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<document id="471D2C02501E4F1C0C7D9E1FB5F5430F" ID-CLB-Dataset="256669" ID-DOI="10.5852/ejt.2023.864.2093" ID-GBIF-Dataset="7f562ba5-17db-46bf-93e8-4d1ba49c2800" ID-ISSN="2118-9773" ID-Zenodo-Dep="7872174" ID-ZooBank="D47B6B04-B2AD-4FDD-B7C4-B71CA6A5BB84" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="diego" IM.treatments_approvedBy="diego" checkinTime="1682513062582" checkinUser="felipe" docAuthor="Brunke, Adam James" docDate="2023" docId="6C38C875FFA05B4EFE2AFCE3E02EFD00" docLanguage="en" docName="ejt-864_117-145.pdf" docOrigin="European Journal of Taxonomy 864" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Quedius kambaitensis Scheerpeltz 1965, syn. nov." docType="treatment" docVersion="5" lastPageNumber="141" masterDocId="9001B00DFFB75B56FFDAFFF6E270FF89" masterDocTitle="Review of Quedius (Coleoptera, Staphylinidae) described from the 1934 expedition by R. Malaise to Myanmar" masterLastPageNumber="145" masterPageNumber="117" pageNumber="140" updateTime="1699514916083" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="9E378A74B28996DBA17EEA9B0D666127">Review of Quedius (Coleoptera, Staphylinidae) described from the 1934 expedition by R. Malaise to Myanmar</mods:title>
</mods:titleInfo>
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<mods:namePart id="FFEB9B077876564918E2BF8B64E98781">Brunke, Adam James</mods:namePart>
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<mods:title id="EA33F225D3AFF927D141D3FBFD39D186">European Journal of Taxonomy</mods:title>
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<mods:date id="9703DC15873838639C2F9064010C68DF">2023</mods:date>
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<mods:number id="6F422E563EBAF5AC0A0430F23B8ABAD0">2023-04-20</mods:number>
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<mods:number id="1DF4227440CF218E106292F919F8CD88">864</mods:number>
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<mods:classification id="C24319DBF98EC9E90CE59941FDA224E8">journal article</mods:classification>
<mods:identifier id="E5F9FC5C3C8E3684ACD7FED457D3BD53" type="CLB-Dataset">256669</mods:identifier>
<mods:identifier id="025624724B898A524441FBCCE80D30B6" type="DOI">10.5852/ejt.2023.864.2093</mods:identifier>
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<mods:identifier id="C56DE5243A4BE0BC905976807047152B" type="ISSN">2118-9773</mods:identifier>
<mods:identifier id="118A7CD898A18F8212E3ED3175671D07" type="Zenodo-Dep">7872174</mods:identifier>
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</mods:mods>
<treatment id="6C38C875FFA05B4EFE2AFCE3E02EFD00" ID-DOI="http://doi.org/10.5281/zenodo.7872319" ID-GBIF-Taxon="209579373" ID-Zenodo-Dep="7872319" LSID="urn:lsid:plazi:treatment:6C38C875FFA05B4EFE2AFCE3E02EFD00" httpUri="http://treatment.plazi.org/id/6C38C875FFA05B4EFE2AFCE3E02EFD00" lastPageId="24" lastPageNumber="141" pageId="23" pageNumber="140">
<subSubSection id="AC8B2AE8FFA05B41FE2AFCE3E634FCB9" box="[496,1092,789,816]" pageId="23" pageNumber="140" type="nomenclature">
<paragraph id="E42E7963FFA05B41FE2AFCE3E634FCB9" blockId="23.[496,1092,789,816]" box="[496,1092,789,816]" pageId="23" pageNumber="140">
<heading id="BF66CE0FFFA05B41FE2AFCE3E634FCB9" box="[496,1092,789,816]" centered="true" fontSize="11" level="2" pageId="23" pageNumber="140" reason="2">
<taxonomicName id="239102E0FFA05B41FE2AFCE3E1A4FCB9" authority="Scheerpeltz, 1965" authorityName="Scheerpeltz" authorityYear="1965" box="[496,980,789,816]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="140" phylum="Arthropoda" rank="species" species="kambaitensis">
<emphasis id="D6E5A571FFA05B41FE2AFCE3E08AFCB9" bold="true" box="[496,762,789,816]" italics="true" pageId="23" pageNumber="140">Quedius kambaitensis</emphasis>
<bibRefCitation id="80000492FFA05B41FCDBFCE3E1A4FCB9" author="Scheerpeltz O." box="[769,980,789,816]" pageId="23" pageNumber="140" pagination="93 - 371" refId="ref13560" refString="Scheerpeltz O. 1965. Wissenschaftliche Ergebnisse der Schwedischen Expedition 1934 nach Indien und Burma. Arkiv for Zoologi 17: 93 - 371." type="journal article" year="1965">Scheerpeltz, 1965</bibRefCitation>
</taxonomicName>
<taxonomicNameLabel id="CDD6180AFFA05B41FC01FCE0E634FCB9" box="[987,1092,790,816]" pageId="23" pageNumber="140">syn. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="AC8B2AE8FFA05B41FF67FCA1E03AF983" pageId="23" pageNumber="140" type="materials_examined">
<paragraph id="E42E7963FFA05B41FF67FCA1E31CFCF8" blockId="23.[189,364,855,881]" box="[189,364,855,881]" pageId="23" pageNumber="140">
<heading id="BF66CE0FFFA05B41FF67FCA1E31CFCF8" bold="true" box="[189,364,855,881]" fontSize="11" level="3" pageId="23" pageNumber="140" reason="3">
<emphasis id="D6E5A571FFA05B41FF67FCA1E31CFCF8" bold="true" box="[189,364,855,881]" pageId="23" pageNumber="140">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="E42E7963FFA05B41FF30FC73E32CFC16" blockId="23.[189,1398,901,1069]" box="[234,348,901,927]" pageId="23" pageNumber="140">
<emphasis id="D6E5A571FFA05B41FF30FC73E32CFC16" bold="true" box="[234,348,901,927]" pageId="23" pageNumber="140">
<typeStatus id="3B2AC7C1FFA05B41FF30FC73E32CFC16" box="[234,348,901,927]" pageId="23" pageNumber="140" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="E42E7963FFA05B41FF67FC5FE121FBA5" blockId="23.[189,1398,901,1069]" pageId="23" pageNumber="140">
<materialsCitation id="54F9733EFFA05B41FF67FC5FE121FBA5" ID-GBIF-Occurrence="4097213303" collectingDate="1934-06-04" collectionCode="NHRS" collectorName="Malaise" country="Myanmar" determinerName="Scheerpeltz" elevation="2000" location="N. E. Burma" municipality="Kambaiti" pageId="23" pageNumber="140" specimenCode="NHRS-JLKB 000073668" specimenCount="1" specimenCount-female="1" stateProvince="Kachin State" typeStatus="holotype">
<collectingCountry id="9C8639F3FFA05B41FF67FC5FE326FC4A" box="[189,342,937,963]" name="Myanmar" pageId="23" pageNumber="140">MYANMAR</collectingCountry>
–
<emphasis id="D6E5A571FFA05B41FEAEFC5EE067FC4B" bold="true" box="[372,535,936,962]" pageId="23" pageNumber="140">
<collectingRegion id="2655B781FFA05B41FEAEFC5EE067FC4B" box="[372,535,936,962]" country="Myanmar" name="Kachin" pageId="23" pageNumber="140">Kachin State</collectingRegion>
</emphasis>
•
<specimenCount id="F297B2EAFFA05B41FDEAFC5FE037FC4A" box="[560,583,937,963]" pageId="23" pageNumber="140" type="female">♀</specimenCount>
; “
<location id="E14E2FB8FFA05B41FDB8FC5FE083FC4A" LSID="urn:lsid:plazi:treatment:6C38C875FFA05B4EFE2AFCE3E02EFD00:E14E2FB8FFA05B41FDB8FC5FE083FC4A" box="[610,755,937,963]" country="Myanmar" municipality="Kambaiti" name="N. E. Burma" pageId="23" pageNumber="140" stateProvince="Kachin State">N.E. Burma</location>
,
<collectingMunicipality id="044AE319FFA05B41FD24FC5EE11FFC4B" box="[766,879,936,962]" pageId="23" pageNumber="140">Kambaiti</collectingMunicipality>
;
<quantity id="2369D486FFA05B41FCA1FC5FE1A7FC4A" box="[891,983,937,963]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="23" pageNumber="140" unit="m" value="2000.0">
<elevation id="6FBC9E50FFA05B41FCA1FC5FE1A7FC4A" box="[891,983,937,963]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="23" pageNumber="140" unit="m" value="2000.0">2000 m</elevation>
</quantity>
; 4/6.1934;
<collectorName id="49641CB5FFA05B41FB81FC5EE6C8FC4B" box="[1115,1208,936,962]" pageId="23" pageNumber="140">Malaise</collectorName>
[printed label] /
<typeStatus id="3B2AC7C1FFA05B41FF67FC3AE31BFC6F" box="[189,363,972,998]" pageId="23" pageNumber="140" type="holotype">HOLOTYPUS</typeStatus>
[red handwritten label] / TYPUS
<taxonomicName id="239102E0FFA05B41FD24FC3AE6B5FC6F" authority=", O. Scheerpeltz" authorityName=", O. Scheerpeltz" authorityYear="1965" box="[766,1221,972,998]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="140" phylum="Arthropoda" rank="species" species="kambaitiensis">Quedius kambaitiensis, O. Scheerpeltz</taxonomicName>
[dark red-pink label] /
<originalDetermination id="C78FC19FFFA05B41FECDFC19E155FB83" box="[279,805,1007,1034]" pageId="23" pageNumber="140">
<taxonomicName id="239102E0FFA05B41FECDFC19E0D6FB80" authorityName=", O. Scheerpeltz" authorityYear="1965" box="[279,678,1007,1034]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="140" phylum="Arthropoda" rank="species" species="kambaitiensis" status="sp. nov." subGenus="Raphirus">Quedius (Raphirus) kambaitiensis</taxonomicName>
<taxonomicNameLabel id="CDD6180AFFA05B41FD74FC06E155FB83" box="[686,805,1008,1034]" pageId="23" pageNumber="140" rank="species">nov. spec.</taxonomicNameLabel>
</originalDetermination>
, det.
<determinerName id="11F295A7FFA05B41FCBEFC19E19DFB80" box="[868,1005,1007,1033]" pageId="23" pageNumber="140">Scheerpeltz</determinerName>
1944 [printed label] / 6599 E91 / [printed blue label] /
<specimenCode id="B437D118FFA05B41FE6FFBE4E095FBA5" box="[437,741,1042,1068]" pageId="23" pageNumber="140">NHRS-JLKB 000073668</specimenCode>
”;
<collectionCode id="8280E1A6FFA05B41FD21FBE4E13DFBA5" box="[763,845,1042,1068]" pageId="23" pageNumber="140">NHRS</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="E42E7963FFA05B41FF30FBA2E32CFBE7" blockId="23.[189,1399,1108,1275]" box="[234,348,1108,1134]" pageId="23" pageNumber="140">
<emphasis id="D6E5A571FFA05B41FF30FBA2E32CFBE7" bold="true" box="[234,348,1108,1134]" pageId="23" pageNumber="140">
<typeStatus id="3B2AC7C1FFA05B41FF30FBA2E32CFBE7" box="[234,348,1108,1134]" pageId="23" pageNumber="140" type="paratype">Paratype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="E42E7963FFA05B41FF67FB81E679FB72" blockId="23.[189,1399,1108,1275]" pageId="23" pageNumber="140">
<materialsCitation id="54F9733EFFA05B41FF67FB81E675FB72" ID-GBIF-Occurrence="4097213326" collectingDate="1934-06-04" collectionCode="NHW" collectorName="Malaise &amp; Scheerpeltz" country="Myanmar" determinerName="A. Smetana" elevation="2000" location="N. E. Burma" municipality="Kambaiti" pageId="23" pageNumber="140" specimenCount="1" specimenCount-male="1" stateProvince="Kachin State" typeStatus="paratype">
<collectingCountry id="9C8639F3FFA05B41FF67FB81E326FB18" box="[189,342,1143,1169]" name="Myanmar" pageId="23" pageNumber="140">MYANMAR</collectingCountry>
–
<emphasis id="D6E5A571FFA05B41FEABFB81E062FB19" bold="true" box="[369,530,1142,1169]" pageId="23" pageNumber="140">
<collectingRegion id="2655B781FFA05B41FEABFB81E062FB19" box="[369,530,1142,1169]" country="Myanmar" name="Kachin" pageId="23" pageNumber="140">Kachin State</collectingRegion>
</emphasis>
•
<specimenCount id="F297B2EAFFA05B41FDF0FB81E028FB18" box="[554,600,1143,1169]" pageId="23" pageNumber="140" type="male">1 ♂</specimenCount>
; “[male symbol on card] /
<location id="E14E2FB8FFA05B41FC51FB81E66AFB18" LSID="urn:lsid:plazi:treatment:6C38C875FFA05B4EFE2AFCE3E02EFD00:E14E2FB8FFA05B41FC51FB81E66AFB18" box="[907,1050,1143,1169]" country="Myanmar" municipality="Kambaiti" name="N. E. Burma" pageId="23" pageNumber="140" stateProvince="Kachin State">N.E. Burma</location>
,
<collectingMunicipality id="044AE319FFA05B41FBFFFB81E6E6FB18" box="[1061,1174,1143,1169]" pageId="23" pageNumber="140">Kambaiti</collectingMunicipality>
;
<quantity id="2369D486FFA05B41FB7AFB81E68BFB18" box="[1184,1275,1143,1169]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="23" pageNumber="140" unit="m" value="2000.0">
<elevation id="6FBC9E50FFA05B41FB7AFB81E68BFB18" box="[1184,1275,1143,1169]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="23" pageNumber="140" unit="m" value="2000.0">2000 m</elevation>
</quantity>
; 4/6.1934;
<collectorName id="49641CB5FFA05B41FF67FB6CE36AFB3D" box="[189,282,1178,1204]" pageId="23" pageNumber="140">Malaise</collectorName>
[printed label] / Schwedisch Indian-Burma Expedition 1934 [printed label] / ex. coll.
<collectorName id="49641CB5FFA05B41FB34FB6CE707FB3D" box="[1262,1399,1178,1204]" pageId="23" pageNumber="140">Scheerpeltz</collectorName>
[printed blue label] / COTYPUS
<taxonomicName id="239102E0FFA05B41FD8FFB4BE66FFB5E" authority="O. Scheerpeltz" authorityName=", O. Scheerpeltz" authorityYear="1965" box="[597,1055,1213,1239]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="140" phylum="Arthropoda" rank="species" species="kambaitiensis">Quedius kambaitiensis O. Scheerpeltz</taxonomicName>
[pink label] / kambaitiensis [orange card] /
<taxonomicName id="239102E0FFA05B41FEABFB16E0EAFB72" authority="Cam." authorityName="Cameron" authorityYear="1932" box="[369,666,1248,1275]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="140" phylum="Arthropoda" rank="species" species="muscicola">Quedius muscicola Cam.</taxonomicName>
, det.
<determinerName id="11F295A7FFA05B41FD0DFB17E12FFB72" box="[727,863,1249,1275]" pageId="23" pageNumber="140">A. Smetana</determinerName>
2018”;
<collectionCode id="8280E1A6FFA05B41FC64FB17E675FB72" box="[958,1029,1249,1275]" pageId="23" pageNumber="140">NHW</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="E42E7963FFA05B41FF30FAD4E3BFFAB5" blockId="23.[187,1399,1314,1411]" box="[234,463,1314,1340]" pageId="23" pageNumber="140">
<emphasis id="D6E5A571FFA05B41FF30FAD4E3BFFAB5" bold="true" box="[234,463,1314,1340]" pageId="23" pageNumber="140">Non-type material</emphasis>
</paragraph>
<paragraph id="E42E7963FFA05B41FF67FAB3E707FA0B" blockId="23.[187,1399,1314,1411]" pageId="23" pageNumber="140">
<materialsCitation id="54F9733EFFA05B41FF67FAB3E707FA0B" ID-GBIF-Occurrence="4097213317" collectingDate="2005-06-19" collectionCode="ASC" collectorName="A. Smetana" country="China" county="Gongshan County" elevation="2000" latitude="27.794167" location="Autonomous Prefecture" longLatPrecision="12" longitude="98.590164" municipality="Nukiang Lisu" pageId="23" pageNumber="140" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan Province">
<collectingCountry id="9C8639F3FFA05B41FF67FAB3E368FAD6" box="[189,280,1349,1375]" name="China" pageId="23" pageNumber="140">CHINA</collectingCountry>
–
<emphasis id="D6E5A571FFA05B41FEF5FAB3E075FAD6" bold="true" box="[303,517,1349,1375]" pageId="23" pageNumber="140">
<collectingRegion id="2655B781FFA05B41FEF5FAB3E075FAD6" box="[303,517,1349,1375]" country="China" name="Yunnan" pageId="23" pageNumber="140">Yunnan Province</collectingRegion>
</emphasis>
•
<specimenCount id="F297B2EAFFA05B41FDC6FAB3E038FAE9" box="[540,584,1349,1376]" pageId="23" pageNumber="140" type="male">1 ♂</specimenCount>
;
<collectingMunicipality id="044AE319FFA05B41FD89FAB3E084FAD6" box="[595,756,1349,1375]" pageId="23" pageNumber="140">Nukiang Lisu</collectingMunicipality>
,
<location id="E14E2FB8FFA05B41FD26FAB3E665FAD6" LSID="urn:lsid:plazi:treatment:6C38C875FFA05B4EFE2AFCE3E02EFD00:E14E2FB8FFA05B41FD26FAB3E665FAD6" box="[764,1045,1349,1375]" country="China" county="Gongshan County" latitude="27.794167" longLatPrecision="12" longitude="98.590164" municipality="Nukiang Lisu" name="Autonomous Prefecture" pageId="23" pageNumber="140" stateProvince="Yunnan Province">Autonomous Prefecture</location>
,
<collectingCounty id="0D4F01EFFFA05B41FBC4FAB3E682FAD6" box="[1054,1266,1349,1375]" pageId="23" pageNumber="140">Gongshan County</collectingCounty>
,
<location id="E14E2FB8FFA05B41FB21FAB3E289FA0B" LSID="urn:lsid:plazi:treatment:6C38C875FFA05B4EFE2AFCE3E02EFD00:E14E2FB8FFA05B41FB21FAB3E289FA0B" country="China" county="Gongshan County" latitude="27.794167" longLatPrecision="12" longitude="98.590164" municipality="Nukiang Lisu" name="Gaoligong Shan" pageId="23" pageNumber="140" stateProvince="Yunnan Province">Gaoligong Shan</location>
, above ranger station;
<geoCoordinate id="81A51FA4FFA05B41FDDAFA91E0E0FA0A" box="[512,656,1383,1411]" degrees="27" direction="north" minutes="47.65" orientation="latitude" pageId="23" pageNumber="140" precision="9" value="27.794167">27°47.65′ N</geoCoordinate>
,
<geoCoordinate id="81A51FA4FFA05B41FD46FA91E159FA0A" box="[668,809,1383,1411]" degrees="98" direction="east" minutes="35.41" orientation="longitude" pageId="23" pageNumber="140" precision="9" value="98.590164">98°35.41′ E</geoCoordinate>
; alt.
<quantity id="2369D486FFA05B41FCBBFA9EE1CFFA0A" box="[865,959,1384,1411]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="23" pageNumber="140" unit="m" value="2000.0">
<elevation id="6FBC9E50FFA05B41FCBBFA9EE1CFFA0A" box="[865,959,1384,1411]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="23" pageNumber="140" unit="m" value="2000.0">2000 m</elevation>
</quantity>
;
<date id="902F5FA3FFA05B41FC11FA9EE614FA0A" box="[971,1124,1384,1411]" pageId="23" pageNumber="140" value="2005-06-19">
<collectingDate id="806BA64BFFA05B41FC11FA9EE614FA0A" box="[971,1124,1384,1411]" pageId="23" pageNumber="140" value="2005-06-19">19 Jun. 2005</collectingDate>
</date>
;
<collectorName id="49641CB5FFA05B41FBB4FA9EE686FA0B" box="[1134,1270,1384,1410]" pageId="23" pageNumber="140">A. Smetana</collectorName>
leg.;
<collectionCode id="8280E1A6FFA05B41FAECFA9EE703FA0B" box="[1334,1395,1384,1410]" pageId="23" pageNumber="140">ASC</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="E42E7963FFA05B41FF67FA5CE03AF983" blockId="23.[189,1399,1449,1546]" pageId="23" pageNumber="140">
<materialsCitation id="54F9733EFFA05B41FF67FA5CE388FA6E" ID-GBIF-Occurrence="4097213321" collectingDate="1984-04-04" collectionCode="ASC" collectorName="Lobl &amp; Smetana" country="Nepal" elevation="2200" location="Chinchilla" municipality="Kosi" pageId="23" pageNumber="140" specimenCount="1" specimenCount-male="1" stateProvince="East Nepal">
<collectingCountry id="9C8639F3FFA05B41FF67FA5CE368FA4D" box="[189,280,1450,1476]" name="Nepal" pageId="23" pageNumber="140">NEPAL</collectingCountry>
–
<emphasis id="D6E5A571FFA05B41FEE9FA5FE055FA4A" bold="true" box="[307,549,1449,1476]" pageId="23" pageNumber="140">Province Number 1</emphasis>
•
<specimenCount id="F297B2EAFFA05B41FDE7FA5CE01BFA4D" box="[573,619,1450,1476]" pageId="23" pageNumber="140" type="male">1 ♂</specimenCount>
;
<collectingRegion id="2655B781FFA05B41FDADFA5CE089FA4D" box="[631,761,1450,1476]" pageId="23" pageNumber="140">East Nepal</collectingRegion>
,
<collectingMunicipality id="044AE319FFA05B41FCDEFA5CE14EFA4D" box="[772,830,1450,1476]" pageId="23" pageNumber="140">Kosi</collectingMunicipality>
,
<location id="E14E2FB8FFA05B41FC93FA5CE1B6FA4D" LSID="urn:lsid:plazi:treatment:6C38C875FFA05B4EFE2AFCE3E02EFD00:E14E2FB8FFA05B41FC93FA5CE1B6FA4D" box="[841,966,1450,1476]" country="Nepal" municipality="Kosi" name="Chinchilla" pageId="23" pageNumber="140" stateProvince="East Nepal">Chinchilla</location>
, “s/Ahale”; alt.
<quantity id="2369D486FFA05B41FBA5FA5CE6AAFA4D" box="[1151,1242,1450,1476]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="23" pageNumber="140" unit="m" value="2200.0">
<elevation id="6FBC9E50FFA05B41FBA5FA5CE6AAFA4D" box="[1151,1242,1450,1476]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="23" pageNumber="140" unit="m" value="2200.0">2200 m</elevation>
</quantity>
;
<date id="902F5FA3FFA05B41FB3CFA5CE702FA4D" box="[1254,1394,1450,1476]" pageId="23" pageNumber="140" value="1984-04-04">
<collectingDate id="806BA64BFFA05B41FB3CFA5CE702FA4D" box="[1254,1394,1450,1476]" pageId="23" pageNumber="140" value="1984-04-04">4 Apr. 1984</collectingDate>
</date>
;
<collectorName id="49641CB5FFA05B41FF67FA3BE284FA6E" box="[189,244,1485,1511]" pageId="23" pageNumber="140">Löbl</collectorName>
&amp;
<collectorName id="49641CB5FFA05B41FEC0FA3BE30FFA6E" box="[282,383,1485,1511]" pageId="23" pageNumber="140">Smetana</collectorName>
leg.;
<collectionCode id="8280E1A6FFA05B41FE65FA3BE388FA6E" box="[447,504,1485,1511]" pageId="23" pageNumber="140">ASC</collectionCode>
</materialsCitation>
<materialsCitation id="54F9733EFFA05B41FE24FA3BE03AF983" ID-GBIF-Occurrence="4097213322" collectingDate="1984-04-05" collectionCode="ASC" collectorName="Smetana &amp; Lobl" elevation="2400" location="forest northeast of Kuwapani" pageId="23" pageNumber="140" specimenCount="1" specimenCount-male="1" stateProvince="Khandbari">
•
<specimenCount id="F297B2EAFFA05B41FDD5FA3BE030FA6E" box="[527,576,1485,1511]" pageId="23" pageNumber="140" type="male">1 ♂</specimenCount>
;
<collectingRegion id="2655B781FFA05B41FD96FA3BE0B8FA6E" box="[588,712,1485,1511]" pageId="23" pageNumber="140">Khandbari</collectingRegion>
District,
<location id="E14E2FB8FFA05B41FCECFA3BE6FFFA6E" LSID="urn:lsid:plazi:treatment:6C38C875FFA05B4EFE2AFCE3E02EFD00:E14E2FB8FFA05B41FCECFA3BE6FFFA6E" box="[822,1167,1485,1511]" name="forest northeast of Kuwapani" pageId="23" pageNumber="140" stateProvince="Khandbari">forest northeast of Kuwapani</location>
; alt.
<quantity id="2369D486FFA05B41FB1DFA3BE752FA6E" box="[1223,1314,1485,1511]" metricMagnitude="3" metricUnit="m" metricValue="2.4" pageId="23" pageNumber="140" unit="m" value="2400.0">
<elevation id="6FBC9E50FFA05B41FB1DFA3BE752FA6E" box="[1223,1314,1485,1511]" metricMagnitude="3" metricUnit="m" metricValue="2.4" pageId="23" pageNumber="140" unit="m" value="2400.0">2400 m</elevation>
</quantity>
;
<date id="902F5FA3FFA05B41FAF5FA3BE28BF983" pageId="23" pageNumber="140" value="1984-04-05">
<collectingDate id="806BA64BFFA05B41FAF5FA3BE28BF983" pageId="23" pageNumber="140" value="1984-04-05">5 Apr. 1984</collectingDate>
</date>
;
<collectorName id="49641CB5FFA05B41FEDDFA06E31CF983" box="[263,364,1520,1546]" pageId="23" pageNumber="140">Smetana</collectorName>
&amp;
<collectorName id="49641CB5FFA05B41FE48FA06E3B9F983" box="[402,457,1520,1546]" pageId="23" pageNumber="140">Löbl</collectorName>
leg.;
<collectionCode id="8280E1A6FFA05B41FDD3FA06E036F983" box="[521,582,1520,1546]" pageId="23" pageNumber="140">ASC</collectionCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="AC8B2AE8FFA05B41FF67F9C7E6ECF914" pageId="23" pageNumber="140" type="description">
<paragraph id="E42E7963FFA05B41FF67F9C7E31BF9C2" blockId="23.[189,363,1585,1611]" box="[189,363,1585,1611]" pageId="23" pageNumber="140">
<heading id="BF66CE0FFFA05B41FF67F9C7E31BF9C2" bold="true" box="[189,363,1585,1611]" fontSize="11" level="3" pageId="23" pageNumber="140" reason="3">
<emphasis id="D6E5A571FFA05B41FF67F9C7E31BF9C2" bold="true" box="[189,363,1585,1611]" pageId="23" pageNumber="140">Redescription</emphasis>
</heading>
</paragraph>
<paragraph id="E42E7963FFA05B41FF67F996E6ECF914" blockId="23.[189,1398,1632,1693]" pageId="23" pageNumber="140">
<taxonomicName id="239102E0FFA05B41FF67F996E3EDF9F3" authorityName="Cameron" authorityYear="1932" box="[189,413,1632,1658]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="140" phylum="Arthropoda" rank="species" species="muscicola">
<emphasis id="D6E5A571FFA05B41FF67F996E3EDF9F3" box="[189,413,1632,1658]" italics="true" pageId="23" pageNumber="140">Quedius muscicola</emphasis>
</taxonomicName>
has been redescribed and diagnosed by
<bibRefCitation id="80000492FFA05B41FCAFF996E631F9F3" author="Smetana A." box="[885,1089,1632,1658]" pageId="23" pageNumber="140" refId="ref13920" refString="Smetana A. 2017 a. Quediine Subtribes of Staphylinini (Coleoptera, Staphylinidae, Staphylininae) of Mainland China. J. Farkac, Prague." type="book" year="2017">Smetana (2017a)</bibRefCitation>
. This redescription is only supplemented by the above diagnosis and illustration of the dorsal habitus (
<figureCitation id="7CAA65E6FFA05B41FBE8F975E6FEF914" box="[1074,1166,1667,1693]" captionStart="Fig" captionStartId="20.[187,226,1931,1955]" captionTargetBox="[190,1397,866,1890]" captionTargetId="figure-183@20.[189,1398,862,1893]" captionTargetPageId="20" captionText="Fig. 3. Dorsal habitus. A. Quedius (Raphirus) semilaeviventris Scheerpeltz, 1965, holotype, ♂ (NHRS). B. Q. (R.) kambaitiensis Scheerpeltz, 1965 syn. nov. (= Q. (R.) muscicola Cameron, 1932), ♀, holotype (NHRS). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7872186" httpUri="https://zenodo.org/record/7872186/files/figure.png" pageId="23" pageNumber="140">Fig. 3B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="AC8B2AE8FFA05B41FF67F932E687F8B8" pageId="23" pageNumber="140" type="distribution">
<paragraph id="E42E7963FFA05B41FF67F932E326F957" blockId="23.[189,342,1732,1758]" box="[189,342,1732,1758]" pageId="23" pageNumber="140">
<heading id="BF66CE0FFFA05B41FF67F932E326F957" bold="true" box="[189,342,1732,1758]" fontSize="11" level="3" pageId="23" pageNumber="140" reason="3">
<emphasis id="D6E5A571FFA05B41FF67F932E326F957" bold="true" box="[189,342,1732,1758]" pageId="23" pageNumber="140">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="E42E7963FFA05B41FF67F905E687F8B8" blockId="23.[189,1398,1779,1841]" pageId="23" pageNumber="140">
This widespread species is known from
<collectingCountry id="9C8639F3FFA05B41FD49F905E0BFF884" box="[659,719,1779,1805]" name="India" pageId="23" pageNumber="140">India</collectingCountry>
(
<collectingRegion id="2655B781FFA05B41FD04F905E109F884" box="[734,889,1779,1805]" country="India" name="West Bengal" pageId="23" pageNumber="140">West Bengal</collectingRegion>
: Darjeeling Distr.,
<collectingRegion id="2655B781FFA05B41FB8CF905E75FF884" box="[1110,1327,1779,1805]" country="India" name="Himachal Pradesh" pageId="23" pageNumber="140">Himachal Pradesh</collectingRegion>
,
<collectingRegion id="2655B781FFA05B41FAE0F902E36FF8B8" country="India" name="Uttar Pradesh" pageId="23" pageNumber="140">Uttar Pradesh</collectingRegion>
),
<collectingCountry id="9C8639F3FFA05B41FEE8F8E0E30BF8B9" box="[306,379,1814,1840]" name="Nepal" pageId="23" pageNumber="140">Nepal</collectingCountry>
,
<collectingCountry id="9C8639F3FFA05B41FE5DF8E1E38BF8B8" box="[391,507,1815,1841]" name="Myanmar" pageId="23" pageNumber="140">Myanmar</collectingCountry>
, and
<collectingCountry id="9C8639F3FFA05B41FDE2F8E0E00EF8B9" box="[568,638,1814,1840]" name="China" pageId="23" pageNumber="140">China</collectingCountry>
(
<collectingRegion id="2655B781FFA05B41FD54F8E1E0ADF8B8" box="[654,733,1815,1841]" country="China" name="Gansu" pageId="23" pageNumber="140">Gansu</collectingRegion>
,
<collectingRegion id="2655B781FFA05B41FD33F8E0E121F8B9" box="[745,849,1814,1840]" country="China" name="Guizhou" pageId="23" pageNumber="140">Guizhou</collectingRegion>
,
<collectingRegion id="2655B781FFA05B41FC87F8E0E1D8F8B9" box="[861,936,1814,1840]" country="China" name="Hubei" pageId="23" pageNumber="140">Hubei</collectingRegion>
,
<collectingRegion id="2655B781FFA05B41FC69F8E0E664F8B9" box="[947,1044,1814,1840]" country="China" name="Shaanxi" pageId="23" pageNumber="140">Shaanxi</collectingRegion>
,
<collectingRegion id="2655B781FFA05B41FBFAF8E0E6F1F8B9" box="[1056,1153,1814,1840]" country="China" name="Sichuan" pageId="23" pageNumber="140">Sichuan</collectingRegion>
,
<collectingRegion id="2655B781FFA05B41FB56F8E1E69BF8B8" box="[1164,1259,1815,1841]" country="China" name="Yunnan" pageId="23" pageNumber="140">Yunnan</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="AC8B2AE8FFA05B41FF67F8AEE366F86E" pageId="23" pageNumber="140" type="biology_ecology">
<paragraph id="E42E7963FFA05B41FF67F8AEE34FF8FB" blockId="23.[189,319,1880,1906]" box="[189,319,1880,1906]" pageId="23" pageNumber="140">
<heading id="BF66CE0FFFA05B41FF67F8AEE34FF8FB" bold="true" box="[189,319,1880,1906]" fontSize="11" level="3" pageId="23" pageNumber="140" reason="3">
<emphasis id="D6E5A571FFA05B41FF67F8AEE34FF8FB" bold="true" box="[189,319,1880,1906]" pageId="23" pageNumber="140">Bionomics</emphasis>
</heading>
</paragraph>
<paragraph id="E42E7963FFA05B41FF67F870E366F86E" blockId="23.[189,1399,1926,2023]" pageId="23" pageNumber="140">
<taxonomicName id="239102E0FFA05B41FF67F870E3D1F829" authorityName="Cameron" authorityYear="1932" box="[189,417,1926,1952]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="140" phylum="Arthropoda" rank="species" species="muscicola">
<emphasis id="D6E5A571FFA05B41FF67F870E3D1F829" box="[189,417,1926,1952]" italics="true" pageId="23" pageNumber="140">Quedius muscicola</emphasis>
</taxonomicName>
has been collected from forests, field edges and other anthropogenic habitats in moist leaf litter, moss and grassy vegetation, rotting wood and bark, compost and flood debris (
<bibRefCitation id="80000492FFA05B41FACAF85CE379F86E" author="Smetana A." pageId="23" pageNumber="140" refId="ref13920" refString="Smetana A. 2017 a. Quediine Subtribes of Staphylinini (Coleoptera, Staphylinidae, Staphylininae) of Mainland China. J. Farkac, Prague." type="book" year="2017">Smetana 2017a</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="AC8B2AE8FFAF5B4EFF67FEFFE02EFD00" pageId="24" pageNumber="141" type="discussion">
<paragraph id="E42E7963FFAF5B4EFF67FEFFE35FFEAA" blockId="24.[189,303,265,291]" box="[189,303,265,291]" pageId="24" pageNumber="141">
<heading id="BF66CE0FFFAF5B4EFF67FEFFE35FFEAA" bold="true" box="[189,303,265,291]" fontSize="11" level="3" pageId="24" pageNumber="141" reason="3">
<emphasis id="D6E5A571FFAF5B4EFF67FEFFE35FFEAA" bold="true" box="[189,303,265,291]" pageId="24" pageNumber="141">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="E42E7963FFAF5B4EFF67FEC1E1BDFD88" blockId="24.[188,1399,311,514]" pageId="24" pageNumber="141">
The male
<typeStatus id="3B2AC7C1FFAF5B4EFEE3FECEE3EEFEDB" box="[313,414,312,338]" pageId="24" pageNumber="141" type="paratype">paratype</typeStatus>
of
<taxonomicName id="239102E0FFAF5B4EFE16FEC1E0E4FED8" authorityName=", O. Scheerpeltz" authorityYear="1965" box="[460,660,311,337]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="141" phylum="Arthropoda" rank="species" species="kambaitiensis">
<emphasis id="D6E5A571FFAF5B4EFE16FEC1E0E4FED8" box="[460,660,311,337]" italics="true" pageId="24" pageNumber="141">Q. kambaitiensis</emphasis>
</taxonomicName>
was dissected to reveal an aedeagus entirely consistent with that of widespread and variable species
<emphasis id="D6E5A571FFAF5B4EFD48FEADE144FEFD" box="[658,820,346,373]" italics="true" pageId="24" pageNumber="141">
<taxonomicName id="239102E0FFAF5B4EFD48FEADE141FEFD" authorityName="Cameron" authorityYear="1932" box="[658,817,346,373]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="141" phylum="Arthropoda" rank="species" species="muscicola">Q. muscicola</taxonomicName>
.
</emphasis>
Therefore
<taxonomicName id="239102E0FFAF5B4EFC6DFEADE60CFEFD" authorityName=", O. Scheerpeltz" authorityYear="1965" box="[951,1148,346,373]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="141" phylum="Arthropoda" rank="species" species="kambaitiensis">
<emphasis id="D6E5A571FFAF5B4EFC6DFEADE60CFEFD" box="[951,1148,346,373]" italics="true" pageId="24" pageNumber="141">Q. kambaitiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CDD6180AFFAF5B4EFB5EFEADE69FFEFC" box="[1156,1263,347,373]" pageId="24" pageNumber="141">syn. nov.</taxonomicNameLabel>
, becomes a synonym of
<emphasis id="D6E5A571FFAF5B4EFE8AFE88E383FE1E" box="[336,499,381,408]" italics="true" pageId="24" pageNumber="141">
<taxonomicName id="239102E0FFAF5B4EFE8AFE88E39FFE1E" authorityName="Cameron" authorityYear="1932" box="[336,495,381,408]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="141" phylum="Arthropoda" rank="species" species="muscicola">Q. muscicola</taxonomicName>
.
</emphasis>
The determination was originally made by A. Smetana in 2018 but it is here confirmed after a comparison with similar species of the group occurring in both the Himalayan region and mainland
<collectingCountry id="9C8639F3FFAF5B4EFEBAFE32E3DAFE57" box="[352,426,452,478]" name="China" pageId="24" pageNumber="141">China</collectingCountry>
. Both the
<typeStatus id="3B2AC7C1FFAF5B4EFDC6FE32E0F3FE57" box="[540,643,452,478]" pageId="24" pageNumber="141" type="holotype">holotype</typeStatus>
and
<typeStatus id="3B2AC7C1FFAF5B4EFD63FE32E16EFE57" box="[697,798,452,478]" pageId="24" pageNumber="141" type="paratype">paratype</typeStatus>
are externally consistent with
<taxonomicName id="239102E0FFAF5B4EFBA7FE32E768FE57" authorityName="Cameron" authorityYear="1932" box="[1149,1304,452,478]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="141" phylum="Arthropoda" rank="species" species="muscicola">
<emphasis id="D6E5A571FFAF5B4EFBA7FE32E768FE57" box="[1149,1304,452,478]" italics="true" pageId="24" pageNumber="141">Q. muscicola</emphasis>
</taxonomicName>
and can be recognized as such by the features given in the diagnosis above.
</paragraph>
<paragraph id="E42E7963FFAF5B4EFF67FDDEE02EFD00" blockId="24.[187,1398,552,649]" pageId="24" pageNumber="141">
<taxonomicName id="239102E0FFAF5B4EFF67FDDEE3EBFDCB" authorityName="Cameron" authorityYear="1932" box="[189,411,552,578]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="141" phylum="Arthropoda" rank="species" species="muscicola">
<emphasis id="D6E5A571FFAF5B4EFF67FDDEE3EBFDCB" box="[189,411,552,578]" italics="true" pageId="24" pageNumber="141">Quedius muscicola</emphasis>
</taxonomicName>
is a rather variable species, especially outside of mainland
<collectingCountry id="9C8639F3FFAF5B4EFBE7FDDEE6F3FDCB" box="[1085,1155,552,578]" name="China" pageId="24" pageNumber="141">China</collectingCountry>
(
<bibRefCitation id="80000492FFAF5B4EFB4AFDDFE738FDCA" author="Smetana A." box="[1168,1352,553,579]" pageId="24" pageNumber="141" refId="ref13920" refString="Smetana A. 2017 a. Quediine Subtribes of Staphylinini (Coleoptera, Staphylinidae, Staphylininae) of Mainland China. J. Farkac, Prague." type="book" year="2017">Smetana 2017a</bibRefCitation>
), so it is not unexpected that
<bibRefCitation id="80000492FFAF5B4EFE0CFDBAE0C4FDEF" author="Scheerpeltz O." box="[470,692,587,614]" pageId="24" pageNumber="141" pagination="93 - 371" refId="ref13560" refString="Scheerpeltz O. 1965. Wissenschaftliche Ergebnisse der Schwedischen Expedition 1934 nach Indien und Burma. Arkiv for Zoologi 17: 93 - 371." type="journal article" year="1965">Scheerpeltz (1965)</bibRefCitation>
also reported
<taxonomicName id="239102E0FFAF5B4EFC83FDBAE183FDEF" authorityName="Cameron" authorityYear="1932" box="[857,1011,588,614]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="141" phylum="Arthropoda" rank="species" species="muscicola">
<emphasis id="D6E5A571FFAF5B4EFC83FDBAE183FDEF" box="[857,1011,588,614]" italics="true" pageId="24" pageNumber="141">Q. muscicola</emphasis>
</taxonomicName>
from Kambaiti, while describing
<taxonomicName id="239102E0FFAF5B4EFF66FD99E3F1FD00" authorityName=", O. Scheerpeltz" authorityYear="1965" box="[188,385,623,649]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="141" phylum="Arthropoda" rank="species" species="kambaitiensis">
<emphasis id="D6E5A571FFAF5B4EFF66FD99E3F1FD00" box="[188,385,623,649]" italics="true" pageId="24" pageNumber="141">Q. kambaitiensis</emphasis>
</taxonomicName>
in the same paper.
</paragraph>
</subSubSection>
</treatment>
</document>
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