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<document id="00C31334AAC6AEDA3985F63E31462413" ID-CLB-Dataset="28407" ID-DOI="10.11646/zootaxa.4565.1.3" ID-GBIF-Dataset="10f99299-a559-4576-aa1a-97a4e21a594f" ID-ISSN="1175-5326" ID-Zenodo-Dep="2589626" ID-ZooBank="F4B6A917-1EA8-40E4-850B-4507F1E92DBE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1552257254448" checkinUser="plazi" docAuthor="Hendrickx, M. E." docDate="2019" docId="5934E94BFFA8FFA7FF21862FFCBEFF48" docLanguage="en" docName="zootaxa.4565.1.3.pdf" docOrigin="Zootaxa 4565 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Heteronika Hendrickx 2019, gen. nov." docType="treatment" docVersion="9" lastPageNumber="50" masterDocId="A50D9133FFA9FFA5FFB68449FFACFFB0" masterDocTitle="Redescription of the rare shrimp Heterocarpus nesisi (Burukovsky, 1986) (Crustacea: Caridea: Pandalidae) rediscovered off western Mexico, with the proposal of a new genus" masterLastPageNumber="60" masterPageNumber="49" pageNumber="50" updateTime="1698701235494" updateUser="ExternalLinkService">
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<mods:title id="F7E12D0C94550E9B2040CA5A96F74CC5">Redescription of the rare shrimp Heterocarpus nesisi (Burukovsky, 1986) (Crustacea: Caridea: Pandalidae) rediscovered off western Mexico, with the proposal of a new genus</mods:title>
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<treatment id="5934E94BFFA8FFA7FF21862FFCBEFF48" ID-DOI="http://doi.org/10.5281/zenodo.5943210" ID-GBIF-Taxon="154366779" ID-Zenodo-Dep="5943210" LSID="urn:lsid:plazi:treatment:5934E94BFFA8FFA7FF21862FFCBEFF48" httpUri="http://treatment.plazi.org/id/5934E94BFFA8FFA7FF21862FFCBEFF48" lastPageId="2" lastPageNumber="50" pageId="1" pageNumber="50">
<subSubSection id="99870BD6FFA8FFA4FF21862FFE38FD30" box="[151,404,614,640]" pageId="1" pageNumber="50" type="nomenclature">
<paragraph id="D122585DFFA8FFA4FF21862FFE38FD30" blockId="1.[151,404,614,640]" box="[151,404,614,640]" pageId="1" pageNumber="50">
<heading id="8A6AEF31FFA8FFA4FF21862FFE38FD30" bold="true" box="[151,404,614,640]" fontSize="11" level="1" pageId="1" pageNumber="50" reason="1">
<emphasis id="E3E9844FFFA8FFA4FF21862FFE38FD30" bold="true" box="[151,404,614,640]" pageId="1" pageNumber="50">
<taxonomicName id="169D23DEFFA8FFA4FF21862FFE8CFD30" ID-CoL="9CLXP" authority="Hendrickx, 2019" authorityName="Hendrickx" authorityYear="2019" box="[151,288,614,640]" class="Malacostraca" family="Pandalidae" genus="Heteronika" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="50" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="E3E9844FFFA8FFA4FF21862FFE8CFD30" bold="true" box="[151,288,614,640]" italics="true" pageId="1" pageNumber="50">Heteronika</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F8DA3934FFA8FFA4FE90862FFE38FD30" box="[294,404,614,640]" pageId="1" pageNumber="50" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="99870BD6FFA8FFA4FF2186E4FC57FC2E" pageId="1" pageNumber="50" type="diagnosis">
<paragraph id="D122585DFFA8FFA4FF2186E4FC57FC2E" blockId="1.[151,1436,685,2006]" pageId="1" pageNumber="50">
<emphasis id="E3E9844FFFA8FFA4FF2186E4FEA6FD76" bold="true" box="[151,266,685,710]" pageId="1" pageNumber="50">Diagnosis</emphasis>
. Rostrum not movable, fixed to carapace. Dorsal and ventral teeth present, subtriangular, large, well separated. No movable spines on prolongation of rostrum on carapace. Carapace with one well marked lateral keel arising from above the antennal spine and extending to at least 2/3 of the carapace length. Carapace dorsally carinated anteriorly, rounded posteriorly. No supraorbital spine. Abdominal somites without dorsal keel, without posterior spine. Cornea wider than eyestalk. Maxilliped 2 terminal segment wider than long. Third maxilliped with exopod. Second pair of pereiopods strongly dissimilar, one stouter and shorter than the other, carpus subdivided in 17–18 segments (left) and (at least) 5 segments (right). No tegumental scales.
</paragraph>
</subSubSection>
<subSubSection id="99870BD6FFA8FFA4FF7187E0FD6EF94E" pageId="1" pageNumber="50" type="discussion">
<paragraph id="D122585DFFA8FFA4FF7187E0FC24FAC2" blockId="1.[151,1436,685,2006]" pageId="1" pageNumber="50">
<emphasis id="E3E9844FFFA8FFA4FF7187E0FE98FC72" bold="true" box="[199,308,937,962]" pageId="1" pageNumber="50">Remarks</emphasis>
. The original description of
<taxonomicName id="169D23DEFFA8FFA4FD2087E3FCD1FC71" authorityName="Burukovsky" authorityYear="1986" box="[662,893,938,961]" class="Malacostraca" family="Pandalidae" genus="Heterocarpus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="50" phylum="Arthropoda" rank="species" species="nesisi">
<emphasis id="E3E9844FFFA8FFA4FD2087E3FCD1FC71" box="[662,893,938,961]" italics="true" pageId="1" pageNumber="50">Heterocarpus nesisi</emphasis>
</taxonomicName>
(see Appendix) was based on a single male (
<typeStatus id="0E26E6FFFFA8FFA4FF288784FEA8FC56" box="[158,260,973,998]" pageId="1" pageNumber="50" type="holotype">holotype</typeStatus>
) of
<quantity id="1665F5B8FFA8FFA4FE808784FE2BFC55" box="[310,391,973,998]" metricMagnitude="-2" metricUnit="m" metricValue="7.6" pageId="1" pageNumber="50" unit="mm" value="76.0">76 mm</quantity>
(measured from the base or rostrum to the end of telson) captured in 1975 on a seamount located off the northeast coast of America (
<bibRefCitation id="B50C25ACFFA8FFA4FD3287B8FCFFFBBA" author="Burukovsky, R. N." box="[644,851,1009,1034]" pageId="1" pageNumber="50" pagination="62 - 73" refId="ref5077" refString="Burukovsky, R. N. (1986) A new shrimp species from the genus Heterocarpus (Crustacea: Decapoda: Pandalidae) and a brief review of species of the genus. Byulleten'Moskovskogo Obshchestva Ispytatelei Prirody, Otdel Biologicheskii, 91, 62 - 73. [in Russian]" type="journal article" year="1986">Burukovsky 1986</bibRefCitation>
). It was later transferred to
<taxonomicName id="169D23DEFFA8FFA4FB3987B8FAA9FBB9" authorityName="Hendrickx" authorityYear="2019" box="[1167,1285,1009,1033]" class="Malacostraca" family="Pandalidae" genus="Plesionika" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="50" phylum="Arthropoda" rank="genus">
<emphasis id="E3E9844FFFA8FFA4FB3987B8FAA9FBB9" box="[1167,1285,1009,1033]" italics="true" pageId="1" pageNumber="50">Plesionika</emphasis>
</taxonomicName>
(see
<bibRefCitation id="B50C25ACFFA8FFA4FAF587B8FE90FB9E" author="Chan, T. Y. &amp; Crosnier, A." pageId="1" pageNumber="50" pagination="187 - 234" refId="ref5171" refString="Chan, T. Y. &amp; Crosnier, A. (1997) Crustacea Decapoda: Deep-sea shrimps of the genus Plesionika Bate, 1888 (Pandalidae) from French Polynesia, with descriptions of five new species. Resultats des Campagnes MUSORSTOM, 18, 187 - 234." type="journal article" year="1997">Chan &amp; Crosnier 1997</bibRefCitation>
). Generic definition within the
<taxonomicName id="169D23DEFFA8FFA4FD13805CFC8CFB9E" authorityName="Haworth" authorityYear="1825" box="[677,800,1045,1070]" class="Malacostraca" family="Pandalidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="50" phylum="Arthropoda" rank="family">Pandalidae</taxonomicName>
is rather unclear. It seems obvious that both
<taxonomicName id="169D23DEFFA8FFA4FA90805CFA30FB9D" authorityName="Hendrickx" authorityYear="2019" box="[1318,1436,1045,1069]" class="Malacostraca" family="Pandalidae" genus="Plesionika" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="50" phylum="Arthropoda" rank="genus">
<emphasis id="E3E9844FFFA8FFA4FA90805CFA30FB9D" box="[1318,1436,1045,1069]" italics="true" pageId="1" pageNumber="50">Plesionika</emphasis>
</taxonomicName>
and
<emphasis id="E3E9844FFFA8FFA4FF7E8073FEFAFBE1" box="[200,342,1082,1105]" italics="true" pageId="1" pageNumber="50">Heterocarpu</emphasis>
s contain several species with morphological characters that do not fit with one another, making both genera heterogeneous. Because it possesses one strong lateral keel &quot;
<taxonomicName id="169D23DEFFA8FFA4FC568017FB6EFBC5" authorityName="Burukovsky" authorityYear="1986" box="[992,1218,1118,1141]" class="Malacostraca" family="Pandalidae" genus="Heterocarpus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="50" phylum="Arthropoda" rank="species" species="nesisi">
<emphasis id="E3E9844FFFA8FFA4FC568017FB6EFBC5" box="[992,1218,1118,1141]" italics="true" pageId="1" pageNumber="50">Heterocarpus nesisi</emphasis>
</taxonomicName>
&quot; cannot be readily classified within the genus
<taxonomicName id="169D23DEFFA8FFA4FE7D80CBFDC8FB29" box="[459,612,1154,1177]" class="Malacostraca" family="Pandalidae" genus="Heterocarpus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="50" phylum="Arthropoda" rank="genus">
<emphasis id="E3E9844FFFA8FFA4FE7D80CBFDC8FB29" box="[459,612,1154,1177]" italics="true" pageId="1" pageNumber="50">Heterocarpus</emphasis>
</taxonomicName>
or within the genus
<taxonomicName id="169D23DEFFA8FFA4FCF880C8FC68FB29" authorityName="Hendrickx" authorityYear="2019" box="[846,964,1153,1177]" class="Malacostraca" family="Pandalidae" genus="Plesionika" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="50" phylum="Arthropoda" rank="genus">
<emphasis id="E3E9844FFFA8FFA4FCF880C8FC68FB29" box="[846,964,1153,1177]" italics="true" pageId="1" pageNumber="50">Plesionika</emphasis>
</taxonomicName>
. Diagnosis provided by
<bibRefCitation id="B50C25ACFFA8FFA4FB6C80C8FF10FB0E" author="Chace, F. A." pageId="1" pageNumber="50" pagination="84 - 88" refId="ref5129" refString="Chace, F. A. (1989) The holotype of Heterocarpus alexandri A. Milne-Edwards (Crustacea: Decapoda: Pandalidae). Proceedings of the Biological Society of Washington, 102 (1), 84 - 88." type="journal article" year="1989">Chace (1989: 19, 43)</bibRefCitation>
and by Holthuis (1993: key to genera) for
<taxonomicName id="169D23DEFFA8FFA4FD1480EFFC97FB0D" box="[674,827,1190,1213]" class="Malacostraca" family="Pandalidae" genus="Heterocarpus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="50" phylum="Arthropoda" rank="genus">
<emphasis id="E3E9844FFFA8FFA4FD1480EFFC97FB0D" box="[674,827,1190,1213]" italics="true" pageId="1" pageNumber="50">Heterocarpus</emphasis>
</taxonomicName>
and
<taxonomicName id="169D23DEFFA8FFA4FCC480ECFC44FB0D" authorityName="Hendrickx" authorityYear="2019" box="[882,1000,1189,1213]" class="Malacostraca" family="Pandalidae" genus="Plesionika" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="50" phylum="Arthropoda" rank="genus">
<emphasis id="E3E9844FFFA8FFA4FCC480ECFC44FB0D" box="[882,1000,1189,1213]" italics="true" pageId="1" pageNumber="50">Plesionika</emphasis>
</taxonomicName>
are very similar, and basic differences are restricted to (
<taxonomicName id="169D23DEFFA8FFA4FEE08083FE43FB51" box="[342,495,1226,1249]" class="Malacostraca" family="Pandalidae" genus="Heterocarpus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="50" phylum="Arthropoda" rank="genus">
<emphasis id="E3E9844FFFA8FFA4FEE08083FE43FB51" box="[342,495,1226,1249]" italics="true" pageId="1" pageNumber="50">Heterocarpus</emphasis>
</taxonomicName>
vs.
<taxonomicName id="169D23DEFFA8FFA4FDAB8080FD3FFB51" authorityName="Hendrickx" authorityYear="2019" box="[541,659,1225,1249]" class="Malacostraca" family="Pandalidae" genus="Plesionika" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="50" phylum="Arthropoda" rank="genus">
<emphasis id="E3E9844FFFA8FFA4FDAB8080FD3FFB51" box="[541,659,1225,1249]" italics="true" pageId="1" pageNumber="50">Plesionika</emphasis>
</taxonomicName>
): the postrostral dorsal carina reaching to nearly posterior margin vs. the postrostral part of the carapace rounded; the presence (1 or 2) vs. absence of lateral keels on carapace; the carpus of second pair of pereopods divided into 7–12 (left) and 18–25 (right) articles vs. divided into more than 3 articles.
<taxonomicName id="169D23DEFFA8FFA4FF4F817CFEC3FAFD" authorityName="Hendrickx" authorityYear="2019" box="[249,367,1333,1357]" class="Malacostraca" family="Pandalidae" genus="Plesionika" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="50" phylum="Arthropoda" rank="genus">
<emphasis id="E3E9844FFFA8FFA4FF4F817CFEC3FAFD" box="[249,367,1333,1357]" italics="true" pageId="1" pageNumber="50">Plesionika</emphasis>
</taxonomicName>
third somite is also unarmed or features a fixed posteromesial spine, while in
<taxonomicName id="169D23DEFFA8FFA4FAB5817FFA30FAFD" box="[1283,1436,1334,1357]" class="Malacostraca" family="Pandalidae" genus="Heterocarpus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="50" phylum="Arthropoda" rank="genus">
<emphasis id="E3E9844FFFA8FFA4FAB5817FFA30FAFD" box="[1283,1436,1334,1357]" italics="true" pageId="1" pageNumber="50">Heterocarpus</emphasis>
</taxonomicName>
several somites are armed with a posteromesial spine (see
<tableCitation id="9C1F6DE6FFA8FFA4FC938110FCDBFAC2" box="[805,887,1369,1394]" captionStart="TABLE 1" captionStartId="2.[151,239,296,319]" captionText="TABLE 1. Diagnostic characters in Heterocarpus, Plesionika, and Heteronika gen. nov. Sources: Chace (1989) and Holthuis (1993)." pageId="1" pageNumber="50">Table 1</tableCitation>
).
</paragraph>
<paragraph id="D122585DFFA8FFA4FF718134FD4DF922" blockId="1.[151,1436,685,2006]" pageId="1" pageNumber="50">
<bibRefCitation id="B50C25ACFFA8FFA4FF718134FE25FA26" author="Chace, F. A." box="[199,393,1405,1430]" pageId="1" pageNumber="50" pagination="84 - 88" refId="ref5129" refString="Chace, F. A. (1989) The holotype of Heterocarpus alexandri A. Milne-Edwards (Crustacea: Decapoda: Pandalidae). Proceedings of the Biological Society of Washington, 102 (1), 84 - 88." type="journal article" year="1989">Chace (1989: 88)</bibRefCitation>
suggested that
<taxonomicName id="169D23DEFFA8FFA4FD8E8137FC15FA26" authority="A. Milne-Edwards, 1883" authorityName="A. Milne-Edwards" authorityYear="1883" box="[568,953,1405,1430]" class="Malacostraca" family="Pandalidae" genus="Heterocarpus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="50" phylum="Arthropoda" rank="species" species="laevis">
<emphasis id="E3E9844FFFA8FFA4FD8E8137FD37FA25" box="[568,667,1405,1429]" italics="true" pageId="1" pageNumber="50">H. laevis</emphasis>
A. Milne-Edwards, 1883
</taxonomicName>
and
<taxonomicName id="169D23DEFFA8FFA4FC468137FA30FA26" authority="A. Milne-Edwards, 1883" authorityName="A. Milne-Edwards" authorityYear="1883" box="[1008,1436,1405,1430]" class="Malacostraca" family="Pandalidae" genus="Heterocarpus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="50" phylum="Arthropoda" rank="species" species="alexandri">
<emphasis id="E3E9844FFFA8FFA4FC468137FBD1FA25" box="[1008,1149,1405,1429]" italics="true" pageId="1" pageNumber="50">H. alexandri</emphasis>
A. Milne-Edwards, 1883
</taxonomicName>
&quot;might eventually constitute a separate genus&quot;. While referring to
<bibRefCitation id="B50C25ACFFA8FFA4FC1581E8FB93FA0A" author="Chace, F. A." box="[931,1087,1441,1466]" pageId="1" pageNumber="50" pagination="84 - 88" refId="ref5129" refString="Chace, F. A. (1989) The holotype of Heterocarpus alexandri A. Milne-Edwards (Crustacea: Decapoda: Pandalidae). Proceedings of the Biological Society of Washington, 102 (1), 84 - 88." type="journal article" year="1989">Chace (1989)</bibRefCitation>
suggestion,
<bibRefCitation id="B50C25ACFFA8FFA4FB6281E8FEB1FA6E" author="Chan, T. Y. &amp; Crosnier, A." pageId="1" pageNumber="50" pagination="187 - 234" refId="ref5171" refString="Chan, T. Y. &amp; Crosnier, A. (1997) Crustacea Decapoda: Deep-sea shrimps of the genus Plesionika Bate, 1888 (Pandalidae) from French Polynesia, with descriptions of five new species. Resultats des Campagnes MUSORSTOM, 18, 187 - 234." type="journal article" year="1997">Chan &amp; Crosnier (1997: 204)</bibRefCitation>
emphasized the fact that &quot;the increasing number of species belonging to the &quot;
<taxonomicName id="169D23DEFFA8FFA4FB1D818FFAA7FA6D" authorityName="A. Milne-Edwards" authorityYear="1883" box="[1195,1291,1477,1501]" class="Malacostraca" family="Pandalidae" genus="Heterocarpus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="50" phylum="Arthropoda" rank="species" species="laevis">
<emphasis id="E3E9844FFFA8FFA4FB1D818FFAA7FA6D" box="[1195,1291,1477,1501]" italics="true" pageId="1" pageNumber="50">P. laevis</emphasis>
</taxonomicName>
&quot; group may prompt some workers to follow
<bibRefCitation id="B50C25ACFFA8FFA4FDB281A0FD01F9B2" author="Chace, F. A." box="[516,685,1513,1538]" pageId="1" pageNumber="50" pagination="84 - 88" refId="ref5129" refString="Chace, F. A. (1989) The holotype of Heterocarpus alexandri A. Milne-Edwards (Crustacea: Decapoda: Pandalidae). Proceedings of the Biological Society of Washington, 102 (1), 84 - 88." type="journal article" year="1989">Chace's (1989)</bibRefCitation>
suggestion and propose a separate genus for them&quot;. In the case of
<taxonomicName id="169D23DEFFA8FFA4FF218247FF7BF995" authorityName="Burukovsky" authorityYear="1986" box="[151,215,1550,1573]" class="Malacostraca" family="Pandalidae" genus="Heterocarpus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="50" phylum="Arthropoda" rank="species" species="nesisi">
<emphasis id="E3E9844FFFA8FFA4FF218247FF7BF995" box="[151,215,1550,1573]" italics="true" pageId="1" pageNumber="50">nesisi</emphasis>
</taxonomicName>
it seems impossible to keep this species within the genus
<taxonomicName id="169D23DEFFA8FFA4FCD88247FBABF995" box="[878,1031,1550,1573]" class="Malacostraca" family="Pandalidae" genus="Heterocarpus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="50" phylum="Arthropoda" rank="genus">
<emphasis id="E3E9844FFFA8FFA4FCD88247FBABF995" box="[878,1031,1550,1573]" italics="true" pageId="1" pageNumber="50">Heterocarpus</emphasis>
</taxonomicName>
mostly because of the absence of a second, well marked lateral keel. Its affinity with
<taxonomicName id="169D23DEFFA8FFA4FD088278FC98F9F9" authorityName="Hendrickx" authorityYear="2019" box="[702,820,1585,1609]" class="Malacostraca" family="Pandalidae" genus="Plesionika" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="50" phylum="Arthropoda" rank="genus">
<emphasis id="E3E9844FFFA8FFA4FD088278FC98F9F9" box="[702,820,1585,1609]" italics="true" pageId="1" pageNumber="50">Plesionika</emphasis>
</taxonomicName>
is rather obscure and is difficult to asses because of the complexity of this genus and the great variety of forms it contains, but differences seem sufficient to consider that he cannot remains in
<taxonomicName id="169D23DEFFA8FFA4FE308230FE50F921" authorityName="Hendrickx" authorityYear="2019" box="[390,508,1657,1681]" class="Malacostraca" family="Pandalidae" genus="Plesionika" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="50" phylum="Arthropoda" rank="genus">
<emphasis id="E3E9844FFFA8FFA4FE308230FE50F921" box="[390,508,1657,1681]" italics="true" pageId="1" pageNumber="50">Plesionika</emphasis>
</taxonomicName>
either (see
<tableCitation id="9C1F6DE6FFA8FFA4FDC88230FD7CF922" box="[638,720,1657,1682]" captionStart="TABLE 1" captionStartId="2.[151,239,296,319]" captionText="TABLE 1. Diagnostic characters in Heterocarpus, Plesionika, and Heteronika gen. nov. Sources: Chace (1989) and Holthuis (1993)." pageId="1" pageNumber="50">Table 1</tableCitation>
).
</paragraph>
<paragraph id="D122585DFFA8FFA4FF7182D4FD6EF94E" blockId="1.[151,1436,685,2006]" pageId="1" pageNumber="50">
<emphasis id="E3E9844FFFA8FFA4FF7182D4FDB2F906" bold="true" box="[199,542,1693,1718]" pageId="1" pageNumber="50">
Ethymology and
<typeStatus id="0E26E6FFFFA8FFA4FE2482D4FE69F906" box="[402,453,1693,1718]" pageId="1" pageNumber="50">type</typeStatus>
species
</emphasis>
. The new generic name denote the intermediate position of
<taxonomicName id="169D23DEFFA8FFA4FB0D82D4FA94F905" authority="Hendrickx, 2019" authorityName="Hendrickx" authorityYear="2019" box="[1211,1336,1693,1717]" class="Malacostraca" family="Pandalidae" genus="Heteronika" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="51" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="E3E9844FFFA8FFA4FB0D82D4FA94F905" box="[1211,1336,1693,1717]" italics="true" pageId="1" pageNumber="50">Heteronika</emphasis>
</taxonomicName>
between
<taxonomicName id="169D23DEFFA8FFA4FF21828BFE9CF969" box="[151,304,1730,1753]" class="Malacostraca" family="Pandalidae" genus="Heterocarpus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="50" phylum="Arthropoda" rank="genus">
<emphasis id="E3E9844FFFA8FFA4FF21828BFE9CF969" box="[151,304,1730,1753]" italics="true" pageId="1" pageNumber="50">Heterocarpus</emphasis>
</taxonomicName>
(Hetero-) and
<taxonomicName id="169D23DEFFA8FFA4FE6D8288FDFDF969" authorityName="Hendrickx" authorityYear="2019" box="[475,593,1729,1753]" class="Malacostraca" family="Pandalidae" genus="Plesionika" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="50" phylum="Arthropoda" rank="genus">
<emphasis id="E3E9844FFFA8FFA4FE6D8288FDFDF969" box="[475,593,1729,1753]" italics="true" pageId="1" pageNumber="50">Plesionika</emphasis>
</taxonomicName>
(-nika), united to form the name
<taxonomicName id="169D23DEFFA8FFA4FC628288FBFDF969" authority="Hendrickx, 2019" authorityName="Hendrickx" authorityYear="2019" box="[980,1105,1729,1753]" class="Malacostraca" family="Pandalidae" genus="Heteronika" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="51" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="E3E9844FFFA8FFA4FC628288FBFDF969" box="[980,1105,1729,1753]" italics="true" pageId="1" pageNumber="50">Heteronika</emphasis>
</taxonomicName>
.
<typeStatus id="0E26E6FFFFA8FFA4FBD7828BFB34F96A" box="[1121,1176,1730,1754]" pageId="1" pageNumber="50">Type</typeStatus>
species:
<taxonomicName id="169D23DEFFA8FFA4FAB5828BFE01F94E" authority="Burukovsky, 1986" authorityName="Burukovsky" authorityYear="1986" class="Malacostraca" family="Pandalidae" genus="Heterocarpus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="50" phylum="Arthropoda" rank="species" species="nesisi">
<emphasis id="E3E9844FFFA8FFA4FAB5828BFF7BF94D" italics="true" pageId="1" pageNumber="50">Heterocarpus nesisi</emphasis>
<bibRefCitation id="B50C25ACFFA8FFA4FF6B82ACFE01F94E" author="Burukovsky, R. N." box="[221,429,1765,1790]" pageId="1" pageNumber="50" pagination="62 - 73" refId="ref5077" refString="Burukovsky, R. N. (1986) A new shrimp species from the genus Heterocarpus (Crustacea: Decapoda: Pandalidae) and a brief review of species of the genus. Byulleten'Moskovskogo Obshchestva Ispytatelei Prirody, Otdel Biologicheskii, 91, 62 - 73. [in Russian]" type="journal article" year="1986">Burukovsky, 1986</bibRefCitation>
</taxonomicName>
, by original designation.
</paragraph>
</subSubSection>
<subSubSection id="99870BD6FFA8FFA7FF718340FCBEFF48" lastPageId="2" lastPageNumber="51" pageId="1" pageNumber="50" type="description">
<paragraph id="D122585DFFA8FFA7FF718340FCBEFF48" blockId="1.[151,1436,685,2006]" lastBlockId="2.[151,1437,151,248]" lastPageId="2" lastPageNumber="51" pageId="1" pageNumber="50">
<emphasis id="E3E9844FFFA8FFA4FF718340FE76F892" bold="true" box="[199,474,1801,1826]" pageId="1" pageNumber="50">Other species included</emphasis>
. The material reported by
<bibRefCitation id="B50C25ACFFA8FFA4FCA58340FB81F892" author="Chan, T. Y. &amp; Crosnier, A." box="[787,1069,1801,1826]" pageId="1" pageNumber="50" pagination="187 - 234" refId="ref5171" refString="Chan, T. Y. &amp; Crosnier, A. (1997) Crustacea Decapoda: Deep-sea shrimps of the genus Plesionika Bate, 1888 (Pandalidae) from French Polynesia, with descriptions of five new species. Resultats des Campagnes MUSORSTOM, 18, 187 - 234." type="journal article" year="1997">Chan &amp; Crosnier (1997)</bibRefCitation>
as &quot;
<taxonomicName id="169D23DEFFA8FFA4FBD58340FA8FF891" baseAuthorityName="Burukovsky" baseAuthorityYear="1996" box="[1123,1315,1801,1825]" class="Malacostraca" family="Pandalidae" genus="Plesionika" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="50" phylum="Arthropoda" rank="species" species="nesisi">
<emphasis id="E3E9844FFFA8FFA4FBD58340FA8FF891" box="[1123,1315,1801,1825]" italics="true" pageId="1" pageNumber="50">Plesionika nesisi</emphasis>
</taxonomicName>
&quot; probably belongs to a different species and should be included in
<taxonomicName id="169D23DEFFA8FFA4FCBC8364FC2BF8F5" authority="Hendrickx, 2019" authorityName="Hendrickx" authorityYear="2019" box="[778,903,1837,1861]" class="Malacostraca" family="Pandalidae" genus="Heteronika" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="50" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="E3E9844FFFA8FFA4FCBC8364FC2BF8F5" box="[778,903,1837,1861]" italics="true" pageId="1" pageNumber="50">Heteronika</emphasis>
</taxonomicName>
<emphasis id="E3E9844FFFA8FFA4FC3B8364FC5AF8F6" bold="true" box="[909,1014,1837,1862]" pageId="1" pageNumber="50">
<taxonomicNameLabel id="F8DA3934FFA8FFA4FC3B8364FC5AF8F6" box="[909,1014,1837,1862]" pageId="1" pageNumber="50" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
once this is confirmed.
<taxonomicName id="169D23DEFFA8FFA4FAB58367FD8CF8DA" authority="A. Milne-Edwards, 1883" authorityName="A. Milne-Edwards" authorityYear="1883" class="Malacostraca" family="Pandalidae" genus="Heterocarpus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="50" phylum="Arthropoda" rank="species" species="alexandri">
<emphasis id="E3E9844FFFA8FFA4FAB58367FEAEF8D9" italics="true" pageId="1" pageNumber="50">Heterocarpus alexandri</emphasis>
A. Milne-Edwards, 1883
</taxonomicName>
, fits well with the diagnosis of
<taxonomicName id="169D23DEFFA8FFA4FCCD8318FC54F8D9" authority="Hendrickx, 2019" authorityName="Hendrickx" authorityYear="2019" box="[891,1016,1873,1897]" class="Malacostraca" family="Pandalidae" genus="Heteronika" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="50" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="E3E9844FFFA8FFA4FCCD8318FC54F8D9" box="[891,1016,1873,1897]" italics="true" pageId="1" pageNumber="50">Heteronika</emphasis>
</taxonomicName>
<emphasis id="E3E9844FFFA8FFA4FC488318FBCBF8DA" bold="true" box="[1022,1127,1873,1898]" pageId="1" pageNumber="50">
<taxonomicNameLabel id="F8DA3934FFA8FFA4FC488318FBCBF8DA" box="[1022,1127,1873,1898]" pageId="1" pageNumber="50" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
and it is herein proposed as a new combination,
<taxonomicName id="169D23DEFFA8FFA4FE36833CFDDDF83D" authorityName="Hendrickx" authorityYear="2019" box="[384,625,1909,1933]" class="Malacostraca" family="Pandalidae" genus="Heteronika" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="50" phylum="Arthropoda" rank="species" species="alexandri" status="comb. nov.">
<emphasis id="E3E9844FFFA8FFA4FE36833CFDDDF83D" box="[384,625,1909,1933]" italics="true" pageId="1" pageNumber="50">Heteronika alexandri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F8DA3934FFA8FFA4FDCC833FFD54F83E" box="[634,760,1909,1934]" pageId="1" pageNumber="50" rank="species">new comb.</taxonomicNameLabel>
The presence of a sharply defined middorsal boss on third somite in
<taxonomicName id="169D23DEFFA8FFA4FEB183D3FDCEF802" authority="(Chace 1989: 85)" authorityPageNumber="85" baseAuthorityName="Chace" baseAuthorityPageNumber="85" baseAuthorityYear="1989" box="[263,610,1945,1970]" class="Malacostraca" family="Pandalidae" genus="Heteronika" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="50" phylum="Arthropoda" rank="species" species="alexandri">
<emphasis id="E3E9844FFFA8FFA4FEB183D3FE3AF801" box="[263,406,1945,1969]" italics="true" pageId="1" pageNumber="50">H. alexandri</emphasis>
(
<bibRefCitation id="B50C25ACFFA8FFA4FE1083D0FDF6F802" author="Chace, F. A." box="[422,602,1945,1970]" pageId="1" pageNumber="50" pagination="84 - 88" refId="ref5129" refString="Chace, F. A. (1989) The holotype of Heterocarpus alexandri A. Milne-Edwards (Crustacea: Decapoda: Pandalidae). Proceedings of the Biological Society of Washington, 102 (1), 84 - 88." type="journal article" year="1989">Chace 1989: 85</bibRefCitation>
)
</taxonomicName>
and in the material identified as &quot;
<taxonomicName id="169D23DEFFA8FFA4FC5383D0FB0FF801" baseAuthorityName="Burukovsky" baseAuthorityYear="1996" box="[997,1187,1945,1969]" class="Malacostraca" family="Pandalidae" genus="Plesionika" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="50" phylum="Arthropoda" rank="species" species="nesisi">
<emphasis id="E3E9844FFFA8FFA4FC5383D0FB0FF801" box="[997,1187,1945,1969]" italics="true" pageId="1" pageNumber="50">Plesionika nesisi</emphasis>
</taxonomicName>
&quot; by
<bibRefCitation id="B50C25ACFFA8FFA4FB6D83D0FF4DF866" author="Chan, T. Y. &amp; Crosnier, A." pageId="1" pageNumber="50" pagination="187 - 234" refId="ref5171" refString="Chan, T. Y. &amp; Crosnier, A. (1997) Crustacea Decapoda: Deep-sea shrimps of the genus Plesionika Bate, 1888 (Pandalidae) from French Polynesia, with descriptions of five new species. Resultats des Campagnes MUSORSTOM, 18, 187 - 234." type="journal article" year="1997">Chan &amp; Crosnier (1997)</bibRefCitation>
matches a similar but weakly defined boss in
<taxonomicName id="169D23DEFFA8FFA4FD4A83F4FC6EF865" authorityName="Burukovsky" authorityYear="1986" box="[764,962,1981,2005]" class="Malacostraca" family="Pandalidae" genus="Heteronika" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="50" phylum="Arthropoda" rank="species" species="nesisi">
<emphasis id="E3E9844FFFA8FFA4FD4A83F4FC6EF865" box="[764,962,1981,2005]" italics="true" pageId="1" pageNumber="50">Heteronika nesisi</emphasis>
</taxonomicName>
. Species of
<taxonomicName id="169D23DEFFA8FFA4FBE483F7FB47F865" box="[1106,1259,1982,2005]" class="Malacostraca" family="Pandalidae" genus="Heterocarpus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="50" phylum="Arthropoda" rank="genus">
<emphasis id="E3E9844FFFA8FFA4FBE483F7FB47F865" box="[1106,1259,1982,2005]" italics="true" pageId="1" pageNumber="50">Heterocarpus</emphasis>
</taxonomicName>
and
<taxonomicName id="169D23DEFFA8FFA4FA9083F4FA30F865" authorityName="Hendrickx" authorityYear="2019" box="[1318,1436,1981,2005]" class="Malacostraca" family="Pandalidae" genus="Plesionika" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="50" phylum="Arthropoda" rank="genus">
<emphasis id="E3E9844FFFA8FFA4FA9083F4FA30F865" box="[1318,1436,1981,2005]" italics="true" pageId="1" pageNumber="50">Plesionika</emphasis>
</taxonomicName>
need to be reviewed in details and once this is achieved it will most probably lead to splitting these heterogeneous groups of species into several genera. As a consequence, some additional species with a single lateral carina on the carapace might then be included in
<taxonomicName id="169D23DEFFABFFA7FD9384A9FD0EFF48" authority="Hendrickx, 2019" authorityName="Hendrickx" authorityYear="2019" box="[549,674,224,248]" class="Malacostraca" family="Pandalidae" genus="Heteronika" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="51" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="E3E9844FFFABFFA7FD9384A9FD0EFF48" box="[549,674,224,248]" italics="true" pageId="2" pageNumber="51">Heteronika</emphasis>
</taxonomicName>
<emphasis id="E3E9844FFFABFFA7FD1F8496FCBEFF48" bold="true" box="[681,786,223,248]" pageId="2" pageNumber="51">
<taxonomicNameLabel id="F8DA3934FFABFFA7FD1F8496FCBEFF48" box="[681,786,223,248]" pageId="2" pageNumber="51" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>
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