treatments-xml/data/4D/4A/93/4D4A9349FFE639221AD2FB7EFA67F948.xml

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<mods:title id="150504041F626ACC5AEC2B153D4C22D5">Five new species in the predaceous midge genus Macrurohelea Ingram &amp; Macfie from Argentina, and descriptions of the previously unknown males of M. kuscheli Wirth and M. monotheca Spinelli &amp; Grogan (Diptera: Ceratopogonidae)</mods:title>
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<mods:namePart id="F5C0375394B6993D3ECDCFD6EB893605">Spinelli, Gustavo R.</mods:namePart>
<mods:affiliation id="6388CBFB0FF910442DA4319B90799393">División Entomología, Museo de La Plata, Paseo del Bosque s / n, 1900 La Plata, Argentina. &amp; Instituto de Limnología “ Dr. Raúl A. Ringuelet ” (Consejo Nacional de Investigaciones Científicas y Técnicas de la Argentina - Universidad Nacional de La Plata), Boulevard 120 y 62 s / n, 1900 La Plata, Buenos Aires, Argentina.</mods:affiliation>
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<mods:namePart id="9C852A80653A649603E72B702026F9BF">Ronderos, Maria M.</mods:namePart>
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<mods:affiliation id="6547BCD0D47BB39CCF206E837078367E">División Entomología, Museo de La Plata, Paseo del Bosque s / n, 1900 La Plata, Argentina. &amp; Centro de Estudios Parasitológicos y de Vectores (Consejo Nacional de Investigaciones Científicas y Técnicas de la Argentina - Universidad Nacional de La Plata), Boulevard 120 y 62 s / n, 1900 La Plata, Buenos Aires, Argentina. &amp; ronderos @ fcnym. unlp. edu. ar; https: // orcid. org / 0000 - 0001 - 8322 - 1471</mods:affiliation>
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<mods:namePart id="28FE1DEA389CC260F62FC30808384D26">Grogan, William L.</mods:namePart>
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<mods:affiliation id="8345A5BB695E1B6DF2679550D9CAC656">Florida State Collection of Arthropods, Florida Department of Agriculture and Consumer Services, Gainesville, Florida 32614 - 7100 USA. &amp; William. Grogan @ fdacs. gov; https: // orcid. org / 0000 - 0002 - 6475 - 8692</mods:affiliation>
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Key to 
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<emphasis id="F797FE4DFFE639211AB4FB7FFE65FB50" bold="true" box="[241,414,1194,1221]" italics="true" pageId="1" pageNumber="446">Macrurohelea</emphasis>
</taxonomicName>
of the Neotropical region
</emphasis>
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(females of 
<taxonomicName id="02E359DCFFE639211B5DFB1AFDACFB7C" authority="Spinelli &amp; Grogan" authorityName="Spinelli &amp; Grogan" authorityYear="1990" box="[280,599,1231,1257]" class="Insecta" family="Ceratopogonidae" genus="Macrurohelea" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="446" phylum="Arthropoda" rank="species" species="similis">
<emphasis id="F797FE4DFFE639211B5DFB1AFED4FB7D" box="[280,303,1231,1256]" italics="true" pageId="1" pageNumber="446">M</emphasis>
. 
<emphasis id="F797FE4DFFE639211B7EFB1AFE7FFB7C" box="[315,388,1231,1257]" italics="true" pageId="1" pageNumber="446">similis</emphasis>
Spinelli &amp; Grogan
</taxonomicName>
, 
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<emphasis id="F797FE4DFFE63921181AFB1AFD8DFB7D" box="[607,630,1231,1256]" italics="true" pageId="1" pageNumber="446">M</emphasis>
. 
<emphasis id="F797FE4DFFE6392118C7FB1AFD25FB7D" box="[642,734,1231,1256]" italics="true" pageId="1" pageNumber="446">morenoi</emphasis>
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<taxonomicNameLabel id="ECA44336FFE6392118A6FB1AFCDBFB7C" box="[739,800,1231,1257]" pageId="1" pageNumber="446" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
and 
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<emphasis id="F797FE4DFFE639211917FB1AFC92FB7D" box="[850,873,1231,1256]" italics="true" pageId="1" pageNumber="446">M</emphasis>
. 
<emphasis id="F797FE4DFFE639211930FB1AFC03FB7D" box="[885,1016,1231,1256]" italics="true" pageId="1" pageNumber="446">ventanensis</emphasis>
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<taxonomicNameLabel id="ECA44336FFE6392119B9FB1AFBC2FB7C" box="[1020,1081,1231,1257]" pageId="1" pageNumber="446" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, and the males of 
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<emphasis id="F797FE4DFFE639211EBEFB1AFAE9FB7D" box="[1275,1298,1231,1256]" italics="true" pageId="1" pageNumber="446">M</emphasis>
. 
<emphasis id="F797FE4DFFE639211F5BFB1AFA67FB7C" box="[1310,1436,1231,1257]" italics="true" pageId="1" pageNumber="446">fuscipennis</emphasis>
Spinelli &amp; Grogan
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and 
<taxonomicName id="02E359DCFFE639211BD9FB26FD20FA98" authority="Spinelli &amp; Grogan" authorityName="Spinelli &amp; Grogan" authorityYear="1984" box="[412,731,1267,1293]" class="Insecta" family="Ceratopogonidae" genus="Macrurohelea" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="446" phylum="Arthropoda" rank="species" species="wirthi">
<emphasis id="F797FE4DFFE639211BD9FB26FE48FA99" box="[412,435,1267,1292]" italics="true" pageId="1" pageNumber="446">M</emphasis>
. 
<emphasis id="F797FE4DFFE639211B87FB26FDFEFA98" box="[450,517,1267,1293]" italics="true" pageId="1" pageNumber="446">wirthi</emphasis>
Spinelli &amp; Grogan
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are unknown)
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<keyLead id="7E12816AFFE639211AD2FAEDFA67FAD8" box="[151,1436,1336,1358]" pageId="1" pageNumber="446">1. Females............................................................................................. 2</keyLead>
</paragraph>
<paragraph id="C55C225FFFE639211AD2FA80FA67FAFE" blockId="1.[151,1437,1336,2032]" box="[151,1436,1365,1387]" pageId="1" pageNumber="446">
<keyLead id="7E12816AFFE639211AD2FA80FA67FAFE" box="[151,1436,1365,1387]" pageId="1" pageNumber="446">- Males............................................................................................. 15</keyLead>
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<paragraph id="C55C225FFFE639211AD2FAA7FA67FA12" blockId="1.[151,1437,1336,2032]" box="[151,1436,1394,1416]" pageId="1" pageNumber="446">
<keyLead id="7E12816AFFE639211AD2FAA7FA67FA12" box="[151,1436,1394,1416]" pageId="1" pageNumber="446">2. One spermatheca......................................................................................3</keyLead>
</paragraph>
<paragraph id="C55C225FFFE639211AD2FA45FA67FA30" blockId="1.[151,1437,1336,2032]" box="[151,1436,1424,1446]" pageId="1" pageNumber="446">
<keyLead id="7E12816AFFE639211AD2FA45FA67FA30" box="[151,1436,1424,1446]" pageId="1" pageNumber="446">
- Two spermathecae (
<figureCitation id="5DD83EDAFFE639211BE7FA45FE1DFA30" box="[418,486,1424,1446]" captionStart="FIGURES 1–7" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[214,1376,188,1920]" captionTargetId="figure-23@4.[210,1378,181,1920]" captionTargetPageId="4" captionText="FIGURES 1–7. Macrurohelea bassoi n. sp. Allotype female, 1–4. Holotype male, 5–7. 1, 5 head. 2 thorax. 3, 6 wing. 4 tip of abdomen. 7 genitalia. Scale bars = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5912783" httpUri="https://zenodo.org/record/5912783/files/figure.png" pageId="1" pageNumber="446">Figs. 4</figureCitation>
, 
<figureCitation id="5DD83EDAFFE639211BB5FA45FDF2FA30" box="[496,521,1424,1445]" captionStart="FIGURES 8–14" captionStartId="6.[151,264,1943,1967]" captionTargetBox="[186,1400,183,1920]" captionTargetId="figure-23@6.[186,1401,181,1920]" captionTargetPageId="6" captionText="FIGURES 8–14. Macrurohelea donatoi n. sp. Allotype female, 8–11. Holotype male, 12–14. 8, 12 head. 9 thorax. 10, 13 wing. 11 tip of abdomen. 14 genitalia. Scale bars = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5912785" httpUri="https://zenodo.org/record/5912785/files/figure.png" pageId="1" pageNumber="446">11</figureCitation>
, 
<figureCitation id="5DD83EDAFFE639211856FA45FDD7FA33" box="[531,556,1424,1446]" captionStart="FIGURES 26–32" captionStartId="14.[151,264,1943,1967]" captionTargetBox="[211,1381,185,1920]" captionTargetId="figure-23@14.[206,1382,181,1920]" captionTargetPageId="14" captionText="FIGURES 26–32. Macrurohelea sirii n. sp. Allotype female, 26–29. Holotype male, 30–32. 26, 30 head. 27 thorax. 28, 31 wing. 29 tip of abdomen. 32 genitalia. Scale bars = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5912791" httpUri="https://zenodo.org/record/5912791/files/figure.png" pageId="1" pageNumber="446">29</figureCitation>
)....................................................................... 4
</keyLead>
</paragraph>
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<keyStep id="7E173AFAFFE639211AD2FA78FA67F98F" pageId="1" pageNumber="446">
<paragraph id="C55C225FFFE639211AD2FA78FA67FA4A" blockId="1.[151,1437,1336,2032]" pageId="1" pageNumber="446">
<keyLead id="7E12816AFFE639211AD2FA78FA67FA4A" pageId="1" pageNumber="446">
3. Large species (wing length 
<quantity id="021B8FBAFFE639211BAEFA78FDBFFA57" box="[491,580,1453,1475]" metricMagnitude="-3" metricUnit="m" metricValue="2.05" pageId="1" pageNumber="446" unit="mm" value="2.05">2.05 mm</quantity>
); wing with intercalary vein in cell r 
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; wing membrane infuscated, veins dark brown............................................................................. 
<taxonomicName id="02E359DCFFE639211E19FA1EFA67FA4A" ID-CoL="3XD2X" authority="Spinelli &amp; Grogan" authorityName="Spinelli &amp; Grogan" authorityYear="1984" box="[1116,1436,1482,1504]" class="Insecta" family="Ceratopogonidae" genus="Macrurohelea" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="446" phylum="Arthropoda" rank="species" species="monotheca">
<emphasis id="F797FE4DFFE639211E19FA1EFB8BFA75" box="[1116,1136,1483,1504]" italics="true" pageId="1" pageNumber="446">M</emphasis>
. 
<emphasis id="F797FE4DFFE639211E39FA1FFB1EFA4A" box="[1148,1253,1482,1503]" italics="true" pageId="1" pageNumber="446">monotheca</emphasis>
Spinelli &amp; Grogan
</taxonomicName>
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<paragraph id="C55C225FFFE639211AD2FA3DFA67F98F" blockId="1.[151,1437,1336,2032]" pageId="1" pageNumber="446">
<keyLead id="7E12816AFFE639211AD2FA3DFA67F98F" pageId="1" pageNumber="446">
- Medium-size species (wing length 
<quantity id="021B8FBAFFE639211873FA32FD3DFA68" box="[566,710,1511,1533]" metricMagnitude="-3" metricUnit="m" metricValue="1.665" metricValueMax="1.74" metricValueMin="1.59" pageId="1" pageNumber="446" unit="mm" value="1.665" valueMax="1.74" valueMin="1.59">1.59–1.74 mm</quantity>
); wing without intercalary vein in cell r 
<subScript id="5967201AFFE639211E13FA22FBA6F996" attach="left" box="[1110,1117,1527,1539]" fontSize="5" pageId="1" pageNumber="446">3</subScript>
; wing membrane hyaline, veins pale.......................................................................... 
<taxonomicName id="02E359DCFFE639211EC4F9D3FA67F98F" ID-CoL="3XD2T" authority="Spinelli &amp; Grogan" authorityName="Spinelli &amp; Grogan" authorityYear="1984" box="[1153,1436,1541,1563]" class="Insecta" family="Ceratopogonidae" genus="Macrurohelea" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="446" phylum="Arthropoda" rank="species" species="gentilii">
<emphasis id="F797FE4DFFE639211EC4F9D3FB6EF98E" box="[1153,1173,1542,1563]" italics="true" pageId="1" pageNumber="446">M</emphasis>
. 
<emphasis id="F797FE4DFFE639211EE4F9D0FB1EF98F" box="[1185,1253,1541,1562]" italics="true" pageId="1" pageNumber="446">gentilii</emphasis>
Spinelli &amp; Grogan
</taxonomicName>
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<keyStep id="7E173AFAFFE639211AD2F9F7FA67F9C0" pageId="1" pageNumber="446">
<paragraph id="C55C225FFFE639211AD2F9F7FA67F9AD" blockId="1.[151,1437,1336,2032]" box="[151,1436,1570,1592]" pageId="1" pageNumber="446">
<keyLead id="7E12816AFFE639211AD2F9F7FA67F9AD" box="[151,1436,1570,1592]" pageId="1" pageNumber="446">
4. Legs with numerous long bristly setae......................................................... 
<taxonomicName id="02E359DCFFE639211F41F9F6FA67F9AD" ID-CoL="3XD2Z" authority="Wirth" authorityName="Wirth" authorityYear="1965" box="[1284,1436,1570,1592]" class="Insecta" family="Ceratopogonidae" genus="Macrurohelea" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="446" phylum="Arthropoda" rank="species" species="setosa">
<emphasis id="F797FE4DFFE639211F41F9F6FAE3F9AD" box="[1284,1304,1571,1592]" italics="true" pageId="1" pageNumber="446">M</emphasis>
. 
<emphasis id="F797FE4DFFE639211F61F9F6FA9BF9AD" box="[1316,1376,1571,1592]" italics="true" pageId="1" pageNumber="446">setosa</emphasis>
Wirth
</taxonomicName>
</keyLead>
</paragraph>
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<keyLead id="7E12816AFFE639211AD2F995FA67F9C0" box="[151,1436,1599,1622]" pageId="1" pageNumber="446">- Legs with vestiture of short setae......................................................................... 5</keyLead>
</paragraph>
</keyStep>
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<paragraph id="C55C225FFFE639211AD2F988FA67F9E6" blockId="1.[151,1437,1336,2032]" box="[151,1436,1629,1651]" pageId="1" pageNumber="446">
<keyLead id="7E12816AFFE639211AD2F988FA67F9E6" box="[151,1436,1629,1651]" pageId="1" pageNumber="446">
5. Large species (wing length 
<quantity id="021B8FBAFFE639211BAEF988FDBFF9E7" box="[491,580,1629,1650]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="1" pageNumber="446" unit="mm" value="2.7">2.70 mm</quantity>
); anal lobe of wing well developed, broad............. 
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. 
<emphasis id="F797FE4DFFE639211ED7F988FB15F9E7" box="[1170,1262,1629,1650]" italics="true" pageId="1" pageNumber="446">thoracica</emphasis>
Ingram &amp; Macfie
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<keyLead id="7E12816AFFE639211AD2F9AEFA67F905" box="[151,1436,1658,1680]" pageId="1" pageNumber="446">
- Smaller species (wing length 
<quantity id="021B8FBAFFE639211BBBF9AFFD76F905" box="[510,653,1658,1680]" metricMagnitude="-3" metricUnit="m" metricValue="1.37" metricValueMax="1.8" metricValueMin="0.94" pageId="1" pageNumber="446" unit="mm" value="1.37" valueMax="1.8" valueMin="0.94">0.94–1.80 mm</quantity>
); anal lobe of wing poorly developed.................................. 6
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<keyLead id="7E12816AFFE639211AD2F94DFA67F93B" box="[151,1436,1688,1710]" pageId="1" pageNumber="446">
6. Tarsal claws with bifid tips..................................................... 
<taxonomicName id="02E359DCFFE639211E1BF94DFA67F93B" ID-CoL="3XD2Y" authority="Grogan &amp; Wirth" authorityName="Grogan &amp; Wirth" box="[1118,1436,1688,1710]" class="Insecta" family="Ceratopogonidae" genus="Macrurohelea" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="446" phylum="Arthropoda" rank="species" species="paracaudata">
<emphasis id="F797FE4DFFE639211E1BF94DFB89F938" box="[1118,1138,1688,1709]" italics="true" pageId="1" pageNumber="446">M</emphasis>
. 
<emphasis id="F797FE4DFFE639211E3BF94DFB02F938" box="[1150,1273,1688,1709]" italics="true" pageId="1" pageNumber="446">paracaudata</emphasis>
Grogan &amp; Wirth
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<keyLead id="7E12816AFFE639211AD2F960FA67F95F" box="[151,1436,1717,1739]" pageId="1" pageNumber="446">- Tarsal claws with entire pointed tips...................................................................... 7</keyLead>
</paragraph>
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<paragraph id="C55C225FFFE639211AD2F906FA67F972" blockId="1.[151,1437,1336,2032]" box="[151,1436,1746,1774]" pageId="1" pageNumber="446">
<keyLead id="7E12816AFFE639211AD2F906FA67F972" box="[151,1436,1746,1774]" pageId="1" pageNumber="446">
7. Wing with intercalary vein in cell r 3 (
<figureCitation id="5DD83EDAFFE639211801F907FD71F97D" box="[580,650,1746,1768]" captionStart="FIGURES 26–32" captionStartId="14.[151,264,1943,1967]" captionTargetBox="[211,1381,185,1920]" captionTargetId="figure-23@14.[206,1382,181,1920]" captionTargetPageId="14" captionText="FIGURES 26–32. Macrurohelea sirii n. sp. Allotype female, 26–29. Holotype male, 30–32. 26, 30 head. 27 thorax. 28, 31 wing. 29 tip of abdomen. 32 genitalia. Scale bars = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5912791" httpUri="https://zenodo.org/record/5912791/files/figure.png" pageId="1" pageNumber="446">Fig. 28</figureCitation>
)............................................................... 8
</keyLead>
</paragraph>
<paragraph id="C55C225FFFE639211AD2F925FA67F890" blockId="1.[151,1437,1336,2032]" box="[151,1436,1776,1803]" pageId="1" pageNumber="446">
<keyLead id="7E12816AFFE639211AD2F925FA67F890" box="[151,1436,1776,1803]" pageId="1" pageNumber="446">
- Wing without intercalary vein in cell r 
<subScript id="5967201AFFE639211814F92AFDA3F89E" attach="left" box="[593,600,1791,1803]" fontSize="5" pageId="1" pageNumber="446">3</subScript>
(
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, 
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)......................................................... 10
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<keyStep id="7E173AFAFFE639211AD2F8D8FA67F8AA" pageId="1" pageNumber="446">
<paragraph id="C55C225FFFE639211AD2F8D8FA66F8B7" blockId="1.[151,1437,1336,2032]" box="[151,1437,1805,1827]" pageId="1" pageNumber="446">
<keyLead id="7E12816AFFE639211AD2F8D8FA66F8B7" box="[151,1437,1805,1827]" pageId="1" pageNumber="446">
8. Intercalary vein very faint, not forked (
<figureCitation id="5DD83EDAFFE63921181DF8D8FD65F8B6" box="[600,670,1805,1827]" captionStart="FIGURES 26–32" captionStartId="14.[151,264,1943,1967]" captionTargetBox="[211,1381,185,1920]" captionTargetId="figure-23@14.[206,1382,181,1920]" captionTargetPageId="14" captionText="FIGURES 26–32. Macrurohelea sirii n. sp. Allotype female, 26–29. Holotype male, 30–32. 26, 30 head. 27 thorax. 28, 31 wing. 29 tip of abdomen. 32 genitalia. Scale bars = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5912791" httpUri="https://zenodo.org/record/5912791/files/figure.png" pageId="1" pageNumber="446">Fig. 28</figureCitation>
)................................................... .. 
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<emphasis id="F797FE4DFFE639211F5FF8DBFAD5F8B6" box="[1306,1326,1806,1827]" italics="true" pageId="1" pageNumber="446">M</emphasis>
. 
<emphasis id="F797FE4DFFE639211F7FF8DBFA9AF8B6" box="[1338,1377,1806,1827]" italics="true" pageId="1" pageNumber="446">sirii</emphasis>
</taxonomicName>
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<taxonomicNameLabel id="ECA44336FFE639211F23F8D8FA66F8B7" box="[1382,1437,1805,1826]" pageId="1" pageNumber="446" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="C55C225FFFE639211AD2F8FEFA67F8AA" blockId="1.[151,1437,1336,2032]" box="[151,1436,1834,1856]" pageId="1" pageNumber="446">
<keyLead id="7E12816AFFE639211AD2F8FEFA67F8AA" box="[151,1436,1834,1856]" pageId="1" pageNumber="446">- Intercalary vein forked................................................................................. 9</keyLead>
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<paragraph id="C55C225FFFE639211AD2F89DFA67F8C8" blockId="1.[151,1437,1336,2032]" box="[151,1436,1864,1886]" pageId="1" pageNumber="446">
<keyLead id="7E12816AFFE639211AD2F89DFA67F8C8" box="[151,1436,1864,1886]" pageId="1" pageNumber="446">
9. Flagellomeres 9–13 greatly elongate, antennal ratio1.61................................. 
<taxonomicName id="02E359DCFFE639211EC9F89DFA67F8C8" ID-CoL="3XD34" authority="Spinelli &amp; Grogan" authorityName="Spinelli &amp; Grogan" authorityYear="1984" box="[1164,1436,1864,1886]" class="Insecta" family="Ceratopogonidae" genus="Macrurohelea" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="446" phylum="Arthropoda" rank="species" species="wirthi">
<emphasis id="F797FE4DFFE639211EC9F89DFB5BF8C8" box="[1164,1184,1864,1885]" italics="true" pageId="1" pageNumber="446">M</emphasis>
. 
<emphasis id="F797FE4DFFE639211EE9F89DFB1EF8C8" box="[1196,1253,1864,1885]" italics="true" pageId="1" pageNumber="446">wirthi</emphasis>
Spinelli &amp; Grogan
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</paragraph>
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<keyLead id="7E12816AFFE639211AD2F8B0FA67F8EE" box="[151,1436,1893,1915]" pageId="1" pageNumber="446">
- Flagellomeres 9–13 very short, antennal ratio 1.00..................................... 
<taxonomicName id="02E359DCFFE639211EC4F8B3FA67F8EE" ID-CoL="3XD2P" authority="Ingram &amp; Macfie" authorityName="Ingram &amp; Macfie" authorityYear="1931" box="[1153,1436,1893,1915]" class="Insecta" family="Ceratopogonidae" genus="Macrurohelea" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="446" phylum="Arthropoda" rank="species" species="caudata">
<emphasis id="F797FE4DFFE639211EC4F8B3FB6EF8EE" box="[1153,1173,1894,1915]" italics="true" pageId="1" pageNumber="446">M</emphasis>
. 
<emphasis id="F797FE4DFFE639211EE4F8B0FB15F8EF" box="[1185,1262,1893,1914]" italics="true" pageId="1" pageNumber="446">caudata</emphasis>
Ingram &amp; Macfie
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<paragraph id="C55C225FFFE639211AD2F857FA67F820" blockId="1.[151,1437,1336,2032]" pageId="1" pageNumber="446">
<keyLead id="7E12816AFFE639211AD2F857FA67F820" pageId="1" pageNumber="446">
10. Proximal 1/3 of wing membrane whitish hyaline, clearly contrasting with distal 2/3 darkly infuscated............................................................................................ 
<taxonomicName id="02E359DCFFE639211E1CF875FA67F820" ID-CoL="3XD2S" authority="Spinelli &amp; Grogan" authorityName="Spinelli &amp; Grogan" authorityYear="1990" box="[1113,1436,1952,1974]" class="Insecta" family="Ceratopogonidae" genus="Macrurohelea" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="446" phylum="Arthropoda" rank="species" species="fuscipennis">
<emphasis id="F797FE4DFFE639211E1CF875FB96F820" box="[1113,1133,1952,1973]" italics="true" pageId="1" pageNumber="446">M</emphasis>
. 
<emphasis id="F797FE4DFFE639211E3CF875FB1EF820" box="[1145,1253,1952,1973]" italics="true" pageId="1" pageNumber="446">fuscipennis</emphasis>
Spinelli &amp; Grogan
</taxonomicName>
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</paragraph>
<paragraph id="C55C225FFFE639211AD2F868FA67F87A" blockId="1.[151,1437,1336,2032]" pageId="1" pageNumber="446">
<keyLead id="7E12816AFFE639211AD2F868FA67F87A" pageId="1" pageNumber="446">
- Wing membrane hyaline or slightly infuscated, but uniformly pigmented (only slightly darkest on distal ½ of cell r 
<quantity id="021B8FBAFFE639211F1DF819FA81F846" box="[1368,1402,1981,2008]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="1" pageNumber="446" unit="in" value="3.0">
<subScript id="5967201AFFE639211F1DF819FAA4F84D" attach="left" box="[1368,1375,1996,2008]" fontSize="5" pageId="1" pageNumber="446">3</subScript>
in
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<taxonomicName id="02E359DCFFE639211FC7F86BFED0F87A" ID-CoL="9HVZP" authority="Spinelli &amp; Ronderos &amp; Grogan, 2022" authorityName="Spinelli &amp; Ronderos &amp; Grogan" authorityYear="2022" class="Insecta" family="Ceratopogonidae" genus="Macrurohelea" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="446" phylum="Arthropoda" rank="species" species="donatoi" status="sp. nov.">
<emphasis id="F797FE4DFFE639211FC7F86BFA6DF846" box="[1410,1430,1982,2003]" italics="true" pageId="1" pageNumber="446">M</emphasis>
. 
<emphasis id="F797FE4DFFE639211AA7F80FFED0F87A" box="[226,299,2010,2031]" italics="true" pageId="1" pageNumber="446">donatoi</emphasis>
</taxonomicName>
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<taxonomicNameLabel id="ECA44336FFE639211B74F80FFE93F87A" box="[305,360,2010,2031]" pageId="1" pageNumber="446" rank="species">n. sp.</taxonomicNameLabel>
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)....................................................................................... 11
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<paragraph id="C55C225FFFE539221AD2FF4DFA67FF38" blockId="2.[151,1437,151,1757]" box="[151,1436,151,173]" pageId="2" pageNumber="447">
<keyLead id="7E12816AFFE539221AD2FF4DFA67FF38" box="[151,1436,151,173]" pageId="2" pageNumber="447">
11. Small species (wing length 
<quantity id="021B8FBAFFE539221BAEFF4DFDBFFF38" box="[491,580,152,173]" metricMagnitude="-4" metricUnit="m" metricValue="9.399999999999999" pageId="2" pageNumber="447" unit="mm" value="0.94">0.94 mm</quantity>
); flagellomeres 9–13 very short (antennal ratio 0.59)............... 
<taxonomicName id="02E359DCFFE539221EB4FF4DFA67FF38" ID-CoL="3XD2V" authority="Wirth" authorityName="Wirth" authorityYear="1965" box="[1265,1436,151,173]" class="Insecta" family="Ceratopogonidae" genus="Macrurohelea" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="447" phylum="Arthropoda" rank="species" species="kuscheli">
<emphasis id="F797FE4DFFE539221EB4FF4DFAFEFF38" box="[1265,1285,152,173]" italics="true" pageId="2" pageNumber="447">M</emphasis>
. 
<emphasis id="F797FE4DFFE539221F54FF42FA9BFF39" box="[1297,1376,151,172]" italics="true" pageId="2" pageNumber="447">kuscheli</emphasis>
Wirth
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<paragraph id="C55C225FFFE539221AD2FF60FA67FF5F" blockId="2.[151,1437,151,1757]" box="[151,1436,181,203]" pageId="2" pageNumber="447">
<keyLead id="7E12816AFFE539221AD2FF60FA67FF5F" box="[151,1436,181,203]" pageId="2" pageNumber="447">
- Larger species (wing length 
<quantity id="021B8FBAFFE539221BB6FF60FD79FF5F" box="[499,642,181,202]" metricMagnitude="-3" metricUnit="m" metricValue="1.535" metricValueMax="1.8" metricValueMin="1.27" pageId="2" pageNumber="447" unit="mm" value="1.535" valueMax="1.8" valueMin="1.27">1.27–1.80 mm</quantity>
); flagellomeres 9–13 elongate (antennal ratio1.10–1.36)................... 12
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<paragraph id="C55C225FFFE539221AD2FF07FA66FF72" blockId="2.[151,1437,151,1757]" box="[151,1437,210,232]" pageId="2" pageNumber="447">
<keyLead id="7E12816AFFE539221AD2FF07FA66FF72" box="[151,1437,210,232]" pageId="2" pageNumber="447">
12. Thorax, excluding legs, dark brown, legs and abdomen yellowish (
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).......................... 
<taxonomicName id="02E359DCFFE539221F46FF06FA9BFF72" ID-CoL="9HVMC" authority="Spinelli &amp; Ronderos &amp; Grogan, 2022" authorityName="Spinelli &amp; Ronderos &amp; Grogan" authorityYear="2022" box="[1283,1376,210,232]" class="Insecta" family="Ceratopogonidae" genus="Macrurohelea" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="447" phylum="Arthropoda" rank="species" species="bassoi" status="sp. nov.">
<emphasis id="F797FE4DFFE539221F46FF06FAECFF7D" box="[1283,1303,211,232]" italics="true" pageId="2" pageNumber="447">M</emphasis>
. 
<emphasis id="F797FE4DFFE539221F66FF07FA9BFF72" box="[1315,1376,210,231]" italics="true" pageId="2" pageNumber="447">bassoi</emphasis>
</taxonomicName>
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<taxonomicNameLabel id="ECA44336FFE539221F23FF07FA66FF72" box="[1382,1437,210,231]" pageId="2" pageNumber="447" rank="species">n. sp.</taxonomicNameLabel>
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</paragraph>
<paragraph id="C55C225FFFE539221AD2FF25FA67FE90" blockId="2.[151,1437,151,1757]" box="[151,1436,239,261]" pageId="2" pageNumber="447">
<keyLead id="7E12816AFFE539221AD2FF25FA67FE90" box="[151,1436,239,261]" pageId="2" pageNumber="447">- Thorax and abdomen with same coloration................................................................ 13</keyLead>
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<keyStep id="7E173AFAFFE539221AD2FED8FA67FEAA" pageId="2" pageNumber="447">
<paragraph id="C55C225FFFE539221AD2FED8FA67FEB6" blockId="2.[151,1437,151,1757]" box="[151,1436,269,291]" pageId="2" pageNumber="447">
<keyLead id="7E12816AFFE539221AD2FED8FA67FEB6" box="[151,1436,269,291]" pageId="2" pageNumber="447">
13. Halter pale; spermathecae spheroidal with short necks.................................... 
<taxonomicName id="02E359DCFFE539221EE5FEDBFA67FEB6" ID-CoL="6R3WW" authority="Grogan &amp; Wirth" authorityName="Grogan &amp; Wirth" box="[1184,1436,269,291]" class="Insecta" family="Ceratopogonidae" genus="Macrurohelea" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="447" phylum="Arthropoda" rank="species" species="irwini">
<emphasis id="F797FE4DFFE539221EE5FEDBFB4FFEB6" box="[1184,1204,270,291]" italics="true" pageId="2" pageNumber="447">M</emphasis>
. 
<emphasis id="F797FE4DFFE539221E85FED8FB02FEB7" box="[1216,1273,269,290]" italics="true" pageId="2" pageNumber="447">irwini</emphasis>
Grogan &amp; Wirth
</taxonomicName>
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<paragraph id="C55C225FFFE539221AD2FEFEFA67FEAA" blockId="2.[151,1437,151,1757]" box="[151,1436,298,320]" pageId="2" pageNumber="447">
<keyLead id="7E12816AFFE539221AD2FEFEFA67FEAA" box="[151,1436,298,320]" pageId="2" pageNumber="447">- Halter light or dark brown; spermathecae ovoid with long, slender necks........................................ 14</keyLead>
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<keyStep id="7E173AFAFFE539221AD2FE9DFA67FE20" pageId="2" pageNumber="447">
<paragraph id="C55C225FFFE539221AD2FE9DFA66FEEF" blockId="2.[151,1437,151,1757]" pageId="2" pageNumber="447">
<keyLead id="7E12816AFFE539221AD2FE9DFA66FEEF" pageId="2" pageNumber="447">
14. Wing membrane infuscated brown, slightly darkest on distal ½ of cell r 
<subScript id="5967201AFFE5392219C2FE82FC75FEF6" attach="left" box="[903,910,343,355]" fontSize="5" pageId="2" pageNumber="447">3</subScript>
(
<figureCitation id="5DD83EDAFFE5392219DFFE92FC19FECB" box="[922,994,327,350]" captionStart="FIGURES 8–14" captionStartId="6.[151,264,1943,1967]" captionTargetBox="[186,1400,183,1920]" captionTargetId="figure-23@6.[186,1401,181,1920]" captionTargetPageId="6" captionText="FIGURES 8–14. Macrurohelea donatoi n. sp. Allotype female, 8–11. Holotype male, 12–14. 8, 12 head. 9 thorax. 10, 13 wing. 11 tip of abdomen. 14 genitalia. Scale bars = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5912785" httpUri="https://zenodo.org/record/5912785/files/figure.png" pageId="2" pageNumber="447">Fig. 10</figureCitation>
); spermathecae large (longest 80–82 µm long) with necks ca. 20 µm long................................................................. 
<taxonomicName id="02E359DCFFE539221EB2FEB3FA9BFEEF" ID-CoL="9HVZP" authority="Spinelli &amp; Ronderos &amp; Grogan, 2022" authorityName="Spinelli &amp; Ronderos &amp; Grogan" authorityYear="2022" box="[1271,1376,357,379]" class="Insecta" family="Ceratopogonidae" genus="Macrurohelea" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="447" phylum="Arthropoda" rank="species" species="donatoi" status="sp. nov.">
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. 
<emphasis id="F797FE4DFFE539221F52FEB0FA9BFEEF" box="[1303,1376,357,378]" italics="true" pageId="2" pageNumber="447">donatoi</emphasis>
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<taxonomicNameLabel id="ECA44336FFE539221F23FEB0FA66FEEF" box="[1382,1437,357,378]" pageId="2" pageNumber="447" rank="species">n. sp.</taxonomicNameLabel>
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<paragraph id="C55C225FFFE539221AD2FE56FA67FE20" blockId="2.[151,1437,151,1757]" pageId="2" pageNumber="447">
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- Wing membrane uniformly lightly infuscated; spermathecae smaller (longest 53 µm long) necks ca. 15 µm long....................................................................................... 
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. 
<emphasis id="F797FE4DFFE539221EDCFE75FB1EFE20" box="[1177,1253,416,437]" italics="true" pageId="2" pageNumber="447">yamana</emphasis>
Spinelli &amp; Grogan
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<keyStep id="7E173AFAFFE539221AD2FE68FA67FE7A" pageId="2" pageNumber="447">
<paragraph id="C55C225FFFE539221AD2FE68FA67FE46" blockId="2.[151,1437,151,1757]" box="[151,1436,445,467]" pageId="2" pageNumber="447">
<keyLead id="7E12816AFFE539221AD2FE68FA67FE46" box="[151,1436,445,467]" pageId="2" pageNumber="447">
15. Tip of parameres with hooked, slender process (
<figureCitation id="5DD83EDAFFE5392218EAFE68FD04FE47" box="[687,767,445,467]" captionStart="FIGURES 37–42" captionStartId="16.[151,264,1913,1937]" captionTargetBox="[214,1370,182,1882]" captionTargetId="figure-23@16.[214,1373,181,1889]" captionTargetPageId="16" captionText="FIGURES 37–42. Macrurohelea bassoi n. sp. Holotype male, 37–38. Macrurohelea donatoi n. sp. Holotype male, 39–40. Macrurohelea kuscheli Wirth, male, 41–42. 37, 39, 41 parameres. 38, 40, 42 aedeagus." figureDoi="http://doi.org/10.5281/zenodo.5912797" httpUri="https://zenodo.org/record/5912797/files/figure.png" pageId="2" pageNumber="447">Figs. 41</figureCitation>
, 
<figureCitation id="5DD83EDAFFE53922194CFE68FCD9FE46" box="[777,802,445,467]" captionStart="FIGURES 47–50" captionStartId="18.[151,264,1227,1251]" captionTargetBox="[250,1333,183,1189]" captionTargetId="figure-23@18.[242,1346,181,1203]" captionTargetPageId="18" captionText="FIGURES 47–50. Macrurohelea sirii n. sp. Holotype male, 47–48. Macrurohelea ventanensis n. sp. Holotype male, 49–50. 47, 49 parameres. 48, 50 aedeagus." figureDoi="http://doi.org/10.5281/zenodo.5912801" httpUri="https://zenodo.org/record/5912801/files/figure.png" pageId="2" pageNumber="447">49</figureCitation>
).................................................. 16
</keyLead>
</paragraph>
<paragraph id="C55C225FFFE539221AD2FE0EFA67FE7A" blockId="2.[151,1437,151,1757]" box="[151,1436,474,496]" pageId="2" pageNumber="447">
<keyLead id="7E12816AFFE539221AD2FE0EFA67FE7A" box="[151,1436,474,496]" pageId="2" pageNumber="447">
- Tip of parameres without hooked process (
<figureCitation id="5DD83EDAFFE539221839FE0FFD0BFE7A" box="[636,752,474,496]" captionStart="FIGURES 37–42" captionStartId="16.[151,264,1913,1937]" captionTargetBox="[214,1370,182,1882]" captionTargetId="figure-23@16.[214,1373,181,1889]" captionTargetPageId="16" captionText="FIGURES 37–42. Macrurohelea bassoi n. sp. Holotype male, 37–38. Macrurohelea donatoi n. sp. Holotype male, 39–40. Macrurohelea kuscheli Wirth, male, 41–42. 37, 39, 41 parameres. 38, 40, 42 aedeagus." figureDoi="http://doi.org/10.5281/zenodo.5912797" httpUri="https://zenodo.org/record/5912797/files/figure.png" pageId="2" pageNumber="447">Figs. 37, 39</figureCitation>
, 
<figureCitation id="5DD83EDAFFE5392218BFFE0FFCC3FE65" box="[762,824,474,496]" captionStart="FIGURES 43–46" captionStartId="17.[151,264,1442,1466]" captionTargetBox="[166,1437,186,1402]" captionTargetId="figure-19@17.[151,1437,181,1418]" captionTargetPageId="17" captionText="FIGURES 43–46. Macrurohelea monotheca Spinelli &amp; Grogan, male, 43–44. Macrurohelea morenoi n. sp. Holotype male, 45–46. 43, 45 parameres. 44, 46 aedeagus." figureDoi="http://doi.org/10.5281/zenodo.5912799" httpUri="https://zenodo.org/record/5912799/files/figure.png" pageId="2" pageNumber="447">43, 45</figureCitation>
, 
<figureCitation id="5DD83EDAFFE539221907FE0FFCA0FE65" box="[834,859,474,496]" captionStart="FIGURES 47–50" captionStartId="18.[151,264,1227,1251]" captionTargetBox="[250,1333,183,1189]" captionTargetId="figure-23@18.[242,1346,181,1203]" captionTargetPageId="18" captionText="FIGURES 47–50. Macrurohelea sirii n. sp. Holotype male, 47–48. Macrurohelea ventanensis n. sp. Holotype male, 49–50. 47, 49 parameres. 48, 50 aedeagus." figureDoi="http://doi.org/10.5281/zenodo.5912801" httpUri="https://zenodo.org/record/5912801/files/figure.png" pageId="2" pageNumber="447">47</figureCitation>
)..............................................17
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<keyStep id="7E173AFAFFE539221AD2FE22FA66FDF0" pageId="2" pageNumber="447">
<paragraph id="C55C225FFFE539221AD2FE22FA67FDBE" blockId="2.[151,1437,151,1757]" pageId="2" pageNumber="447">
<keyLead id="7E12816AFFE539221AD2FE22FA67FDBE" pageId="2" pageNumber="447">
16. Antennal flagellomeres 4–9 fused (
<figureCitation id="5DD83EDAFFE539221870FE22FD87FD98" box="[565,636,503,525]" captionStart="FIGURES 15–21" captionStartId="8.[151,264,1944,1968]" captionTargetBox="[193,1399,188,1907]" captionTargetId="figure-23@8.[189,1399,181,1920]" captionTargetPageId="8" captionText="FIGURES 15–21. Macrurohelea kuscheli Wirth, male, 15–18. Macrurohelea monotheca Spinelli &amp; Grogan, male, 19–21. 15, 19 head. 16 thorax. 17, 20 wing. 18-21 genitalia. Scale bars = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5912787" httpUri="https://zenodo.org/record/5912787/files/figure.png" pageId="2" pageNumber="447">Fig. 15</figureCitation>
); second radial cell 0.95–1.76x longer than first; distal portion of aedeagus heavily sclerotized, slender, with bifid tip (
<figureCitation id="5DD83EDAFFE539221874FDC0FD8CFDBE" box="[561,631,533,555]" captionStart="FIGURES 37–42" captionStartId="16.[151,264,1913,1937]" captionTargetBox="[214,1370,182,1882]" captionTargetId="figure-23@16.[214,1373,181,1889]" captionTargetPageId="16" captionText="FIGURES 37–42. Macrurohelea bassoi n. sp. Holotype male, 37–38. Macrurohelea donatoi n. sp. Holotype male, 39–40. Macrurohelea kuscheli Wirth, male, 41–42. 37, 39, 41 parameres. 38, 40, 42 aedeagus." figureDoi="http://doi.org/10.5281/zenodo.5912797" httpUri="https://zenodo.org/record/5912797/files/figure.png" pageId="2" pageNumber="447">Fig. 42</figureCitation>
).................................................... 
<taxonomicName id="02E359DCFFE539221EB4FDC3FA67FDBE" ID-CoL="3XD2V" authority="Wirth" authorityName="Wirth" authorityYear="1965" box="[1265,1436,533,555]" class="Insecta" family="Ceratopogonidae" genus="Macrurohelea" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="447" phylum="Arthropoda" rank="species" species="kuscheli">
<emphasis id="F797FE4DFFE539221EB4FDC3FAFEFDBE" box="[1265,1285,534,555]" italics="true" pageId="2" pageNumber="447">M</emphasis>
. 
<emphasis id="F797FE4DFFE539221F54FDC0FA9BFDBF" box="[1297,1376,533,554]" italics="true" pageId="2" pageNumber="447">kuscheli</emphasis>
Wirth
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</paragraph>
<paragraph id="C55C225FFFE539221AD2FDE6FA66FDF0" blockId="2.[151,1437,151,1757]" pageId="2" pageNumber="447">
<keyLead id="7E12816AFFE539221AD2FDE6FA66FDF0" pageId="2" pageNumber="447">
- Antennal flagellomeres 4–9 separated (
<figureCitation id="5DD83EDAFFE53922181AFDE7FD53FDDD" box="[607,680,562,584]" captionStart="FIGURES 33–36" captionStartId="15.[151,264,1943,1967]" captionTargetBox="[240,1349,181,1917]" captionTargetId="figure-19@15.[237,1350,181,1920]" captionTargetPageId="15" captionText="FIGURES 33–36. Macrurohelea ventanensis n. sp. Holotype male. 33 head. 34 thorax. 35 wing. 36 genitalia. Scale bars = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5912793" httpUri="https://zenodo.org/record/5912793/files/figure.png" pageId="2" pageNumber="447">Fig. 33</figureCitation>
); second radial cell twice as long as first (
<figureCitation id="5DD83EDAFFE539221E00FDE7FB75FDDD" box="[1093,1166,562,584]" captionStart="FIGURES 33–36" captionStartId="15.[151,264,1943,1967]" captionTargetBox="[240,1349,181,1917]" captionTargetId="figure-19@15.[237,1350,181,1920]" captionTargetPageId="15" captionText="FIGURES 33–36. Macrurohelea ventanensis n. sp. Holotype male. 33 head. 34 thorax. 35 wing. 36 genitalia. Scale bars = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5912793" httpUri="https://zenodo.org/record/5912793/files/figure.png" pageId="2" pageNumber="447">Fig. 35</figureCitation>
); distal portion of aedeagus lightly sclerotized, ending in a quadrate hyaline process (
<figureCitation id="5DD83EDAFFE539221943FD9AFCADFDF0" box="[774,854,591,613]" captionStart="FIGURES 33–36" captionStartId="15.[151,264,1943,1967]" captionTargetBox="[240,1349,181,1917]" captionTargetId="figure-19@15.[237,1350,181,1920]" captionTargetPageId="15" captionText="FIGURES 33–36. Macrurohelea ventanensis n. sp. Holotype male. 33 head. 34 thorax. 35 wing. 36 genitalia. Scale bars = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5912793" httpUri="https://zenodo.org/record/5912793/files/figure.png" pageId="2" pageNumber="447">Figs. 36</figureCitation>
, 
<figureCitation id="5DD83EDAFFE539221925FD9AFC82FDF0" box="[864,889,591,613]" captionStart="FIGURES 47–50" captionStartId="18.[151,264,1227,1251]" captionTargetBox="[250,1333,183,1189]" captionTargetId="figure-23@18.[242,1346,181,1203]" captionTargetPageId="18" captionText="FIGURES 47–50. Macrurohelea sirii n. sp. Holotype male, 47–48. Macrurohelea ventanensis n. sp. Holotype male, 49–50. 47, 49 parameres. 48, 50 aedeagus." figureDoi="http://doi.org/10.5281/zenodo.5912801" httpUri="https://zenodo.org/record/5912801/files/figure.png" pageId="2" pageNumber="447">50</figureCitation>
)........................... 
<taxonomicName id="02E359DCFFE539221E95FD85FA9BFDF0" ID-CoL="9HXXG" authority="Spinelli &amp; Ronderos &amp; Grogan, 2022" authorityName="Spinelli &amp; Ronderos &amp; Grogan" authorityYear="2022" box="[1232,1376,592,613]" class="Insecta" family="Ceratopogonidae" genus="Macrurohelea" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="447" phylum="Arthropoda" rank="species" species="ventanensis" status="sp. nov.">
<emphasis id="F797FE4DFFE539221E95FD85FB1FFDF0" box="[1232,1252,592,613]" italics="true" pageId="2" pageNumber="447">M</emphasis>
. 
<emphasis id="F797FE4DFFE539221EB5FD85FA9BFDF0" box="[1264,1376,592,613]" italics="true" pageId="2" pageNumber="447">ventanensis</emphasis>
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<taxonomicNameLabel id="ECA44336FFE539221F23FD85FA66FDF0" box="[1382,1437,592,613]" pageId="2" pageNumber="447" rank="species">n. sp.</taxonomicNameLabel>
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<keyStep id="7E173AFAFFE539221AD2FDB8FA67FD0A" pageId="2" pageNumber="447">
<paragraph id="C55C225FFFE539221AD2FDB8FA67FD16" blockId="2.[151,1437,151,1757]" box="[151,1436,621,643]" pageId="2" pageNumber="447">
<keyLead id="7E12816AFFE539221AD2FDB8FA67FD16" box="[151,1436,621,643]" pageId="2" pageNumber="447">
17. Legs with numerous long bristly setae.......................................................... 
<taxonomicName id="02E359DCFFE539221F41FDBBFA67FD16" ID-CoL="3XD2Z" authority="Wirth" authorityName="Wirth" authorityYear="1965" box="[1284,1436,621,643]" class="Insecta" family="Ceratopogonidae" genus="Macrurohelea" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="447" phylum="Arthropoda" rank="species" species="setosa">
<emphasis id="F797FE4DFFE539221F41FDBBFAE3FD16" box="[1284,1304,622,643]" italics="true" pageId="2" pageNumber="447">M</emphasis>
. 
<emphasis id="F797FE4DFFE539221F61FDBBFA9BFD16" box="[1316,1376,622,643]" italics="true" pageId="2" pageNumber="447">setosa</emphasis>
Wirth
</taxonomicName>
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</paragraph>
<paragraph id="C55C225FFFE539221AD2FD5EFA67FD0A" blockId="2.[151,1437,151,1757]" box="[151,1436,650,672]" pageId="2" pageNumber="447">
<keyLead id="7E12816AFFE539221AD2FD5EFA67FD0A" box="[151,1436,650,672]" pageId="2" pageNumber="447">- Legs with vestiture of short setae........................................................................ 18</keyLead>
</paragraph>
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<paragraph id="C55C225FFFE539221AD2FD7DFA67FD28" blockId="2.[151,1437,151,1757]" box="[151,1436,679,701]" pageId="2" pageNumber="447">
<keyLead id="7E12816AFFE539221AD2FD7DFA67FD28" box="[151,1436,679,701]" pageId="2" pageNumber="447">
18. Gonostylus bent abruptly subapically at 90°.......................................... 
<taxonomicName id="02E359DCFFE539221EC4FD7DFA67FD28" ID-CoL="3XD2T" authority="Spinelli &amp; Grogan" authorityName="Spinelli &amp; Grogan" authorityYear="1984" box="[1153,1436,679,701]" class="Insecta" family="Ceratopogonidae" genus="Macrurohelea" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="447" phylum="Arthropoda" rank="species" species="gentilii">
<emphasis id="F797FE4DFFE539221EC4FD7DFB6EFD28" box="[1153,1173,680,701]" italics="true" pageId="2" pageNumber="447">M</emphasis>
. 
<emphasis id="F797FE4DFFE539221EE4FD72FB1EFD29" box="[1185,1253,679,700]" italics="true" pageId="2" pageNumber="447">gentilii</emphasis>
Spinelli &amp; Grogan
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</paragraph>
<paragraph id="C55C225FFFE539221AD2FD10FA67FD4F" blockId="2.[151,1437,151,1757]" box="[151,1436,709,731]" pageId="2" pageNumber="447">
<keyLead id="7E12816AFFE539221AD2FD10FA67FD4F" box="[151,1436,709,731]" pageId="2" pageNumber="447">- Gonostylus curved or straight, but not bent abruptly subapically at 90°.......................................... 19</keyLead>
</paragraph>
</keyStep>
<keyStep id="7E173AFAFFE539221AD2FD37FA67FC80" pageId="2" pageNumber="447">
<paragraph id="C55C225FFFE539221AD2FD37FA67FD62" blockId="2.[151,1437,151,1757]" box="[151,1436,738,760]" pageId="2" pageNumber="447">
<keyLead id="7E12816AFFE539221AD2FD37FA67FD62" box="[151,1436,738,760]" pageId="2" pageNumber="447">
19. Aedeagus Y-shaped or V-shaped, basal arch stout or slender, distal portion short or elongate (
<figureCitation id="5DD83EDAFFE539221E1BFD37FB55FD6D" box="[1118,1198,738,760]" captionStart="FIGURES 43–46" captionStartId="17.[151,264,1442,1466]" captionTargetBox="[166,1437,186,1402]" captionTargetId="figure-19@17.[151,1437,181,1418]" captionTargetPageId="17" captionText="FIGURES 43–46. Macrurohelea monotheca Spinelli &amp; Grogan, male, 43–44. Macrurohelea morenoi n. sp. Holotype male, 45–46. 43, 45 parameres. 44, 46 aedeagus." figureDoi="http://doi.org/10.5281/zenodo.5912799" httpUri="https://zenodo.org/record/5912799/files/figure.png" pageId="2" pageNumber="447">Figs. 46</figureCitation>
, 
<figureCitation id="5DD83EDAFFE539221EFDFD37FB2AFD6D" box="[1208,1233,738,760]" captionStart="FIGURES 47–50" captionStartId="18.[151,264,1227,1251]" captionTargetBox="[250,1333,183,1189]" captionTargetId="figure-23@18.[242,1346,181,1203]" captionTargetPageId="18" captionText="FIGURES 47–50. Macrurohelea sirii n. sp. Holotype male, 47–48. Macrurohelea ventanensis n. sp. Holotype male, 49–50. 47, 49 parameres. 48, 50 aedeagus." figureDoi="http://doi.org/10.5281/zenodo.5912801" httpUri="https://zenodo.org/record/5912801/files/figure.png" pageId="2" pageNumber="447">48</figureCitation>
).............. 20
</keyLead>
</paragraph>
<paragraph id="C55C225FFFE539221AD2FCD5FA67FC80" blockId="2.[151,1437,151,1757]" box="[151,1436,767,789]" pageId="2" pageNumber="447">
<keyLead id="7E12816AFFE539221AD2FCD5FA67FC80" box="[151,1436,767,789]" pageId="2" pageNumber="447">
- Aedeagus triangular or tongue-shaped (
<figureCitation id="5DD83EDAFFE539221818FD2AFD2AFC80" box="[605,721,767,789]" captionStart="FIGURES 37–42" captionStartId="16.[151,264,1913,1937]" captionTargetBox="[214,1370,182,1882]" captionTargetId="figure-23@16.[214,1373,181,1889]" captionTargetPageId="16" captionText="FIGURES 37–42. Macrurohelea bassoi n. sp. Holotype male, 37–38. Macrurohelea donatoi n. sp. Holotype male, 39–40. Macrurohelea kuscheli Wirth, male, 41–42. 37, 39, 41 parameres. 38, 40, 42 aedeagus." figureDoi="http://doi.org/10.5281/zenodo.5912797" httpUri="https://zenodo.org/record/5912797/files/figure.png" pageId="2" pageNumber="447">Figs. 38, 40</figureCitation>
, 
<figureCitation id="5DD83EDAFFE53922189EFCD5FD0EFC80" box="[731,757,768,789]" captionStart="FIGURES 43–46" captionStartId="17.[151,264,1442,1466]" captionTargetBox="[166,1437,186,1402]" captionTargetId="figure-19@17.[151,1437,181,1418]" captionTargetPageId="17" captionText="FIGURES 43–46. Macrurohelea monotheca Spinelli &amp; Grogan, male, 43–44. Macrurohelea morenoi n. sp. Holotype male, 45–46. 43, 45 parameres. 44, 46 aedeagus." figureDoi="http://doi.org/10.5281/zenodo.5912799" httpUri="https://zenodo.org/record/5912799/files/figure.png" pageId="2" pageNumber="447">44</figureCitation>
, 
<figureCitation id="5DD83EDAFFE5392218BAFD2AFCE3FC80" box="[767,792,767,789]" captionStart="FIGURES 47–50" captionStartId="18.[151,264,1227,1251]" captionTargetBox="[250,1333,183,1189]" captionTargetId="figure-23@18.[242,1346,181,1203]" captionTargetPageId="18" captionText="FIGURES 47–50. Macrurohelea sirii n. sp. Holotype male, 47–48. Macrurohelea ventanensis n. sp. Holotype male, 49–50. 47, 49 parameres. 48, 50 aedeagus." figureDoi="http://doi.org/10.5281/zenodo.5912801" httpUri="https://zenodo.org/record/5912801/files/figure.png" pageId="2" pageNumber="447">50</figureCitation>
)................................................... 23
</keyLead>
</paragraph>
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<keyStep id="7E173AFAFFE539221AD2FCC8FA67FC1F" pageId="2" pageNumber="447">
<paragraph id="C55C225FFFE539221AD2FCC8FA67FCC5" blockId="2.[151,1437,151,1757]" pageId="2" pageNumber="447">
<keyLead id="7E12816AFFE539221AD2FCC8FA67FCC5" pageId="2" pageNumber="447">
20. Large species (wing length 
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); aedeagus Y-shaped, with slender basal arch and distal portion elongate, slender; scutum, scutellum and post-scutellum golden brown; anal lobe of wing well developed, broad....... 
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. 
<emphasis id="F797FE4DFFE539221ED7FCEFFB15FCDA" box="[1170,1262,826,847]" italics="true" pageId="2" pageNumber="447">thoracica</emphasis>
Ingram &amp; Macfie
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<keyLead id="7E12816AFFE539221AD2FC8DFA67FC1F" pageId="2" pageNumber="447">
- Smaller species (wing length 
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); aedeagus with stout basal arch and distal portion short; scutum, scutellum and post-scutellum dark brown; anal lobe of wing poorly developed............................................... 21
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<keyLead id="7E12816AFFE539221AD2FC47FA67FC3D" box="[151,1436,914,936]" pageId="2" pageNumber="447">
21. Sternite 9 with posteromedian excavation......................................... 
<taxonomicName id="02E359DCFFE539221E1BFC46FA67FC3D" ID-CoL="3XD2Y" authority="Grogan &amp; Wirth" authorityName="Grogan &amp; Wirth" box="[1118,1436,914,936]" class="Insecta" family="Ceratopogonidae" genus="Macrurohelea" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="447" phylum="Arthropoda" rank="species" species="paracaudata">
<emphasis id="F797FE4DFFE539221E1BFC46FB89FC3D" box="[1118,1138,915,936]" italics="true" pageId="2" pageNumber="447">M</emphasis>
. 
<emphasis id="F797FE4DFFE539221E3BFC47FB02FC32" box="[1150,1273,914,935]" italics="true" pageId="2" pageNumber="447">paracaudata</emphasis>
Grogan &amp; Wirth
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<paragraph id="C55C225FFFE539221AD2FC65FA67FC50" blockId="2.[151,1437,151,1757]" box="[151,1436,943,966]" pageId="2" pageNumber="447">
<keyLead id="7E12816AFFE539221AD2FC65FA67FC50" box="[151,1436,943,966]" pageId="2" pageNumber="447">
- Sternite 9 without posteromedian excavation (
<figureCitation id="5DD83EDAFFE5392218D9FC7AFD17FC50" box="[668,748,943,965]" captionStart="FIGURES 22–25" captionStartId="11.[151,264,1971,1995]" captionTargetBox="[230,1356,229,1875]" captionTargetId="figure-19@11.[169,1417,181,1947]" captionTargetPageId="11" captionText="FIGURES 22–25. Macrurohelea morenoi n. sp. Holotype male. 22, head. 23 thorax. 24 wing. 25 genitalia. Scale bars = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5912789" httpUri="https://zenodo.org/record/5912789/files/figure.png" pageId="2" pageNumber="447">Figs. 25</figureCitation>
, 
<figureCitation id="5DD83EDAFFE5392218B3FC65FCF4FC53" box="[758,783,944,966]" captionStart="FIGURES 26–32" captionStartId="14.[151,264,1943,1967]" captionTargetBox="[211,1381,185,1920]" captionTargetId="figure-23@14.[206,1382,181,1920]" captionTargetPageId="14" captionText="FIGURES 26–32. Macrurohelea sirii n. sp. Allotype female, 26–29. Holotype male, 30–32. 26, 30 head. 27 thorax. 28, 31 wing. 29 tip of abdomen. 32 genitalia. Scale bars = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5912791" httpUri="https://zenodo.org/record/5912791/files/figure.png" pageId="2" pageNumber="447">32</figureCitation>
)................................................... 22
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<keyStep id="7E173AFAFFE539221AD2FC18FA66FBC2" pageId="2" pageNumber="447">
<paragraph id="C55C225FFFE539221AD2FC18FA66FB88" blockId="2.[151,1437,151,1757]" pageId="2" pageNumber="447">
<keyLead id="7E12816AFFE539221AD2FC18FA66FB88" pageId="2" pageNumber="447">
22. Aedeagus V-shaped, very broad basally, broader than bases of parameres, distal portion triangular, divided medially, apices slightly divergent (
<figureCitation id="5DD83EDAFFE539221BDEFC3FFE1EFB95" box="[411,485,1002,1024]" captionStart="FIGURES 43–46" captionStartId="17.[151,264,1442,1466]" captionTargetBox="[166,1437,186,1402]" captionTargetId="figure-19@17.[151,1437,181,1418]" captionTargetPageId="17" captionText="FIGURES 43–46. Macrurohelea monotheca Spinelli &amp; Grogan, male, 43–44. Macrurohelea morenoi n. sp. Holotype male, 45–46. 43, 45 parameres. 44, 46 aedeagus." figureDoi="http://doi.org/10.5281/zenodo.5912799" httpUri="https://zenodo.org/record/5912799/files/figure.png" pageId="2" pageNumber="447">Fig. 46</figureCitation>
); distal portion of parameres stout, nearly straight, slightly divergent distally, apex rounded (
<figureCitation id="5DD83EDAFFE539221F30FC3FFF00FB88" captionStart="FIGURES 43–46" captionStartId="17.[151,264,1442,1466]" captionTargetBox="[166,1437,186,1402]" captionTargetId="figure-19@17.[151,1437,181,1418]" captionTargetPageId="17" captionText="FIGURES 43–46. Macrurohelea monotheca Spinelli &amp; Grogan, male, 43–44. Macrurohelea morenoi n. sp. Holotype male, 45–46. 43, 45 parameres. 44, 46 aedeagus." figureDoi="http://doi.org/10.5281/zenodo.5912799" httpUri="https://zenodo.org/record/5912799/files/figure.png" pageId="2" pageNumber="447">Fig. 45</figureCitation>
)................................................................................... 
<taxonomicName id="02E359DCFFE539221EB4FBDDFA9BFB88" ID-CoL="9HYPY" authority="Spinelli &amp; Ronderos &amp; Grogan, 2022" authorityName="Spinelli &amp; Ronderos &amp; Grogan" authorityYear="2022" box="[1265,1376,1032,1053]" class="Insecta" family="Ceratopogonidae" genus="Macrurohelea" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="447" phylum="Arthropoda" rank="species" species="morenoi" status="sp. nov.">
<emphasis id="F797FE4DFFE539221EB4FBDDFAFEFB88" box="[1265,1285,1032,1053]" italics="true" pageId="2" pageNumber="447">M</emphasis>
. 
<emphasis id="F797FE4DFFE539221F54FBDDFA9BFB88" box="[1297,1376,1032,1053]" italics="true" pageId="2" pageNumber="447">morenoi</emphasis>
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<emphasis id="F797FE4DFFE539221F23FBDDFA66FB88" bold="true" box="[1382,1437,1032,1053]" pageId="2" pageNumber="447">
<taxonomicNameLabel id="ECA44336FFE539221F23FBDDFA66FB88" box="[1382,1437,1032,1053]" pageId="2" pageNumber="447" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
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</paragraph>
<paragraph id="C55C225FFFE539221AD2FBF0FA66FBC2" blockId="2.[151,1437,151,1757]" pageId="2" pageNumber="447">
<keyLead id="7E12816AFFE539221AD2FBF0FA66FBC2" pageId="2" pageNumber="447">
- Aedeagus Y-shaped, moderately narrow basally, slightly narrower than bases of parameres, distal portion slender, apex entire, pointed (
<figureCitation id="5DD83EDAFFE539221B72FB97FE84FBCD" box="[311,383,1090,1112]" captionStart="FIGURES 47–50" captionStartId="18.[151,264,1227,1251]" captionTargetBox="[250,1333,183,1189]" captionTargetId="figure-23@18.[242,1346,181,1203]" captionTargetPageId="18" captionText="FIGURES 47–50. Macrurohelea sirii n. sp. Holotype male, 47–48. Macrurohelea ventanensis n. sp. Holotype male, 49–50. 47, 49 parameres. 48, 50 aedeagus." figureDoi="http://doi.org/10.5281/zenodo.5912801" httpUri="https://zenodo.org/record/5912801/files/figure.png" pageId="2" pageNumber="447">Fig. 48</figureCitation>
); distal portion of parameres slender, apices tapered, divergent, with pointed tips (
<figureCitation id="5DD83EDAFFE539221ED7FB97FB23FBCD" box="[1170,1240,1090,1112]" captionStart="FIGURES 47–50" captionStartId="18.[151,264,1227,1251]" captionTargetBox="[250,1333,183,1189]" captionTargetId="figure-23@18.[242,1346,181,1203]" captionTargetPageId="18" captionText="FIGURES 47–50. Macrurohelea sirii n. sp. Holotype male, 47–48. Macrurohelea ventanensis n. sp. Holotype male, 49–50. 47, 49 parameres. 48, 50 aedeagus." figureDoi="http://doi.org/10.5281/zenodo.5912801" httpUri="https://zenodo.org/record/5912801/files/figure.png" pageId="2" pageNumber="447">Fig. 47</figureCitation>
).... 
<taxonomicName id="02E359DCFFE539221F5FFB96FA9AFBCD" ID-CoL="9HSG3" authority="Spinelli &amp; Ronderos &amp; Grogan, 2022" authorityName="Spinelli &amp; Ronderos &amp; Grogan" authorityYear="2022" box="[1306,1377,1091,1112]" class="Insecta" family="Ceratopogonidae" genus="Macrurohelea" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="447" phylum="Arthropoda" rank="species" species="sirii" status="sp. nov.">
<emphasis id="F797FE4DFFE539221F5FFB96FAD5FBCD" box="[1306,1326,1091,1112]" italics="true" pageId="2" pageNumber="447">M</emphasis>
. 
<emphasis id="F797FE4DFFE539221F7FFB96FA9AFBCD" box="[1338,1377,1091,1112]" italics="true" pageId="2" pageNumber="447">sirii</emphasis>
</taxonomicName>
<emphasis id="F797FE4DFFE539221F23FB97FA66FBC2" bold="true" box="[1382,1437,1090,1111]" pageId="2" pageNumber="447">
<taxonomicNameLabel id="ECA44336FFE539221F23FB97FA66FBC2" box="[1382,1437,1090,1111]" pageId="2" pageNumber="447" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
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</keyStep>
<keyStep id="7E173AFAFFE539221AD2FBB5FA67FB58" pageId="2" pageNumber="447">
<paragraph id="C55C225FFFE539221AD2FBB5FA66FB07" blockId="2.[151,1437,151,1757]" pageId="2" pageNumber="447">
<keyLead id="7E12816AFFE539221AD2FBB5FA66FB07" pageId="2" pageNumber="447">
23. Gonostylus abruptly curved at mid-length, distal ½ highly modified with basomesal hump and broad foot-shaped apex (
<figureCitation id="5DD83EDAFFE539221F10FB8AFA74FBE0" box="[1365,1423,1119,1141]" captionStart="FIGURES 1–7" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[214,1376,188,1920]" captionTargetId="figure-23@4.[210,1378,181,1920]" captionTargetPageId="4" captionText="FIGURES 1–7. Macrurohelea bassoi n. sp. Allotype female, 1–4. Holotype male, 5–7. 1, 5 head. 2 thorax. 3, 6 wing. 4 tip of abdomen. 7 genitalia. Scale bars = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5912783" httpUri="https://zenodo.org/record/5912783/files/figure.png" pageId="2" pageNumber="447">Fig. 7</figureCitation>
); distal portion of parameres with bulbous tips (
<figureCitation id="5DD83EDAFFE5392218D9FBA8FD1AFB07" box="[668,737,1149,1171]" captionStart="FIGURES 1–7" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[214,1376,188,1920]" captionTargetId="figure-23@4.[210,1378,181,1920]" captionTargetPageId="4" captionText="FIGURES 1–7. Macrurohelea bassoi n. sp. Allotype female, 1–4. Holotype male, 5–7. 1, 5 head. 2 thorax. 3, 6 wing. 4 tip of abdomen. 7 genitalia. Scale bars = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5912783" httpUri="https://zenodo.org/record/5912783/files/figure.png" pageId="2" pageNumber="447">Figs. 7</figureCitation>
, 
<figureCitation id="5DD83EDAFFE5392218AEFBA8FCFFFB06" box="[747,772,1149,1171]" captionStart="FIGURES 37–42" captionStartId="16.[151,264,1913,1937]" captionTargetBox="[214,1370,182,1882]" captionTargetId="figure-23@16.[214,1373,181,1889]" captionTargetPageId="16" captionText="FIGURES 37–42. Macrurohelea bassoi n. sp. Holotype male, 37–38. Macrurohelea donatoi n. sp. Holotype male, 39–40. Macrurohelea kuscheli Wirth, male, 41–42. 37, 39, 41 parameres. 38, 40, 42 aedeagus." figureDoi="http://doi.org/10.5281/zenodo.5912797" httpUri="https://zenodo.org/record/5912797/files/figure.png" pageId="2" pageNumber="447">37</figureCitation>
)......................................... 
<taxonomicName id="02E359DCFFE539221F46FBABFA9BFB07" ID-CoL="9HVMC" authority="Spinelli &amp; Ronderos &amp; Grogan, 2022" authorityName="Spinelli &amp; Ronderos &amp; Grogan" authorityYear="2022" box="[1283,1376,1149,1171]" class="Insecta" family="Ceratopogonidae" genus="Macrurohelea" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="447" phylum="Arthropoda" rank="species" species="bassoi" status="sp. nov.">
<emphasis id="F797FE4DFFE539221F46FBABFAECFB06" box="[1283,1303,1150,1171]" italics="true" pageId="2" pageNumber="447">M</emphasis>
. 
<emphasis id="F797FE4DFFE539221F66FBA8FA9BFB07" box="[1315,1376,1149,1170]" italics="true" pageId="2" pageNumber="447">bassoi</emphasis>
</taxonomicName>
<emphasis id="F797FE4DFFE539221F23FBA8FA66FB07" bold="true" box="[1382,1437,1149,1170]" pageId="2" pageNumber="447">
<taxonomicNameLabel id="ECA44336FFE539221F23FBA8FA66FB07" box="[1382,1437,1149,1170]" pageId="2" pageNumber="447" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
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</paragraph>
<paragraph id="C55C225FFFE539221AD2FB4EFA67FB58" blockId="2.[151,1437,151,1757]" pageId="2" pageNumber="447">
<keyLead id="7E12816AFFE539221AD2FB4EFA67FB58" pageId="2" pageNumber="447">
- Gonostylus curved or nearly straight, distal ½ not modified (
<figureCitation id="5DD83EDAFFE539221969FB4FFC86FB3A" box="[812,893,1178,1200]" captionStart="FIGURES 8–14" captionStartId="6.[151,264,1943,1967]" captionTargetBox="[186,1400,183,1920]" captionTargetId="figure-23@6.[186,1401,181,1920]" captionTargetPageId="6" captionText="FIGURES 8–14. Macrurohelea donatoi n. sp. Allotype female, 8–11. Holotype male, 12–14. 8, 12 head. 9 thorax. 10, 13 wing. 11 tip of abdomen. 14 genitalia. Scale bars = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5912785" httpUri="https://zenodo.org/record/5912785/files/figure.png" pageId="2" pageNumber="447">Figs. 14</figureCitation>
, 
<figureCitation id="5DD83EDAFFE5392219CDFB4FFC58FB25" box="[904,931,1178,1200]" captionStart="FIGURES 15–21" captionStartId="8.[151,264,1944,1968]" captionTargetBox="[193,1399,188,1907]" captionTargetId="figure-23@8.[189,1399,181,1920]" captionTargetPageId="8" captionText="FIGURES 15–21. Macrurohelea kuscheli Wirth, male, 15–18. Macrurohelea monotheca Spinelli &amp; Grogan, male, 19–21. 15, 19 head. 16 thorax. 17, 20 wing. 18-21 genitalia. Scale bars = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5912787" httpUri="https://zenodo.org/record/5912787/files/figure.png" pageId="2" pageNumber="447">21</figureCitation>
); distal portion of parameres nearly straight, slender or stout, without bulbous tip (
<figureCitation id="5DD83EDAFFE539221BBBFB62FDB4FB58" box="[510,591,1207,1229]" captionStart="FIGURES 37–42" captionStartId="16.[151,264,1913,1937]" captionTargetBox="[214,1370,182,1882]" captionTargetId="figure-23@16.[214,1373,181,1889]" captionTargetPageId="16" captionText="FIGURES 37–42. Macrurohelea bassoi n. sp. Holotype male, 37–38. Macrurohelea donatoi n. sp. Holotype male, 39–40. Macrurohelea kuscheli Wirth, male, 41–42. 37, 39, 41 parameres. 38, 40, 42 aedeagus." figureDoi="http://doi.org/10.5281/zenodo.5912797" httpUri="https://zenodo.org/record/5912797/files/figure.png" pageId="2" pageNumber="447">Figs. 39</figureCitation>
, 
<figureCitation id="5DD83EDAFFE53922181CFB6DFD89FB5B" box="[601,626,1208,1230]" captionStart="FIGURES 43–46" captionStartId="17.[151,264,1442,1466]" captionTargetBox="[166,1437,186,1402]" captionTargetId="figure-19@17.[151,1437,181,1418]" captionTargetPageId="17" captionText="FIGURES 43–46. Macrurohelea monotheca Spinelli &amp; Grogan, male, 43–44. Macrurohelea morenoi n. sp. Holotype male, 45–46. 43, 45 parameres. 44, 46 aedeagus." figureDoi="http://doi.org/10.5281/zenodo.5912799" httpUri="https://zenodo.org/record/5912799/files/figure.png" pageId="2" pageNumber="447">43</figureCitation>
)................................................................ 24
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<keyStep id="7E173AFAFFE539221AD2FB00FA67FA92" pageId="2" pageNumber="447">
<paragraph id="C55C225FFFE539221AD2FB00FA67FB7E" blockId="2.[151,1437,151,1757]" box="[151,1436,1237,1259]" pageId="2" pageNumber="447">
<keyLead id="7E12816AFFE539221AD2FB00FA67FB7E" box="[151,1436,1237,1259]" pageId="2" pageNumber="447">
24. Parameres disjunct (
<figureCitation id="5DD83EDAFFE539221BE0FB00FE0EFB7F" box="[421,501,1237,1259]" captionStart="FIGURES 37–42" captionStartId="16.[151,264,1913,1937]" captionTargetBox="[214,1370,182,1882]" captionTargetId="figure-23@16.[214,1373,181,1889]" captionTargetPageId="16" captionText="FIGURES 37–42. Macrurohelea bassoi n. sp. Holotype male, 37–38. Macrurohelea donatoi n. sp. Holotype male, 39–40. Macrurohelea kuscheli Wirth, male, 41–42. 37, 39, 41 parameres. 38, 40, 42 aedeagus." figureDoi="http://doi.org/10.5281/zenodo.5912797" httpUri="https://zenodo.org/record/5912797/files/figure.png" pageId="2" pageNumber="447">Figs. 39</figureCitation>
, 
<figureCitation id="5DD83EDAFFE539221BBAFB00FDE1FB7E" box="[511,538,1237,1259]" captionStart="FIGURES 43–46" captionStartId="17.[151,264,1442,1466]" captionTargetBox="[166,1437,186,1402]" captionTargetId="figure-19@17.[151,1437,181,1418]" captionTargetPageId="17" captionText="FIGURES 43–46. Macrurohelea monotheca Spinelli &amp; Grogan, male, 43–44. Macrurohelea morenoi n. sp. Holotype male, 45–46. 43, 45 parameres. 44, 46 aedeagus." figureDoi="http://doi.org/10.5281/zenodo.5912799" httpUri="https://zenodo.org/record/5912799/files/figure.png" pageId="2" pageNumber="447">43</figureCitation>
); gonostylus moderately curved with broad tip (
<figureCitation id="5DD83EDAFFE5392219A2FB00FBCCFB7F" box="[999,1079,1237,1259]" captionStart="FIGURES 8–14" captionStartId="6.[151,264,1943,1967]" captionTargetBox="[186,1400,183,1920]" captionTargetId="figure-23@6.[186,1401,181,1920]" captionTargetPageId="6" captionText="FIGURES 8–14. Macrurohelea donatoi n. sp. Allotype female, 8–11. Holotype male, 12–14. 8, 12 head. 9 thorax. 10, 13 wing. 11 tip of abdomen. 14 genitalia. Scale bars = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5912785" httpUri="https://zenodo.org/record/5912785/files/figure.png" pageId="2" pageNumber="447">Figs. 14</figureCitation>
, 
<figureCitation id="5DD83EDAFFE539221E04FB00FBA1FB7E" box="[1089,1114,1237,1259]" captionStart="FIGURES 15–21" captionStartId="8.[151,264,1944,1968]" captionTargetBox="[193,1399,188,1907]" captionTargetId="figure-23@8.[189,1399,181,1920]" captionTargetPageId="8" captionText="FIGURES 15–21. Macrurohelea kuscheli Wirth, male, 15–18. Macrurohelea monotheca Spinelli &amp; Grogan, male, 19–21. 15, 19 head. 16 thorax. 17, 20 wing. 18-21 genitalia. Scale bars = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5912787" httpUri="https://zenodo.org/record/5912787/files/figure.png" pageId="2" pageNumber="447">21</figureCitation>
)....................... 25
</keyLead>
</paragraph>
<paragraph id="C55C225FFFE539221AD2FB26FA67FA92" blockId="2.[151,1437,151,1757]" box="[151,1436,1266,1288]" pageId="2" pageNumber="447">
<keyLead id="7E12816AFFE539221AD2FB26FA67FA92" box="[151,1436,1266,1288]" pageId="2" pageNumber="447">- Parameres entire; gonostylus greatly curved with pointed tip.................................................. 27</keyLead>
</paragraph>
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<keyStep id="7E173AFAFFE539221AD2FADAFA67FAD6" pageId="2" pageNumber="447">
<paragraph id="C55C225FFFE539221AD2FADAFA67FAB0" blockId="2.[151,1437,151,1757]" box="[151,1436,1295,1317]" pageId="2" pageNumber="447">
<keyLead id="7E12816AFFE539221AD2FADAFA67FAB0" box="[151,1436,1295,1317]" pageId="2" pageNumber="447">
25. Distal portion of parameres disjunct at mid-length.................................... 
<taxonomicName id="02E359DCFFE539221E3CFAC5FA67FAB0" ID-CoL="3XD35" authority="Spinelli &amp; Grogan" authorityName="Spinelli &amp; Grogan" authorityYear="1999" box="[1145,1436,1295,1317]" class="Insecta" family="Ceratopogonidae" genus="Macrurohelea" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="447" phylum="Arthropoda" rank="species" species="yamana">
<emphasis id="F797FE4DFFE539221E3CFAC5FB76FAB0" box="[1145,1165,1296,1317]" italics="true" pageId="2" pageNumber="447">M</emphasis>
. 
<emphasis id="F797FE4DFFE539221EDCFAC5FB1EFAB0" box="[1177,1253,1296,1317]" italics="true" pageId="2" pageNumber="447">yamana</emphasis>
Spinelli &amp; Grogan
</taxonomicName>
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</paragraph>
<paragraph id="C55C225FFFE539221AD2FAF8FA67FAD6" blockId="2.[151,1437,151,1757]" box="[151,1436,1325,1347]" pageId="2" pageNumber="447">
<keyLead id="7E12816AFFE539221AD2FAF8FA67FAD6" box="[151,1436,1325,1347]" pageId="2" pageNumber="447">
- Distal portion of parameres disjunct subapically (
<figureCitation id="5DD83EDAFFE5392218F9FAF8FCF7FAD7" box="[700,780,1325,1347]" captionStart="FIGURES 37–42" captionStartId="16.[151,264,1913,1937]" captionTargetBox="[214,1370,182,1882]" captionTargetId="figure-23@16.[214,1373,181,1889]" captionTargetPageId="16" captionText="FIGURES 37–42. Macrurohelea bassoi n. sp. Holotype male, 37–38. Macrurohelea donatoi n. sp. Holotype male, 39–40. Macrurohelea kuscheli Wirth, male, 41–42. 37, 39, 41 parameres. 38, 40, 42 aedeagus." figureDoi="http://doi.org/10.5281/zenodo.5912797" httpUri="https://zenodo.org/record/5912797/files/figure.png" pageId="2" pageNumber="447">Figs. 39</figureCitation>
, 
<figureCitation id="5DD83EDAFFE539221953FAF8FCD4FAD6" box="[790,815,1325,1347]" captionStart="FIGURES 43–46" captionStartId="17.[151,264,1442,1466]" captionTargetBox="[166,1437,186,1402]" captionTargetId="figure-19@17.[151,1437,181,1418]" captionTargetPageId="17" captionText="FIGURES 43–46. Macrurohelea monotheca Spinelli &amp; Grogan, male, 43–44. Macrurohelea morenoi n. sp. Holotype male, 45–46. 43, 45 parameres. 44, 46 aedeagus." figureDoi="http://doi.org/10.5281/zenodo.5912799" httpUri="https://zenodo.org/record/5912799/files/figure.png" pageId="2" pageNumber="447">43</figureCitation>
)................................................. 26
</keyLead>
</paragraph>
</keyStep>
<keyStep id="7E173AFAFFE539221AD2FA9FFA67FA67" pageId="2" pageNumber="447">
<paragraph id="C55C225FFFE539221AD2FA9FFA66FA0F" blockId="2.[151,1437,151,1757]" pageId="2" pageNumber="447">
<keyLead id="7E12816AFFE539221AD2FA9FFA66FA0F" pageId="2" pageNumber="447">
26. Apicolateral process of tergite10 with short inner peg; sternite 9 with narrow shallow posteromedian excavation; gonostylus 0.85 length of gonocoxite (
<figureCitation id="5DD83EDAFFE539221BA3FAB2FDD5FAEB" box="[486,558,1383,1406]" captionStart="FIGURES 8–14" captionStartId="6.[151,264,1943,1967]" captionTargetBox="[186,1400,183,1920]" captionTargetId="figure-23@6.[186,1401,181,1920]" captionTargetPageId="6" captionText="FIGURES 8–14. Macrurohelea donatoi n. sp. Allotype female, 8–11. Holotype male, 12–14. 8, 12 head. 9 thorax. 10, 13 wing. 11 tip of abdomen. 14 genitalia. Scale bars = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5912785" httpUri="https://zenodo.org/record/5912785/files/figure.png" pageId="2" pageNumber="447">Fig. 14</figureCitation>
); aedeagus with convex lateral margins and slender basal arms (
<figureCitation id="5DD83EDAFFE539221ED0FAB2FB20FAEB" box="[1173,1243,1383,1406]" captionStart="FIGURES 37–42" captionStartId="16.[151,264,1913,1937]" captionTargetBox="[214,1370,182,1882]" captionTargetId="figure-23@16.[214,1373,181,1889]" captionTargetPageId="16" captionText="FIGURES 37–42. Macrurohelea bassoi n. sp. Holotype male, 37–38. Macrurohelea donatoi n. sp. Holotype male, 39–40. Macrurohelea kuscheli Wirth, male, 41–42. 37, 39, 41 parameres. 38, 40, 42 aedeagus." figureDoi="http://doi.org/10.5281/zenodo.5912797" httpUri="https://zenodo.org/record/5912797/files/figure.png" pageId="2" pageNumber="447">Fig. 40</figureCitation>
).................................................................................................... 
<taxonomicName id="02E359DCFFE539221EB2FA53FA9BFA0F" ID-CoL="9HVZP" authority="Spinelli &amp; Ronderos &amp; Grogan, 2022" authorityName="Spinelli &amp; Ronderos &amp; Grogan" authorityYear="2022" box="[1271,1376,1413,1435]" class="Insecta" family="Ceratopogonidae" genus="Macrurohelea" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="447" phylum="Arthropoda" rank="species" species="donatoi" status="sp. nov.">
<emphasis id="F797FE4DFFE539221EB2FA53FAF0FA0E" box="[1271,1291,1414,1435]" italics="true" pageId="2" pageNumber="447">M</emphasis>
. 
<emphasis id="F797FE4DFFE539221F52FA50FA9BFA0F" box="[1303,1376,1413,1434]" italics="true" pageId="2" pageNumber="447">donatoi</emphasis>
</taxonomicName>
<emphasis id="F797FE4DFFE539221F23FA50FA66FA0F" bold="true" box="[1382,1437,1413,1434]" pageId="2" pageNumber="447">
<taxonomicNameLabel id="ECA44336FFE539221F23FA50FA66FA0F" box="[1382,1437,1413,1434]" pageId="2" pageNumber="447" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="C55C225FFFE539221AD2FA76FA67FA67" blockId="2.[151,1437,151,1757]" pageId="2" pageNumber="447">
<keyLead id="7E12816AFFE539221AD2FA76FA67FA67" pageId="2" pageNumber="447">
- Apicolateral process of tergite 10 without short peg; sternite 9 with deep posteromedian excavation; gonostylus 0.69 length of gonocoxite (
<figureCitation id="5DD83EDAFFE539221B1FFA6AFE59FA43" box="[346,418,1471,1494]" captionStart="FIGURES 15–21" captionStartId="8.[151,264,1944,1968]" captionTargetBox="[193,1399,188,1907]" captionTargetId="figure-23@8.[189,1399,181,1920]" captionTargetPageId="8" captionText="FIGURES 15–21. Macrurohelea kuscheli Wirth, male, 15–18. Macrurohelea monotheca Spinelli &amp; Grogan, male, 19–21. 15, 19 head. 16 thorax. 17, 20 wing. 18-21 genitalia. Scale bars = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5912787" httpUri="https://zenodo.org/record/5912787/files/figure.png" pageId="2" pageNumber="447">Fig. 21</figureCitation>
); aedeagus with straight lateral margins and stout basal arms with truncate apices (
<figureCitation id="5DD83EDAFFE539221EFEFA6AFAFAFA43" box="[1211,1281,1471,1494]" captionStart="FIGURES 43–46" captionStartId="17.[151,264,1442,1466]" captionTargetBox="[166,1437,186,1402]" captionTargetId="figure-19@17.[151,1437,181,1418]" captionTargetPageId="17" captionText="FIGURES 43–46. Macrurohelea monotheca Spinelli &amp; Grogan, male, 43–44. Macrurohelea morenoi n. sp. Holotype male, 45–46. 43, 45 parameres. 44, 46 aedeagus." figureDoi="http://doi.org/10.5281/zenodo.5912799" httpUri="https://zenodo.org/record/5912799/files/figure.png" pageId="2" pageNumber="447">Fig. 44</figureCitation>
)...................................................................................... 
<taxonomicName id="02E359DCFFE539221E19FA0BFA67FA67" ID-CoL="3XD2X" authority="Spinelli &amp; Grogan" authorityName="Spinelli &amp; Grogan" authorityYear="1984" box="[1116,1436,1501,1523]" class="Insecta" family="Ceratopogonidae" genus="Macrurohelea" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="447" phylum="Arthropoda" rank="species" species="monotheca">
<emphasis id="F797FE4DFFE539221E19FA0BFB8BFA66" box="[1116,1136,1502,1523]" italics="true" pageId="2" pageNumber="447">M</emphasis>
. 
<emphasis id="F797FE4DFFE539221E39FA08FB1EFA67" box="[1148,1253,1501,1522]" italics="true" pageId="2" pageNumber="447">monotheca</emphasis>
Spinelli &amp; Grogan
</taxonomicName>
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</paragraph>
</keyStep>
<keyStep id="7E173AFAFFE539221AD2FA2FFA67F9F2" pageId="2" pageNumber="447">
<paragraph id="C55C225FFFE539221AD2FA2FFA67F9B8" blockId="2.[151,1437,151,1757]" pageId="2" pageNumber="447">
<keyLead id="7E12816AFFE539221AD2FA2FFA67F9B8" pageId="2" pageNumber="447">
27. Gonostylus nearly straight, moderately short (length 
<quantity id="021B8FBAFFE539221898FA2FFCCDF985" box="[733,822,1530,1552]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="447" unit="mm" value="0.5">0.50 mm</quantity>
); distal portion of parameres with tips bent 90°............................................................................................ 
<taxonomicName id="02E359DCFFE539221EE5F9CDFA67F9B8" ID-CoL="6R3WW" authority="Grogan &amp; Wirth" authorityName="Grogan &amp; Wirth" box="[1184,1436,1559,1581]" class="Insecta" family="Ceratopogonidae" genus="Macrurohelea" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="447" phylum="Arthropoda" rank="species" species="irwini">
<emphasis id="F797FE4DFFE539221EE5F9CDFB4FF9B8" box="[1184,1204,1560,1581]" italics="true" pageId="2" pageNumber="447">M</emphasis>
. 
<emphasis id="F797FE4DFFE539221E85F9CDFB02F9B8" box="[1216,1273,1560,1581]" italics="true" pageId="2" pageNumber="447">irwini</emphasis>
Grogan &amp; Wirth
</taxonomicName>
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</paragraph>
<paragraph id="C55C225FFFE539221AD2F9E0FA67F9F2" blockId="2.[151,1437,151,1757]" pageId="2" pageNumber="447">
<keyLead id="7E12816AFFE539221AD2F9E0FA67F9F2" pageId="2" pageNumber="447">
- Gonostylus greatly curved, elongate (length 
<quantity id="021B8FBAFFE5392218CAF9E0FCE4F9DF" box="[655,799,1589,1611]" metricMagnitude="-4" metricUnit="m" metricValue="8.1" metricValueMax="8.5" metricValueMin="7.7" pageId="2" pageNumber="447" unit="mm" value="0.81" valueMax="0.85" valueMin="0.77">0.77–0.85 mm</quantity>
); distal portion of parameres tapering near apex with slightly bent pointed tips......................................................................................... 28
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<paragraph id="C55C225FFFE539221AD2F9A5FA67F937" blockId="2.[151,1437,151,1757]" pageId="2" pageNumber="447">
<keyLead id="7E12816AFFE539221AD2F9A5FA67F937" pageId="2" pageNumber="447">
28. Cerci elongate, extending beyond tergite 10; aedeagus triangular, basal arch extending ½ of total length, tip pointed; distal portions of parameres divergent.................................................... 
<taxonomicName id="02E359DCFFE539221EC3F95BFA67F937" ID-CoL="3XD32" authority="Spinelli &amp; Grogan" authorityName="Spinelli &amp; Grogan" authorityYear="1990" box="[1158,1436,1677,1699]" class="Insecta" family="Ceratopogonidae" genus="Macrurohelea" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="447" phylum="Arthropoda" rank="species" species="similis">
<emphasis id="F797FE4DFFE539221EC3F95BFB61F936" box="[1158,1178,1678,1699]" italics="true" pageId="2" pageNumber="447">M</emphasis>
. 
<emphasis id="F797FE4DFFE539221EE3F958FB1EF937" box="[1190,1253,1677,1698]" italics="true" pageId="2" pageNumber="447">similis</emphasis>
Spinelli &amp; Grogan
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="C55C225FFFE539221AD2F97EFA67F948" blockId="2.[151,1437,151,1757]" pageId="2" pageNumber="447">
<keyLead id="7E12816AFFE539221AD2F97EFA67F948" pageId="2" pageNumber="447">
- Cerci shorter, not extending beyond tergite 10; aedeagus with very shallow basal arch, tip truncate; distal portions of parameres parallel....................................................................... 
<taxonomicName id="02E359DCFFE539221EC4F91DFA67F948" ID-CoL="3XD2P" authority="Ingram &amp; Macfie" authorityName="Ingram &amp; Macfie" authorityYear="1931" box="[1153,1436,1735,1757]" class="Insecta" family="Ceratopogonidae" genus="Macrurohelea" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="447" phylum="Arthropoda" rank="species" species="caudata">
<emphasis id="F797FE4DFFE539221EC4F91DFB6EF948" box="[1153,1173,1736,1757]" italics="true" pageId="2" pageNumber="447">M</emphasis>
. 
<emphasis id="F797FE4DFFE539221EE4F912FB15F949" box="[1185,1262,1735,1756]" italics="true" pageId="2" pageNumber="447">caudata</emphasis>
Ingram &amp; Macfie
</taxonomicName>
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</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>