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<document id="9BF2B729E638B33CAFE3D412F76C84A5" ID-CLB-Dataset="25190" ID-DOI="10.11646/zootaxa.4687.1.1" ID-GBIF-Dataset="de00ca6c-2843-4ae1-a39c-a29603947612" ID-ISSN="1175-5326" ID-Zenodo-Dep="3508098" ID-ZooBank="CE73264D-C234-4B82-A634-CAD6254C5957" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1571388220410" checkinUser="plazi" docAuthor="Ho, - Z." docDate="2019" docId="6C3DA91C516D1801FF0EF9F5EAA41845" docLanguage="en" docName="zootaxa.4687.1.1.pdf.imf" docOrigin="Zootaxa 4687 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Luciola Ho 2019, s. str." docType="key" docVersion="19" lastPageNumber="88" masterDocId="9004D164513B1856FF99FF9FEF381C47" masterDocTitle="The Luciolinae of S. E. Asia and the Australopacific region: a revisionary checklist (Coleoptera: Lampyridae) including description of three new genera and 13 new species" masterLastPageNumber="174" masterPageNumber="1" pageNumber="87" updateTime="1698760145766" updateUser="ExternalLinkService">
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<mods:title id="326F8413602C578DF095749472CC40F1">The Luciolinae of S. E. Asia and the Australopacific region: a revisionary checklist (Coleoptera: Lampyridae) including description of three new genera and 13 new species</mods:title>
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<treatment id="6C3DA91C516D1801FF0EF9F5EAA41845" ID-DOI="http://doi.org/10.5281/zenodo.4688869" ID-GBIF-Taxon="159757343" ID-Zenodo-Dep="4688869" LSID="urn:lsid:plazi:treatment:6C3DA91C516D1801FF0EF9F5EAA41845" httpUri="http://treatment.plazi.org/id/6C3DA91C516D1801FF0EF9F5EAA41845" lastPageId="87" lastPageNumber="88" pageId="86" pageNumber="87">
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<paragraph id="E42B180A516D1800FF0EF9F5EBA21AC3" blockId="86.[151,1178,1641,1668]" box="[151,1178,1641,1668]" pageId="86" pageNumber="87">
<heading id="BF63AF66516D1800FF0EF9F5EBA21AC3" bold="true" box="[151,1178,1641,1668]" fontSize="11" level="1" pageId="86" pageNumber="87" reason="1">
<emphasis id="D6E0C418516D1800FF0EF9F5EBA21AC3" bold="true" box="[151,1178,1641,1668]" pageId="86" pageNumber="87">
Key to species of
<taxonomicName id="23946389516D1800FEF7F9F6EEF31AC3" ID-CoL="5H69" authority="Ho, 2019" authorityName="Ho" authorityYear="2019" box="[366,459,1641,1668]" class="Insecta" family="Lampyridae" genus="Luciola" kingdom="Animalia" order="Coleoptera" pageId="86" pageNumber="87" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="D6E0C418516D1800FEF7F9F6EEF31AC3" bold="true" box="[366,459,1641,1668]" italics="true" pageId="86" pageNumber="87">Luciola</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CDD37963516D1800FE4BF9F5ED2B1AC3" box="[466,531,1642,1668]" pageId="86" pageNumber="87" sensu="stricto">s. str.</taxonomicNameLabel>
as defined here, using males and associated females
</emphasis>
</heading>
</paragraph>
</subSubSection>
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<paragraph id="E42B180A516D1800FF0EF931EAA41AA6" blockId="86.[151,1436,1710,2026]" pageId="86" pageNumber="87">
1. Male...............................................................................................
<specimenCount id="F292D383516D1800FA09F931EE111AA5" pageId="86" pageNumber="87" type="female">2 Female</specimenCount>
............................................................................................ 14
</paragraph>
<paragraph id="E42B180A516D1800FF0EF976EAA41B7E" blockId="86.[151,1436,1710,2026]" pageId="86" pageNumber="87">
2. Light organs absent; very dark brown to black species with pronounced posterolateral eye emargination; known only from
<collectingCountry id="9C83589A516D1800FAF6F976EE7B1B5B" name="New Caledonia" pageId="86" pageNumber="87">New Caledonia</collectingCountry>
(
<bibRefCitation id="800565FB516D1800FEC9F899ED5C1B5C" author="Ballantyne &amp; Lambkin" box="[336,612,1798,1820]" firstAuthor="Ballantyne" pageId="86" pageNumber="87" pagination="1 - 162" refId="ref114597" refString="Ballantyne, L. A. &amp; Lambkin, C. L. (2013) Systematics and phylogenetics of Indo-Pacific Luciolinae fireflies (Coleoptera: Lampyridae) and the description of new genera. Zootaxa, 3653 (1), 1 - 162. https: // doi. org / 10.11646 / zootaxa. 3653.1.1" type="journal article" year="2013">Ballantyne &amp; Lambkin 2013</bibRefCitation>
figs 120 121, 124, 125)...............................
<taxonomicName id="23946389516D1800FB54F899EE2E1B7D" ID-CoL="6R2BZ" authority="Ballantyne Light" authorityName="Ballantyne Light" authorityYear="2013" class="Insecta" family="Lampyridae" genus="Luciola" kingdom="Animalia" order="Coleoptera" pageId="86" pageNumber="87" phylum="Arthropoda" rank="species" species="oculofissa">
<emphasis id="D6E0C418516D1800FB54F899EA161B5C" box="[1229,1326,1798,1819]" italics="true" pageId="86" pageNumber="87">oculofissa</emphasis>
Ballantyne Light
</taxonomicName>
organs present in at least one abdominal ventrite; eye emargination present or not............................. 3
</paragraph>
<paragraph id="E42B180A516D1800FF0EF8DEEAA41B33" blockId="86.[151,1436,1710,2026]" pageId="86" pageNumber="87">
3. Eyes with strong posterolateral emargination clearly visible in resting head (
<bibRefCitation id="800565FB516D1800FC24F8DEEBEA1B11" author="Ballantyne &amp; Lambkin" box="[957,1234,1857,1879]" firstAuthor="Ballantyne" pageId="86" pageNumber="87" pagination="1 - 162" refId="ref114597" refString="Ballantyne, L. A. &amp; Lambkin, C. L. (2013) Systematics and phylogenetics of Indo-Pacific Luciolinae fireflies (Coleoptera: Lampyridae) and the description of new genera. Zootaxa, 3653 (1), 1 - 162. https: // doi. org / 10.11646 / zootaxa. 3653.1.1" type="journal article" year="2013">Ballantyne &amp; Lambkin 2013</bibRefCitation>
fig. 110)....... 4 Eyes without a visible posterolateral emargination, or with a weak emargination usually covered by the pronotum........ 5
</paragraph>
<paragraph id="E42B180A516D1800FF0EF8E3EAA41B8B" blockId="86.[151,1436,1710,2026]" pageId="86" pageNumber="87">
4. No light organs in
<date id="902A3ECA516D1800FE0CF8E3EE8D1BD6" box="[405,437,1916,1937]" pageId="86" pageNumber="87">V7</date>
; light organs in
<date id="902A3ECA516D1800FDD6F8E3ED541BD5" box="[591,620,1916,1938]" pageId="86" pageNumber="87">V6</date>
reduced to anterolateral plaques; known only from
<collectingCountry id="9C83589A516D1800FBC4F8E3EB461BD5" box="[1117,1150,1916,1938]" name="Fiji" pageId="86" pageNumber="87">Fiji</collectingCountry>
.........
<taxonomicName id="23946389516D1800FB6FF8E3EE2E1BE8" ID-CoL="3WCRY" authority="Olivier Light" authorityName="Olivier Light" authorityYear="1888" class="Insecta" family="Lampyridae" genus="Luciola" kingdom="Animalia" order="Coleoptera" pageId="86" pageNumber="87" phylum="Arthropoda" rank="species" species="hypocrita">
<emphasis id="D6E0C418516D1800FB6FF8E3EA6A1BD6" box="[1270,1362,1916,1937]" italics="true" pageId="86" pageNumber="87">hypocrita</emphasis>
Olivier Light
</taxonomicName>
organs in both
<date id="902A3ECA516D1800FE33F806ED3A1BE9" box="[426,514,1945,1967]" pageId="86" pageNumber="87">V6 and 7</date>
entire, may be retracted from lateral margins (
<bibRefCitation id="800565FB516D1800FC58F806EBEE1BE9" author="Ballantyne &amp; Lambkin" box="[961,1238,1945,1967]" firstAuthor="Ballantyne" pageId="86" pageNumber="87" pagination="1 - 162" refId="ref114597" refString="Ballantyne, L. A. &amp; Lambkin, C. L. (2013) Systematics and phylogenetics of Indo-Pacific Luciolinae fireflies (Coleoptera: Lampyridae) and the description of new genera. Zootaxa, 3653 (1), 1 - 162. https: // doi. org / 10.11646 / zootaxa. 3653.1.1" type="journal article" year="2013">Ballantyne &amp; Lambkin 2013</bibRefCitation>
figs 114, 119).......................................................................................
<taxonomicName id="23946389516D1800FB54F829EAA41B8B" ID-CoL="3WCN7" authority="Bourgeois" authorityName="Bourgeois" authorityYear="1884" box="[1229,1436,1974,1996]" class="Insecta" family="Lampyridae" genus="Luciola" kingdom="Animalia" order="Coleoptera" pageId="86" pageNumber="87" phylum="Arthropoda" rank="species" species="antipodum">
<emphasis id="D6E0C418516D1800FB54F829EA0C1B8C" box="[1229,1332,1974,1995]" italics="true" pageId="86" pageNumber="87">antipodum</emphasis>
Bourgeois
</taxonomicName>
</paragraph>
<paragraph id="E42B180A516D1801FF0EF84BEAA41CE8" blockId="86.[151,1436,1710,2026]" lastBlockId="87.[151,1437,153,1026]" lastPageId="87" lastPageNumber="88" pageId="86" pageNumber="87">5. Dorsal surface completely black or very dark brown.......................................................... 6 Dorsal surface never completely dark brown; pronotum usually pale coloured sometimes with a median dark marking..... 7</paragraph>
<paragraph id="E42B180A516C1801FF0EFF28EAA51D05" blockId="87.[151,1437,153,1026]" pageId="87" pageNumber="88">
6. Known only from
<collectingCountry id="9C83589A516C1801FE0EFF28ED081C8A" box="[407,560,183,205]" name="New Caledonia" pageId="87" pageNumber="88">New Caledonia</collectingCountry>
; aedeagal LL widely separated along their inner dorsal margins with a rounded projection along inner dorsal margins approximately half their length from their apices (
<bibRefCitation id="800565FB516C1801FC5EFF4BEBE41CAE" author="Ballantyne &amp; Lambkin" box="[967,1244,212,234]" firstAuthor="Ballantyne" pageId="87" pageNumber="88" pagination="1 - 162" refId="ref114597" refString="Ballantyne, L. A. &amp; Lambkin, C. L. (2013) Systematics and phylogenetics of Indo-Pacific Luciolinae fireflies (Coleoptera: Lampyridae) and the description of new genera. Zootaxa, 3653 (1), 1 - 162. https: // doi. org / 10.11646 / zootaxa. 3653.1.1" type="journal article" year="2013">Ballantyne &amp; Lambkin 2013</bibRefCitation>
figs 111, 112)......................................................................................
<taxonomicName id="23946389516C1801FB27FF6EEE111D63" ID-CoL="3WCNB" authority="Ballantyne A S. E." authorityName="Ballantyne A S. E." class="Insecta" family="Lampyridae" genus="Luciola" kingdom="Animalia" order="Coleoptera" pageId="87" pageNumber="88" phylum="Arthropoda" rank="species" species="aquilaclara">
<emphasis id="D6E0C418516C1801FB27FF6EEA161D41" box="[1214,1326,241,262]" italics="true" pageId="87" pageNumber="88">aquilaclara</emphasis>
Ballantyne A S. E.
</taxonomicName>
Asian species sometimes from high altitudes; aedeagal LL not widely separated along their inner dorsal margins and without rounded projection as described above (
<figureCitation id="7CAF048F516C1801FD29FEB3EC391D05" box="[688,769,300,322]" captionStart="FIGURES 305308" captionStartId="98.[151,264,950,975]" captionTargetBox="[151,1435,166,924]" captionTargetId="figure@98.[151,1435,166,930]" captionTargetPageId="98" captionText="FIGURES 305308. Luciola pallidipes male. 305308 aedeagal sheath: 305 dorsal (arrow indicates prolonged margin of sheath tergite; aedeagus removed); 306308 aedeagal sheath with aedeagus: 306 right lateral (arrow indicates prolonged margin of sheath tergite); 307 dorsal; 308 ventral (arrows indicate leaf like lobes from LL of aedeagus). Scale lines are 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3508218" httpUri="https://zenodo.org/record/3508218/files/figure.png" pageId="87" pageNumber="88">Fig. 307</figureCitation>
)...........................................
<taxonomicName id="23946389516C1801FA80FEB3EAA51D05" ID-CoL="72QC9" authority="Pic" authorityName="Pic" authorityYear="1928" box="[1305,1437,300,322]" class="Insecta" family="Lampyridae" genus="Luciola" kingdom="Animalia" order="Coleoptera" pageId="87" pageNumber="88" phylum="Arthropoda" rank="species" species="pallidipes">
<emphasis id="D6E0C418516C1801FA80FEB3EA401D06" box="[1305,1400,300,321]" italics="true" pageId="87" pageNumber="88">pallidipes</emphasis>
Pic
</taxonomicName>
</paragraph>
<paragraph id="E42B180A516C1801FF0EFED5EAA41D3B" blockId="87.[151,1437,153,1026]" pageId="87" pageNumber="88">7. Dorsal surface pale yellowish brown with dark elytral apices....................................................8 Dorsal surface not as above; pronotum may be black; elytra usually dark brown to black............................. 9</paragraph>
<paragraph id="E42B180A516C1801FF0EFE1BEAA41DB6" blockId="87.[151,1437,153,1026]" pageId="87" pageNumber="88">
8. Ventral abdomen with median brown markings on
<date id="902A3ECA516C1801FD50FE1BEDC61DDD" box="[713,766,388,410]" pageId="87" pageNumber="88">V45</date>
with lateral margins converging anteriorly (
<figureCitation id="7CAF048F516C1801FB3CFE1BEA131DDD" box="[1189,1323,388,410]" captionStart="FIGURES 270279" captionStartId="93.[151,264,1924,1949]" captionTargetBox="[203,1385,194,1903]" captionTargetId="figure@93.[203,1385,166,1905]" captionTargetPageId="93" captionText="FIGURES 270279. Luciola jengai sp. nov. type males. 270, 273 habitus dorsal; 271, 272 habitus ventral; 274, 275 abdomen: 274 dorsal; 275 ventral; 276 tergite 8 and aedeagal sheath ventral; 277279 aedeagus: 277 dorsal; 279 ventral; 279 left lateral. Scale lines are 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3508206" httpUri="https://zenodo.org/record/3508206/files/figure.png" pageId="87" pageNumber="88">Figs 271, 275</figureCitation>
)...........................................................................................
<taxonomicName id="23946389516C1801FB4CFE3DEA721DF0" ID-CoL="8NC6S" authority="Nada" authorityName="Ho" authorityYear="2019" box="[1237,1354,417,439]" class="Insecta" family="Lampyridae" genus="Luciola" kingdom="Animalia" order="Coleoptera" pageId="87" pageNumber="88" phylum="Arthropoda" rank="species" species="jengai" status="sp. nov.">
<emphasis id="D6E0C418516C1801FB4CFE3DEA291DF0" box="[1237,1297,418,439]" italics="true" pageId="87" pageNumber="88">jengai</emphasis>
Nada
</taxonomicName>
<emphasis id="D6E0C418516C1801FAC9FE3DEAA41DF0" bold="true" box="[1360,1436,418,439]" pageId="87" pageNumber="88">
<taxonomicNameLabel id="CDD37963516C1801FAC9FE3DEAA41DF0" box="[1360,1436,418,439]" pageId="87" pageNumber="88" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Ventral abdomen not coloured as above; ventrites anterior to the LO yellowish except for dark posterior margin of
<date id="902A3ECA516C1801FAD0FE20EA5F1D92" box="[1353,1383,447,469]" pageId="87" pageNumber="88">V5</date>
(Fig. 285).................................................................................
<taxonomicName id="23946389516C1801FB7BFE43EA721DB5" ID-CoL="8NC2B" authority="Jusoh" authorityName="Ho" authorityYear="2019" box="[1250,1354,476,498]" class="Insecta" family="Lampyridae" genus="Luciola" kingdom="Animalia" order="Coleoptera" pageId="87" pageNumber="88" phylum="Arthropoda" rank="species" species="niah" status="sp. nov.">
<emphasis id="D6E0C418516C1801FB7BFE43EA351DB6" box="[1250,1293,476,497]" italics="true" pageId="87" pageNumber="88">niah</emphasis>
Jusoh
</taxonomicName>
<emphasis id="D6E0C418516C1801FAC9FE43EAA41DB6" bold="true" box="[1360,1436,476,497]" pageId="87" pageNumber="88">
<taxonomicNameLabel id="CDD37963516C1801FAC9FE43EAA41DB6" box="[1360,1436,476,497]" pageId="87" pageNumber="88" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="E42B180A516C1801FF0EFE65EAA41E0E" blockId="87.[151,1437,153,1026]" pageId="87" pageNumber="88">
9. Pronotum MN and MS black; elytra light brown with extensive darker brown markings at base and apex (
<figureCitation id="7CAF048F516C1801FB40FE66EA671E57" box="[1241,1375,505,528]" captionStart="FIGURES 309314" captionStartId="100.[151,264,1754,1779]" captionTargetBox="[152,1435,166,1723]" captionTargetId="figure@100.[151,1435,166,1734]" captionTargetPageId="100" captionText="FIGURES 309314. Luciola tiomana sp. nov. type males. 309, 310 habitus dorsal (309 Prague; 310 FRIM); 311 habitus ventral; 312, 313 abdomen: 312 ventral; 313 dorsal; 314 tergite 8, aedeagal sheath and aedeagus, dorsal. Scale line is 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3508222" httpUri="https://zenodo.org/record/3508222/files/figure.png" pageId="87" pageNumber="88">Figs 309, 310</figureCitation>
)................................................................................
<taxonomicName id="23946389516C1801FB15FD88EA701E6A" ID-CoL="8NBWZ" authority="Ballantyne" authorityName="Ho" authorityYear="2019" box="[1164,1352,535,557]" class="Insecta" family="Lampyridae" genus="Luciola" kingdom="Animalia" order="Coleoptera" pageId="87" pageNumber="88" phylum="Arthropoda" rank="species" species="tiomana" status="sp. nov.">
<emphasis id="D6E0C418516C1801FB15FD88EBE31E6B" box="[1164,1243,535,556]" italics="true" pageId="87" pageNumber="88">tiomana</emphasis>
Ballantyne
</taxonomicName>
<taxonomicNameLabel id="CDD37963516C1801FAD7FD88EAA41E6B" box="[1358,1436,535,556]" pageId="87" pageNumber="88" rank="species">
<emphasis id="D6E0C418516C1801FAD7FD88EAAE1E6B" bold="true" box="[1358,1430,535,556]" pageId="87" pageNumber="88">sp. nov</emphasis>
.
</taxonomicNameLabel>
Dorsal surface with elytra black and pronotum pale coloured sometimes with a median dark marking.................. 10
</paragraph>
<paragraph id="E42B180A516C1801FF0EFDCDEAA51EC3" blockId="87.[151,1437,153,1026]" pageId="87" pageNumber="88">10. Pronotum uniformly pale coloured with no darker markings.................................................. 11 Pronotum always with median darker markings.............................................................12</paragraph>
<paragraph id="E42B180A516C1801FF0EFD13EAA41EF8" blockId="87.[151,1437,153,1026]" pageId="87" pageNumber="88">
11. Large thick bodied, 10.5
<quantity id="236CB5EF516C1801FE45FD13ED251EE5" box="[476,541,652,674]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="87" pageNumber="88" unit="mm" value="15.0">15 mm</quantity>
long (
<bibRefCitation id="800565FB516C1801FDC2FD13EDCF1EE5" author="Chen" box="[603,759,652,674]" firstAuthor="Chen" pageId="87" pageNumber="88" refId="ref115752" refString="Chen, T. R. (2003) The fireflies of Taiwan. Field Image Publisher, Taipei, 256 pp. [in Chinese]" type="book" year="2003">Chen 2003: 168</bibRefCitation>
)...................................... ..
<taxonomicName id="23946389516C1801FB45FD13EE161EF8" ID-CoL="3WCSP" authority="Matsumura Smaller" authorityName="Matsumura Smaller" authorityYear="1928" class="Insecta" family="Lampyridae" genus="Luciola" kingdom="Animalia" order="Coleoptera" pageId="87" pageNumber="88" phylum="Arthropoda" rank="species" species="kagiana">
<emphasis id="D6E0C418516C1801FB45FD13EA111EE6" box="[1244,1321,652,673]" italics="true" pageId="87" pageNumber="88">kagiana</emphasis>
Matsumura Smaller
</taxonomicName>
, 6.5
<quantity id="236CB5EF516C1801FEFDFD36EE9E1EF8" box="[356,422,681,703]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="87" pageNumber="88" unit="mm" value="10.0">10 mm</quantity>
long (
<bibRefCitation id="800565FB516C1801FE7DFD36ED461EF8" author="Chen" box="[484,638,681,703]" firstAuthor="Chen" pageId="87" pageNumber="88" refId="ref115752" refString="Chen, T. R. (2003) The fireflies of Taiwan. Field Image Publisher, Taipei, 256 pp. [in Chinese]" type="book" year="2003">Chen 2003: 169</bibRefCitation>
as
<taxonomicName id="23946389516C1801FD07FD36EDDF1EF9" ID-CoL="5H69" box="[670,743,681,702]" class="Insecta" family="Lampyridae" genus="Luciola" kingdom="Animalia" order="Coleoptera" pageId="87" pageNumber="88" phylum="Arthropoda" rank="genus">
<emphasis id="D6E0C418516C1801FD07FD36EDDF1EF9" box="[670,743,681,702]" italics="true" pageId="87" pageNumber="88">Luciola</emphasis>
</taxonomicName>
sp. 3)........................................
<taxonomicName id="23946389516C1801FA9FFD35EAA41EF8" ID-CoL="4QP8W" authority="Jeng et al." authorityName="Jeng" box="[1286,1436,681,703]" class="Insecta" family="Lampyridae" genus="Pygoluciola" kingdom="Animalia" order="Coleoptera" pageId="87" pageNumber="88" phylum="Arthropoda" rank="species" species="satoi">
<emphasis id="D6E0C418516C1801FA9FFD35EA0D1EF8" box="[1286,1333,682,703]" italics="true" pageId="87" pageNumber="88">satoi</emphasis>
Jeng
<emphasis id="D6E0C418516C1801FAF5FD35EAAF1EF9" box="[1388,1431,681,703]" italics="true" pageId="87" pageNumber="88">et al</emphasis>
.
</taxonomicName>
</paragraph>
<paragraph id="E42B180A516C1801FF0EFD58EAA41F50" blockId="87.[151,1437,153,1026]" pageId="87" pageNumber="88">
12. Pronotum orange with darker median markings; pronounced tubercles along posterior margin (
<bibRefCitation id="800565FB516C1801FBF6FD58EBF11E9B" author="Yiu" box="[1135,1225,711,733]" firstAuthor="Yiu" pageId="87" pageNumber="88" pagination="67 - 119" refId="ref122609" refString="Yiu, V. (2017) A Study of Rhagophthalmidae and Lampyridae in Hong Kong with descriptions of new species (Coleoptera): Part 2. Lampyrid, 4, 67 - 119." type="journal article" year="2017">Yiu 2017</bibRefCitation>
fig. 22f)................................................................................................ ..
<taxonomicName id="23946389516C1801FA9EFD7BEFCA1F50" ID-CoL="8N98Z" authority="Yiu If" authorityName="Yiu If" authorityYear="2017" class="Insecta" family="Lampyridae" genus="Luciola" kingdom="Animalia" order="Coleoptera" pageId="87" pageNumber="88" phylum="Arthropoda" rank="species" species="tuberculata">
<emphasis id="D6E0C418516C1801FA9EFD7BEA4D1EBE" box="[1287,1397,740,761]" italics="true" pageId="87" pageNumber="88">tuberculata</emphasis>
Yiu If
</taxonomicName>
pronotum with darker median markings then no tubercles across posterior margin............................... 13
</paragraph>
<paragraph id="E42B180A516C1801FF0EFC80EAA41FCB" blockId="87.[151,1437,153,1026]" pageId="87" pageNumber="88">
13. Elytra dark brown to black with dark apices;
<quantity id="236CB5EF516C1801FD0FFC80EDF61F73" box="[662,718,799,821]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="87" pageNumber="88" unit="mm" value="6.0">6 mm</quantity>
long species from
<collectingCountry id="9C83589A516C1801FC1FFC80ECDD1F72" box="[902,997,799,821]" name="Sri Lanka" pageId="87" pageNumber="88">Sri Lanka</collectingCountry>
with dark brown elytra and orange pronotum having a wide median dark band (
<figureCitation id="7CAF048F516C1801FDBEFCA3EDE51F15" box="[551,733,828,850]" captionStart="FIGURES 258269" captionStartId="91.[151,264,1894,1919]" captionTargetBox="[213,1373,166,1875]" captionTargetId="figure@91.[213,1375,166,1875]" captionTargetPageId="91" captionText="FIGURES 258269. Luciola horni males. 258, 259, 269 type male; 260265, 267, 268 Halupahani male (NHML); 266 India male (Prague). 258, 266, 267 habitus dorsal; 259 habitus ventral; 260, 261, 265 aedeagal sheath: 260 ventral; 261, 265 dorsal (260, 261 under transmitted light from below); 262, 263 aedeagus: 262 ventral; 263 right lateral; 268 tergites 7, 8 dorsal; 269 label. Scale lines are 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3508204" httpUri="https://zenodo.org/record/3508204/files/figure.png" pageId="87" pageNumber="88">Figs 258, 266, 267</figureCitation>
)............................................
<taxonomicName id="23946389516C1801FA99FCA3EE251F28" ID-CoL="72QCH" authority="Bourgeois Elytra" authorityName="Bourgeois Elytra" authorityYear="1905" class="Insecta" family="Lampyridae" genus="Luciola" kingdom="Animalia" order="Coleoptera" pageId="87" pageNumber="88" phylum="Arthropoda" rank="species" species="horni">
<emphasis id="D6E0C418516C1801FA99FCA3EA0C1F16" box="[1280,1332,828,849]" italics="true" pageId="87" pageNumber="88">horni</emphasis>
Bourgeois Elytra
</taxonomicName>
pale brown with white apices;
<quantity id="236CB5EF516C1801FDD3FCC5EDB61F28" box="[586,654,858,879]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="87" pageNumber="88" unit="mm" value="10.0">10 mm</quantity>
long; known from
<collectingCountry id="9C83589A516C1801FCD1FCC6ECA41F28" box="[840,924,857,879]" name="Vietnam" pageId="87" pageNumber="88">Vietnam</collectingCountry>
; pronotum pale yellowish, may have pale brown median marking (
<figureCitation id="7CAF048F516C1801FEF7FCE8EECE1FCA" box="[366,502,887,909]" captionStart="FIGURES 246257" captionStartId="88.[151,264,1892,1917]" captionTargetBox="[214,1374,166,1872]" captionTargetId="figure@88.[214,1374,166,1872]" captionTargetPageId="88" captionText="FIGURES 246257. Luciola chapaensis males. 246, 253 Luciola atripes type MNHN; 247 Luciola chapaensis type male MNHN; 248251, 254257 Malaysia Kelantan males; 246 elytra; 247, 248 habitus dorsal; 249 habitus ventral; 250, 251 abdo- men: 250 ventral; 251 dorsal; 254256 aedeagus: 254 ventral; 255 dorsal; 256 left lateral; 257 aedeagal sheath dorsal (arrow indicates area of prolongation of sheath tergite). Scale lines are 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3508202" httpUri="https://zenodo.org/record/3508202/files/figure.png" pageId="87" pageNumber="88">Figs 246247</figureCitation>
)...........................................................
<taxonomicName id="23946389516C1801FB27FCE8EA111FCB" ID-CoL="3WCPG" authorityName="Ho" authorityYear="2019" box="[1214,1321,887,908]" class="Insecta" family="Lampyridae" genus="Luciola" kingdom="Animalia" order="Coleoptera" pageId="87" pageNumber="88" phylum="Arthropoda" rank="species" species="chapaensis" status="comb. nov.">
<emphasis id="D6E0C418516C1801FB27FCE8EA111FCB" box="[1214,1321,887,908]" italics="true" pageId="87" pageNumber="88">chapaensis</emphasis>
</taxonomicName>
<emphasis id="D6E0C418516C1801FAB7FCE8EAA41FCB" bold="true" box="[1326,1436,887,908]" pageId="87" pageNumber="88">
<taxonomicNameLabel id="CDD37963516C1801FAB7FCE8EAA41FCB" box="[1326,1436,887,908]" pageId="87" pageNumber="88" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="E42B180A516C1801FF0EFC0BEAA41F80" blockId="87.[151,1437,153,1026]" pageId="87" pageNumber="88">
14. Female fully winged and flight capable (Chen: 172)............................................
<taxonomicName id="23946389516C1801FB7FFC0BEAA41FED" ID-CoL="3WCQ9" authority="Olivier" authorityName="Olivier" box="[1254,1436,916,938]" class="Insecta" family="Lampyridae" genus="Luciola" kingdom="Animalia" order="Coleoptera" pageId="87" pageNumber="88" phylum="Arthropoda" rank="species" species="curtithorax">
<emphasis id="D6E0C418516C1801FB7FFC0BEA6A1FEE" box="[1254,1362,916,937]" italics="true" pageId="87" pageNumber="88">curtithorax</emphasis>
Olivier
</taxonomicName>
Female flightless..................................................................................... 15
</paragraph>
<paragraph id="E42B180A516C1801FF0EFC50EAA41845" blockId="87.[151,1437,153,1026]" pageId="87" pageNumber="88">
15. Female with elytra usually covering abdomen and hind wings absent............................
<taxonomicName id="23946389516C1801FB54FC50EAA51FA2" ID-CoL="6QQBY" authority="Kiesenwetter" authorityName="Kiesenwetter" authorityYear="1874" box="[1229,1437,975,997]" class="Insecta" family="Lampyridae" genus="Luciola" kingdom="Animalia" order="Coleoptera" pageId="87" pageNumber="88" phylum="Arthropoda" rank="species" species="parvula">
<emphasis id="D6E0C418516C1801FB54FC50EA201FA3" box="[1229,1304,975,996]" italics="true" pageId="87" pageNumber="88">parvula</emphasis>
Kiesenwetter
</taxonomicName>
Female with short elytra up to twice as long as median pronotal length (Chen: 170, 171)................
<taxonomicName id="23946389516C1801FB63FC73EAA41845" ID-CoL="3WCQY" authority="Olivier" authorityName="Olivier" authorityYear="1913" box="[1274,1436,1004,1026]" class="Insecta" family="Lampyridae" genus="Luciola" kingdom="Animalia" order="Coleoptera" pageId="87" pageNumber="88" phylum="Arthropoda" rank="species" species="filiformis">
<emphasis id="D6E0C418516C1801FB63FC73EA6A1846" box="[1274,1362,1004,1025]" italics="true" pageId="87" pageNumber="88">filiformis</emphasis>
Olivier
</taxonomicName>
</paragraph>
</subSubSection>
</treatment>
</document>