bold="true" box="[595,709,1893,1917]" pageId="135" pageNumber="136">
<taxonomicNameLabel id="CDD3796351BC18D1FDCAF8FAEDFD1B3A" box="[595,709,1893,1917]" pageId="135" pageNumber="136" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
430432 male (Sarawak coll. Polunin; ANIC); 433436 female (Ranau; NHML). 430432 aedeagus: 430 ventral; 431 dorsal; 432 left lateral; 433 habitus dorsal; 434 habitus ventral; 435 abdominal ventrite 8, ventral; 436 reproductive system, ovipositor to right, spermatophore digesting gland and spermatheca to left (arrow indicates area of bursa and possible thin bursa plates). Scale lines are 1 mm in 430, 435 and 2 mm in 433.
</paragraph>
</caption>
<paragraph id="E42B180A51B318DEFF5EFD46EC93188F" blockId="136.[151,1437,153,1619]" pageId="136" pageNumber="137">
<emphasis id="D6E0C41851B318DEFF5EFD46EE261EB4" bold="true" box="[199,286,729,755]" pageId="136" pageNumber="137">Female</emphasis>
.
<figureCitation id="7CAF048F51B318DEFEB7FD46EEF61EB4" box="[302,462,729,755]" captionStart="FIGURES 430436" captionStartId="135.[151,264,1893,1918]" captionTargetBox="[218,1364,166,1867]" captionTargetId="figure@135.[218,1369,166,1873]" captionTargetPageId="135" captionText="FIGURES 430436. Pygoluciola nitescens comb. nov. 430432 male (Sarawak coll. Polunin;ANIC); 433436 female (Ranau; NHML). 430432 aedeagus: 430 ventral; 431 dorsal; 432 left lateral; 433 habitus dorsal; 434 habitus ventral; 435 abdominal ventrite 8, ventral; 436 reproductive system, ovipositor to right, spermatophore digesting gland and spermatheca to left (arrow indicates area of bursa and possible thin bursa plates). Scale lines are 1 mm in 430, 435 and 2 mm in 433." figureDoi="http://doi.org/10.5281/zenodo.3508264" httpUri="https://zenodo.org/record/3508264/files/figure.png" pageId="136" pageNumber="137">Figs 433436</figureCitation>
. Assigned on the basis of its size, locality and distinctive colour.
<quantity id="236CB5EF51B318DEFB2CFD46EA3D1EB4" box="[1205,1285,729,755]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" pageId="136" pageNumber="137" unit="mm" value="18.0">18 mm</quantity>
long. Colour (
<figureCitation id="7CAF048F51B318DEFF06FD62EE781F50" box="[159,320,765,791]" captionStart="FIGURES 430436" captionStartId="135.[151,264,1893,1918]" captionTargetBox="[218,1364,166,1867]" captionTargetId="figure@135.[218,1369,166,1873]" captionTargetPageId="135" captionText="FIGURES 430436. Pygoluciola nitescens comb. nov. 430432 male (Sarawak coll. Polunin;ANIC); 433436 female (Ranau; NHML). 430432 aedeagus: 430 ventral; 431 dorsal; 432 left lateral; 433 habitus dorsal; 434 habitus ventral; 435 abdominal ventrite 8, ventral; 436 reproductive system, ovipositor to right, spermatophore digesting gland and spermatheca to left (arrow indicates area of bursa and possible thin bursa plates). Scale lines are 1 mm in 430, 435 and 2 mm in 433." figureDoi="http://doi.org/10.5281/zenodo.3508264" httpUri="https://zenodo.org/record/3508264/files/figure.png" pageId="136" pageNumber="137">Figs 433, 434</figureCitation>
): pronotum dingy yellowish with faint traces of very small median brown markings; underlying fat body is retracted and confuses interpretation of colour; MS, MN dingy yellow; elytra black shiny; head, antennae and palpi black; venter of thorax and abdominal ventrites anterior to LO brown; legs brown except for dark brown tibiae and tarsi; ventrites 25 semitransparent and underlying fat body is visible beneath cuticle; LO in
<date id="902A3ECA51B318DEFAB9FCF6EA7A1FC4" box="[1312,1346,873,899]" pageId="136" pageNumber="137">V6</date>
creamy white retracted from anterior margin of
<date id="902A3ECA51B318DEFDC9FC12ED4E1FE0" box="[592,630,909,935]" pageId="136" pageNumber="137">V6</date>
;
<date id="902A3ECA51B318DEFD19FC12ED9C1FE0" box="[640,676,909,935]" pageId="136" pageNumber="137">V7</date>
, 8 semitransparent orange yellow; all abdominal tergites orange yellow, semitransparent. Pronotum (
<figureCitation id="7CAF048F51B318DEFD9FFC2EED511F8C" box="[518,617,945,971]" captionStart="FIGURES 430436" captionStartId="135.[151,264,1893,1918]" captionTargetBox="[218,1364,166,1867]" captionTargetId="figure@135.[218,1369,166,1873]" captionTargetPageId="135" captionText="FIGURES 430436. Pygoluciola nitescens comb. nov. 430432 male (Sarawak coll. Polunin;ANIC); 433436 female (Ranau; NHML). 430432 aedeagus: 430 ventral; 431 dorsal; 432 left lateral; 433 habitus dorsal; 434 habitus ventral; 435 abdominal ventrite 8, ventral; 436 reproductive system, ovipositor to right, spermatophore digesting gland and spermatheca to left (arrow indicates area of bursa and possible thin bursa plates). Scale lines are 1 mm in 430, 435 and 2 mm in 433." figureDoi="http://doi.org/10.5281/zenodo.3508264" httpUri="https://zenodo.org/record/3508264/files/figure.png" pageId="136" pageNumber="137">Fig. 433</figureCitation>
): median anterior margin not produced anteriorly; lateral margins strongly divergent; posterolateral corners rounded and produced beyond midposterior margin; width greater than humeral width. Elytron: slightly convex sided; interstitial line 3 irregularly defined. Abdomen (
<figureCitation id="7CAF048F51B318DEFBC4FC66EBC51854" box="[1117,1277,1017,1043]" captionStart="FIGURES 430436" captionStartId="135.[151,264,1893,1918]" captionTargetBox="[218,1364,166,1867]" captionTargetId="figure@135.[218,1369,166,1873]" captionTargetPageId="135" captionText="FIGURES 430436. Pygoluciola nitescens comb. nov. 430432 male (Sarawak coll. Polunin;ANIC); 433436 female (Ranau; NHML). 430432 aedeagus: 430 ventral; 431 dorsal; 432 left lateral; 433 habitus dorsal; 434 habitus ventral; 435 abdominal ventrite 8, ventral; 436 reproductive system, ovipositor to right, spermatophore digesting gland and spermatheca to left (arrow indicates area of bursa and possible thin bursa plates). Scale lines are 1 mm in 430, 435 and 2 mm in 433." figureDoi="http://doi.org/10.5281/zenodo.3508264" httpUri="https://zenodo.org/record/3508264/files/figure.png" pageId="136" pageNumber="137">Figs 433, 434</figureCitation>
):
<date id="902A3ECA51B318DEFA89FC66EA091854" box="[1296,1329,1017,1043]" pageId="136" pageNumber="137">V7</date>
posterior margin broadly emarginated with posterolateral corners rounded;
<date id="902A3ECA51B318DEFCE1FB82ECA11870" box="[888,921,1053,1079]" pageId="136" pageNumber="137">V8</date>
posterior margin emarginated (
<figureCitation id="7CAF048F51B318DEFB62FB82EA641870" box="[1275,1372,1052,1079]" captionStart="FIGURES 430436" captionStartId="135.[151,264,1893,1918]" captionTargetBox="[218,1364,166,1867]" captionTargetId="figure@135.[218,1369,166,1873]" captionTargetPageId="135" captionText="FIGURES 430436. Pygoluciola nitescens comb. nov. 430432 male (Sarawak coll. Polunin;ANIC); 433436 female (Ranau; NHML). 430432 aedeagus: 430 ventral; 431 dorsal; 432 left lateral; 433 habitus dorsal; 434 habitus ventral; 435 abdominal ventrite 8, ventral; 436 reproductive system, ovipositor to right, spermatophore digesting gland and spermatheca to left (arrow indicates area of bursa and possible thin bursa plates). Scale lines are 1 mm in 430, 435 and 2 mm in 433." figureDoi="http://doi.org/10.5281/zenodo.3508264" httpUri="https://zenodo.org/record/3508264/files/figure.png" pageId="136" pageNumber="137">Fig. 435</figureCitation>
) with anterior margin well coloured and separated from rest of ventrite by a clear membranous position. Genitalia (
<figureCitation id="7CAF048F51B318DEFAF6FBDEEFFE1838" captionStart="FIGURES 437448" captionStartId="137.[151,264,1892,1917]" captionTargetBox="[202,1361,166,1873]" captionTargetId="figure@137.[202,1362,166,1873]" captionTargetPageId="137" captionText="FIGURES 437448. Pygoluciola phupan sp. nov. type males. 437, 438 habitus dorsal; 439, 440 tergite 8: 439 ventral; 440 right lateral (projections arrowed); 441 habitus ventral; 442 444 abdomen: 442 ventral; 443 dorsal; 444 right lateral; 445, 446 aedeagus: 445 ventral; 446 dorsal (transmitted light); 447, 448 aedeagal sheath: 447 dorsal; 448 ventral. Scale lines are 1 mm unless otherwise indicated." figureDoi="http://doi.org/10.5281/zenodo.3508269" httpUri="https://zenodo.org/record/3508269/files/figure.png" pageId="136" pageNumber="137">Fig. 437</figureCitation>
): styli with both segments elongated slender, basal segment slightly longer than apical. Reproductive system (
<figureCitation id="7CAF048F51B318DEFF39FB16EE3B18E3" box="[160,259,1161,1188]" captionStart="FIGURES 430436" captionStartId="135.[151,264,1893,1918]" captionTargetBox="[218,1364,166,1867]" captionTargetId="figure@135.[218,1369,166,1873]" captionTargetPageId="135" captionText="FIGURES 430436. Pygoluciola nitescens comb. nov. 430432 male (Sarawak coll. Polunin;ANIC); 433436 female (Ranau; NHML). 430432 aedeagus: 430 ventral; 431 dorsal; 432 left lateral; 433 habitus dorsal; 434 habitus ventral; 435 abdominal ventrite 8, ventral; 436 reproductive system, ovipositor to right, spermatophore digesting gland and spermatheca to left (arrow indicates area of bursa and possible thin bursa plates). Scale lines are 1 mm in 430, 435 and 2 mm in 433." figureDoi="http://doi.org/10.5281/zenodo.3508264" httpUri="https://zenodo.org/record/3508264/files/figure.png" pageId="136" pageNumber="137">Fig. 436</figureCitation>
): bursa, SDG and spermatheca observed through abdomen soaked for an hour in cold KOH; bursa plates may be present but not clearly observed (white arrow indicates bursa).
</paragraph>
</subSubSection>
<subSubSection id="AC8E4B8151B318DEFF5EFB4EEE801A14" pageId="136" pageNumber="137" type="discussion">
<paragraph id="E42B180A51B318DEFF5EFB4EED241974" blockId="136.[151,1437,153,1619]" pageId="136" pageNumber="137">
<emphasis id="D6E0C41851B318DEFF5EFB4EEE0C18AC" bold="true" box="[199,308,1233,1259]" pageId="136" pageNumber="137">Remarks</emphasis>
.
<bibRefCitation id="800565FB51B318DEFED8FB4EEED618AB" author="Olivier" box="[321,494,1233,1260]" firstAuthor="Olivier" pageId="136" pageNumber="137" pagination="19 - 20" refId="ref120510" refString="Olivier, E. (1903 b) Coleopteres Lampyrides captures ae Dardjilling par M. le Dr. Harmand. Bulletin de Museum d'histoire naturelle de Paris, 9, 19 - 20." type="journal article" year="1903" yearSuffix="b">Olivier (1903b)</bibRefCitation>
described this species from
<collectingCountry id="9C83589A51B318DEFCBEFB4EEC5C18AC" box="[807,868,1233,1259]" name="India" pageId="136" pageNumber="137">India</collectingCountry>
. We are unable to locate Indian specimens. Howev- er we believe the male described here conforms with those external features we have determined from the Bornean male described by Olivier in 1905.
</paragraph>
<paragraph id="E42B180A51B318DEFF5EFAA2EE801A14" blockId="136.[151,1437,153,1619]" pageId="136" pageNumber="137">
The assignment to
<taxonomicName id="2394638951B318DEFE3AFAA2ED121910" authorityName="Wittmer" authorityYear="1939" box="[419,554,1341,1367]" class="Insecta" family="Lampyridae" genus="Pygoluciola" kingdom="Animalia" order="Coleoptera" pageId="136" pageNumber="137" phylum="Arthropoda" rank="genus">
<emphasis id="D6E0C41851B318DEFE3AFAA2ED121910" box="[419,554,1341,1367]" italics="true" pageId="136" pageNumber="137">Pygoluciola</emphasis>
</taxonomicName>
is justified by the general outline of the aedeagal sheath sternite and the aedeagus pattern. This is the first time however that an aedeagal sheath in this genus was found with a separate plate lying presumably anterior to the sheath tergite. This is a feature of several species in
<taxonomicName id="2394638951B318DEFBDAFA1AEBF919D8" authorityName="Ballantyne Lambkin &amp; Fu" authorityYear="2013" box="[1091,1217,1413,1439]" class="Insecta" family="Lampyridae" genus="Abscondita" kingdom="Animalia" order="Coleoptera" pageId="136" pageNumber="137" phylum="Arthropoda" rank="genus">
<emphasis id="D6E0C41851B318DEFBDAFA1AEBF919D8" box="[1091,1217,1413,1439]" italics="true" pageId="136" pageNumber="137">Abscondita</emphasis>
</taxonomicName>
which our analyses show to be closely related to
<taxonomicName id="2394638951B318DEFE47FA36ED5D1984" authorityName="Wittmer" authorityYear="1939" box="[478,613,1449,1475]" class="Insecta" family="Lampyridae" genus="Pygoluciola" kingdom="Animalia" order="Coleoptera" pageId="136" pageNumber="137" phylum="Arthropoda" rank="genus">
<emphasis id="D6E0C41851B318DEFE47FA36ED5D1984" box="[478,613,1449,1475]" italics="true" pageId="136" pageNumber="137">Pygoluciola</emphasis>
</taxonomicName>
. The pronotal outline, with strongly developed posterolateral corners and divergent lateral margins, and the distinctively shaped T8, which is parallel sided with long slender anterolateral prolongations, are more easily recognised features. However there is nothing about the outline of the terminal abdominal ventrite, which has a rounded MPP, which would suggest
<taxonomicName id="2394638951B318DEFC08F98AEB201A68" authorityName="Wittmer" authorityYear="1939" box="[913,1048,1557,1583]" class="Insecta" family="Lampyridae" genus="Pygoluciola" kingdom="Animalia" order="Coleoptera" pageId="136" pageNumber="137" phylum="Arthropoda" rank="genus">
<emphasis id="D6E0C41851B318DEFC08F98AEB201A68" box="[913,1048,1557,1583]" italics="true" pageId="136" pageNumber="137">Pygoluciola</emphasis>
</taxonomicName>
, and the shape is more associated with species of
<taxonomicName id="2394638951B318DEFEDFF9A6EE891A14" authorityName="Olliff" authorityYear="1890" box="[326,433,1593,1619]" class="Insecta" family="Lampyridae" genus="Atyphella" kingdom="Animalia" order="Coleoptera" pageId="136" pageNumber="137" phylum="Arthropoda" rank="genus">
<emphasis id="D6E0C41851B318DEFEDFF9A6EE891A14" box="[326,433,1593,1619]" italics="true" pageId="136" pageNumber="137">Atyphella</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>