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<document id="93EB1065C4EDF7B3B7846BAB903C971E" ID-DOI="10.5252/z2012n2a10" ID-ISSN="1638-9387" ID-Zenodo-Dep="5165469" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628213698565" checkinUser="felipe" docAuthor="Mclean, James Hamilton" docDate="2012" docId="6C26A814FF9F1472DA207A926CEFF112" docLanguage="en" docName="Zoosystema.34.2.343-376.pdf" docOrigin="Zoosystema 34 (2)" docSource="http://dx.doi.org/10.5252/z2012n2a10" docStyle="DocumentStyle:EB3C2C73672D2E883259A050BD27978B.6:Zoosystema.2011-2012.journal_article" docStyleId="EB3C2C73672D2E883259A050BD27978B" docStyleName="Zoosystema.2011-2012.journal_article" docStyleVersion="6" docTitle="Rhombipoma rowleyana Mclean 2012, n. sp." docType="treatment" docVersion="3" lastPageNumber="372" masterDocId="901FD06CFF84146FD92A7F586D0EF518" masterDocTitle="New species and genera of colloniids from Indo-Pacific coral reefs, with the definition of a new subfamily Liotipomatinae n. subfam. (Turbinoidea, Colloniidae)" masterLastPageNumber="376" masterPageNumber="343" pageNumber="370" updateTime="1699035883793" updateUser="plazi" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="D3E1D9E24AA64CDC5401322179DBAAE9" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="3F06AF8A8A7487AA7568A38608345286">
<mods:title id="F61ECB1AC40A702FA42DC08364DDB9F1">New species and genera of colloniids from Indo-Pacific coral reefs, with the definition of a new subfamily Liotipomatinae n. subfam. (Turbinoidea, Colloniidae)</mods:title>
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<mods:name id="E17AD22742C57F5A1AB2C809E614143D" type="personal">
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<mods:roleTerm id="EE1D1A25B4EC83724982AD2E7A55B617">Author</mods:roleTerm>
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<mods:namePart id="DADBA45D3A8594C9C07F9A7AC3A052BC">Mclean, James Hamilton</mods:namePart>
<mods:affiliation id="56B056F5877C8F4A272669230DD621C9">Natural History Museum of Los Angeles County, 900 Exposition Blvd, Los Angeles, California 90007 (USA) jmclean @ nhm. org</mods:affiliation>
<mods:nameIdentifier id="2555AADF4F3B3D6E0F65E6DD96D8C970" type="email">jmclean@nhm.org</mods:nameIdentifier>
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<mods:typeOfResource id="C11CECF786416F31C4F4902321A15793">text</mods:typeOfResource>
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<mods:title id="BCA50B8AB0BC0F5ABEFDF8B0F00B0B9D">Zoosystema</mods:title>
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<mods:part id="C1BE4333D2149D741C9C894EBD029318">
<mods:date id="6F5FF2B780EFC81A76778B8376AD8BCA">2012</mods:date>
<mods:detail id="B0924B82948FA6C2A2FF5050BFDB5A14" type="pubDate">
<mods:number id="8CB7CB40B8DB9FD714C35594B29DB312">2012-06-30</mods:number>
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<mods:number id="520A156249963557B226816BAC11A994">34</mods:number>
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<mods:number id="E8638D9A627CB6862FC195127B761888">2</mods:number>
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<mods:start id="F86CD1D781E296A2F830BE5AC25A1060">343</mods:start>
<mods:end id="F005F52C9ED00ADCE24504256A8EDFA3">376</mods:end>
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<mods:location id="5B16F75253975FA3742AF6047B73FA6F">
<mods:url id="DF604A87F5547E4EAEFBA9A1DEDD7323">http://dx.doi.org/10.5252/z2012n2a10</mods:url>
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<mods:classification id="7BEC82047A41F7F118077AC50EAE7F12">journal article</mods:classification>
<mods:identifier id="3255A867D826E7D272391CF847A3299B" type="DOI">10.5252/z2012n2a10</mods:identifier>
<mods:identifier id="42226EC7A08AF5552F48653CD660895F" type="ISSN">1638-9387</mods:identifier>
<mods:identifier id="C06AD1DBB6FF23B97D9C7F8386BB790A" type="Zenodo-Dep">5165469</mods:identifier>
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<treatment id="6C26A814FF9F1472DA207A926CEFF112" LSID="urn:lsid:plazi:treatment:6C26A814FF9F1472DA207A926CEFF112" httpUri="http://treatment.plazi.org/id/6C26A814FF9F1472DA207A926CEFF112" lastPageId="29" lastPageNumber="372" pageId="27" pageNumber="370">
<subSubSection id="AC954A89FF9F1474DA207A92695AF0FE" box="[778,1108,1482,1510]" pageId="27" pageNumber="370" type="nomenclature">
<paragraph id="E4301902FF9F1474DA207A92695AF0FE" blockId="27.[778,1108,1482,1541]" box="[778,1108,1482,1510]" pageId="27" pageNumber="370">
<heading id="BF78AE6EFF9F1474DA207A92695AF0FE" box="[778,1108,1482,1510]" centered="true" fontSize="11" level="2" pageId="27" pageNumber="370" reason="2">
<taxonomicName id="238F6281FF9F1474DA207A92691CF0FD" ID-CoL="4SJYR" authority="Mclean, 2012" authorityName="Mclean" authorityYear="2012" box="[778,1042,1482,1509]" class="Gastropoda" family="Colloniidae" genus="Rhombipoma" kingdom="Animalia" order="Trochida" pageId="27" pageNumber="370" phylum="Mollusca" rank="species" species="rowleyana" status="sp. nov.">
<emphasis id="D6FBC510FF9F1474DA207A92691CF0FD" bold="true" box="[778,1042,1482,1509]" italics="true" pageId="27" pageNumber="370">Rhombipoma rowleyana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CDC8786BFF9F1474DD337A94695AF0FE" box="[1049,1108,1484,1510]" pageId="27" pageNumber="370" rank="species">n. sp.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="AC954A89FF9F1474DA437AB36EFBF31D" box="[873,1013,1514,1541]" pageId="27" pageNumber="370" type="description">
<paragraph id="E4301902FF9F1474DA437AB36EFBF31D" blockId="27.[778,1108,1482,1541]" box="[873,1013,1514,1541]" pageId="27" pageNumber="370">
(
<figureCitation id="7CB40587FF9F1474DA5B7AB36EB5F31D" box="[881,955,1514,1541]" captionStart="FIG" captionStartId="30.[113,124,1299,1316]" captionTargetBox="[112,1208,206,1260]" captionTargetId="figure-243@30.[478,844,493,781]" captionTargetPageId="30" captionText="FIG. 15.— A-C, Rhombipoma rowleyana n. gen, n. sp. holotype (AMS C.212474,ex LACM stn 86-250),N end of Mermaid Reef, Rowley Shoals,off northern Western Australia,18-20 m,H 1.5,D 3.2 mm (base of shell broken and filled with agglutinated sand);D-F, Paraliotipoma sirenkoi n. gen., n. sp. holotype (ZISP 61624/1), Sea Horse Shoal, South China Sea, 10-320 m, H. 2.7, D 4.2 mm. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5165499" httpUri="https://zenodo.org/record/5165499/files/figure.png" pageId="27" pageNumber="370">Fig. 15</figureCitation>
A-C)
</paragraph>
</subSubSection>
<subSubSection id="AC954A89FF9F1473DB8F797A6CAFF35C" lastPageId="28" lastPageNumber="371" pageId="27" pageNumber="370" type="materials_examined">
<paragraph id="E4301902FF9F1473DB8F797A6DEDF0E2" blockId="27.[675,1211,1569,1699]" lastBlockId="28.[113,648,1453,1530]" lastPageId="28" lastPageNumber="371" pageId="27" pageNumber="370">
<materialsCitation id="54E7135FFF9F1473DB8F797A6DEDF0E2" collectingDate="1986-08-20" collectionCode="AMS, LACM" collectorName="Rowley Shoals, N" elevation="19" lastPageId="28" lastPageNumber="371" latitude="-17.05" location="Mermaid Reef" longLatPrecision="1279" longitude="119.6" pageId="27" pageNumber="370" specimenCount="1" stateProvince="Western Australia" typeStatus="holotype">
TYPE MATERIAL. —
<typeStatus id="3B34A7A0FF9F1474DA5D79796ED3F321" box="[887,989,1569,1593]" pageId="27" pageNumber="370" type="holotype">Holotype</typeStatus>
: stn 86-250,
<collectorName id="497A7CD4FF9F1474DD4779796E06F34B" pageId="27" pageNumber="370">Rowley Shoals, N</collectorName>
end
<location id="E1504FD9FF9F1474DA1579636EDBF34B" LSID="urn:lsid:plazi:treatment:6C26A814FF9F1472DA207A926CEFF112:E1504FD9FF9F1474DA1579636EDBF34B" box="[831,981,1595,1619]" latitude="-17.05" longLatPrecision="1279" longitude="119.6" name="Mermaid Reef" pageId="27" pageNumber="370" stateProvince="Western Australia">Mermaid Reef</location>
, off northern
<collectingRegion id="264BD7E0FF9F1474DD4D79646FF1F376" country="Australia" name="Western Australia" pageId="27" pageNumber="370">Western Australia</collectingRegion>
,
<geoCoordinate id="81BB7FC5FF9F1474DA20790E6E57F376" box="[778,857,1622,1646]" degrees="17" direction="south" minutes="03" orientation="latitude" pageId="27" pageNumber="370" precision="925" value="-17.05">17°03S</geoCoordinate>
,
<geoCoordinate id="81BB7FC5FF9F1474DA4E790E6ECDF376" box="[868,963,1622,1646]" degrees="119" direction="east" minutes="36" orientation="longitude" pageId="27" pageNumber="370" precision="925" value="119.6">119°36E</geoCoordinate>
,
<quantity id="2377B4E7FF9F1474DAE4790E6928F375" box="[974,1062,1622,1646]" metricMagnitude="1" metricUnit="m" metricValue="1.9" metricValueMax="2.0" metricValueMin="1.8" pageId="27" pageNumber="370" unit="m" value="19.0" valueMax="20.0" valueMin="18.0">
<elevation id="6FA2FE31FF9F1474DAE4790E6928F375" box="[974,1062,1622,1646]" metricMagnitude="1" metricUnit="m" metricValue="1.9" metricValueMax="2.0" metricValueMin="1.8" pageId="27" pageNumber="370" unit="m" value="19.0" valueMax="20.0" valueMin="18.0">18-20 m</elevation>
</quantity>
,
<date id="90313FC2FF9F1474DD1B790E69B9F376" box="[1073,1207,1622,1646]" pageId="27" pageNumber="370" value="1986-08-20">
<collectingDate id="8075C62AFF9F1474DD1B790E69B9F376" box="[1073,1207,1622,1646]" pageId="27" pageNumber="370" value="1986-08-20">20.VIII.1986</collectingDate>
</date>
, Twila Bratcher, in shell grit sample, 1 damaged shell with first whorl eroded and with agglutinated sand filling aperture and exposed final quarter whorl, in which base of shell is missing, H 1.5, D
<quantity id="2377B4E7FF981473D8AD7A906CDBF0C7" box="[391,469,1480,1504]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="28" pageNumber="371" unit="mm" value="3.2">3.2 mm</quantity>
(
<collectionCode id="829E81C7FF981473D8CB7A906F18F0F8" box="[481,534,1480,1504]" pageId="28" pageNumber="371">AMS</collectionCode>
C.212474, ex
<collectionCode id="829E81C7FF981473D9A77ABA6DD6F0E2" box="[141,216,1506,1530]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="28" pageNumber="371" type="Museum">LACM</collectionCode>
).
</materialsCitation>
</paragraph>
<caption id="B0F0498AFF981473D95B7A7A6C04F078" ID-DOI="http://doi.org/10.5281/zenodo.5165497" ID-Zenodo-Dep="5165497" httpUri="https://zenodo.org/record/5165497/files/figure.png" pageId="28" pageNumber="371" startId="28.[113,124,1314,1331]" targetBox="[112,1210,214,1271]" targetPageId="28">
<paragraph id="E4301902FF981473D95B7A7A6C04F078" blockId="28.[113,1211,1314,1376]" pageId="28" pageNumber="371">
FIG. 14. —
<emphasis id="D6FBC510FF981473D9ED7A7A6DE7F02B" bold="true" box="[199,233,1314,1331]" pageId="28" pageNumber="371">A-C</emphasis>
,
<taxonomicName id="238F6281FF981473D9D87A7A6CCEF02B" authority="Mclean, 2012" authorityName="Mclean" authorityYear="2012" box="[242,448,1314,1331]" class="Gastropoda" family="Colloniidae" genus="Depressipoma" kingdom="Animalia" order="Trochida" pageId="28" pageNumber="371" phylum="Mollusca" rank="species" species="kwajaleina" status="n. gen., n. sp.">
<emphasis id="D6FBC510FF981473D9D87A7A6CCEF02B" box="[242,448,1314,1331]" italics="true" pageId="28" pageNumber="371">Depressipoma kwajaleina</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CDC8786BFF981473D8EE7A7A6F3DF02B" box="[452,563,1314,1331]" pageId="28" pageNumber="371" rank="species">n. gen., n. sp.</taxonomicNameLabel>
holotype (LACM 3213),Kwajalein Atoll,Marshall Islands,20 m,H 1.7,D 3.2 mm;
<emphasis id="D6FBC510FF981473D95B7A606D9FF051" bold="true" box="[113,145,1336,1353]" pageId="28" pageNumber="371">D-F</emphasis>
,
<taxonomicName id="238F6281FF981473D9B77A606C4CF051" authority="Mclean, 2012" authorityName="Mclean" authorityYear="2012" box="[157,322,1336,1353]" class="Gastropoda" family="Colloniidae" genus="Depressipoma" kingdom="Animalia" order="Trochida" pageId="28" pageNumber="371" phylum="Mollusca" rank="species" species="laddi" status="n. gen., n. sp.">
<emphasis id="D6FBC510FF981473D9B77A606C4CF051" box="[157,322,1336,1353]" italics="true" pageId="28" pageNumber="371">Depressipoma laddi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CDC8786BFF981473D86D7A606CB2F051" box="[327,444,1336,1353]" pageId="28" pageNumber="371" rank="species">n. gen., n. sp.</taxonomicNameLabel>
holotype (USNM 648301), Elugelab Islet, Eniwetok Atoll, Marshall Islands, H 1.4, D 3.3 mm. Scale bars: 1 mm.
</paragraph>
</caption>
<paragraph id="E4301902FF981473D95B794A6CAFF35C" blockId="28.[112,647,1553,1604]" pageId="28" pageNumber="371">
<typeStatus id="3B34A7A0FF981473D95B794A6DACF33F" box="[113,162,1554,1577]" pageId="28" pageNumber="371">TYPE</typeStatus>
LOCALITY. —Rowley Shoals, N end Mermaid Reef, off northern
<collectingRegion id="264BD7E0FF981473D9C779746C93F35C" box="[237,413,1580,1604]" country="Australia" name="Western Australia" pageId="28" pageNumber="371">Western Australia</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="AC954A89FF981473D95B79046F6DF36B" box="[113,611,1627,1651]" pageId="28" pageNumber="371" type="distribution">
<paragraph id="E4301902FF981473D95B79046F6DF36B" blockId="28.[113,611,1627,1651]" box="[113,611,1627,1651]" pageId="28" pageNumber="371">
DISTRIBUTION. — Known only from the
<typeStatus id="3B34A7A0FF981473DB2C79036F51F36B" box="[518,607,1627,1651]" pageId="28" pageNumber="371" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="AC954A89FF981473D95B79D46F89F3BB" box="[113,647,1675,1700]" pageId="28" pageNumber="371" type="etymology">
<paragraph id="E4301902FF981473D95B79D46F89F3BB" blockId="28.[113,647,1675,1700]" box="[113,647,1675,1700]" pageId="28" pageNumber="371">
ETYMOLOGY. — After the
<typeStatus id="3B34A7A0FF981473D85979D46C93F3BC" box="[371,413,1676,1700]" pageId="28" pageNumber="371">type</typeStatus>
locality, Rowley Shoals.
</paragraph>
</subSubSection>
<subSubSection id="AC954A89FF981472DB8F7AF46DE8F756" lastPageId="29" lastPageNumber="372" pageId="28" pageNumber="371" type="description">
<paragraph id="E4301902FF981473DB8F7AF46E3BF0DC" blockId="28.[675,1211,1452,1700]" box="[677,821,1452,1478]" pageId="28" pageNumber="371">DESCRIPTION</paragraph>
<paragraph id="E4301902FF981473DB8F7A936E68F0FD" blockId="28.[675,1211,1452,1700]" box="[677,870,1483,1509]" pageId="28" pageNumber="371">
<emphasis id="D6FBC510FF981473DB8F7A936E68F0FD" box="[677,870,1483,1509]" italics="true" pageId="28" pageNumber="371">
Shell (
<figureCitation id="7CB40587FF981473DBCF7A936E3EF0FD" box="[741,816,1483,1509]" captionStart="FIG" captionStartId="30.[113,124,1299,1316]" captionTargetBox="[112,1208,206,1260]" captionTargetId="figure-243@30.[478,844,493,781]" captionTargetPageId="30" captionText="FIG. 15.— A-C, Rhombipoma rowleyana n. gen, n. sp. holotype (AMS C.212474,ex LACM stn 86-250),N end of Mermaid Reef, Rowley Shoals,off northern Western Australia,18-20 m,H 1.5,D 3.2 mm (base of shell broken and filled with agglutinated sand);D-F, Paraliotipoma sirenkoi n. gen., n. sp. holotype (ZISP 61624/1), Sea Horse Shoal, South China Sea, 10-320 m, H. 2.7, D 4.2 mm. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5165499" httpUri="https://zenodo.org/record/5165499/files/figure.png" pageId="28" pageNumber="371">Fig. 15</figureCitation>
A-C)
</emphasis>
</paragraph>
<paragraph id="E4301902FF981472DB8F7AB36DE4F4E8" blockId="28.[675,1211,1452,1700]" lastBlockId="29.[111,648,216,496]" lastPageId="29" lastPageNumber="372" pageId="28" pageNumber="371">
Of about three whorls, profile very low, whorls rounded. Axial ribs
<quantity id="2377B4E7FF981473DAAC79536ECCF33C" box="[902,962,1546,1573]" metricMagnitude="-1" metricUnit="m" metricValue="5.588" pageId="28" pageNumber="371" unit="in" value="22.0">22 in</quantity>
basal view, plus seven more closely spaced axials of lesser strength in advance of final lip. Spiral sculpture of two broad peripheral cords arising at final stage and extending over varix of final lip; spiral sculpture lacking on upper part of whorl; spiral sculpture on base of broad, low subtending cord; last half whorl rapidly expanding in diameter. Final lip comprised of about eight closely spaced axials of similar strength to those that precede the lip; final lip filling about ⅓ of final quarter whorl; base of final lip broken in
<typeStatus id="3B34A7A0FF991472D9B87ECE6DF8F4A8" box="[146,246,406,432]" pageId="29" pageNumber="372" type="holotype">holotype</typeStatus>
, but tip of tongue intact, club-like, extending forward of aperture. H 1.5, D
<quantity id="2377B4E7FF991472DB187EEE6F86F4C8" box="[562,648,438,464]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="29" pageNumber="372" unit="mm" value="3.2">3.2 mm</quantity>
(
<typeStatus id="3B34A7A0FF991472D9537E8D6DD0F4E8" box="[121,222,469,496]" pageId="29" pageNumber="372" type="holotype">holotype</typeStatus>
).
</paragraph>
<paragraph id="E4301902FF991472D95B7D4D6DEBF737" blockId="29.[113,230,533,590]" box="[113,229,533,559]" pageId="29" pageNumber="372">
<emphasis id="D6FBC510FF991472D95B7D4D6DEBF737" box="[113,229,533,559]" italics="true" pageId="29" pageNumber="372">Operculum</emphasis>
</paragraph>
<paragraph id="E4301902FF991472D95B7D6C6DE8F756" blockId="29.[113,230,533,590]" box="[113,230,564,590]" pageId="29" pageNumber="372">Unknown.</paragraph>
</subSubSection>
<subSubSection id="AC954A89FF991472D95B7D2C6CEFF112" pageId="29" pageNumber="372" type="discussion">
<paragraph id="E4301902FF991472D95B7D2C6DDBF794" blockId="29.[111,648,628,1034]" box="[113,213,628,654]" pageId="29" pageNumber="372">REMARKS</paragraph>
<paragraph id="E4301902FF991472D95B7DCB6CEFF112" blockId="29.[111,648,628,1034]" pageId="29" pageNumber="372">
<taxonomicName id="238F6281FF991472D95B7DCB6C6AF7B5" authority="Mclean, 2012" authorityName="Mclean" authorityYear="2012" box="[113,356,659,685]" class="Gastropoda" family="Colloniidae" genus="Rhombipoma" kingdom="Animalia" order="Trochida" pageId="29" pageNumber="372" phylum="Mollusca" rank="species" species="rowleyana" status="n. gen., n. sp.">
<emphasis id="D6FBC510FF991472D95B7DCB6C6AF7B5" box="[113,356,659,685]" italics="true" pageId="29" pageNumber="372">Rhombipoma rowleyana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CDC8786BFF991472D8437DCC6CFDF7B6" box="[361,499,660,686]" pageId="29" pageNumber="372" rank="species">n. gen., n. sp.</taxonomicNameLabel>
is comparable to
<taxonomicName id="238F6281FF991472D9A47DEC6DEEF7D5" authority="Mclean, 2012" authorityName="Mclean" authorityYear="2012" box="[142,224,691,717]" class="Gastropoda" family="Colloniidae" genus="Depressipoma" kingdom="Animalia" order="Trochida" pageId="29" pageNumber="372" phylum="Mollusca" rank="species" species="laddi" status="sp. nov.">
<emphasis id="D6FBC510FF991472D9A47DEC6DEEF7D5" box="[142,224,691,717]" italics="true" pageId="29" pageNumber="372">L. laddi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CDC8786BFF991472D9CC7DEC6C2EF7D6" box="[230,288,692,718]" pageId="29" pageNumber="372" rank="species">n. sp.</taxonomicNameLabel>
(
<figureCitation id="7CB40587FF991472D8077DEB6C79F7D5" box="[301,375,691,717]" captionStart="FIG" captionStartId="28.[113,124,1314,1331]" captionTargetBox="[112,1210,214,1271]" captionTargetId="figure-205@28.[478,845,478,744]" captionTargetPageId="28" captionText="FIG. 14. — A-C, Depressipoma kwajaleina n. gen., n. sp.holotype (LACM 3213),Kwajalein Atoll,Marshall Islands,20 m,H 1.7,D 3.2 mm; D-F, Depressipoma laddi n. gen., n. sp. holotype (USNM 648301), Elugelab Islet, Eniwetok Atoll, Marshall Islands, H 1.4, D 3.3 mm. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5165497" httpUri="https://zenodo.org/record/5165497/files/figure.png" pageId="29" pageNumber="372">Fig. 14</figureCitation>
D-F) in its extremely low profile, and even more rounded final whorl, but differs in lacking the prominent spiral cords that produce the clathrate sculpture of that species. The full morphology of the umbilical area is not clear because the base of the shell is broken, but if this species is sexually dimorphic, it is unlikely that the
<typeStatus id="3B34A7A0FF991472D95A7CC96DC3F6B3" box="[112,205,913,939]" pageId="29" pageNumber="372" type="holotype">holotype</typeStatus>
is a female shell because there seems not to be an umbilical cavity bordered by a raised rib. It is likely that the extreme lenticular profile precludes sexual dimorphism in this species.
</paragraph>
</subSubSection>
</treatment>
</document>