304 lines
36 KiB
XML
304 lines
36 KiB
XML
<document ID-CLB-Dataset="53340" ID-DOI="http://dx.doi.org/10.3897/zookeys.1163.101230" ID-GBIF-Dataset="1a8c80e6-f90d-40b7-a267-fa4973298147" ID-PMC="PMC10230278" ID-Pensoft-Pub="1313-2970-1163-143" ID-Pensoft-UUID="1B22BC4EE557576DB977D4F770CF3198" ID-PubMed="37266516" ID-ZooBank="9D07C59D7AF64428BB1A79D029368CA9" ModsDocID="1313-2970-1163-143" checkinTime="1684941535047" checkinUser="pensoft" docAuthor="Luu, Vinh Quang, Nguyen, Thuong Huyen, Le, Minh Duc, Grismer, Jesse L., Ha, Hong Bich, Sitthivong, Saly, Hoang, Tuoi Thi & Grismer, L. Lee" docDate="2023" docId="63620ADCA9D55D0D8FE4143C79AC4A51" docLanguage="en" docName="ZooKeys 1163: 143-176" docOrigin="ZooKeys 1163" docPubDate="2023-05-23" docSource="http://dx.doi.org/10.3897/zookeys.1163.101230" docTitle="Dixonius gialaiensis Luu & Nguyen & Le & Grismer & Ha & Sitthivong & Hoang & Grismer 2023, sp. nov." docType="treatment" docUuid="10BF67E1-8059-47CE-891C-B219BD7AA9C1" docUuidSource="ZooBank" docVersion="5" id="1B22BC4EE557576DB977D4F770CF3198" lastPageNumber="143" masterDocId="1B22BC4EE557576DB977D4F770CF3198" masterDocTitle="Two new species of Dixonius from Vietnam and Laos with a discussion of the taxonomy of Dixonius (Squamata, Gekkonidae)" masterLastPageNumber="176" masterPageNumber="143" pageNumber="143" updateTime="1732727107732" updateUser="ExternalLinkService">
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<mods:title id="4704E1A97F7412C6939AD14EC45C6609">Two new species of Dixonius from Vietnam and Laos with a discussion of the taxonomy of Dixonius (Squamata, Gekkonidae)</mods:title>
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<mods:namePart id="195A1F54EDC7ED0508CBB27DE8DF5E65">Luu, Vinh Quang</mods:namePart>
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<mods:affiliation id="00D83E88623D73579772F634E20E7C7E">Faculty of Forest Resources and Environmental Management, Vietnam National University of Forestry, Xuan Mai, Chuong My, Hanoi, Vietnam & Herpetology Laboratory, Department of Biology, La Sierra University, 4500 Riverwalk Parkway, Riverside, California 92505, USA</mods:affiliation>
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<mods:nameIdentifier id="618ACFF50E3272BDF9511D26426633B0" type="email">qvinhfuv@yahoo.com.au</mods:nameIdentifier>
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<mods:namePart id="FFC7EB854E70E8F73E322974E592D098">Nguyen, Thuong Huyen</mods:namePart>
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<mods:affiliation id="481755649613E953DB2814C5C67DA656">Faculty of Forest Resources and Environmental Management, Vietnam National University of Forestry, Xuan Mai, Chuong My, Hanoi, Vietnam</mods:affiliation>
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<mods:namePart id="2D6B5424BC648F75D33696027359007D">Le, Minh Duc</mods:namePart>
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<mods:affiliation id="B1E024556C45154C7A2D4C6ECAD10B1B">Faculty of Environmental Sciences, University of Science, Vietnam National University, Hanoi, 334 Nguyen Trai Road, Hanoi, Vietnam & Central Institute for Natural Resources and Environmental Studies, Vietnam National University, Hanoi, 19 Le Thanh Tong, Hanoi, Vietnam</mods:affiliation>
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<mods:namePart id="AB045C780FD86D955B78AA8252C18D6D">Grismer, Jesse L.</mods:namePart>
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<mods:affiliation id="AB2F10332A99E8EFC9961242E7A44A96">Herpetology Laboratory, Department of Biology, La Sierra University, 4500 Riverwalk Parkway, Riverside, California 92505, USA</mods:affiliation>
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<mods:namePart id="EFDAB77BD870890470FD01C1AE268A06">Ha, Hong Bich</mods:namePart>
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<mods:affiliation id="AED70F848424841694A557B10DAF9FEF">Faculty of Forest Resources and Environmental Management, Vietnam National University of Forestry, Xuan Mai, Chuong My, Hanoi, Vietnam</mods:affiliation>
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<mods:namePart id="D32620A7DB9510FE255B29030B7AD040">Sitthivong, Saly</mods:namePart>
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<mods:affiliation id="8B139C301713F5724BF14E3578E5428C">Department of Herpetology, American Museum of Natural History, Central Park West at 79</mods:affiliation>
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<mods:namePart id="1C611C796309CC05C354806E652800D4">Hoang, Tuoi Thi</mods:namePart>
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<mods:affiliation id="549575FD4E55100F27B1FF0F7326329B">Faculty of Forest Resources and Environmental Management, Vietnam National University of Forestry, Xuan Mai, Chuong My, Hanoi, Vietnam</mods:affiliation>
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<mods:namePart id="9B2CCFD6D3D8EFBA73BEEF51CFAA71A9">Grismer, L. Lee</mods:namePart>
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<mods:affiliation id="6C03EC15CEFB15C0817A746410E55FCE">Herpetology Laboratory, Department of Biology, La Sierra University, 4500 Riverwalk Parkway, Riverside, California 92505, USA & th</mods:affiliation>
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<mods:nameIdentifier id="0DBC494D73343E0C483CEAD1533D04F9" type="email">lgrismer@lasierra.edu</mods:nameIdentifier>
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<mods:typeOfResource id="1FDC84114FEE35A74592D86337644902">text</mods:typeOfResource>
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<mods:title id="10B7815226FA0F5B0BAC3E8B41E4AC26">ZooKeys</mods:title>
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<mods:part id="93F262A790DB845A7C710DCC6ED85BAB">
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<mods:date id="8AE6DC296A1872696798BDD10B90E223">2023</mods:date>
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<mods:number id="C6F8510C83820DB19ED8C123EDC0530A">2023-05-23</mods:number>
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<mods:number id="E514AACA0194669F1E9D3849390F8588">1163</mods:number>
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<mods:start id="52CA211ADA2974C6299BDC420C0BE6F5">143</mods:start>
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<mods:url id="66152DDF2B0EC4B7424A336663118C21">http://dx.doi.org/10.3897/zookeys.1163.101230</mods:url>
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<mods:classification id="0D1CD639A17B19586E2374A09042007F">journal article</mods:classification>
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<mods:identifier id="EE449410BC412E0B0EA6268B741311AD" type="DOI">http://dx.doi.org/10.3897/zookeys.1163.101230</mods:identifier>
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<treatment id="63620ADCA9D55D0D8FE4143C79AC4A51" ID-GBIF-Taxon="209629486" LSID="urn:lsid:zoobank.org:act:10BF67E1-8059-47CE-891C-B219BD7AA9C1" httpUri="http://treatment.plazi.org/id/63620ADCA9D55D0D8FE4143C79AC4A51" lastPageNumber="143" pageId="0" pageNumber="143" scope_class="Squamata" scope_family="Gekkonidae">
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<subSubSection id="4D9A1944061F2DF3599A7E3BFF51D645" pageId="0" pageNumber="143" type="nomenclature">
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<paragraph id="007659C1CF7566A48C1E210E0E107BE0" pageId="0" pageNumber="143">
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<taxonomicName id="EAD72643E019B8158C747C175879445C" LSID="https://zoobank.org/10BF67E1-8059-47CE-891C-B219BD7AA9C1" authority="Luu & Nguyen & Le & Grismer & Ha & Sitthivong & Hoang & Grismer, 2023" authorityName="Luu & Nguyen & Le & Grismer & Ha & Sitthivong & Hoang & Grismer" authorityYear="2023" family="Gekkonidae" genus="Dixonius" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Dixonius gialaiensis" order="Squamata" pageId="0" pageNumber="143" rank="species" species="gialaiensis" status="sp. nov.">Dixonius gialaiensis</taxonomicName>
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<taxonomicNameLabel id="4D14C6CDAC398EE3202A176727D43419" pageId="0" pageNumber="143">sp. nov.</taxonomicNameLabel>
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</paragraph>
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<subSubSection id="7740825A44332F16A75EACD09BB7B57D" pageId="0" pageNumber="143" type="description">
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<paragraph id="02D2A8143AA4F7EC432BAC239B7B24B5" pageId="0" pageNumber="143">
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<figureCitation id="2830B7AFCA81F968AAB1E470CC348304" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Dorsal views of Dixonius gialaiensis sp. nov. A adult male holotype VNUF R. 2020.22 (Field no. GL 02) B adult female paratype VNUF R. 2020.33 (Field No. GL 03) C juvenile male paratype VNUF R. 2020.44 (Field No. GL 04) in Chu Se Mountain Pass, Hbong Commune, Chu Se District, Gia Lai Province." figureDoi="10.3897/zookeys.1163.101230.figure8" httpUri="https://binary.pensoft.net/fig/853913" pageId="0" pageNumber="143">Fig. 8 Gialai leaf-toed gecko</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="010DEEB976EA66B74EE686D161957928" pageId="0" pageNumber="143" type="materials_examined">
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<paragraph id="339B14BDAEB7C31D969F4D66A4D49667" pageId="0" pageNumber="143">Material examined.</paragraph>
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<paragraph id="ACEC5197EA2E5C8075C26553FBDE7438" pageId="0" pageNumber="143">
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<materialsCitation id="D3EF260DC54A8A0A011B69230E78DCB0" collectingDate="2020-02-15" collectorName="Oanh Van Lo, Khanh Quoc Nguyen" county="Gia Lai Province" elevation="330" latitude="13.578972" location="Adult" longLatPrecision="1" longitude="108.23214" municipality="Chu Se District" specimenCount="1" typeStatus="Holotype">
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||
<emphasis id="D1139C19CF16B735FAFF9F8CD8D7DF08" bold="true" pageId="0" pageNumber="143">
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||
<emphasis id="723170B742C065FE6AC95C1B44FF663E" italics="true" pageId="0" pageNumber="143">
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||
<typeStatus id="60FD2DD7024ED9FBB337F3CB62AE8DF2">Holotype</typeStatus>
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||
</emphasis>
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||
.
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</emphasis>
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||
<location id="3A28A964696D5911C38D89413AF5C237" LSID="urn:lsid:plazi:treatment:63620ADCA9D55D0D8FE4143C79AC4A51:3A28A964696D5911C38D89413AF5C237" county="Gia Lai Province" latitude="13.578972" longLatPrecision="1" longitude="108.23214" municipality="Chu Se District" name="Adult">Adult</location>
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male, VNUF R.2020.22 (Field no. GL02) in
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<location id="0A04817F884A3E87A82D718C3540AC2A" LSID="urn:lsid:plazi:treatment:63620ADCA9D55D0D8FE4143C79AC4A51:0A04817F884A3E87A82D718C3540AC2A" county="Gia Lai Province" latitude="13.578972" longLatPrecision="1" longitude="108.23214" municipality="Chu Se District" name="Chu Se Mountain Pass">
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||
<location id="671EC8E9E9A70D7BAEAD340F46F6CDFF" LSID="urn:lsid:plazi:treatment:63620ADCA9D55D0D8FE4143C79AC4A51:671EC8E9E9A70D7BAEAD340F46F6CDFF" county="Gia Lai Province" latitude="13.578972" longLatPrecision="1" longitude="108.23214" municipality="Chu Se District" name="Chu Se Mountain">
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||
Chu Se
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<location id="1B4D71E721BA3CEECC29804B83614572" LSID="urn:lsid:plazi:treatment:63620ADCA9D55D0D8FE4143C79AC4A51:1B4D71E721BA3CEECC29804B83614572" county="Gia Lai Province" latitude="13.578972" longLatPrecision="1" longitude="108.23214" municipality="Chu Se District" name="Mountain Pass">Mountain Pass</location>
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</location>
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||
</location>
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,
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<location id="1A56B5F9A0A9B97EB8E9088B67032E95" LSID="urn:lsid:plazi:treatment:63620ADCA9D55D0D8FE4143C79AC4A51:1A56B5F9A0A9B97EB8E9088B67032E95" county="Gia Lai Province" latitude="13.578972" longLatPrecision="1" longitude="108.23214" municipality="Chu Se District" name="H'Bong Commune">
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||
<normalizedToken id="58D1BD035D707768CEFB970FC2A6A5FA" originalValue="H’Bong">H'Bong</normalizedToken>
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||
Commune
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||
</location>
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||
,
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||
<collectingMunicipality id="C12CF4AE05677478EEE4491FC627EA70">Chu Se District</collectingMunicipality>
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||
,
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||
<collectingCounty id="9A626932750551A9A18CA53D51F76F03">Gia Lai Province</collectingCounty>
|
||
(
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||
<geoCoordinate id="15062FBF8838D5D0B65620D2E977579F" degrees="13" direction="north" minutes="34" orientation="latitude" precision="1" seconds="44.3" value="13.578972">13°34'44.3"N</geoCoordinate>
|
||
,
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||
<geoCoordinate id="2EEF6CFE188D3758D34D105411B7603B" degrees="108" direction="east" minutes="13" orientation="longitude" precision="1" seconds="55.7" value="108.23214">108°13'55.7"E</geoCoordinate>
|
||
;
|
||
<elevation id="3E55248ED21043E24EB9BCB04105FC7F" metricMagnitude="2" metricUnit="m" metricValue="3.3" unit="m" value="330.0">
|
||
<quantity id="9BC74605A3FF8C661954857B938C80F3" metricMagnitude="2" metricUnit="m" metricValue="3.3" unit="m" value="330.0">330 m</quantity>
|
||
a.s.l.
|
||
</elevation>
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||
), collected by
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||
<collectorName id="6F7F454B2DC9D5FB8B8BDEA0D8E00604">Oanh Van Lo</collectorName>
|
||
and
|
||
<collectorName id="174D7E2D87CD020B526AE2D831508E83">Khanh Quoc Nguyen</collectorName>
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||
on
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||
<collectingDate id="3B5FCD9DD8518A4742030D739723E59A" value="2020-02-15">15 February 2020</collectingDate>
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||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="DADD10C9F6310B80D8D85E0C69752B64" collectingDate="2023-01-01" collectingDateMax="2023-12-31" collectingDateMin="2023-01-01" collectorName="Field No. GL" county="Gia Lai Province" elevation="330" latitude="13.578972" location="Adult" longLatPrecision="1" longitude="108.23214" municipality="Chu Se District" specimenCount="1" typeStatus="Paratypes">
|
||
<emphasis id="92A69A95F6D1C87C218E8EEBF7A4BD89" bold="true" pageId="0" pageNumber="143">
|
||
<emphasis id="37D6DFBD5E8F4DAED6A271ADE2B1FCE5" italics="true" pageId="0" pageNumber="143">
|
||
<typeStatus id="C28B4938E63A3281A48AA428DF881804">Paratypes</typeStatus>
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||
</emphasis>
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||
.
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||
</emphasis>
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VNUF R.2020.44 (
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||
<collectorName id="E65E57AAB117661F6CA466D63BCCB57E">Field No. GL</collectorName>
|
||
04), juvenile male, and VNUF R.2020.33 (
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||
<collectorName id="59F92F8436F91A0CDA6357797C8851BC">Field No. GL</collectorName>
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03), adult female; the same data as the holotype
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||
</materialsCitation>
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||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="552C6AD60042ECA3CF40E50F0AF2AC72" pageId="0" pageNumber="143" type="diagnosis">
|
||
<paragraph id="1785E32BB8C1222CFC4EDE6B4FEECD0F" pageId="0" pageNumber="143">Diagnosis.</paragraph>
|
||
<paragraph id="E2D9D8FCDF0B0979CE5CF65ABD12572C" pageId="0" pageNumber="143">
|
||
<taxonomicName id="EFA2863A3CBDC3AD0213769BAD356736" authorityName="Luu & Nguyen & Le & Grismer & Ha & Sitthivong & Hoang & Grismer" authorityYear="2023" family="Gekkonidae" genus="Dixonius" kingdom="Animalia" lsidName="Dixonius gialaiensis" order="Squamata" pageId="0" pageNumber="143" rank="species" species="gialaiensis">
|
||
<emphasis id="9A8E39C7FE56BA814CD1F91D34C5B747" italics="true" pageId="0" pageNumber="143">Dixonius gialaiensis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. can be separated from all other species of
|
||
<taxonomicName id="191EE904F069596C1AB38107F8D5DB30" authorityName="Bauer, Good & Branch" authorityYear="1997" family="Gekkonidae" genus="Dixonius" kingdom="Animalia" lsidName="Dixonius" order="Squamata" pageId="0" pageNumber="143" rank="genus">
|
||
<emphasis id="B8036007833944885E6FF5FC71CCA28E" italics="true" pageId="0" pageNumber="143">Dixonius</emphasis>
|
||
</taxonomicName>
|
||
by possessing the unique combination of having a maximum SVL of 47.4 mm; 19 longitudinal rows of dorsal tubercles at midbody; 19-21 longitudinal rows of ventrals across the abdomen; 7 or 8 supralabials, sixth in at midorbital position; 6 or 7 infralabials; 7 interorbital scales; 7 or 8 precloacal pores in males, femoral pores lacking; precloacal and femoral pores absent in female; 13-15 lamellae on fourth toe; dorsum olive grey color with more round brown blotches; canthal stripe continues behind orbit to back of head; lips with dark bars; two regularly disposed whitish tubercles along the sides near the flanks to tail tip. These characters are scored across all
|
||
<taxonomicName id="A8E56D381AEB5420F85D8B2873D6D0BE" authorityName="Bauer, Good & Branch" authorityYear="1997" family="Gekkonidae" genus="Dixonius" kingdom="Animalia" lsidName="Dixonius" order="Squamata" pageId="0" pageNumber="143" rank="genus">
|
||
<emphasis id="571147300D591D092137D41F8CF6AD6C" italics="true" pageId="0" pageNumber="143">Dixonius</emphasis>
|
||
</taxonomicName>
|
||
species from Vietnam and Laos in Tables
|
||
<tableCitation id="02E7CBB1ADBE306C66FF648EF147F918" captionStart="Table 6" captionStartId="T6" captionText="Table 6. Summary statistics of morphometric and meristic characters among the Dixonius species." httpUri="http://table.plazi.org/id/6B2B624F5CF43D86210AF0D3589F4317" pageId="0" pageNumber="143" tableUuid="6B2B624F5CF43D86210AF0D3589F4317">6</tableCitation>
|
||
,
|
||
<tableCitation id="C27FCAEFE76CD7A6857F04547C8E29C2" captionStart="Table 7" captionStartId="T7" captionText="Table 7. Significant p - values from the results of the ANOVA and TukeyHDS analyses comparing all combinations of species pairs. Character abbreviations are listed in the Materials and methods." httpUri="http://table.plazi.org/id/476E3BE45C50629A057F66E779CD961D" pageId="0" pageNumber="143" tableUuid="476E3BE45C50629A057F66E779CD961D">7</tableCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="39F9344C3E8580827ECB3C2AA4EC1A82" pageId="0" pageNumber="143" type="description">
|
||
<paragraph id="428DFC7C8C5E434C13CFE05025102BC3" pageId="0" pageNumber="143">Description of the holotype.</paragraph>
|
||
<paragraph id="917059DB18CD39A0E330195441891106" pageId="0" pageNumber="143">Adult male, SVL 41.2 mm; head moderate in length (HL/SVL 0.28), wide (HW/HL 0.66), depressed (HD/HL 0.44), distinct from neck; prefrontal region concave; canthus rostralis rounded; snout elongate (ES/HL 0.37), rounded in dorsal profile; eye moderate size (ED/HL 0.25); ear opening oval, obliquely oriented, moderate in size; diameter of eye slightly smaller than eye to ear distance (ED/EE 0.88); rostral rectangular, partially divided dorsally by straight rostral groove, bordered posteriorly by large left and right supranasals, bordered laterally by first supralabials; external nares bordered anteriorly by rostral, dorsally by large supranasal, posteriorly by two smaller postnasals, bordered ventrally by first supralabial; 8,7 (R,L) rectangular supralabials extending to below and slightly past posterior margin of eye, sixth in midorbital position; 6,6 (R,L), infralabials tapering smoothly to just below midpoint of eye, decreasing gradually in size; scales of rostrum and lores flat to domed, larger than granular scales on top of head and occiput; scales of occiput intermixed with distinct, small, conical tubercles; superciliaries elongate, largest anteriorly; mental triangular, bordered laterally by first infralabials and posteriorly by large left and right trapezoidal postmentals contacting medially for 60% of their length posterior to mental; gular and throat scales small, granular, grading anteriorly into slightly larger, flatter, smooth, imbricate, pectoral and ventral scales.</paragraph>
|
||
<paragraph id="A41AF112E694148B4E61A86E35DA7251" pageId="0" pageNumber="143">Body relatively short (AG/SVL 0.38); dorsal scales small, granular interspersed with larger, conical, regularly arranged, keeled tubercles; tubercles extend from top of head onto posterior haft of tail forming longitudinal rows, terminating at last portion of tail; smaller tubercles extend anteriorly onto nape and occiput, diminishing in size and distinction on top of head; 19 longitudinal rows of tubercles at midbody; 33 paravertebral scales, number of scales in a paravertebral row from first scale posterior to parietal scale to last scale at the level of vent opening; 23 paravertebral scales in a row between limb insertions; 21 flat, imbricate, ventral scales much larger than dorsal scales; 7 enlarge, pore-bearing, precloacal scales in an angular series; and no deep precloacal groove or depression.</paragraph>
|
||
<paragraph id="475100B50E0B5220698C281F50B56D1E" pageId="0" pageNumber="143">Forelimbs moderate in stature, relatively short (FA/SVL 0.15); granular scales of forearm slightly larger than those on body, interspersed with small tubercles; hind limbs more robust than forelimbs, moderate in length (TBL/SVL 0.17), covered dorsally by granular scales interspersed with large, and small conical tubercles; ventral scales of thigh flat, imbricate, larger than dorsals; subtibial scales flat, imbricate; proximal femoral scales smaller than distal femorals; femoral pores absent; digits relatively long with 14 lamellae on fourth toe; and claws well developed.</paragraph>
|
||
<paragraph id="E923FD37DE866DDDB79A44EF3E156BBE" pageId="0" pageNumber="143">Tail 108.4 mm in length, 4.5 mm in width at base, tapering to a point; dorsal scales of flat, square with conical, keeled tubercles in anterior part; median row of transversely expanded subcaudal scales, significantly larger than dorsal caudal scales on original portion; base of tail bearing hemipenal swellings; and postcloacal scales flat, imbricate.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="FA452DA94D30B8788BFABD897003EF67" pageId="0" pageNumber="143" type="coloration in life">
|
||
<paragraph id="FF8DA11D8E6BBCF0CBEBF484113F7672" pageId="0" pageNumber="143">Coloration in life</paragraph>
|
||
<paragraph id="987943B094EC39FE7167019F6F149D6F" pageId="0" pageNumber="143">
|
||
<emphasis id="ABE9F1879820C03AA1F07B455212F723" bold="true" pageId="0" pageNumber="143">
|
||
(Fig.
|
||
<figureCitation id="20932549B18DF69263EBA48E4F85DC90" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Dorsal views of Dixonius gialaiensis sp. nov. A adult male holotype VNUF R. 2020.22 (Field no. GL 02) B adult female paratype VNUF R. 2020.33 (Field No. GL 03) C juvenile male paratype VNUF R. 2020.44 (Field No. GL 04) in Chu Se Mountain Pass, Hbong Commune, Chu Se District, Gia Lai Province." figureDoi="10.3897/zookeys.1163.101230.figure8" httpUri="https://binary.pensoft.net/fig/853913" pageId="0" pageNumber="143">8</figureCitation>
|
||
).
|
||
</emphasis>
|
||
Ground color of dorsal head and dorsum grey brown with rounded black-brown blotches, decreasing gradually in size from head to body; canthal stripe continues behind orbit to back of head; dark bars on the lips; uneven light spots running from postorbital along the flanks to tip tail; upper surface of fore- and hindlimbs uniformly light grey with black-brown spots; dorsum of tail covered with some small rounded black-brown blotches; ventral surface beige uniformly as the belly and the throat.
|
||
</paragraph>
|
||
<caption id="BDA9AE8E77853E0ACE6843AE67CB3FE3" doi="10.3897/zookeys.1163.101230.figure8" httpUri="https://binary.pensoft.net/fig/853913" pageId="0" pageNumber="143" start="Figure 8" startId="F8">
|
||
<paragraph id="02CC46E117415FEAFD424723B7C4A399" pageId="0" pageNumber="143">
|
||
<emphasis id="19A22793805ED7210D1BCEC86EB24F0A" bold="true" pageId="0" pageNumber="143">Figure 8.</emphasis>
|
||
Dorsal views of
|
||
<taxonomicName id="D806A27FFE5BEA874FB6AFF4356FC50E" authorityName="Luu & Nguyen & Le & Grismer & Ha & Sitthivong & Hoang & Grismer" authorityYear="2023" family="Gekkonidae" genus="Dixonius" kingdom="Animalia" lsidName="Dixonius gialaiensis" order="Squamata" pageId="0" pageNumber="143" rank="species" species="gialaiensis">
|
||
<emphasis id="072F1D01AB97F4841A64638029EDF645" italics="true" pageId="0" pageNumber="143">Dixonius gialaiensis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.
|
||
<emphasis id="1F435CFCDB1255B8536211A4694DBF32" bold="true" pageId="0" pageNumber="143">A</emphasis>
|
||
adult male holotype VNUF R.2020.22 (Field no. GL02)
|
||
<emphasis id="5897657ADF3774E44ED6136EE4AB48A7" bold="true" pageId="0" pageNumber="143">B</emphasis>
|
||
adult female paratype VNUF R.2020.33 (Field No. GL03)
|
||
<emphasis id="E69B9A5CCDFD9820D15AC0001B7A1D99" bold="true" pageId="0" pageNumber="143">C</emphasis>
|
||
juvenile male paratype VNUF R.2020.44 (Field No. GL04) in Chu Se Mountain Pass, Hbong Commune, Chu Se District, Gia Lai Province.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="1C4FAB2669A84E90C80A41FECC4009CA" pageId="0" pageNumber="143" type="variation">
|
||
<paragraph id="D806B413FDD54143405A05CD7A0DF363" pageId="0" pageNumber="143">Variation</paragraph>
|
||
<paragraph id="20FA1CBA15CFA449C5816B76B2A66884" pageId="0" pageNumber="143">
|
||
<emphasis id="CAF9A0EF5ECAD7CB82A60C8854071D54" bold="true" pageId="0" pageNumber="143">
|
||
(Fig.
|
||
<figureCitation id="557616427B781D714011C2EA2CE4EA8E" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Dorsal views of Dixonius gialaiensis sp. nov. A adult male holotype VNUF R. 2020.22 (Field no. GL 02) B adult female paratype VNUF R. 2020.33 (Field No. GL 03) C juvenile male paratype VNUF R. 2020.44 (Field No. GL 04) in Chu Se Mountain Pass, Hbong Commune, Chu Se District, Gia Lai Province." figureDoi="10.3897/zookeys.1163.101230.figure8" httpUri="https://binary.pensoft.net/fig/853913" pageId="0" pageNumber="143">8</figureCitation>
|
||
).
|
||
</emphasis>
|
||
The female paratype (VNUF R.2020.33) generally has more dark brown blotches on head and dorsum, and uniformly black on the new regenerated tail. The dorsum of the of head and body of the juvenile male paratype (VNUF R.2020.44) pale brown with pale-colored blotches on granulose skin arranged along its sides extending from the flanks to the tail tip. Further measurements are summarized in Tables
|
||
<tableCitation id="EF6DBF96FCCBB7C00A0290CC660A21C1" captionStart="Table 2" captionStartId="T2" captionText="Table 2. Sex and raw meristic and categorical data used in the analyses from specimens of Dixonius from Vietnam and Laos. m = male; f = female; j = juvenile; r / l = right / left." httpUri="http://table.plazi.org/id/801CA6CB7150A0718E9859850689A939" pageId="0" pageNumber="143" tableUuid="801CA6CB7150A0718E9859850689A939">2</tableCitation>
|
||
-
|
||
<tableCitation id="85BDE79AA9DA2245CFF6A5123CB7B9C5" captionStart="Table 4" captionStartId="T4" captionText="Table 4. Mean percentages of uncorrected pairwise sequence divergence (p-distances) among the species of Dixonius. Intraspecific p-distance are in bold font, n / a = data not applicable." httpUri="http://table.plazi.org/id/DD1407F166BCF9B4DB0F908399E1D770" pageId="0" pageNumber="143" tableUuid="DD1407F166BCF9B4DB0F908399E1D770">4</tableCitation>
|
||
, Suppl. material 1: table S1.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="05F80A3EBF2D529C936EFAF9B85CEB18" pageId="0" pageNumber="143" type="distribution">
|
||
<paragraph id="8BDB0C58D3F61BBFCD917ECA305D5120" pageId="0" pageNumber="143">Distribution.</paragraph>
|
||
<paragraph id="03DAE014A1F5DE2E8B2834D4A8BA8A8C" pageId="0" pageNumber="143">
|
||
<taxonomicName id="2C25395715C4CF61B1EE611361192FCC" authorityName="Luu & Nguyen & Le & Grismer & Ha & Sitthivong & Hoang & Grismer" authorityYear="2023" family="Gekkonidae" genus="Dixonius" kingdom="Animalia" lsidName="Dixonius gialaiensis" order="Squamata" pageId="0" pageNumber="143" rank="species" species="gialaiensis">
|
||
<emphasis id="E96E7BF1030CA8C55EE89B0E46BEA048" italics="true" pageId="0" pageNumber="143">Dixonius gialaiensis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. currently is only known from the type locality of Chu Se Mountain Pass,
|
||
<normalizedToken id="CD7B60C965E3C30C9EAFAAB2FC1ACC55" originalValue="H’Bong">H'Bong</normalizedToken>
|
||
Commune, Chu Se District, Gia Lai Province, Central Highlands, Vietnam (Fig.
|
||
<figureCitation id="F394F71DE65C232EAB8FADA60A852760" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Location of the type localities of all known species of Dixonius. The inset delimits the study area. 1 Dixonius aaronbaueri from Ninh Thuan Province, Vietnam; 2 D. dulayaphitakorum from Ranong Province, Thailand; 3 D. mekongensis from Ubon Ratchathani Province, Thailand; 4 D. hangseesom from Kanchanaburi Province, Thailand; 5 D. kaweesaki from Prachuap Khiri Khan Province, Thailand; 6 D. pawangkhananti from Phetchaburi Province, Thailand; 7 D. lao from Khammouane Province, Laos; 8 D. melanostictus from Nakhon Ratchasima Province, Thailand; 9 D. minhlei from Dong Nai Province, Vietnam; 10 D. siamensis from SaraBuri and Nakhon Ratchasima provinces, Thailand; 11 D. somchanhae from Vientiane Capital, Laos; 12 D. taoi from Binh Thuan Province, Vietnam; 13 D. vietnamensis from Khanh Hoa Province, Vietnam; 14 D. muangfuangensis sp. nov. from Vientiane Province, Laos; 15 D. gialaiensis sp. nov. from Gia Lai Province, Vietnam." figureDoi="10.3897/zookeys.1163.101230.figure1" httpUri="https://binary.pensoft.net/fig/853906" pageId="0" pageNumber="143">1</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="DEE66019D0268CF2E68E1B78D989CCC4" pageId="0" pageNumber="143" type="natural history">
|
||
<paragraph id="F99E64FE8CB72964577C9A8BC02B0627" pageId="0" pageNumber="143">Natural history.</paragraph>
|
||
<paragraph id="30A62A4D4F67F8CC318C7F45DFD09CEC" pageId="0" pageNumber="143">
|
||
The specimens were found at night, between 19:45 and 21:00 h, on the ground in an area along the National Highway 25. The surrounding habitat was secondary montane forest with woody trees. The temperature and humidity were approximately 32.6 °C and 57% (Fig.
|
||
<figureCitation id="289AFC093C43647B1CBD7734E024BF92" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Habitat of Dixonius gialaiensis sp. nov. HBong Commune, Chu Se District, Gia Lai Province, Central Highlands, Vietnam." figureDoi="10.3897/zookeys.1163.101230.figure9" httpUri="https://binary.pensoft.net/fig/853914" pageId="0" pageNumber="143">9</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="C6111E2B9FBD5773E50C79865B09E453" doi="10.3897/zookeys.1163.101230.figure9" httpUri="https://binary.pensoft.net/fig/853914" pageId="0" pageNumber="143" start="Figure 9" startId="F9">
|
||
<paragraph id="B0AF947C2F64E185493FED97B243D6BD" pageId="0" pageNumber="143">
|
||
<emphasis id="9535F68BF533F482863BE9C99651D5F6" bold="true" pageId="0" pageNumber="143">Figure 9.</emphasis>
|
||
Habitat of
|
||
<taxonomicName id="0A2E52DD3D8F8F0FC7C59E970CD095D1" authorityName="Luu & Nguyen & Le & Grismer & Ha & Sitthivong & Hoang & Grismer" authorityYear="2023" family="Gekkonidae" genus="Dixonius" kingdom="Animalia" lsidName="Dixonius gialaiensis" order="Squamata" pageId="0" pageNumber="143" rank="species" species="gialaiensis">
|
||
<emphasis id="62C79E3E1FFFF2501CEA1DD4BF61614C" italics="true" pageId="0" pageNumber="143">Dixonius gialaiensis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. HBong Commune, Chu Se District, Gia Lai Province, Central Highlands, Vietnam.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="AC2B99E5B7D4651C614DC2141A1694A1" pageId="0" pageNumber="143" type="etymology">
|
||
<paragraph id="9084ED46AC344C997D0DD343A04A0A78" pageId="0" pageNumber="143">Etymology.</paragraph>
|
||
<paragraph id="07DE17E774A1D835EF6D4C4CB8535212" pageId="0" pageNumber="143">The new species is named after the type locality of Gia Lai Province, Central Highlands, Vietnam.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="4AAFBA79FA1FBFA6D33FFF65263C62FB" pageId="0" pageNumber="143" type="comparisons">
|
||
<paragraph id="F99658D75650DD2C21E68A998F8CDE02" pageId="0" pageNumber="143">Comparisons.</paragraph>
|
||
<paragraph id="8C7167BC1BE4049A197D24EEB6FD777E" pageId="0" pageNumber="143">
|
||
<taxonomicName id="152AC284448006DC38B60822EFEBA64C" authorityName="Luu & Nguyen & Le & Grismer & Ha & Sitthivong & Hoang & Grismer" authorityYear="2023" family="Gekkonidae" genus="Dixonius" kingdom="Animalia" lsidName="Dixonius gialaiensis" order="Squamata" pageId="0" pageNumber="143" rank="species" species="gialaiensis">
|
||
<emphasis id="5BD156AA342CF40CD487CD8E3A6EE4F6" italics="true" pageId="0" pageNumber="143">Dixonius gialaiensis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. is the sister species to
|
||
<taxonomicName id="DDE11AF3B43985038ABFD9223A60E52D" class="Reptilia" kingdom="Animalia" lsidName="D. minhlei" order="Squamata" pageId="0" pageNumber="143" phylum="Chordata" rank="species" species="minhlei">
|
||
<emphasis id="8A49B1280A2120B95D5C3CDE48610CA5" italics="true" pageId="0" pageNumber="143">D. minhlei</emphasis>
|
||
</taxonomicName>
|
||
(Fig.
|
||
<figureCitation id="546179C8CB563360B6CFF87B76F2BDFB" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Maximum likelihood topology of the Dixonius species from Vietnam and Laos with ultrafast bootstrap values (UFB) and Bayesian posterior probabilities (BPP) at the nodes, respectively." figureDoi="10.3897/zookeys.1163.101230.figure2" httpUri="https://binary.pensoft.net/fig/853907" pageId="0" pageNumber="143">2</figureCitation>
|
||
) from which it differs by an uncorrected pairwise sequence divergence of 3.60% (Table
|
||
<tableCitation id="3BB6C2CCEC47E0C57268E2A69A9F8BB4" captionStart="Table 4" captionStartId="T4" captionText="Table 4. Mean percentages of uncorrected pairwise sequence divergence (p-distances) among the species of Dixonius. Intraspecific p-distance are in bold font, n / a = data not applicable." httpUri="http://table.plazi.org/id/DD1407F166BCF9B4DB0F908399E1D770" pageId="0" pageNumber="143" tableUuid="DD1407F166BCF9B4DB0F908399E1D770">4</tableCitation>
|
||
). It is differentiated from it morphologically by having a significantly higher mean number of head length (HL), head width (HW), and axilla to groin length (AG). In addition, it differs from
|
||
<taxonomicName id="42E9AB33119821146AAAA69D9FC6ED35" class="Reptilia" kingdom="Animalia" lsidName="D. minhlei" order="Squamata" pageId="0" pageNumber="143" phylum="Chordata" rank="species" species="minhlei">
|
||
<emphasis id="3CF84EEEE1E785591CA1BCAFE6352B9E" italics="true" pageId="0" pageNumber="143">D. minhlei</emphasis>
|
||
</taxonomicName>
|
||
in color pattern (grey-brown dorsum with more round black-brown blotches versus olive gray dorsum with round brownish olive blotches). Statistically significant and discrete categorical differences between
|
||
<taxonomicName id="093DAB8935F80EB43081FE7F4F93FB79" authorityName="Luu & Nguyen & Le & Grismer & Ha & Sitthivong & Hoang & Grismer" authorityYear="2023" family="Gekkonidae" genus="Dixonius" kingdom="Animalia" lsidName="Dixonius gialaiensis" order="Squamata" pageId="0" pageNumber="143" rank="species" species="gialaiensis">
|
||
<emphasis id="4868577ACE418DAF98B902FB8B08A665" italics="true" pageId="0" pageNumber="143">Dixonius gialaiensis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. and all other species and populations are presented in Tables
|
||
<tableCitation id="F612201D7851F38FE3961082ED3B1262" captionStart="Table 5" captionStartId="T5" captionText="Table 5. Summary statistics of the principal component analysis of Dixonius species. Abbreviations are listed in the Materials and methods." httpUri="http://table.plazi.org/id/EE1BF5A11E5A2D773A5B0C585D97364F" pageId="0" pageNumber="143" tableUuid="EE1BF5A11E5A2D773A5B0C585D97364F">5</tableCitation>
|
||
-
|
||
<tableCitation id="A940F9ACFC1092FEFAE09635594A6E3B" captionStart="Table 7" captionStartId="T7" captionText="Table 7. Significant p - values from the results of the ANOVA and TukeyHDS analyses comparing all combinations of species pairs. Character abbreviations are listed in the Materials and methods." httpUri="http://table.plazi.org/id/476E3BE45C50629A057F66E779CD961D" pageId="0" pageNumber="143" tableUuid="476E3BE45C50629A057F66E779CD961D">7</tableCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |