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<document ID-CLB-Dataset="265156" ID-DOI="http://dx.doi.org/10.3897/zookeys.1178.108929" ID-GBIF-Dataset="9d81d7e5-4459-4d19-9953-b65e6b42fc25" ID-PMC="PMC10492037" ID-Pensoft-Pub="1313-2970-1178-69" ID-Pensoft-UUID="0D0038AB24DB57888D1D81AFDFF6DB43" ID-PubMed="37692917" ID-ZooBank="8D54500C9C9D4D8B8BC7547CF1A70A68" ModsDocID="1313-2970-1178-69" checkinTime="1693642879942" checkinUser="pensoft" docAuthor="Claridge, Brandon R., Kaunisto, Kari M. &amp; Saeaeksjaervi, Ilari E." docDate="2023" docId="46CCB75E629F56DCBA5A762996FAC673" docLanguage="en" docName="ZooKeys 1178: 69-76" docOrigin="ZooKeys 1178" docPubDate="2023-09-01" docSource="http://dx.doi.org/10.3897/zookeys.1178.108929" docTitle="Capitojoppa Claridge, Saeaeksjaervi &amp; Kaunisto 2023, gen. nov." docType="treatment" docUuid="DCE30183-1262-42CE-A34C-6348D04AF9B3" docUuidSource="ZooBank" docVersion="4" id="0D0038AB24DB57888D1D81AFDFF6DB43" lastPageNumber="69" masterDocId="0D0038AB24DB57888D1D81AFDFF6DB43" masterDocTitle="Capitojoppa, a new genus of Ichneumoninae (Hymenoptera, Ichneumonidae) from Peruvian Amazonia" masterLastPageNumber="76" masterPageNumber="69" pageNumber="69" updateTime="1732725628027" updateUser="ExternalLinkService">
<mods:mods id="751AAC3CAB1778A5C5E7F9445290865A" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="5A3849DB467899F1AC3515A52A318027">
<mods:title id="3DEEDA9669D891CD41A14543120A0CA8">Capitojoppa, a new genus of Ichneumoninae (Hymenoptera, Ichneumonidae) from Peruvian Amazonia</mods:title>
</mods:titleInfo>
<mods:name id="97D7359193B674E71BBB616F2A9C6A54" type="personal">
<mods:role id="70050EF745EA8BE50AB09A9AC77B406A">
<mods:roleTerm id="B7E44485C49EFD7255294CBC663EBB18">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="017A679CB36A0D9055246F033B021168">Claridge, Brandon R.</mods:namePart>
<mods:nameIdentifier id="92503F1F5E0E3E77D41B73AC362BECE3" type="ORCID">https://orcid.org/0000-0002-2222-326X</mods:nameIdentifier>
<mods:affiliation id="835D3ABCE89210B23D0D603F794B702E">Utah State University, 5305 Old Main Hill, Logan, UT, 84322 USA</mods:affiliation>
<mods:nameIdentifier id="9BBBB84E98CD793B3B31018A4B35471A" type="email">brandonclaridge1@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="D8078B79E99665C6A4776206BDCB3FD8" type="personal">
<mods:role id="1CAB6EBB1E1ED0849F11149961F76A57">
<mods:roleTerm id="6575E4879A36F604C9AE78DC98DC9FFB">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E143962B5A833908CE28D9444B42A334">Kaunisto, Kari M.</mods:namePart>
<mods:nameIdentifier id="CC9D6D8D85E721622EBF26E812B5BB9D" type="ORCID">https://orcid.org/0000-0001-6665-0047</mods:nameIdentifier>
<mods:affiliation id="8A827E143796AC3B4E3021C929ECD0B3">Biodiversity Unit, Zoological Museum, University of Turku, 20014 Turku, Finland</mods:affiliation>
</mods:name>
<mods:name id="6ED250304FA6D55FAB3FEFB8CC578A45" type="personal">
<mods:role id="EA054A684E11DA60087D7F5E0709EB33">
<mods:roleTerm id="4A29C1DAFA7139ADC16097F85F4B301A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="114DB394E0F7D900DE0821BF1D8278FE">Saeaeksjaervi, Ilari E.</mods:namePart>
<mods:nameIdentifier id="F5DB47337B5D0FA98F3606237D52EA1B" type="ORCID">https://orcid.org/0000-0002-8107-5607</mods:nameIdentifier>
<mods:affiliation id="FD8502B032DB8AAA9C47023A7E117D66">Biodiversity Unit, Zoological Museum, University of Turku, 20014 Turku, Finland</mods:affiliation>
</mods:name>
<mods:typeOfResource id="C7B7064B465621A4F555E92190F5B367">text</mods:typeOfResource>
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<mods:titleInfo id="6B8D3C7BC6805F09CBA589E0DFF2109E">
<mods:title id="8A2BC1195A973A4C1157CCCE181B0553">ZooKeys</mods:title>
</mods:titleInfo>
<mods:part id="75A12CFEF341824E3D7E6CD8709AAE4E">
<mods:date id="4CA0A294B51E4AF01B9060E68D82E0E5">2023</mods:date>
<mods:detail id="B9281D2F72E65DBEA60DA88F330ABF7E" type="pubDate">
<mods:number id="C740BC2D899664C4544BBD50078C6B78">2023-09-01</mods:number>
</mods:detail>
<mods:detail id="6BBF3391FBF22C1685F316032F464944" type="volume">
<mods:number id="3E712A6A90D4045B507D7F6CFB8CECA9">1178</mods:number>
</mods:detail>
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<mods:start id="37D58BF21FFEEDBF369AD8DD84CB2A00">69</mods:start>
<mods:end id="D89C276C1BB5BD2AE45636E2EBC5DE61">76</mods:end>
</mods:extent>
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<mods:location id="E9AB6F26D0D3518894C256B80F3E1C1E">
<mods:url id="D3B8584ADD3F0EB6D7EAAC0CEB36F1B7">http://dx.doi.org/10.3897/zookeys.1178.108929</mods:url>
</mods:location>
<mods:classification id="8EBEA3F80C1393375FB01C2962A5867C">journal article</mods:classification>
<mods:identifier id="B1A5170806ED7CD0D926FCFEB14BF05E" type="DOI">http://dx.doi.org/10.3897/zookeys.1178.108929</mods:identifier>
<mods:identifier id="C7CCE6EE4715ADF1D3765594EF0B3EC1" type="Pensoft-Pub">1313-2970-1178-69</mods:identifier>
<mods:identifier id="A5D5BD36CA56799C52EE102362999F7E" type="ZooBank">8D54500C9C9D4D8B8BC7547CF1A70A68</mods:identifier>
<mods:identifier id="737E9A600574748A2EF6E9217BAD30A3" type="Pensoft-UUID">0D0038AB24DB57888D1D81AFDFF6DB43</mods:identifier>
</mods:mods>
<treatment id="46CCB75E629F56DCBA5A762996FAC673" ID-GBIF-Taxon="212746804" LSID="urn:lsid:zoobank.org:act:DCE30183-1262-42CE-A34C-6348D04AF9B3" httpUri="http://treatment.plazi.org/id/46CCB75E629F56DCBA5A762996FAC673" lastPageNumber="69" pageId="0" pageNumber="69" scope_family="Ichneumonidae" scope_order="Hymenoptera">
<subSubSection id="2FFBEAAA9001A6E766B1DF7245B0CECE" pageId="0" pageNumber="69" type="nomenclature">
<paragraph id="8C93E1C1889740FA1903A12BB68C6F8B" pageId="0" pageNumber="69">
<taxonomicName id="95D168B4BC33B30C931D19AE7B7FC07F" LSID="https://zoobank.org/DCE30183-1262-42CE-A34C-6348D04AF9B3" authority="Claridge, Sääksjärvi &amp; Kaunisto" authorityName="Claridge, Saeaeksjaervi &amp; Kaunisto" authorityYear="2023" class="Insecta" family="Ichneumonidae" genus="Capitojoppa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capitojoppa" order="Hymenoptera" pageId="0" pageNumber="69" phylum="Arthropoda" rank="genus" status="gen. nov.">
Capitojoppa Claridge,
<normalizedToken id="4E993E21067A500800CBE186C4FA887F" originalValue="Sääksjärvi">Saeaeksjaervi</normalizedToken>
&amp; Kaunisto
</taxonomicName>
<taxonomicNameLabel id="7C712583232E6CB939216C5DCBBD6C09" pageId="0" pageNumber="69">gen. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="3393991BA271B0BD395EF5EE2156684B" pageId="0" pageNumber="69" type="description">
<paragraph id="F254C7E59DA0E944E33D40DDA5D42EA3" pageId="0" pageNumber="69">
<figureCitation id="3DEFABFC265900DC19DD8FAA35979540" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Capitojoppa amazonica sp. nov., holotype female A habitus B mesosoma, lateral view C head, anterior view D head, ventral view E mesosoma, dorsal view. Scale bars: 5 mm (A, B, E); 2 mm (C, D)." figureDoi="10.3897/zookeys.1178.108929.figure1" httpUri="https://binary.pensoft.net/fig/900094" pageId="0" pageNumber="69">Figs 1</figureCitation>
<figureCitation id="6C3A9E3F3658D9D16CE0D817C1CC483F" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Capitojoppa amazonica sp. nov., paratype male A habitus B head, lateral view C propodeum, dorsal view D hind coxae and metasoma, dorsal view. Scale bars: 5 mm (A, D); 2 mm (B, C)." figureDoi="10.3897/zookeys.1178.108929.figure2" httpUri="https://binary.pensoft.net/fig/900095" pageId="0" pageNumber="69">, 2</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="0561BB9545E7B8553EC9ED4021D58F4F" pageId="0" pageNumber="69" type="type species">
<paragraph id="C5FBC12687E07F651461553CD96E251D" pageId="0" pageNumber="69">Type species.</paragraph>
<paragraph id="6017B3F69049F94C5B3A4604C3C243F1" pageId="0" pageNumber="69">
<taxonomicName id="CD461554994337C948AE8FF86F4A375B" authorityName="Claridge, Saeaeksjaervi &amp; Kaunisto" authorityYear="2023" class="Insecta" family="Ichneumonidae" genus="Capitojoppa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capitojoppa amazonica" order="Hymenoptera" pageId="0" pageNumber="69" phylum="Arthropoda" rank="species" species="amazonica">
<emphasis id="962A729B4C0F577FFA9832A541491BFA" italics="true" pageId="0" pageNumber="69">Capitojoppa amazonica</emphasis>
</taxonomicName>
Claridge,
<normalizedToken id="CBCF690952316384162DDC109B940D6C" originalValue="Sääksjärvi">Saeaeksjaervi</normalizedToken>
&amp; Kaunisto, sp. nov.
</paragraph>
</subSubSection>
<subSubSection id="C0FEF7CC2E4F8A525930DAB5CF974B05" pageId="0" pageNumber="69" type="diagnosis">
<paragraph id="0EC9EF3602903C14188D56BD36499B1D" pageId="0" pageNumber="69">Diagnosis.</paragraph>
<paragraph id="6438AE7411B18E9A7C89CA369706F669" pageId="0" pageNumber="69">
<taxonomicName id="6C2A16937568EFCD5E246CB315BB73C3" authorityName="Claridge, Saeaeksjaervi &amp; Kaunisto" authorityYear="2023" class="Insecta" family="Ichneumonidae" genus="Capitojoppa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capitojoppa" order="Hymenoptera" pageId="0" pageNumber="69" phylum="Arthropoda" rank="genus">
<emphasis id="4BA4212D9FB7874F734D2FEB445A7F46" italics="true" pageId="0" pageNumber="69">Capitojoppa</emphasis>
</taxonomicName>
belongs to the tribe
<taxonomicName id="9D78EB7B3CE0A946B5138EC76CE32FCF" authorityName="Latreille" authorityYear="1802" lsidName="" pageId="0" pageNumber="69" rank="tribe" tribe="Ichneumonini">Ichneumonini</taxonomicName>
and is diagnosed by the following combination of characters: overall robust body with an enlarged head and a wide gena; large mandibles with widely spaced, subequal teeth; labrum exposed and visible in anterior view; supraclypeal area and clypeus without lateral ridge as in
<taxonomicName id="0C4BDF9F32D882F791435D60F67749F7" class="Insecta" family="Ichneumonidae" genus="Joppa" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Joppa" order="Hymenoptera" pageId="0" pageNumber="69" phylum="Arthropoda" rank="genus">
<emphasis id="F2CB637D01A018B961B79FCEB42C548F" italics="true" pageId="0" pageNumber="69">Joppa</emphasis>
</taxonomicName>
Fabricius and
<taxonomicName id="42C470FB18D706840DB0672C511F5D37" authorityName="Townes" authorityYear="1936" class="Insecta" family="Ichneumonidae" genus="Projoppa" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Projoppa" order="Hymenoptera" pageId="0" pageNumber="69" phylum="Arthropoda" rank="genus">
<emphasis id="4900DC1126A859C202ABB5C0343C0EE3" italics="true" pageId="0" pageNumber="69">Projoppa</emphasis>
</taxonomicName>
Townes; occipital carina complete and meeting hypostomal carina at mandibular base; wing membrane clear; propodeum without distinct dorsal and posterior faces (anterior margin strongly sloping and propodeum gently sloping posteriorly); dorsomedial surface of hind coxa coarsely striate; hind coxa without scopa; T1 longitudinally striate; T2 longitudinally striate medially; gastrocoelus deeply impressed; thyridium well developed; and female metasomal apex oxypygous.
</paragraph>
<paragraph id="EE570EC10ABA44FECFA3518B1436C8C4" pageId="0" pageNumber="69">
Overall,
<taxonomicName id="28D89273A137E1A81111941C93CF2983" authorityName="Claridge, Saeaeksjaervi &amp; Kaunisto" authorityYear="2023" class="Insecta" family="Ichneumonidae" genus="Capitojoppa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capitojoppa" order="Hymenoptera" pageId="0" pageNumber="69" phylum="Arthropoda" rank="genus">
<emphasis id="1085C047EDC7A518E53C1B21970D67D3" italics="true" pageId="0" pageNumber="69">Capitojoppa</emphasis>
</taxonomicName>
is most similar to
<taxonomicName id="88EC53E758E518FCAA0E7BDF0FAAE843" class="Insecta" family="Ichneumonidae" genus="Joppa" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Joppa" order="Hymenoptera" pageId="0" pageNumber="69" phylum="Arthropoda" rank="genus">
<emphasis id="7DC196CD65A238D4FBDB4925E435F6BF" italics="true" pageId="0" pageNumber="69">Joppa</emphasis>
</taxonomicName>
due to the following characters: mandibles large with subequal teeth; propodeum lacking distinct dorsal and posterior faces due to both the anterior and posterior margins sloping (variable in
<taxonomicName id="CFF038FD4FA73C64EF06AB9234231058" class="Insecta" family="Ichneumonidae" genus="Joppa" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Joppa" order="Hymenoptera" pageId="0" pageNumber="69" phylum="Arthropoda" rank="genus">
<emphasis id="0E287BB5F0C674F1307912B15B245E20" italics="true" pageId="0" pageNumber="69">Joppa</emphasis>
</taxonomicName>
but usually as in
<taxonomicName id="1AD07FB098650725622884EDFD8693D0" authorityName="Claridge, Saeaeksjaervi &amp; Kaunisto" authorityYear="2023" class="Insecta" family="Ichneumonidae" genus="Capitojoppa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capitojoppa" order="Hymenoptera" pageId="0" pageNumber="69" phylum="Arthropoda" rank="genus">
<emphasis id="204DCD0AEA767A556D9E0ED70B2895FC" italics="true" pageId="0" pageNumber="69">Capitojoppa</emphasis>
</taxonomicName>
); gastrocoelus deeply impressed; thyridium well developed; and T1-2 coarsely, longitudinally striate.
<taxonomicName id="F8785B553F05936F168717C65F02561E" authorityName="Claridge, Saeaeksjaervi &amp; Kaunisto" authorityYear="2023" class="Insecta" family="Ichneumonidae" genus="Capitojoppa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capitojoppa" order="Hymenoptera" pageId="0" pageNumber="69" phylum="Arthropoda" rank="genus">
<emphasis id="0FD12684D7252903E730C5FCDB49A4ED" italics="true" pageId="0" pageNumber="69">Capitojoppa</emphasis>
</taxonomicName>
primarily differs from
<taxonomicName id="42766F81B809B7D08380A6754696683D" class="Insecta" family="Ichneumonidae" genus="Joppa" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Joppa" order="Hymenoptera" pageId="0" pageNumber="69" phylum="Arthropoda" rank="genus">
<emphasis id="3202186D8E50D926B8B2A60D84B378EB" italics="true" pageId="0" pageNumber="69">Joppa</emphasis>
</taxonomicName>
by the exposed labrum which is visible in anterior view (concealed by the elongated clypeus in
<taxonomicName id="3FB0157E97B15E117AD33DA8E2D89935" class="Insecta" family="Ichneumonidae" genus="Joppa" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Joppa" order="Hymenoptera" pageId="0" pageNumber="69" phylum="Arthropoda" rank="genus">
<emphasis id="EDCF410E970150B194DC63EB86919C8C" italics="true" pageId="0" pageNumber="69">Joppa</emphasis>
</taxonomicName>
); the lack of distinct lateral ridges on the supraclypeal area and clypeus (a defining character in
<taxonomicName id="191139E876955B62292FBF88A389AC55" class="Insecta" family="Ichneumonidae" genus="Joppa" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Joppa" order="Hymenoptera" pageId="0" pageNumber="69" phylum="Arthropoda" rank="genus">
<emphasis id="1114437DE13A0CBFEC6E6B5FD4725D43" italics="true" pageId="0" pageNumber="69">Joppa</emphasis>
</taxonomicName>
); the coarse, longitudinal striation on the supra-antennal area (smooth and without striation or rugosity in
<taxonomicName id="294728216D2AB4135E308E60E52CCCBE" class="Insecta" family="Ichneumonidae" genus="Joppa" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Joppa" order="Hymenoptera" pageId="0" pageNumber="69" phylum="Arthropoda" rank="genus">
<emphasis id="1EC7D6CC61E1502ADF8D82D9CFB3B584" italics="true" pageId="0" pageNumber="69">Joppa</emphasis>
</taxonomicName>
); complete occipital carina (obsolete ventrally in
<taxonomicName id="E9F779B78A9CAF56E607A723A700B40C" class="Insecta" family="Ichneumonidae" genus="Joppa" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Joppa" order="Hymenoptera" pageId="0" pageNumber="69" phylum="Arthropoda" rank="genus">
<emphasis id="A27240DAE9F3DB1D722924B9E3A0A3D0" italics="true" pageId="0" pageNumber="69">Joppa</emphasis>
</taxonomicName>
); clear wings (yellow with black markings in
<taxonomicName id="31F497F31EE6B1B6844E787412167045" class="Insecta" family="Ichneumonidae" genus="Joppa" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Joppa" order="Hymenoptera" pageId="0" pageNumber="69" phylum="Arthropoda" rank="genus">
<emphasis id="9F080056DD1BB986D91C43F4DB2A860F" italics="true" pageId="0" pageNumber="69">Joppa</emphasis>
</taxonomicName>
); and coarse striation on dorsomedial surface of the hind coxa (smooth and without striation or rugosity in
<taxonomicName id="2172CE1F9B247F3B1CCA1726F76DC34C" class="Insecta" family="Ichneumonidae" genus="Joppa" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Joppa" order="Hymenoptera" pageId="0" pageNumber="69" phylum="Arthropoda" rank="genus">
<emphasis id="F01231B12EE038A94B103E4CBE7096CC" italics="true" pageId="0" pageNumber="69">Joppa</emphasis>
</taxonomicName>
). Also, the habitus is considerably more robust than any
<taxonomicName id="7A38D3432E7C4878CC0F9D47C245BC7D" class="Insecta" family="Ichneumonidae" genus="Joppa" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Joppa" order="Hymenoptera" pageId="0" pageNumber="69" phylum="Arthropoda" rank="genus">
<emphasis id="ED985B084918CE3F8D61A276B57BC819" italics="true" pageId="0" pageNumber="69">Joppa</emphasis>
</taxonomicName>
species examined.
</paragraph>
</subSubSection>
<subSubSection id="D477C35131558DE87A752CDC14EFEE48" pageId="0" pageNumber="69" type="description">
<paragraph id="3B3FF26A211CBFF41F4F8529AB0D6B4E" pageId="0" pageNumber="69">Description.</paragraph>
<paragraph id="09517D81753E8C84A55BF75E0F1B386E" pageId="0" pageNumber="69">
<emphasis id="5DE949486E64B3D6FC57F1D158827AC1" bold="true" pageId="0" pageNumber="69">
<emphasis id="285C90A4422A679D3FCCAE083EA85FF3" italics="true" pageId="0" pageNumber="69">Head</emphasis>
.
</emphasis>
Mandible large with subequal, widely separated teeth. Clypeus flat and with ventral margin straight. Epistomal suture subobsolete. Malar space ~0.7
<normalizedToken id="883AC9C82CE9473F44924D87485E241D" originalValue="×">x</normalizedToken>
basal mandibular width. Clypeus and supraclypeal area simple, without sublateral ridges. Dorsal margin of median field with small medial tubercle. Supra-antennal area coarsely, longitudinally striate. Gena wide. Female flagellum bristle-shaped (flattened and widened past midpoint before acutely tapering apically); basal flagellomeres moderately slender (flagellomere 1 ~3.0
<normalizedToken id="56BF36686F7A2738767D09D392A47D3C" originalValue="×">x</normalizedToken>
as wide as long). Male flagellum with tyloids present and beginning on flagellomere 10 (endpoint of tyloids unknown); bristle ridges present beginning on flagellomere 11 (development of bristle ridges uncertain due to missing flagellum past flagellomere 15 on male paratype).
</paragraph>
<paragraph id="AF72ED15A09F77A9944346DE336D3322" pageId="0" pageNumber="69">
<emphasis id="9269D6E9D233CDEC6B8418786686ED8A" bold="true" pageId="0" pageNumber="69">
<emphasis id="6FC5C05F88A85049C3A46C388CEF1D77" italics="true" pageId="0" pageNumber="69">Mesosoma</emphasis>
.
</emphasis>
Epomia present. Notaulus obsolete. Scutellum flat, not significantly elevated above postscutellum; not laterally carinate. Sternaulus obsolete. Posterior transverse carinae of the mesothoracic venter only present laterally. Juxtacoxal carina present. Propodeal carinae largely obsolete with only anterior transverse carina well developed medially and longitudinal carinae developed at extreme posterior (carinae more strongly developed in male specimen). Anterior margin of propodeum without medial tubercle. Legs stout. Fore and mid tibiae with well-developed, decumbent, conical spines (more strongly developed in female). Hind coxa without scopa and with dorsomedial surface coarsely striate. Tarsal claws simple, not pectinate.
</paragraph>
<paragraph id="073C7EF36902E8BCF1D4C72DA6FA0C42" pageId="0" pageNumber="69">
<emphasis id="3D540E78679626CFEF31CDAC06DAA263" bold="true" pageId="0" pageNumber="69">
<emphasis id="E0A3D9C54AA5A774EB0C92FC889BA6CD" italics="true" pageId="0" pageNumber="69">Wings</emphasis>
.
</emphasis>
Areolet pentagonal. Distal abscissa of 1/Cu present.
</paragraph>
<paragraph id="71AF9D170B2D32AB1B61585443E59D23" pageId="0" pageNumber="69">
<emphasis id="77DDA66356D9488C61803295FF05B47A" bold="true" pageId="0" pageNumber="69">
<emphasis id="1EFCB044293EA9AC5E18CA735213EEB6" italics="true" pageId="0" pageNumber="69">Metasoma</emphasis>
.
</emphasis>
Metasoma rather stout, but linear and weakly tapering apically. T1 coarsely, longitudinally striate, median field indistinct. T2 longitudinally striate medially with striate area becoming narrower posteriorly. Thyridium well developed and wide (thyridium ~1.2
<normalizedToken id="C7ED13AA1D73C8A0F0ECC57DE248C3D7" originalValue="×">x</normalizedToken>
interthyridial width). Gastrocoelus deeply impressed. Female metasomal apex oxypygous. Ovipositor not significantly projecting past metasomal apex. Male hypopygium simple, not medially elongated. Gonoforceps simple, not enlarged.
</paragraph>
</subSubSection>
<subSubSection id="F253D43E4DCA40E0FD236F03B837E6C3" pageId="0" pageNumber="69" type="etymology">
<paragraph id="8B3819819F0AB6E5576CFEDAC4556221" pageId="0" pageNumber="69">Etymology.</paragraph>
<paragraph id="F2A7F5630347E26A04AEC0344DB42890" pageId="0" pageNumber="69">
Derived from
<emphasis id="4650AB3D7985397A9E771CD4B05C72DD" italics="true" pageId="0" pageNumber="69">capito</emphasis>
, meaning big-headed in medieval Latin, combined with the generic name,
<emphasis id="EB17D96B0568635C41625A882C48A00C" italics="true" pageId="0" pageNumber="69">Joppa</emphasis>
. In addition,
<emphasis id="380DA988EB9D0C4A4F44D593059DC667" italics="true" pageId="0" pageNumber="69">Capito</emphasis>
refers to the Neotropical bird genus
<emphasis id="0E370A377BAD26A6925FF65B478C242E" italics="true" pageId="0" pageNumber="69">Capito</emphasis>
(barbets), including species of stocky birds with heavy bills. Gender feminine.
</paragraph>
</subSubSection>
<subSubSection id="42D8110C9771D78926CA1820F28D69F4" pageId="0" pageNumber="69" type="comments">
<paragraph id="1AF52365438666D83C86DA9EEE1956DD" pageId="0" pageNumber="69">Comments.</paragraph>
<paragraph id="584CCD47CF73AD32DA00E4D44D10254A" pageId="0" pageNumber="69">Sexual dimorphism is minimal with only minor colour and surface sculpture differences between the female and male specimens.</paragraph>
<paragraph id="80FE3D918F5FA46A43F7F36A5371D4B6" pageId="0" pageNumber="69">
The similarity to
<taxonomicName id="9A6FE6578D0F786F5C24FD9EA49C2B81" class="Insecta" family="Ichneumonidae" genus="Joppa" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Joppa" order="Hymenoptera" pageId="0" pageNumber="69" phylum="Arthropoda" rank="genus">
<emphasis id="1CA03710B48FF1639501294E331973B9" italics="true" pageId="0" pageNumber="69">Joppa</emphasis>
</taxonomicName>
and other characters typical of the group (females with bristle-shaped antennae and males with bristle-ridges) suggests that
<taxonomicName id="A40D9233395ABDECE1FE5DE7B71961C8" authorityName="Claridge, Saeaeksjaervi &amp; Kaunisto" authorityYear="2023" class="Insecta" family="Ichneumonidae" genus="Capitojoppa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capitojoppa" order="Hymenoptera" pageId="0" pageNumber="69" phylum="Arthropoda" rank="genus">
<emphasis id="F7D95E1CD8CCAC9C3B711080BC76883F" italics="true" pageId="0" pageNumber="69">Capitojoppa</emphasis>
</taxonomicName>
belongs to a clade of Neotropical genera that includes
<taxonomicName id="41AB9D16A3A90DBA8E9CC532909F059E" authorityName="Hancock" authorityYear="1926" class="Insecta" family="Ichneumonidae" genus="Carinodes" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Carinodes" order="Hymenoptera" pageId="0" pageNumber="69" phylum="Arthropoda" rank="genus">
<emphasis id="BEE67226EB85CEDF0D56F2A795D7649C" italics="true" pageId="0" pageNumber="69">Carinodes</emphasis>
</taxonomicName>
Hancock,
<taxonomicName id="34EC83434B038E2E03E43D99E77F0E5B" class="Insecta" family="Ichneumonidae" genus="Joppa" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Joppa" order="Hymenoptera" pageId="0" pageNumber="69" phylum="Arthropoda" rank="genus">
<emphasis id="905C93A7DF94B974A4DCC859DE7D59FB" italics="true" pageId="0" pageNumber="69">Joppa</emphasis>
</taxonomicName>
,
<taxonomicName id="C53B626066E6C456DCFC8CF0DF981D70" authorityName="Viereck" authorityYear="1913" class="Insecta" family="Ichneumonidae" genus="Joppocryptus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Joppocryptus" order="Hymenoptera" pageId="0" pageNumber="69" phylum="Arthropoda" rank="genus">
<emphasis id="F9503710B40A772B5A56DB1C5455F92D" italics="true" pageId="0" pageNumber="69">Joppocryptus</emphasis>
</taxonomicName>
Viereck,
<taxonomicName id="866030ABCB49249FD36E6E3AEC5A7473" authorityName="Townes" authorityYear="1946" class="Insecta" family="Ichneumonidae" genus="Limonethe" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Limonethe" order="Hymenoptera" pageId="0" pageNumber="69" phylum="Arthropoda" rank="genus">
<emphasis id="8C750A74D2EA9F1728BD891E750083D9" italics="true" pageId="0" pageNumber="69">Limonethe</emphasis>
</taxonomicName>
Townes,
<taxonomicName id="03BB873A7291D955C97BAEFFBB2AC412" authorityName="Townes" authorityYear="1946" class="Insecta" family="Ichneumonidae" genus="Lobaegis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Lobaegis" order="Hymenoptera" pageId="0" pageNumber="69" phylum="Arthropoda" rank="genus">
<emphasis id="5EFD7646B101A05895FAB6B80EC7AEAC" italics="true" pageId="0" pageNumber="69">Lobaegis</emphasis>
</taxonomicName>
Townes,
<taxonomicName id="C7E67CE3839077001168408A8F135538" authorityName="Brethes" authorityYear="1927" class="Insecta" family="Ichneumonidae" genus="Lophojoppa" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Lophojoppa" order="Hymenoptera" pageId="0" pageNumber="69" phylum="Arthropoda" rank="genus">
<emphasis id="FD74E3F1C933BE2075003EE4F29C7902" italics="true" pageId="0" pageNumber="69">Lophojoppa</emphasis>
</taxonomicName>
<normalizedToken id="BA75B7FCCB7160E6E04AEBC8F1FCF9E2" originalValue="Brèthes">Brethes</normalizedToken>
,
<taxonomicName id="B9F3977F2CDFB8C89EB31912EDF85401" authorityName="Townes" authorityYear="1946" class="Insecta" family="Ichneumonidae" genus="Narthecura" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Narthecura" order="Hymenoptera" pageId="0" pageNumber="69" phylum="Arthropoda" rank="genus">
<emphasis id="03D9B55168BF5D8244E2380D47A79CF3" italics="true" pageId="0" pageNumber="69">Narthecura</emphasis>
</taxonomicName>
Townes,
<taxonomicName id="0299B998AA17D9655B76654BEEABE259" authorityName="Townes" authorityYear="1946" class="Insecta" family="Ichneumonidae" genus="Notacma" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Notacma" order="Hymenoptera" pageId="0" pageNumber="69" phylum="Arthropoda" rank="genus">
<emphasis id="D59EF0559552B8A9156F3EE4C7F970D2" italics="true" pageId="0" pageNumber="69">Notacma</emphasis>
</taxonomicName>
Townes,
<taxonomicName id="9CEB2AC93AC5AF3DA17F0705DE9F0E49" authorityName="Ozdikmen &amp; Turgut" authorityYear="2006" class="Insecta" family="Ichneumonidae" genus="Oezdemirus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Oezdemirus" order="Hymenoptera" pageId="0" pageNumber="69" phylum="Arthropoda" rank="genus">
<emphasis id="FB75124BA0F4957737162D0FED64016E" italics="true" pageId="0" pageNumber="69">Oezdemirus</emphasis>
</taxonomicName>
Ozdikmen &amp; Turgut,
<taxonomicName id="5F82694ABE195CBBD3C3B1AD5522D660" authorityName="Cresson" authorityYear="1873" class="Insecta" family="Ichneumonidae" genus="Ortezia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Ortezia" order="Hymenoptera" pageId="0" pageNumber="69" phylum="Arthropoda" rank="genus">
<emphasis id="6DF8469AC31A3FC09101ADF0CC9EB2B9" italics="true" pageId="0" pageNumber="69">Ortezia</emphasis>
</taxonomicName>
Cresson, and
<taxonomicName id="AC2F5B7E09A628024A4821EA21440CBA" authorityName="Ashmead" authorityYear="1900" class="Insecta" family="Ichneumonidae" genus="Trogomorpha" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Trogomorpha" order="Hymenoptera" pageId="0" pageNumber="69" phylum="Arthropoda" rank="genus">
<emphasis id="8D935D03496CC1A045B607D30C1CD29D" italics="true" pageId="0" pageNumber="69">Trogomorpha</emphasis>
</taxonomicName>
Ashmead (
<bibRefCitation id="B94F538CA616268E53AC2C3E0B58401D" DOI="https://doi.org/10.1111/syen.12484" author="Santos, BF" journalOrPublisher="Systematic Entomology" pageId="0" pageNumber="69" pagination="704 - 724" refId="B8" refString="Santos, BF, Wahl, DB, Rousse, P, Bennett, AMR, Kula, R, Brady, SG, 2021. Phylogenomics of Ichneumoninae (Hymenoptera, Ichneumonidae) reveals pervasive morphological convergence and the shortcomings of previous classifications. Systematic Entomology 46 (3): 704 - 724, DOI: https://doi.org/10.1111/syen.12484" title="Phylogenomics of Ichneumoninae (Hymenoptera, Ichneumonidae) reveals pervasive morphological convergence and the shortcomings of previous classifications." url="https://doi.org/10.1111/syen.12484" volume="46" year="2021">Santos et al. 2021</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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