325 lines
37 KiB
XML
325 lines
37 KiB
XML
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<mods:title id="D83ADE971F9BF6E2C56321CC4F11EFB9">The Luciolinae of S. E. Asia and the Australopacific region: a revisionary checklist (Coleoptera: Lampyridae) including description of three new genera and 13 new species</mods:title>
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<mods:namePart id="F9DD56CDC30D0E79A042A99007D3DAD9">Ho, - Z.</mods:namePart>
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<mods:title id="4F8C6BC98F7C3766A8B772F985ED429E">Zootaxa</mods:title>
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<mods:date id="6D13BA863D6F3FC7792DFD56F42046AF">2019</mods:date>
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<treatment id="6C3DA91C51761818FF0EFD9EEE571AA4" ID-DOI="http://doi.org/10.5281/zenodo.4688855" ID-GBIF-Taxon="159757306" ID-Zenodo-Dep="4688855" LSID="urn:lsid:plazi:treatment:6C3DA91C51761818FF0EFD9EEE571AA4" httpUri="http://treatment.plazi.org/id/6C3DA91C51761818FF0EFD9EEE571AA4" lastPageId="78" lastPageNumber="79" pageId="77" pageNumber="78">
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<subSubSection id="AC8E4B815176181BFF0EFD9EED141E5C" box="[151,556,513,539]" pageId="77" pageNumber="78" type="nomenclature">
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<paragraph id="E42B180A5176181BFF0EFD9EED141E5C" blockId="77.[151,556,513,575]" box="[151,556,513,539]" pageId="77" pageNumber="78">
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<heading id="BF63AF665176181BFF0EFD9EED141E5C" bold="true" box="[151,556,513,539]" fontSize="11" level="1" pageId="77" pageNumber="78" reason="1">
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<emphasis id="D6E0C4185176181BFF0EFD9EED141E5C" bold="true" box="[151,556,513,539]" pageId="77" pageNumber="78">
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<taxonomicName id="239463895176181BFF0EFD9EEE8F1E5C" ID-CoL="4B5C" authority="Ballantyne" authorityName="Ho" authorityYear="2019" box="[151,439,513,539]" class="Aves" family="Muscicapidae" genus="Emarginata" kingdom="Animalia" order="Passeriformes" pageId="77" pageNumber="78" phylum="Chordata" rank="genus" status="gen. nov.">
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<emphasis id="D6E0C4185176181BFF0EFD9EEE101E5C" bold="true" box="[151,296,513,539]" italics="true" pageId="77" pageNumber="78">Emarginata</emphasis>
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Ballantyne
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</taxonomicName>
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<taxonomicNameLabel id="CDD379635176181BFE27FD9EED141E5C" box="[446,556,513,539]" pageId="77" pageNumber="78" rank="genus">gen. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="AC8E4B815176181BFF0EFDBAEE2E1E78" box="[151,278,548,575]" pageId="77" pageNumber="78" type="description">
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<paragraph id="E42B180A5176181BFF0EFDBAEE2E1E78" blockId="77.[151,556,513,575]" box="[151,278,548,575]" pageId="77" pageNumber="78">
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<figureCitation id="7CAF048F5176181BFF0EFDBAEE2E1E78" box="[151,278,548,575]" captionStart="FIGURES 70–77" captionStartId="21.[151,264,1907,1932]" captionTargetBox="[184,1404,166,1879]" captionTargetId="figure@21.[184,1404,166,1888]" captionTargetPageId="21" captionText="FIGURES 70–77. Key to genera characters 11. 70–72 Serratia sibuyania gen. et sp. nov.; 73–75 Emarginata trilucida gen. nov. et comb. nov.; 76, 77 Colophotia praeusta. 70, 72, 73 dorsal habitus (70 anterior body only); 74 ventral habitus; 71 dissected terminal abdomen ventral; 75 ventrite 7 and elytral apices from below; 76, 77 ventrites 6, 7 from left (76) and below (77). Figure legends: MC median carina; MPP median posterior projection; PLP posterolateral projections." figureDoi="http://doi.org/10.5281/zenodo.3508131" httpUri="https://zenodo.org/record/3508131/files/figure.png" pageId="77" pageNumber="78">Figs 73−75</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="AC8E4B815176181BFF0EFDF6EC251EC5" box="[151,797,617,642]" pageId="77" pageNumber="78" type="type_taxon">
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<paragraph id="E42B180A5176181BFF0EFDF6EC251EC5" blockId="77.[151,797,617,642]" box="[151,797,617,642]" pageId="77" pageNumber="78">
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<emphasis id="D6E0C4185176181BFF0EFDF6EE1E1EC5" bold="true" box="[151,294,617,642]" pageId="77" pageNumber="78">
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<typeStatus id="3B2FA6A85176181BFF0EFDF6EFF41EC6" box="[151,204,617,641]" pageId="77" pageNumber="78">Type</typeStatus>
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species:
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</emphasis>
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<taxonomicName id="239463895176181BFEB5FDF6EDB21EC5" ID-CoL="8MYPW" authority="Jeng et al. 2003 b" authorityName="Jeng" authorityYear="2003" box="[300,650,617,642]" class="Insecta" family="Lampyridae" genus="Luciola" kingdom="Animalia" order="Coleoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="trilucida">
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<emphasis id="D6E0C4185176181BFEB5FDF6EEEF1EC6" box="[300,471,617,641]" italics="true" pageId="77" pageNumber="78">Luciola trilucida</emphasis>
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<bibRefCitation id="800565FB5176181BFE44FDF5EDB21EC5" author="Jeng" box="[477,650,617,642]" etAl="et al." firstAuthor="Jeng" pageId="77" pageNumber="78" pagination="247 - 262" refId="ref118190" refString="Jeng, M. L., Yang, P. S. & Lai, J. (2003 b) Notes on the genus Luciola (Coleoptera, Lampyridae, Luciolinae) of Taiwan. Special Bulletin of the Japanese Society of Coleopterolog y, 6, 247 - 262." type="journal article" year="2003" yearSuffix="b">
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Jeng
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<emphasis id="D6E0C4185176181BFD8BFDF5ED7D1EC6" box="[530,581,617,641]" italics="true" pageId="77" pageNumber="78">et al.</emphasis>
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2003b
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</bibRefCitation>
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</taxonomicName>
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by monotypy.
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</paragraph>
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</subSubSection>
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<subSubSection id="AC8E4B8151761818FF0EFD36ED1F1A14" lastPageId="78" lastPageNumber="79" pageId="77" pageNumber="78" type="diagnosis">
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<paragraph id="E42B180A5176181BFF0EFD36ECD21833" blockId="77.[151,1437,681,2041]" pageId="77" pageNumber="78">
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<emphasis id="D6E0C4185176181BFF0EFD36EE321E83" bold="true" box="[151,266,681,708]" pageId="77" pageNumber="78">Diagnosis</emphasis>
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.
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<taxonomicName id="239463895176181BFE8EFD34EEA41E83" authority="Ho, 2019" authorityName="Ho" authorityYear="2019" box="[279,412,683,708]" class="Aves" family="Muscicapidae" genus="Emarginata" kingdom="Animalia" order="Passeriformes" pageId="77" pageNumber="78" phylum="Chordata" rank="genus" status="gen. nov.">
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<emphasis id="D6E0C4185176181BFE8EFD34EEA41E83" box="[279,412,683,708]" italics="true" pageId="77" pageNumber="78">Emarginata</emphasis>
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</taxonomicName>
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<emphasis id="D6E0C4185176181BFE3BFD35ED321E83" bold="true" box="[418,522,682,708]" pageId="77" pageNumber="78">
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<taxonomicNameLabel id="CDD379635176181BFE3BFD35ED321E83" box="[418,522,682,708]" pageId="77" pageNumber="78" rank="genus">gen. nov.</taxonomicNameLabel>
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</emphasis>
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is a S. E. Asian genus having orange pronotum, and black elytra having the unique feature of a slight emargination at their apices, and from which it derives its name. It belongs in a group of genera characterized by: an elongate slender aedeagus with LL largely concealed behind the ML when viewed from beneath; aedeagal sheath elongate slender, widest across middle, without bulbous paraprocts, and with both sides of posterior half of sheath sternite tapering evenly towards a narrow entire apex; pronotal width less than width across elytral humeri, parallel-sided elytra, no MFC. It is distinguished from all other
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<taxonomicName id="239463895176181BFB97FCC1EBBE1F3F" authorityName="Lacordaire" authorityYear="1857" box="[1038,1158,862,888]" class="Insecta" family="Lampyridae" genus="Luciolinae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="77" pageNumber="166" phylum="Arthropoda" rank="genus">Luciolinae</taxonomicName>
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genera by the emarginations of the posterior end of the elytra in males and from which it derives its name. Additionally males are distinguished from
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<taxonomicName id="239463895176181BFEABFC39EE881F87" authorityName="Motschulsky" authorityYear="1853" box="[306,432,934,960]" class="Insecta" family="Lampyridae" genus="Colophotia" kingdom="Animalia" order="Coleoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="genus">
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<emphasis id="D6E0C4185176181BFEABFC39EE881F87" box="[306,432,934,960]" italics="true" pageId="77" pageNumber="78">Colophotia</emphasis>
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</taxonomicName>
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in having no median carina on
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<date id="902A3ECA5176181BFC94FC38EC0A1F86" box="[781,818,935,961]" pageId="77" pageNumber="78">V7</date>
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, nor expanded and oblique PLP, or bipartite LOs in
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<date id="902A3ECA5176181BFAEBFC38EAAF1F86" box="[1394,1431,935,961]" pageId="77" pageNumber="78">V7</date>
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; from
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<taxonomicName id="239463895176181BFF4EFC54EE7B1FA3" authorityName="Olivier" authorityYear="1902" box="[215,323,971,996]" class="Insecta" family="Lampyridae" genus="Pteroptyx" kingdom="Animalia" order="Coleoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="genus">
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<emphasis id="D6E0C4185176181BFF4EFC54EE7B1FA3" box="[215,323,971,996]" italics="true" pageId="77" pageNumber="78">Pteroptyx</emphasis>
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</taxonomicName>
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in having no MFC, nor deflexed elytral apices, bulbous aedeagal sheath paraprocts and bipartite LOs in
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<date id="902A3ECA5176181BFF70FC70EE36184E" box="[233,270,1007,1033]" pageId="77" pageNumber="78">V7</date>
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; from
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<taxonomicName id="239463895176181BFECFFC71EEE3184F" authorityName="Olivier" authorityYear="1885" box="[342,475,1006,1032]" class="Insecta" family="Lampyridae" genus="Pyrophanes" kingdom="Animalia" order="Coleoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="genus">
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<emphasis id="D6E0C4185176181BFECFFC71EEE3184F" box="[342,475,1006,1032]" italics="true" pageId="77" pageNumber="78">Pyrophanes</emphasis>
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</taxonomicName>
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(which has a MFC) and
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<taxonomicName id="239463895176181BFD74FC70EC5C184F" box="[749,868,1007,1032]" class="Insecta" family="Lampyridae" genus="Pteroptyx" kingdom="Animalia" order="Coleoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="testacea">
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<emphasis id="D6E0C4185176181BFD74FC70EC5C184F" box="[749,868,1007,1032]" italics="true" pageId="77" pageNumber="78">P. testacea</emphasis>
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</taxonomicName>
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(which has no MFC) in not having incurving lobes along
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<date id="902A3ECA5176181BFF44FB8CEE39186A" box="[221,257,1043,1069]" pageId="77" pageNumber="78">V7</date>
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, or bipartite LO in
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<date id="902A3ECA5176181BFE44FB8CED3A186A" box="[477,514,1043,1069]" pageId="77" pageNumber="78">V7</date>
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; from
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<taxonomicName id="239463895176181BFDD7FB8CEDF9186B" authorityName="Ballantyne in Ballantyne & Lambkin" authorityYear="2013" box="[590,705,1043,1068]" class="Insecta" family="Lampyridae" genus="Trisinuata" kingdom="Animalia" order="Coleoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="genus">
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<emphasis id="D6E0C4185176181BFDD7FB8CEDF9186B" box="[590,705,1043,1068]" italics="true" pageId="77" pageNumber="78">Trisinuata</emphasis>
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</taxonomicName>
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by the entire LOs in
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<date id="902A3ECA5176181BFC2DFB8CECEE186A" box="[948,982,1043,1069]" pageId="77" pageNumber="78">V7</date>
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(those of
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<taxonomicName id="239463895176181BFBD3FB8CEB85186B" authorityName="Ballantyne in Ballantyne & Lambkin" authorityYear="2013" box="[1098,1213,1043,1068]" class="Insecta" family="Lampyridae" genus="Trisinuata" kingdom="Animalia" order="Coleoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="genus">
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<emphasis id="D6E0C4185176181BFBD3FB8CEB85186B" box="[1098,1213,1043,1068]" italics="true" pageId="77" pageNumber="78">Trisinuata</emphasis>
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</taxonomicName>
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are bipartite); from most
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<taxonomicName id="239463895176181BFF4AFBA9EE5D1817" authorityName="Ballantyne in Ballantyne & Lambkin" authorityYear="2013" box="[211,357,1078,1104]" class="Insecta" family="Lampyridae" genus="Medeopteryx" kingdom="Animalia" order="Coleoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="genus">
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<emphasis id="D6E0C4185176181BFF4AFBA9EE5D1817" box="[211,357,1078,1104]" italics="true" pageId="77" pageNumber="78">Medeopteryx</emphasis>
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</taxonomicName>
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in not having deflexed elytral apices and trisinuate
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<date id="902A3ECA5176181BFC07FBA8ECFF1816" box="[926,967,1079,1105]" pageId="77" pageNumber="78">V7.</date>
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It differs from
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<taxonomicName id="239463895176181BFBF7FBA9EA3D1817" baseAuthorityName="Motschulsky" baseAuthorityYear="1854" box="[1134,1285,1078,1104]" class="Insecta" family="Lampyridae" genus="Inflata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="indica">
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<emphasis id="D6E0C4185176181BFBF7FBA9EA3D1817" box="[1134,1285,1078,1104]" italics="true" pageId="77" pageNumber="78">Inflata indica</emphasis>
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</taxonomicName>
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in not having either the bulbous ML of the aedeagus or paraprocts on the aedeagal sheath.
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</paragraph>
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<paragraph id="E42B180A5176181BFF5EFBE1EAAA190B" blockId="77.[151,1437,681,2041]" pageId="77" pageNumber="78">
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<emphasis id="D6E0C4185176181BFF5EFBE1EE3C18DF" bold="true" box="[199,260,1150,1176]" pageId="77" pageNumber="78">Male</emphasis>
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. Pronotum (
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<bibRefCitation id="800565FB5176181BFE14FBE1ED7418DF" author="Jeng" box="[397,588,1150,1177]" etAl="et al." firstAuthor="Jeng" pageId="77" pageNumber="78" pagination="247 - 262" refId="ref118190" refString="Jeng, M. L., Yang, P. S. & Lai, J. (2003 b) Notes on the genus Luciola (Coleoptera, Lampyridae, Luciolinae) of Taiwan. Special Bulletin of the Japanese Society of Coleopterolog y, 6, 247 - 262." type="journal article" year="2003" yearSuffix="b">
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||
Jeng
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<emphasis id="D6E0C4185176181BFE50FBE0EEC118DF" box="[457,505,1150,1176]" italics="true" pageId="77" pageNumber="78">et al</emphasis>
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||
. 2003b
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</bibRefCitation>
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fig.1): dorsal surface without irregularities in posterolateral areas and longitudinal groove in lateral areas; punctation dense; anterior margin not explanate; lateral margins diverging posteriorly along most of their length (C> A, B); width <humeral width; anterolateral corners rounded obtuse; lateral margins without indentation at mid-point, and sinuosity in either horizontal or vertical plane; without indentation in lateral margin near posterolateral corner, and irregularities at corner; posterolateral corners angulate or rounded obtuse; posterolateral corners not projecting as far as median posterior margin; separated from it by scarce emarginations.
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</paragraph>
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<paragraph id="E42B180A5176181BFF5EFAC8ED8F19D3" blockId="77.[151,1437,681,2041]" pageId="77" pageNumber="78">Hypomera: closed; median area of hypomeron not elevated in vertical direction; median area more widely flattened than elsewhere; pronotal width/ GHW 1.2.</paragraph>
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<paragraph id="E42B180A5176181BFF5EFA01EE191AF3" blockId="77.[151,1437,681,2041]" pageId="77" pageNumber="78">
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Elytron (
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<bibRefCitation id="800565FB5176181BFEB3FA01EED419FF" author="Jeng" box="[298,492,1438,1465]" etAl="et al." firstAuthor="Jeng" pageId="77" pageNumber="78" pagination="247 - 262" refId="ref118190" refString="Jeng, M. L., Yang, P. S. & Lai, J. (2003 b) Notes on the genus Luciola (Coleoptera, Lampyridae, Luciolinae) of Taiwan. Special Bulletin of the Japanese Society of Coleopterolog y, 6, 247 - 262." type="journal article" year="2003" yearSuffix="b">
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Jeng
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<emphasis id="D6E0C4185176181BFEFFFA00EEA019FF" box="[358,408,1438,1464]" italics="true" pageId="77" pageNumber="78">et al</emphasis>
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. 2003b
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</bibRefCitation>
|
||
fig.2): punctation dense, not linear, not as large as that of pronotum, nor widely and evenly spaced; apices not deflexed; apices emarginated; epipleuron and sutural ridge extend beyond mid-point, almost to apex but not extending around emarginated apex, neither thickened in apical half; no interstitial lines; elytral carina absent; in horizontal specimen viewed from below epipleuron at elytral base wide, covering humerus; viewed from above anterior margin of epipleuron arises anterior to posterior margin of MS; epipleuron developed as a lateral ridge along most of length; sutural margins approximate along most of length in closed elytra; lateral margins parallel-sided; apex narrowly emarginated transversely with the outer edge (the lateral margin) forming a small hook..
|
||
</paragraph>
|
||
<paragraph id="E42B180A5176181BFF5EF921EA021BBF" blockId="77.[151,1437,681,2041]" pageId="77" pageNumber="78">
|
||
Head: moderately depressed between eyes; well exposed in front of pronotum, not capable of complete retraction within prothoracic cavity; eyes moderately separated beneath at level of posterior margin of mouthpart complex; eyes above labrum close to moderately separated; frons-vertex junction rounded, without median elevation; posterolateral eye excavation not strongly developed, not visible in resting head position; antennal sockets on head between eyes, not contiguous, separated by <ASW; clypeolabral suture present, flexible, not in front of anterior eye margin when head viewed with labrum horizontal; outer edges of labrum reach inner edges of closed mandibles. Mouthparts: functional; apical labial palpomere flattened, shaped like a moderately broad triangle (widest at base and L 2xW), with inner edge entire (
|
||
<bibRefCitation id="800565FB5176181BFDADF825EDC01B93" author="Jeng" box="[564,760,1978,2005]" etAl="et al." firstAuthor="Jeng" pageId="77" pageNumber="78" pagination="247 - 262" refId="ref118190" refString="Jeng, M. L., Yang, P. S. & Lai, J. (2003 b) Notes on the genus Luciola (Coleoptera, Lampyridae, Luciolinae) of Taiwan. Special Bulletin of the Japanese Society of Coleopterolog y, 6, 247 - 262." type="journal article" year="2003" yearSuffix="b">
|
||
Jeng
|
||
<emphasis id="D6E0C4185176181BFDE8F824ED9B1B93" box="[625,675,1978,2004]" italics="true" pageId="77" pageNumber="78">et al</emphasis>
|
||
. 2003b
|
||
</bibRefCitation>
|
||
fig. 3). Antennae 11 segmented; length> GHW up to twice GHW; no segments flattened, shortened, or expanded; pedicel not produced; FS1 not shorter than pedicel.
|
||
</paragraph>
|
||
<paragraph id="E42B180A51751818FF5EFF06ED1F1C90" blockId="78.[151,1437,153,1763]" pageId="78" pageNumber="79">Legs: with inner tarsal claw not split; without MFC; femora 3 not swollen and curved and tibiae 3 not curved; no basitarsi expanded or excavated.</paragraph>
|
||
<paragraph id="E42B180A51751818FF5EFF7EEB6A1F7C" blockId="78.[151,1437,153,1763]" pageId="78" pageNumber="79">
|
||
Abdomen (
|
||
<bibRefCitation id="800565FB51751818FEDDFF7EED3D1CBC" author="Jeng" box="[324,517,225,251]" etAl="et al." firstAuthor="Jeng" pageId="78" pageNumber="79" pagination="247 - 262" refId="ref118190" refString="Jeng, M. L., Yang, P. S. & Lai, J. (2003 b) Notes on the genus Luciola (Coleoptera, Lampyridae, Luciolinae) of Taiwan. Special Bulletin of the Japanese Society of Coleopterolog y, 6, 247 - 262." type="journal article" year="2003" yearSuffix="b">
|
||
Jeng
|
||
<emphasis id="D6E0C41851751818FE19FF7DEE891CBC" box="[384,433,225,251]" italics="true" pageId="78" pageNumber="79">et al</emphasis>
|
||
. 2003b
|
||
</bibRefCitation>
|
||
figs 5, 7; figs): without cuticular remnants in association with aedeagal sheath; no ventrites with curved posterior margins nor extending anteriorly into emarginated posterior margin of anterior segment; LO in
|
||
<date id="902A3ECA51751818FEB3FEB6EE741D04" box="[298,332,297,323]" pageId="78" pageNumber="79">V7</date>
|
||
entire, occupying almost all of
|
||
<date id="902A3ECA51751818FD2CFEB6EDE21D04" box="[693,730,297,323]" pageId="78" pageNumber="79">V7</date>
|
||
, and reaching to sides and posterior margin (
|
||
<bibRefCitation id="800565FB51751818FB41FEB6EAA41D04" author="Jeng" box="[1240,1436,297,323]" etAl="et al." firstAuthor="Jeng" pageId="78" pageNumber="79" pagination="247 - 262" refId="ref118190" refString="Jeng, M. L., Yang, P. S. & Lai, J. (2003 b) Notes on the genus Luciola (Coleoptera, Lampyridae, Luciolinae) of Taiwan. Special Bulletin of the Japanese Society of Coleopterolog y, 6, 247 - 262." type="journal article" year="2003" yearSuffix="b">
|
||
Jeng
|
||
<emphasis id="D6E0C41851751818FA8CFEB5EA7F1D04" box="[1301,1351,297,323]" italics="true" pageId="78" pageNumber="79">et al</emphasis>
|
||
. 2003b
|
||
</bibRefCitation>
|
||
fig.7); PLP not developed; posterior half of
|
||
<date id="902A3ECA51751818FD1EFED2ED911D20" box="[647,681,333,359]" pageId="78" pageNumber="79">V7</date>
|
||
<materialsCitation id="54FC125751751818FD28FED2ECD41DCC" ID-GBIF-Occurrence="2427485704" collectingDate="V 6" collectionCode="LO" pageId="78" pageNumber="79" specimenCount="1">
|
||
not arched or swollen, muscle impressions not visible in this area; neither anterior nor posterior margin of
|
||
<collectionCode id="828580CF51751818FDFDFEEEEDB11DCC" box="[612,649,369,395]" country="Sweden" name="Type Collection" pageId="78" pageNumber="79">LO</collectionCode>
|
||
emarginate;
|
||
<collectionCode id="828580CF51751818FCB9FEEEEC7D1DCC" box="[800,837,369,395]" country="Sweden" name="Type Collection" pageId="78" pageNumber="79">LO</collectionCode>
|
||
present in
|
||
<date id="902A3ECA51751818FC51FEEEECD41DCC" box="[968,1004,369,395]" pageId="78" pageNumber="79">
|
||
<collectingDate id="806EC72251751818FC51FEEEECD41DCC" box="[968,1004,369,395]" pageId="78" pageNumber="79">V6</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
, occupying almost all
|
||
<date id="902A3ECA51751818FB6BFEEEEA231DCC" box="[1266,1307,369,395]" pageId="78" pageNumber="79">V6.</date>
|
||
<materialsCitation id="54FC125751751818FABDFEEEEEB61DE8" ID-GBIF-Occurrence="2427485691" collectingDate="V 5." collectionCode="LO" pageId="78" pageNumber="79" specimenCount="1">
|
||
Functional
|
||
<collectionCode id="828580CF51751818FF0EFE0AEF841DE8" box="[151,188,405,431]" country="Sweden" name="Type Collection" pageId="78" pageNumber="79">LO</collectionCode>
|
||
not present in
|
||
<date id="902A3ECA51751818FEFCFE0BEEB61DE8" box="[357,398,404,431]" pageId="78" pageNumber="79">
|
||
<collectingDate id="806EC72251751818FEFCFE0BEEB61DE8" box="[357,398,404,431]" pageId="78" pageNumber="79">V5.</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
<materialsCitation id="54FC125751751818FE0FFE0AEDB31DB0" ID-GBIF-Occurrence="2427485709" collectingDate="V 7" collectionCode="MPP" pageId="78" pageNumber="79" specimenCount="1">
|
||
<collectionCode id="828580CF51751818FE0FFE0AEEF51DE8" box="[406,461,405,431]" pageId="78" pageNumber="79">MPP</collectionCode>
|
||
present, symmetrical, apex slightly medianly emarginated (
|
||
<bibRefCitation id="800565FB51751818FBF7FE0AEA091DE8" author="Jeng" box="[1134,1329,405,431]" etAl="et al." firstAuthor="Jeng" pageId="78" pageNumber="79" pagination="247 - 262" refId="ref118190" refString="Jeng, M. L., Yang, P. S. & Lai, J. (2003 b) Notes on the genus Luciola (Coleoptera, Lampyridae, Luciolinae) of Taiwan. Special Bulletin of the Japanese Society of Coleopterolog y, 6, 247 - 262." type="journal article" year="2003" yearSuffix="b">
|
||
Jeng
|
||
<emphasis id="D6E0C41851751818FB33FE09EBE41DE8" box="[1194,1244,405,431]" italics="true" pageId="78" pageNumber="79">et al</emphasis>
|
||
. 2003b
|
||
</bibRefCitation>
|
||
fig. 7) or squarely truncate, not laterally compressed, short (
|
||
<collectionCode id="828580CF51751818FD4FFE26EDD11D94" box="[726,745,441,467]" pageId="78" pageNumber="79">L</collectionCode>
|
||
=
|
||
<collectionCode id="828580CF51751818FD60FE26EC2F1D94" box="[761,791,441,467]" pageId="78" pageNumber="79">W</collectionCode>
|
||
), not inclined dorsally nor engulfed by T8 apex, without dorsal ridge, median longitudinal trough.
|
||
<date id="902A3ECA51751818FDF0FE42EDB31DB0" box="[617,651,477,503]" pageId="78" pageNumber="79">
|
||
<collectingDate id="806EC72251751818FDF0FE42EDB31DB0" box="[617,651,477,503]" pageId="78" pageNumber="79">V7</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
<materialsCitation id="54FC125751751818FD0BFE42EFE01EEC" ID-GBIF-Occurrence="2427485676" collectingDate="V 7" collectionCode="LO" pageId="78" pageNumber="79" specimenCount="1">
|
||
without median carina, median longitudinal trough, anteromedian depression on face of
|
||
<collectionCode id="828580CF51751818FEE8FD9EEEA11E5C" box="[369,409,513,539]" country="Sweden" name="Type Collection" pageId="78" pageNumber="79">LO</collectionCode>
|
||
, incurving lobes or pointed projections, median ‘dimple’, or reflexed lobes. T7 without prolonged anterolateral corners. T8: (
|
||
<bibRefCitation id="800565FB51751818FD8DFDBAEDEE1E78" author="Jeng" box="[532,726,549,575]" etAl="et al." firstAuthor="Jeng" pageId="78" pageNumber="79" pagination="247 - 262" refId="ref118190" refString="Jeng, M. L., Yang, P. S. & Lai, J. (2003 b) Notes on the genus Luciola (Coleoptera, Lampyridae, Luciolinae) of Taiwan. Special Bulletin of the Japanese Society of Coleopterolog y, 6, 247 - 262." type="journal article" year="2003" yearSuffix="b">
|
||
Jeng
|
||
<emphasis id="D6E0C41851751818FDC9FDB9EDBA1E78" box="[592,642,549,575]" italics="true" pageId="78" pageNumber="79">et al</emphasis>
|
||
. 2003b
|
||
</bibRefCitation>
|
||
fig. 6) symmetrical, W=L, visible posterior area not narrowing abruptly, median posterior margin shallowly and narrowly emarginate; T8 lateral margins subparallel-sided; without prolonged posterolateral corners, median posterior projections, not inclined ventrally nor engulfing posterior margin of
|
||
<date id="902A3ECA51751818FF2FFD0EEFE01EEC" box="[182,216,657,683]" pageId="78" pageNumber="79">
|
||
<collectingDate id="806EC72251751818FF2FFD0EEFE01EEC" box="[182,216,657,683]" pageId="78" pageNumber="79">V7</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
<materialsCitation id="54FC125751751818FF78FD0EECC61EEC" ID-GBIF-Occurrence="2427485674" box="[225,1022,657,683]" collectingDate="V 7" collectionCode="MPP" pageId="78" pageNumber="79" specimenCount="1">
|
||
nor
|
||
<collectionCode id="828580CF51751818FE97FD0EEE7E1EEC" box="[270,326,657,683]" pageId="78" pageNumber="79">MPP</collectionCode>
|
||
, not extending conspicuously beyond posterior margin of
|
||
<date id="902A3ECA51751818FC40FD0EECC61EEC" box="[985,1022,657,683]" pageId="78" pageNumber="79">
|
||
<collectingDate id="806EC72251751818FC40FD0EECC61EEC" box="[985,1022,657,683]" pageId="78" pageNumber="79">V7</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
; T8 ventral surface without well-developed median longitudinal trough, without lateral depressed troughs, asymmetrical projections, median posterior ridge; concealed anterolateral arms of T8 very short and wide (
|
||
<bibRefCitation id="800565FB51751818FCFBFD46EB1E1EB4" author="Jeng" box="[866,1062,729,755]" etAl="et al." firstAuthor="Jeng" pageId="78" pageNumber="79" pagination="247 - 262" refId="ref118190" refString="Jeng, M. L., Yang, P. S. & Lai, J. (2003 b) Notes on the genus Luciola (Coleoptera, Lampyridae, Luciolinae) of Taiwan. Special Bulletin of the Japanese Society of Coleopterolog y, 6, 247 - 262." type="journal article" year="2003" yearSuffix="b">
|
||
Jeng
|
||
<emphasis id="D6E0C41851751818FC06FD45ECE91EB4" box="[927,977,729,755]" italics="true" pageId="78" pageNumber="79">et al</emphasis>
|
||
. 2003b
|
||
</bibRefCitation>
|
||
fig. 6), not laterally emarginated before their origins, not expanded dorsoventrally, expanded only in horizontal plane; without bifurcation of inner margin and ventrally directed pieces; lateral margins of T8 not enfolding sides of
|
||
<date id="902A3ECA51751818FBB1FCBEEB6A1F7C" box="[1064,1106,801,827]" pageId="78" pageNumber="79">V7.</date>
|
||
</paragraph>
|
||
<paragraph id="E42B180A51751818FF5EFCDAEBE91F8C" blockId="78.[151,1437,153,1763]" pageId="78" pageNumber="79">
|
||
Aedeagal sheath: (
|
||
<bibRefCitation id="800565FB51751818FE0FFCDAED6F1F18" author="Jeng" box="[406,599,837,863]" etAl="et al." firstAuthor="Jeng" pageId="78" pageNumber="79" pagination="247 - 262" refId="ref118190" refString="Jeng, M. L., Yang, P. S. & Lai, J. (2003 b) Notes on the genus Luciola (Coleoptera, Lampyridae, Luciolinae) of Taiwan. Special Bulletin of the Japanese Society of Coleopterolog y, 6, 247 - 262." type="journal article" year="2003" yearSuffix="b">
|
||
Jeng
|
||
<emphasis id="D6E0C41851751818FE4BFCD9ED3B1F18" box="[466,515,837,863]" italics="true" pageId="78" pageNumber="79">et al</emphasis>
|
||
. 2003b
|
||
</bibRefCitation>
|
||
fig. 10) approx. 4 times as long as wide; without bulbous paraprocts; symmetrical in posterior area where sheath sternite tapers evenly to a narrow, rounded apex; anterior half of sternite relatively narrow, apically rounded; tergite without lateral arms extending anteriorly at sides of sheath sternite; tergite without projecting pieces along posterior margin of T9, anterior margin without transverse band.
|
||
</paragraph>
|
||
<paragraph id="E42B180A51751818FF5EFC4AEB1B1948" blockId="78.[151,1437,153,1763]" pageId="78" pageNumber="79">
|
||
Aedeagus: (
|
||
<bibRefCitation id="800565FB51751818FED4FC4AED371FA8" author="Jeng" box="[333,527,981,1007]" etAl="et al." firstAuthor="Jeng" pageId="78" pageNumber="79" pagination="247 - 262" refId="ref118190" refString="Jeng, M. L., Yang, P. S. & Lai, J. (2003 b) Notes on the genus Luciola (Coleoptera, Lampyridae, Luciolinae) of Taiwan. Special Bulletin of the Japanese Society of Coleopterolog y, 6, 247 - 262." type="journal article" year="2003" yearSuffix="b">
|
||
Jeng
|
||
<emphasis id="D6E0C41851751818FE10FC49EE831FA8" box="[393,443,981,1007]" italics="true" pageId="78" pageNumber="79">et al</emphasis>
|
||
. 2003b
|
||
</bibRefCitation>
|
||
figs 11A, B) L/W 3.0; LL lack lateral appendages; apices of LL not visible from beneath at sides of ML, LL/ML narrow; LL of equal length, slightly shorter than ML, contiguous or closely approaching along inner dorsal margins; LL narrowly separated longitudinally by most of their length; LL base width not = LL apex width which is slightly narrower than that of ML; LL apices not expanded in horizontal plane; dorsal base of LL symmetrical, not excavated; LL without lateral hairy appendages along their outer ventral margins, not produced preapically nor narrowly on inner apical margin, apices of LL not inturned, nor out-turned; without projection on left LL; inner margins without slender leaf-like projection; ML symmetrical, without paired lateral teeth and tooth to left side, not strongly arched, apex not shaped like arrowhead, not bulbous, not inclined ventrally; BP not strongly sclerotised, not hooded, not strongly emarginated along anterior margin.
|
||
</paragraph>
|
||
<paragraph id="E42B180A51751818FF5EFA86EB3F1A68" blockId="78.[151,1437,153,1763]" pageId="78" pageNumber="79">
|
||
<emphasis id="D6E0C41851751818FF5EFA86EE261974" bold="true" box="[199,286,1305,1331]" pageId="78" pageNumber="79">Female</emphasis>
|
||
(
|
||
<bibRefCitation id="800565FB51751818FEB4FA86EED41974" author="Jeng" box="[301,492,1305,1331]" etAl="et al." firstAuthor="Jeng" pageId="78" pageNumber="79" pagination="247 - 262" refId="ref118190" refString="Jeng, M. L., Yang, P. S. & Lai, J. (2003 b) Notes on the genus Luciola (Coleoptera, Lampyridae, Luciolinae) of Taiwan. Special Bulletin of the Japanese Society of Coleopterolog y, 6, 247 - 262." type="journal article" year="2003" yearSuffix="b">
|
||
Jeng
|
||
<emphasis id="D6E0C41851751818FEF1FA85EEA11974" box="[360,409,1305,1331]" italics="true" pageId="78" pageNumber="79">et al</emphasis>
|
||
. 2003b
|
||
</bibRefCitation>
|
||
fig. 8). Macropterous and observed in flight; (
|
||
<bibRefCitation id="800565FB51751818FC69FA86EBE21974" author="Jeng" box="[1008,1242,1304,1331]" etAl="et al." firstAuthor="Jeng" pageId="78" pageNumber="79" pagination="1 - 572" refId="ref118113" refString="Jeng, M. L., Lai, J. & Yang, P. S. (2003 a) Lampyridae: A synopsis of aquatic fireflies with description of a new species. In: Jach, M. A. & Ji, L. (Eds.), Water Beetles of China III. Zoologisch-Botanische Gesellschaft in Osterreich, and Wiener Coleopterologenverein, Wien, pp. 1 - 572." type="book chapter" year="2003">
|
||
Jeng
|
||
<emphasis id="D6E0C41851751818FBB5FA85EB651974" box="[1068,1117,1305,1331]" italics="true" pageId="78" pageNumber="79">et al</emphasis>
|
||
. 2003: 251
|
||
</bibRefCitation>
|
||
observed a flight period beginning just after sunset, with peak numbers reached after about 25 minutes and cessation of activity after another five minutes). Pronotum without irregularities in posterolateral areas; punctation moderate to dense; pronotal width less than humeral width; without indentation of lateral margin, irregularities at posterolateral corner; outline similar to that of male. Elytral punctation not as large as that of pronotum, nor evenly spaced; no interstitial lines; elytral carina absent. No legs or parts thereof swollen and /or curved. LO in
|
||
<date id="902A3ECA51751818FBAFFA52EB6019A0" box="[1078,1112,1485,1511]" pageId="78" pageNumber="79">V6</date>
|
||
only, without any elevations or depressions or ridges on
|
||
<date id="902A3ECA51751818FE53FA6EEED71A4C" box="[458,495,1521,1547]" pageId="78" pageNumber="79">V7</date>
|
||
; median posterior margin of
|
||
<date id="902A3ECA51751818FCA8FA6EEC6B1A4C" box="[817,851,1521,1547]" pageId="78" pageNumber="79">V7</date>
|
||
widely and very shallowly emarginate; median posterior margin of
|
||
<date id="902A3ECA51751818FED7F98AEE481A68" box="[334,368,1557,1583]" pageId="78" pageNumber="79">V8</date>
|
||
very shallowly emarginated. Bursa plates not investigated.
|
||
</paragraph>
|
||
<paragraph id="E42B180A51751818FF5EF9A6ED1F1A14" blockId="78.[151,1437,153,1763]" box="[199,551,1593,1619]" pageId="78" pageNumber="79">
|
||
<emphasis id="D6E0C41851751818FF5EF9A6EE2F1A14" bold="true" box="[199,279,1593,1619]" pageId="78" pageNumber="79">Larva.</emphasis>
|
||
Not reliably associated.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="AC8E4B8151751818FF5EF9C2EDB61ADC" pageId="78" pageNumber="79" type="etymology">
|
||
<paragraph id="E42B180A51751818FF5EF9C2EDB61ADC" blockId="78.[151,1437,153,1763]" pageId="78" pageNumber="79">
|
||
<emphasis id="D6E0C41851751818FF5EF9C2EE701A30" bold="true" box="[199,328,1629,1655]" pageId="78" pageNumber="79">Etymology</emphasis>
|
||
.
|
||
<taxonomicName id="2394638951751818FECEF9C1EEE41A30" authorityName="Ho" authorityYear="2019" box="[343,476,1630,1655]" class="Aves" family="Muscicapidae" genus="Emarginata" kingdom="Animalia" order="Passeriformes" pageId="78" pageNumber="79" phylum="Chordata" rank="genus">
|
||
<emphasis id="D6E0C41851751818FECEF9C1EEE41A30" box="[343,476,1630,1655]" italics="true" pageId="78" pageNumber="79">Emarginata</emphasis>
|
||
</taxonomicName>
|
||
(feminine) is a noun latinised from the English word emarginate reflecting the nature of the emarginated elytral apices in the male.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="AC8E4B8151751818FF5EF93AEE571AA4" pageId="78" pageNumber="79" type="discussion">
|
||
<paragraph id="E42B180A51751818FF5EF93AEE571AA4" blockId="78.[151,1437,153,1763]" pageId="78" pageNumber="79">
|
||
<emphasis id="D6E0C41851751818FF5EF93AEE031AF8" bold="true" box="[199,315,1701,1727]" pageId="78" pageNumber="79">Remarks.</emphasis>
|
||
The distinctiveness of this species was established from phylogenetic inference from prior analysis (
|
||
<bibRefCitation id="800565FB51751818FF06F956EE5B1AA4" author="Jusoh" box="[159,355,1737,1763]" etAl="et al." firstAuthor="Jusoh" pageId="78" pageNumber="79" pagination="1 - 71" refId="ref118381" refString="Jusoh, W. F. A., Ballantyne, L., Lambkin, C. L., Hashim, N. R. & Wahlberg, N. (2018) The firefly genus Pteroptyx Olivier revisited (Coleoptera: Lampyridae: Luciolinae). Zootaxa, 4456 (1), 1 - 71. https: // doi. org / 10.11646 / zootaxa. 4456.1.1" type="journal article" year="2018">
|
||
Jusoh
|
||
<emphasis id="D6E0C41851751818FF7EF955EE181AA4" box="[231,288,1737,1763]" italics="true" pageId="78" pageNumber="79">et al.</emphasis>
|
||
2018
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |