treatments-xml/data/4D/4A/93/4D4A9349FFEB392D1AD2FDEEFAC1FD64.xml

<document id="3BBCFE93ABD1FFE6993DCA780ADAA909" ID-CLB-Dataset="20928" ID-DOI="10.11646/zootaxa.5093.4.4" ID-GBIF-Dataset="d6297960-52dd-4b0a-a51b-05bda0e7d69f" ID-ISSN="1175-5326" ID-Zenodo-Dep="5912781" ID-ZooBank="4FE93B11-0714-400F-B9A6-41A54E2FCDC9" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1643354449005" checkinUser="plazi" docAuthor="Spinelli, Gustavo R., Ronderos, Maria M. &amp; Grogan, William L." docDate="2022" docId="4D4A9349FFEB392D1AD2FDEEFAC1FD64" docLanguage="en" docName="zootaxa.5093.4.4.pdf" docOrigin="Zootaxa 5093 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Macrurohelea sirii Spinelli &amp; Ronderos &amp; Grogan 2022, n. sp." docType="treatment" docVersion="7" lastPageNumber="458" masterDocId="B173EB31FFE739201A45FFD5FFFBFF95" masterDocTitle="Five new species in the predaceous midge genus Macrurohelea Ingram &amp; Macfie from Argentina, and descriptions of the previously unknown males of M. kuscheli Wirth and M. monotheca Spinelli &amp; Grogan (Diptera: Ceratopogonidae)" masterLastPageNumber="464" masterPageNumber="445" pageNumber="457" updateTime="1699304617883" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="9CAE82D0AD84755A5B72DBDF2BE02B7E">Five new species in the predaceous midge genus Macrurohelea Ingram &amp; Macfie from Argentina, and descriptions of the previously unknown males of M. kuscheli Wirth and M. monotheca Spinelli &amp; Grogan (Diptera: Ceratopogonidae)</mods:title>
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<mods:namePart id="E245194A209822E4E9337F9E02661321">Spinelli, Gustavo R.</mods:namePart>
<mods:affiliation id="73E6EAD654FF62E191BE3EF69A3339A3">División Entomología, Museo de La Plata, Paseo del Bosque s / n, 1900 La Plata, Argentina. &amp; Instituto de Limnología “ Dr. Raúl A. Ringuelet ” (Consejo Nacional de Investigaciones Científicas y Técnicas de la Argentina - Universidad Nacional de La Plata), Boulevard 120 y 62 s / n, 1900 La Plata, Buenos Aires, Argentina.</mods:affiliation>
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<mods:namePart id="B1A6D59765409CB30EB92E2A833C4024">Ronderos, Maria M.</mods:namePart>
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<mods:affiliation id="2A9278B7E1B9863264007CF06029DBA4">División Entomología, Museo de La Plata, Paseo del Bosque s / n, 1900 La Plata, Argentina. &amp; Centro de Estudios Parasitológicos y de Vectores (Consejo Nacional de Investigaciones Científicas y Técnicas de la Argentina - Universidad Nacional de La Plata), Boulevard 120 y 62 s / n, 1900 La Plata, Buenos Aires, Argentina. &amp; ronderos @ fcnym. unlp. edu. ar; https: // orcid. org / 0000 - 0001 - 8322 - 1471</mods:affiliation>
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<mods:affiliation id="259F7A6EBCD67F91E3463BD341C9284F">Florida State Collection of Arthropods, Florida Department of Agriculture and Consumer Services, Gainesville, Florida 32614 - 7100 USA. &amp; William. Grogan @ fdacs. gov; https: // orcid. org / 0000 - 0002 - 6475 - 8692</mods:affiliation>
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<emphasis id="F797FE4DFFEB392C1AD2FDEEFE81FDC3" bold="true" box="[151,378,571,598]" italics="true" pageId="12" pageNumber="457">Macrurohelea sirii</emphasis>
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<paragraph id="C55C225FFFEB392C1ADBFDB5FE72FDEF" blockId="12.[151,1065,571,671]" box="[158,393,608,634]" pageId="12" pageNumber="457">
(
<figureCitation id="5DD83EDAFFEB392C1AE3FDB5FED4FDEF" box="[166,303,608,634]" captionStart="FIGURES 26–32" captionStartId="14.[151,264,1943,1967]" captionTargetBox="[211,1381,185,1920]" captionTargetId="figure-23@14.[206,1382,181,1920]" captionTargetPageId="14" captionText="FIGURES 26–32. Macrurohelea sirii n. sp. Allotype female, 26–29. Holotype male, 30–32. 26, 30 head. 27 thorax. 28, 31 wing. 29 tip of abdomen. 32 genitalia. Scale bars = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5912791" httpUri="https://zenodo.org/record/5912791/files/figure.png" pageId="12" pageNumber="457">Figs. 26–32</figureCitation>
, 
<figureCitation id="5DD83EDAFFEB392C1B7FFDB5FE7AFDEF" box="[314,385,608,634]" captionStart="FIGURES 47–50" captionStartId="18.[151,264,1227,1251]" captionTargetBox="[250,1333,183,1189]" captionTargetId="figure-23@18.[242,1346,181,1203]" captionTargetPageId="18" captionText="FIGURES 47–50. Macrurohelea sirii n. sp. Holotype male, 47–48. Macrurohelea ventanensis n. sp. Holotype male, 49–50. 47, 49 parameres. 48, 50 aedeagus." figureDoi="http://doi.org/10.5281/zenodo.5912801" httpUri="https://zenodo.org/record/5912801/files/figure.png" pageId="12" pageNumber="457">47–48</figureCitation>
)
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Zoobank 
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urn:lsid:zoobank.org:act: 
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<subSubSection id="8DF971D4FFEB392C1AD2FD19FC58F863" pageId="12" pageNumber="457" type="diagnosis">
<paragraph id="C55C225FFFEB392C1AD2FD19FC93FC2B" blockId="12.[151,1437,716,2038]" pageId="12" pageNumber="457">
<emphasis id="F797FE4DFFEB392C1AD2FD19FEE9FD73" bold="true" box="[151,274,716,742]" pageId="12" pageNumber="457">Diagnosis.</emphasis>
Females: the only species with flagellomeres 2–8 short, barrel-shaped, 9–12 vasiform, very slightly elongate, 13 longest; thorax dark brown; wing membrane hyaline, veins brown, cell r 
<subScript id="5967201AFFEB392C1E1AFCD7FB9CFC85" attach="left" box="[1119,1127,770,784]" fontSize="6" pageId="12" pageNumber="457">3</subScript>
with faint intercalary vein; halter pale; posteromedian excavation of sternite 8 heavily sclerotized, dark brown; two round, slightly unequal-size spermathecae with short, slender necks. Males: the only species with tergite 10 with lateral margins convergent, apex truncate; apicolateral process minute with medium-size seta; distal portion of parameres slender, slightly divergent distally, apices tapered with outer pointed tip; aedeagus broadly Y-shaped, with a stout, curved medial basal arch and apices short, slender, with adpressed pointed tips.
</paragraph>
<paragraph id="C55C225FFFEB392C1A82FC1DFC73FA4F" blockId="12.[151,1437,716,2038]" pageId="12" pageNumber="457">
<emphasis id="F797FE4DFFEB392C1A82FC1DFEDEFC77" bold="true" box="[199,293,968,994]" pageId="12" pageNumber="457">Female.</emphasis>
Head (
<figureCitation id="5DD83EDAFFEB392C1B37FC1DFE3AFC76" box="[370,449,968,995]" captionStart="FIGURES 26–32" captionStartId="14.[151,264,1943,1967]" captionTargetBox="[211,1381,185,1920]" captionTargetId="figure-23@14.[206,1382,181,1920]" captionTargetPageId="14" captionText="FIGURES 26–32. Macrurohelea sirii n. sp. Allotype female, 26–29. Holotype male, 30–32. 26, 30 head. 27 thorax. 28, 31 wing. 29 tip of abdomen. 32 genitalia. Scale bars = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5912791" httpUri="https://zenodo.org/record/5912791/files/figure.png" pageId="12" pageNumber="457">Fig. 26</figureCitation>
) dark brown. Eyes separated medially by width of 2 ommatidia with short interommatidial spicules. Antennal flagellum uniformly light brown; flagellomere 1 massive with pair of apical sensilla coeloconica; flagellomeres 2–8 short, barrel-shaped, 9–12 slightly elongate, vasiform, 13 longest, broader than 2–12; antennal ratio 0.74–0.93 (0.84, n = 4). Palpus elongate, light brown; segment 3 with well developed subapical sensory pit; segment 4 short, segment 5 slightly longer than segment 3; palpal ratio 2.00 (n = 4). Mandible with 8–10 coarse teeth. Thorax (
<figureCitation id="5DD83EDAFFEB392C1AB2FBA9FEB2FB03" box="[247,329,1148,1174]" captionStart="FIGURES 26–32" captionStartId="14.[151,264,1943,1967]" captionTargetBox="[211,1381,185,1920]" captionTargetId="figure-23@14.[206,1382,181,1920]" captionTargetPageId="14" captionText="FIGURES 26–32. Macrurohelea sirii n. sp. Allotype female, 26–29. Holotype male, 30–32. 26, 30 head. 27 thorax. 28, 31 wing. 29 tip of abdomen. 32 genitalia. Scale bars = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5912791" httpUri="https://zenodo.org/record/5912791/files/figure.png" pageId="12" pageNumber="457">Fig. 27</figureCitation>
) uniformly dark brown; scutum with 3 stout prealar setae, one postalar seta; scutellum with 3 stout setae. Legs uniformly medium brown; palisade setae on tarsomere 1 of hind leg; hind tarsal ratio 2.12–2.33 (2.21, n = 4); tarsomeres 4 cordiform; tarsomeres 5 with pair of small, slightly curved tarsal claws. Wing (
<figureCitation id="5DD83EDAFFEB392C1E8DFB11FAE3FB4B" box="[1224,1304,1220,1246]" captionStart="FIGURES 26–32" captionStartId="14.[151,264,1943,1967]" captionTargetBox="[211,1381,185,1920]" captionTargetId="figure-23@14.[206,1382,181,1920]" captionTargetPageId="14" captionText="FIGURES 26–32. Macrurohelea sirii n. sp. Allotype female, 26–29. Holotype male, 30–32. 26, 30 head. 27 thorax. 28, 31 wing. 29 tip of abdomen. 32 genitalia. Scale bars = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5912791" httpUri="https://zenodo.org/record/5912791/files/figure.png" pageId="12" pageNumber="457">Fig. 28</figureCitation>
) membrane hyaline with numerous microtrichiae; anterior veins dark brown, posterior veins light brown; costa with long setae; second radial cell 1.80–2.63 (2.20, n = 4)x longer than first; cell r 
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with very faint intercalary vein; r-m crossvein as long as petiole of M; medium-size macrotrichiae on costa; wing length 0.94–1.21 (1.09, n = 4) mm, width 0.42–0.54 (0.49, n = 4) mm; costal ratio 0.72–0.75 (0.73, n = 4). Halter pale. Abdomen (
<figureCitation id="5DD83EDAFFEB392C1E54FA81FB9FFAFB" box="[1041,1124,1364,1390]" captionStart="FIGURES 26–32" captionStartId="14.[151,264,1943,1967]" captionTargetBox="[211,1381,185,1920]" captionTargetId="figure-23@14.[206,1382,181,1920]" captionTargetPageId="14" captionText="FIGURES 26–32. Macrurohelea sirii n. sp. Allotype female, 26–29. Holotype male, 30–32. 26, 30 head. 27 thorax. 28, 31 wing. 29 tip of abdomen. 32 genitalia. Scale bars = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5912791" httpUri="https://zenodo.org/record/5912791/files/figure.png" pageId="12" pageNumber="457">Fig. 29</figureCitation>
) medium brown. Segments 9–10 elongated, bent forward ventrally; sternite 8 stout with broad, rounded posteromedian excavation with heavily sclerotized borders. Two round, slightly unequal-size spermathecae with short, slender necks, diameters 44–56 (49, n = 3) and 40–52 (45, n = 3) µm, necks approximately 10 µm long.
</paragraph>
<paragraph id="C55C225FFFEB392C1A82FA31FC58F863" blockId="12.[151,1437,716,2038]" pageId="12" pageNumber="457">
<emphasis id="F797FE4DFFEB392C1A82FA31FEF0FA6B" bold="true" box="[199,267,1508,1534]" pageId="12" pageNumber="457">Male.</emphasis>
Coloration similar to female with the usual sexual differences. Head (
<figureCitation id="5DD83EDAFFEB392C1E6EFA31FB7AFA6B" box="[1067,1153,1508,1534]" captionStart="FIGURES 26–32" captionStartId="14.[151,264,1943,1967]" captionTargetBox="[211,1381,185,1920]" captionTargetId="figure-23@14.[206,1382,181,1920]" captionTargetPageId="14" captionText="FIGURES 26–32. Macrurohelea sirii n. sp. Allotype female, 26–29. Holotype male, 30–32. 26, 30 head. 27 thorax. 28, 31 wing. 29 tip of abdomen. 32 genitalia. Scale bars = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5912791" httpUri="https://zenodo.org/record/5912791/files/figure.png" pageId="12" pageNumber="457">Fig. 30</figureCitation>
). Eyes broadly separated medially by width of 3 ommatidia. Antennal flagellum with flagellomeres distinctly separated; antennal ratio 0.51–0.59 (0.56, n =6). Palpus slender, elongate, segment 3 slightly longer than segment 5. Tarsal claws small, nearly straight with bifid tips. Wing (
<figureCitation id="5DD83EDAFFEB392C180CF985FD66F9FF" box="[585,669,1616,1642]" captionStart="FIGURES 26–32" captionStartId="14.[151,264,1943,1967]" captionTargetBox="[211,1381,185,1920]" captionTargetId="figure-23@14.[206,1382,181,1920]" captionTargetPageId="14" captionText="FIGURES 26–32. Macrurohelea sirii n. sp. Allotype female, 26–29. Holotype male, 30–32. 26, 30 head. 27 thorax. 28, 31 wing. 29 tip of abdomen. 32 genitalia. Scale bars = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5912791" httpUri="https://zenodo.org/record/5912791/files/figure.png" pageId="12" pageNumber="457">Fig. 31</figureCitation>
) veins brown; 2 
<superScript id="32968F17FFEB392C1919F985FC97F9CB" attach="left" box="[860,876,1616,1630]" fontSize="6" pageId="12" pageNumber="457">nd</superScript>
radial cell 1.22–1.41 (1.32, n = 7)x longer than 1st; a very faint intercalary vein in cell r 
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; wing length 0.86–1.06 (1.00, n = 7) mm, width 0.32–0.42 (0.40, n = 7) mm; costal ratio 0.63–0.70 (0.67, n = 7). Abdomen brown. Genitalia (
<figureCitation id="5DD83EDAFFEB392C19FFF94DFBE2F927" box="[954,1049,1688,1714]" captionStart="FIGURES 26–32" captionStartId="14.[151,264,1943,1967]" captionTargetBox="[211,1381,185,1920]" captionTargetId="figure-23@14.[206,1382,181,1920]" captionTargetPageId="14" captionText="FIGURES 26–32. Macrurohelea sirii n. sp. Allotype female, 26–29. Holotype male, 30–32. 26, 30 head. 27 thorax. 28, 31 wing. 29 tip of abdomen. 32 genitalia. Scale bars = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5912791" httpUri="https://zenodo.org/record/5912791/files/figure.png" pageId="12" pageNumber="457">Figs. 32</figureCitation>
, 
<figureCitation id="5DD83EDAFFEB392C1E62F94DFB8AF927" box="[1063,1137,1688,1714]" captionStart="FIGURES 47–50" captionStartId="18.[151,264,1227,1251]" captionTargetBox="[250,1333,183,1189]" captionTargetId="figure-23@18.[242,1346,181,1203]" captionTargetPageId="18" captionText="FIGURES 47–50. Macrurohelea sirii n. sp. Holotype male, 47–48. Macrurohelea ventanensis n. sp. Holotype male, 49–50. 47, 49 parameres. 48, 50 aedeagus." figureDoi="http://doi.org/10.5281/zenodo.5912801" httpUri="https://zenodo.org/record/5912801/files/figure.png" pageId="12" pageNumber="457">47–48</figureCitation>
). Tergite 9 extending 0.55 length of gonocoxite, abruptly tapering at mid-length with rounded apex. Tergite 10 with lateral margins slightly convergent, apex truncate, extending 0.70 length of gonocoxite; apicolateral process minute, with medium-size seta; cercus slender, finger-like, slightly divergent. Sternite 9 2.5x broader than long, posteromedian margin slightly convex. Gonocoxite stout, 1.60x longer than sub-basal width, narrowing gradually distally; gonostylus slender, 0.75 length of gonocoxite, curved distally, tip pointed. Parameres (
<figureCitation id="5DD83EDAFFEB392C1902F899FC63F8F3" box="[839,920,1868,1894]" captionStart="FIGURES 47–50" captionStartId="18.[151,264,1227,1251]" captionTargetBox="[250,1333,183,1189]" captionTargetId="figure-23@18.[242,1346,181,1203]" captionTargetPageId="18" captionText="FIGURES 47–50. Macrurohelea sirii n. sp. Holotype male, 47–48. Macrurohelea ventanensis n. sp. Holotype male, 49–50. 47, 49 parameres. 48, 50 aedeagus." figureDoi="http://doi.org/10.5281/zenodo.5912801" httpUri="https://zenodo.org/record/5912801/files/figure.png" pageId="12" pageNumber="457">Fig. 47</figureCitation>
) separate; basal apodeme very stout, recurved 120°, with short, heavily sclerotized, posteriorly directed basal lobe; distal portions slender, closely approximated anteriorly, divergent at mid-length, each half tapering slightly distally with pointed, outer tip. Aedeagus (
<figureCitation id="5DD83EDAFFEB392C1F04F841FA6FF83B" box="[1345,1428,1940,1966]" captionStart="FIGURES 47–50" captionStartId="18.[151,264,1227,1251]" captionTargetBox="[250,1333,183,1189]" captionTargetId="figure-23@18.[242,1346,181,1203]" captionTargetPageId="18" captionText="FIGURES 47–50. Macrurohelea sirii n. sp. Holotype male, 47–48. Macrurohelea ventanensis n. sp. Holotype male, 49–50. 47, 49 parameres. 48, 50 aedeagus." figureDoi="http://doi.org/10.5281/zenodo.5912801" httpUri="https://zenodo.org/record/5912801/files/figure.png" pageId="12" pageNumber="457">Fig. 48</figureCitation>
) broadly Y-shaped, 1.3x broader than long, with stout, slightly concave basal arch, which extends 1/3 of total length; distal extensions short, slender, tapering slightly distally, tips pointed.
</paragraph>
</subSubSection>
<subSubSection id="8DF971D4FFEA392D1A82FF42FE23FE44" pageId="13" pageNumber="458" type="materials_examined">
<paragraph id="C55C225FFFEA392D1A82FF42FF1BFEF0" blockId="13.[151,1437,150,753]" pageId="13" pageNumber="458">
<emphasis id="F797FE4DFFEA392D1A82FF42FE8DFF25" bold="true" box="[199,374,150,177]" pageId="13" pageNumber="458">Type material</emphasis>
. 
<materialsCitation id="758B2802FFEA392D1BCEFF42FC6CFF6C" ID-GBIF-Occurrence="3458732313" collectingDate="2007-12-12" collectingDateMax="2008-01-03" collectingDateMin="2007-12-12" collectionCode="MLPA" collectorName="A. Garre &amp; F. Montes de Oca" country="Argentina" elevation="762" latitude="-40.68986" location="Montes de Oca" longLatPrecision="1" longitude="-71.29256" municipality="rio Minero" pageId="13" pageNumber="458" specimenCount="3" stateProvince="Neuquen" typeStatus="holotype">
<typeStatus id="1A589CFDFFEA392D1BCEFF42FE08FF24" box="[395,499,151,177]" pageId="13" pageNumber="458" type="holotype">Holotype</typeStatus>
male, labeled “ 
<typeStatus id="1A589CFDFFEA392D18FDFF42FCD8FF24" box="[696,803,151,177]" pageId="13" pageNumber="458" type="holotype">Holotype</typeStatus>
<taxonomicName id="02E359DCFFEA392D1977FF42FA75FF24" authority="Spinelli, Ronderos and Grogan" authorityName="Spinelli, Ronderos and Grogan" box="[818,1422,151,177]" class="Insecta" family="Ceratopogonidae" genus="Macrurohelea" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="458" phylum="Arthropoda" rank="species" species="sirii">Macrurohelea sirii Spinelli, Ronderos and Grogan</taxonomicName>
”, “ 
<collectingCountry id="BDF462CFFFEA392D1AE4FF6EFEEDFF40" box="[161,278,187,213]" name="Argentina" pageId="13" pageNumber="458">Argentina</collectingCountry>
, 
<collectingRegion id="0727ECBDFFEA392D1B66FF6EFE70FF40" box="[291,395,187,213]" country="Argentina" name="Neuquen" pageId="13" pageNumber="458">Neuquén</collectingRegion>
, Parque Nacional Nahuel Huapi, arroyo León, 
<geoCoordinate id="A0D74498FFEA392D19E3FF6EFBD5FF40" box="[934,1070,187,213]" degrees="40" direction="south" minutes="43" orientation="latitude" pageId="13" pageNumber="458" precision="15" seconds="49" value="-40.730278">40º43’49’’S</geoCoordinate>
, 
<geoCoordinate id="A0D74498FFEA392D1E79FF6EFB1AFF40" box="[1084,1249,187,213]" degrees="71" direction="west" minutes="09" orientation="longitude" pageId="13" pageNumber="458" precision="1" seconds="11.6" value="-71.15322">71º09’11.6’’W</geoCoordinate>
, 
<quantity id="021B8FBAFFEA392D1EABFF6EFAC2FF40" box="[1262,1337,187,213]" metricMagnitude="2" metricUnit="m" metricValue="7.62" pageId="13" pageNumber="458" unit="m" value="762.0">
<elevation id="4ECEC56CFFEA392D1EABFF6EFAC2FF40" box="[1262,1337,187,213]" metricMagnitude="2" metricUnit="m" metricValue="7.62" pageId="13" pageNumber="458" unit="m" value="762.0">762 m</elevation>
</quantity>
, 
<date id="B15D049FFFEA392D1F02FF6EFF29FF6C" pageId="13" pageNumber="458" value="2007-12-12">
<collectingDate id="A119FD77FFEA392D1F02FF6EFF29FF6C" pageId="13" pageNumber="458" value="2007-12-12">12-XII- 2007</collectingDate>
</date>
/ 
<date id="B15D049FFFEA392D1A9EFF0AFEC7FF6C" box="[219,316,223,249]" pageId="13" pageNumber="458" value="2008-01-03">
<collectingDate id="A119FD77FFEA392D1A9EFF0AFEC7FF6C" box="[219,316,223,249]" pageId="13" pageNumber="458" value="2008-01-03">3-I-2008</collectingDate>
</date>
, 
<collectorName id="68164789FFEA392D1B02FF0AFE50FF6C" box="[327,427,223,249]" pageId="13" pageNumber="458">A. Garré</collectorName>
– 
<collectorName id="68164789FFEA392D1BFFFF0AFD7FFF6C" box="[442,644,223,249]" pageId="13" pageNumber="458">F. Montes de Oca</collectorName>
, 
<collectingMethod id="1CA25A48FFEA392D18D5FF0AFCE4FF6C" box="[656,799,223,249]" pageId="13" pageNumber="458">Malaise trap</collectingMethod>
” (
<collectionCode id="A3F2BA9AFFEA392D197CFF0AFC70FF6C" box="[825,907,223,249]" pageId="13" pageNumber="458">MLPA</collectionCode>
);
</materialsCitation>
<materialsCitation id="758B2802FFEA392D19DAFF0AFF3DFED4" ID-GBIF-Occurrence="3458732305" collectingDate="2007-12-12" collectingDateMax="2008-01-03" collectingDateMin="2007-12-12" collectionCode="MLPA" collectorName="A. Garre &amp; F. Montes de Oca" country="Argentina" elevation="762" latitude="-40.68986" location="Montes de Oca" longLatPrecision="1" longitude="-71.29256" municipality="rio Minero" pageId="13" pageNumber="458" specimenCount="3" stateProvince="Neuquen" typeStatus="allotype">
<typeStatus id="1A589CFDFFEA392D19DAFF0AFC02FF6C" box="[927,1017,223,249]" pageId="13" pageNumber="458" type="allotype">allotype</typeStatus>
female, 
<collectingCountry id="BDF462CFFFEA392D1E1EFF0AFB36FF6C" box="[1115,1229,223,249]" name="Argentina" pageId="13" pageNumber="458">Argentina</collectingCountry>
, 
<collectingRegion id="0727ECBDFFEA392D1E9FFF0AFAB9FF6C" box="[1242,1346,223,249]" country="Argentina" name="Neuquen" pageId="13" pageNumber="458">Neuquén</collectingRegion>
, Parque Nacional Nahuel Huapi, 
<collectingMunicipality id="2538B825FFEA392D1BF6FED6FDCBFE88" box="[435,560,259,285]" pageId="13" pageNumber="458">río Minero</collectingMunicipality>
, 
<geoCoordinate id="A0D74498FFEA392D1878FED7FD26FE88" box="[573,733,258,285]" degrees="40" direction="south" minutes="41" orientation="latitude" pageId="13" pageNumber="458" precision="1" seconds="23.5" value="-40.68986">40°41’23.5’’S</geoCoordinate>
, 
<geoCoordinate id="A0D74498FFEA392D18AEFED6FC68FE88" box="[747,915,259,285]" degrees="71" direction="west" minutes="17" orientation="longitude" pageId="13" pageNumber="458" precision="1" seconds="33.2" value="-71.29256">71°17’33.2’’W</geoCoordinate>
, 
<quantity id="021B8FBAFFEA392D19E4FED6FC10FE88" box="[929,1003,259,285]" metricMagnitude="2" metricUnit="m" metricValue="8.32" pageId="13" pageNumber="458" unit="m" value="832.0">
<elevation id="4ECEC56CFFEA392D19E4FED6FC10FE88" box="[929,1003,259,285]" metricMagnitude="2" metricUnit="m" metricValue="8.32" pageId="13" pageNumber="458" unit="m" value="832.0">832 m</elevation>
</quantity>
, 
<collectorName id="68164789FFEA392D19BDFED6FBA6FE88" box="[1016,1117,259,285]" pageId="13" pageNumber="458">A. Garré</collectorName>
– 
<collectorName id="68164789FFEA392D1E2EFED6FACCFE88" box="[1131,1335,259,285]" pageId="13" pageNumber="458">
F. 
<location id="C03C7484FFEA392D1EC2FED6FACCFE88" LSID="urn:lsid:plazi:treatment:4D4A9349FFEB392D1AD2FDEEFAC1FD64:C03C7484FFEA392D1EC2FED6FACCFE88" box="[1159,1335,259,285]" country="Argentina" latitude="-40.68986" longLatPrecision="1" longitude="-71.29256" municipality="rio Minero" name="Montes de Oca" pageId="13" pageNumber="458" stateProvince="Neuquen">Montes de Oca</location>
</collectorName>
, 
<collectingMethod id="1CA25A48FFEA392D1F01FED6FF3DFED4" pageId="13" pageNumber="458">Malaise trap</collectingMethod>
</materialsCitation>
. 
<materialsCitation id="758B2802FFEA392D1A94FEF2FC5DFED4" ID-GBIF-Occurrence="3458732311" box="[209,934,295,321]" collectingDate="2007-12-12" collectingDateMax="2008-01-03" collectingDateMin="2007-12-12" collectionCode="MLPA" collectorName="A. Garre &amp; F. Montes de Oca" country="Argentina" elevation="762" latitude="-40.68986" location="Montes de Oca" longLatPrecision="1" longitude="-71.29256" municipality="rio Minero" pageId="13" pageNumber="458" specimenCount="9" specimenCount-female="3" specimenCount-male="6" stateProvince="Neuquen" typeStatus="paratype">
<typeStatus id="1A589CFDFFEA392D1A94FEF2FEBAFED4" box="[209,321,295,321]" pageId="13" pageNumber="458" type="paratype">Paratypes</typeStatus>
, 
<specimenCount id="D3E5E9D6FFEA392D1B09FEF2FE5EFED4" box="[332,421,295,321]" pageId="13" pageNumber="458" type="male">6 males</specimenCount>
, 
<specimenCount id="D3E5E9D6FFEA392D1BF5FEF2FDDBFED4" box="[432,544,295,321]" pageId="13" pageNumber="458" type="female">3 females</specimenCount>
, as follows: same data as holotype
</materialsCitation>
, 
<materialsCitation id="758B2802FFEA392D19F4FEF2FA78FED4" ID-GBIF-Occurrence="3458732303" box="[945,1411,295,321]" collectingDate="2007-12-12" collectingDateMax="2008-01-03" collectingDateMin="2007-12-12" collectionCode="MLPA" collectorName="A. Garre &amp; F. Montes de Oca" country="Argentina" elevation="762" latitude="-40.68986" location="Montes de Oca" longLatPrecision="1" longitude="-71.29256" municipality="rio Minero" pageId="13" pageNumber="458" specimenCount="7" specimenCount-female="3" specimenCount-male="4" stateProvince="Neuquen" typeStatus="paratype">
<specimenCount id="D3E5E9D6FFEA392D19F4FEF2FBF1FED4" box="[945,1034,295,321]" pageId="13" pageNumber="458" type="male">4 males</specimenCount>
, 
<specimenCount id="D3E5E9D6FFEA392D1E53FEF2FB7DFED4" box="[1046,1158,295,321]" pageId="13" pageNumber="458" type="female">3 females</specimenCount>
; same data as allotype
</materialsCitation>
, 
<specimenCount id="D3E5E9D6FFEA392D1FCBFEF2FF27FEF0" pageId="13" pageNumber="458" type="male">2 males</specimenCount>
.
</paragraph>
<paragraph id="C55C225FFFEA392D1A82FEBBFE23FE44" blockId="13.[151,1437,150,753]" pageId="13" pageNumber="458">
<emphasis id="F797FE4DFFEA392D1A82FEBBFDDDFE1D" bold="true" box="[199,550,366,392]" pageId="13" pageNumber="458">Derivation of specific epithet.</emphasis>
We are pleased to name this new species after Dr. Augusto Siri of the Instituto de Limnología “Dr. Raúl A. Ringuelet” La Plata, 
<collectingCountry id="BDF462CFFFEA392D188EFE46FCC6FE38" box="[715,829,403,429]" name="Argentina" pageId="13" pageNumber="458">Argentina</collectingCountry>
, for his friendship and companionship during several collecting trips to Patagonia.
</paragraph>
</subSubSection>
<subSubSection id="8DF971D4FFEA392D1A82FE0FFC2BFE60" box="[199,976,474,501]" pageId="13" pageNumber="458" type="distribution">
<paragraph id="C55C225FFFEA392D1A82FE0FFC2BFE60" blockId="13.[151,1437,150,753]" box="[199,976,474,501]" pageId="13" pageNumber="458">
<emphasis id="F797FE4DFFEA392D1A82FE0FFE9AFE61" bold="true" box="[199,353,474,500]" pageId="13" pageNumber="458">Distribution.</emphasis>
<collectingCountry id="BDF462CFFFEA392D1B22FE0EFE22FE60" box="[359,473,475,501]" name="Argentina" pageId="13" pageNumber="458">Argentina</collectingCountry>
, in 
<taxonomicName id="02E359DCFFEA392D1847FE0EFD78FE60" box="[514,643,475,501]" class="Magnoliopsida" family="Nothofagaceae" genus="Nothofagus" kingdom="Plantae" order="Fagales" pageId="13" pageNumber="458" phylum="Tracheophyta" rank="genus">
<emphasis id="F797FE4DFFEA392D1847FE0EFD78FE60" box="[514,643,475,501]" italics="true" pageId="13" pageNumber="458">Nothofagus</emphasis>
</taxonomicName>
forests of 
<collectingRegion id="0727ECBDFFEA392D18BFFE0EFC37FE60" box="[762,972,475,501]" country="Argentina" name="Neuquen" pageId="13" pageNumber="458">Neuquén province</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="8DF971D4FFEA392D1A82FE2BFAC1FD64" pageId="13" pageNumber="458" type="discussion">
<paragraph id="C55C225FFFEA392D1A82FE2BFAC1FD64" blockId="13.[151,1437,150,753]" pageId="13" pageNumber="458">
<emphasis id="F797FE4DFFEA392D1A82FE2BFEB0FD8D" bold="true" box="[199,331,510,536]" pageId="13" pageNumber="458">Discussion.</emphasis>
Females of this new species are very similar to those of 
<taxonomicName id="02E359DCFFEA392D1989FE2AFAF5FD8C" authority="Spinelli &amp; Grogan" authorityName="Spinelli &amp; Grogan" authorityYear="1984" box="[972,1294,511,537]" class="Insecta" family="Ceratopogonidae" genus="Macrurohelea" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="458" phylum="Arthropoda" rank="species" species="wirthi">
<emphasis id="F797FE4DFFEA392D1989FE2AFC18FD8D" box="[972,995,511,536]" italics="true" pageId="13" pageNumber="458">M</emphasis>
. 
<emphasis id="F797FE4DFFEA392D19B7FE2AFBCFFD8C" box="[1010,1076,511,537]" italics="true" pageId="13" pageNumber="458">wirthi</emphasis>
Spinelli &amp; Grogan
</taxonomicName>
, from which they can be distinguished by the distinctly longer distal flagellomeres in 
<taxonomicName id="02E359DCFFEA392D19FFFDF6FBDAFDA8" authorityName="Spinelli &amp; Grogan" authorityYear="1984" box="[954,1057,547,573]" class="Insecta" family="Ceratopogonidae" genus="Macrurohelea" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="458" phylum="Arthropoda" rank="species" species="wirthi">
<emphasis id="F797FE4DFFEA392D19FFFDF6FC2AFDA9" box="[954,977,547,572]" italics="true" pageId="13" pageNumber="458">M</emphasis>
. 
<emphasis id="F797FE4DFFEA392D199AFDF6FBDAFDA8" box="[991,1057,547,573]" italics="true" pageId="13" pageNumber="458">wirthi</emphasis>
</taxonomicName>
and by the intercalary vein of cell r 
<subScript id="5967201AFFEA392D1AE5FD8CFF53FDF2" attach="left" box="[160,168,601,615]" fontSize="6" pageId="13" pageNumber="458">3</subScript>
that is fainter in 
<taxonomicName id="02E359DCFFEA392D1B27FD92FE32FDF4" authorityName="Spinelli &amp; Grogan" authorityYear="1984" box="[354,457,583,609]" class="Insecta" family="Ceratopogonidae" genus="Macrurohelea" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="458" phylum="Arthropoda" rank="species" species="wirthi">
<emphasis id="F797FE4DFFEA392D1B27FD92FE82FDF5" box="[354,377,583,608]" italics="true" pageId="13" pageNumber="458">M</emphasis>
. 
<emphasis id="F797FE4DFFEA392D1BC2FD92FE32FDF4" box="[391,457,583,609]" italics="true" pageId="13" pageNumber="458">wirthi</emphasis>
</taxonomicName>
. Males resemble males of 
<taxonomicName id="02E359DCFFEA392D18B6FD92FBCCFDF4" authority="Spinelli &amp; Grogan" authorityName="Spinelli &amp; Grogan" authorityYear="1990" box="[755,1079,583,609]" class="Insecta" family="Ceratopogonidae" genus="Macrurohelea" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="458" phylum="Arthropoda" rank="species" species="similis">
<emphasis id="F797FE4DFFEA392D18B6FD92FCF1FDF5" box="[755,778,583,608]" italics="true" pageId="13" pageNumber="458">M</emphasis>
. 
<emphasis id="F797FE4DFFEA392D195DFD92FC9BFDF4" box="[792,864,583,609]" italics="true" pageId="13" pageNumber="458">similis</emphasis>
Spinelli &amp; Grogan
</taxonomicName>
, but in that species the posterior margin of sternite 9 is straight, the gonocoxite is curved, and the aedeagus is triangular, and its distal portion tapers to a moderately pointed tip. It is also similar to 
<taxonomicName id="02E359DCFFEA392D18C7FD5AFCFAFD3D" authority="Spinelli &amp; Ronderos &amp; Grogan, 2022" authorityName="Spinelli &amp; Ronderos &amp; Grogan" authorityYear="2022" box="[642,769,655,681]" class="Insecta" family="Ceratopogonidae" genus="Macrurohelea" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="458" phylum="Arthropoda" rank="species" species="morenoi" status="sp. nov.">
<emphasis id="F797FE4DFFEA392D18C7FD5AFD62FD3D" box="[642,665,655,680]" italics="true" pageId="13" pageNumber="458">M</emphasis>
. 
<emphasis id="F797FE4DFFEA392D18E3FD5AFCFAFD3D" box="[678,769,655,680]" italics="true" pageId="13" pageNumber="458">morenoi</emphasis>
</taxonomicName>
<emphasis id="F797FE4DFFEA392D194DFD5AFCBEFD3C" bold="true" box="[776,837,655,681]" pageId="13" pageNumber="458">
<taxonomicNameLabel id="ECA44336FFEA392D194DFD5AFCBEFD3C" box="[776,837,655,681]" pageId="13" pageNumber="458" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, but in that species the base of the aedeagus is broader than the bases of parameres and the distal portion is triangular and divided medially with apices slightly divergent, and the distal portion of parameres is stout, nearly straight and slightly divergent distally with rounded tips.
</paragraph>
</subSubSection>
</treatment>
</document>