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<mods:title id="406CC34F02A1BAB82507D05AB7BD9679">The Luciolinae of S. E. Asia and the Australopacific region: a revisionary checklist (Coleoptera: Lampyridae) including description of three new genera and 13 new species</mods:title>
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<emphasis id="D6E0C4185153183EFF0EFDF2EE231ECF" bold="true" box="[151,283,621,648]" italics="true" pageId="104" pageNumber="105">Magnalata</emphasis>
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<figureCitation id="7CAF048F5153183EFF0EFD0EEFDF1EEC" box="[151,231,657,683]" captionStart="FIGURES 6469" captionStartId="20.[151,264,1850,1875]" captionTargetBox="[157,1431,186,1824]" captionTargetId="figure@20.[151,1435,166,1831]" captionTargetPageId="20" captionText="FIGURES 6469. Key to genera characters 10. 64 Aquilonia costata; 65, 66 Aq. similismessoria comb. nov.; 67 Magnalata limbata; 68 Atyphella flammans; 69 Aty. aphrogeneia." figureDoi="http://doi.org/10.5281/zenodo.3508127" httpUri="https://zenodo.org/record/3508127/files/figure.png" pageId="104" pageNumber="105">Fig. 67</figureCitation>
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<emphasis id="D6E0C4185153183EFF0EFD4AEE3E1EAA" box="[151,262,725,749]" italics="true" pageId="104" pageNumber="105">Magnalata</emphasis>
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Ballantyne
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in
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figs 17, 18, 225245.
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Jusoh
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2018: 10
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species
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:
<taxonomicName id="239463895153183EFEB3FD6BED4B1F4B" ID-CoL="3WCT6" authority="Blanchard 1853" authorityName="Blanchard" authorityYear="1853" box="[298,627,755,780]" class="Insecta" family="Lampyridae" genus="Luciola" kingdom="Animalia" order="Coleoptera" pageId="104" pageNumber="105" phylum="Arthropoda" rank="species" species="limbata">
<emphasis id="D6E0C4185153183EFEB3FD6BEEF31F4B" box="[298,459,756,780]" italics="true" pageId="104" pageNumber="105">Luciola limbata</emphasis>
<bibRefCitation id="800565FB5153183EFE4BFD6CED4B1F4B" author="Blanchard" box="[466,627,755,780]" firstAuthor="Blanchard" pageId="104" pageNumber="105" refId="ref115321" refString="Blanchard, E. (1853) Description des insectes In: Voyage au Pole sud et dans l'Oceanie sur les corvettes l'Astrolabe et la Zelee execute par ordre du roi pendant des annees 1837 - 1838 - 1839 - 1840 sous le commandement de M. J. Dumont-Durville. Zoologie 4. Gide et Baudry, Paris, 422 pp." type="book" year="1853">Blanchard 1853</bibRefCitation>
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designated by
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.
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<emphasis id="D6E0C4185153183EFF0EFCA6EE321F13" bold="true" box="[151,266,825,852]" pageId="104" pageNumber="105">Diagnosis</emphasis>
. A large wide distinctively coloured genus known from the
<collectingCountry id="9C83589A5153183EFC20FCA5EB421F13" box="[953,1146,826,852]" name="Solomon Islands" pageId="104" pageNumber="105">Solomon Islands</collectingCountry>
; up to
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long and
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wide; with orange pronotum and black elytra which are orange margined laterally only. One of the
<taxonomicName id="239463895153183EFAA8FCC1EAA41F3F" authorityName="Olliff" authorityYear="1890" box="[1329,1436,862,888]" class="Insecta" family="Lampyridae" genus="Atyphella" kingdom="Animalia" order="Coleoptera" pageId="104" pageNumber="105" phylum="Arthropoda" rank="genus">
<emphasis id="D6E0C4185153183EFAA8FCC1EAA41F3F" box="[1329,1436,862,888]" italics="true" pageId="104" pageNumber="105">Atyphella</emphasis>
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complex, having LL of the aedeagus visible beside the ML when viewed from beneath; distinguished by its size and colouration and occurrence in the Solomons. As addressed here
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<emphasis id="D6E0C4185153183EFC0DFC39EB371F87" box="[916,1039,934,960]" italics="true" pageId="104" pageNumber="105">Magnalata</emphasis>
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is monotypic.
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<emphasis id="D6E0C4185153183EFF0EFC72EE9F184F" bold="true" box="[151,423,1005,1032]" pageId="104" pageNumber="105">Redescription of genus</emphasis>
.
</paragraph>
<paragraph id="E42B180A5153183EFF5EFB8DEEB21943" blockId="104.[151,1437,1005,2041]" pageId="104" pageNumber="105">
<emphasis id="D6E0C4185153183EFF5EFB8DEE3C186B" bold="true" box="[199,260,1042,1068]" pageId="104" pageNumber="105">Male</emphasis>
. Pronotum: wider than long; dorsal surface without irregularities in posterolateral areas (1) and longitudinal groove in lateral areas (2); punctation moderately dense (3); anterior margin not explanate (6); lateral margins diverging posteriorly along most of their length (C&gt;A, B); width&gt; humeral width; anterolateral corners rounded obtuse; lateral margins without indentation at mid-point, and sinuosity in either horizontal or vertical plane (15, 16); without indentation in lateral margin near posterolateral corner, and irregularities at corner (17, 18); posterolateral corners rounded obtuse posterolateral corners projecting beyond median posterior margin; separated from it by scarce emarginations.
</paragraph>
<paragraph id="E42B180A5153183EFF5EFA90EC46190B" blockId="104.[151,1437,1005,2041]" pageId="104" pageNumber="105">Hypomera: closed; median area of hypomeron not elevated in vertical direction; posterior area widely flat but not strongly adpressed; pronotal width/ GHW 1.6 in other species.</paragraph>
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Elytron: lateral margins parallel-sided; apices not deflexed; punctation dense (34, 35), not linear (36), not as large as that of pronotum, nor widely and evenly spaced; epipleuron and sutural ridge extend beyond mid-point almost to apex but not extending around apex, neither thickened in apical half; no ridge around apex as in some
<emphasis id="D6E0C4185153183EFF0EFA5DEE34199B" box="[151,268,1474,1500]" italics="true" pageId="104" pageNumber="105">
<taxonomicName id="239463895153183EFF0EFA5DEE3D199B" authorityName="Olliff" authorityYear="1890" box="[151,261,1474,1500]" class="Insecta" family="Lampyridae" genus="Atyphella" kingdom="Animalia" order="Coleoptera" pageId="104" pageNumber="105" phylum="Arthropoda" rank="genus">Atyphella</taxonomicName>
;
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fourinterstitial lines well defined; elytral carina absent; in horizontal specimen viewed from below epipleuron at elytral base wide, covering humerus or almost so; viewed from above in horizontal specimen the anterior margin of epipleuron arises level with or anterior to posterior margin of MS; epipleuron developed as a lateral ridge along most of length; sutural margins approximate along most of length in closed elytra.
</paragraph>
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Head: moderately depressed between eyes; moderately well exposed in front of pronotum, and capable of partial retraction within prothoracic cavity; eyes with moderate separation beneath at level of posterior margin of mouthpart complex; eyes above labrum close; frons-vertex junction rounded, without median elevation; posterolateral eye excavation not strongly developed, not visible in resting head position; antennal sockets on head between eyes, not contiguous, always separated by &lt;ASW; clypeolabral suture present, flexible, not in front of anterior eye margin when head viewed with labrum horizontal; outer edges of labrum reach inner edges of closed mandibles or beyond. Mouthparts: functional; apical labial palpomere flattened, shaped like a broad triangle with inner edge irregular, not dentate, and at least half as long as apical maxillary palpomere (
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listed variations in shape of inner margin). Antennae 11 segmented; length&gt;GHW up to slightly less than twice GHW; FS1 subequal in length to, or longer than, pedicel; FS 79 not conspicuously shorter than rest; no FS expanded or laterally produced; all FS including FS 1 longer than wide.
</paragraph>
<paragraph id="E42B180A5153183EFF5EF840EA261BBF" blockId="104.[151,1437,1005,2041]" box="[199,1310,2014,2041]" pageId="104" pageNumber="105">Legs: with inner tarsal claw entire; without MFC; no leg segments swollen, curved or emarginated.</paragraph>
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Abdomen: without cuticular remnants in association with aedeagal sheath (131, 132); no ventrites with curved posterior margins, nor extending anteriorly into emarginated posterior margin of anterior segment (106108).
<date id="902A3ECA5152183FFAE3FF22EAA41C90" box="[1402,1436,189,215]" pageId="105" pageNumber="106">V6</date>
with LO present, occupying almost ventrite surface.
<date id="902A3ECA5152183FFD7EFF7EEC311CBC" box="[743,777,225,251]" pageId="105" pageNumber="106">V7</date>
with LO entire, occupying most of the surface of ventrite, and reaching to sides, but not into MPP; MPP moderately broad apically rounded; no PLP. T8 well sclerotised, symmetrical, visible posterior area either widest across median area (W&gt;L) or W=L; median posterior margin not emarginate; T8 not engulfing posterior margin of
<date id="902A3ECA5152183FFD25FED2EDE61D20" box="[700,734,333,359]" pageId="105" pageNumber="106">V7</date>
nor MPP (133, 134), without well-defined median longitudinal trough; concealed anterolateral arms of T8 very short, not as long as entire posterior portion; not laterally emarginated before their origins, dorsoventral expansions absent, expanded only in horizontal plane; without bifurcation of inner margin and ventrally directed pieces (217); lateral margins of T8 not enfolding sides of
<date id="902A3ECA5152183FFB35FE26EBF61D94" box="[1196,1230,441,467]" pageId="105" pageNumber="106">V7</date>
(190); T7 without prolonged anterolateral corners.
</paragraph>
<paragraph id="E42B180A5152183FFF5EFD9EEE881EC0" blockId="105.[151,1437,153,1079]" pageId="105" pageNumber="106">Aedeagal sheath: asymmetrical; sheath sternite evenly emarginated on its right side from the point of attachment of the tergite; anterior half of sternite broad, apically rounded; tergite without lateral arms extending anteriorly at sides of sheath sternite; tergite without projecting pieces along posterior margin of T9, anterior margin without separate transverse band.</paragraph>
<paragraph id="E42B180A5152183FFF5EFD0EED971F7C" blockId="105.[151,1437,153,1079]" pageId="105" pageNumber="106">Aedeagus: L/W 3/1; symmetrical with apices of LL visible from beneath at sides of ML, LL/ML moderate; LL of equal length, slightly shorter than ML, separated along inner dorsal margins but not divergent in basal half; with apices narrower than ML; preapical inner margin of LL obliquely truncate, with obvious tooth; dorsal base of LL produced anteriorly, apically acute and medially entire; ML symmetrical; BP not strongly sclerotised, not hooded, not strongly emarginated along anterior margin.</paragraph>
<paragraph id="E42B180A5152183FFF5EFCDAEDBE1FC4" blockId="105.[151,1437,153,1079]" pageId="105" pageNumber="106">
<emphasis id="D6E0C4185152183FFF5EFCDAEE261F18" bold="true" box="[199,286,837,863]" pageId="105" pageNumber="106">Female</emphasis>
. Macropterous and probably capable of flight. Pinned females were not further investigated.Associated by label data, size and dorsal colour pattern.
</paragraph>
<paragraph id="E42B180A5152183FFF5EFC12ED2C1F8C" blockId="105.[151,1437,153,1079]" pageId="105" pageNumber="106">
<emphasis id="D6E0C4185152183FFF5EFC12EE281FE0" bold="true" box="[199,272,909,935]" pageId="105" pageNumber="106">Larva</emphasis>
. Not reliably associated.
<bibRefCitation id="800565FB5152183FFDB2FC12EC9F1FEF" author="Ballantyne &amp; Lambkin" box="[555,935,909,936]" firstAuthor="Ballantyne" pageId="105" pageNumber="106" pagination="1 - 188" refId="ref114542" refString="Ballantyne, L. A. &amp; Lambkin, C. L. (2009) Systematics of Indo-Pacific fireflies with a redefinition of Australasian Atyphella Olliff, Madagascan Photuroluciola Pic, and description of seven new genera from the Luciolinae (Coleoptera: Lampyridae). Zootaxa, 1997, 1 - 188." type="journal article" year="2009">Ballantyne &amp; Lambkin (2009: 67)</bibRefCitation>
described a distinctively coloured larva with laterally explanate tergal margins.
</paragraph>
</subSubSection>
<subSubSection id="AC8E4B815152183FFF5EFC4AEFA619D8" pageId="105" pageNumber="106" type="discussion">
<paragraph id="E42B180A5152183FFF5EFC4AEB261870" blockId="105.[151,1437,153,1079]" pageId="105" pageNumber="106">
<emphasis id="D6E0C4185152183FFF5EFC4AEE0C1FA8" bold="true" box="[199,308,981,1007]" pageId="105" pageNumber="106">Remarks</emphasis>
.
<bibRefCitation id="800565FB5152183FFEDCFC4AEDE81FA8" author="Ballantyne &amp; Lambkin" box="[325,720,980,1007]" firstAuthor="Ballantyne" pageId="105" pageNumber="106" pagination="1 - 188" refId="ref114542" refString="Ballantyne, L. A. &amp; Lambkin, C. L. (2009) Systematics of Indo-Pacific fireflies with a redefinition of Australasian Atyphella Olliff, Madagascan Photuroluciola Pic, and description of seven new genera from the Luciolinae (Coleoptera: Lampyridae). Zootaxa, 1997, 1 - 188." type="journal article" year="2009">Ballantyne &amp; Lambkin (2009: 65)</bibRefCitation>
characterised this genus from three species of which two,
<taxonomicName id="239463895152183FFAE7FC49EE601854" authority="Olivier" authorityName="Olivier" class="Insecta" family="Lampyridae" genus="Magnalata" kingdom="Animalia" order="Coleoptera" pageId="105" pageNumber="106" phylum="Arthropoda" rank="species" species="carolinae">
<emphasis id="D6E0C4185152183FFAE7FC49EE391854" italics="true" pageId="105" pageNumber="106">M. carolinae</emphasis>
Olivier
</taxonomicName>
and
<taxonomicName id="239463895152183FFE08FC65EDA41854" authority="Ballantyne" authorityName="Ballantyne 2009 in Ballantyne &amp; Lambkin" authorityYear="2009" box="[401,668,1017,1043]" class="Insecta" family="Lampyridae" genus="Magnalata" kingdom="Animalia" order="Coleoptera" pageId="105" pageNumber="106" phylum="Arthropoda" rank="species" species="rennellia">
<emphasis id="D6E0C4185152183FFE08FC65ED231854" box="[401,539,1017,1043]" italics="true" pageId="105" pageNumber="106">M. rennellia</emphasis>
Ballantyne
</taxonomicName>
are removed to
<taxonomicName id="239463895152183FFCCFFC66ECF91854" authorityName="Olliff" authorityYear="1890" box="[854,961,1017,1043]" class="Insecta" family="Lampyridae" genus="Atyphella" kingdom="Animalia" order="Coleoptera" pageId="105" pageNumber="106" phylum="Arthropoda" rank="genus">
<emphasis id="D6E0C4185152183FFCCFFC66ECF91854" box="[854,961,1017,1043]" italics="true" pageId="105" pageNumber="106">Atyphella</emphasis>
</taxonomicName>
here.
<taxonomicName id="239463895152183FFB90FC66EBDA1854" authorityName="Ho" authorityYear="2019" baseAuthorityName="Blanchard. McDermott" baseAuthorityYear="2019" box="[1033,1250,1017,1043]" class="Insecta" family="Lampyridae" genus="Magnalata" kingdom="Animalia" order="Coleoptera" pageId="105" pageNumber="106" phylum="Arthropoda" rank="species" species="limbata">
<emphasis id="D6E0C4185152183FFB90FC66EBDA1854" box="[1033,1250,1017,1043]" italics="true" pageId="105" pageNumber="106">Magnalata limbata</emphasis>
</taxonomicName>
was extensively described from many specimens ranging over the length of the
<collectingCountry id="9C83589A5152183FFCC2FB82EB221870" box="[859,1050,1053,1079]" name="Solomon Islands" pageId="105" pageNumber="106">Solomon Islands</collectingCountry>
.
</paragraph>
<caption id="B0EB48825152183FFF0EFBF9EC8618E4" ID-Table-UUID="B0EB48825152183FFF0EFBF9EC8618E4" httpUri="http://table.plazi.org/id/B0EB48825152183FFF0EFBF9EC8618E4" pageId="105" pageNumber="106" targetBox="[166,1345,1198,1400]" targetIsTable="true" targetPageId="105">
<paragraph id="E42B180A5152183FFF0EFBF9EC8618E4" blockId="105.[151,958,1126,1187]" pageId="105" pageNumber="106">
<emphasis id="D6E0C4185152183FFF0EFBF9EC251839" bold="true" box="[151,797,1126,1151]" pageId="105" pageNumber="106">
TABLE 19. List of species of
<taxonomicName id="239463895152183FFE79FBF9EC201839" authority="Ballantyne 2009" authorityName="Ballantyne 2009 in Ballantyne &amp; Lambkin" authorityYear="2009" box="[480,792,1126,1151]" class="Insecta" family="Lampyridae" genus="Magnalata" kingdom="Animalia" order="Coleoptera" pageId="105" pageNumber="106" phylum="Arthropoda" rank="genus">
<emphasis id="D6E0C4185152183FFE79FBF9ED601839" bold="true" box="[480,600,1126,1150]" italics="true" pageId="105" pageNumber="106">Magnalata</emphasis>
<bibRefCitation id="800565FB5152183FFDC6FBF8EC201839" author="Ballantyne" box="[607,792,1126,1151]" firstAuthor="Ballantyne" pageId="105" pageNumber="106" pagination="1 - 188" refId="ref114542" refString="Ballantyne, L. A. &amp; Lambkin, C. L. (2009) Systematics of Indo-Pacific fireflies with a redefinition of Australasian Atyphella Olliff, Madagascan Photuroluciola Pic, and description of seven new genera from the Luciolinae (Coleoptera: Lampyridae). Zootaxa, 1997, 1 - 188." type="journal article" year="2009">Ballantyne 2009</bibRefCitation>
</taxonomicName>
.
</emphasis>
(*Reference: 1 =
<bibRefCitation id="800565FB5152183FFEC8FB14EDBF18E4" author="Ballantyne &amp; Lambkin" box="[337,647,1163,1187]" firstAuthor="Ballantyne" pageId="105" pageNumber="106" pagination="1 - 188" refId="ref114542" refString="Ballantyne, L. A. &amp; Lambkin, C. L. (2009) Systematics of Indo-Pacific fireflies with a redefinition of Australasian Atyphella Olliff, Madagascan Photuroluciola Pic, and description of seven new genera from the Luciolinae (Coleoptera: Lampyridae). Zootaxa, 1997, 1 - 188." type="journal article" year="2009">Ballantyne &amp; Lambkin 2009</bibRefCitation>
. Red block = type not found)
</paragraph>
</caption>
<paragraph id="E42B180A5152183FFF3FFB31EBD3193F" pageId="105" pageNumber="106">
<table id="9694EAAA5152E7A9FF3FFB31EA79193F" box="[166,1345,1198,1400]" gridcols="5" gridrows="2" pageId="105" pageNumber="106">
<tr id="5AA41A485152E7A9FF3FFB31EA791917" box="[166,1345,1198,1360]" gridrow="0" pageId="105" pageNumber="106">
<th id="197573345152E7A9FF3FFB31EE8D1917" box="[166,437,1198,1360]" gridcol="0" gridrow="0" pageId="105" pageNumber="106">Species</th>
<td id="197573345152E7A9FD6CFB31ECA01917" box="[757,920,1198,1360]" gridcol="1" gridrow="0" pageId="105" pageNumber="106">
Solomon Islands Location of
<typeStatus id="3B2FA6A85152183FFCE6FABCECA01908" box="[895,920,1315,1359]" pageId="105" pageNumber="106">type</typeStatus>
</td>
<td id="197573345152E7A9FC49FB31ECD21917" box="[976,1002,1198,1360]" gridcol="2" gridrow="0" pageId="105" pageNumber="106">Females</td>
<td id="197573345152E7A9FBC1FB31EB491917" box="[1112,1137,1198,1360]" gridcol="3" gridrow="0" pageId="105" pageNumber="106">Larvae</td>
<td id="197573345152E7A9FB46FB31EA791917" box="[1247,1345,1198,1360]" gridcol="4" gridrow="0" pageId="105" pageNumber="106">
<collectingCountry id="9C83589A5152183FFB79FB40EA791944" box="[1248,1345,1211,1360]" name="Solomon Islands" pageId="105" pageNumber="106">References for illustrations of species *</collectingCountry>
</td>
</tr>
<tr id="5AA41A485152E7A9FF3FFAC1EA79193F" box="[166,1345,1374,1400]" gridrow="1" pageId="105" pageNumber="106">
<th id="197573345152E7A9FF3FFAC1EE8D193F" box="[166,437,1374,1400]" gridcol="0" gridrow="1" pageId="105" pageNumber="106">
<emphasis id="D6E0C4185152183FFF3FFAFFEFCB193F" box="[166,243,1376,1400]" italics="true" pageId="105" pageNumber="106">limbata</emphasis>
<emphasis id="D6E0C4185152183FFF60FAC0EE8D193F" bold="true" box="[249,437,1375,1400]" pageId="105" pageNumber="106">
(
<bibRefCitation id="800565FB5152183FFE99FAC0EE96193F" author="Blanchard" box="[256,430,1375,1400]" firstAuthor="Blanchard" pageId="105" pageNumber="106" refId="ref115321" refString="Blanchard, E. (1853) Description des insectes In: Voyage au Pole sud et dans l'Oceanie sur les corvettes l'Astrolabe et la Zelee execute par ordre du roi pendant des annees 1837 - 1838 - 1839 - 1840 sous le commandement de M. J. Dumont-Durville. Zoologie 4. Gide et Baudry, Paris, 422 pp." type="book" year="1853">Blanchard 1853</bibRefCitation>
)
</emphasis>
</th>
<td id="197573345152E7A9FD6CFAC1ECA0193F" box="[757,920,1374,1400]" gridcol="1" gridrow="1" pageId="105" pageNumber="106">*</td>
<td id="197573345152E7A9FC49FAC1ECD2193F" box="[976,1002,1374,1400]" gridcol="2" gridrow="1" pageId="105" pageNumber="106"></td>
<td id="197573345152E7A9FBC1FAC1EB49193F" box="[1112,1137,1374,1400]" gridcol="3" gridrow="1" pageId="105" pageNumber="106"></td>
<td id="197573345152E7A9FB46FAC1EA79193F" box="[1247,1345,1374,1400]" gridcol="4" gridrow="1" pageId="105" pageNumber="106">1</td>
</tr>
</table>
</paragraph>
<paragraph id="E42B180A5152183FFF0EFA1AEFA619D8" blockId="105.[151,158,1413,1439]" box="[151,158,1413,1439]" pageId="105" pageNumber="106">.</paragraph>
</subSubSection>
</treatment>
</document>