treatments-xml/data/6A/59/87/6A598782FFF6ED1CFF1FF951EE4145B9.xml

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<mods:title id="6F551D70A6D4D462081BA0B961545F6D">New species of Cyrtodactylus (Squamata: Gekkonidae) from Karkar Island, Papua New Guinea</mods:title>
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<mods:namePart id="EF4C05B919BD530E6D397356B7A02426">Kraus, Fred</mods:namePart>
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<mods:namePart id="4C937D094283FD8CB1B869373A1EE3A1">Weijola, Valter</mods:namePart>
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<mods:date id="99CB9A89C472C340AFF6D2F696B6731B">2019</mods:date>
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<mods:number id="44092784F5EA650F3536D77013E4A8D1">2019-11-13</mods:number>
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<mods:number id="5399FA4E8FCDAC9FE5D2639CC45BB067">4695</mods:number>
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<treatment id="6A598782FFF6ED1CFF1FF951EE4145B9" ID-GBIF-Taxon="160406215" LSID="urn:lsid:zoobank.org:act:DBED7B72-5A6C-486B-945C-522B3729FA52" httpUri="http://treatment.plazi.org/id/6A598782FFF6ED1CFF1FF951EE4145B9" lastPageId="7" lastPageNumber="536" pageId="1" pageNumber="530">
<subSubSection id="AAEA651FFFF6ED1AFF1FF951EF6142EA" box="[151,523,1737,1764]" pageId="1" pageNumber="530" type="nomenclature">
<paragraph id="E24F3694FFF6ED1AFF1FF951EF6142EA" blockId="1.[151,523,1737,1799]" box="[151,523,1737,1764]" pageId="1" pageNumber="530">
<heading id="B90781F8FFF6ED1AFF1FF951EF6142EA" bold="true" box="[151,523,1737,1764]" fontSize="11" level="1" pageId="1" pageNumber="530" reason="1">
<emphasis id="D084EA86FFF6ED1AFF1FF951EF6142EA" bold="true" box="[151,523,1737,1764]" pageId="1" pageNumber="530">
<taxonomicName id="25F04D17FFF6ED1AFF1FF951ECCD42EA" authority="Kraus &amp; Weijola, 2019" authorityName="Kraus &amp; Weijola" authorityYear="2019" box="[151,423,1737,1764]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="530" phylum="Chordata" rank="species" species="atremus" status="sp. nov.">
<emphasis id="D084EA86FFF6ED1AFF1FF951ECCD42EA" bold="true" box="[151,423,1737,1764]" italics="true" pageId="1" pageNumber="530">Cyrtodactylus atremus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CBB757FDFFF6ED1AFE26F951EF6142EA" box="[430,523,1737,1763]" pageId="1" pageNumber="530" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="AAEA651FFFF6ED1AFF1FF975EF1D4340" pageId="1" pageNumber="530" type="description">
<paragraph id="E24F3694FFF6ED1AFF1FF975EC33430E" blockId="1.[151,523,1737,1799]" box="[151,345,1772,1799]" pageId="1" pageNumber="530">
<figureCitation id="7ACB2A11FFF6ED1AFF1FF975EDB3430E" box="[151,217,1773,1799]" captionStart="FIGURE 1" captionStartId="3.[151,250,1935,1960]" captionTargetBox="[162,1424,456,1901]" captionTargetId="figure@3.[151,1436,445,1916]" captionTargetPageId="3" captionText="FIGURE 1. (A) Top of head, (B) side of head, (C) mid-dorsum, and (D) left leg of holotype of Cyrtodactylus atremus sp. nov., UMMZ 247747, Karkar Island, Madang Province, PNG." figureDoi="http://doi.org/10.5281/zenodo.3542679" httpUri="https://zenodo.org/record/3542679/files/figure.png" pageId="1" pageNumber="530">Fig. 1</figureCitation>
, 
<figureCitation id="7ACB2A11FFF6ED1AFF6FF975EC61430E" box="[231,267,1773,1799]" captionStart="FIGURE 3" captionStartId="6.[151,250,1184,1209]" captionTargetBox="[156,1431,307,1157]" captionTargetId="figure@6.[151,1436,301,1164]" captionTargetPageId="6" captionText="FIGURE 3. Portraits in life of (A) paratype of Cyrtodactylus atremus sp. nov., UMMZ 247746, Karkar Island, Madang Province, PNG; (B) Cyrtodactylus sermowaiensis, BPBM 23317, Parkop, Torricelli Mts., West Sepik Province, PNG; (C) Cyrtodactylus sermowaiensis, BPBM 23320, base of Mt. Sapau, Torricelli Mts., West Sepik Province, PNG; and (D) Cyrtodactylus sermowaiensis, BPBM 35598, Yefehi, Prince Alexander Mts., East Sepik Province, PNG." figureDoi="http://doi.org/10.5281/zenodo.3542685" httpUri="https://zenodo.org/record/3542685/files/figure.png" pageId="1" pageNumber="530">3A</figureCitation>
, 
<figureCitation id="7ACB2A11FFF6ED1AFE9FF974EC33430E" box="[279,345,1772,1799]" captionStart="FIGURE 5" captionStartId="8.[151,250,1578,1603]" captionTargetBox="[160,1427,379,1550]" captionTargetId="figure@8.[151,1436,373,1558]" captionTargetPageId="8" captionText="FIGURE 5. (A), (B) Habitat of Cyrtodactylus atremus sp. nov. on the lower northwestern slopes of Mt. Uluman, Karkar Island, Madang Province, PNG; (C) UMMZ 247751, and (D) UMMZ 247750 in situ showing their head-down resting position during the daytime." figureDoi="http://doi.org/10.5281/zenodo.3542689" httpUri="https://zenodo.org/record/3542689/files/figure.png" pageId="1" pageNumber="530">5C, D</figureCitation>
</paragraph>
<paragraph id="E24F3694FFF6ED1AFF1FF8A9EF1D4340" blockId="1.[151,631,1840,1865]" box="[151,631,1840,1865]" pageId="1" pageNumber="530">
<taxonomicName id="25F04D17FFF6ED1AFF1FF8A9EC4F4340" authorityName="Gray" authorityYear="1827" box="[151,293,1841,1865]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="530" phylum="Chordata" rank="genus">
<emphasis id="D084EA86FFF6ED1AFF1FF8A9EC4F4340" box="[151,293,1841,1865]" italics="true" pageId="1" pageNumber="530">Cyrtodactylus</emphasis>
</taxonomicName>
sp. ‘Karkar’ 
<bibRefCitation id="86614B65FFF6ED1AFE23F8A8EF1D4340" author="Tallowin, O. J. S. &amp; Tamar, K. &amp; Meiri, S. &amp; Allison, A. &amp; Kraus, F. &amp; Richards, S. J. &amp; Oliver, P. M." box="[427,631,1840,1865]" pageId="1" pageNumber="530" pagination="29 - 39" refId="ref8115" refString="Tallowin, O. J. S., Tamar, K., Meiri, S., Allison, A., Kraus, F., Richards, S. J. &amp; Oliver, P. M. (2018) Early insularity and mountain uplift were complrmentary drivers of diversification in a Melanesian lizard radiation (Gekkonidae: Cyrtodactylus). Molecular Phylogenetics and Evolution, 125, 29 - 39. https: // doi. org / 10.1016 / j. ympev. 2018.03.020" type="journal article" year="2018">
Tallowin 
<emphasis id="D084EA86FFF6ED1AFD83F8AAEF544340" box="[523,574,1841,1865]" italics="true" pageId="1" pageNumber="530">et al.</emphasis>
2018
</bibRefCitation>
</paragraph>
</subSubSection>
<subSubSection id="AAEA651FFFF6ED19FF1FF8EFEC6544F5" lastPageId="2" lastPageNumber="531" pageId="1" pageNumber="530" type="materials_examined">
<paragraph id="E24F3694FFF6ED1AFF1FF8EFE84943BF" blockId="1.[151,1436,1911,2010]" pageId="1" pageNumber="530">
<materialsCitation id="52983CC9FFF6ED1AFF1FF8EFE84943BF" ID-GBIF-Occurrence="2451575187" collectingDate="2018-02-26" collectionCode="UMMZ, FK" collectorName="V. Weijola &amp; Mom" country="Papua New Guinea" elevation="573" latitude="-4.6248" location="Karkar Island" longLatPrecision="7" longitude="145.9378" pageId="1" pageNumber="530" specimenCode="UMMZ 247747, FK 17685" specimenCount="1" stateProvince="Madang" typeStatus="holotype">
<emphasis id="D084EA86FFF6ED1AFF1FF8EFEC6C439B" bold="true" box="[151,262,1911,1938]" pageId="1" pageNumber="530">
<emphasis id="D084EA86FFF6ED1AFF1FF8EFED95439B" bold="true" box="[151,255,1911,1938]" italics="true" pageId="1" pageNumber="530">
<typeStatus id="3D4B8836FFF6ED1AFF1FF8EFED95439B" box="[151,255,1911,1938]" pageId="1" pageNumber="530" type="holotype">Holotype</typeStatus>
</emphasis>
.
</emphasis>
<specimenCode id="B2569EEFFFF6ED1AFE9AF8E0ECA2439B" box="[274,456,1912,1938]" collectionCode="UMMZ" country="USA" httpUri="http://grbio.org/cool/ra64-3eva" name="University of Michigan, Museum of Zoology" pageId="1" pageNumber="530" type="University or college">UMMZ 247747</specimenCode>
(field tag 
<specimenCode id="B2569EEFFFF6ED1AFDC1F8E0EFA8439B" box="[585,706,1911,1938]" collectionCode="FK" pageId="1" pageNumber="530">FK 17685</specimenCode>
), mature male collected by 
<collectorName id="4F055342FFF6ED1AFB84F8E0E9EE4398" box="[1036,1156,1911,1938]" pageId="1" pageNumber="530">V. Weijola</collectorName>
above 
<collectorName id="4F055342FFF6ED1AFB56F8E0E874439B" box="[1246,1310,1912,1938]" pageId="1" pageNumber="530">Mom</collectorName>
, 
<geoCoordinate id="87C45053FFF6ED1AFAA5F8EFE8F24398" box="[1325,1432,1911,1937]" degrees="4.6248" direction="south" orientation="latitude" pageId="1" pageNumber="530" precision="5" value="-4.6248">4.6248°S</geoCoordinate>
, 
<geoCoordinate id="87C45053FFF6ED1AFF1FF803EC7543BF" box="[151,287,1947,1974]" degrees="145.9378" direction="east" orientation="longitude" pageId="1" pageNumber="530" precision="5" value="145.9378">145.9378°E</geoCoordinate>
, 
<elevation id="69DDD1A7FFF6ED1AFEA2F803EC8C43BC" box="[298,486,1947,1974]" metricMagnitude="2" metricUnit="m" metricValue="5.725" metricValueMax="6.45" metricValueMin="5.0" pageId="1" pageNumber="530" unit="m" value="572.5" valueMax="645.0" valueMin="500.0">
<quantity id="25089B71FFF6ED1AFEA2F803ECC343BF" box="[298,425,1947,1974]" metricMagnitude="2" metricUnit="m" metricValue="5.725" metricValueMax="6.45" metricValueMin="5.0" pageId="1" pageNumber="530" unit="m" value="572.5" valueMax="645.0" valueMin="500.0">500–645 m</quantity>
a.s.l.
</elevation>
, 
<location id="E72F604FFFF6ED1AFE7AF803EFE443BC" LSID="urn:lsid:plazi:treatment:6A598782FFF6ED1CFF1FF951EE4145B9:E72F604FFFF6ED1AFE7AF803EFE443BC" box="[498,654,1947,1973]" country="Papua New Guinea" latitude="-4.6248" longLatPrecision="7" longitude="145.9378" name="Karkar Island" pageId="1" pageNumber="530" stateProvince="Madang">Karkar Island</location>
, 
<collectingRegion id="2034F876FFF6ED1AFD12F803EE0843BC" box="[666,866,1947,1973]" country="Papua New Guinea" name="Madang" pageId="1" pageNumber="530">Madang Province</collectingRegion>
, 
<collectingCountry id="9AE77604FFF6ED1AFCE6F804E92343BC" box="[878,1097,1947,1974]" name="Papua New Guinea" pageId="1" pageNumber="530">Papua New Guinea</collectingCountry>
, 
<date id="964E1054FFF6ED1AFBDCF803E87543BF" box="[1108,1311,1947,1974]" pageId="1" pageNumber="530" value="2018-02-26">
<collectingDate id="860AE9BCFFF6ED1AFBDCF803E87543BF" box="[1108,1311,1947,1974]" pageId="1" pageNumber="530" value="2018-02-26">26 February 2018</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
<paragraph id="E24F3694FFF6ED19FF4FF858EC6544F5" blockId="1.[151,1436,1911,2010]" lastBlockId="2.[151,1437,153,2015]" lastPageId="2" lastPageNumber="531" pageId="1" pageNumber="530">
<materialsCitation id="52983CC9FFF6ED19FF4FF858ED9244BD" ID-GBIF-Occurrence="2451575188" collectingDate="2018-02-26" collectionCode="UMMZ" collectorName="V. Weijola &amp; Mom" country="Papua New Guinea" elevation="573" lastPageId="2" lastPageNumber="531" latitude="-4.6248" location="Karkar Island" longLatPrecision="7" longitude="145.9378" municipality="Male" pageId="1" pageNumber="530" specimenCode="UMMZ 247746" specimenCount="1" stateProvince="Madang" typeStatus="paratype">
<emphasis id="D084EA86FFF6ED1AFF4FF858ECFE43D3" box="[199,404,1984,2010]" italics="true" pageId="1" pageNumber="530">
<emphasis id="D084EA86FFF6ED1AFF4FF858EC5343D3" bold="true" box="[199,313,1984,2010]" italics="true" pageId="1" pageNumber="530">
<typeStatus id="3D4B8836FFF6ED1AFF4FF858EC5343D3" box="[199,313,1984,2010]" pageId="1" pageNumber="530" type="paratype">Paratypes</typeStatus>
</emphasis>
(n = 8).
</emphasis>
<collectingMunicipality id="022BACEEFFF6ED1AFE12F827ECB243D0" box="[410,472,1983,2009]" pageId="1" pageNumber="530">Male</collectingMunicipality>
: 
<collectingCountry id="9AE77604FFF6ED1AFE6AF858EFD743D0" box="[482,701,1983,2010]" name="Papua New Guinea" pageId="1" pageNumber="530">Papua New Guinea</collectingCountry>
: 
<collectingRegion id="2034F876FFF6ED1AFD4FF827EEFB43D0" box="[711,913,1983,2009]" country="Papua New Guinea" name="Madang" pageId="1" pageNumber="530">Madang Province</collectingRegion>
: 
<location id="E72F604FFFF6ED1AFC13F827E95243D0" LSID="urn:lsid:plazi:treatment:6A598782FFF6ED1CFF1FF951EE4145B9:E72F604FFFF6ED1AFC13F827E95243D0" box="[923,1080,1983,2009]" country="Papua New Guinea" municipality="Male" name="Karkar Island" pageId="1" pageNumber="530" stateProvince="Madang">Karkar Island</location>
: same data as holotype (
<specimenCode id="B2569EEFFFF6ED19FACBF858ED9A44BD" collectionCode="UMMZ" country="USA" httpUri="http://grbio.org/cool/ra64-3eva" lastPageId="2" lastPageNumber="531" name="University of Michigan, Museum of Zoology" pageId="1" pageNumber="530" type="University or college">UMMZ 247746</specimenCode>
)
</materialsCitation>
; 
<materialsCitation id="52983CC9FFF5ED19FE8BFF01EDD544DE" ID-GBIF-Occurrence="2451575185" collectingDate="2018-02-26" collectionCode="UMMZ" collectorName="V. Weijola &amp; Mom" country="Papua New Guinea" elevation="573" latitude="-4.6248" location="Karkar Island" longLatPrecision="7" longitude="145.9378" municipality="Females" pageId="2" pageNumber="531" specimenCode="UMMZ 247748- 51" specimenCount="1" stateProvince="Madang" typeStatus="paratype">
<collectingMunicipality id="022BACEEFFF5ED19FE8BFF01EC0F44BA" box="[259,357,153,179]" pageId="2" pageNumber="531">Females</collectingMunicipality>
: 
<collectingCountry id="9AE77604FFF5ED19FEF8FF01EF2644BA" box="[368,588,153,179]" name="Papua New Guinea" pageId="2" pageNumber="531">Papua New Guinea</collectingCountry>
: 
<collectingRegion id="2034F876FFF5ED19FDDFFF01EE4844BA" box="[599,802,153,179]" country="Papua New Guinea" name="Madang" pageId="2" pageNumber="531">Madang Province</collectingRegion>
: 
<location id="E72F604FFFF5ED19FCA4FF01EEA044BA" LSID="urn:lsid:plazi:treatment:6A598782FFF6ED1CFF1FF951EE4145B9:E72F604FFFF5ED19FCA4FF01EEA044BA" box="[812,970,153,179]" country="Papua New Guinea" municipality="Females" name="Karkar Island" pageId="2" pageNumber="531" stateProvince="Madang">Karkar Island</location>
: same data as holotype (
<specimenCode id="B2569EEFFFF5ED19FB52FF01EDDF44DE" collectionCode="UMMZ" country="USA" httpUri="http://grbio.org/cool/ra64-3eva" name="University of Michigan, Museum of Zoology" pageId="2" pageNumber="531" type="University or college">UMMZ 247748– 51</specimenCode>
)
</materialsCitation>
, 
<materialsCitation id="52983CC9FFF5ED19FF44FF25EEFE44DE" ID-GBIF-Occurrence="2451575189" box="[204,916,188,215]" collectionCode="AMS" country="Papua New Guinea" elevation="290" latitude="-4.61" longLatPrecision="783" longitude="145.92" pageId="2" pageNumber="531" specimenCount="1" stateProvince="Madang" typeStatus="paratype">
Mom, 
<geoCoordinate id="87C45053FFF5ED19FE90FF25EC0C44DE" box="[280,358,189,215]" degrees="4.61" direction="south" orientation="latitude" pageId="2" pageNumber="531" precision="555" value="-4.61">4.61°S</geoCoordinate>
, 
<geoCoordinate id="87C45053FFF5ED19FEFBFF24ECB544DE" box="[371,479,188,215]" degrees="145.92" direction="east" orientation="longitude" pageId="2" pageNumber="531" precision="555" value="145.92">145.92°E</geoCoordinate>
, 
<elevation id="69DDD1A7FFF5ED19FE64FF25EF1B44DE" box="[492,625,189,215]" metricMagnitude="2" metricUnit="m" metricValue="2.9" pageId="2" pageNumber="531" unit="m" value="290.0">
<quantity id="25089B71FFF5ED19FE64FF25EF5E44DE" box="[492,564,189,215]" metricMagnitude="2" metricUnit="m" metricValue="2.9" pageId="2" pageNumber="531" unit="m" value="290.0">290 m</quantity>
a.s.l.
</elevation>
(
<collectionCode id="84E1AE51FFF5ED19FD08FF25EFD444DE" box="[640,702,189,215]" pageId="2" pageNumber="531">AMS</collectionCode>
R24744, R28901)
</materialsCitation>
, 
<materialsCitation id="52983CC9FFF5ED19FC28FF25EC6144F5" ID-GBIF-Occurrence="2451575186" collectionCode="AMS" country="Papua New Guinea" elevation="130" latitude="-4.7" location="Kevasob" longLatPrecision="7843" longitude="145.9" pageId="2" pageNumber="531" specimenCode="R124604" specimenCount="1" stateProvince="Madang" typeStatus="paratype">
<location id="E72F604FFFF5ED19FC28FF25E96F44DE" LSID="urn:lsid:plazi:treatment:6A598782FFF6ED1CFF1FF951EE4145B9:E72F604FFFF5ED19FC28FF25E96F44DE" box="[928,1029,189,215]" country="Papua New Guinea" latitude="-4.7" longLatPrecision="7843" longitude="145.9" name="Kevasob" pageId="2" pageNumber="531" stateProvince="Madang">Kevasob</location>
, 
<geoCoordinate id="87C45053FFF5ED19FB9AFF25E93844DE" box="[1042,1106,189,215]" degrees="4.7" direction="south" orientation="latitude" pageId="2" pageNumber="531" precision="5555" value="-4.7">4.7°S</geoCoordinate>
, 
<geoCoordinate id="87C45053FFF5ED19FBD7FF24E9D644DE" box="[1119,1212,188,215]" degrees="145.9" direction="east" orientation="longitude" pageId="2" pageNumber="531" precision="5555" value="145.9">145.9°E</geoCoordinate>
, 
<elevation id="69DDD1A7FFF5ED19FB41FF25E82444DE" box="[1225,1358,189,215]" metricMagnitude="2" metricUnit="m" metricValue="1.3" pageId="2" pageNumber="531" unit="m" value="130.0">
<quantity id="25089B71FFF5ED19FB41FF25E87844DE" box="[1225,1298,189,215]" metricMagnitude="2" metricUnit="m" metricValue="1.3" pageId="2" pageNumber="531" unit="m" value="130.0">130 m</quantity>
a.s.l.
</elevation>
(
<collectionCode id="84E1AE51FFF5ED19FAD6FF25E8F644DE" box="[1374,1436,189,215]" pageId="2" pageNumber="531">AMS</collectionCode>
<specimenCode id="B2569EEFFFF5ED19FF1FFF79EC6944F5" box="[151,259,225,252]" collectionCode="AMS" pageId="2" pageNumber="531">R124604</specimenCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="AAEA651FFFF5ED19FF4FFE9DEEC04756" pageId="2" pageNumber="531" type="diagnosis">
<paragraph id="E24F3694FFF5ED19FF4FFE9DE81C45DA" blockId="2.[151,1437,153,2015]" pageId="2" pageNumber="531">
<emphasis id="D084EA86FFF5ED19FF4FFE9DEC2B4516" bold="true" box="[199,321,261,287]" pageId="2" pageNumber="531">
<emphasis id="D084EA86FFF5ED19FF4FFE9DEC504516" bold="true" box="[199,314,261,287]" italics="true" pageId="2" pageNumber="531">Diagnosis</emphasis>
.
</emphasis>
A moderately large (adult SVL = 
<quantity id="25089B71FFF5ED19FD5EFE9DEE044516" box="[726,878,261,287]" metricMagnitude="-1" metricUnit="m" metricValue="1.13" metricValueMax="1.18" metricValueMin="1.08" pageId="2" pageNumber="531" unit="mm" value="113.0" valueMax="118.0" valueMin="108.0">108–118 mm</quantity>
) species of 
<taxonomicName id="25F04D17FFF5ED19FC75FE9DE9F04516" authorityName="Gray" authorityYear="1827" box="[1021,1178,261,287]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="531" phylum="Chordata" rank="genus">
<emphasis id="D084EA86FFF5ED19FC75FE9DE9F04516" box="[1021,1178,261,287]" italics="true" pageId="2" pageNumber="531">Cyrtodactylus</emphasis>
</taxonomicName>
with a small patch of somewhat enlarged precloacal scales (ca. 2–3 times size of nearby femoral scales), no enlarged femoral scales, no precloacal/femoral pores in males, and having a highly tuberculate dorsum, 22–24 rows of dorsal tubercles between the lateral skin folds, dorsal tubercles 3–5 times size of adjacent granules, 24–31 scale rows below the fourth toe, 9–11 supralabials to the center of eye, 10–13 infralabials to rictus, 40–47 mid-belly scale rows, 5–6 dark dorsal crossbands to the base of the tail, uniformly brown top of head, and lips dirty white with some brown stippling.
</paragraph>
<paragraph id="E24F3694FFF5ED19FF4FFE45EEC04756" blockId="2.[151,1437,153,2015]" pageId="2" pageNumber="531">
<emphasis id="D084EA86FFF5ED19FF4FFE45EF5245FE" bold="true" box="[199,568,476,503]" pageId="2" pageNumber="531">
<emphasis id="D084EA86FFF5ED19FF4FFE45EF5B45FE" bold="true" box="[199,561,476,503]" italics="true" pageId="2" pageNumber="531">Comparisons with other species</emphasis>
.
</emphasis>
The absence of enlarged femoral scales and precloacal/femoral pores in males distinguishes the new species from all other Papuan 
<taxonomicName id="25F04D17FFF5ED19FD79FD99EEE44612" authorityName="Gray" authorityYear="1827" box="[753,910,513,539]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="531" phylum="Chordata" rank="genus">
<emphasis id="D084EA86FFF5ED19FD79FD99EEE44612" box="[753,910,513,539]" italics="true" pageId="2" pageNumber="531">Cyrtodactylus</emphasis>
</taxonomicName>
except 
<taxonomicName id="25F04D17FFF5ED19FC61FD99E8394612" authority="(Kraus 2008)" authorityName="Kraus" authorityYear="2008" baseAuthorityName="Kraus" baseAuthorityYear="2008" box="[1001,1363,513,539]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="531" phylum="Chordata" rank="species" species="sermowaiensis">
<emphasis id="D084EA86FFF5ED19FC61FD99E9DA4612" box="[1001,1200,513,539]" italics="true" pageId="2" pageNumber="531">C. sermowaiensis</emphasis>
(
<bibRefCitation id="86614B65FFF5ED19FB49FD99E8214612" author="Kraus, F." box="[1217,1355,513,539]" pageId="2" pageNumber="531" pagination="1 - 27" refId="ref7448" refString="Kraus, F. (2008) Taxonomic partitioning of Cyrtodactylus louisiadensis (Lacertilia: Gekkonidae) from Papua New Guinea. Zootaxa, 1883, 1 - 27. https: // doi. org / 10.11646 / zootaxa. 1883.1.1" type="journal article" year="2008">Kraus 2008</bibRefCitation>
)
</taxonomicName>
. From that species, 
<taxonomicName id="25F04D17FFF5ED19FEAFFDBDECCB4636" authorityName="Kraus &amp; Weijola" authorityYear="2019" box="[295,417,549,575]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="531" phylum="Chordata" rank="species" species="atremus">
<emphasis id="D084EA86FFF5ED19FEAFFDBDECCB4636" box="[295,417,549,575]" italics="true" pageId="2" pageNumber="531">C. atremus</emphasis>
</taxonomicName>
differs in its larger size (adult SVL = 
<quantity id="25089B71FFF5ED19FCC2FDBDEEB44636" box="[842,990,549,575]" metricMagnitude="-1" metricUnit="m" metricValue="1.13" metricValueMax="1.18" metricValueMin="1.08" pageId="2" pageNumber="531" unit="mm" value="113.0" valueMax="118.0" valueMin="108.0">108–118 mm</quantity>
<emphasis id="D084EA86FFF5ED19FC6CFDBEE9464636" box="[996,1068,550,575]" italics="true" pageId="2" pageNumber="531">versus</emphasis>
<quantity id="25089B71FFF5ED19FBBBFDBCE9C64636" box="[1075,1196,548,575]" metricMagnitude="-2" metricUnit="m" metricValue="8.95" metricValueMax="9.4" metricValueMin="8.5" pageId="2" pageNumber="531" unit="mm" value="89.5" valueMax="94.0" valueMin="85.0">85–94 mm</quantity>
in 
<taxonomicName id="25F04D17FFF5ED19FB47FDBDE8FF4636" baseAuthorityName="Kraus" baseAuthorityYear="2008" box="[1231,1429,549,575]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="531" phylum="Chordata" rank="species" species="sermowaiensis">
<emphasis id="D084EA86FFF5ED19FB47FDBDE8FF4636" box="[1231,1429,549,575]" italics="true" pageId="2" pageNumber="531">C. sermowaiensis</emphasis>
</taxonomicName>
; adult mass = 
<quantity id="25089B71FFF5ED19FEA6FDD0ECDE466A" box="[302,436,584,611]" metricMagnitude="-2" metricUnit="kg" metricValue="2.505" metricValueMax="2.58" metricValueMin="2.43" pageId="2" pageNumber="531" unit="g" value="25.05" valueMax="25.8" valueMin="24.3">24.3–25.8 g</quantity>
<emphasis id="D084EA86FFF5ED19FE33FDD2EF69466A" box="[443,515,586,611]" italics="true" pageId="2" pageNumber="531">versus</emphasis>
<quantity id="25089B71FFF5ED19FD83FDD0EFFA466A" box="[523,656,584,611]" metricMagnitude="-2" metricUnit="kg" metricValue="1.305" metricValueMax="1.58" metricValueMin="1.03" pageId="2" pageNumber="531" unit="g" value="13.05" valueMax="15.8" valueMin="10.3">10.3–15.8 g</quantity>
in 
<taxonomicName id="25F04D17FFF5ED19FD3EFDD1EE16466A" baseAuthorityName="Kraus" baseAuthorityYear="2008" box="[694,892,585,611]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="531" phylum="Chordata" rank="species" species="sermowaiensis">
<emphasis id="D084EA86FFF5ED19FD3EFDD1EE16466A" box="[694,892,585,611]" italics="true" pageId="2" pageNumber="531">C. sermowaiensis</emphasis>
</taxonomicName>
); more tuberculate dorsum, limbs, and head (cf. 
<figureCitation id="7ACB2A11FFF5ED19FF1FFDF5EDB0468E" box="[151,218,621,647]" captionStart="FIGURE 1" captionStartId="3.[151,250,1935,1960]" captionTargetBox="[162,1424,456,1901]" captionTargetId="figure@3.[151,1436,445,1916]" captionTargetPageId="3" captionText="FIGURE 1. (A) Top of head, (B) side of head, (C) mid-dorsum, and (D) left leg of holotype of Cyrtodactylus atremus sp. nov., UMMZ 247747, Karkar Island, Madang Province, PNG." figureDoi="http://doi.org/10.5281/zenodo.3542679" httpUri="https://zenodo.org/record/3542679/files/figure.png" pageId="2" pageNumber="531">Fig. 1</figureCitation>
<emphasis id="D084EA86FFF5ED19FF6BFDF6EC41468E" box="[227,299,622,647]" italics="true" pageId="2" pageNumber="531">versus</emphasis>
<figureCitation id="7ACB2A11FFF5ED19FEBCFDF5EC13468E" box="[308,377,621,647]" captionStart="FIGURE 2" captionStartId="4.[151,250,1636,1661]" captionTargetBox="[164,1423,187,1601]" captionTargetId="figure@4.[151,1436,174,1617]" captionTargetPageId="4" captionText="FIGURE 2. (A) Top of head, (B) side of head, (C) mid-dorsum, and (D) left leg of Cyrtodactylus sermowaiensis, BPBM 35618, Waven, Adelbert Mts., Madang Province, PNG." figureDoi="http://doi.org/10.5281/zenodo.3542683" httpUri="https://zenodo.org/record/3542683/files/figure.png" pageId="2" pageNumber="531">Fig. 2</figureCitation>
), with tubercles 3–5 times size of adjacent granules (
<emphasis id="D084EA86FFF5ED19FC54FDF6E94E468E" box="[988,1060,622,647]" italics="true" pageId="2" pageNumber="531">versus</emphasis>
2–3 times in 
<taxonomicName id="25F04D17FFF5ED19FB4CFDF5E8E1468E" baseAuthorityName="Kraus" baseAuthorityYear="2008" box="[1220,1419,621,647]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="531" phylum="Chordata" rank="species" species="sermowaiensis">
<emphasis id="D084EA86FFF5ED19FB4CFDF5E8E1468E" box="[1220,1419,621,647]" italics="true" pageId="2" pageNumber="531">C. sermowaiensis</emphasis>
</taxonomicName>
); greater average number of rows of dorsal tubercles between the lateral skin folds (mean = 22.6, range = 22–24, SD = 0.799 
<emphasis id="D084EA86FFF5ED19FF54FD2EEC4E46C6" box="[220,292,694,719]" italics="true" pageId="2" pageNumber="531">versus</emphasis>
mean = 20.3, range = 18–25, SD = 
<quantity id="25089B71FFF5ED19FD21FD2DEE6E46C6" box="[681,772,693,719]" metricMagnitude="-2" metricUnit="m" metricValue="2.14122" pageId="2" pageNumber="531" unit="in" value="0.843">0.843 in</quantity>
<taxonomicName id="25F04D17FFF5ED19FC81FD2DEEA746C6" baseAuthorityName="Kraus" baseAuthorityYear="2008" box="[777,973,693,719]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="531" phylum="Chordata" rank="species" species="sermowaiensis">
<emphasis id="D084EA86FFF5ED19FC81FD2DEEA746C6" box="[777,973,693,719]" italics="true" pageId="2" pageNumber="531">C. sermowaiensis</emphasis>
</taxonomicName>
); fewer dark dorsal crossbands from head to tail base (mean = 5.8, range = 5–6, SD = 0.167 
<emphasis id="D084EA86FFF5ED19FD36FD42EE6C46FA" box="[702,774,730,755]" italics="true" pageId="2" pageNumber="531">versus</emphasis>
mean = 8.0, range = 7–9, SD = 
<quantity id="25089B71FFF5ED19FBEDFD40E9AA46FA" box="[1125,1216,728,755]" metricMagnitude="-3" metricUnit="m" metricValue="9.271" pageId="2" pageNumber="531" unit="in" value="0.365">0.365 in</quantity>
<taxonomicName id="25F04D17FFF5ED19FB4EFD41E8E146FA" baseAuthorityName="Kraus" baseAuthorityYear="2008" box="[1222,1419,729,755]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="531" phylum="Chordata" rank="species" species="sermowaiensis">
<emphasis id="D084EA86FFF5ED19FB4EFD41E8E146FA" box="[1222,1419,729,755]" italics="true" pageId="2" pageNumber="531">C. sermowaiensis</emphasis>
</taxonomicName>
); top of head uniformly brown (cf. 
<figureCitation id="7ACB2A11FFF5ED19FD9CFD65EF06471E" box="[532,620,765,791]" captionStart="FIGURE 1" captionStartId="3.[151,250,1935,1960]" captionTargetBox="[162,1424,456,1901]" captionTargetId="figure@3.[151,1436,445,1916]" captionTargetPageId="3" captionText="FIGURE 1. (A) Top of head, (B) side of head, (C) mid-dorsum, and (D) left leg of holotype of Cyrtodactylus atremus sp. nov., UMMZ 247747, Karkar Island, Madang Province, PNG." figureDoi="http://doi.org/10.5281/zenodo.3542679" httpUri="https://zenodo.org/record/3542679/files/figure.png" pageId="2" pageNumber="531">Fig. 1A</figureCitation>
, 
<figureCitation id="7ACB2A11FFF5ED19FDF1FD65EFF3471E" box="[633,665,765,791]" captionStart="FIGURE 3" captionStartId="6.[151,250,1184,1209]" captionTargetBox="[156,1431,307,1157]" captionTargetId="figure@6.[151,1436,301,1164]" captionTargetPageId="6" captionText="FIGURE 3. Portraits in life of (A) paratype of Cyrtodactylus atremus sp. nov., UMMZ 247746, Karkar Island, Madang Province, PNG; (B) Cyrtodactylus sermowaiensis, BPBM 23317, Parkop, Torricelli Mts., West Sepik Province, PNG; (C) Cyrtodactylus sermowaiensis, BPBM 23320, base of Mt. Sapau, Torricelli Mts., West Sepik Province, PNG; and (D) Cyrtodactylus sermowaiensis, BPBM 35598, Yefehi, Prince Alexander Mts., East Sepik Province, PNG." figureDoi="http://doi.org/10.5281/zenodo.3542685" httpUri="https://zenodo.org/record/3542685/files/figure.png" pageId="2" pageNumber="531">3A</figureCitation>
<emphasis id="D084EA86FFF5ED19FD29FD66EF83471E" box="[673,745,766,791]" italics="true" pageId="2" pageNumber="531">versus</emphasis>
heavily patterned with dark-brown blotches, cf. 
<figureCitation id="7ACB2A11FFF5ED19FA87FD65E802471E" box="[1295,1384,765,791]" captionStart="FIGURE 2" captionStartId="4.[151,250,1636,1661]" captionTargetBox="[164,1423,187,1601]" captionTargetId="figure@4.[151,1436,174,1617]" captionTargetPageId="4" captionText="FIGURE 2. (A) Top of head, (B) side of head, (C) mid-dorsum, and (D) left leg of Cyrtodactylus sermowaiensis, BPBM 35618, Waven, Adelbert Mts., Madang Province, PNG." figureDoi="http://doi.org/10.5281/zenodo.3542683" httpUri="https://zenodo.org/record/3542683/files/figure.png" pageId="2" pageNumber="531">Fig. 2A</figureCitation>
, 
<figureCitation id="7ACB2A11FFF5ED19FAFDFD65EDA64732" captionStart="FIGURE 3" captionStartId="6.[151,250,1184,1209]" captionTargetBox="[156,1431,307,1157]" captionTargetId="figure@6.[151,1436,301,1164]" captionTargetPageId="6" captionText="FIGURE 3. Portraits in life of (A) paratype of Cyrtodactylus atremus sp. nov., UMMZ 247746, Karkar Island, Madang Province, PNG; (B) Cyrtodactylus sermowaiensis, BPBM 23317, Parkop, Torricelli Mts., West Sepik Province, PNG; (C) Cyrtodactylus sermowaiensis, BPBM 23320, base of Mt. Sapau, Torricelli Mts., West Sepik Province, PNG; and (D) Cyrtodactylus sermowaiensis, BPBM 35598, Yefehi, Prince Alexander Mts., East Sepik Province, PNG." figureDoi="http://doi.org/10.5281/zenodo.3542685" httpUri="https://zenodo.org/record/3542685/files/figure.png" pageId="2" pageNumber="531">3B, C, D</figureCitation>
in 
<taxonomicName id="25F04D17FFF5ED19FF78FCB9ECDF4732" baseAuthorityName="Kraus" baseAuthorityYear="2008" box="[240,437,801,827]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="531" phylum="Chordata" rank="species" species="sermowaiensis">
<emphasis id="D084EA86FFF5ED19FF78FCB9ECDF4732" box="[240,437,801,827]" italics="true" pageId="2" pageNumber="531">C. sermowaiensis</emphasis>
</taxonomicName>
); and lips dirty white with some brown stippling (cf. 
<figureCitation id="7ACB2A11FFF5ED19FB84FCB9E90B4732" box="[1036,1121,801,827]" captionStart="FIGURE 1" captionStartId="3.[151,250,1935,1960]" captionTargetBox="[162,1424,456,1901]" captionTargetId="figure@3.[151,1436,445,1916]" captionTargetPageId="3" captionText="FIGURE 1. (A) Top of head, (B) side of head, (C) mid-dorsum, and (D) left leg of holotype of Cyrtodactylus atremus sp. nov., UMMZ 247747, Karkar Island, Madang Province, PNG." figureDoi="http://doi.org/10.5281/zenodo.3542679" httpUri="https://zenodo.org/record/3542679/files/figure.png" pageId="2" pageNumber="531">Fig. 1B</figureCitation>
, 
<figureCitation id="7ACB2A11FFF5ED19FBE5FCB9E9E44732" box="[1133,1166,801,827]" captionStart="FIGURE 3" captionStartId="6.[151,250,1184,1209]" captionTargetBox="[156,1431,307,1157]" captionTargetId="figure@6.[151,1436,301,1164]" captionTargetPageId="6" captionText="FIGURE 3. Portraits in life of (A) paratype of Cyrtodactylus atremus sp. nov., UMMZ 247746, Karkar Island, Madang Province, PNG; (B) Cyrtodactylus sermowaiensis, BPBM 23317, Parkop, Torricelli Mts., West Sepik Province, PNG; (C) Cyrtodactylus sermowaiensis, BPBM 23320, base of Mt. Sapau, Torricelli Mts., West Sepik Province, PNG; and (D) Cyrtodactylus sermowaiensis, BPBM 35598, Yefehi, Prince Alexander Mts., East Sepik Province, PNG." figureDoi="http://doi.org/10.5281/zenodo.3542685" httpUri="https://zenodo.org/record/3542685/files/figure.png" pageId="2" pageNumber="531">3A</figureCitation>
<emphasis id="D084EA86FFF5ED19FB1DFCBAE9B74732" box="[1173,1245,802,827]" italics="true" pageId="2" pageNumber="531">versus</emphasis>
dark brown with few white or yellow spots, cf. 
<figureCitation id="7ACB2A11FFF5ED19FE62FCDDEF2B4756" box="[490,577,837,863]" captionStart="FIGURE 2" captionStartId="4.[151,250,1636,1661]" captionTargetBox="[164,1423,187,1601]" captionTargetId="figure@4.[151,1436,174,1617]" captionTargetPageId="4" captionText="FIGURE 2. (A) Top of head, (B) side of head, (C) mid-dorsum, and (D) left leg of Cyrtodactylus sermowaiensis, BPBM 35618, Waven, Adelbert Mts., Madang Province, PNG." figureDoi="http://doi.org/10.5281/zenodo.3542683" httpUri="https://zenodo.org/record/3542683/files/figure.png" pageId="2" pageNumber="531">Fig. 2B</figureCitation>
, 
<figureCitation id="7ACB2A11FFF5ED19FDC4FCDDEFDA4756" box="[588,688,837,863]" captionStart="FIGURE 3" captionStartId="6.[151,250,1184,1209]" captionTargetBox="[156,1431,307,1157]" captionTargetId="figure@6.[151,1436,301,1164]" captionTargetPageId="6" captionText="FIGURE 3. Portraits in life of (A) paratype of Cyrtodactylus atremus sp. nov., UMMZ 247746, Karkar Island, Madang Province, PNG; (B) Cyrtodactylus sermowaiensis, BPBM 23317, Parkop, Torricelli Mts., West Sepik Province, PNG; (C) Cyrtodactylus sermowaiensis, BPBM 23320, base of Mt. Sapau, Torricelli Mts., West Sepik Province, PNG; and (D) Cyrtodactylus sermowaiensis, BPBM 35598, Yefehi, Prince Alexander Mts., East Sepik Province, PNG." figureDoi="http://doi.org/10.5281/zenodo.3542685" httpUri="https://zenodo.org/record/3542685/files/figure.png" pageId="2" pageNumber="531">3B, C, D</figureCitation>
in 
<taxonomicName id="25F04D17FFF5ED19FD5CFCDDEEF34756" baseAuthorityName="Kraus" baseAuthorityYear="2008" box="[724,921,837,863]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="531" phylum="Chordata" rank="species" species="sermowaiensis">
<emphasis id="D084EA86FFF5ED19FD5CFCDDEEF34756" box="[724,921,837,863]" italics="true" pageId="2" pageNumber="531">C. sermowaiensis</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="AAEA651FFFF5ED1EFF4FFCF1EE274376" lastPageId="5" lastPageNumber="534" pageId="2" pageNumber="531" type="description">
<paragraph id="E24F3694FFF5ED19FF4FFCF1ECB941CA" blockId="2.[151,1437,153,2015]" pageId="2" pageNumber="531">
<emphasis id="D084EA86FFF5ED19FF4FFCF1ECB6478A" bold="true" box="[199,476,872,899]" pageId="2" pageNumber="531">
<emphasis id="D084EA86FFF5ED19FF4FFCF1ECBF478A" bold="true" box="[199,469,872,899]" italics="true" pageId="2" pageNumber="531">
Description of 
<typeStatus id="3D4B8836FFF5ED19FEFBFCF0ECBF478A" box="[371,469,872,899]" pageId="2" pageNumber="531" type="holotype">holotype</typeStatus>
</emphasis>
.
</emphasis>
A mature male, with right-lateral incision. Animal large (SVL = 
<quantity id="25089B71FFF5ED19FB32FCF1E87D478A" box="[1210,1303,873,899]" metricMagnitude="-1" metricUnit="m" metricValue="1.1099999999999999" pageId="2" pageNumber="531" unit="mm" value="111.0">111 mm</quantity>
, TrL = 
<quantity id="25089B71FFF5ED19FAE3FCF1EDAD47AE" metricMagnitude="-2" metricUnit="m" metricValue="4.99" pageId="2" pageNumber="531" unit="mm" value="49.9">49.9 mm</quantity>
); preserved with tail folded back on itself and unamenable to accurate measurement in preservative, but 
<quantity id="25089B71FFF5ED19FACAFC15E8F747AE" box="[1346,1437,909,935]" metricMagnitude="-1" metricUnit="m" metricValue="1.13" pageId="2" pageNumber="531" unit="mm" value="113.0">113 mm</quantity>
in life. Head relatively long (HL/SVL = 0.28) and wide (HW/HL = 0.66), not depressed (HH/HL = 0.43), distinct from neck (
<figureCitation id="7ACB2A11FFF5ED19FE91FC4DEC1847E6" box="[281,370,981,1007]" captionStart="FIGURE 1" captionStartId="3.[151,250,1935,1960]" captionTargetBox="[162,1424,456,1901]" captionTargetId="figure@3.[151,1436,445,1916]" captionTargetPageId="3" captionText="FIGURE 1. (A) Top of head, (B) side of head, (C) mid-dorsum, and (D) left leg of holotype of Cyrtodactylus atremus sp. nov., UMMZ 247747, Karkar Island, Madang Province, PNG." figureDoi="http://doi.org/10.5281/zenodo.3542679" httpUri="https://zenodo.org/record/3542679/files/figure.png" pageId="2" pageNumber="531">Fig. 1A</figureCitation>
). Loreal region slightly inflated; canthus rostralis poorly defined. Interorbital region and top of snout concave. Snout relatively long (SN/HL = 0.40), much longer than eye diameter (SN/EY = 1.7). Eye relatively large (EY/HL = 0.23); pupil vertical, with weakly crenellated margins; supraciliaries large, frill-like, well-differentiated from adjacent granules. Ear opening small (Ear/HL = 0.058), taller than wide (
<figureCitation id="7ACB2A11FFF5ED19FBCCFBD9E9F14052" box="[1092,1179,1089,1115]" captionStart="FIGURE 1" captionStartId="3.[151,250,1935,1960]" captionTargetBox="[162,1424,456,1901]" captionTargetId="figure@3.[151,1436,445,1916]" captionTargetPageId="3" captionText="FIGURE 1. (A) Top of head, (B) side of head, (C) mid-dorsum, and (D) left leg of holotype of Cyrtodactylus atremus sp. nov., UMMZ 247747, Karkar Island, Madang Province, PNG." figureDoi="http://doi.org/10.5281/zenodo.3542679" httpUri="https://zenodo.org/record/3542679/files/figure.png" pageId="2" pageNumber="531">Fig. 1B</figureCitation>
); distance between ear and eye larger than eye diameter (EE/EY = 1.3). Rostral wider (
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) than high (
<quantity id="25089B71FFF5ED19FBB6FBFCE9F34076" box="[1086,1177,1124,1151]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="2" pageNumber="531" unit="mm" value="2.5">2.5 mm</quantity>
), highest just medial to nares, lower between these points, length 
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; dorsal surface with weak (
<quantity id="25089B71FFF5ED19FB8FFB11E90B40AA" box="[1031,1121,1161,1187]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="531" unit="mm" value="0.2">0.2 mm</quantity>
) medial groove at posterior end. Two enlarged supranasals separated by single internasal; internasal bordered anteriorly by supernumerary scale otherwise surrounded by posterior lobes of rostral. Rostral in contact with first supralabials, two supranasals, one internasal, and one postnasal (on right). External nares circular; each bordered by rostral, supranasal, first supralabial, and four postnasals. Three rows of small scales separate orbit from supralabials at narrowest point. Mental shield shaped, posterior margins shallowly concave, wider (4.0 mm) than long (
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). Mental bordered posteriorly by two enlarged postmentals. Infralabials bordered by slightly enlarged, elongate scales that gradually decrease in size posteriorly. Supralabials to mid-orbital position nine on right side, 11 on left; to angle of jaw 12 on right, 14 on left. Infralabials 12 on each side.
</paragraph>
<paragraph id="E24F3694FFF5ED19FF4FFA55E9C442B6" blockId="2.[151,1437,153,2015]" pageId="2" pageNumber="531">
Body moderately robust (TrL/SVL = 0.45), with lateral skin folds consisting of scales approximately same size as adjacent ventral scales and dorsal granules on its ventral and dorsal surfaces, respectively, and with larger scales along its lateralmost margin. Dorsal scales on head, body, and limbs tiny, juxtaposed granules. Both raised and flattened tubercles intermixed and dense on head, body, and limbs (
<figureCitation id="7ACB2A11FFF5ED19FC5AF9A1E97F425A" box="[978,1045,1593,1619]" captionStart="FIGURE 1" captionStartId="3.[151,250,1935,1960]" captionTargetBox="[162,1424,456,1901]" captionTargetId="figure@3.[151,1436,445,1916]" captionTargetPageId="3" captionText="FIGURE 1. (A) Top of head, (B) side of head, (C) mid-dorsum, and (D) left leg of holotype of Cyrtodactylus atremus sp. nov., UMMZ 247747, Karkar Island, Madang Province, PNG." figureDoi="http://doi.org/10.5281/zenodo.3542679" httpUri="https://zenodo.org/record/3542679/files/figure.png" pageId="2" pageNumber="531">Fig. 1</figureCitation>
); 3–5 times areal size of adjacent granules; distinct from granules in profile; each with small dorsal keel, highest posteriorly. Tubercles arranged in approximately 23 rows between the lateral folds. Ventral scales flat and smooth; those on chin and throat smaller and juxtaposed; those on abdomen larger and subimbricate, arrayed in 41 rows across mid-venter.
</paragraph>
<paragraph id="E24F3694FFF5ED19FF4FF951EFEF4346" blockId="2.[151,1437,153,2015]" pageId="2" pageNumber="531">Precloacal and femoral pores absent. Scales of femoral regions of approximately uniform size, smaller than those on abdomen. A diamond-shaped patch of slightly enlarged precloacal scales (ca. 2–3 times larger than nearby femoral scales) approximately six rows deep mid-ventrally and 17 scales across at its widest. No precloacal groove. Scales on palms and soles rounded, smooth.</paragraph>
<paragraph id="E24F3694FFF5ED18FF4FF8C1EC9744F2" blockId="2.[151,1437,153,2015]" lastBlockId="3.[151,1436,152,431]" lastPageId="3" lastPageNumber="532" pageId="2" pageNumber="531">Fore- and hindlimbs relatively well-developed (FA/SVL = 0.15, CS/SVL = 0.18). Digits well-developed, inflected at basal interphalangeal joints; all with robust, recurved claws surrounded by two scales. Subdigital scales consisting of several rows of basal lamellae, followed distally by ca. 6–8 rows of small granular scales, and terminally by several additional rows of lamellae not as expanded as those basally. Subdigital scales of manus 16 (tip missing)–22–24–26–22 on right, 18–23–25–26–22 on left; of pes 18–22–29–31–26 on right, 17–23–27–31–25 on left. Relative lengths of digits on manus I &lt;V &lt;II &lt;III &lt;IV; on pes I &lt;II &lt;V &lt;III &lt;IV. Slight interdigital webbing on manus, slightly more on pes.</paragraph>
<paragraph id="E24F3694FFF4ED18FF4FFE9DECCE45A6" blockId="3.[151,1436,152,431]" pageId="3" pageNumber="532">Tail regenerated, with rather narrow base (TW/SVL = 0.067), tapering to a blunt point. Cloacal sacs swollen, with small external orifices situated near lateral margins of vent; midventral scales of sac not distinctly larger than those ventrolaterally. Scales of tail small, flat, smooth, subimbricate, slightly larger ventrally than dorsally; with approximately eight rows of enlarged dorsal tubercles just before regenerated tail; four enlarged postcloacal spurs on each side of tailbase.</paragraph>
<caption id="B68F661CFFF4ED18FF1FF817EFBC43C5" ID-DOI="http://doi.org/10.5281/zenodo.3542679" ID-Zenodo-Dep="3542679" httpUri="https://zenodo.org/record/3542679/files/figure.png" pageId="3" pageNumber="532" startId="3.[151,250,1935,1960]" targetBox="[162,1424,456,1901]" targetPageId="3">
<paragraph id="E24F3694FFF4ED18FF1FF817EFBC43C5" blockId="3.[151,1436,1935,1996]" pageId="3" pageNumber="532">
<emphasis id="D084EA86FFF4ED18FF1FF817EC7943A1" bold="true" box="[151,275,1935,1960]" pageId="3" pageNumber="532">FIGURE 1.</emphasis>
(A) Top of head, (B) side of head, (C) mid-dorsum, and (D) left leg of holotype of 
<taxonomicName id="25F04D17FFF4ED18FBD2F808E85543A1" authority="Kraus &amp; Weijola, 2019" authorityName="Kraus &amp; Weijola" authorityYear="2019" box="[1114,1343,1936,1960]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="532" phylum="Chordata" rank="species" species="atremus" status="sp. nov.">
<emphasis id="D084EA86FFF4ED18FBD2F808E85543A1" box="[1114,1343,1936,1960]" italics="true" pageId="3" pageNumber="532">Cyrtodactylus atremus</emphasis>
</taxonomicName>
<emphasis id="D084EA86FFF4ED18FACDF808E8FF43A1" bold="true" box="[1349,1429,1936,1960]" pageId="3" pageNumber="532">
<taxonomicNameLabel id="CBB757FDFFF4ED18FACDF808E8FF43A1" box="[1349,1429,1936,1960]" pageId="3" pageNumber="532" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, UMMZ 247747, Karkar Island, Madang Province, PNG.
</paragraph>
</caption>
<caption id="B68F661CFFF3ED1FFF1FF9FCEF1642A8" ID-DOI="http://doi.org/10.5281/zenodo.3542683" ID-Zenodo-Dep="3542683" httpUri="https://zenodo.org/record/3542683/files/figure.png" pageId="4" pageNumber="533" startId="4.[151,250,1636,1661]" targetBox="[164,1423,187,1601]" targetPageId="4">
<paragraph id="E24F3694FFF3ED1FFF1FF9FCEF1642A8" blockId="4.[151,1436,1636,1697]" pageId="4" pageNumber="533">
<emphasis id="D084EA86FFF3ED1FFF1FF9FCEC784274" bold="true" box="[151,274,1636,1661]" pageId="4" pageNumber="533">FIGURE 2.</emphasis>
(A) Top of head, (B) side of head, (C) mid-dorsum, and (D) left leg of 
<taxonomicName id="25F04D17FFF3ED1FFC5EF9FDE9954274" baseAuthorityName="Kraus" baseAuthorityYear="2008" box="[982,1279,1637,1661]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="533" phylum="Chordata" rank="species" species="sermowaiensis">
<emphasis id="D084EA86FFF3ED1FFC5EF9FDE9954274" box="[982,1279,1637,1661]" italics="true" pageId="4" pageNumber="533">Cyrtodactylus sermowaiensis</emphasis>
</taxonomicName>
, BPBM 35618, Waven, Adelbert Mts., Madang Province, PNG.
</paragraph>
</caption>
<paragraph id="E24F3694FFF3ED1FFF4FF957EC5743EF" blockId="4.[151,1437,1743,2022]" pageId="4" pageNumber="533">
<emphasis id="D084EA86FFF3ED1FFF4FF957ECDF42E0" box="[199,437,1743,1769]" italics="true" pageId="4" pageNumber="533">Color in preservative</emphasis>
: Dorsal ground color brown with six dark-brown bands including nuchal collar and band above hindlimb insertion. Lateral ends of the dark scapular band extend posteriorly as narrow stripes to almost meet the following band; last three bands narrowly divided vertebrally by ground color. Nuchal collar narrowly indented posteromedially (
<figureCitation id="7ACB2A11FFF3ED1FFED6F8A3ECD3435F" box="[350,441,1851,1878]" captionStart="FIGURE 1" captionStartId="3.[151,250,1935,1960]" captionTargetBox="[162,1424,456,1901]" captionTargetId="figure@3.[151,1436,445,1916]" captionTargetPageId="3" captionText="FIGURE 1. (A) Top of head, (B) side of head, (C) mid-dorsum, and (D) left leg of holotype of Cyrtodactylus atremus sp. nov., UMMZ 247747, Karkar Island, Madang Province, PNG." figureDoi="http://doi.org/10.5281/zenodo.3542679" httpUri="https://zenodo.org/record/3542679/files/figure.png" pageId="4" pageNumber="533">Fig. 1A</figureCitation>
). Regenerated tail uniformly brown. Dorsal tubercles dirty white stippled with brown. Head and limbs brown vaguely mottled with dark brown (
<figureCitation id="7ACB2A11FFF3ED1FFC95F8C7EEF24373" box="[797,920,1887,1914]" captionStart="FIGURE 1" captionStartId="3.[151,250,1935,1960]" captionTargetBox="[162,1424,456,1901]" captionTargetId="figure@3.[151,1436,445,1916]" captionTargetPageId="3" captionText="FIGURE 1. (A) Top of head, (B) side of head, (C) mid-dorsum, and (D) left leg of holotype of Cyrtodactylus atremus sp. nov., UMMZ 247747, Karkar Island, Madang Province, PNG." figureDoi="http://doi.org/10.5281/zenodo.3542679" httpUri="https://zenodo.org/record/3542679/files/figure.png" pageId="4" pageNumber="533">Fig. 1A, D</figureCitation>
); labials white stippled with brown, paler than head (
<figureCitation id="7ACB2A11FFF3ED1FFF54F81BEC594397" box="[220,307,1923,1950]" captionStart="FIGURE 1" captionStartId="3.[151,250,1935,1960]" captionTargetBox="[162,1424,456,1901]" captionTargetId="figure@3.[151,1436,445,1916]" captionTargetPageId="3" captionText="FIGURE 1. (A) Top of head, (B) side of head, (C) mid-dorsum, and (D) left leg of holotype of Cyrtodactylus atremus sp. nov., UMMZ 247747, Karkar Island, Madang Province, PNG." figureDoi="http://doi.org/10.5281/zenodo.3542679" httpUri="https://zenodo.org/record/3542679/files/figure.png" pageId="4" pageNumber="533">Fig. 1B</figureCitation>
). Venter pale brown to naked eye; each scale dirty white punctated with brown under magnification. Palmar and plantar surfaces pale brown but basal lamellae dirty white. Iris copper, heavily veined with brown; pupil gray (
<figureCitation id="7ACB2A11FFF3ED1FFF50F853EC4543EF" box="[216,303,1995,2022]" captionStart="FIGURE 1" captionStartId="3.[151,250,1935,1960]" captionTargetBox="[162,1424,456,1901]" captionTargetId="figure@3.[151,1436,445,1916]" captionTargetPageId="3" captionText="FIGURE 1. (A) Top of head, (B) side of head, (C) mid-dorsum, and (D) left leg of holotype of Cyrtodactylus atremus sp. nov., UMMZ 247747, Karkar Island, Madang Province, PNG." figureDoi="http://doi.org/10.5281/zenodo.3542679" httpUri="https://zenodo.org/record/3542679/files/figure.png" pageId="4" pageNumber="533">Fig. 1B</figureCitation>
).
</paragraph>
<paragraph id="E24F3694FFF2ED1EFF4FFF02E9C444DE" blockId="5.[151,1437,153,467]" pageId="5" pageNumber="534">
<emphasis id="D084EA86FFF2ED1EFF4FFF02ECA244BA" box="[199,456,153,179]" italics="true" pageId="5" pageNumber="534">Measurements (in mm)</emphasis>
. SVL = 111, TrL = 49.9, TW = 7.4, FA = 16.2, CS = 19.6, HL = 31.0, HW = 20.6, HH = 13.3, Ear = 1.8, EE = 9.6, EY = 7.2, SN = 12.3, EN = 10.4, IN = 4.4, IO = 8.3, mass = 
<quantity id="25089B71FFF2ED1EFBE9FF25E9C044DE" box="[1121,1194,189,215]" metricMagnitude="-2" metricUnit="kg" metricValue="2.43" pageId="5" pageNumber="534" unit="g" value="24.3">24.3 g</quantity>
.
</paragraph>
<paragraph id="E24F3694FFF2ED1EFF4FFF79EFF745DA" blockId="5.[151,1437,153,467]" pageId="5" pageNumber="534">
<emphasis id="D084EA86FFF2ED1EFF4FFF79EC5344F2" bold="true" box="[199,313,225,251]" italics="true" pageId="5" pageNumber="534">Variation.</emphasis>
There appear to be no meaningful morphometric differences between the sexes, though there are suggestions in the small samples available that juveniles may have relatively larger eyes (EY/SVL, EY/EN) and narrower heads (IN/SVL, IO/SVL, HW/HL, EN/IN) (
<tableCitation id="AF72032FFFF2ED1EFD74FEB1EE38454A" box="[764,850,297,323]" captionStart="TABLE 1" captionStartId="5.[151,239,514,538]" captionTargetPageId="5" captionText="TABLE 1. Mensural and meristic data for the type series of Cyrtodactylus atremus sp. nov." httpUri="http://table.plazi.org/id/B68F661CFFF2ED1EFF1FFD9AE9024612" pageId="5" pageNumber="534" tableUuid="B68F661CFFF2ED1EFF1FFD9AE9024612">Table 1</tableCitation>
). The dark crossbands often have a few dark spots along their anterior or posterior margins, and some of them (usually the posterior ones) may be narrowly divided by ground color vertebrally. In UMMZ 247749, one band is split into anterior and posterior portions on the right side, and one band in UMMZ 247748 is also offset anteriorly on the right side. UMMZ 247748 also has more ventrolateral brown stippling than the other specimens.
</paragraph>
<caption id="B68F661CFFF2ED1EFF1FFD9AE9024612" ID-Table-UUID="B68F661CFFF2ED1EFF1FFD9AE9024612" box="[151,1128,514,539]" httpUri="http://table.plazi.org/id/B68F661CFFF2ED1EFF1FFD9AE9024612" pageId="5" pageNumber="534" targetBox="[166,1373,550,1664]" targetIsTable="true" targetPageId="5">
<paragraph id="E24F3694FFF2ED1EFF1FFD9AE9024612" blockId="5.[151,1128,514,539]" box="[151,1128,514,539]" pageId="5" pageNumber="534">
<emphasis id="D084EA86FFF2ED1EFF1FFD9AEC604612" bold="true" box="[151,266,514,539]" pageId="5" pageNumber="534">TABLE 1.</emphasis>
Mensural and meristic data for the type series of 
<taxonomicName id="25F04D17FFF2ED1EFC93FD9BE9664613" authority="Kraus &amp; Weijola, 2019" authorityName="Kraus &amp; Weijola" authorityYear="2019" box="[795,1036,515,539]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="534" phylum="Chordata" rank="species" species="atremus" status="sp. nov.">
<emphasis id="D084EA86FFF2ED1EFC93FD9BE9664613" box="[795,1036,515,539]" italics="true" pageId="5" pageNumber="534">Cyrtodactylus atremus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CBB757FDFFF2ED1EFB9BFD9BE9024612" box="[1043,1128,515,539]" pageId="5" pageNumber="534" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="E24F3694FFF2ED1EFF2EFDBEE9814289" pageId="5" pageNumber="534">
<table id="90F0C434FFF212E4FF2EFDBEE8374289" box="[166,1373,550,1664]" gridcols="7" gridrows="30" pageId="5" pageNumber="534">
<tr id="5CC034D6FFF212E4FF2EFDBEE8374636" box="[166,1373,550,575]" gridrow="0" pageId="5" pageNumber="534" rowspan-2="1" rowspan-4="1">
<th id="1F115DAAFFF212E4FF2EFDBEEC194636" box="[166,371,550,575]" gridcol="0" gridrow="0" pageId="5" pageNumber="534">Character</th>
<th id="1F115DAAFFF212E4FE53FDBEEF4B4636" box="[475,545,550,575]" gridcol="1" gridrow="0" pageId="5" pageNumber="534">Males</th>
<th id="1F115DAAFFF212E4FCA8FDBEEE1E4636" box="[800,884,550,575]" gridcol="3" gridrow="0" pageId="5" pageNumber="534">Females</th>
<th id="1F115DAAFFF212E4FBEDFDBEE8374636" box="[1125,1373,550,575]" colspan="2" colspanRight="1" gridcol="5" gridrow="0" pageId="5" pageNumber="534">Immature females</th>
</tr>
<tr id="5CC034D6FFF212E4FF2EFDD2E837466A" box="[166,1373,586,611]" gridrow="1" pageId="5" pageNumber="534" rowspan-0="1" rowspan-2="1" rowspan-4="1" rowspan-6="1">
<td id="1F115DAAFFF212E4FE53FDD2EF4B466A" box="[475,545,586,611]" gridcol="1" gridrow="1" pageId="5" pageNumber="534">(n = 2)</td>
<td id="1F115DAAFFF212E4FCA8FDD2EE1E466A" box="[800,884,586,611]" gridcol="3" gridrow="1" pageId="5" pageNumber="534">(n = 5)</td>
<td id="1F115DAAFFF212E4FBEDFDD2E9C0466A" box="[1125,1194,586,611]" gridcol="5" gridrow="1" pageId="5" pageNumber="534">(n = 2)</td>
</tr>
<tr id="5CC034D6FFF212E4FF2EFDF4E837468D" box="[166,1373,620,644]" gridrow="2" pageId="5" pageNumber="534" rowspan-0="1">
<td id="1F115DAAFFF212E4FE53FDF4EF4B468D" box="[475,545,620,644]" gridcol="1" gridrow="2" pageId="5" pageNumber="534">Mean</td>
<td id="1F115DAAFFF212E4FDDDFDF4EFBE468D" box="[597,724,620,644]" gridcol="2" gridrow="2" pageId="5" pageNumber="534">Range</td>
<td id="1F115DAAFFF212E4FCA8FDF4EE1E468D" box="[800,884,620,644]" gridcol="3" gridrow="2" pageId="5" pageNumber="534">Mean</td>
<td id="1F115DAAFFF212E4FC12FDF4E973468D" box="[922,1049,620,644]" gridcol="4" gridrow="2" pageId="5" pageNumber="534">Range</td>
<td id="1F115DAAFFF212E4FBEDFDF4E9C0468D" box="[1125,1194,620,644]" gridcol="5" gridrow="2" pageId="5" pageNumber="534">Mean</td>
<td id="1F115DAAFFF212E4FB56FDF4E837468D" box="[1246,1373,620,644]" gridcol="6" gridrow="2" pageId="5" pageNumber="534">Range</td>
</tr>
<tr id="5CC034D6FFF212E4FF2EFD09E83746A3" box="[166,1373,657,682]" gridrow="3" pageId="5" pageNumber="534">
<th id="1F115DAAFFF212E4FF2EFD09EC1946A3" box="[166,371,657,682]" gridcol="0" gridrow="3" pageId="5" pageNumber="534">SVL (mm)</th>
<td id="1F115DAAFFF212E4FE53FD09EF4B46A3" box="[475,545,657,682]" gridcol="1" gridrow="3" pageId="5" pageNumber="534">114.5</td>
<td id="1F115DAAFFF212E4FDDDFD09EFBE46A3" box="[597,724,657,682]" gridcol="2" gridrow="3" pageId="5" pageNumber="534">111.0–118.0</td>
<td id="1F115DAAFFF212E4FCA8FD09EE1E46A3" box="[800,884,657,682]" gridcol="3" gridrow="3" pageId="5" pageNumber="534">110.3</td>
<td id="1F115DAAFFF212E4FC12FD09E97346A3" box="[922,1049,657,682]" gridcol="4" gridrow="3" pageId="5" pageNumber="534">108–112.5</td>
<td id="1F115DAAFFF212E4FBEDFD09E9C046A3" box="[1125,1194,657,682]" gridcol="5" gridrow="3" pageId="5" pageNumber="534">69.5</td>
<td id="1F115DAAFFF212E4FB56FD09E83746A3" box="[1246,1373,657,682]" gridcol="6" gridrow="3" pageId="5" pageNumber="534">66–73</td>
</tr>
<tr id="5CC034D6FFF212E4FF2EFD2FE83746D9" box="[166,1373,695,720]" gridrow="4" pageId="5" pageNumber="534">
<th id="1F115DAAFFF212E4FF2EFD2FEC1946D9" box="[166,371,695,720]" gridcol="0" gridrow="4" pageId="5" pageNumber="534">TrL/SVL</th>
<td id="1F115DAAFFF212E4FE53FD2FEF4B46D9" box="[475,545,695,720]" gridcol="1" gridrow="4" pageId="5" pageNumber="534">0.45</td>
<td id="1F115DAAFFF212E4FDDDFD2FEFBE46D9" box="[597,724,695,720]" gridcol="2" gridrow="4" pageId="5" pageNumber="534">0.45</td>
<td id="1F115DAAFFF212E4FCA8FD2FEE1E46D9" box="[800,884,695,720]" gridcol="3" gridrow="4" pageId="5" pageNumber="534">0.48</td>
<td id="1F115DAAFFF212E4FC12FD2FE97346D9" box="[922,1049,695,720]" gridcol="4" gridrow="4" pageId="5" pageNumber="534">0.46–0.52</td>
<td id="1F115DAAFFF212E4FBEDFD2FE9C046D9" box="[1125,1194,695,720]" gridcol="5" gridrow="4" pageId="5" pageNumber="534">0.44</td>
<td id="1F115DAAFFF212E4FB56FD2FE83746D9" box="[1246,1373,695,720]" gridcol="6" gridrow="4" pageId="5" pageNumber="534">0.43–0.44</td>
</tr>
<tr id="5CC034D6FFF212E4FF2EFD45E83746FC" box="[166,1373,733,757]" gridrow="5" pageId="5" pageNumber="534">
<th id="1F115DAAFFF212E4FF2EFD45EC1946FC" box="[166,371,733,757]" gridcol="0" gridrow="5" pageId="5" pageNumber="534">CS/SVL</th>
<td id="1F115DAAFFF212E4FE53FD45EF4B46FC" box="[475,545,733,757]" gridcol="1" gridrow="5" pageId="5" pageNumber="534">0.17</td>
<td id="1F115DAAFFF212E4FDDDFD45EFBE46FC" box="[597,724,733,757]" gridcol="2" gridrow="5" pageId="5" pageNumber="534">0.17–0.18</td>
<td id="1F115DAAFFF212E4FCA8FD45EE1E46FC" box="[800,884,733,757]" gridcol="3" gridrow="5" pageId="5" pageNumber="534">0.18</td>
<td id="1F115DAAFFF212E4FC12FD45E97346FC" box="[922,1049,733,757]" gridcol="4" gridrow="5" pageId="5" pageNumber="534">0.18–0.20</td>
<td id="1F115DAAFFF212E4FBEDFD45E9C046FC" box="[1125,1194,733,757]" gridcol="5" gridrow="5" pageId="5" pageNumber="534">0.18</td>
<td id="1F115DAAFFF212E4FB56FD45E83746FC" box="[1246,1373,733,757]" gridcol="6" gridrow="5" pageId="5" pageNumber="534">0.18–0.19</td>
</tr>
<tr id="5CC034D6FFF212E4FF2EFC9AE8374712" box="[166,1373,770,795]" gridrow="6" pageId="5" pageNumber="534">
<th id="1F115DAAFFF212E4FF2EFC9AEC194712" box="[166,371,770,795]" gridcol="0" gridrow="6" pageId="5" pageNumber="534">TL/SVL</th>
<td id="1F115DAAFFF212E4FE53FC9AEF4B4712" box="[475,545,770,795]" gridcol="1" gridrow="6" pageId="5" pageNumber="534">0.66</td>
<td id="1F115DAAFFF212E4FDDDFC9AEFBE4712" box="[597,724,770,795]" gridcol="2" gridrow="6" pageId="5" pageNumber="534">0.66</td>
<td id="1F115DAAFFF212E4FCA8FC9AEE1E4712" box="[800,884,770,795]" gridcol="3" gridrow="6" pageId="5" pageNumber="534">0.84</td>
<td id="1F115DAAFFF212E4FC12FC9AE9734712" box="[922,1049,770,795]" gridcol="4" gridrow="6" pageId="5" pageNumber="534">0.69–0.98</td>
<td id="1F115DAAFFF212E4FBEDFC9AE9C04712" box="[1125,1194,770,795]" gridcol="5" gridrow="6" pageId="5" pageNumber="534">1.23</td>
<td id="1F115DAAFFF212E4FB56FC9AE8374712" box="[1246,1373,770,795]" gridcol="6" gridrow="6" pageId="5" pageNumber="534">1.23</td>
</tr>
<tr id="5CC034D6FFF212E4FF2EFCB0E8374748" box="[166,1373,808,833]" gridrow="7" pageId="5" pageNumber="534">
<th id="1F115DAAFFF212E4FF2EFCB0EC194748" box="[166,371,808,833]" gridcol="0" gridrow="7" pageId="5" pageNumber="534">TW/SVL</th>
<td id="1F115DAAFFF212E4FE53FCB0EF4B4748" box="[475,545,808,833]" gridcol="1" gridrow="7" pageId="5" pageNumber="534">0.066</td>
<td id="1F115DAAFFF212E4FDDDFCB0EFBE4748" box="[597,724,808,833]" gridcol="2" gridrow="7" pageId="5" pageNumber="534">0.064–0.067</td>
<td id="1F115DAAFFF212E4FCA8FCB0EE1E4748" box="[800,884,808,833]" gridcol="3" gridrow="7" pageId="5" pageNumber="534">0.059</td>
<td id="1F115DAAFFF212E4FC12FCB0E9734748" box="[922,1049,808,833]" gridcol="4" gridrow="7" pageId="5" pageNumber="534">0.050–0.068</td>
<td id="1F115DAAFFF212E4FBEDFCB0E9C04748" box="[1125,1194,808,833]" gridcol="5" gridrow="7" pageId="5" pageNumber="534">0.051</td>
<td id="1F115DAAFFF212E4FB56FCB0E8374748" box="[1246,1373,808,833]" gridcol="6" gridrow="7" pageId="5" pageNumber="534">0.050–0.052</td>
</tr>
<tr id="5CC034D6FFF212E4FF2EFCD7E837476E" box="[166,1373,847,871]" gridrow="8" pageId="5" pageNumber="534">
<th id="1F115DAAFFF212E4FF2EFCD7EC19476E" box="[166,371,847,871]" gridcol="0" gridrow="8" pageId="5" pageNumber="534">HL/SVL</th>
<td id="1F115DAAFFF212E4FE53FCD7EF4B476E" box="[475,545,847,871]" gridcol="1" gridrow="8" pageId="5" pageNumber="534">0.27</td>
<td id="1F115DAAFFF212E4FDDDFCD7EFBE476E" box="[597,724,847,871]" gridcol="2" gridrow="8" pageId="5" pageNumber="534">0.27–0.28</td>
<td id="1F115DAAFFF212E4FCA8FCD7EE1E476E" box="[800,884,847,871]" gridcol="3" gridrow="8" pageId="5" pageNumber="534">0.29</td>
<td id="1F115DAAFFF212E4FC12FCD7E973476E" box="[922,1049,847,871]" gridcol="4" gridrow="8" pageId="5" pageNumber="534">0.27–0.30</td>
<td id="1F115DAAFFF212E4FBEDFCD7E9C0476E" box="[1125,1194,847,871]" gridcol="5" gridrow="8" pageId="5" pageNumber="534">0.30</td>
<td id="1F115DAAFFF212E4FB56FCD7E837476E" box="[1246,1373,847,871]" gridcol="6" gridrow="8" pageId="5" pageNumber="534">0.29–0.31</td>
</tr>
<tr id="5CC034D6FFF212E4FF2EFCECE8374785" box="[166,1373,884,908]" gridrow="9" pageId="5" pageNumber="534">
<th id="1F115DAAFFF212E4FF2EFCECEC194785" box="[166,371,884,908]" gridcol="0" gridrow="9" pageId="5" pageNumber="534">HW/SVL</th>
<td id="1F115DAAFFF212E4FE53FCECEF4B4785" box="[475,545,884,908]" gridcol="1" gridrow="9" pageId="5" pageNumber="534">0.18</td>
<td id="1F115DAAFFF212E4FDDDFCECEFBE4785" box="[597,724,884,908]" gridcol="2" gridrow="9" pageId="5" pageNumber="534">0.18–0.19</td>
<td id="1F115DAAFFF212E4FCA8FCECEE1E4785" box="[800,884,884,908]" gridcol="3" gridrow="9" pageId="5" pageNumber="534">0.19</td>
<td id="1F115DAAFFF212E4FC12FCECE9734785" box="[922,1049,884,908]" gridcol="4" gridrow="9" pageId="5" pageNumber="534">0.19–0.20</td>
<td id="1F115DAAFFF212E4FBEDFCECE9C04785" box="[1125,1194,884,908]" gridcol="5" gridrow="9" pageId="5" pageNumber="534">0.19</td>
<td id="1F115DAAFFF212E4FB56FCECE8374785" box="[1246,1373,884,908]" gridcol="6" gridrow="9" pageId="5" pageNumber="534">0.19</td>
</tr>
<tr id="5CC034D6FFF212E4FF2EFC02E83747BB" box="[166,1373,922,946]" gridrow="10" pageId="5" pageNumber="534">
<th id="1F115DAAFFF212E4FF2EFC02EC1947BB" box="[166,371,922,946]" gridcol="0" gridrow="10" pageId="5" pageNumber="534">HH/SVL</th>
<td id="1F115DAAFFF212E4FE53FC02EF4B47BB" box="[475,545,922,946]" gridcol="1" gridrow="10" pageId="5" pageNumber="534">0.11</td>
<td id="1F115DAAFFF212E4FDDDFC02EFBE47BB" box="[597,724,922,946]" gridcol="2" gridrow="10" pageId="5" pageNumber="534">0.11–0.12</td>
<td id="1F115DAAFFF212E4FCA8FC02EE1E47BB" box="[800,884,922,946]" gridcol="3" gridrow="10" pageId="5" pageNumber="534">0.11</td>
<td id="1F115DAAFFF212E4FC12FC02E97347BB" box="[922,1049,922,946]" gridcol="4" gridrow="10" pageId="5" pageNumber="534">0.11–0.13</td>
<td id="1F115DAAFFF212E4FBEDFC02E9C047BB" box="[1125,1194,922,946]" gridcol="5" gridrow="10" pageId="5" pageNumber="534">0.12</td>
<td id="1F115DAAFFF212E4FB56FC02E83747BB" box="[1246,1373,922,946]" gridcol="6" gridrow="10" pageId="5" pageNumber="534">0.11–0.13</td>
</tr>
<tr id="5CC034D6FFF212E4FF2EFC27E83747D1" box="[166,1373,959,984]" gridrow="11" pageId="5" pageNumber="534">
<th id="1F115DAAFFF212E4FF2EFC27EC1947D1" box="[166,371,959,984]" gridcol="0" gridrow="11" pageId="5" pageNumber="534">FA/SVL</th>
<td id="1F115DAAFFF212E4FE53FC27EF4B47D1" box="[475,545,959,984]" gridcol="1" gridrow="11" pageId="5" pageNumber="534">0.14</td>
<td id="1F115DAAFFF212E4FDDDFC27EFBE47D1" box="[597,724,959,984]" gridcol="2" gridrow="11" pageId="5" pageNumber="534">0.14–0.15</td>
<td id="1F115DAAFFF212E4FCA8FC27EE1E47D1" box="[800,884,959,984]" gridcol="3" gridrow="11" pageId="5" pageNumber="534">0.15</td>
<td id="1F115DAAFFF212E4FC12FC27E97347D1" box="[922,1049,959,984]" gridcol="4" gridrow="11" pageId="5" pageNumber="534">0.14–0.15</td>
<td id="1F115DAAFFF212E4FBEDFC27E9C047D1" box="[1125,1194,959,984]" gridcol="5" gridrow="11" pageId="5" pageNumber="534">0.15</td>
<td id="1F115DAAFFF212E4FB56FC27E83747D1" box="[1246,1373,959,984]" gridcol="6" gridrow="11" pageId="5" pageNumber="534">0.15–0.16</td>
</tr>
<tr id="5CC034D6FFF212E4FF2EFC7DE83747F7" box="[166,1373,997,1022]" gridrow="12" pageId="5" pageNumber="534">
<th id="1F115DAAFFF212E4FF2EFC7DEC1947F7" box="[166,371,997,1022]" gridcol="0" gridrow="12" pageId="5" pageNumber="534">EY/SVL</th>
<td id="1F115DAAFFF212E4FE53FC7DEF4B47F7" box="[475,545,997,1022]" gridcol="1" gridrow="12" pageId="5" pageNumber="534">0.063</td>
<td id="1F115DAAFFF212E4FDDDFC7DEFBE47F7" box="[597,724,997,1022]" gridcol="2" gridrow="12" pageId="5" pageNumber="534">0.060–0.065</td>
<td id="1F115DAAFFF212E4FCA8FC7DEE1E47F7" box="[800,884,997,1022]" gridcol="3" gridrow="12" pageId="5" pageNumber="534">0.066</td>
<td id="1F115DAAFFF212E4FC12FC7DE97347F7" box="[922,1049,997,1022]" gridcol="4" gridrow="12" pageId="5" pageNumber="534">0.063–0.073</td>
<td id="1F115DAAFFF212E4FBEDFC7DE9C047F7" box="[1125,1194,997,1022]" gridcol="5" gridrow="12" pageId="5" pageNumber="534">0.074</td>
<td id="1F115DAAFFF212E4FB56FC7DE83747F7" box="[1246,1373,997,1022]" gridcol="6" gridrow="12" pageId="5" pageNumber="534">0.071–0.077</td>
</tr>
<tr id="5CC034D6FFF212E4FF2EFB93E837402D" box="[166,1373,1035,1060]" gridrow="13" pageId="5" pageNumber="534">
<th id="1F115DAAFFF212E4FF2EFB93EC19402D" box="[166,371,1035,1060]" gridcol="0" gridrow="13" pageId="5" pageNumber="534">EN/SVL</th>
<td id="1F115DAAFFF212E4FE53FB93EF4B402D" box="[475,545,1035,1060]" gridcol="1" gridrow="13" pageId="5" pageNumber="534">0.092</td>
<td id="1F115DAAFFF212E4FDDDFB93EFBE402D" box="[597,724,1035,1060]" gridcol="2" gridrow="13" pageId="5" pageNumber="534">0.090–0.094</td>
<td id="1F115DAAFFF212E4FCA8FB93EE1E402D" box="[800,884,1035,1060]" gridcol="3" gridrow="13" pageId="5" pageNumber="534">0.095</td>
<td id="1F115DAAFFF212E4FC12FB93E973402D" box="[922,1049,1035,1060]" gridcol="4" gridrow="13" pageId="5" pageNumber="534">0.092–0.100</td>
<td id="1F115DAAFFF212E4FBEDFB93E9C0402D" box="[1125,1194,1035,1060]" gridcol="5" gridrow="13" pageId="5" pageNumber="534">0.094</td>
<td id="1F115DAAFFF212E4FB56FB93E837402D" box="[1246,1373,1035,1060]" gridcol="6" gridrow="13" pageId="5" pageNumber="534">0.091–0.097</td>
</tr>
<tr id="5CC034D6FFF212E4FF2EFBA9E8374043" box="[166,1373,1073,1098]" gridrow="14" pageId="5" pageNumber="534">
<th id="1F115DAAFFF212E4FF2EFBA9EC194043" box="[166,371,1073,1098]" gridcol="0" gridrow="14" pageId="5" pageNumber="534">IN/SVL</th>
<td id="1F115DAAFFF212E4FE53FBA9EF4B4043" box="[475,545,1073,1098]" gridcol="1" gridrow="14" pageId="5" pageNumber="534">0.038</td>
<td id="1F115DAAFFF212E4FDDDFBA9EFBE4043" box="[597,724,1073,1098]" gridcol="2" gridrow="14" pageId="5" pageNumber="534">0.036–0.040</td>
<td id="1F115DAAFFF212E4FCA8FBA9EE1E4043" box="[800,884,1073,1098]" gridcol="3" gridrow="14" pageId="5" pageNumber="534">0.038</td>
<td id="1F115DAAFFF212E4FC12FBA9E9734043" box="[922,1049,1073,1098]" gridcol="4" gridrow="14" pageId="5" pageNumber="534">0.032–0.041</td>
<td id="1F115DAAFFF212E4FBEDFBA9E9C04043" box="[1125,1194,1073,1098]" gridcol="5" gridrow="14" pageId="5" pageNumber="534">0.034</td>
<td id="1F115DAAFFF212E4FB56FBA9E8374043" box="[1246,1373,1073,1098]" gridcol="6" gridrow="14" pageId="5" pageNumber="534">0.033–0.034</td>
</tr>
<tr id="5CC034D6FFF212E4FF2EFBCFE8374066" box="[166,1373,1111,1135]" gridrow="15" pageId="5" pageNumber="534">
<th id="1F115DAAFFF212E4FF2EFBCFEC194066" box="[166,371,1111,1135]" gridcol="0" gridrow="15" pageId="5" pageNumber="534">SN/SVL</th>
<td id="1F115DAAFFF212E4FE53FBCFEF4B4066" box="[475,545,1111,1135]" gridcol="1" gridrow="15" pageId="5" pageNumber="534">0.11</td>
<td id="1F115DAAFFF212E4FDDDFBCFEFBE4066" box="[597,724,1111,1135]" gridcol="2" gridrow="15" pageId="5" pageNumber="534">0.10–0.11</td>
<td id="1F115DAAFFF212E4FCA8FBCFEE1E4066" box="[800,884,1111,1135]" gridcol="3" gridrow="15" pageId="5" pageNumber="534">0.11</td>
<td id="1F115DAAFFF212E4FC12FBCFE9734066" box="[922,1049,1111,1135]" gridcol="4" gridrow="15" pageId="5" pageNumber="534">0.11–0.12</td>
<td id="1F115DAAFFF212E4FBEDFBCFE9C04066" box="[1125,1194,1111,1135]" gridcol="5" gridrow="15" pageId="5" pageNumber="534">0.12</td>
<td id="1F115DAAFFF212E4FB56FBCFE8374066" box="[1246,1373,1111,1135]" gridcol="6" gridrow="15" pageId="5" pageNumber="534">0.12</td>
</tr>
<tr id="5CC034D6FFF212E4FF2EFBE4E837409C" box="[166,1373,1148,1173]" gridrow="16" pageId="5" pageNumber="534">
<th id="1F115DAAFFF212E4FF2EFBE4EC19409C" box="[166,371,1148,1173]" gridcol="0" gridrow="16" pageId="5" pageNumber="534">EE/SVL</th>
<td id="1F115DAAFFF212E4FE53FBE4EF4B409C" box="[475,545,1148,1173]" gridcol="1" gridrow="16" pageId="5" pageNumber="534">0.085</td>
<td id="1F115DAAFFF212E4FDDDFBE4EFBE409C" box="[597,724,1148,1173]" gridcol="2" gridrow="16" pageId="5" pageNumber="534">0.084–0.086</td>
<td id="1F115DAAFFF212E4FCA8FBE4EE1E409C" box="[800,884,1148,1173]" gridcol="3" gridrow="16" pageId="5" pageNumber="534">0.086</td>
<td id="1F115DAAFFF212E4FC12FBE4E973409C" box="[922,1049,1148,1173]" gridcol="4" gridrow="16" pageId="5" pageNumber="534">0.080–0.092</td>
<td id="1F115DAAFFF212E4FBEDFBE4E9C0409C" box="[1125,1194,1148,1173]" gridcol="5" gridrow="16" pageId="5" pageNumber="534">0.078</td>
<td id="1F115DAAFFF212E4FB56FBE4E837409C" box="[1246,1373,1148,1173]" gridcol="6" gridrow="16" pageId="5" pageNumber="534">0.078–0.079</td>
</tr>
<tr id="5CC034D6FFF212E4FF2EFB3AE83740B2" box="[166,1373,1186,1211]" gridrow="17" pageId="5" pageNumber="534">
<th id="1F115DAAFFF212E4FF2EFB3AEC1940B2" box="[166,371,1186,1211]" gridcol="0" gridrow="17" pageId="5" pageNumber="534">IO/SVL</th>
<td id="1F115DAAFFF212E4FE53FB3AEF4B40B2" box="[475,545,1186,1211]" gridcol="1" gridrow="17" pageId="5" pageNumber="534">0.073</td>
<td id="1F115DAAFFF212E4FDDDFB3AEFBE40B2" box="[597,724,1186,1211]" gridcol="2" gridrow="17" pageId="5" pageNumber="534">0.071–0.075</td>
<td id="1F115DAAFFF212E4FCA8FB3AEE1E40B2" box="[800,884,1186,1211]" gridcol="3" gridrow="17" pageId="5" pageNumber="534">0.073</td>
<td id="1F115DAAFFF212E4FC12FB3AE97340B2" box="[922,1049,1186,1211]" gridcol="4" gridrow="17" pageId="5" pageNumber="534">0.061–0.081</td>
<td id="1F115DAAFFF212E4FBEDFB3AE9C040B2" box="[1125,1194,1186,1211]" gridcol="5" gridrow="17" pageId="5" pageNumber="534">0.069</td>
<td id="1F115DAAFFF212E4FB56FB3AE83740B2" box="[1246,1373,1186,1211]" gridcol="6" gridrow="17" pageId="5" pageNumber="534">0.067–0.071</td>
</tr>
<tr id="5CC034D6FFF212E4FF2EFB50E83740E8" box="[166,1373,1224,1249]" gridrow="18" pageId="5" pageNumber="534">
<th id="1F115DAAFFF212E4FF2EFB50EC1940E8" box="[166,371,1224,1249]" gridcol="0" gridrow="18" pageId="5" pageNumber="534">HW/HL</th>
<td id="1F115DAAFFF212E4FE53FB50EF4B40E8" box="[475,545,1224,1249]" gridcol="1" gridrow="18" pageId="5" pageNumber="534">0.66</td>
<td id="1F115DAAFFF212E4FDDDFB50EFBE40E8" box="[597,724,1224,1249]" gridcol="2" gridrow="18" pageId="5" pageNumber="534">0.66</td>
<td id="1F115DAAFFF212E4FCA8FB50EE1E40E8" box="[800,884,1224,1249]" gridcol="3" gridrow="18" pageId="5" pageNumber="534">0.67</td>
<td id="1F115DAAFFF212E4FC12FB50E97340E8" box="[922,1049,1224,1249]" gridcol="4" gridrow="18" pageId="5" pageNumber="534">0.63–0.69</td>
<td id="1F115DAAFFF212E4FBEDFB50E9C040E8" box="[1125,1194,1224,1249]" gridcol="5" gridrow="18" pageId="5" pageNumber="534">0.63</td>
<td id="1F115DAAFFF212E4FB56FB50E83740E8" box="[1246,1373,1224,1249]" gridcol="6" gridrow="18" pageId="5" pageNumber="534">0.61–0.64</td>
</tr>
<tr id="5CC034D6FFF212E4FF2EFB76E837410E" box="[166,1373,1262,1287]" gridrow="19" pageId="5" pageNumber="534">
<th id="1F115DAAFFF212E4FF2EFB76EC19410E" box="[166,371,1262,1287]" gridcol="0" gridrow="19" pageId="5" pageNumber="534">EY/EN</th>
<td id="1F115DAAFFF212E4FE53FB76EF4B410E" box="[475,545,1262,1287]" gridcol="1" gridrow="19" pageId="5" pageNumber="534">0.68</td>
<td id="1F115DAAFFF212E4FDDDFB76EFBE410E" box="[597,724,1262,1287]" gridcol="2" gridrow="19" pageId="5" pageNumber="534">0.67–0.69</td>
<td id="1F115DAAFFF212E4FCA8FB76EE1E410E" box="[800,884,1262,1287]" gridcol="3" gridrow="19" pageId="5" pageNumber="534">0.69</td>
<td id="1F115DAAFFF212E4FC12FB76E973410E" box="[922,1049,1262,1287]" gridcol="4" gridrow="19" pageId="5" pageNumber="534">0.64–0.77</td>
<td id="1F115DAAFFF212E4FBEDFB76E9C0410E" box="[1125,1194,1262,1287]" gridcol="5" gridrow="19" pageId="5" pageNumber="534">0.79</td>
<td id="1F115DAAFFF212E4FB56FB76E837410E" box="[1246,1373,1262,1287]" gridcol="6" gridrow="19" pageId="5" pageNumber="534">0.73–0.85</td>
</tr>
<tr id="5CC034D6FFF212E4FF2EFA8BE8374125" box="[166,1373,1299,1324]" gridrow="20" pageId="5" pageNumber="534">
<th id="1F115DAAFFF212E4FF2EFA8BEC194125" box="[166,371,1299,1324]" gridcol="0" gridrow="20" pageId="5" pageNumber="534">EN/IN</th>
<td id="1F115DAAFFF212E4FE53FA8BEF4B4125" box="[475,545,1299,1324]" gridcol="1" gridrow="20" pageId="5" pageNumber="534">2.4</td>
<td id="1F115DAAFFF212E4FDDDFA8BEFBE4125" box="[597,724,1299,1324]" gridcol="2" gridrow="20" pageId="5" pageNumber="534">2.4–2.5</td>
<td id="1F115DAAFFF212E4FCA8FA8BEE1E4125" box="[800,884,1299,1324]" gridcol="3" gridrow="20" pageId="5" pageNumber="534">2.5</td>
<td id="1F115DAAFFF212E4FC12FA8BE9734125" box="[922,1049,1299,1324]" gridcol="4" gridrow="20" pageId="5" pageNumber="534">2.3–2.9</td>
<td id="1F115DAAFFF212E4FBEDFA8BE9C04125" box="[1125,1194,1299,1324]" gridcol="5" gridrow="20" pageId="5" pageNumber="534">2.8</td>
<td id="1F115DAAFFF212E4FB56FA8BE8374125" box="[1246,1373,1299,1324]" gridcol="6" gridrow="20" pageId="5" pageNumber="534">2.7–2.8</td>
</tr>
<tr id="5CC034D6FFF212E4FF2EFAA2E837415B" box="[166,1373,1338,1362]" gridrow="21" pageId="5" pageNumber="534">
<th id="1F115DAAFFF212E4FF2EFAA2EC19415B" box="[166,371,1338,1362]" gridcol="0" gridrow="21" pageId="5" pageNumber="534">SN/HL</th>
<td id="1F115DAAFFF212E4FE53FAA2EF4B415B" box="[475,545,1338,1362]" gridcol="1" gridrow="21" pageId="5" pageNumber="534">0.39</td>
<td id="1F115DAAFFF212E4FDDDFAA2EFBE415B" box="[597,724,1338,1362]" gridcol="2" gridrow="21" pageId="5" pageNumber="534">0.38–0.40</td>
<td id="1F115DAAFFF212E4FCA8FAA2EE1E415B" box="[800,884,1338,1362]" gridcol="3" gridrow="21" pageId="5" pageNumber="534">0.39</td>
<td id="1F115DAAFFF212E4FC12FAA2E973415B" box="[922,1049,1338,1362]" gridcol="4" gridrow="21" pageId="5" pageNumber="534">0.38–0.41</td>
<td id="1F115DAAFFF212E4FBEDFAA2E9C0415B" box="[1125,1194,1338,1362]" gridcol="5" gridrow="21" pageId="5" pageNumber="534">0.41</td>
<td id="1F115DAAFFF212E4FB56FAA2E837415B" box="[1246,1373,1338,1362]" gridcol="6" gridrow="21" pageId="5" pageNumber="534">0.40–0.42</td>
</tr>
<tr id="5CC034D6FFF212E4FF2EFAC7E8374171" box="[166,1373,1375,1400]" gridrow="22" pageId="5" pageNumber="534">
<th id="1F115DAAFFF212E4FF2EFAC7EC194171" box="[166,371,1375,1400]" gridcol="0" gridrow="22" pageId="5" pageNumber="534">HH/HL</th>
<td id="1F115DAAFFF212E4FE53FAC7EF4B4171" box="[475,545,1375,1400]" gridcol="1" gridrow="22" pageId="5" pageNumber="534">0.42</td>
<td id="1F115DAAFFF212E4FDDDFAC7EFBE4171" box="[597,724,1375,1400]" gridcol="2" gridrow="22" pageId="5" pageNumber="534">0.40–0.43</td>
<td id="1F115DAAFFF212E4FCA8FAC7EE1E4171" box="[800,884,1375,1400]" gridcol="3" gridrow="22" pageId="5" pageNumber="534">0.40</td>
<td id="1F115DAAFFF212E4FC12FAC7E9734171" box="[922,1049,1375,1400]" gridcol="4" gridrow="22" pageId="5" pageNumber="534">0.37–0.42</td>
<td id="1F115DAAFFF212E4FBEDFAC7E9C04171" box="[1125,1194,1375,1400]" gridcol="5" gridrow="22" pageId="5" pageNumber="534">0.40</td>
<td id="1F115DAAFFF212E4FB56FAC7E8374171" box="[1246,1373,1375,1400]" gridcol="6" gridrow="22" pageId="5" pageNumber="534">0.36–0.44</td>
</tr>
<tr id="5CC034D6FFF212E4FF2EFA1DE8374197" box="[166,1373,1413,1438]" gridrow="23" pageId="5" pageNumber="534">
<th id="1F115DAAFFF212E4FF2EFA1DEC194197" box="[166,371,1413,1438]" gridcol="0" gridrow="23" pageId="5" pageNumber="534">SL to mid-eye</th>
<td id="1F115DAAFFF212E4FE53FA1DEF4B4197" box="[475,545,1413,1438]" gridcol="1" gridrow="23" pageId="5" pageNumber="534">10.0</td>
<td id="1F115DAAFFF212E4FDDDFA1DEFBE4197" box="[597,724,1413,1438]" gridcol="2" gridrow="23" pageId="5" pageNumber="534">9–11</td>
<td id="1F115DAAFFF212E4FCA8FA1DEE1E4197" box="[800,884,1413,1438]" gridcol="3" gridrow="23" pageId="5" pageNumber="534">9.9</td>
<td id="1F115DAAFFF212E4FC12FA1DE9734197" box="[922,1049,1413,1438]" gridcol="4" gridrow="23" pageId="5" pageNumber="534">9–11</td>
<td id="1F115DAAFFF212E4FBEDFA1DE9C04197" box="[1125,1194,1413,1438]" gridcol="5" gridrow="23" pageId="5" pageNumber="534">11</td>
<td id="1F115DAAFFF212E4FB56FA1DE8374197" box="[1246,1373,1413,1438]" gridcol="6" gridrow="23" pageId="5" pageNumber="534">11</td>
</tr>
<tr id="5CC034D6FFF212E4FF2EFA32E83741CA" box="[166,1373,1450,1475]" gridrow="24" pageId="5" pageNumber="534">
<th id="1F115DAAFFF212E4FF2EFA32EC1941CA" box="[166,371,1450,1475]" gridcol="0" gridrow="24" pageId="5" pageNumber="534">SL to jaw</th>
<td id="1F115DAAFFF212E4FE53FA32EF4B41CA" box="[475,545,1450,1475]" gridcol="1" gridrow="24" pageId="5" pageNumber="534">12.7</td>
<td id="1F115DAAFFF212E4FDDDFA32EFBE41CA" box="[597,724,1450,1475]" gridcol="2" gridrow="24" pageId="5" pageNumber="534">12–14</td>
<td id="1F115DAAFFF212E4FCA8FA32EE1E41CA" box="[800,884,1450,1475]" gridcol="3" gridrow="24" pageId="5" pageNumber="534">12.7</td>
<td id="1F115DAAFFF212E4FC12FA32E97341CA" box="[922,1049,1450,1475]" gridcol="4" gridrow="24" pageId="5" pageNumber="534">11–14</td>
<td id="1F115DAAFFF212E4FBEDFA32E9C041CA" box="[1125,1194,1450,1475]" gridcol="5" gridrow="24" pageId="5" pageNumber="534">13.2</td>
<td id="1F115DAAFFF212E4FB56FA32E83741CA" box="[1246,1373,1450,1475]" gridcol="6" gridrow="24" pageId="5" pageNumber="534">13–14</td>
</tr>
<tr id="5CC034D6FFF212E4FF2EFA48E83741E0" box="[166,1373,1488,1513]" gridrow="25" pageId="5" pageNumber="534">
<th id="1F115DAAFFF212E4FF2EFA48EC1941E0" box="[166,371,1488,1513]" gridcol="0" gridrow="25" pageId="5" pageNumber="534">Infralabials</th>
<td id="1F115DAAFFF212E4FE53FA48EF4B41E0" box="[475,545,1488,1513]" gridcol="1" gridrow="25" pageId="5" pageNumber="534">11.0</td>
<td id="1F115DAAFFF212E4FDDDFA48EFBE41E0" box="[597,724,1488,1513]" gridcol="2" gridrow="25" pageId="5" pageNumber="534">10–12</td>
<td id="1F115DAAFFF212E4FCA8FA48EE1E41E0" box="[800,884,1488,1513]" gridcol="3" gridrow="25" pageId="5" pageNumber="534">11.6</td>
<td id="1F115DAAFFF212E4FC12FA48E97341E0" box="[922,1049,1488,1513]" gridcol="4" gridrow="25" pageId="5" pageNumber="534">11–12</td>
<td id="1F115DAAFFF212E4FBEDFA48E9C041E0" box="[1125,1194,1488,1513]" gridcol="5" gridrow="25" pageId="5" pageNumber="534">12.0</td>
<td id="1F115DAAFFF212E4FB56FA48E83741E0" box="[1246,1373,1488,1513]" gridcol="6" gridrow="25" pageId="5" pageNumber="534">11–13</td>
</tr>
<tr id="5CC034D6FFF212E4FF2EFA6EE8374206" box="[166,1373,1526,1551]" gridrow="26" pageId="5" pageNumber="534">
<th id="1F115DAAFFF212E4FF2EFA6EEC194206" box="[166,371,1526,1551]" gridcol="0" gridrow="26" pageId="5" pageNumber="534">T4 scales</th>
<td id="1F115DAAFFF212E4FE53FA6EEF4B4206" box="[475,545,1526,1551]" gridcol="1" gridrow="26" pageId="5" pageNumber="534">28.5</td>
<td id="1F115DAAFFF212E4FDDDFA6EEFBE4206" box="[597,724,1526,1551]" gridcol="2" gridrow="26" pageId="5" pageNumber="534">25–31</td>
<td id="1F115DAAFFF212E4FCA8FA6EEE1E4206" box="[800,884,1526,1551]" gridcol="3" gridrow="26" pageId="5" pageNumber="534">25.8</td>
<td id="1F115DAAFFF212E4FC12FA6EE9734206" box="[922,1049,1526,1551]" gridcol="4" gridrow="26" pageId="5" pageNumber="534">24–28</td>
<td id="1F115DAAFFF212E4FBEDFA6EE9C04206" box="[1125,1194,1526,1551]" gridcol="5" gridrow="26" pageId="5" pageNumber="534">28.2</td>
<td id="1F115DAAFFF212E4FB56FA6EE8374206" box="[1246,1373,1526,1551]" gridcol="6" gridrow="26" pageId="5" pageNumber="534">28–29</td>
</tr>
<tr id="5CC034D6FFF212E4FF2EF984E837423C" box="[166,1373,1564,1589]" gridrow="27" pageId="5" pageNumber="534">
<th id="1F115DAAFFF212E4FF2EF984EC19423C" box="[166,371,1564,1589]" gridcol="0" gridrow="27" pageId="5" pageNumber="534">Midbelly scale rows</th>
<td id="1F115DAAFFF212E4FE53F984EF4B423C" box="[475,545,1564,1589]" gridcol="1" gridrow="27" pageId="5" pageNumber="534">43.0</td>
<td id="1F115DAAFFF212E4FDDDF984EFBE423C" box="[597,724,1564,1589]" gridcol="2" gridrow="27" pageId="5" pageNumber="534">41–45</td>
<td id="1F115DAAFFF212E4FCA8F984EE1E423C" box="[800,884,1564,1589]" gridcol="3" gridrow="27" pageId="5" pageNumber="534">43.2</td>
<td id="1F115DAAFFF212E4FC12F984E973423C" box="[922,1049,1564,1589]" gridcol="4" gridrow="27" pageId="5" pageNumber="534">41–47</td>
<td id="1F115DAAFFF212E4FBEDF984E9C0423C" box="[1125,1194,1564,1589]" gridcol="5" gridrow="27" pageId="5" pageNumber="534">42.5</td>
<td id="1F115DAAFFF212E4FB56F984E837423C" box="[1246,1373,1564,1589]" gridcol="6" gridrow="27" pageId="5" pageNumber="534">40–45</td>
</tr>
<tr id="5CC034D6FFF212E4FF2EF9DAE8374252" box="[166,1373,1602,1627]" gridrow="28" pageId="5" pageNumber="534">
<th id="1F115DAAFFF212E4FF2EF9DAEC194252" box="[166,371,1602,1627]" gridcol="0" gridrow="28" pageId="5" pageNumber="534">Tubercle rows</th>
<td id="1F115DAAFFF212E4FE53F9DAEF4B4252" box="[475,545,1602,1627]" gridcol="1" gridrow="28" pageId="5" pageNumber="534">22.5</td>
<td id="1F115DAAFFF212E4FDDDF9DAEFBE4252" box="[597,724,1602,1627]" gridcol="2" gridrow="28" pageId="5" pageNumber="534">22–23</td>
<td id="1F115DAAFFF212E4FCA8F9DAEE1E4252" box="[800,884,1602,1627]" gridcol="3" gridrow="28" pageId="5" pageNumber="534">22.8</td>
<td id="1F115DAAFFF212E4FC12F9DAE9734252" box="[922,1049,1602,1627]" gridcol="4" gridrow="28" pageId="5" pageNumber="534">22–24</td>
<td id="1F115DAAFFF212E4FBEDF9DAE9C04252" box="[1125,1194,1602,1627]" gridcol="5" gridrow="28" pageId="5" pageNumber="534">22</td>
<td id="1F115DAAFFF212E4FB56F9DAE8374252" box="[1246,1373,1602,1627]" gridcol="6" gridrow="28" pageId="5" pageNumber="534">22</td>
</tr>
<tr id="5CC034D6FFF212E4FF2EF9FFE8374289" box="[166,1373,1639,1664]" gridrow="29" pageId="5" pageNumber="534">
<th id="1F115DAAFFF212E4FF2EF9FFEC194289" box="[166,371,1639,1664]" gridcol="0" gridrow="29" pageId="5" pageNumber="534">Dark dorsal bands</th>
<td id="1F115DAAFFF212E4FE53F9FFEF4B4289" box="[475,545,1639,1664]" gridcol="1" gridrow="29" pageId="5" pageNumber="534">6</td>
<td id="1F115DAAFFF212E4FDDDF9FFEFBE4289" box="[597,724,1639,1664]" gridcol="2" gridrow="29" pageId="5" pageNumber="534">6</td>
<td id="1F115DAAFFF212E4FCA8F9FFEE1E4289" box="[800,884,1639,1664]" gridcol="3" gridrow="29" pageId="5" pageNumber="534">5.8</td>
<td id="1F115DAAFFF212E4FC12F9FFE9734289" box="[922,1049,1639,1664]" gridcol="4" gridrow="29" pageId="5" pageNumber="534">5–6</td>
<td id="1F115DAAFFF212E4FBEDF9FFE9C04289" box="[1125,1194,1639,1664]" gridcol="5" gridrow="29" pageId="5" pageNumber="534">6</td>
<td id="1F115DAAFFF212E4FB56F9FFE8374289" box="[1246,1373,1639,1664]" gridcol="6" gridrow="29" pageId="5" pageNumber="534">6</td>
</tr>
</table>
</paragraph>
<paragraph id="E24F3694FFF2ED1EFF4FF928EE274376" blockId="5.[151,1436,1712,2028]" pageId="5" pageNumber="534">
<emphasis id="D084EA86FFF2ED1EFF4FF928EC3542C2" bold="true" box="[199,351,1712,1739]" italics="true" pageId="5" pageNumber="534">Color in life.</emphasis>
Field notes for UMMZ 247749 recorded the animal as golden brown with dark-brown bands, each partially and narrowly margined in pale golden white; legs with many pale golden spots; head marbled golden white and brown; iris brass; venter pale golden white. UMMZ 247750 differed from this in being brown above with dark-brown bands, no pale golden spots, iris darker brass and brown, and venter pale purplish. UMMZ 247746 had a medium-brown ground color, with darker-brown bands having a few chocolate-brown spots along the margins and superciliaries that were yellowish with brown spots (
<figureCitation id="7ACB2A11FFF2ED1EFD6EF8FDEE2A4376" box="[742,832,1893,1919]" captionStart="FIGURE 3" captionStartId="6.[151,250,1184,1209]" captionTargetBox="[156,1431,307,1157]" captionTargetId="figure@6.[151,1436,301,1164]" captionTargetPageId="6" captionText="FIGURE 3. Portraits in life of (A) paratype of Cyrtodactylus atremus sp. nov., UMMZ 247746, Karkar Island, Madang Province, PNG; (B) Cyrtodactylus sermowaiensis, BPBM 23317, Parkop, Torricelli Mts., West Sepik Province, PNG; (C) Cyrtodactylus sermowaiensis, BPBM 23320, base of Mt. Sapau, Torricelli Mts., West Sepik Province, PNG; and (D) Cyrtodactylus sermowaiensis, BPBM 35598, Yefehi, Prince Alexander Mts., East Sepik Province, PNG." figureDoi="http://doi.org/10.5281/zenodo.3542685" httpUri="https://zenodo.org/record/3542685/files/figure.png" pageId="5" pageNumber="534">Fig. 3A</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="AAEA651FFFF2ED1DFF4FF810EC774516" lastPageId="6" lastPageNumber="535" pageId="5" pageNumber="534" type="etymology">
<paragraph id="E24F3694FFF2ED1EFF4FF810E9B143CE" blockId="5.[151,1436,1712,2028]" pageId="5" pageNumber="534">
<emphasis id="D084EA86FFF2ED1EFF4FF810EC2343AA" bold="true" box="[199,329,1928,1955]" pageId="5" pageNumber="534">
<emphasis id="D084EA86FFF2ED1EFF4FF810EC2843AA" bold="true" box="[199,322,1928,1955]" italics="true" pageId="5" pageNumber="534">Etymology</emphasis>
.
</emphasis>
The name is a Latinized masculine compound adjective formed from the Greek 
<emphasis id="D084EA86FFF2ED1EFB48F812E9A443AA" box="[1216,1230,1930,1955]" italics="true" pageId="5" pageNumber="534">a</emphasis>
, meaning 
<emphasis id="D084EA86FFF2ED1EFAC9F811E8FF43AA" box="[1345,1429,1929,1955]" italics="true" pageId="5" pageNumber="534">without</emphasis>
, and 
<emphasis id="D084EA86FFF2ED1EFF4FF836EC6D43CE" box="[199,263,1966,1991]" italics="true" pageId="5" pageNumber="534">trema</emphasis>
, meaning 
<emphasis id="D084EA86FFF2ED1EFEF4F835ECC643CE" box="[380,428,1965,1991]" italics="true" pageId="5" pageNumber="534">hole</emphasis>
. It refers to the lack of precloacal/femoral pores in males of this species.
</paragraph>
<paragraph id="E24F3694FFF2ED1DFF4FF84AEC774516" blockId="5.[151,1436,1712,2028]" lastBlockId="6.[151,1437,153,287]" lastPageId="6" lastPageNumber="535" pageId="5" pageNumber="534">
<emphasis id="D084EA86FFF2ED1EFF4FF84AEC7243E5" bold="true" box="[199,280,2002,2028]" italics="true" pageId="5" pageNumber="534">Range.</emphasis>
Known from the western slopes of Mount Uluman at altitudes between 130–650 meters on Karkar Island, but no doubt ranges across the lower elevations of the entire island (
<figureCitation id="7ACB2A11FFF1ED1DFC7EFF01E95144BA" box="[1014,1083,153,179]" captionStart="FIGURE 4" captionStartId="7.[151,250,1379,1404]" captionTargetBox="[153,1435,445,1359]" captionTargetId="figure@7.[151,1435,445,1359]" captionTargetPageId="7" captionText="FIGURE 4. Map of New Guinea and surrounding islands showing known localities taken from American and Australian museum catalogues for Cyrtodactylus atremus sp. nov. (star), C. sermowaiensis (dots), and an undescribed species of Cyrtodactylus in the C. sermowaiensis species group (diamond). The type locality for C. sermowaiensis is along the Sermowai River in Indonesian New Guinea, approximately 200 km WNW of the westernmost locality shown on this map. Contour intervals are 0 (white), 500 (light gray), and 1000 (dark gray) meters a.s.l." figureDoi="http://doi.org/10.5281/zenodo.3542687" httpUri="https://zenodo.org/record/3542687/files/figure.png" pageId="6" pageNumber="535">Fig. 4</figureCitation>
) where suitable habitat occurs. Karkar attains of maximum elevation of 
<quantity id="25089B71FFF1ED1DFDD0FF25EFC444DE" box="[600,686,189,215]" metricMagnitude="3" metricUnit="m" metricValue="1.8399999999999999" pageId="6" pageNumber="535" unit="m" value="1840.0">1840 m</quantity>
, and it is likely that this species will only occur up to approximately 
<quantity id="25089B71FFF1ED1DFF1FFF79EC5F44F2" box="[151,309,225,251]" metricMagnitude="3" metricUnit="m" metricValue="1.1" metricValueMax="1.2" metricValueMin="1.0" pageId="6" pageNumber="535" unit="m" value="1100.0" valueMax="1200.0" valueMin="1000.0">1000–1200 m</quantity>
elevation—depending on the elevation at which cloud forest begins—although that remains to be determined.
</paragraph>
</subSubSection>
<caption id="B68F661CFFF1ED1DFF1FFB38E949412D" ID-DOI="http://doi.org/10.5281/zenodo.3542685" ID-Zenodo-Dep="3542685" httpUri="https://zenodo.org/record/3542685/files/figure.png" pageId="6" pageNumber="535" startId="6.[151,250,1184,1209]" targetBox="[156,1431,307,1157]" targetPageId="6">
<paragraph id="E24F3694FFF1ED1DFF1FFB38E949412D" blockId="6.[151,1436,1184,1317]" pageId="6" pageNumber="535">
<emphasis id="D084EA86FFF1ED1DFF1FFB38EC7940B0" bold="true" box="[151,275,1184,1209]" pageId="6" pageNumber="535">FIGURE 3.</emphasis>
Portraits in life of (A) paratype of 
<taxonomicName id="25F04D17FFF1ED1DFDF1FB38EE3540B1" authority="Kraus &amp; Weijola, 2019" authorityName="Kraus &amp; Weijola" authorityYear="2019" box="[633,863,1184,1208]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="535" phylum="Chordata" rank="species" species="atremus" status="sp. nov.">
<emphasis id="D084EA86FFF1ED1DFDF1FB38EE3540B1" box="[633,863,1184,1208]" italics="true" pageId="6" pageNumber="535">Cyrtodactylus atremus</emphasis>
</taxonomicName>
<emphasis id="D084EA86FFF1ED1DFCEDFB39EEDC40B0" bold="true" box="[869,950,1185,1209]" pageId="6" pageNumber="535">
<taxonomicNameLabel id="CBB757FDFFF1ED1DFCEDFB39EEDC40B0" box="[869,950,1185,1209]" pageId="6" pageNumber="535" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, UMMZ 247746, Karkar Island, Madang Province, PNG; (B) 
<taxonomicName id="25F04D17FFF1ED1DFEB4FB5CEF0D40D5" baseAuthorityName="Kraus" baseAuthorityYear="2008" box="[316,615,1220,1244]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="535" phylum="Chordata" rank="species" species="sermowaiensis">
<emphasis id="D084EA86FFF1ED1DFEB4FB5CEF0D40D5" box="[316,615,1220,1244]" italics="true" pageId="6" pageNumber="535">Cyrtodactylus sermowaiensis</emphasis>
</taxonomicName>
, BPBM 23317, Parkop, Torricelli Mts., West Sepik Province, PNG; (C) 
<taxonomicName id="25F04D17FFF1ED1DFAD2FB5DECE24109" baseAuthorityName="Kraus" baseAuthorityYear="2008" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="535" phylum="Chordata" rank="species" species="sermowaiensis">
<emphasis id="D084EA86FFF1ED1DFAD2FB5DECE24109" italics="true" pageId="6" pageNumber="535">Cyrtodactylus sermowaiensis</emphasis>
</taxonomicName>
, BPBM 23320, base of Mt. Sapau, Torricelli Mts., West Sepik Province, PNG; and (D) 
<taxonomicName id="25F04D17FFF1ED1DFA86FB70EC46412D" baseAuthorityName="Kraus" baseAuthorityYear="2008" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="535" phylum="Chordata" rank="species" species="sermowaiensis">
<emphasis id="D084EA86FFF1ED1DFA86FB70EC46412D" italics="true" pageId="6" pageNumber="535">Cyrtodactylus sermowaiensis</emphasis>
</taxonomicName>
, BPBM 35598, Yefehi, Prince Alexander Mts., East Sepik Province, PNG.
</paragraph>
</caption>
<subSubSection id="AAEA651FFFF1ED1CFF4FFACAEE4145B9" lastPageId="7" lastPageNumber="536" pageId="6" pageNumber="535" type="biology_ecology">
<paragraph id="E24F3694FFF1ED1DFF4FFACAEE8342DC" blockId="6.[151,1437,1362,2037]" pageId="6" pageNumber="535">
<emphasis id="D084EA86FFF1ED1DFF4FFACAEC414164" bold="true" box="[199,299,1362,1389]" pageId="6" pageNumber="535">
<emphasis id="D084EA86FFF1ED1DFF4FFACAEC4E4164" bold="true" box="[199,292,1362,1389]" italics="true" pageId="6" pageNumber="535">Ecology</emphasis>
.
</emphasis>
The six UMMZ specimens were all found uphill from the village of Mom on the western slopes of Mount Uluman at altitudes between 
<quantity id="25089B71FFF1ED1DFDBDFAEEEFDF4198" box="[565,693,1398,1425]" metricMagnitude="2" metricUnit="m" metricValue="5.75" metricValueMax="6.5" metricValueMin="5.0" pageId="6" pageNumber="535" unit="m" value="575.0" valueMax="650.0" valueMin="500.0">500–650 m</quantity>
a.s.l. The tropical rainforests in this area are still fairly intact and recently undisturbed though appearing open enough to be of secondary development (
<figureCitation id="7ACB2A11FFF1ED1DFBECFA03E98A41BC" box="[1124,1248,1434,1461]" captionStart="FIGURE 5" captionStartId="8.[151,250,1578,1603]" captionTargetBox="[160,1427,379,1550]" captionTargetId="figure@8.[151,1436,373,1558]" captionTargetPageId="8" captionText="FIGURE 5. (A), (B) Habitat of Cyrtodactylus atremus sp. nov. on the lower northwestern slopes of Mt. Uluman, Karkar Island, Madang Province, PNG; (C) UMMZ 247751, and (D) UMMZ 247750 in situ showing their head-down resting position during the daytime." figureDoi="http://doi.org/10.5281/zenodo.3542689" httpUri="https://zenodo.org/record/3542689/files/figure.png" pageId="6" pageNumber="535">Fig. 5A, B</figureCitation>
). All individuals were caught during daytime head down on larger tree trunks or saplings, one to three meters from the ground (
<figureCitation id="7ACB2A11FFF1ED1DFAE7FA27EDB441F4" captionStart="FIGURE 5" captionStartId="8.[151,250,1578,1603]" captionTargetBox="[160,1427,379,1550]" captionTargetId="figure@8.[151,1436,373,1558]" captionTargetPageId="8" captionText="FIGURE 5. (A), (B) Habitat of Cyrtodactylus atremus sp. nov. on the lower northwestern slopes of Mt. Uluman, Karkar Island, Madang Province, PNG; (C) UMMZ 247751, and (D) UMMZ 247750 in situ showing their head-down resting position during the daytime." figureDoi="http://doi.org/10.5281/zenodo.3542689" httpUri="https://zenodo.org/record/3542689/files/figure.png" pageId="6" pageNumber="535">Fig. 5C, D</figureCitation>
). All were easy to approach and capture, which is not usual for 
<taxonomicName id="25F04D17FFF1ED1DFC2DFA7BE92841F4" authorityName="Gray" authorityYear="1827" box="[933,1090,1507,1533]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="535" phylum="Chordata" rank="genus">
<emphasis id="D084EA86FFF1ED1DFC2DFA7BE92841F4" box="[933,1090,1507,1533]" italics="true" pageId="6" pageNumber="535">Cyrtodactylus</emphasis>
</taxonomicName>
active at night, so we presume they were sleeping, though we have never seen such exposed resting behavior in the several other 
<taxonomicName id="25F04D17FFF1ED1DFB77F99FE8F64228" authorityName="Gray" authorityYear="1827" box="[1279,1436,1543,1569]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="535" phylum="Chordata" rank="genus">
<emphasis id="D084EA86FFF1ED1DFB77F99FE8F64228" box="[1279,1436,1543,1569]" italics="true" pageId="6" pageNumber="535">Cyrtodactylus</emphasis>
</taxonomicName>
species we have collected. Day- and night-time searches in disturbed habitats along the coast and lowland coconut plantations did not yield any specimens. Data for the AMS specimens are for the villages of Mom and Kevasob (ca. 270 and 
<quantity id="25089B71FFF1ED1DFF74F9EBEC2F4284" box="[252,325,1651,1677]" metricMagnitude="2" metricUnit="m" metricValue="1.3" pageId="6" pageNumber="535" unit="m" value="130.0">130 m</quantity>
a.s.l., respectively). We have never previously found 
<taxonomicName id="25F04D17FFF1ED1DFC27F9EBE9264284" authorityName="Gray" authorityYear="1827" box="[943,1100,1651,1677]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="535" phylum="Chordata" rank="genus">
<emphasis id="D084EA86FFF1ED1DFC27F9EBE9264284" box="[943,1100,1651,1677]" italics="true" pageId="6" pageNumber="535">Cyrtodactylus</emphasis>
</taxonomicName>
species in villages per se, so we presume that those lizards were collected in the forests adjacent to the villages. In any event, like several other Papuan 
<taxonomicName id="25F04D17FFF1ED1DFF79F923ECE442DC" authorityName="Gray" authorityYear="1827" box="[241,398,1723,1749]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="535" phylum="Chordata" rank="genus">
<emphasis id="D084EA86FFF1ED1DFF79F923ECE442DC" box="[241,398,1723,1749]" italics="true" pageId="6" pageNumber="535">Cyrtodactylus</emphasis>
</taxonomicName>
, 
<taxonomicName id="25F04D17FFF1ED1DFE14F923EF7F42DC" authorityName="Kraus &amp; Weijola" authorityYear="2019" box="[412,533,1723,1749]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="535" phylum="Chordata" rank="species" species="atremus">
<emphasis id="D084EA86FFF1ED1DFE14F923EF7F42DC" box="[412,533,1723,1749]" italics="true" pageId="6" pageNumber="535">C. atremus</emphasis>
</taxonomicName>
seems to persist well in secondary forest.
</paragraph>
<paragraph id="E24F3694FFF1ED1DFF4FF947E9184380" blockId="6.[151,1437,1362,2037]" pageId="6" pageNumber="535">
Other geckos present on Karkar include 
<taxonomicName id="25F04D17FFF1ED1DFD1AF947EE3842F0" box="[658,850,1759,1785]" class="Reptilia" family="Gekkonidae" genus="Gehyra" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="535" phylum="Chordata" rank="species" species="oceanica">
<emphasis id="D084EA86FFF1ED1DFD1AF947EE3842F0" box="[658,850,1759,1785]" italics="true" pageId="6" pageNumber="535">Gehyra oceanica</emphasis>
</taxonomicName>
, 
<taxonomicName id="25F04D17FFF1ED1DFCEAF947E96942F0" authorityName="Houttuyn" authorityYear="1782" box="[866,1027,1759,1785]" class="Reptilia" family="Gekkonidae" genus="Gekko" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="535" phylum="Chordata" rank="species" species="vittatus">
<emphasis id="D084EA86FFF1ED1DFCEAF947E96942F0" box="[866,1027,1759,1785]" italics="true" pageId="6" pageNumber="535">Gekko vittatus</emphasis>
</taxonomicName>
, 
<taxonomicName id="25F04D17FFF1ED1DFB9AF947E86542F0" box="[1042,1295,1759,1785]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="535" phylum="Chordata" rank="species" species="frenatus">
<emphasis id="D084EA86FFF1ED1DFB9AF947E86542F0" box="[1042,1295,1759,1785]" italics="true" pageId="6" pageNumber="535">Hemidactylus frenatus</emphasis>
</taxonomicName>
, and 
<taxonomicName id="25F04D17FFF1ED1DFAC6F978E8F642F0" authorityName="Kluge" authorityYear="1983" box="[1358,1436,1760,1785]" class="Reptilia" family="Gekkonidae" genus="Nactus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="535" phylum="Chordata" rank="genus">
<emphasis id="D084EA86FFF1ED1DFAC6F978E8F642F0" box="[1358,1436,1760,1785]" italics="true" pageId="6" pageNumber="535">Nactus</emphasis>
</taxonomicName>
sp. Of these, 
<taxonomicName id="25F04D17FFF1ED1DFEA5F89CEC114314" authorityName="Kluge" authorityYear="1983" box="[301,379,1796,1821]" class="Reptilia" family="Gekkonidae" genus="Nactus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="535" phylum="Chordata" rank="genus">
<emphasis id="D084EA86FFF1ED1DFEA5F89CEC114314" box="[301,379,1796,1821]" italics="true" pageId="6" pageNumber="535">Nactus</emphasis>
</taxonomicName>
sp. is an inhabitant of forest interiors, though it lives under debris on the forest floor, and the arboreal 
<taxonomicName id="25F04D17FFF1ED1DFF72F8BFECE84348" box="[250,386,1831,1857]" class="Reptilia" family="Gekkonidae" genus="Gehyra" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="535" phylum="Chordata" rank="species" species="oceanica">
<emphasis id="D084EA86FFF1ED1DFF72F8BFECE84348" box="[250,386,1831,1857]" italics="true" pageId="6" pageNumber="535">G. oceanica</emphasis>
</taxonomicName>
was found in the forest in the same vicinity as our specimens of 
<taxonomicName id="25F04D17FFF1ED1DFBD4F8BFE9BC4348" authorityName="Kraus &amp; Weijola" authorityYear="2019" box="[1116,1238,1831,1857]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="535" phylum="Chordata" rank="species" species="atremus">
<emphasis id="D084EA86FFF1ED1DFBD4F8BFE9BC4348" box="[1116,1238,1831,1857]" italics="true" pageId="6" pageNumber="535">C. atremus</emphasis>
</taxonomicName>
though hidden in leaf axils and under bark. It is unclear to what extent the other species might share the forest habitat of 
<taxonomicName id="25F04D17FFF1ED1DFA93F8D3E8FF436C" authorityName="Kraus &amp; Weijola" authorityYear="2019" box="[1307,1429,1867,1893]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="535" phylum="Chordata" rank="species" species="atremus">
<emphasis id="D084EA86FFF1ED1DFA93F8D3E8FF436C" box="[1307,1429,1867,1893]" italics="true" pageId="6" pageNumber="535">C. atremus</emphasis>
</taxonomicName>
, though overlap is probably not great as each typically occurs in more disturbed habitats.
</paragraph>
<paragraph id="E24F3694FFF1ED1CFF4FF80BEDB744DE" blockId="6.[151,1437,1362,2037]" lastBlockId="7.[151,1436,153,432]" lastPageId="7" lastPageNumber="536" pageId="6" pageNumber="535">
Among the females captured in late February, two (UMMZ 247749–50) are of adult size (
<quantity id="25089B71FFF1ED1DFB48F80BE83043A4" box="[1216,1370,1939,1965]" metricMagnitude="-1" metricUnit="m" metricValue="1.105" metricValueMax="1.1199999999999999" metricValueMin="1.09" pageId="6" pageNumber="535" unit="mm" value="110.5" valueMax="112.0" valueMin="109.0">109–112 mm</quantity>
), lack enlarged ova, but have small oocytes, suggesting that they recently laid eggs prior to capture. The other two females captured the same day (UMMZ 247748, 247751) each have two enlarged white eggs in their abdomens that were presumably near to deposition. AMS R24744 and R124604 are immature at 
<quantity id="25089B71FFF0ED1CFC79FF01E92A44BA" box="[1009,1088,153,179]" metricMagnitude="-2" metricUnit="m" metricValue="7.7" pageId="7" pageNumber="536" unit="mm" value="77.0">77 mm</quantity>
SVL and 
<quantity id="25089B71FFF0ED1CFB3AFF01E86B44BA" box="[1202,1281,153,179]" metricMagnitude="-2" metricUnit="m" metricValue="6.6" pageId="7" pageNumber="536" unit="mm" value="66.0">66 mm</quantity>
SVL, respectively.
</paragraph>
<paragraph id="E24F3694FFF0ED1CFF4FFF79EE4145B9" blockId="7.[151,1436,153,432]" pageId="7" pageNumber="536">
One specimen (UMMZ 247749) has numerous nymphs of the lungworm 
<taxonomicName id="25F04D17FFF0ED1CFB8DFF79E98744F2" baseAuthorityName="Stephens" baseAuthorityYear="1908" box="[1029,1261,225,251]" class="Maxillopoda" family="Porocephalidae" genus="Kiricephalus" kingdom="Animalia" order="Porocephalida" pageId="7" pageNumber="536" phylum="Arthropoda" rank="species" species="pattoni">
<emphasis id="D084EA86FFF0ED1CFB8DFF79E98744F2" box="[1029,1261,225,251]" italics="true" pageId="7" pageNumber="536">Kiricephalus pattoni</emphasis>
</taxonomicName>
(Pentastomida, catalogued as HWML- 
<date id="964E1054FFF0ED1CFE11FE9DEC864516" box="[409,492,261,287]" pageId="7" pageNumber="536" value="19-11-10">111019</date>
) between the skin and underlying musculature of the trunk; some of these have their heads protruding into the body cavity. This species has previously been reported in 
<collectingCountry id="9AE77604FFF0ED1CFBC7FEB1E840454A" box="[1103,1322,297,323]" name="Papua New Guinea" pageId="7" pageNumber="536">Papua New Guinea</collectingCountry>
under the name 
<taxonomicName id="25F04D17FFF0ED1CFF51FED5EC00456E" authorityName="Sambon" authorityYear="1922" box="[217,362,333,359]" class="Maxillopoda" family="Porocephalidae" genus="Kiricephalus" kingdom="Animalia" order="Porocephalida" pageId="7" pageNumber="536" phylum="Arthropoda" rank="genus">
<emphasis id="D084EA86FFF0ED1CFF51FED5EC00456E" box="[217,362,333,359]" italics="true" pageId="7" pageNumber="536">Kiricephalus</emphasis>
</taxonomicName>
sp. from the lizards 
<taxonomicName id="25F04D17FFF0ED1CFDC2FED5EE14456E" baseAuthorityName="De Vis" baseAuthorityYear="1892" box="[586,894,333,359]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="536" phylum="Chordata" rank="species" species="louisiadensis">
<emphasis id="D084EA86FFF0ED1CFDC2FED5EE14456E" box="[586,894,333,359]" italics="true" pageId="7" pageNumber="536">Cyrtodactylus louisiadensis</emphasis>
</taxonomicName>
, 
<taxonomicName id="25F04D17FFF0ED1CFC03FED5E9C3456E" baseAuthorityName="Meyer" baseAuthorityYear="1874" box="[907,1193,333,359]" class="Reptilia" family="Scincidae" genus="Sphenomorphus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="536" phylum="Chordata" rank="species" species="jobiensis">
<emphasis id="D084EA86FFF0ED1CFC03FED5E9C3456E" box="[907,1193,333,359]" italics="true" pageId="7" pageNumber="536">Sphenomorphus jobiensis</emphasis>
</taxonomicName>
, and 
<taxonomicName id="25F04D17FFF0ED1CFB6BFED5E854456E" baseAuthorityName="Sauvage" baseAuthorityYear="1879" box="[1251,1342,333,359]" class="Reptilia" family="Scincidae" genus="Sphenomorphus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="536" phylum="Chordata" rank="species" species="simus">
<emphasis id="D084EA86FFF0ED1CFB6BFED5E854456E" box="[1251,1342,333,359]" italics="true" pageId="7" pageNumber="536">S. simus</emphasis>
</taxonomicName>
(
<bibRefCitation id="86614B65FFF0ED1CFAC3FED5EC714582" author="Bursey, C. R. &amp; Goldberg, S. R. &amp; Kraus, F." pageId="7" pageNumber="536" pagination="1385 - 1394" refId="ref6947" refString="Bursey, C. R., Goldberg, S. R. &amp; Kraus, F. (2005) Endoparasites in Sphenomorphus jobiensis (Sauria: Scincidae) from Papua New Guinea with descriptions of three new species. Journal of Parasitology, 91, 1385 - 1394. https: // doi. org / 10.1645 / GE- 3502.1" type="journal article" year="2005">
Bursey 
<emphasis id="D084EA86FFF0ED1CFF1FFEEAEDBB4582" box="[151,209,369,395]" italics="true" pageId="7" pageNumber="536">et al.</emphasis>
, 2005
</bibRefCitation>
, 
<bibRefCitation id="86614B65FFF0ED1CFEA0FEE9EC1A4582" author="Bursey, C. R. &amp; Goldberg, S. R. &amp; Kraus, F." box="[296,368,369,395]" pageId="7" pageNumber="536" pagination="41 - 45" refId="ref7009" refString="Bursey, C. R., Goldberg, S. R. &amp; Kraus, F. (2008 a) A new species of Pharyngodon (Nematoda: Pharyngodonidae) and other helminths in Cyrtodactylus louisiadensis (Squamata: Gekkonidae) from Papua New Guinea. Acta Parasitologica, 53, 41 - 45. https: // doi. org / 10.2478 / s 11686 - 008 - 0005 - z" type="journal article" year="2008">2008a</bibRefCitation>
; 
<bibRefCitation id="86614B65FFF0ED1CFEF4FEE9EF174582" author="Goldberg, S. R. &amp; Bursey, C. R. &amp; Kraus, F." box="[380,637,369,395]" pageId="7" pageNumber="536" pagination="58 - 83" refId="ref7164" refString="Goldberg, S. R., Bursey, C. R. &amp; Kraus, F. (2009 a) Endoparasites in 12 species of Sphenomorphus (Squamata: Scincidae) from Papua New Guinea. Comparative Parasitology, 76, 58 - 83. https: // doi. org / 10.1654 / 4358.1" type="journal article" year="2009">
Goldberg 
<emphasis id="D084EA86FFF0ED1CFE66FEEAEF424582" box="[494,552,369,395]" italics="true" pageId="7" pageNumber="536">et al.</emphasis>
, 2009a
</bibRefCitation>
) as well as the frogs 
<taxonomicName id="25F04D17FFF0ED1CFCFBFEE9E9314582" authorityName="Boettger" authorityYear="1892" box="[883,1115,369,395]" class="Amphibia" family="Ceratobatrachidae" genus="Cornufer" higherTaxonomySource="CoL" kingdom="Animalia" order="Anura" pageId="7" pageNumber="536" phylum="Chordata" rank="species" species="boulengeri">
<emphasis id="D084EA86FFF0ED1CFCFBFEE9E9314582" box="[883,1115,369,395]" italics="true" pageId="7" pageNumber="536">Cornufer boulengeri</emphasis>
</taxonomicName>
, 
<taxonomicName id="25F04D17FFF0ED1CFBE3FEE9E9864582" baseAuthorityName="Brown &amp; Tyler" baseAuthorityYear="1968" box="[1131,1260,369,395]" class="Amphibia" family="Ceratobatrachidae" genus="Cornufer" higherTaxonomySource="CoL" kingdom="Animalia" order="Anura" pageId="7" pageNumber="536" phylum="Chordata" rank="species" species="schmidti">
<emphasis id="D084EA86FFF0ED1CFBE3FEE9E9864582" box="[1131,1260,369,395]" italics="true" pageId="7" pageNumber="536">C. schmidti</emphasis>
</taxonomicName>
, and 
<taxonomicName id="25F04D17FFF0ED1CFAA4FEEAEC7745A6" baseAuthorityName="Menzies" baseAuthorityYear="1987" class="Amphibia" family="Ranidae" genus="Papurana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="7" pageNumber="536" phylum="Chordata" rank="species" species="supragrisea">
<emphasis id="D084EA86FFF0ED1CFAA4FEEAEC7745A6" italics="true" pageId="7" pageNumber="536">Papurana supragrisea</emphasis>
</taxonomicName>
(
<bibRefCitation id="86614B65FFF0ED1CFEA4FE0DEF7945A6" author="Bursey, C. R. &amp; Goldberg, S. R. &amp; Kraus, F." box="[300,531,405,431]" pageId="7" pageNumber="536" pagination="33 - 48" refId="ref7083" refString="Bursey, C. R., Goldberg, S. R. &amp; Kraus, F. (2008 b) A new species of Proteocephalus (Cestoda: Proteocephalidae), description of the male of Desmogathiema papuensis (Nematoda: Quimperiidae), and other endoparasites in Sylvirana supragrisesa (Anura: Ranidae) from Papua New Guinea. Comparative Parasitology, 75, 33 - 48. https: // doi. org / 10.1654 / 4291.1" type="journal article" year="2008">
Bursey 
<emphasis id="D084EA86FFF0ED1CFE0CFE0EECD645A6" box="[388,444,405,431]" italics="true" pageId="7" pageNumber="536">et al.</emphasis>
, 2008b
</bibRefCitation>
; 
<bibRefCitation id="86614B65FFF0ED1CFD96FE0DEE4A45B9" author="Goldberg, S. R. &amp; Bursey, C. R. &amp; Kraus, F." box="[542,800,405,432]" pageId="7" pageNumber="536" pagination="327 - 337" refId="ref7221" refString="Goldberg, S. R., Bursey, C. R. &amp; Kraus, F. (2009 b) Endoparasites in eleven species of ranid frogs (Anura: Ranidae) from Papua New Guinea. Pacific Science, 63, 327 - 337. https: // doi. org / 10.2984 / 049.063.0303" type="journal article" year="2009">
Goldberg 
<emphasis id="D084EA86FFF0ED1CFD07FE0EEFAD45A6" box="[655,711,405,431]" italics="true" pageId="7" pageNumber="536">et al.</emphasis>
, 2009b
</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>