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<document id="E96722F92E2DF9EA346AABA4BB4FA8F8" ID-CLB-Dataset="25190" ID-DOI="10.11646/zootaxa.4687.1.1" ID-GBIF-Dataset="de00ca6c-2843-4ae1-a39c-a29603947612" ID-ISSN="1175-5326" ID-Zenodo-Dep="3508098" ID-ZooBank="CE73264D-C234-4B82-A634-CAD6254C5957" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1571388220410" checkinUser="plazi" docAuthor="Ho, - Z." docDate="2019" docId="6C3DA91C51B818D3FF0EFC12ED1E18BA" docLanguage="en" docName="zootaxa.4687.1.1.pdf.imf" docOrigin="Zootaxa 4687 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Pygoluciola matalangao Ho 2019, sp. nov." docType="treatment" docVersion="19" lastPageNumber="134" masterDocId="9004D164513B1856FF99FF9FEF381C47" masterDocTitle="The Luciolinae of S. E. Asia and the Australopacific region: a revisionary checklist (Coleoptera: Lampyridae) including description of three new genera and 13 new species" masterLastPageNumber="174" masterPageNumber="1" pageNumber="132" updateTime="1698760145766" updateUser="ExternalLinkService">
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<mods:title id="07EBA33E12A0A93DA3C3A51D0E1C7594">The Luciolinae of S. E. Asia and the Australopacific region: a revisionary checklist (Coleoptera: Lampyridae) including description of three new genera and 13 new species</mods:title>
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<mods:namePart id="604365ED393AF44F7F07755EA750A6F8">Ho, - Z.</mods:namePart>
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<mods:date id="DB2892A486876C50ABD587DF38E18CA3">2019</mods:date>
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<mods:number id="BB3B7B17ECF091E4534C51ED84265E80">2019-10-18</mods:number>
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<mods:number id="18F0E435F2444B9F85B389C171FA2C44">4687</mods:number>
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<mods:classification id="0639A492494906DCF55A90A598DCB508">journal article</mods:classification>
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<mods:identifier id="DC447A446C370DDB75DFA8C42984D7CB" type="DOI">10.11646/zootaxa.4687.1.1</mods:identifier>
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<treatment id="6C3DA91C51B818D3FF0EFC12ED1E18BA" ID-DOI="http://doi.org/10.5281/zenodo.4688935" ID-GBIF-Taxon="159757401" ID-Zenodo-Dep="4688935" LSID="urn:lsid:plazi:treatment:6C3DA91C51B818D3FF0EFC12ED1E18BA" httpUri="http://treatment.plazi.org/id/6C3DA91C51B818D3FF0EFC12ED1E18BA" lastPageId="133" lastPageNumber="134" pageId="131" pageNumber="132">
<subSubSection id="AC8E4B8151B818D5FF0EFC12ED891FE0" box="[151,689,909,936]" pageId="131" pageNumber="132" type="nomenclature">
<paragraph id="E42B180A51B818D5FF0EFC12ED891FE0" blockId="131.[151,689,909,971]" box="[151,689,909,936]" pageId="131" pageNumber="132">
<heading id="BF63AF6651B818D5FF0EFC12ED891FE0" bold="true" box="[151,689,909,936]" fontSize="11" level="1" pageId="131" pageNumber="132" reason="1">
<emphasis id="D6E0C41851B818D5FF0EFC12ED891FE0" bold="true" box="[151,689,909,936]" pageId="131" pageNumber="132">
<taxonomicName id="2394638951B818D5FF0EFC12ED741FE0" ID-CoL="8NBSP" authority="Ballantyne" authorityName="Ho" authorityYear="2019" box="[151,588,909,936]" class="Insecta" family="Lampyridae" genus="Pygoluciola" kingdom="Animalia" order="Coleoptera" pageId="131" pageNumber="132" phylum="Arthropoda" rank="species" species="matalangao" status="sp. nov.">
<emphasis id="D6E0C41851B818D5FF0EFC12EE851FEF" bold="true" box="[151,445,909,936]" italics="true" pageId="131" pageNumber="132">Pygoluciola matalangao</emphasis>
Ballantyne
</taxonomicName>
<taxonomicNameLabel id="CDD3796351B818D5FDCAFC12ED891FE0" box="[595,689,909,935]" pageId="131" pageNumber="132" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="AC8E4B8151B818D5FF0EFC2EEE0B1F8C" box="[151,307,944,971]" pageId="131" pageNumber="132" type="description">
<paragraph id="E42B180A51B818D5FF0EFC2EEE0B1F8C" blockId="131.[151,689,909,971]" box="[151,307,944,971]" pageId="131" pageNumber="132">
<figureCitation id="7CAF048F51B818D5FF0EFC2EEE0B1F8C" box="[151,307,944,971]" captionStart-0="FIGURES 405417" captionStart-1="FIGURES 418421" captionStartId-0="132.[151,264,1885,1910]" captionStartId-1="133.[151,264,820,845]" captionTargetBox-0="[216,1366,166,1859]" captionTargetBox-1="[151,1435,174,792]" captionTargetId-0="figure@132.[216,1372,166,1865]" captionTargetId-1="figure@133.[151,1435,174,800]" captionTargetPageId-0="132" captionTargetPageId-1="133" captionText-0="FIGURES 405417. Pygoluciola matalangao sp. nov. type males. 405 habitus dorsal; 406 habitus ventral; 407 head anterior; 408, 409 dorsal pronotum; 410 abdomen ventral tergite 8 removed; 411 terminal abdominal segments dorsal, tergite 8 removed (arrows indicate dorsally reflexed margins of ventrite 7); 412 tergite 8 ventral; 413415 aedeagus: 413 ventral; 414 left lateral; 415 dorsal; 416, 417 aedeagal sheath: 416 dorsal; 417 ventral." captionText-1="FIGURES 418421. Pygoluciola matalangao sp. nov. females. 418 habitus dorsal; 419 habitus ventral; 420, 420 pronotum and base of elytra, dorsal." figureDoi-0="http://doi.org/10.5281/zenodo.3508254" figureDoi-1="http://doi.org/10.5281/zenodo.3508260" httpUri-0="https://zenodo.org/record/3508254/files/figure.png" httpUri-1="https://zenodo.org/record/3508260/files/figure.png" pageId="131" pageNumber="132">Figs 405421</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="AC8E4B8151B818D5FF0EFC6AEB8B180C" pageId="131" pageNumber="132" type="materials_examined">
<paragraph id="E42B180A51B818D5FF0EFC6AEE25186B" blockId="131.[151,1436,1012,1099]" pageId="131" pageNumber="132">
<materialsCitation id="54FC125751B818D5FF0EFC6AEE25186B" ID-GBIF-Occurrence="2449324225" pageId="131" pageNumber="132" typeStatus="holotype">
<emphasis id="D6E0C41851B818D5FF0EFC6AEFEA184A" bold="true" box="[151,210,1013,1037]" pageId="131" pageNumber="132">
<typeStatus id="3B2FA6A851B818D5FF0EFC6AEFF6184A" box="[151,206,1013,1037]" pageId="131" pageNumber="132">Type</typeStatus>
.
</emphasis>
Male.
<emphasis id="D6E0C41851B818D5FE86FC6BEEF9184A" bold="true" box="[287,449,1012,1037]" pageId="131" pageNumber="132">
<collectingCountry id="9C83589A51B818D5FE86FC6BEEF9184A" box="[287,449,1012,1037]" name="Philippines" pageId="131" pageNumber="132">PHILIPPINES</collectingCountry>
</emphasis>
. Island of
<collectingRegion id="2650D6E851B818D5FDACFC6BEDB5184A" box="[565,653,1012,1037]" country="Philippines" name="Palawan" pageId="131" pageNumber="132">Palawan</collectingRegion>
, Province of
<collectingRegion id="2650D6E851B818D5FC85FC6BEC4C184A" box="[796,884,1012,1037]" country="Philippines" name="Palawan" pageId="131" pageNumber="132">Palawan</collectingRegion>
, Matalangao nr Roxas,
<quantity id="236CB5EF51B818D5FBF6FC6BEB8C184A" box="[1135,1204,1012,1037]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="131" pageNumber="132" unit="m" value="150.0">150 m</quantity>
,
<date id="902A3ECA51B818D5FB58FC6BEA04184A" box="[1217,1340,1012,1037]" pageId="131" pageNumber="132" value="1985-08-28">28.viii.1985</date>
M Sakai (
<collectionCode id="828580CF51B818D5FF57FB8BEE2A186B" box="[206,274,1044,1068]" collectionName="Australia, Australian Capital Territory, Canberra City, CSIRO, Australian National Insect Collection" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="131" pageNumber="132">ANIC</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="E42B180A51B818D5FF0EFBACEB8B180C" blockId="131.[151,1436,1012,1099]" box="[151,1203,1074,1099]" pageId="131" pageNumber="132">
<materialsCitation id="54FC125751B818D5FF0EFBACEB97180C" ID-GBIF-Occurrence="2449324226" box="[151,1199,1074,1099]" collectionCode="ANIC" pageId="131" pageNumber="132" specimenCount="2" specimenCount-female="2" typeStatus="paratype">
<emphasis id="D6E0C41851B818D5FF0EFBACEE3B180C" bold="true" box="[151,259,1075,1099]" pageId="131" pageNumber="132">
<typeStatus id="3B2FA6A851B818D5FF0EFBACEE3B180C" box="[151,259,1075,1099]" pageId="131" pageNumber="132" type="paratype">Paratypes</typeStatus>
</emphasis>
(6). Same locality and date as
<typeStatus id="3B2FA6A851B818D5FDA5FBADEDAD180C" box="[572,661,1074,1099]" pageId="131" pageNumber="132" type="holotype">holotype</typeStatus>
<specimenCount id="F292D38351B818D5FD02FBACEDD4180C" box="[667,748,1074,1099]" pageId="131" pageNumber="132" type="male">3 males</specimenCount>
;
<quantity id="236CB5EF51B818D5FD6FFBACEC08180C" box="[758,816,1075,1099]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="131" pageNumber="132" unit="m" value="200.0">200m</quantity>
<date id="902A3ECA51B818D5FCAFFBADEC89180C" box="[822,945,1074,1099]" pageId="131" pageNumber="132" value="1985-08-30">30.viii.1985</date>
male,
<specimenCount id="F292D38351B818D5FC6CFBACEB6F180C" box="[1013,1111,1074,1099]" pageId="131" pageNumber="132" type="female">2 females</specimenCount>
(
<collectionCode id="828580CF51B818D5FBFDFBACEB90180C" box="[1124,1192,1075,1099]" collectionName="Australia, Australian Capital Territory, Canberra City, CSIRO, Australian National Insect Collection" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="131" pageNumber="132">ANIC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="AC8E4B8151B818D5FF0EFBE7ED2E18B8" pageId="131" pageNumber="132" type="diagnosis">
<paragraph id="E42B180A51B818D5FF0EFBE7ED2E18B8" blockId="131.[151,1437,1144,1999]" pageId="131" pageNumber="132">
<emphasis id="D6E0C41851B818D5FF0EFBE7EE2A18D4" bold="true" box="[151,274,1144,1171]" pageId="131" pageNumber="132">Diagnosis.</emphasis>
8.49.8 mm long; slender body; with dingy orange pronotum having median dark markings, and medi- um brown elytra with narrowly paler interstitial lines 1, 2 and suture. Dorsally reflexed lateral margins of
<date id="902A3ECA51B818D5FAB5FB02EA7618F0" box="[1324,1358,1181,1207]" pageId="131" pageNumber="132">V7</date>
enfold sides of T8. Belongs to that section of
<taxonomicName id="2394638951B818D5FDCBFB5EEDE1189C" authorityName="Wittmer" authorityYear="1939" box="[594,729,1217,1243]" class="Insecta" family="Lampyridae" genus="Pygoluciola" kingdom="Animalia" order="Coleoptera" pageId="131" pageNumber="132" phylum="Arthropoda" rank="genus">
<emphasis id="D6E0C41851B818D5FDCBFB5EEDE1189C" box="[594,729,1217,1243]" italics="true" pageId="131" pageNumber="132">Pygoluciola</emphasis>
</taxonomicName>
where neither
<date id="902A3ECA51B818D5FC1CFB5EEC9F189C" box="[901,935,1217,1243]" pageId="131" pageNumber="132">V7</date>
nor T8 are prolonged and apically inclined. Known only from the
<collectingCountry id="9C83589A51B818D5FE09FB7AED2A18B8" box="[400,530,1253,1279]" name="Philippines" pageId="131" pageNumber="132">Philippines</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="AC8E4B8151B818D3FF5EFA97ED7C180E" lastPageId="133" lastPageNumber="134" pageId="131" pageNumber="132" type="description">
<paragraph id="E42B180A51B818D5FF5EFA97EDA11B88" blockId="131.[151,1437,1144,1999]" pageId="131" pageNumber="132">
<emphasis id="D6E0C41851B818D5FF5EFA97EE891964" bold="true" box="[199,433,1288,1315]" pageId="131" pageNumber="132">Description of male</emphasis>
. 8.49.8 mm long; 3.2 mm wide. Colour (
<figureCitation id="7CAF048F51B818D5FC17FA96EB081964" box="[910,1072,1288,1315]" captionStart="FIGURES 405417" captionStartId="132.[151,264,1885,1910]" captionTargetBox="[216,1366,166,1859]" captionTargetId="figure@132.[216,1372,166,1865]" captionTargetPageId="132" captionText="FIGURES 405417. Pygoluciola matalangao sp. nov. type males. 405 habitus dorsal; 406 habitus ventral; 407 head anterior; 408, 409 dorsal pronotum; 410 abdomen ventral tergite 8 removed; 411 terminal abdominal segments dorsal, tergite 8 removed (arrows indicate dorsally reflexed margins of ventrite 7); 412 tergite 8 ventral; 413415 aedeagus: 413 ventral; 414 left lateral; 415 dorsal; 416, 417 aedeagal sheath: 416 dorsal; 417 ventral." figureDoi="http://doi.org/10.5281/zenodo.3508254" httpUri="https://zenodo.org/record/3508254/files/figure.png" pageId="131" pageNumber="132">Figs 405409</figureCitation>
): (the specimens were originally preserved in 70% ethanol and remounted). Pronotum yellowish brown with paired median areas beside midline darker (
<typeStatus id="3B2FA6A851B818D5FF76FACEEE6B192C" box="[239,339,1361,1387]" pageId="131" pageNumber="132" type="holotype">holotype</typeStatus>
male
<figureCitation id="7CAF048F51B818D5FE02FACEEEC6192C" box="[411,510,1361,1387]" captionStart="FIGURES 405417" captionStartId="132.[151,264,1885,1910]" captionTargetBox="[216,1366,166,1859]" captionTargetId="figure@132.[216,1372,166,1865]" captionTargetPageId="132" captionText="FIGURES 405417. Pygoluciola matalangao sp. nov. type males. 405 habitus dorsal; 406 habitus ventral; 407 head anterior; 408, 409 dorsal pronotum; 410 abdomen ventral tergite 8 removed; 411 terminal abdominal segments dorsal, tergite 8 removed (arrows indicate dorsally reflexed margins of ventrite 7); 412 tergite 8 ventral; 413415 aedeagus: 413 ventral; 414 left lateral; 415 dorsal; 416, 417 aedeagal sheath: 416 dorsal; 417 ventral." figureDoi="http://doi.org/10.5281/zenodo.3508254" httpUri="https://zenodo.org/record/3508254/files/figure.png" pageId="131" pageNumber="132">Fig. 409</figureCitation>
); or dingy brown with mid to dark brown markings beside mid line not reaching the narrowly pale posterior margin, and anterolateral and posterolateral corners paler dingy brown, and median line widely paler yellowish brown (
<figureCitation id="7CAF048F51B818D5FE6CFA06EDAD19F4" box="[501,661,1432,1459]" captionStart="FIGURES 405417" captionStartId="132.[151,264,1885,1910]" captionTargetBox="[216,1366,166,1859]" captionTargetId="figure@132.[216,1372,166,1865]" captionTargetPageId="132" captionText="FIGURES 405417. Pygoluciola matalangao sp. nov. type males. 405 habitus dorsal; 406 habitus ventral; 407 head anterior; 408, 409 dorsal pronotum; 410 abdomen ventral tergite 8 removed; 411 terminal abdominal segments dorsal, tergite 8 removed (arrows indicate dorsally reflexed margins of ventrite 7); 412 tergite 8 ventral; 413415 aedeagus: 413 ventral; 414 left lateral; 415 dorsal; 416, 417 aedeagal sheath: 416 dorsal; 417 ventral." figureDoi="http://doi.org/10.5281/zenodo.3508254" httpUri="https://zenodo.org/record/3508254/files/figure.png" pageId="131" pageNumber="132">Figs 405, 408</figureCitation>
); MN and MS paler than pronotum, pale cream to pale dingy brown; elytra medium brown with interstitial lines 1, 2 and suture narrowly paler brown; elytral punctation black; head antennae and labrum very dark brown; ventral aspect pro and mesothorax brown, of metathorax dark brown; all legs with basal segments light brown, tibiae and tarsi 1 mid brown; legs 2 light brown except for darker brown tarsi; all of legs 3 light brown; basal abdominal ventrites brown,
<date id="902A3ECA51B818D5FC9FF9B6EC101A04" box="[774,808,1577,1603]" pageId="131" pageNumber="132">V6</date>
very dark brown, LO creamy yellow; basal tergites pale brown, T7, 8 pale yellowish. Pronotum: narrower than humeral width. Elytron: subparallel sided. Abdomen (
<figureCitation id="7CAF048F51B818D5FAF2F9D2EE011ACC" captionStart="FIGURES 405417" captionStartId="132.[151,264,1885,1910]" captionTargetBox="[216,1366,166,1859]" captionTargetId="figure@132.[216,1372,166,1865]" captionTargetPageId="132" captionText="FIGURES 405417. Pygoluciola matalangao sp. nov. type males. 405 habitus dorsal; 406 habitus ventral; 407 head anterior; 408, 409 dorsal pronotum; 410 abdomen ventral tergite 8 removed; 411 terminal abdominal segments dorsal, tergite 8 removed (arrows indicate dorsally reflexed margins of ventrite 7); 412 tergite 8 ventral; 413415 aedeagus: 413 ventral; 414 left lateral; 415 dorsal; 416, 417 aedeagal sheath: 416 dorsal; 417 ventral." figureDoi="http://doi.org/10.5281/zenodo.3508254" httpUri="https://zenodo.org/record/3508254/files/figure.png" pageId="131" pageNumber="132">Figs 406, 410412</figureCitation>
): LO entire in both
<date id="902A3ECA51B818D5FD8EF9EEED471ACC" box="[535,639,1649,1675]" pageId="131" pageNumber="132">V6 and 7</date>
occupying all of
<date id="902A3ECA51B818D5FCDFF9EEEC501ACC" box="[838,872,1649,1675]" pageId="131" pageNumber="132">V6</date>
and extending to sides and into MPP of
<date id="902A3ECA51B818D5FAADF9EEEA601ACC" box="[1332,1368,1649,1675]" pageId="131" pageNumber="132">V7</date>
; MPP narrow and apically acute, not inclined dorsally; dorsally reflexed margins of
<date id="902A3ECA51B818D5FB93F90AEB141AE8" box="[1034,1068,1685,1711]" pageId="131" pageNumber="132">V7</date>
enfold the sides of the posterior portion of T8 (
<figureCitation id="7CAF048F51B818D5FEDBF926EE991A94" box="[322,417,1721,1747]" captionStart="FIGURES 405417" captionStartId="132.[151,264,1885,1910]" captionTargetBox="[216,1366,166,1859]" captionTargetId="figure@132.[216,1372,166,1865]" captionTargetPageId="132" captionText="FIGURES 405417. Pygoluciola matalangao sp. nov. type males. 405 habitus dorsal; 406 habitus ventral; 407 head anterior; 408, 409 dorsal pronotum; 410 abdomen ventral tergite 8 removed; 411 terminal abdominal segments dorsal, tergite 8 removed (arrows indicate dorsally reflexed margins of ventrite 7); 412 tergite 8 ventral; 413415 aedeagus: 413 ventral; 414 left lateral; 415 dorsal; 416, 417 aedeagal sheath: 416 dorsal; 417 ventral." figureDoi="http://doi.org/10.5281/zenodo.3508254" httpUri="https://zenodo.org/record/3508254/files/figure.png" pageId="131" pageNumber="132">Fig. 411</figureCitation>
arrowed). T8 (
<figureCitation id="7CAF048F51B818D5FDD6F926ED8D1A94" box="[591,693,1721,1747]" captionStart="FIGURES 405417" captionStartId="132.[151,264,1885,1910]" captionTargetBox="[216,1366,166,1859]" captionTargetId="figure@132.[216,1372,166,1865]" captionTargetPageId="132" captionText="FIGURES 405417. Pygoluciola matalangao sp. nov. type males. 405 habitus dorsal; 406 habitus ventral; 407 head anterior; 408, 409 dorsal pronotum; 410 abdomen ventral tergite 8 removed; 411 terminal abdominal segments dorsal, tergite 8 removed (arrows indicate dorsally reflexed margins of ventrite 7); 412 tergite 8 ventral; 413415 aedeagus: 413 ventral; 414 left lateral; 415 dorsal; 416, 417 aedeagal sheath: 416 dorsal; 417 ventral." figureDoi="http://doi.org/10.5281/zenodo.3508254" httpUri="https://zenodo.org/record/3508254/files/figure.png" pageId="131" pageNumber="132">Fig. 412</figureCitation>
): about half as wide as
<date id="902A3ECA51B818D5FC5FF926ECD21A94" box="[966,1002,1721,1747]" pageId="131" pageNumber="132">V7</date>
, subparallel sided with posterior margin straight (depending on orientation this margin may appear emarginated); anterolateral arms narrowed, slightly expanded in vertical plane and shorter than posterior entire portion; without ventrally directed pieces arising from bases of anterolateral arms. Aedeagal sheath (
<figureCitation id="7CAF048F51B818D5FD00F8BAEC021B78" box="[665,826,1829,1855]" captionStart="FIGURES 405417" captionStartId="132.[151,264,1885,1910]" captionTargetBox="[216,1366,166,1859]" captionTargetId="figure@132.[216,1372,166,1865]" captionTargetPageId="132" captionText="FIGURES 405417. Pygoluciola matalangao sp. nov. type males. 405 habitus dorsal; 406 habitus ventral; 407 head anterior; 408, 409 dorsal pronotum; 410 abdomen ventral tergite 8 removed; 411 terminal abdominal segments dorsal, tergite 8 removed (arrows indicate dorsally reflexed margins of ventrite 7); 412 tergite 8 ventral; 413415 aedeagus: 413 ventral; 414 left lateral; 415 dorsal; 416, 417 aedeagal sheath: 416 dorsal; 417 ventral." figureDoi="http://doi.org/10.5281/zenodo.3508254" httpUri="https://zenodo.org/record/3508254/files/figure.png" pageId="131" pageNumber="132">Figs 416, 417</figureCitation>
): with elongate very slender anterior portion of sheath sternite; anterior margin of dorsal portion of sheath tergite gently emarginated. Aedeagus (
<figureCitation id="7CAF048F51B818D5FB38F8D6EA7D1B24" box="[1185,1349,1864,1891]" captionStart="FIGURES 405417" captionStartId="132.[151,264,1885,1910]" captionTargetBox="[216,1366,166,1859]" captionTargetId="figure@132.[216,1372,166,1865]" captionTargetPageId="132" captionText="FIGURES 405417. Pygoluciola matalangao sp. nov. type males. 405 habitus dorsal; 406 habitus ventral; 407 head anterior; 408, 409 dorsal pronotum; 410 abdomen ventral tergite 8 removed; 411 terminal abdominal segments dorsal, tergite 8 removed (arrows indicate dorsally reflexed margins of ventrite 7); 412 tergite 8 ventral; 413415 aedeagus: 413 ventral; 414 left lateral; 415 dorsal; 416, 417 aedeagal sheath: 416 dorsal; 417 ventral." figureDoi="http://doi.org/10.5281/zenodo.3508254" httpUri="https://zenodo.org/record/3508254/files/figure.png" pageId="131" pageNumber="132">Figs 413415</figureCitation>
): apical portion of LL membranous and extending well beyond acute apex of ML with margins clothed with a single line of hairs; basal sclerotised portion of LL separated along half its dorsal length; anterior dorsal basal margin of LL irregularly prolonged to the left, and rounded.
</paragraph>
<caption id="B0EB488251BF18D2FF0EF8C2EC341BA5" ID-DOI="http://doi.org/10.5281/zenodo.3508254" ID-Zenodo-Dep="3508254" httpUri="https://zenodo.org/record/3508254/files/figure.png" pageId="132" pageNumber="133" startId="132.[151,264,1885,1910]" targetBox="[216,1366,166,1859]" targetPageId="132">
<paragraph id="E42B180A51BF18D2FF0EF8C2EC341BA5" blockId="132.[151,1437,1885,2018]" pageId="132" pageNumber="133">
<emphasis id="D6E0C41851BF18D2FF0EF8C2EE481B31" bold="true" box="[151,368,1885,1910]" pageId="132" pageNumber="133">FIGURES 405417.</emphasis>
<taxonomicName id="2394638951BF18D2FEEFF8C1ED481B31" authority="Ho, 2019" authorityName="Ho" authorityYear="2019" box="[374,624,1886,1910]" class="Insecta" family="Lampyridae" genus="Pygoluciola" kingdom="Animalia" order="Coleoptera" pageId="132" pageNumber="133" phylum="Arthropoda" rank="species" species="matalangao" status="sp. nov.">
<emphasis id="D6E0C41851BF18D2FEEFF8C1ED481B31" box="[374,624,1886,1910]" italics="true" pageId="132" pageNumber="133">Pygoluciola matalangao</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CDD3796351BF18D2FDEEF8C1EDF11B32" box="[631,713,1886,1910]" pageId="132" pageNumber="133" rank="species">
<emphasis id="D6E0C41851BF18D2FDEEF8C1EDFB1B31" bold="true" box="[631,707,1886,1910]" pageId="132" pageNumber="133">sp. nov</emphasis>
.
</taxonomicNameLabel>
type males. 405 habitus dorsal; 406 habitus ventral; 407 head anterior; 408, 409 dorsal pronotum; 410 abdomen ventral tergite 8 removed; 411 terminal abdominal segments dorsal, tergite 8 removed (arrows indicate dorsally reflexed margins of ventrite 7); 412 tergite 8 ventral; 413415 aedeagus: 413 ventral; 414 left lateral; 415 dorsal; 416, 417 aedeagal sheath: 416 dorsal; 417 ventral.
</paragraph>
</caption>
<caption id="B0EB488251BE18D3FF0EFCABEE4A1F36" ID-DOI="http://doi.org/10.5281/zenodo.3508260" ID-Zenodo-Dep="3508260" httpUri="https://zenodo.org/record/3508260/files/figure.png" pageId="133" pageNumber="134" startId="133.[151,264,820,845]" targetBox="[151,1435,174,792]" targetPageId="133">
<paragraph id="E42B180A51BE18D3FF0EFCABEE4A1F36" blockId="133.[151,1437,820,881]" pageId="133" pageNumber="134">
<emphasis id="D6E0C41851BE18D3FF0EFCABEE561F0A" bold="true" box="[151,366,820,845]" pageId="133" pageNumber="134">FIGURES 418421.</emphasis>
<taxonomicName id="2394638951BE18D3FEEAFCABED541F0B" authority="Ho, 2019" authorityName="Ho" authorityYear="2019" box="[371,620,820,844]" class="Insecta" family="Lampyridae" genus="Pygoluciola" kingdom="Animalia" order="Coleoptera" pageId="133" pageNumber="134" phylum="Arthropoda" rank="species" species="matalangao" status="sp. nov.">
<emphasis id="D6E0C41851BE18D3FEEAFCABED541F0B" box="[371,620,820,844]" italics="true" pageId="133" pageNumber="134">Pygoluciola matalangao</emphasis>
</taxonomicName>
<emphasis id="D6E0C41851BE18D3FDEBFCAAEDF91F0A" bold="true" box="[626,705,821,845]" pageId="133" pageNumber="134">
<taxonomicNameLabel id="CDD3796351BE18D3FDEBFCAAEDF91F0A" box="[626,705,821,845]" pageId="133" pageNumber="134" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
females. 418 habitus dorsal; 419 habitus ventral; 420, 420 pronotum and base of elytra, dorsal.
</paragraph>
</caption>
<paragraph id="E42B180A51BE18D3FF5EFC00ED7C180E" blockId="133.[151,1437,927,1277]" pageId="133" pageNumber="134">
<emphasis id="D6E0C41851BE18D3FF5EFC00EE261FFE" bold="true" box="[199,286,927,953]" pageId="133" pageNumber="134">Female</emphasis>
.
<figureCitation id="7CAF048F51BE18D3FEB5FC00EEF31FFE" box="[300,459,927,953]" captionStart="FIGURES 418421" captionStartId="133.[151,264,820,845]" captionTargetBox="[151,1435,174,792]" captionTargetId="figure@133.[151,1435,174,800]" captionTargetPageId="133" captionText="FIGURES 418421. Pygoluciola matalangao sp. nov. females. 418 habitus dorsal; 419 habitus ventral; 420, 420 pronotum and base of elytra, dorsal." figureDoi="http://doi.org/10.5281/zenodo.3508260" httpUri="https://zenodo.org/record/3508260/files/figure.png" pageId="133" pageNumber="134">Figs 418421</figureCitation>
); coloured as for male except for: pronotum orange with extensive darker markings and narrow paler yellow median line; MN and MS pale dingy brown; abdomen mid brown with creamy LO restricted to
<date id="902A3ECA51BE18D3FF2CFC78EFE11846" box="[181,217,999,1025]" pageId="133" pageNumber="134">V6</date>
,
<date id="902A3ECA51BE18D3FF7FFC78EE301846" box="[230,264,999,1025]" pageId="133" pageNumber="134">V7</date>
semitransparent pale, with creamy fat body visible at sides; posterior margin of
<date id="902A3ECA51BE18D3FB38FC78EBFB1846" box="[1185,1219,999,1025]" pageId="133" pageNumber="134">V7</date>
widely and deeply emarginated;
<date id="902A3ECA51BE18D3FEB6FB94EE691862" box="[303,337,1035,1061]" pageId="133" pageNumber="134">V8</date>
brown, with posterior margin rounded. Without any ridges or depressions on either
<date id="902A3ECA51BE18D3FB60FB94EA251862" box="[1273,1309,1035,1061]" pageId="133" pageNumber="134">V7</date>
, 8 or T7, 8. Reproductive system not investigated.
</paragraph>
</subSubSection>
<subSubSection id="AC8E4B8151BE18D3FF5EFBCCEB1F182A" box="[199,1063,1107,1133]" pageId="133" pageNumber="134" type="etymology">
<paragraph id="E42B180A51BE18D3FF5EFBCCEB1F182A" blockId="133.[151,1437,927,1277]" box="[199,1063,1107,1133]" pageId="133" pageNumber="134">
<emphasis id="D6E0C41851BE18D3FF5EFBCCEE70182A" bold="true" box="[199,328,1107,1133]" pageId="133" pageNumber="134">Etymology</emphasis>
. The specific name uses the
<typeStatus id="3B2FA6A851BE18D3FD13FBCCED82182A" box="[650,698,1107,1133]" pageId="133" pageNumber="134">type</typeStatus>
locality as a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="AC8E4B8151BE18D3FF5EFBE9ED1E18BA" pageId="133" pageNumber="134" type="discussion">
<paragraph id="E42B180A51BE18D3FF5EFBE9ED1E18BA" blockId="133.[151,1437,927,1277]" pageId="133" pageNumber="134">
<emphasis id="D6E0C41851BE18D3FF5EFBE9EE7018D6" bold="true" box="[199,328,1142,1169]" pageId="133" pageNumber="134">Comments</emphasis>
. The distinctiveness of this species and its placement in
<taxonomicName id="2394638951BE18D3FC20FBE8EB7818D6" authorityName="Wittmer" authorityYear="1939" box="[953,1088,1143,1169]" class="Insecta" family="Lampyridae" genus="Pygoluciola" kingdom="Animalia" order="Coleoptera" pageId="133" pageNumber="134" phylum="Arthropoda" rank="genus">
<emphasis id="D6E0C41851BE18D3FC20FBE8EB7818D6" box="[953,1088,1143,1169]" italics="true" pageId="133" pageNumber="134">Pygoluciola</emphasis>
</taxonomicName>
was recognised in several phylogenetic analyses (Fu Ballantyne &amp; Lambkin 2012: 7;
<bibRefCitation id="800565FB51BE18D3FC9CFB04EB4D18F2" author="Ballantyne &amp; Lambkin" box="[773,1141,1179,1205]" firstAuthor="Ballantyne" pageId="133" pageNumber="134" pagination="1 - 162" refId="ref114597" refString="Ballantyne, L. A. &amp; Lambkin, C. L. (2013) Systematics and phylogenetics of Indo-Pacific Luciolinae fireflies (Coleoptera: Lampyridae) and the description of new genera. Zootaxa, 3653 (1), 1 - 162. https: // doi. org / 10.11646 / zootaxa. 3653.1.1" type="journal article" year="2013">Ballantyne &amp; Lambkin 2013: 10</bibRefCitation>
;
<bibRefCitation id="800565FB51BE18D3FB19FB04EAAE18F2" author="Ballantyne" box="[1152,1430,1178,1205]" etAl="et al." firstAuthor="Ballantyne" pageId="133" pageNumber="134" pagination="1 - 84" refId="ref114655" refString="Ballantyne, L., Lambkin, C. L., Boontop, Y. &amp; Jusoh, W. F. A. (2015) Revisional studies on the Luciolinae (Coleoptera: Lampyridae): 1. The firefly genus Pyrophanes Olivier with two new species. 2. Four new species of Pteroptyx Olivier. 3. Inflata gen. nov. Boontop, with redescription of Luciola indica (Motsch.) as Inflata indica comb. nov. Zootaxa, 3959 (1), 1 - 84. https: // doi. org / 10.11646 / zootaxa. 3959.1.1" type="journal article" year="2015">
Ballantyne
<emphasis id="D6E0C41851BE18D3FA98FB03EA0118F2" box="[1281,1337,1179,1205]" italics="true" pageId="133" pageNumber="134">et al.</emphasis>
2015: 8
</bibRefCitation>
;
<bibRefCitation id="800565FB51BE18D3FF0EFB20EE32189E" author="Ballantyne" box="[151,266,1215,1241]" firstAuthor="Ballantyne" pageId="133" pageNumber="134" pagination="201 - 249" refId="ref114763" refString="Ballantyne, L. A., Lambkin, C. L., Luan, X., Boontop, Y., Nak-eiam, S., Pimpasalee, S., Silalom, S. &amp; Thancharoen, A. (2016) Further studies on south eastern Asian Luciolinae: 1. Sclerotia Ballantyne, a new genus of fireflies with back swimming larvae. 2. Triangulara Pimpasalee, a new genus from Thailand (Coleoptera: Lampyridae). Zootaxa, 4170 (2), 201 - 249. https: // doi. org / 10.11646 / zootaxa. 4170.2.1" type="journal article" year="2016">2016: 204</bibRefCitation>
) where it was scored under the code name “Jeng Matalanga”. We thank Ming-Luen Jeng for making the specimens available to us for study.
</paragraph>
</subSubSection>
</treatment>
</document>