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<document id="4271B607841D0601EC2734CA4874E7CE" ID-CLB-Dataset="22469" ID-DOI="10.5281/zenodo.3776470" ID-GBIF-Dataset="95dd362c-59b3-409a-8846-a51b6e839ff1" ID-ISSN="0253-116X" ID-Zenodo-Dep="3776470" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1588185693586" checkinUser="valdenar" docAuthor="Assing, Volker" docDate="2018" docId="423287F433385C2AC7E1FA25AE60E2C3" docLanguage="en" docName="Linzer.biol.Beitr.50.2.977-996.pdf.imf" docOrigin="Linzer biologische Beiträge 50 (2)" docStyle="DocumentStyle:24E5EB772D355DE2EE23B1E8D07853E0.9:LinzerbiolBeitr.2006-.journal_article" docStyleId="24E5EB772D355DE2EE23B1E8D07853E0" docStyleName="LinzerbiolBeitr.2006-.journal_article" docStyleVersion="9" docTitle="Scymbalium rossii Assing 2018, nov.sp." docType="treatment" docVersion="6" lastPageNumber="980" masterDocId="BE0BFF8C333A5C29C753FF9BAC47E722" masterDocTitle="A revision of Palaearctic and Oriental Scymbalium and Micrillus III. New species, new combinations, and additional records (Coleoptera: Staphylinidae: Paederinae)" masterLastPageNumber="996" masterPageNumber="977" pageNumber="979" updateTime="1698838054887" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="1B882FE25E4988C3D7AA38BCB31872E9">A revision of Palaearctic and Oriental Scymbalium and Micrillus III. New species, new combinations, and additional records (Coleoptera: Staphylinidae: Paederinae)</mods:title>
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<mods:namePart id="2714585A15104067724C790FCA94BE42">Assing, Volker</mods:namePart>
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<mods:title id="0B273FA9F765BB89D1662AA56FA30674">Linzer biologische Beiträge</mods:title>
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<mods:date id="EABC26AEE576F992111AAC1A0145FF1D">2018</mods:date>
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<treatment id="423287F433385C2AC7E1FA25AE60E2C3" ID-DOI="http://doi.org/10.5281/zenodo.3796183" ID-GBIF-Taxon="163877104" ID-Zenodo-Dep="3796183" LSID="urn:lsid:plazi:treatment:423287F433385C2AC7E1FA25AE60E2C3" httpUri="http://treatment.plazi.org/id/423287F433385C2AC7E1FA25AE60E2C3" lastPageId="3" lastPageNumber="980" pageId="2" pageNumber="979">
<subSubSection id="8281656933385C2BC7E1FA25AD86E11F" pageId="2" pageNumber="979" type="nomenclature">
<paragraph id="CA2436E233385C2BC7E1FA25AE68E2F6" blockId="2.[178,559,1470,1492]" box="[178,559,1470,1492]" pageId="2" pageNumber="979">
<heading id="916C818E33385C2BC7E1FA25AE68E2F6" box="[178,559,1470,1492]" fontSize="8" level="2" pageId="2" pageNumber="979" reason="2">
<emphasis id="F8EFEAF033385C2BC7E1FA25AD83E2F1" bold="true" box="[178,452,1470,1492]" pageId="2" pageNumber="979">
<taxonomicName id="0D9B4D6133385C2BC7E1FA25AD36E2F1" ID-CoL="9V8WJ" authority="Assing, 2018" authorityName="Assing" authorityYear="2018" box="[178,369,1470,1492]" class="Insecta" family="Staphylinidae" genus="Scymbalium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="979" phylum="Arthropoda" rank="species" species="rossii" status="sp. nov.">
<emphasis id="F8EFEAF033385C2BC7E1FA25AD36E2F1" bold="true" box="[178,369,1470,1492]" italics="true" pageId="2" pageNumber="979">Scymbalium rossii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E3DC578B33385C2BC62BFA25AD83E2F1" box="[376,452,1470,1491]" pageId="2" pageNumber="979" rank="species">nov.sp.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="52A02A6733385C2BC682FA25AE61E2F6" box="[465,550,1470,1492]" captionStart-0="Figs 1-8" captionStart-1="Fig" captionStartId-0="4.[178,218,1522,1544]" captionStartId-1="5.[178,211,909,931]" captionTargetBox-0="[180,1084,227,1502]" captionTargetBox-1="[177,1085,214,895]" captionTargetId-0="figure@4.[177,1084,227,1512]" captionTargetId-1="figure@5.[177,1085,214,895]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionText-0="Figs 1-8: Scymbalium rossii: (1) habitus; (2) forebody; (3) abdomen; (4) male sternite VIII; (5-6) aedeagus in lateral and in ventral view; (7-8) apex of aedeagus in lateral and in ventral view. Scale bars: 1-3: 1.0 mm; 4: 0.5 mm; 5-6: 0.2 mm; 7-8: 0.1 mm." captionText-1="Fig. 9: Type locality of Scymbalium rossii, Micrillus virgatus, M. kongi, and M. coloratus. Photo: Walter Rossi." figureDoi-0="http://doi.org/10.5281/zenodo.3776472" figureDoi-1="http://doi.org/10.5281/zenodo.3776474" httpUri-0="https://zenodo.org/record/3776472/files/figure.png" httpUri-1="https://zenodo.org/record/3776474/files/figure.png" pageId="2" pageNumber="979">Figs 1-9</figureCitation>
)
</heading>
</paragraph>
<paragraph id="CA2436E233385C2BC7E1FA78AD86E11F" blockId="2.[178,1086,1507,1597]" pageId="2" pageNumber="979">
T y p e m a t e r i a l:
<materialsCitation id="7AF33CBF33385C2BC6E1FA78AD86E11F" ID-GBIF-Occurrence="2609445623" collectingDate="2018-05-21" collectorName="Bernardi" country="Cambodia" determinerName="V. Assing" latitude="12.237223" location="Toul Ompel" longLatPrecision="21" longitude="104.6875" pageId="2" pageNumber="979" specimenCount="4" specimenCount-female="1" specimenCount-male="2" stateProvince="Kampong Chhnang prov." typeStatus="holotype">
<typeStatus id="1520884033385C2BC6E1FA78AE40E2DB" box="[434,519,1507,1529]" pageId="2" pageNumber="979" type="holotype">Holotype</typeStatus>
<specimenCount id="DC9DFD6B33385C2BC540FA78AE60E2DA" box="[531,551,1507,1528]" pageId="2" pageNumber="979" type="male"></specimenCount>
: &quot;
<collectingCountry id="B28C767233385C2BC56DFA78AEFBE2DA" box="[574,700,1507,1528]" name="Cambodia" pageId="2" pageNumber="979">CAMBODIA</collectingCountry>
-
<collectingRegion id="085FF80033385C2BC58EFA78AF93E2DA" box="[733,980,1507,1529]" country="Cambodia" name="Kampong Chhnang" pageId="2" pageNumber="979">Kampong Chhnang prov.</collectingRegion>
,
<collectingRegion id="085FF80033385C2BC4B0FA78AD43E12D" country="Cambodia" name="Kampong Chhnang" pageId="2" pageNumber="979">Kampong Chhnang</collectingRegion>
env.,
<location id="CF44603933385C2BC617FA62ADFEE12D" LSID="urn:lsid:plazi:treatment:423287F433385C2AC7E1FA25AE60E2C3:CF44603933385C2BC617FA62ADFEE12D" box="[324,441,1529,1551]" country="Cambodia" latitude="12.237223" longLatPrecision="21" longitude="104.6875" name="Toul Ompel" pageId="2" pageNumber="979" stateProvince="Kampong Chhnang prov.">Toul Ompel</location>
,
<geoCoordinate id="AFAF502533385C2BC694FA62AE74E12C" box="[455,563,1529,1550]" degrees="12" direction="north" minutes="14" orientation="latitude" pageId="2" pageNumber="979" precision="15" seconds="14" value="12.237223">12°14'14''N</geoCoordinate>
,
<geoCoordinate id="AFAF502533385C2BC513FA62AEF2E12D" box="[576,693,1529,1551]" degrees="104" direction="east" minutes="41" orientation="longitude" pageId="2" pageNumber="979" precision="15" seconds="15" value="104.6875">104°41'15''E</geoCoordinate>
, light trap, 19+
<date id="BE25102233385C2BC41EFA62AFE8E12C" box="[845,943,1529,1550]" pageId="2" pageNumber="979" value="2018-05-21">
<collectingDate id="AE61E9CA33385C2BC41EFA62AFE8E12C" box="[845,943,1529,1550]" pageId="2" pageNumber="979" value="2018-05-21">21.V.2018</collectingDate>
</date>
, leg.
<collectorName id="676E533433385C2BC4B4FA62A87EE12D" box="[999,1081,1529,1551]" pageId="2" pageNumber="979">Bernardi</collectorName>
,
<location id="CF44603933385C2BC7E1F98BAD70E104" LSID="urn:lsid:plazi:treatment:423287F433385C2AC7E1FA25AE60E2C3:CF44603933385C2BC7E1F98BAD70E104" box="[178,311,1552,1574]" country="Cambodia" latitude="12.237223" longLatPrecision="21" longitude="104.6875" name="Kong &amp; Rossi" pageId="2" pageNumber="979" stateProvince="Kampong Chhnang prov.">Kong &amp; Rossi</location>
/
<typeStatus id="1520884033385C2BC61EF98BADEBE104" box="[333,428,1552,1574]" pageId="2" pageNumber="979" type="holotype">Holotypus</typeStatus>
<specimenCount id="DC9DFD6B33385C2BC6E0F98BAD83E107" box="[435,452,1552,1573]" pageId="2" pageNumber="979" type="male"></specimenCount>
<taxonomicName id="0D9B4D6133385C2BC698F98BAE32E104" authority="Assing, 2018" authorityName="Assing" authorityYear="2018" box="[459,629,1552,1574]" class="Insecta" family="Staphylinidae" genus="Scymbalium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="979" phylum="Arthropoda" rank="species" species="rossii" status="sp. nov.">
<emphasis id="F8EFEAF033385C2BC698F98BAE32E104" box="[459,629,1552,1574]" italics="true" pageId="2" pageNumber="979">
<location id="CF44603933385C2BC698F98BAE7DE107" LSID="urn:lsid:plazi:treatment:423287F433385C2AC7E1FA25AE60E2C3:CF44603933385C2BC698F98BAE7DE107" box="[459,570,1552,1573]" country="Cambodia" latitude="12.237223" longLatPrecision="21" longitude="104.6875" name="Scymbalium" pageId="2" pageNumber="979" stateProvince="Kampong Chhnang prov.">Scymbalium</location>
rossii
</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E3DC578B33385C2BC52FF98BAEF7E107" box="[636,688,1552,1573]" pageId="2" pageNumber="979" rank="species">
<specimenCount id="DC9DFD6B33385C2BC52FF98BAED1E107" box="[636,662,1552,1573]" pageId="2" pageNumber="979" type="generic">sp.</specimenCount>
n.
</taxonomicNameLabel>
, det.
<determinerName id="3FF8DA2633385C2BC5B7F98BAF07E104" box="[740,832,1552,1574]" pageId="2" pageNumber="979">V. Assing</determinerName>
2018&quot; (cAss).
<typeStatus id="1520884033385C2BC49DF98BA85AE107" box="[974,1053,1552,1573]" pageId="2" pageNumber="979" type="paratype">Paratype</typeStatus>
<specimenCount id="DC9DFD6B33385C2BC376F98BA87FE107" box="[1061,1080,1552,1573]" pageId="2" pageNumber="979" type="female"></specimenCount>
: same data as
<typeStatus id="1520884033385C2BC679F9BCAD3DE11F" box="[298,378,1575,1597]" pageId="2" pageNumber="979" type="holotype">holotype</typeStatus>
(cAss).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="8281656933395C2AC7E1FF40AE60E2C3" pageId="3" pageNumber="980" type="description">
<paragraph id="CA2436E233395C2AC7E1FF40AFB1E62F" blockId="3.[178,1085,219,270]" pageId="3" pageNumber="980">E t y m o l o g y: This species is dedicated to Walter Rossi (L'Aquila), to whom I am grateful for the generous gift of all the Cambodian specimens treated in this paper.</paragraph>
<paragraph id="CA2436E233395C2AC7E1FE85AFE0E6A9" blockId="3.[178,1085,285,395]" pageId="3" pageNumber="980">
D e s c r i p t i o n: Body length 7.0-
<quantity id="0D639B0733395C2AC516FE86AED0E611" box="[581,663,285,307]" metricMagnitude="-3" metricUnit="m" metricValue="7.7" pageId="3" pageNumber="980" unit="mm" value="7.7">7.7 mm</quantity>
; length of forebody
<quantity id="0D639B0733395C2AC439FE86AFA4E611" box="[874,995,285,307]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" metricValueMax="4.1" metricValueMin="3.7" pageId="3" pageNumber="980" unit="mm" value="3.9" valueMax="4.1" valueMin="3.7">3.7-4.1 mm</quantity>
. Habitus as in
<figureCitation id="52A02A6733395C2AC7B8FEA0AD6DE672" box="[235,298,315,337]" captionStart="Figs 1-8" captionStartId="4.[178,218,1522,1544]" captionTargetBox="[180,1084,227,1502]" captionTargetId="figure@4.[177,1084,227,1512]" captionTargetPageId="4" captionText="Figs 1-8: Scymbalium rossii: (1) habitus; (2) forebody; (3) abdomen; (4) male sternite VIII; (5-6) aedeagus in lateral and in ventral view; (7-8) apex of aedeagus in lateral and in ventral view. Scale bars: 1-3: 1.0 mm; 4: 0.5 mm; 5-6: 0.2 mm; 7-8: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3776472" httpUri="https://zenodo.org/record/3776472/files/figure.png" pageId="3" pageNumber="980">Fig. 1</figureCitation>
. Body somewhat depressed. Coloration: head and pronotum reddish-brown; elytra reddish; abdomen dark-brown to blackish-brown with the apex (segments VIII-X and the posterior portion of VII) reddish; legs yellowish-red; antennae red.
</paragraph>
<paragraph id="CA2436E233395C2AC7E1FE01AF5DE55F" blockId="3.[178,1085,410,637]" pageId="3" pageNumber="980">
Head (
<figureCitation id="52A02A6733395C2AC7A5FE01AD74E693" box="[246,307,410,433]" captionStart="Figs 1-8" captionStartId="4.[178,218,1522,1544]" captionTargetBox="[180,1084,227,1502]" captionTargetId="figure@4.[177,1084,227,1512]" captionTargetPageId="4" captionText="Figs 1-8: Scymbalium rossii: (1) habitus; (2) forebody; (3) abdomen; (4) male sternite VIII; (5-6) aedeagus in lateral and in ventral view; (7-8) apex of aedeagus in lateral and in ventral view. Scale bars: 1-3: 1.0 mm; 4: 0.5 mm; 5-6: 0.2 mm; 7-8: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3776472" httpUri="https://zenodo.org/record/3776472/files/figure.png" pageId="3" pageNumber="980">Fig. 2</figureCitation>
) 1.10-1.15 times as broad as long; integument with double punctation, with coarse and dense macropunctures and with interspersed micropunctures; median dorsal portion with somewhat sparser punctation; interstices without microsculpture. Eyes approximately as long as postocular region in dorsal view. Antenna long and very slen- der, approximately 3.0 mm long; antennomeres IV-XI at least approximately twice as long as broad and club-shaped, i.e., basally very thin and apically distincty dilated, broadest anterior to apex. Maxillary palpus with short and conical apical palpomere, this palpomere approximately one-third as long as palpomere III.
</paragraph>
<paragraph id="CA2436E233395C2AC7E1FD16ADF2E43A" blockId="3.[178,1085,652,792]" pageId="3" pageNumber="980">
Pronotum (
<figureCitation id="52A02A6733395C2AC675FD16AD22E581" box="[294,357,653,675]" captionStart="Figs 1-8" captionStartId="4.[178,218,1522,1544]" captionTargetBox="[180,1084,227,1502]" captionTargetId="figure@4.[177,1084,227,1512]" captionTargetPageId="4" captionText="Figs 1-8: Scymbalium rossii: (1) habitus; (2) forebody; (3) abdomen; (4) male sternite VIII; (5-6) aedeagus in lateral and in ventral view; (7-8) apex of aedeagus in lateral and in ventral view. Scale bars: 1-3: 1.0 mm; 4: 0.5 mm; 5-6: 0.2 mm; 7-8: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3776472" httpUri="https://zenodo.org/record/3776472/files/figure.png" pageId="3" pageNumber="980">Fig. 2</figureCitation>
) 1.12-1.15 times as broad as long, broadest near anterior angles, distinctly tapering posteriad; lateral margins straight or weakly sinuate in the middle; posterior margin weakly concave in the middle; macropunctation dense, slightly less coarse than that of head; micropunctation scattered, sparser than that of head; midline moderately broadly impunctate.
</paragraph>
<paragraph id="CA2436E233395C2AC7E1FCBCAFC7E455" blockId="3.[178,1085,807,888]" pageId="3" pageNumber="980">
Elytra (
<figureCitation id="52A02A6733395C2AC651FCBCAD04E41F" box="[258,323,807,829]" captionStart="Figs 1-8" captionStartId="4.[178,218,1522,1544]" captionTargetBox="[180,1084,227,1502]" captionTargetId="figure@4.[177,1084,227,1512]" captionTargetPageId="4" captionText="Figs 1-8: Scymbalium rossii: (1) habitus; (2) forebody; (3) abdomen; (4) male sternite VIII; (5-6) aedeagus in lateral and in ventral view; (7-8) apex of aedeagus in lateral and in ventral view. Scale bars: 1-3: 1.0 mm; 4: 0.5 mm; 5-6: 0.2 mm; 7-8: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3776472" httpUri="https://zenodo.org/record/3776472/files/figure.png" pageId="3" pageNumber="980">Fig. 2</figureCitation>
) approximately as long as pronotum; punctation rather fine and dense. Hind wings fully developed. Protarsomeres IIV distinctly dilated in both sexes. Metatarsomere I approximately as long as the combined length of II and III.
</paragraph>
<paragraph id="CA2436E233395C2AC7E1FC1CAE17E30D" blockId="3.[178,1085,903,1071]" pageId="3" pageNumber="980">
Abdomen (
<figureCitation id="52A02A6733395C2AC676FC1CAD23E4BF" box="[293,356,903,925]" captionStart="Figs 1-8" captionStartId="4.[178,218,1522,1544]" captionTargetBox="[180,1084,227,1502]" captionTargetId="figure@4.[177,1084,227,1512]" captionTargetPageId="4" captionText="Figs 1-8: Scymbalium rossii: (1) habitus; (2) forebody; (3) abdomen; (4) male sternite VIII; (5-6) aedeagus in lateral and in ventral view; (7-8) apex of aedeagus in lateral and in ventral view. Scale bars: 1-3: 1.0 mm; 4: 0.5 mm; 5-6: 0.2 mm; 7-8: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3776472" httpUri="https://zenodo.org/record/3776472/files/figure.png" pageId="3" pageNumber="980">Fig. 3</figureCitation>
) approximately as broad as elytra; punctation distinct and very dense, similar to that of elytra; interstices glossy, with extremely fine and nearly obsolete transverse microsculpture visible only at high magnification (100 x); posterior margin of tergite VII with palisade fringe; tergite X and sternite IX unmodified, not of reduced size; postero-lateral processes of segment IX approximately 1.5 times as long as tergite X, apically with a spine curved upwards.
</paragraph>
<paragraph id="CA2436E233395C2AC7E1FBA4AED5E3AD" blockId="3.[178,1085,1086,1167]" pageId="3" pageNumber="980">
<specimenCount id="DC9DFD6B33395C2AC7E1FBA4AC8FE376" box="[178,200,1087,1108]" pageId="3" pageNumber="980" type="male"></specimenCount>
: posterior excision of sternite VIII narrow and short, about one-eighth a long as length of sternite (
<figureCitation id="52A02A6733395C2AC678FBC7AD2BE350" box="[299,364,1116,1138]" captionStart="Figs 1-8" captionStartId="4.[178,218,1522,1544]" captionTargetBox="[180,1084,227,1502]" captionTargetId="figure@4.[177,1084,227,1512]" captionTargetPageId="4" captionText="Figs 1-8: Scymbalium rossii: (1) habitus; (2) forebody; (3) abdomen; (4) male sternite VIII; (5-6) aedeagus in lateral and in ventral view; (7-8) apex of aedeagus in lateral and in ventral view. Scale bars: 1-3: 1.0 mm; 4: 0.5 mm; 5-6: 0.2 mm; 7-8: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3776472" httpUri="https://zenodo.org/record/3776472/files/figure.png" pageId="3" pageNumber="980">Fig. 4</figureCitation>
); aedeagus (
<figureCitation id="52A02A6733395C2AC6BDFBC7AE01E350" box="[494,582,1116,1138]" captionStart="Figs 1-8" captionStartId="4.[178,218,1522,1544]" captionTargetBox="[180,1084,227,1502]" captionTargetId="figure@4.[177,1084,227,1512]" captionTargetPageId="4" captionText="Figs 1-8: Scymbalium rossii: (1) habitus; (2) forebody; (3) abdomen; (4) male sternite VIII; (5-6) aedeagus in lateral and in ventral view; (7-8) apex of aedeagus in lateral and in ventral view. Scale bars: 1-3: 1.0 mm; 4: 0.5 mm; 5-6: 0.2 mm; 7-8: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3776472" httpUri="https://zenodo.org/record/3776472/files/figure.png" pageId="3" pageNumber="980">Figs 5-8</figureCitation>
) slender, with short hooked apex (lateral view), and with heavily sclerotized internal structures.
</paragraph>
<paragraph id="CA2436E233395C2AC7E1FB04AF6BE246" blockId="3.[178,1085,1182,1380]" pageId="3" pageNumber="980">
C o m p a r a t i v e n o t e s: This species is evidently closely allied to
<taxonomicName id="0D9B4D6133395C2AC4EDFB04A857E396" baseAuthorityName="Nordmann" baseAuthorityYear="1837" box="[958,1040,1183,1204]" class="Insecta" family="Staphylinidae" genus="Scymbalium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="980" phylum="Arthropoda" rank="species" species="anale">
<emphasis id="F8EFEAF033395C2AC4EDFB04A857E396" box="[958,1040,1183,1204]" italics="true" pageId="3" pageNumber="980">S. anale</emphasis>
</taxonomicName>
and
<taxonomicName id="0D9B4D6133395C2AC7E1FB27AD6DE3F3" baseAuthorityName="Cameron" baseAuthorityYear="1940" box="[178,298,1212,1233]" class="Insecta" family="Staphylinidae" genus="Scymbalium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="980" phylum="Arthropoda" rank="species" species="persimile">
<emphasis id="F8EFEAF033395C2AC7E1FB27AD6DE3F3" box="[178,298,1212,1233]" italics="true" pageId="3" pageNumber="980">S. persimile</emphasis>
</taxonomicName>
. It is distinguished from both of them by smaller size, paler coloration of the forebody, and coarser punctation of the abdomen. It additionally differs from
<taxonomicName id="0D9B4D6133395C2AC499FB42A85BE3CC" baseAuthorityName="Nordmann" baseAuthorityYear="1837" box="[970,1052,1241,1262]" class="Insecta" family="Staphylinidae" genus="Scymbalium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="980" phylum="Arthropoda" rank="species" species="anale">
<emphasis id="F8EFEAF033395C2AC499FB42A85BE3CC" box="[970,1052,1241,1262]" italics="true" pageId="3" pageNumber="980">S. anale</emphasis>
</taxonomicName>
by a posteriorly less strongly tapering pronotum, relatively larger eyes, a much shorter and narrower posterior excision of the male sternite VIII, and by a much smaller aedeagus (
<taxonomicName id="0D9B4D6133395C2AC379FA8FACADE264" baseAuthorityName="Nordmann" baseAuthorityYear="1837" class="Insecta" family="Staphylinidae" genus="Scymbalium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="980" phylum="Arthropoda" rank="species" species="anale">
<emphasis id="F8EFEAF033395C2AC379FA8FACADE264" italics="true" pageId="3" pageNumber="980">S. anale</emphasis>
</taxonomicName>
: aedeagus approximately 2.0 mm long) with an apex of different shape in lateral view. The male sexual characters of
<taxonomicName id="0D9B4D6133395C2AC577FAD4AEDCE241" baseAuthorityName="Cameron" baseAuthorityYear="1940" box="[548,667,1358,1380]" class="Insecta" family="Staphylinidae" genus="Scymbalium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="980" phylum="Arthropoda" rank="species" species="persimile">
<emphasis id="F8EFEAF033395C2AC577FAD4AEDCE241" box="[548,667,1358,1380]" italics="true" pageId="3" pageNumber="980">S. persimile</emphasis>
</taxonomicName>
are unknown.
</paragraph>
<paragraph id="CA2436E233395C2AC7E1FAEFAE60E2C3" blockId="3.[178,1085,1395,1505]" pageId="3" pageNumber="980">
D i s t r i b u t i o n a n d n a t u r a l h i s t o r y: The
<typeStatus id="1520884033395C2AC414FAEFAF34E2AB" box="[839,883,1396,1417]" pageId="3" pageNumber="980">type</typeStatus>
locality is situated near
<collectingRegion id="085FF80033395C2AC7B5FA0AADEAE285" box="[230,429,1425,1447]" country="Cambodia" name="Kampong Chhnang" pageId="3" pageNumber="980">Kampong Chhnang</collectingRegion>
in the estuary of the effluent of Tonle Sap lake (
<figureCitation id="52A02A6733395C2AC4E3FA0AAFA8E285" box="[944,1007,1425,1447]" captionStart="Fig" captionStartId="5.[178,211,909,931]" captionTargetBox="[177,1085,214,895]" captionTargetId="figure@5.[177,1085,214,895]" captionTargetPageId="5" captionText="Fig. 9: Type locality of Scymbalium rossii, Micrillus virgatus, M. kongi, and M. coloratus. Photo: Walter Rossi." figureDoi="http://doi.org/10.5281/zenodo.3776474" httpUri="https://zenodo.org/record/3776474/files/figure.png" pageId="3" pageNumber="980">Fig. 9</figureCitation>
),
<collectingCountry id="B28C767233395C2AC356FA0AACAAE2E6" name="Cambodia" pageId="3" pageNumber="980">Cambodia</collectingCountry>
. The specimens were collected with a light trap, together with seven species of
<taxonomicName id="0D9B4D6133395C2AC7E1FA50AD48E2C2" box="[178,271,1483,1504]" class="Insecta" family="Staphylinidae" genus="Micrillus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="980" phylum="Arthropoda" rank="genus">
<emphasis id="F8EFEAF033395C2AC7E1FA50AD48E2C2" box="[178,271,1483,1504]" italics="true" pageId="3" pageNumber="980">Micrillus</emphasis>
</taxonomicName>
(one of them undescribed).
</paragraph>
</subSubSection>
</treatment>
</document>