433 lines
66 KiB
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433 lines
66 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/jhr.84.68709" ID-GBIF-Dataset="b9e61bc3-4158-4bee-8d57-a65655588f70" ID-Pensoft-Pub="1314-2607-84-145" ID-Pensoft-UUID="AEDB6967BB17574BB6988187C56EE6CA" ID-Zenodo-Dep="5349694" ID-ZooBank="5F83EC637C7947FDBD29E7A2B9328518" ModsDocID="1314-2607-84-145" checkinTime="1629847073322" checkinUser="pensoft" docAuthor="Okayasu, Juriya" docDate="2021" docId="6A19D29C19245A50A33A7EB56087CE48" docLanguage="en" docName="JourHymenoptRes 84: 145-168" docOrigin="Journal of Hymenoptera Research 84" docPubDate="2021-08-24" docSource="http://dx.doi.org/10.3897/jhr.84.68709" docTitle="Arkaditilla Okayasu 2021, gen. nov." docType="treatment" docUuid="A07954E6-26EB-4494-860C-51F96F27D2C1" docUuidSource="ZooBank" docVersion="4" id="AEDB6967BB17574BB6988187C56EE6CA" lastPageNumber="145" masterDocId="AEDB6967BB17574BB6988187C56EE6CA" masterDocTitle="A new velvet ant genus Arkaditilla (Hymenoptera, Mutillidae, Trogaspidiini) from the Oriental Region, with review of species" masterLastPageNumber="168" masterPageNumber="145" pageNumber="145" updateTime="1678756233000" updateUser="pensoft">
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<mods:titleInfo>
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<mods:title>A new velvet ant genus Arkaditilla (Hymenoptera, Mutillidae, Trogaspidiini) from the Oriental Region, with review of species</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Okayasu, Juriya</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-6307-2939</mods:nameIdentifier>
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<mods:affiliation>Entomological Laboratory, Faculty of Agriculture, Ehime University, Tarumi 3 - 5 - 7, Matsuyama, 790 - 8566, Japan & Systematic Entomology, Graduate School of Agriculture, Hokkaido University, Sapporo, 060 - 8589, Japan</mods:affiliation>
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<mods:nameIdentifier type="email">mutiphiidae@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>Journal of Hymenoptera Research</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2021</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2021-08-24</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>84</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>145</mods:start>
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<mods:end>168</mods:end>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/jhr.84.68709</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/jhr.84.68709</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1314-2607-84-145</mods:identifier>
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<mods:identifier type="ZooBank">5F83EC637C7947FDBD29E7A2B9328518</mods:identifier>
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<mods:identifier type="Zenodo-Dep">5349694</mods:identifier>
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<treatment ID-GBIF-Taxon="184941292" LSID="urn:lsid:zoobank.org:act:A07954E6-26EB-4494-860C-51F96F27D2C1" httpUri="http://treatment.plazi.org/id/6A19D29C19245A50A33A7EB56087CE48" lastPageNumber="145" pageId="0" pageNumber="145">
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<subSubSection pageId="0" pageNumber="145" type="nomenclature">
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<paragraph pageId="0" pageNumber="145">
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<taxonomicName LSID="http://zoobank.org/A07954E6-26EB-4494-860C-51F96F27D2C1" authority="Okayasu, 2021" authorityName="Okayasu" authorityYear="2021" class="Insecta" family="Mutillidae" genus="Arkaditilla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Arkaditilla" order="Hymenoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus" status="gen. nov.">Arkaditilla</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="145">gen. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="145" type="description">
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<paragraph pageId="0" pageNumber="145">
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<figureCitation captionStart="Figures 1–6" captionStartId="F1" captionText="Figures 1 - 6. Arkaditilla spp., ♂, habitus 1, 3, 5 dorsal view 2, 4, 6 lateral view 1, 2 A. bagrada (Cameron) 3, 4 A. depressicornis (Mickel) 5, 6 A. frim sp. nov., holotype. Scale bars: 3 mm." figureDoi="10.3897/jhr.84.68709.figures1-6" httpUri="https://binary.pensoft.net/fig/579480" pageId="0" pageNumber="145">Figures 1-6</figureCitation>
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<figureCitation captionStart="Figures 7–12" captionStartId="F2" captionText="Figures 7 - 12. Arkaditilla spp., ♂, habitus 7, 9, 11 dorsal view 8, 10, 12 lateral view 7, 8 A. gibba sp. nov., holotype 9, 10 A. leleji sp. nov., holotype 11, 12 A. nallinia (Zavattari). Scale bars: 3 mm." figureDoi="10.3897/jhr.84.68709.figures7-12" httpUri="https://binary.pensoft.net/fig/579481" pageId="0" pageNumber="145">, 7-12</figureCitation>
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<figureCitation captionStart="Figures 13–18" captionStartId="F3" captionText="Figures 13 - 18. Arkaditilla spp., ♂, face 13 A. bagrada (Cameron) 14 A. depressicornis (Mickel) 15 A. frim sp. nov., holotype 16 A. gibba sp. nov., paratype 17 A. leleji sp. nov., holotype 18 A. nallinia (Zavattari)." figureDoi="10.3897/jhr.84.68709.figures13-18" httpUri="https://binary.pensoft.net/fig/579482" pageId="0" pageNumber="145">, 13-18</figureCitation>
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<figureCitation captionStart="Figures 19–36" captionStartId="F4" captionText="Figures 19 - 36. Arkaditilla spp., ♂ 19 - 24 mandible, lateral view 25, 27, 29, 31, 33, 35 scape, ventral view 26, 28, 30, 32, 34, 36 pedicel, F 1, and F 2, lateral view 19, 25, 26 A. bagrada (Cameron) 20, 27, 28 A. depressicornis (Mickel) 21, 29, 30 A. frim sp. nov., holotype 22, 31, 32 A. gibba sp. nov., holotype 23, 33, 34 A. leleji sp. nov., holotype 24, 35, 36 A. nallinia (Zavattari)." figureDoi="10.3897/jhr.84.68709.figures19-36" httpUri="https://binary.pensoft.net/fig/579483" pageId="0" pageNumber="145">, 19-36</figureCitation>
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<figureCitation captionStart="Figures 37–48" captionStartId="F5" captionText="Figures 37 - 48. Arkaditilla spp., ♂, genitalia 37, 39, 41, 43, 45, 47 dorsal view 38, 40, 42, 44, 46, 48 ventral view 37, 38 A. bagrada (Cameron) 39, 40 A. depressicornis (Mickel) 41, 42 A. frim sp. nov., holotype 43, 44 A. gibba sp. nov., holotype 45, 46 A. leleji sp. nov., holotype 47, 48 A. nallinia (Zavattari)." figureDoi="10.3897/jhr.84.68709.figures37-48" httpUri="https://binary.pensoft.net/fig/579484" pageId="0" pageNumber="145">, 37-48</figureCitation>
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<figureCitation captionStart="Figures 49–66" captionStartId="F6" captionText="Figures 49 - 66. Arkaditilla spp., ♂, genitalia 49, 52, 55, 58, 61, 64 genital capsule and volsella, inner view 50, 53, 56, 59, 62, 65 left penis valve 51, 54, 57, 60, 63, 66 right penis valve 49 - 51 A. bagrada (Cameron) 52 - 54 A. depressicornis (Mickel) 55 - 57 A. frim sp. nov., holotype 58 - 60 A. gibba sp. nov., holotype 61 - 63 A. leleji sp. nov., holotype 64 - 66 A. nallinia (Zavattari)." figureDoi="10.3897/jhr.84.68709.figures49-66" httpUri="https://binary.pensoft.net/fig/579485" pageId="0" pageNumber="145">, 49-66</figureCitation>
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<figureCitation captionStart="Figures 67–72" captionStartId="F7" captionText="Figures 67 - 72. Arkaditilla spp., ♂, genital volsella, ventro-inner view 67 A. bagrada (Cameron) 68 A. depressicornis (Mickel) 69 A. frim sp. nov., holotype 70 A. gibba sp. nov., holotype 71 A. leleji sp. nov., holotype 72 A. nallinia (Zavattari). White arrows indicate the paracuspis." figureDoi="10.3897/jhr.84.68709.figures67-72" httpUri="https://binary.pensoft.net/fig/579486" pageId="0" pageNumber="145">, 67-72</figureCitation>
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<figureCitation captionStart="Figure 73–76" captionStartId="F8" captionText="Figure 73 - 76. Distributional map of Arkaditilla spp. based on specimens examined herein and the literature (Cameron 1902; Lelej 1996 b; Mickel 1935, 1937; Zavattari 1914) 73 A. bagrada (Cameron) (red circle) 74 A. depressicornis (Mickel) (blue circle) 75 A. frim sp. nov. (yellow circle), A. gibba sp. nov. (purple triangle), and A. leleji sp. nov. (light blue square) 76 A. nallinia (Zavattari) (orange circle)." figureDoi="10.3897/jhr.84.68709.figure73-76" httpUri="https://binary.pensoft.net/fig/579487" pageId="0" pageNumber="145">, 73-76</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="145" type="reference_group">
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<paragraph pageId="0" pageNumber="145">
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<taxonomicName authority="Okayasu, 2021" authorityName="Okayasu" authorityYear="2021" class="Insecta" family="Mutillidae" genus="Mutilla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mutilla" order="Hymenoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus" status="gen. nov.">Mutilla</taxonomicName>
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(in part):
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<bibRefCitation author="Cameron, P" journalOrPublisher="Journal of the Straits Branch of the Royal Asiatic Society" pageId="0" pageNumber="145" pagination="29 - 131" refId="B2" refString="Cameron, P, 1902. On the Hymenoptera collected by Mr. Robert Shelford at Sarawak, and on the Hymenoptera of the Sarawak Museum. Journal of the Straits Branch of the Royal Asiatic Society 37: 29 - 131" title="On the Hymenoptera collected by Mr. Robert Shelford at Sarawak, and on the Hymenoptera of the Sarawak Museum." volume="37" year="1902">Cameron 1902</bibRefCitation>
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: 76, ♂;
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<bibRefCitation author="Zavattari, E" journalOrPublisher="Bollettino della Societa Entomologica Italiana" pageId="0" pageNumber="145" pagination="61 - 114" refId="B26" refString="Zavattari, E, 1914. Mutille Austro-Malesi. Bollettino della Societa Entomologica Italiana 45: 61 - 114" title="Mutille Austro-Malesi." volume="45" year="1914">Zavattari 1914</bibRefCitation>
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: 100, ♂.
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</paragraph>
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<paragraph pageId="0" pageNumber="145">
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<taxonomicName authority=")" authorityName="Okayasu" authorityYear="2021" class="Insecta" family="Mutillidae" genus="Timulla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Timulla (Trogaspidia)" order="Hymenoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="subGenus" status="gen. nov." subGenus="Trogaspidia">Timulla (Trogaspidia)</taxonomicName>
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(in part):
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<bibRefCitation DOI="https://doi.org/10.1111/j.1365-2311.1935.tb01207.x" author="Mickel, CE" journalOrPublisher="Transactions of the Royal Entomological Society of London" pageId="0" pageNumber="145" pagination="177 - 312" refId="B20" refString="Mickel, CE, 1935. The mutillid wasps of the islands of the Pacific Ocean (Hymenoptera, Mutillidae). Transactions of the Royal Entomological Society of London 83 (2): 177 - 312, DOI: https://doi.org/10.1111/j.1365-2311.1935.tb01207.x" title="The mutillid wasps of the islands of the Pacific Ocean (Hymenoptera, Mutillidae)." url="https://doi.org/10.1111/j.1365-2311.1935.tb01207.x" volume="83" year="1935">Mickel 1935</bibRefCitation>
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: 263, 264, ♂; 1937: 449, ♂.
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</paragraph>
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<paragraph pageId="0" pageNumber="145">
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<taxonomicName authority="Okayasu, 2021" authorityName="Okayasu" authorityYear="2021" class="Insecta" family="Mutillidae" genus="Krombeinidia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Krombeinidia" order="Hymenoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus" status="gen. nov.">Krombeinidia</taxonomicName>
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(in part):
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<bibRefCitation author="Lelej, AS" journalOrPublisher="Far Eastern Entomologist" pageId="0" pageNumber="145" pagination="1 - 24" refId="B10" refString="Lelej, AS, 1996a. To the knowledge of the East Asian species of the tribe Trogaspidiini Bischoff, 1920 (Hymenoptera, Mutillidae) with description of eight new genera and two new species. Far Eastern Entomologist 30: 1 - 24" title="To the knowledge of the East Asian species of the tribe Trogaspidiini Bischoff, 1920 (Hymenoptera, Mutillidae) with description of eight new genera and two new species." volume="30" year="1996 a">Lelej 1996a</bibRefCitation>
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: 11, ♂.
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</paragraph>
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<paragraph pageId="0" pageNumber="145">
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<taxonomicName authority="Okayasu, 2021" authorityName="Okayasu" authorityYear="2021" class="Insecta" family="Mutillidae" genus="Petersenidia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Petersenidia" order="Hymenoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus" status="gen. nov.">Petersenidia</taxonomicName>
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(in part):
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<bibRefCitation DOI="https://doi.org/10.3759/tropics.6.91" author="Lelej, AS" journalOrPublisher="Tropics" pageId="0" pageNumber="145" pagination="91 - 104" refId="B11" refString="Lelej, AS, 1996b. Mutillid wasps collected in Malaysia and Indonesia by Dr. Sk. Yamane (Hymenoptera, Mutillidae). Tropics 6 (1-2): 91 - 104, DOI: https://doi.org/10.3759/tropics.6.91" title="Mutillid wasps collected in Malaysia and Indonesia by Dr. Sk. Yamane (Hymenoptera, Mutillidae)." url="https://doi.org/10.3759/tropics.6.91" volume="6" year="1996 b">Lelej 1996b</bibRefCitation>
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: 94, 95, ♂.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="145" type="diagnosis">
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<paragraph pageId="0" pageNumber="145">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="145">
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<emphasis bold="true" pageId="0" pageNumber="145">Male.</emphasis>
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Frons punctate; vertex roundly sloping posteriorly; postgenal bridge flattened; hypostomal carina simple, without denticle; clypeus dorsally elevated; prementum without tubercle. Mandible apically bidentate, ventrally excised into basal tooth (Figs
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<figureCitation captionStart="Figures 19–36" captionStartId="F4" captionText="Figures 19 - 36. Arkaditilla spp., ♂ 19 - 24 mandible, lateral view 25, 27, 29, 31, 33, 35 scape, ventral view 26, 28, 30, 32, 34, 36 pedicel, F 1, and F 2, lateral view 19, 25, 26 A. bagrada (Cameron) 20, 27, 28 A. depressicornis (Mickel) 21, 29, 30 A. frim sp. nov., holotype 22, 31, 32 A. gibba sp. nov., holotype 23, 33, 34 A. leleji sp. nov., holotype 24, 35, 36 A. nallinia (Zavattari)." figureDoi="10.3897/jhr.84.68709.figures19-36" httpUri="https://binary.pensoft.net/fig/579483" pageId="0" pageNumber="145">19-24</figureCitation>
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); inner margin expanded, forming subbasal tooth (Figs
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<figureCitation captionStart="Figures 13–18" captionStartId="F3" captionText="Figures 13 - 18. Arkaditilla spp., ♂, face 13 A. bagrada (Cameron) 14 A. depressicornis (Mickel) 15 A. frim sp. nov., holotype 16 A. gibba sp. nov., paratype 17 A. leleji sp. nov., holotype 18 A. nallinia (Zavattari)." figureDoi="10.3897/jhr.84.68709.figures13-18" httpUri="https://binary.pensoft.net/fig/579482" pageId="0" pageNumber="145">13-18</figureCitation>
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); dorso-lateral carina sharp, not crooked. Scape ventrally with two longitudinal carinae, densely punctate or mostly smooth with sparse punctures apically between carinae (Figs
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<figureCitation captionStart="Figures 19–36" captionStartId="F4" captionText="Figures 19 - 36. Arkaditilla spp., ♂ 19 - 24 mandible, lateral view 25, 27, 29, 31, 33, 35 scape, ventral view 26, 28, 30, 32, 34, 36 pedicel, F 1, and F 2, lateral view 19, 25, 26 A. bagrada (Cameron) 20, 27, 28 A. depressicornis (Mickel) 21, 29, 30 A. frim sp. nov., holotype 22, 31, 32 A. gibba sp. nov., holotype 23, 33, 34 A. leleji sp. nov., holotype 24, 35, 36 A. nallinia (Zavattari)." figureDoi="10.3897/jhr.84.68709.figures19-36" httpUri="https://binary.pensoft.net/fig/579483" pageId="0" pageNumber="145">25</figureCitation>
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,
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<figureCitation captionStart="Figures 19–36" captionStartId="F4" captionText="Figures 19 - 36. Arkaditilla spp., ♂ 19 - 24 mandible, lateral view 25, 27, 29, 31, 33, 35 scape, ventral view 26, 28, 30, 32, 34, 36 pedicel, F 1, and F 2, lateral view 19, 25, 26 A. bagrada (Cameron) 20, 27, 28 A. depressicornis (Mickel) 21, 29, 30 A. frim sp. nov., holotype 22, 31, 32 A. gibba sp. nov., holotype 23, 33, 34 A. leleji sp. nov., holotype 24, 35, 36 A. nallinia (Zavattari)." figureDoi="10.3897/jhr.84.68709.figures19-36" httpUri="https://binary.pensoft.net/fig/579483" pageId="0" pageNumber="145">27</figureCitation>
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,
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<figureCitation captionStart="Figures 19–36" captionStartId="F4" captionText="Figures 19 - 36. Arkaditilla spp., ♂ 19 - 24 mandible, lateral view 25, 27, 29, 31, 33, 35 scape, ventral view 26, 28, 30, 32, 34, 36 pedicel, F 1, and F 2, lateral view 19, 25, 26 A. bagrada (Cameron) 20, 27, 28 A. depressicornis (Mickel) 21, 29, 30 A. frim sp. nov., holotype 22, 31, 32 A. gibba sp. nov., holotype 23, 33, 34 A. leleji sp. nov., holotype 24, 35, 36 A. nallinia (Zavattari)." figureDoi="10.3897/jhr.84.68709.figures19-36" httpUri="https://binary.pensoft.net/fig/579483" pageId="0" pageNumber="145">29</figureCitation>
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,
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<figureCitation captionStart="Figures 19–36" captionStartId="F4" captionText="Figures 19 - 36. Arkaditilla spp., ♂ 19 - 24 mandible, lateral view 25, 27, 29, 31, 33, 35 scape, ventral view 26, 28, 30, 32, 34, 36 pedicel, F 1, and F 2, lateral view 19, 25, 26 A. bagrada (Cameron) 20, 27, 28 A. depressicornis (Mickel) 21, 29, 30 A. frim sp. nov., holotype 22, 31, 32 A. gibba sp. nov., holotype 23, 33, 34 A. leleji sp. nov., holotype 24, 35, 36 A. nallinia (Zavattari)." figureDoi="10.3897/jhr.84.68709.figures19-36" httpUri="https://binary.pensoft.net/fig/579483" pageId="0" pageNumber="145">31</figureCitation>
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,
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<figureCitation captionStart="Figures 19–36" captionStartId="F4" captionText="Figures 19 - 36. Arkaditilla spp., ♂ 19 - 24 mandible, lateral view 25, 27, 29, 31, 33, 35 scape, ventral view 26, 28, 30, 32, 34, 36 pedicel, F 1, and F 2, lateral view 19, 25, 26 A. bagrada (Cameron) 20, 27, 28 A. depressicornis (Mickel) 21, 29, 30 A. frim sp. nov., holotype 22, 31, 32 A. gibba sp. nov., holotype 23, 33, 34 A. leleji sp. nov., holotype 24, 35, 36 A. nallinia (Zavattari)." figureDoi="10.3897/jhr.84.68709.figures19-36" httpUri="https://binary.pensoft.net/fig/579483" pageId="0" pageNumber="145">33</figureCitation>
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,
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<figureCitation captionStart="Figures 19–36" captionStartId="F4" captionText="Figures 19 - 36. Arkaditilla spp., ♂ 19 - 24 mandible, lateral view 25, 27, 29, 31, 33, 35 scape, ventral view 26, 28, 30, 32, 34, 36 pedicel, F 1, and F 2, lateral view 19, 25, 26 A. bagrada (Cameron) 20, 27, 28 A. depressicornis (Mickel) 21, 29, 30 A. frim sp. nov., holotype 22, 31, 32 A. gibba sp. nov., holotype 23, 33, 34 A. leleji sp. nov., holotype 24, 35, 36 A. nallinia (Zavattari)." figureDoi="10.3897/jhr.84.68709.figures19-36" httpUri="https://binary.pensoft.net/fig/579483" pageId="0" pageNumber="145">35</figureCitation>
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); flagellum with yellow or reddish brown ventral markings; F1 longer than F2, depressed (Figs
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<figureCitation captionStart="Figures 19–36" captionStartId="F4" captionText="Figures 19 - 36. Arkaditilla spp., ♂ 19 - 24 mandible, lateral view 25, 27, 29, 31, 33, 35 scape, ventral view 26, 28, 30, 32, 34, 36 pedicel, F 1, and F 2, lateral view 19, 25, 26 A. bagrada (Cameron) 20, 27, 28 A. depressicornis (Mickel) 21, 29, 30 A. frim sp. nov., holotype 22, 31, 32 A. gibba sp. nov., holotype 23, 33, 34 A. leleji sp. nov., holotype 24, 35, 36 A. nallinia (Zavattari)." figureDoi="10.3897/jhr.84.68709.figures19-36" httpUri="https://binary.pensoft.net/fig/579483" pageId="0" pageNumber="145">28</figureCitation>
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,
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<figureCitation captionStart="Figures 19–36" captionStartId="F4" captionText="Figures 19 - 36. Arkaditilla spp., ♂ 19 - 24 mandible, lateral view 25, 27, 29, 31, 33, 35 scape, ventral view 26, 28, 30, 32, 34, 36 pedicel, F 1, and F 2, lateral view 19, 25, 26 A. bagrada (Cameron) 20, 27, 28 A. depressicornis (Mickel) 21, 29, 30 A. frim sp. nov., holotype 22, 31, 32 A. gibba sp. nov., holotype 23, 33, 34 A. leleji sp. nov., holotype 24, 35, 36 A. nallinia (Zavattari)." figureDoi="10.3897/jhr.84.68709.figures19-36" httpUri="https://binary.pensoft.net/fig/579483" pageId="0" pageNumber="145">30</figureCitation>
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,
|
||
<figureCitation captionStart="Figures 19–36" captionStartId="F4" captionText="Figures 19 - 36. Arkaditilla spp., ♂ 19 - 24 mandible, lateral view 25, 27, 29, 31, 33, 35 scape, ventral view 26, 28, 30, 32, 34, 36 pedicel, F 1, and F 2, lateral view 19, 25, 26 A. bagrada (Cameron) 20, 27, 28 A. depressicornis (Mickel) 21, 29, 30 A. frim sp. nov., holotype 22, 31, 32 A. gibba sp. nov., holotype 23, 33, 34 A. leleji sp. nov., holotype 24, 35, 36 A. nallinia (Zavattari)." figureDoi="10.3897/jhr.84.68709.figures19-36" httpUri="https://binary.pensoft.net/fig/579483" pageId="0" pageNumber="145">32</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figures 19–36" captionStartId="F4" captionText="Figures 19 - 36. Arkaditilla spp., ♂ 19 - 24 mandible, lateral view 25, 27, 29, 31, 33, 35 scape, ventral view 26, 28, 30, 32, 34, 36 pedicel, F 1, and F 2, lateral view 19, 25, 26 A. bagrada (Cameron) 20, 27, 28 A. depressicornis (Mickel) 21, 29, 30 A. frim sp. nov., holotype 22, 31, 32 A. gibba sp. nov., holotype 23, 33, 34 A. leleji sp. nov., holotype 24, 35, 36 A. nallinia (Zavattari)." figureDoi="10.3897/jhr.84.68709.figures19-36" httpUri="https://binary.pensoft.net/fig/579483" pageId="0" pageNumber="145">34</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figures 19–36" captionStartId="F4" captionText="Figures 19 - 36. Arkaditilla spp., ♂ 19 - 24 mandible, lateral view 25, 27, 29, 31, 33, 35 scape, ventral view 26, 28, 30, 32, 34, 36 pedicel, F 1, and F 2, lateral view 19, 25, 26 A. bagrada (Cameron) 20, 27, 28 A. depressicornis (Mickel) 21, 29, 30 A. frim sp. nov., holotype 22, 31, 32 A. gibba sp. nov., holotype 23, 33, 34 A. leleji sp. nov., holotype 24, 35, 36 A. nallinia (Zavattari)." figureDoi="10.3897/jhr.84.68709.figures19-36" httpUri="https://binary.pensoft.net/fig/579483" pageId="0" pageNumber="145">36</figureCitation>
|
||
) except
|
||
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="A. bagrada" order="Hymenoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="bagrada">
|
||
<emphasis italics="true" pageId="0" pageNumber="145">A. bagrada</emphasis>
|
||
</taxonomicName>
|
||
with F1 cylindrical (Fig.
|
||
<figureCitation captionStart="Figures 19–36" captionStartId="F4" captionText="Figures 19 - 36. Arkaditilla spp., ♂ 19 - 24 mandible, lateral view 25, 27, 29, 31, 33, 35 scape, ventral view 26, 28, 30, 32, 34, 36 pedicel, F 1, and F 2, lateral view 19, 25, 26 A. bagrada (Cameron) 20, 27, 28 A. depressicornis (Mickel) 21, 29, 30 A. frim sp. nov., holotype 22, 31, 32 A. gibba sp. nov., holotype 23, 33, 34 A. leleji sp. nov., holotype 24, 35, 36 A. nallinia (Zavattari)." figureDoi="10.3897/jhr.84.68709.figures19-36" httpUri="https://binary.pensoft.net/fig/579483" pageId="0" pageNumber="145">26</figureCitation>
|
||
). Mesosoma black; notaulus present, reaching anterior 1/3 to 1/2 of mesoscutum; parascutal carina forming weakly elevated, rounded tubercle posteriorly; tegula posterior margin slightly protruding over transscutal articulation, not recurved; mesoscutellum evenly convex, puncto-reticulate; mesopleuron divided into dorsal and ventral convexities by transverse groove; propodeum gradually sloping posteriorly, without sublateral longitudinal carina; propodeal dorso-lateral margin without denticles. Pterostigma length subequal to distance between origin of RS on vein SC and proximal base of pterostigma. Protibial spur slender; mesocoxa evenly convex; metacoxa evenly convex with long sparse recumbent setae, without tubercle on inner margin. Metasoma black with segments 1-2 or 3 orange; S2 swollen anteriorly, without lateral felt line (Figs
|
||
<figureCitation captionStart="Figures 1–6" captionStartId="F1" captionText="Figures 1 - 6. Arkaditilla spp., ♂, habitus 1, 3, 5 dorsal view 2, 4, 6 lateral view 1, 2 A. bagrada (Cameron) 3, 4 A. depressicornis (Mickel) 5, 6 A. frim sp. nov., holotype. Scale bars: 3 mm." figureDoi="10.3897/jhr.84.68709.figures1-6" httpUri="https://binary.pensoft.net/fig/579480" pageId="0" pageNumber="145">2</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figures 1–6" captionStartId="F1" captionText="Figures 1 - 6. Arkaditilla spp., ♂, habitus 1, 3, 5 dorsal view 2, 4, 6 lateral view 1, 2 A. bagrada (Cameron) 3, 4 A. depressicornis (Mickel) 5, 6 A. frim sp. nov., holotype. Scale bars: 3 mm." figureDoi="10.3897/jhr.84.68709.figures1-6" httpUri="https://binary.pensoft.net/fig/579480" pageId="0" pageNumber="145">4</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figures 1–6" captionStartId="F1" captionText="Figures 1 - 6. Arkaditilla spp., ♂, habitus 1, 3, 5 dorsal view 2, 4, 6 lateral view 1, 2 A. bagrada (Cameron) 3, 4 A. depressicornis (Mickel) 5, 6 A. frim sp. nov., holotype. Scale bars: 3 mm." figureDoi="10.3897/jhr.84.68709.figures1-6" httpUri="https://binary.pensoft.net/fig/579480" pageId="0" pageNumber="145">6</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figures 7–12" captionStartId="F2" captionText="Figures 7 - 12. Arkaditilla spp., ♂, habitus 7, 9, 11 dorsal view 8, 10, 12 lateral view 7, 8 A. gibba sp. nov., holotype 9, 10 A. leleji sp. nov., holotype 11, 12 A. nallinia (Zavattari). Scale bars: 3 mm." figureDoi="10.3897/jhr.84.68709.figures7-12" httpUri="https://binary.pensoft.net/fig/579481" pageId="0" pageNumber="145">8</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figures 7–12" captionStartId="F2" captionText="Figures 7 - 12. Arkaditilla spp., ♂, habitus 7, 9, 11 dorsal view 8, 10, 12 lateral view 7, 8 A. gibba sp. nov., holotype 9, 10 A. leleji sp. nov., holotype 11, 12 A. nallinia (Zavattari). Scale bars: 3 mm." figureDoi="10.3897/jhr.84.68709.figures7-12" httpUri="https://binary.pensoft.net/fig/579481" pageId="0" pageNumber="145">10</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figures 7–12" captionStartId="F2" captionText="Figures 7 - 12. Arkaditilla spp., ♂, habitus 7, 9, 11 dorsal view 8, 10, 12 lateral view 7, 8 A. gibba sp. nov., holotype 9, 10 A. leleji sp. nov., holotype 11, 12 A. nallinia (Zavattari). Scale bars: 3 mm." figureDoi="10.3897/jhr.84.68709.figures7-12" httpUri="https://binary.pensoft.net/fig/579481" pageId="0" pageNumber="145">12</figureCitation>
|
||
); T7 medially with slender longitudinal smooth convex line; S6-S8 without lateral carina; S7 entirely exposed, 0.75
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
as long as S8; S8 mostly sclerotized with posterior membranous area evenly convex posteriorly; left and right basiparameres ventro-medial margins connected to form postero-medial tongue-like tubercle (Figs
|
||
<figureCitation captionStart="Figures 37–48" captionStartId="F5" captionText="Figures 37 - 48. Arkaditilla spp., ♂, genitalia 37, 39, 41, 43, 45, 47 dorsal view 38, 40, 42, 44, 46, 48 ventral view 37, 38 A. bagrada (Cameron) 39, 40 A. depressicornis (Mickel) 41, 42 A. frim sp. nov., holotype 43, 44 A. gibba sp. nov., holotype 45, 46 A. leleji sp. nov., holotype 47, 48 A. nallinia (Zavattari)." figureDoi="10.3897/jhr.84.68709.figures37-48" httpUri="https://binary.pensoft.net/fig/579484" pageId="0" pageNumber="145">38</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figures 37–48" captionStartId="F5" captionText="Figures 37 - 48. Arkaditilla spp., ♂, genitalia 37, 39, 41, 43, 45, 47 dorsal view 38, 40, 42, 44, 46, 48 ventral view 37, 38 A. bagrada (Cameron) 39, 40 A. depressicornis (Mickel) 41, 42 A. frim sp. nov., holotype 43, 44 A. gibba sp. nov., holotype 45, 46 A. leleji sp. nov., holotype 47, 48 A. nallinia (Zavattari)." figureDoi="10.3897/jhr.84.68709.figures37-48" httpUri="https://binary.pensoft.net/fig/579484" pageId="0" pageNumber="145">40</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figures 37–48" captionStartId="F5" captionText="Figures 37 - 48. Arkaditilla spp., ♂, genitalia 37, 39, 41, 43, 45, 47 dorsal view 38, 40, 42, 44, 46, 48 ventral view 37, 38 A. bagrada (Cameron) 39, 40 A. depressicornis (Mickel) 41, 42 A. frim sp. nov., holotype 43, 44 A. gibba sp. nov., holotype 45, 46 A. leleji sp. nov., holotype 47, 48 A. nallinia (Zavattari)." figureDoi="10.3897/jhr.84.68709.figures37-48" httpUri="https://binary.pensoft.net/fig/579484" pageId="0" pageNumber="145">42</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figures 37–48" captionStartId="F5" captionText="Figures 37 - 48. Arkaditilla spp., ♂, genitalia 37, 39, 41, 43, 45, 47 dorsal view 38, 40, 42, 44, 46, 48 ventral view 37, 38 A. bagrada (Cameron) 39, 40 A. depressicornis (Mickel) 41, 42 A. frim sp. nov., holotype 43, 44 A. gibba sp. nov., holotype 45, 46 A. leleji sp. nov., holotype 47, 48 A. nallinia (Zavattari)." figureDoi="10.3897/jhr.84.68709.figures37-48" httpUri="https://binary.pensoft.net/fig/579484" pageId="0" pageNumber="145">46</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figures 37–48" captionStartId="F5" captionText="Figures 37 - 48. Arkaditilla spp., ♂, genitalia 37, 39, 41, 43, 45, 47 dorsal view 38, 40, 42, 44, 46, 48 ventral view 37, 38 A. bagrada (Cameron) 39, 40 A. depressicornis (Mickel) 41, 42 A. frim sp. nov., holotype 43, 44 A. gibba sp. nov., holotype 45, 46 A. leleji sp. nov., holotype 47, 48 A. nallinia (Zavattari)." figureDoi="10.3897/jhr.84.68709.figures37-48" httpUri="https://binary.pensoft.net/fig/579484" pageId="0" pageNumber="145">48</figureCitation>
|
||
); parapenial lobe spatulate, not flexed; ventral portion of basiparamere strongly projecting posteriorly to connect volsella (Figs
|
||
<figureCitation captionStart="Figures 49–66" captionStartId="F6" captionText="Figures 49 - 66. Arkaditilla spp., ♂, genitalia 49, 52, 55, 58, 61, 64 genital capsule and volsella, inner view 50, 53, 56, 59, 62, 65 left penis valve 51, 54, 57, 60, 63, 66 right penis valve 49 - 51 A. bagrada (Cameron) 52 - 54 A. depressicornis (Mickel) 55 - 57 A. frim sp. nov., holotype 58 - 60 A. gibba sp. nov., holotype 61 - 63 A. leleji sp. nov., holotype 64 - 66 A. nallinia (Zavattari)." figureDoi="10.3897/jhr.84.68709.figures49-66" httpUri="https://binary.pensoft.net/fig/579485" pageId="0" pageNumber="145">49</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figures 49–66" captionStartId="F6" captionText="Figures 49 - 66. Arkaditilla spp., ♂, genitalia 49, 52, 55, 58, 61, 64 genital capsule and volsella, inner view 50, 53, 56, 59, 62, 65 left penis valve 51, 54, 57, 60, 63, 66 right penis valve 49 - 51 A. bagrada (Cameron) 52 - 54 A. depressicornis (Mickel) 55 - 57 A. frim sp. nov., holotype 58 - 60 A. gibba sp. nov., holotype 61 - 63 A. leleji sp. nov., holotype 64 - 66 A. nallinia (Zavattari)." figureDoi="10.3897/jhr.84.68709.figures49-66" httpUri="https://binary.pensoft.net/fig/579485" pageId="0" pageNumber="145">52</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figures 49–66" captionStartId="F6" captionText="Figures 49 - 66. Arkaditilla spp., ♂, genitalia 49, 52, 55, 58, 61, 64 genital capsule and volsella, inner view 50, 53, 56, 59, 62, 65 left penis valve 51, 54, 57, 60, 63, 66 right penis valve 49 - 51 A. bagrada (Cameron) 52 - 54 A. depressicornis (Mickel) 55 - 57 A. frim sp. nov., holotype 58 - 60 A. gibba sp. nov., holotype 61 - 63 A. leleji sp. nov., holotype 64 - 66 A. nallinia (Zavattari)." figureDoi="10.3897/jhr.84.68709.figures49-66" httpUri="https://binary.pensoft.net/fig/579485" pageId="0" pageNumber="145">55</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figures 49–66" captionStartId="F6" captionText="Figures 49 - 66. Arkaditilla spp., ♂, genitalia 49, 52, 55, 58, 61, 64 genital capsule and volsella, inner view 50, 53, 56, 59, 62, 65 left penis valve 51, 54, 57, 60, 63, 66 right penis valve 49 - 51 A. bagrada (Cameron) 52 - 54 A. depressicornis (Mickel) 55 - 57 A. frim sp. nov., holotype 58 - 60 A. gibba sp. nov., holotype 61 - 63 A. leleji sp. nov., holotype 64 - 66 A. nallinia (Zavattari)." figureDoi="10.3897/jhr.84.68709.figures49-66" httpUri="https://binary.pensoft.net/fig/579485" pageId="0" pageNumber="145">58</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figures 49–66" captionStartId="F6" captionText="Figures 49 - 66. Arkaditilla spp., ♂, genitalia 49, 52, 55, 58, 61, 64 genital capsule and volsella, inner view 50, 53, 56, 59, 62, 65 left penis valve 51, 54, 57, 60, 63, 66 right penis valve 49 - 51 A. bagrada (Cameron) 52 - 54 A. depressicornis (Mickel) 55 - 57 A. frim sp. nov., holotype 58 - 60 A. gibba sp. nov., holotype 61 - 63 A. leleji sp. nov., holotype 64 - 66 A. nallinia (Zavattari)." figureDoi="10.3897/jhr.84.68709.figures49-66" httpUri="https://binary.pensoft.net/fig/579485" pageId="0" pageNumber="145">61</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figures 49–66" captionStartId="F6" captionText="Figures 49 - 66. Arkaditilla spp., ♂, genitalia 49, 52, 55, 58, 61, 64 genital capsule and volsella, inner view 50, 53, 56, 59, 62, 65 left penis valve 51, 54, 57, 60, 63, 66 right penis valve 49 - 51 A. bagrada (Cameron) 52 - 54 A. depressicornis (Mickel) 55 - 57 A. frim sp. nov., holotype 58 - 60 A. gibba sp. nov., holotype 61 - 63 A. leleji sp. nov., holotype 64 - 66 A. nallinia (Zavattari)." figureDoi="10.3897/jhr.84.68709.figures49-66" httpUri="https://binary.pensoft.net/fig/579485" pageId="0" pageNumber="145">64</figureCitation>
|
||
); paramere antero-ventrally serrate (Figs
|
||
<figureCitation captionStart="Figures 49–66" captionStartId="F6" captionText="Figures 49 - 66. Arkaditilla spp., ♂, genitalia 49, 52, 55, 58, 61, 64 genital capsule and volsella, inner view 50, 53, 56, 59, 62, 65 left penis valve 51, 54, 57, 60, 63, 66 right penis valve 49 - 51 A. bagrada (Cameron) 52 - 54 A. depressicornis (Mickel) 55 - 57 A. frim sp. nov., holotype 58 - 60 A. gibba sp. nov., holotype 61 - 63 A. leleji sp. nov., holotype 64 - 66 A. nallinia (Zavattari)." figureDoi="10.3897/jhr.84.68709.figures49-66" httpUri="https://binary.pensoft.net/fig/579485" pageId="0" pageNumber="145">49</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figures 49–66" captionStartId="F6" captionText="Figures 49 - 66. Arkaditilla spp., ♂, genitalia 49, 52, 55, 58, 61, 64 genital capsule and volsella, inner view 50, 53, 56, 59, 62, 65 left penis valve 51, 54, 57, 60, 63, 66 right penis valve 49 - 51 A. bagrada (Cameron) 52 - 54 A. depressicornis (Mickel) 55 - 57 A. frim sp. nov., holotype 58 - 60 A. gibba sp. nov., holotype 61 - 63 A. leleji sp. nov., holotype 64 - 66 A. nallinia (Zavattari)." figureDoi="10.3897/jhr.84.68709.figures49-66" httpUri="https://binary.pensoft.net/fig/579485" pageId="0" pageNumber="145">52</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figures 49–66" captionStartId="F6" captionText="Figures 49 - 66. Arkaditilla spp., ♂, genitalia 49, 52, 55, 58, 61, 64 genital capsule and volsella, inner view 50, 53, 56, 59, 62, 65 left penis valve 51, 54, 57, 60, 63, 66 right penis valve 49 - 51 A. bagrada (Cameron) 52 - 54 A. depressicornis (Mickel) 55 - 57 A. frim sp. nov., holotype 58 - 60 A. gibba sp. nov., holotype 61 - 63 A. leleji sp. nov., holotype 64 - 66 A. nallinia (Zavattari)." figureDoi="10.3897/jhr.84.68709.figures49-66" httpUri="https://binary.pensoft.net/fig/579485" pageId="0" pageNumber="145">55</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figures 49–66" captionStartId="F6" captionText="Figures 49 - 66. Arkaditilla spp., ♂, genitalia 49, 52, 55, 58, 61, 64 genital capsule and volsella, inner view 50, 53, 56, 59, 62, 65 left penis valve 51, 54, 57, 60, 63, 66 right penis valve 49 - 51 A. bagrada (Cameron) 52 - 54 A. depressicornis (Mickel) 55 - 57 A. frim sp. nov., holotype 58 - 60 A. gibba sp. nov., holotype 61 - 63 A. leleji sp. nov., holotype 64 - 66 A. nallinia (Zavattari)." figureDoi="10.3897/jhr.84.68709.figures49-66" httpUri="https://binary.pensoft.net/fig/579485" pageId="0" pageNumber="145">58</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figures 49–66" captionStartId="F6" captionText="Figures 49 - 66. Arkaditilla spp., ♂, genitalia 49, 52, 55, 58, 61, 64 genital capsule and volsella, inner view 50, 53, 56, 59, 62, 65 left penis valve 51, 54, 57, 60, 63, 66 right penis valve 49 - 51 A. bagrada (Cameron) 52 - 54 A. depressicornis (Mickel) 55 - 57 A. frim sp. nov., holotype 58 - 60 A. gibba sp. nov., holotype 61 - 63 A. leleji sp. nov., holotype 64 - 66 A. nallinia (Zavattari)." figureDoi="10.3897/jhr.84.68709.figures49-66" httpUri="https://binary.pensoft.net/fig/579485" pageId="0" pageNumber="145">61</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figures 49–66" captionStartId="F6" captionText="Figures 49 - 66. Arkaditilla spp., ♂, genitalia 49, 52, 55, 58, 61, 64 genital capsule and volsella, inner view 50, 53, 56, 59, 62, 65 left penis valve 51, 54, 57, 60, 63, 66 right penis valve 49 - 51 A. bagrada (Cameron) 52 - 54 A. depressicornis (Mickel) 55 - 57 A. frim sp. nov., holotype 58 - 60 A. gibba sp. nov., holotype 61 - 63 A. leleji sp. nov., holotype 64 - 66 A. nallinia (Zavattari)." figureDoi="10.3897/jhr.84.68709.figures49-66" httpUri="https://binary.pensoft.net/fig/579485" pageId="0" pageNumber="145">64</figureCitation>
|
||
); digitus straight, dorsal margin lamellate (Figs
|
||
<figureCitation captionStart="Figures 67–72" captionStartId="F7" captionText="Figures 67 - 72. Arkaditilla spp., ♂, genital volsella, ventro-inner view 67 A. bagrada (Cameron) 68 A. depressicornis (Mickel) 69 A. frim sp. nov., holotype 70 A. gibba sp. nov., holotype 71 A. leleji sp. nov., holotype 72 A. nallinia (Zavattari). White arrows indicate the paracuspis." figureDoi="10.3897/jhr.84.68709.figures67-72" httpUri="https://binary.pensoft.net/fig/579486" pageId="0" pageNumber="145">67-72</figureCitation>
|
||
); paracuspis absent, tuberculate or stick-like; penis valves symmetrical, with posterior hook (Figs
|
||
<figureCitation captionStart="Figures 49–66" captionStartId="F6" captionText="Figures 49 - 66. Arkaditilla spp., ♂, genitalia 49, 52, 55, 58, 61, 64 genital capsule and volsella, inner view 50, 53, 56, 59, 62, 65 left penis valve 51, 54, 57, 60, 63, 66 right penis valve 49 - 51 A. bagrada (Cameron) 52 - 54 A. depressicornis (Mickel) 55 - 57 A. frim sp. nov., holotype 58 - 60 A. gibba sp. nov., holotype 61 - 63 A. leleji sp. nov., holotype 64 - 66 A. nallinia (Zavattari)." figureDoi="10.3897/jhr.84.68709.figures49-66" httpUri="https://binary.pensoft.net/fig/579485" pageId="0" pageNumber="145">50</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figures 49–66" captionStartId="F6" captionText="Figures 49 - 66. Arkaditilla spp., ♂, genitalia 49, 52, 55, 58, 61, 64 genital capsule and volsella, inner view 50, 53, 56, 59, 62, 65 left penis valve 51, 54, 57, 60, 63, 66 right penis valve 49 - 51 A. bagrada (Cameron) 52 - 54 A. depressicornis (Mickel) 55 - 57 A. frim sp. nov., holotype 58 - 60 A. gibba sp. nov., holotype 61 - 63 A. leleji sp. nov., holotype 64 - 66 A. nallinia (Zavattari)." figureDoi="10.3897/jhr.84.68709.figures49-66" httpUri="https://binary.pensoft.net/fig/579485" pageId="0" pageNumber="145">51</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figures 49–66" captionStartId="F6" captionText="Figures 49 - 66. Arkaditilla spp., ♂, genitalia 49, 52, 55, 58, 61, 64 genital capsule and volsella, inner view 50, 53, 56, 59, 62, 65 left penis valve 51, 54, 57, 60, 63, 66 right penis valve 49 - 51 A. bagrada (Cameron) 52 - 54 A. depressicornis (Mickel) 55 - 57 A. frim sp. nov., holotype 58 - 60 A. gibba sp. nov., holotype 61 - 63 A. leleji sp. nov., holotype 64 - 66 A. nallinia (Zavattari)." figureDoi="10.3897/jhr.84.68709.figures49-66" httpUri="https://binary.pensoft.net/fig/579485" pageId="0" pageNumber="145">53</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figures 49–66" captionStartId="F6" captionText="Figures 49 - 66. Arkaditilla spp., ♂, genitalia 49, 52, 55, 58, 61, 64 genital capsule and volsella, inner view 50, 53, 56, 59, 62, 65 left penis valve 51, 54, 57, 60, 63, 66 right penis valve 49 - 51 A. bagrada (Cameron) 52 - 54 A. depressicornis (Mickel) 55 - 57 A. frim sp. nov., holotype 58 - 60 A. gibba sp. nov., holotype 61 - 63 A. leleji sp. nov., holotype 64 - 66 A. nallinia (Zavattari)." figureDoi="10.3897/jhr.84.68709.figures49-66" httpUri="https://binary.pensoft.net/fig/579485" pageId="0" pageNumber="145">54</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figures 49–66" captionStartId="F6" captionText="Figures 49 - 66. Arkaditilla spp., ♂, genitalia 49, 52, 55, 58, 61, 64 genital capsule and volsella, inner view 50, 53, 56, 59, 62, 65 left penis valve 51, 54, 57, 60, 63, 66 right penis valve 49 - 51 A. bagrada (Cameron) 52 - 54 A. depressicornis (Mickel) 55 - 57 A. frim sp. nov., holotype 58 - 60 A. gibba sp. nov., holotype 61 - 63 A. leleji sp. nov., holotype 64 - 66 A. nallinia (Zavattari)." figureDoi="10.3897/jhr.84.68709.figures49-66" httpUri="https://binary.pensoft.net/fig/579485" pageId="0" pageNumber="145">56</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figures 49–66" captionStartId="F6" captionText="Figures 49 - 66. Arkaditilla spp., ♂, genitalia 49, 52, 55, 58, 61, 64 genital capsule and volsella, inner view 50, 53, 56, 59, 62, 65 left penis valve 51, 54, 57, 60, 63, 66 right penis valve 49 - 51 A. bagrada (Cameron) 52 - 54 A. depressicornis (Mickel) 55 - 57 A. frim sp. nov., holotype 58 - 60 A. gibba sp. nov., holotype 61 - 63 A. leleji sp. nov., holotype 64 - 66 A. nallinia (Zavattari)." figureDoi="10.3897/jhr.84.68709.figures49-66" httpUri="https://binary.pensoft.net/fig/579485" pageId="0" pageNumber="145">57</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figures 49–66" captionStartId="F6" captionText="Figures 49 - 66. Arkaditilla spp., ♂, genitalia 49, 52, 55, 58, 61, 64 genital capsule and volsella, inner view 50, 53, 56, 59, 62, 65 left penis valve 51, 54, 57, 60, 63, 66 right penis valve 49 - 51 A. bagrada (Cameron) 52 - 54 A. depressicornis (Mickel) 55 - 57 A. frim sp. nov., holotype 58 - 60 A. gibba sp. nov., holotype 61 - 63 A. leleji sp. nov., holotype 64 - 66 A. nallinia (Zavattari)." figureDoi="10.3897/jhr.84.68709.figures49-66" httpUri="https://binary.pensoft.net/fig/579485" pageId="0" pageNumber="145">59</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figures 49–66" captionStartId="F6" captionText="Figures 49 - 66. Arkaditilla spp., ♂, genitalia 49, 52, 55, 58, 61, 64 genital capsule and volsella, inner view 50, 53, 56, 59, 62, 65 left penis valve 51, 54, 57, 60, 63, 66 right penis valve 49 - 51 A. bagrada (Cameron) 52 - 54 A. depressicornis (Mickel) 55 - 57 A. frim sp. nov., holotype 58 - 60 A. gibba sp. nov., holotype 61 - 63 A. leleji sp. nov., holotype 64 - 66 A. nallinia (Zavattari)." figureDoi="10.3897/jhr.84.68709.figures49-66" httpUri="https://binary.pensoft.net/fig/579485" pageId="0" pageNumber="145">60</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figures 49–66" captionStartId="F6" captionText="Figures 49 - 66. Arkaditilla spp., ♂, genitalia 49, 52, 55, 58, 61, 64 genital capsule and volsella, inner view 50, 53, 56, 59, 62, 65 left penis valve 51, 54, 57, 60, 63, 66 right penis valve 49 - 51 A. bagrada (Cameron) 52 - 54 A. depressicornis (Mickel) 55 - 57 A. frim sp. nov., holotype 58 - 60 A. gibba sp. nov., holotype 61 - 63 A. leleji sp. nov., holotype 64 - 66 A. nallinia (Zavattari)." figureDoi="10.3897/jhr.84.68709.figures49-66" httpUri="https://binary.pensoft.net/fig/579485" pageId="0" pageNumber="145">62</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figures 49–66" captionStartId="F6" captionText="Figures 49 - 66. Arkaditilla spp., ♂, genitalia 49, 52, 55, 58, 61, 64 genital capsule and volsella, inner view 50, 53, 56, 59, 62, 65 left penis valve 51, 54, 57, 60, 63, 66 right penis valve 49 - 51 A. bagrada (Cameron) 52 - 54 A. depressicornis (Mickel) 55 - 57 A. frim sp. nov., holotype 58 - 60 A. gibba sp. nov., holotype 61 - 63 A. leleji sp. nov., holotype 64 - 66 A. nallinia (Zavattari)." figureDoi="10.3897/jhr.84.68709.figures49-66" httpUri="https://binary.pensoft.net/fig/579485" pageId="0" pageNumber="145">63</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figures 49–66" captionStartId="F6" captionText="Figures 49 - 66. Arkaditilla spp., ♂, genitalia 49, 52, 55, 58, 61, 64 genital capsule and volsella, inner view 50, 53, 56, 59, 62, 65 left penis valve 51, 54, 57, 60, 63, 66 right penis valve 49 - 51 A. bagrada (Cameron) 52 - 54 A. depressicornis (Mickel) 55 - 57 A. frim sp. nov., holotype 58 - 60 A. gibba sp. nov., holotype 61 - 63 A. leleji sp. nov., holotype 64 - 66 A. nallinia (Zavattari)." figureDoi="10.3897/jhr.84.68709.figures49-66" httpUri="https://binary.pensoft.net/fig/579485" pageId="0" pageNumber="145">65</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figures 49–66" captionStartId="F6" captionText="Figures 49 - 66. Arkaditilla spp., ♂, genitalia 49, 52, 55, 58, 61, 64 genital capsule and volsella, inner view 50, 53, 56, 59, 62, 65 left penis valve 51, 54, 57, 60, 63, 66 right penis valve 49 - 51 A. bagrada (Cameron) 52 - 54 A. depressicornis (Mickel) 55 - 57 A. frim sp. nov., holotype 58 - 60 A. gibba sp. nov., holotype 61 - 63 A. leleji sp. nov., holotype 64 - 66 A. nallinia (Zavattari)." figureDoi="10.3897/jhr.84.68709.figures49-66" httpUri="https://binary.pensoft.net/fig/579485" pageId="0" pageNumber="145">66</figureCitation>
|
||
).
|
||
<emphasis bold="true" pageId="0" pageNumber="145">Female.</emphasis>
|
||
Unknown.
|
||
</paragraph>
|
||
<caption ID-Zenodo-Dep="5349696" doi="10.3897/jhr.84.68709.figures1-6" httpUri="https://binary.pensoft.net/fig/579480" pageId="0" pageNumber="145" start="Figures 1–6" startId="F1">
|
||
<paragraph pageId="0" pageNumber="145">
|
||
<emphasis bold="true" pageId="0" pageNumber="145">Figures 1-6.</emphasis>
|
||
<taxonomicName authorityName="Okayasu" authorityYear="2021" class="Insecta" family="Mutillidae" genus="Arkaditilla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Arkaditilla" order="Hymenoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="145">Arkaditilla</emphasis>
|
||
</taxonomicName>
|
||
spp., ♂, habitus
|
||
<emphasis bold="true" pageId="0" pageNumber="145">1, 3, 5</emphasis>
|
||
dorsal view
|
||
<emphasis bold="true" pageId="0" pageNumber="145">2, 4, 6</emphasis>
|
||
lateral view
|
||
<emphasis bold="true" pageId="0" pageNumber="145">1, 2</emphasis>
|
||
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="A. bagrada" order="Hymenoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="bagrada">
|
||
<emphasis italics="true" pageId="0" pageNumber="145">A. bagrada</emphasis>
|
||
</taxonomicName>
|
||
(Cameron)
|
||
<emphasis bold="true" pageId="0" pageNumber="145">3, 4</emphasis>
|
||
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="A. depressicornis" order="Hymenoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="depressicornis">
|
||
<emphasis italics="true" pageId="0" pageNumber="145">A. depressicornis</emphasis>
|
||
</taxonomicName>
|
||
(Mickel)
|
||
<emphasis bold="true" pageId="0" pageNumber="145">5, 6</emphasis>
|
||
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="A. frim" order="Hymenoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="frim">
|
||
<emphasis italics="true" pageId="0" pageNumber="145">A. frim</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., holotype. Scale bars: 3 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption ID-Zenodo-Dep="5349698" doi="10.3897/jhr.84.68709.figures7-12" httpUri="https://binary.pensoft.net/fig/579481" pageId="0" pageNumber="145" start="Figures 7–12" startId="F2">
|
||
<paragraph pageId="0" pageNumber="145">
|
||
<emphasis bold="true" pageId="0" pageNumber="145">Figures 7-12.</emphasis>
|
||
<taxonomicName authorityName="Okayasu" authorityYear="2021" class="Insecta" family="Mutillidae" genus="Arkaditilla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Arkaditilla" order="Hymenoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="145">Arkaditilla</emphasis>
|
||
</taxonomicName>
|
||
spp., ♂, habitus
|
||
<emphasis bold="true" pageId="0" pageNumber="145">7, 9, 11</emphasis>
|
||
dorsal view
|
||
<emphasis bold="true" pageId="0" pageNumber="145">8, 10, 12</emphasis>
|
||
lateral view
|
||
<emphasis bold="true" pageId="0" pageNumber="145">7, 8</emphasis>
|
||
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="A. gibba" order="Hymenoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="gibba">
|
||
<emphasis italics="true" pageId="0" pageNumber="145">A. gibba</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., holotype
|
||
<emphasis bold="true" pageId="0" pageNumber="145">9, 10</emphasis>
|
||
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="A. leleji" order="Hymenoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="leleji">
|
||
<emphasis italics="true" pageId="0" pageNumber="145">A. leleji</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., holotype
|
||
<emphasis bold="true" pageId="0" pageNumber="145">11, 12</emphasis>
|
||
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="A. nallinia" order="Hymenoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="nallinia">
|
||
<emphasis italics="true" pageId="0" pageNumber="145">A. nallinia</emphasis>
|
||
</taxonomicName>
|
||
(Zavattari). Scale bars: 3 mm.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="145" type="type species">
|
||
<paragraph pageId="0" pageNumber="145">Type species.</paragraph>
|
||
<paragraph pageId="0" pageNumber="145">
|
||
<taxonomicName authorityName="Okayasu" authorityYear="2021" class="Insecta" family="Mutillidae" genus="Arkaditilla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Arkaditilla leleji" order="Hymenoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="leleji">
|
||
<emphasis italics="true" pageId="0" pageNumber="145">Arkaditilla leleji</emphasis>
|
||
</taxonomicName>
|
||
gen. et sp. nov., designated here.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="145" type="species included">
|
||
<paragraph pageId="0" pageNumber="145">Species included.</paragraph>
|
||
<paragraph pageId="0" pageNumber="145">
|
||
This genus includes the following six species:
|
||
<taxonomicName authorityName="Okayasu" authorityYear="2021" baseAuthorityName="Cameron" baseAuthorityYear="1902" class="Insecta" family="Mutillidae" genus="Arkaditilla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Arkaditilla bagrada" order="Hymenoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="bagrada">
|
||
<emphasis italics="true" pageId="0" pageNumber="145">Arkaditilla bagrada</emphasis>
|
||
</taxonomicName>
|
||
(Cameron, 1902), comb. nov., ♂ (Indonesia, Malaysia);
|
||
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="A. depressicornis" order="Hymenoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="depressicornis">
|
||
<emphasis italics="true" pageId="0" pageNumber="145">A. depressicornis</emphasis>
|
||
</taxonomicName>
|
||
(Mickel, 1935), comb. nov., ♂ (Malaysia);
|
||
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="A. frim" order="Hymenoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="frim">
|
||
<emphasis italics="true" pageId="0" pageNumber="145">A. frim</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., ♂ (Malaysia);
|
||
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="A. gibba" order="Hymenoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="gibba">
|
||
<emphasis italics="true" pageId="0" pageNumber="145">A. gibba</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., ♂ (Indonesia);
|
||
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="A. leleji" order="Hymenoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="leleji">
|
||
<emphasis italics="true" pageId="0" pageNumber="145">A. leleji</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., ♂ (Indonesia);
|
||
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="A. nallinia" order="Hymenoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="nallinia">
|
||
<emphasis italics="true" pageId="0" pageNumber="145">A. nallinia</emphasis>
|
||
</taxonomicName>
|
||
(Zavattari, 1914), comb. nov., ♂ (Indonesia).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="145" type="distribution">
|
||
<paragraph pageId="0" pageNumber="145">Distribution.</paragraph>
|
||
<paragraph pageId="0" pageNumber="145">
|
||
Widespread in Sundaland, the Oriental Region (Figs
|
||
<figureCitation captionStart="Figure 73–76" captionStartId="F8" captionText="Figure 73 - 76. Distributional map of Arkaditilla spp. based on specimens examined herein and the literature (Cameron 1902; Lelej 1996 b; Mickel 1935, 1937; Zavattari 1914) 73 A. bagrada (Cameron) (red circle) 74 A. depressicornis (Mickel) (blue circle) 75 A. frim sp. nov. (yellow circle), A. gibba sp. nov. (purple triangle), and A. leleji sp. nov. (light blue square) 76 A. nallinia (Zavattari) (orange circle)." figureDoi="10.3897/jhr.84.68709.figure73-76" httpUri="https://binary.pensoft.net/fig/579487" pageId="0" pageNumber="145">73-76</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="145" type="etymology">
|
||
<paragraph pageId="0" pageNumber="145">Etymology.</paragraph>
|
||
<paragraph pageId="0" pageNumber="145">
|
||
I am delighted to name this new genus in honor of Arkady Stepanovich Lelej, a fellow mutillidologist, pioneer of Old World mutillid systematics, and friend, to celebrate his 75th birthday and 50th anniversary of his career since the publication of
|
||
<bibRefCitation author="Lelej, AS" journalOrPublisher="Entomologicheskoe Obozrenie" pageId="0" pageNumber="145" pagination="676 - 680" refId="B8" refString="Lelej, AS, 1971. New and little-known species of velvet ants (Hymenoptera, Mutillidae) from Middle Asia and Kazakhstan. Entomologicheskoe Obozrenie 50 (3): 676 - 680" title="New and little-known species of velvet ants (Hymenoptera, Mutillidae) from Middle Asia and Kazakhstan." volume="50" year="1971">Lelej (1971)</bibRefCitation>
|
||
, with a common suffix for mutillid genera. Gender feminine.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="145" type="biology_ecology">
|
||
<paragraph pageId="0" pageNumber="145">Biology.</paragraph>
|
||
<paragraph pageId="0" pageNumber="145">
|
||
This genus is rare in collections; only twelve specimens of
|
||
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="A. bagrada" order="Hymenoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="bagrada">
|
||
<emphasis italics="true" pageId="0" pageNumber="145">A. bagrada</emphasis>
|
||
</taxonomicName>
|
||
, nine specimens of
|
||
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="A. depressicornis" order="Hymenoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="depressicornis">
|
||
<emphasis italics="true" pageId="0" pageNumber="145">A. depressicornis</emphasis>
|
||
</taxonomicName>
|
||
, and seven specimens of
|
||
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="A. nallinia" order="Hymenoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="nallinia">
|
||
<emphasis italics="true" pageId="0" pageNumber="145">A. nallinia</emphasis>
|
||
</taxonomicName>
|
||
have been reported by previous authors (
|
||
<bibRefCitation author="Cameron, P" journalOrPublisher="Journal of the Straits Branch of the Royal Asiatic Society" pageId="0" pageNumber="145" pagination="29 - 131" refId="B2" refString="Cameron, P, 1902. On the Hymenoptera collected by Mr. Robert Shelford at Sarawak, and on the Hymenoptera of the Sarawak Museum. Journal of the Straits Branch of the Royal Asiatic Society 37: 29 - 131" title="On the Hymenoptera collected by Mr. Robert Shelford at Sarawak, and on the Hymenoptera of the Sarawak Museum." volume="37" year="1902">Cameron 1902</bibRefCitation>
|
||
;
|
||
<bibRefCitation author="Zavattari, E" journalOrPublisher="Bollettino della Societa Entomologica Italiana" pageId="0" pageNumber="145" pagination="61 - 114" refId="B26" refString="Zavattari, E, 1914. Mutille Austro-Malesi. Bollettino della Societa Entomologica Italiana 45: 61 - 114" title="Mutille Austro-Malesi." volume="45" year="1914">Zavattari 1914</bibRefCitation>
|
||
;
|
||
<bibRefCitation DOI="https://doi.org/10.1111/j.1365-2311.1935.tb01207.x" author="Mickel, CE" journalOrPublisher="Transactions of the Royal Entomological Society of London" pageId="0" pageNumber="145" pagination="177 - 312" refId="B20" refString="Mickel, CE, 1935. The mutillid wasps of the islands of the Pacific Ocean (Hymenoptera, Mutillidae). Transactions of the Royal Entomological Society of London 83 (2): 177 - 312, DOI: https://doi.org/10.1111/j.1365-2311.1935.tb01207.x" title="The mutillid wasps of the islands of the Pacific Ocean (Hymenoptera, Mutillidae)." url="https://doi.org/10.1111/j.1365-2311.1935.tb01207.x" volume="83" year="1935">Mickel 1935</bibRefCitation>
|
||
,
|
||
<bibRefCitation DOI="https://doi.org/10.1080/00222933708655284" author="Mickel, CE" journalOrPublisher="The Annals and Magazine of Natural History" pageId="0" pageNumber="145" pagination="441 - 456" refId="B21" refString="Mickel, CE, 1937. New species and records of Mutillidae (Hymenoptera) from Borneo and the Solomon Islands. The Annals and Magazine of Natural History 10 (19): 441 - 456, DOI: https://doi.org/10.1080/00222933708655284" title="New species and records of Mutillidae (Hymenoptera) from Borneo and the Solomon Islands." url="https://doi.org/10.1080/00222933708655284" volume="10" year="1937">1937</bibRefCitation>
|
||
;
|
||
<bibRefCitation DOI="https://doi.org/10.3759/tropics.6.91" author="Lelej, AS" journalOrPublisher="Tropics" pageId="0" pageNumber="145" pagination="91 - 104" refId="B11" refString="Lelej, AS, 1996b. Mutillid wasps collected in Malaysia and Indonesia by Dr. Sk. Yamane (Hymenoptera, Mutillidae). Tropics 6 (1-2): 91 - 104, DOI: https://doi.org/10.3759/tropics.6.91" title="Mutillid wasps collected in Malaysia and Indonesia by Dr. Sk. Yamane (Hymenoptera, Mutillidae)." url="https://doi.org/10.3759/tropics.6.91" volume="6" year="1996 b">Lelej 1996b</bibRefCitation>
|
||
). In this study, however, a series of 28 specimens collected in Malaise traps placed in a lowland rain forest at Pasoh Forest Reserve was examined, suggesting greater abundance in forested habitats. See
|
||
<bibRefCitation author="Maeto, K" journalOrPublisher="Journal of Tropical Forest Science" pageId="0" pageNumber="145" pagination="537 - 547" refId="B16" refString="Maeto, K, Fukuyama, K, Kirton, LG, 1999. Edge effects on ambrosia beetle assemblages in a lowland rain forest, bordering oil palm plantations, in peninsular Malaysia. Journal of Tropical Forest Science 11 (3): 537 - 547" title="Edge effects on ambrosia beetle assemblages in a lowland rain forest, bordering oil palm plantations, in peninsular Malaysia." volume="11" year="1999">
|
||
<normalizedToken originalValue="Maetô">Maeto</normalizedToken>
|
||
et al. (1999)
|
||
</bibRefCitation>
|
||
for sampling sites.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="145" type="remarks">
|
||
<paragraph pageId="0" pageNumber="145">Remarks.</paragraph>
|
||
<paragraph pageId="0" pageNumber="145">
|
||
This new genus belongs to the former
|
||
<taxonomicName authorityName="Lelej" authorityYear="1996" lsidName="" pageId="0" pageNumber="145" rank="tribe" tribe="Petersenidiini">Petersenidiini</taxonomicName>
|
||
Lelej, 1996 by having the penis valves symmetrical, and the species of this genus were formerly placed in
|
||
<taxonomicName authorityName="Okayasu" authorityYear="2021" class="Insecta" family="Mutillidae" genus="Krombeinidia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Krombeinidia" order="Hymenoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="145">Krombeinidia</emphasis>
|
||
</taxonomicName>
|
||
Lelej, 1996 (
|
||
<bibRefCitation author="Lelej, AS" journalOrPublisher="Far Eastern Entomologist" pageId="0" pageNumber="145" pagination="1 - 24" refId="B10" refString="Lelej, AS, 1996a. To the knowledge of the East Asian species of the tribe Trogaspidiini Bischoff, 1920 (Hymenoptera, Mutillidae) with description of eight new genera and two new species. Far Eastern Entomologist 30: 1 - 24" title="To the knowledge of the East Asian species of the tribe Trogaspidiini Bischoff, 1920 (Hymenoptera, Mutillidae) with description of eight new genera and two new species." volume="30" year="1996 a">Lelej 1996a</bibRefCitation>
|
||
) or
|
||
<taxonomicName authorityName="Okayasu" authorityYear="2021" class="Insecta" family="Mutillidae" genus="Petersenidia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Petersenidia" order="Hymenoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="145">Petersenidia</emphasis>
|
||
</taxonomicName>
|
||
Lelej, 1992 (
|
||
<bibRefCitation DOI="https://doi.org/10.3759/tropics.6.91" author="Lelej, AS" journalOrPublisher="Tropics" pageId="0" pageNumber="145" pagination="91 - 104" refId="B11" refString="Lelej, AS, 1996b. Mutillid wasps collected in Malaysia and Indonesia by Dr. Sk. Yamane (Hymenoptera, Mutillidae). Tropics 6 (1-2): 91 - 104, DOI: https://doi.org/10.3759/tropics.6.91" title="Mutillid wasps collected in Malaysia and Indonesia by Dr. Sk. Yamane (Hymenoptera, Mutillidae)." url="https://doi.org/10.3759/tropics.6.91" volume="6" year="1996 b">Lelej 1996b</bibRefCitation>
|
||
). However, the males of
|
||
<taxonomicName authorityName="Okayasu" authorityYear="2021" class="Insecta" family="Mutillidae" genus="Arkaditilla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Arkaditilla" order="Hymenoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="145">Arkaditilla</emphasis>
|
||
</taxonomicName>
|
||
differ from those of
|
||
<taxonomicName authorityName="Okayasu" authorityYear="2021" class="Insecta" family="Mutillidae" genus="Krombeinidia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Krombeinidia" order="Hymenoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="145">Krombeinidia</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName authorityName="Okayasu" authorityYear="2021" class="Insecta" family="Mutillidae" genus="Petersenidia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Petersenidia" order="Hymenoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="145">Petersenidia</emphasis>
|
||
</taxonomicName>
|
||
by having the mandible with a subbasal inner tooth (mandibular inner margin entire in
|
||
<taxonomicName authorityName="Okayasu" authorityYear="2021" class="Insecta" family="Mutillidae" genus="Krombeinidia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Krombeinidia" order="Hymenoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="145">Krombeinidia</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName authorityName="Okayasu" authorityYear="2021" class="Insecta" family="Mutillidae" genus="Petersenidia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Petersenidia" order="Hymenoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="145">Petersenidia</emphasis>
|
||
</taxonomicName>
|
||
), the basiparamere antero-ventrally serrate (entire in
|
||
<taxonomicName authorityName="Okayasu" authorityYear="2021" class="Insecta" family="Mutillidae" genus="Krombeinidia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Krombeinidia" order="Hymenoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="145">Krombeinidia</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName authorityName="Okayasu" authorityYear="2021" class="Insecta" family="Mutillidae" genus="Petersenidia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Petersenidia" order="Hymenoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="145">Petersenidia</emphasis>
|
||
</taxonomicName>
|
||
), and the digitus dorsal margin lamellate (digitus cylindrical in
|
||
<taxonomicName authorityName="Okayasu" authorityYear="2021" class="Insecta" family="Mutillidae" genus="Krombeinidia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Krombeinidia" order="Hymenoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="145">Krombeinidia</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName authorityName="Okayasu" authorityYear="2021" class="Insecta" family="Mutillidae" genus="Petersenidia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Petersenidia" order="Hymenoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="145">Petersenidia</emphasis>
|
||
</taxonomicName>
|
||
). Within
|
||
<taxonomicName authorityName="Bischoff" authorityYear="1920" lsidName="" pageId="0" pageNumber="145" rank="tribe" tribe="Trogaspidiini">Trogaspidiini</taxonomicName>
|
||
, the males of this new genus and
|
||
<taxonomicName authorityName="Lelej" authorityYear="2005" class="Insecta" family="Mutillidae" genus="Serendibiella" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Serendibiella" order="Hymenoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="145">Serendibiella</emphasis>
|
||
</taxonomicName>
|
||
Lelej, 2005 share the mandible with subbasal inner tooth, but the former differs from the latter by having the hypostomal carina without a tubercle (present in
|
||
<taxonomicName authorityName="Lelej" authorityYear="2005" class="Insecta" family="Mutillidae" genus="Serendibiella" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Serendibiella" order="Hymenoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="145">Serendibiella</emphasis>
|
||
</taxonomicName>
|
||
), the mesocoxa evenly convex (laterally swollen in
|
||
<taxonomicName authorityName="Lelej" authorityYear="2005" class="Insecta" family="Mutillidae" genus="Serendibiella" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Serendibiella" order="Hymenoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="145">Serendibiella</emphasis>
|
||
</taxonomicName>
|
||
), S2 without a lateral felt line (present in
|
||
<taxonomicName authorityName="Lelej" authorityYear="2005" class="Insecta" family="Mutillidae" genus="Serendibiella" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Serendibiella" order="Hymenoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="145">Serendibiella</emphasis>
|
||
</taxonomicName>
|
||
), S8 flattened (with sublateral longitudinal carina in
|
||
<taxonomicName authorityName="Lelej" authorityYear="2005" class="Insecta" family="Mutillidae" genus="Serendibiella" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Serendibiella" order="Hymenoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="145">Serendibiella</emphasis>
|
||
</taxonomicName>
|
||
), and the penis valves symmetrical (right penis valve longer than left in
|
||
<taxonomicName authorityName="Lelej" authorityYear="2005" class="Insecta" family="Mutillidae" genus="Serendibiella" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Serendibiella" order="Hymenoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="145">Serendibiella</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |