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<document id="C5616D351B2CA334CFAE210D29532E26" ID-CLB-Dataset="9065" ID-DOI="10.11646/zootaxa.4897.1.1" ID-GBIF-Dataset="44450b6f-59be-485c-8fc3-8e673f8b03f5" ID-ISSN="1175-5326" ID-Zenodo-Dep="4392347" ID-ZooBank="1C8AA826-36BA-4F48-8F67-6DCD62DFC20F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1608799780834" checkinUser="plazi" docAuthor="Lepeco, Anderson &amp; Gonçalves, Rodrigo Barbosa" docDate="2020" docId="507B87CDAD43B570FF72292CCE82FECB" docLanguage="en" docName="zootaxa.4897.1.1.pdf" docOrigin="Zootaxa 4897 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Augochlora (Augochlora) helena Lepeco &amp; Gonçalves 2020, sp. nov." docType="treatment" docUuid="2C9EFC50-793E-4FC9-BBC2-9AA377F15259" docUuidSource="ZooBank" docVersion="8" lastPageNumber="48" masterDocId="AC42FFB5AD6EB55FFFE52B0ECA11FFD6" masterDocTitle="A revision of the bee genus Augochlora Smith (Hymenoptera; Apoidea) in Southern South America" masterLastPageNumber="97" masterPageNumber="1" pageNumber="46" updateTime="1698847960660" updateUser="ExternalLinkService">
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<mods:title id="1F9072FDAC9F475C8A0420E089D910CF">A revision of the bee genus Augochlora Smith (Hymenoptera; Apoidea) in Southern South America</mods:title>
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<mods:namePart id="110AE1A45FF03B3481BB4D38E3267F0A">Lepeco, Anderson</mods:namePart>
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<mods:namePart id="8FFA3006262F74D8E7C33C9B1992A2DC">Gonçalves, Rodrigo Barbosa</mods:namePart>
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<mods:nameIdentifier id="89DEE3AFE55FAE813895949D9686B2B7" type="email">al.lepeco@gmail.com</mods:nameIdentifier>
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<mods:title id="36CFEA7CB68A72F225B6A8F2C15B2AC3">Zootaxa</mods:title>
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<mods:date id="8840A602CB7414960ED9729D1C16EA9C">2020</mods:date>
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<treatment id="507B87CDAD43B570FF72292CCE82FECB" ID-DOI="http://doi.org/10.5281/zenodo.4392429" ID-GBIF-Taxon="176435633" ID-Zenodo-Dep="4392429" LSID="urn:lsid:zoobank.org:act:2C9EFC50-793E-4FC9-BBC2-9AA377F15259" httpUri="http://treatment.plazi.org/id/507B87CDAD43B570FF72292CCE82FECB" lastPageId="47" lastPageNumber="48" pageId="45" pageNumber="46">
<subSubSection id="90C86550AD43B572FF72292CC89EFDEB" box="[151,655,546,573]" pageId="45" pageNumber="46" type="nomenclature">
<paragraph id="D86D36DBAD43B572FF72292CC89EFDEB" blockId="45.[151,655,546,609]" box="[151,655,546,573]" pageId="45" pageNumber="46">
<heading id="832581B7AD43B572FF72292CC89EFDEB" bold="true" box="[151,655,546,573]" fontSize="11" level="2" pageId="45" pageNumber="46" reason="4">
<emphasis id="EAA6EAC9AD43B572FF72292CC89EFDEB" bold="true" box="[151,655,546,573]" pageId="45" pageNumber="46">
<taxonomicName id="1FD24D58AD43B572FF72292CC838FDEB" authority="Lepeco &amp; Gonçalves, 2020" authorityName="Lepeco &amp; Gonçalves" authorityYear="2020" box="[151,553,546,573]" class="Insecta" family="Halictidae" genus="Augochlora" kingdom="Animalia" order="Hymenoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="helena" status="sp. nov." subGenus="Augochlora">
<emphasis id="EAA6EAC9AD43B572FF72292CC838FDEB" bold="true" box="[151,553,546,573]" italics="true" pageId="45" pageNumber="46">Augochlora (Augochlora) helena</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F19557B2AD43B572FDD4292DC89EFDEB" box="[561,655,547,573]" pageId="45" pageNumber="46" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="90C86550AD43B572FF722949CB5CFDB7" box="[151,333,583,609]" pageId="45" pageNumber="46" type="description">
<paragraph id="D86D36DBAD43B572FF722949CB5CFDB7" blockId="45.[151,655,546,609]" box="[151,333,583,609]" pageId="45" pageNumber="46">
(
<figureCitation id="40E92A5EAD43B572FF7A2949CB1DFDB7" box="[159,268,583,609]" captionStart="FIGURE 23" captionStartId="46.[151,250,1295,1319]" captionTargetBox="[151,1435,181,1270]" captionTargetId="figure-19@46.[151,1435,181,1270]" captionTargetPageId="46" captionText="FIGURE 23. Holotype female of Augochlora helena sp. nov. A) habitus, B) frontal view of head, C) dorsal view of mesosoma, D) dorsal view of metasoma. Scale bar: 1.0 mm, all at same scale." figureDoi="http://doi.org/10.5281/zenodo.4392405" httpUri="https://zenodo.org/record/4392405/files/figure.png" pageId="45" pageNumber="46">Figure 23</figureCitation>
AD)
</paragraph>
</subSubSection>
<subSubSection id="90C86550AD43B572FF722986CF8AFCD9" pageId="45" pageNumber="46" type="diagnosis">
<paragraph id="D86D36DBAD43B572FF722986CF8AFCD9" blockId="45.[151,1437,648,2043]" pageId="45" pageNumber="46">
<emphasis id="EAA6EAC9AD43B572FF722986CB03FD74" bold="true" box="[151,274,648,674]" pageId="45" pageNumber="46">Diagnosis.</emphasis>
This is the only species in the studied area with characteristics typical of the
<taxonomicName id="1FD24D58AD43B572FB462986CF50FD74" authorityName="Vachal" authorityYear="1911" box="[1187,1345,648,674]" class="Insecta" family="Halictidae" genus="Halictus" kingdom="Animalia" order="Hymenoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="repandirostris">
<emphasis id="EAA6EAC9AD43B572FB462986CF50FD74" box="[1187,1345,648,674]" italics="true" pageId="45" pageNumber="46">repandirostris</emphasis>
</taxonomicName>
species group: epistomal angle weakly protruding over clypeus, clypeus apex prolonged over apical border, frons slightly protuberant above antennal sockets, and overall body and appendages with swollen appearance. Besides this,
<taxonomicName id="1FD24D58AD43B572FA6129DECAF3FCD8" authority="Lepeco &amp; Gonçalves, 2020" authorityName="Lepeco &amp; Gonçalves" authorityYear="2020" class="Insecta" family="Halictidae" genus="Augochlora" kingdom="Animalia" order="Hymenoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="helena" status="sp. nov.">
<emphasis id="EAA6EAC9AD43B572FA6129DECAF3FCD8" italics="true" pageId="45" pageNumber="46">A. helena</emphasis>
</taxonomicName>
<emphasis id="EAA6EAC9AD43B572FF0C29FACB53FCD8" bold="true" box="[233,322,756,782]" pageId="45" pageNumber="46">
<taxonomicNameLabel id="F19557B2AD43B572FF0C29FACB53FCD8" box="[233,322,756,782]" pageId="45" pageNumber="46" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is the only
<emphasis id="EAA6EAC9AD43B572FE2029FAC89AFCD8" box="[453,651,756,782]" italics="true" pageId="45" pageNumber="46">
<taxonomicName id="1FD24D58AD43B572FE2029FAC858FCD8" authorityName="Smith" authorityYear="1853" box="[453,585,756,782]" class="Insecta" family="Halictidae" genus="Augochlora" kingdom="Animalia" order="Hymenoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="genus" sensu="stricto">Augochlora</taxonomicName>
<taxonomicNameLabel id="F19557B2AD43B572FDB529FBC89AFCD8" box="[592,651,757,782]" pageId="45" pageNumber="46" sensu="stricto">s. str.</taxonomicNameLabel>
</emphasis>
in the study area with smooth medioapical margin of metapostnotum.
</paragraph>
</subSubSection>
<subSubSection id="90C86550AD43B572FF222816C823FCAC" pageId="45" pageNumber="46" type="discussion">
<paragraph id="D86D36DBAD43B572FF222816C823FCAC" blockId="45.[151,1437,648,2043]" pageId="45" pageNumber="46">
<emphasis id="EAA6EAC9AD43B572FF222816CB5EFCE4" bold="true" box="[199,335,792,818]" pageId="45" pageNumber="46">Comments.</emphasis>
Species of the
<taxonomicName id="1FD24D58AD43B572FE1E2816C888FCE4" authorityName="Vachal" authorityYear="1911" box="[507,665,792,818]" class="Insecta" family="Halictidae" genus="Halictus" kingdom="Animalia" order="Hymenoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="repandirostris">
<emphasis id="EAA6EAC9AD43B572FE1E2816C888FCE4" box="[507,665,792,818]" italics="true" pageId="45" pageNumber="46">repandirostris</emphasis>
</taxonomicName>
species group are known from Amazon forest and from Panamanian forest, with no species described for the southern region so far. Although a single specimen was found, it does not correspond to any described species.
</paragraph>
</subSubSection>
<subSubSection id="90C86550AD43B571FF22288ACB27F866" lastPageId="46" lastPageNumber="47" pageId="45" pageNumber="46" type="description">
<paragraph id="D86D36DBAD43B572FF22288AC8B7FC30" blockId="45.[151,1437,648,2043]" pageId="45" pageNumber="46">
<emphasis id="EAA6EAC9AD43B572FF22288AC828FC48" bold="true" box="[199,569,900,928]" pageId="45" pageNumber="46">
Description.
<specimenCount id="CED4FD52AD43B572FE85288BCB64FC76" box="[352,373,901,928]" pageId="45" pageNumber="46" type="female"></specimenCount>
. Measurements.
</emphasis>
Maximum interocular distance: 1.49; head length: 1.88; clypeus width: 0.60; clypeus length: 0.42; clypeoantennal distance: 0.40; clypeus ocellar distance: 1.12; intertegular distance: 1.55; T1 width: 2.36; T2 width: 2.56; body length: 7.00.
</paragraph>
<paragraph id="D86D36DBAD43B572FF2228FEC93FFAA4" blockId="45.[151,1437,648,2043]" pageId="45" pageNumber="46">
<emphasis id="EAA6EAC9AD43B572FF2228FECB1FFBDC" bold="true" box="[199,270,1008,1034]" pageId="45" pageNumber="46">Head.</emphasis>
Labrum basal surface black; basal elevation transverse. Mandible mostly dark brown, apex black; medial portion not strongly constricted, minimum width between 0.7x and 0.9x basal width; preapical tooth large and sharp, produced near mandible apex; distal part of adductor ridge weakly swollen. Clypeus mostly green with golden reflection, apical black band = OD; punctate, punctures large and weak, smooth in between; apex prolonged over apical border. Epistomal angle acute, weakly protruding over clypeus. Supraclypeal area green with golden reflection; densely punctate, loosely imbricate in between. Lower paraocular area with golden reflection; contiguously punctate; long setae (12 OD) intermixed with tiny setae. Antennae: scape black, pedicel dark brown, flagellum dark brown, ventrally pale. Frons with golden and green reflections intermixed; contiguously punctate; slightly protuberant above antennal sockets. Preoccipital area strongly lamellate, lamella expanded near post gena. Gena with golden reflection. Post gena with copper reflection; microreticulate, some scattered weak punctures; setae length &lt;2 OD. Hypostomal carina not projected anteriorly.
</paragraph>
<paragraph id="D86D36DBAD43B572FF222E72CF77F82D" blockId="45.[151,1437,648,2043]" pageId="45" pageNumber="46">
<emphasis id="EAA6EAC9AD43B572FF222E72CB5DFA40" bold="true" box="[199,332,1404,1430]" pageId="45" pageNumber="46">Mesosoma.</emphasis>
Pronotum dorsolateral angle orthogonal. Fore leg: coxa dark brown, weak green reflection; trochanter dark brown; femur dark brown with weak green reflection; ventral surface strongly rounded; long plumose setae (&lt;3 OD); tibia brown; tarsus light brown; white to yellowish setae on both. Mesoscutum green with golden reflection; densely punctate on disc, I &lt;PD, sparser posterior to median line, smooth in between, contiguously punctate anteriorly and on lateral surfaces; setae length &lt;OD. Scutellum medial furrow inconspicuous; green with golden reflection; punctate, I = PD, with larger punctures in between; with dense tiny setae intermixed with longer white setae (3 OD). Mesepisternum green; preepisternum densely punctate near pronotal lobe; hypoepimeral area densely punctate, I &lt;0.5 PD, smooth in between; below scrobe densely punctate, I &lt;0.5 PD, smooth in between, ventrally crowded punctate; with long setae (2 OD). Mid leg: coxa dark brown with weak green reflection; trochanter dark brown; ventral margin not swollen, almost straight; femur dark brown, flattened, mesofemoral brush yellowish; tibia brown; tarsus light brown; white to yellowish setae on both. Tegula dark brown with anterior green spot. Metanotum with dense tiny setae intermixed with longer white setae (= 3 OD). Metepisternum green with golden reflection; densely to crowded punctate, punctures weak. Hind leg: coxa green, anterior and posterior surfaces ventrally separated by a smoothly curved area; trochanter dark brown; setae length = 2 OD; femur dark brown; tibia brown; tarsus brown; outer surfaces without dark setae. Metapostnotum green with golden reflection; shorter than scutellum; basal area with radiating carinae, medioapical area microreticulate to smooth. Propodeum green with golden reflection; with tiny and long (3 OD) setae intermixed; posterior surface with tiny and larger punctures intermixed, smooth in between; lateral carinae ending below propodeum half; lateral surfaces imbricate among punctures.
</paragraph>
<caption id="8CAD6653AD40B571FF722E01C925FA9D" ID-DOI="http://doi.org/10.5281/zenodo.4392405" ID-Zenodo-Dep="4392405" httpUri="https://zenodo.org/record/4392405/files/figure.png" pageId="46" pageNumber="47" startId="46.[151,250,1295,1319]" targetBox="[151,1435,181,1270]" targetPageId="46">
<paragraph id="D86D36DBAD40B571FF722E01C925FA9D" blockId="46.[151,1437,1295,1355]" pageId="46" pageNumber="47">
<emphasis id="EAA6EAC9AD40B571FF722E01CB0EFAF1" bold="true" box="[151,287,1295,1319]" pageId="46" pageNumber="47">FIGURE 23.</emphasis>
Holotype female of
<taxonomicName id="1FD24D58AD40B571FE0B2E01C8A1FAF1" authority="Lepeco &amp; Gonçalves, 2020" authorityName="Lepeco &amp; Gonçalves" authorityYear="2020" box="[494,688,1295,1319]" class="Insecta" family="Halictidae" genus="Augochlora" kingdom="Animalia" order="Hymenoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="helena" status="sp. nov.">
<emphasis id="EAA6EAC9AD40B571FE0B2E01C8A1FAF1" box="[494,688,1295,1319]" italics="true" pageId="46" pageNumber="47">Augochlora helena</emphasis>
</taxonomicName>
<emphasis id="EAA6EAC9AD40B571FD502E01C914FAF1" bold="true" box="[693,773,1295,1319]" pageId="46" pageNumber="47">
<taxonomicNameLabel id="F19557B2AD40B571FD502E01C914FAF1" box="[693,773,1295,1319]" pageId="46" pageNumber="47" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
A) habitus, B) frontal view of head, C) dorsal view of mesosoma, D) dorsal view of metasoma. Scale bar: 1.0 mm, all at same scale.
</paragraph>
</caption>
<paragraph id="D86D36DBAD40B571FF222E93CB27F866" blockId="46.[151,1437,1437,2040]" pageId="46" pageNumber="47">
<emphasis id="EAA6EAC9AD40B571FF222E93CB5AFA61" bold="true" box="[199,331,1437,1463]" pageId="46" pageNumber="47">Metasoma.</emphasis>
T1 anterior surface densely punctate, punctures weak, smooth in between; mostly dark brown with weak green reflection; with few tiny decumbent setae among long setae. T1 dorsal surface weakly puncticulate on disc, smooth in between, coarser punctation on lateral surfaces; mostly brown, lateral surfaces green; mostly glabrous, bristles up to 2 OD on lateral surfaces; apex inflexed, black. T2 mostly green with golden reflection, dorsally brown; minutely punctate on disc, punctures weak, coarser and denser punctation on lateral surfaces, mostly imbricate in between, smooth on lateral surfaces; marginal area with punctate portion shorter than apical black band on sublateral surfaces; tiny decumbent setae near base and on lateral surfaces, long setae (2 OD) on lateral surfaces. T3 weakly punctate, imbricate in between; mostly green, base and apical band brown; tiny setae apex not reaching apical margin sublaterally. T4 weakly punctate, imbricate in between; mostly green, apical band brown; mostly with tiny decumbent setae, long setae (2 OD) scattered on disc and lateral surfaces. T5 black; pseudopygidial area not covered by scale-like setae, colliculate integument exposed. Pygidial plate black, apex slightly rounded. S1 without elevation; brown with weak green reflection; long setae (&gt; 2 OD) medially, margin tiny decumbent setae. S2 mostly dark brown, apical band yellowish; medioapical margin sharp; punctate, microreticulate in between; long setae (&gt; 1 OD). S34 as S2. S5 apical margin widely rounded; dark brown; long setae (&gt; 1 OD). S6 dark brown; long dark setae on apex.
</paragraph>
</subSubSection>
<subSubSection id="90C86550AD40B571FF222CB7C9D6F805" box="[199,967,1977,2004]" pageId="46" pageNumber="47" type="distribution">
<paragraph id="D86D36DBAD40B571FF222CB7C9D6F805" blockId="46.[151,1437,1437,2040]" box="[199,967,1977,2004]" pageId="46" pageNumber="47">
<emphasis id="EAA6EAC9AD40B571FF222CB7CB70F805" bold="true" box="[199,353,1977,2003]" pageId="46" pageNumber="47">Distribution.</emphasis>
The only specimen found is from Balsa Nova,
<collectingRegion id="1A16F839AD40B571FC962CB7C9D2F805" box="[883,963,1977,2003]" country="Brazil" name="Parana" pageId="46" pageNumber="47">Paraná</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="90C86550AD40B570FF222CD3CEA0FF03" lastPageId="47" lastPageNumber="48" pageId="46" pageNumber="47" type="etymology">
<paragraph id="D86D36DBAD40B570FF222CD3CEA0FF03" blockId="46.[151,1437,1437,2040]" lastBlockId="47.[151,1436,151,287]" lastPageId="47" lastPageNumber="48" pageId="46" pageNumber="47">
<emphasis id="EAA6EAC9AD40B571FF222CD3CB5FF821" bold="true" box="[199,334,2013,2039]" pageId="46" pageNumber="47">Etymology.</emphasis>
Helena is a Latinate form of the name Hélène, who was cast away from her homeland by Paris in the Greek mythology. This is a reference to the fact that this is the first species in the
<taxonomicName id="1FD24D58AD41B570FBB82B99CEEAFF67" authorityName="Vachal" authorityYear="1911" box="[1117,1275,151,177]" class="Insecta" family="Halictidae" genus="Halictus" kingdom="Animalia" order="Hymenoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="repandirostris">
<emphasis id="EAA6EAC9AD41B570FBB82B99CEEAFF67" box="[1117,1275,151,177]" italics="true" pageId="47" pageNumber="48">repandirostris</emphasis>
</taxonomicName>
species group found away from tropical forests, where other species of the group occur. Noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="90C86550AD41B570FF222BD1CE82FECB" pageId="47" pageNumber="48" type="materials_examined">
<paragraph id="D86D36DBAD41B570FF222BD1CE82FECB" blockId="47.[151,1436,151,287]" pageId="47" pageNumber="48">
<emphasis id="EAA6EAC9AD41B570FF222BD1CB67FF2E" bold="true" box="[199,374,222,249]" pageId="47" pageNumber="48">Type material.</emphasis>
<materialsCitation id="68BA3C86AD41B570FE9B2BD1CE82FECB" ID-GBIF-Occurrence="3009686334" collectingDate="2007-02-24" collectionCode="DZUP" collectorName="A. Martins &amp; J. Gillung" country="Brazil" county="Balsa Nova" latitude="-25.469723" location="Rio" longLatPrecision="21" longitude="-49.769722" municipality="Campos" pageId="47" pageNumber="48" specimenCount="1" specimenCount-female="1" stateProvince="Parana" typeStatus="holotype">
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,
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\565271” “
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,
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\
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próximos ao
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\
<location id="DD0D6000AD41B570FF722A0DCB1AFECB" LSID="urn:lsid:plazi:treatment:507B87CDAD43B570FF72292CCE82FECB:DD0D6000AD41B570FF722A0DCB1AFECB" box="[151,267,259,285]" country="Brazil" county="Balsa Nova" latitude="-25.469723" longLatPrecision="21" longitude="-49.769722" municipality="Campos" name="Papagaios" pageId="47" pageNumber="48" stateProvince="Parana">Papagaios</location>
,
<date id="AC6C101BAD41B570FEF22A0DCB9AFECB" box="[279,395,259,285]" pageId="47" pageNumber="48" value="2007-02-24">
<collectingDate id="BC28E9F3AD41B570FEF22A0DCB9AFECB" box="[279,395,259,285]" pageId="47" pageNumber="48" value="2007-02-24">
24.ii.
<quantity id="1F2A9B3EAD41B570FEA82A0DCB9AFECB" box="[333,395,259,285]" metricMagnitude="6" metricUnit="m" metricValue="2.007943" pageId="47" pageNumber="48" unit="m" value="2007943.0">2007</quantity>
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, 943m\
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e
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, A.\Aguiar,
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,
<collectorName id="7527530DAD41B570FBF42A0DCE94FECB" box="[1041,1157,259,285]" pageId="47" pageNumber="48">J. Gillung</collectorName>
”.
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</paragraph>
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</treatment>
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