140 lines
15 KiB
XML
140 lines
15 KiB
XML
<document id="5AF9FFF1DC7E55AFEBF377D5C0526BBD" ID="10.11646/zootaxa.4117.4.1" ID-CLB-Dataset="38943" ID-DOI="10.11646/zootaxa.4117.4.1" ID-GBIF-Dataset="3ac9c002-f8fd-430b-8989-ff203416a264" ID-ISSN="1175-5326" ID-Zenodo-Dep="271659" ID-ZooBank="E52D10EF-B462-4F18-9C41-A7366170DFF7" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1464847700924" checkinUser="plazi" docAuthor="Last, Peter R., Séret, Bernard & Naylor, Gavin J. P." docDate="2016" docId="397087F0FFF40207FF69EC63FB49FB2F" docLanguage="en" docName="zootaxa.4117.4.1.pdf" docOrigin="Zootaxa 4117 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Rhinobatidae Bonaparte 1835" docType="treatment" docVersion="8" lastPageNumber="465" masterDocId="C549FF88FFFA0209FFFEED4BFFE2FFCE" masterDocTitle="A new species of guitarfish, Rhinobatos borneensis sp. nov. with a redefinition of the family-level classification in the order Rhinopristiformes (Chondrichthyes: Batoidea)" masterLastPageNumber="475" masterPageNumber="451" pageNumber="465" updateTime="1698662679303" updateUser="ExternalLinkService">
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<mods:mods id="914E946288C179C53AC26941C387FAF7" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="A9ADD134B2AB60E905D9F7FD9432AA7D">
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<mods:title id="3476C3825E7888617A24EE43F2A4B442">A new species of guitarfish, Rhinobatos borneensis sp. nov. with a redefinition of the family-level classification in the order Rhinopristiformes (Chondrichthyes: Batoidea)</mods:title>
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</mods:titleInfo>
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<mods:name id="EA183358441815B2817B0AE4C3F26675" type="personal">
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<mods:role id="D19246DF976ECC34FCD76A74E9125F0A">
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<mods:roleTerm id="F5E003247BCFA7D8421F29BD9E1E9EC0">Author</mods:roleTerm>
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<mods:namePart id="9172D34C96E2B5B7CD48D6649F99A566">Last, Peter R.</mods:namePart>
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</mods:name>
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<mods:name id="9226B966C475677B83EE211974D87185" type="personal">
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<mods:role id="7B624E35E1E88AB80191AAC747DBADAC">
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<mods:roleTerm id="F7403F94053E41B92216B485A43C8E0A">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="DAF78F583901D76D6750F0515005E109">Séret, Bernard</mods:namePart>
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</mods:name>
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<mods:name id="FDE36ED203740C63E598AED232929E16" type="personal">
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<mods:role id="05DE355594D2870C1ABB2CA8E6CACDDE">
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<mods:roleTerm id="36CD9A5A759265B471D6D001275AFF8D">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="66970E2EF2E68CE23DDCBD84BCB99EEA">Naylor, Gavin J. P.</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="71DBC64854F63110A3CF49CCF6256759">text</mods:typeOfResource>
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<mods:relatedItem id="F127D4A31F8A46F2450B72EEFFDEE026" type="host">
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<mods:titleInfo id="798E6BCC3B23F9BD790BBBDE9A48468E">
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<mods:title id="66BE4376E9BA03158D9779A2CEC58A78">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="0814EBC2D79BC4ACD345E0A440630AD9">
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<mods:date id="0D4DBF956E224FA233A8E69A9FC18F1E">2016</mods:date>
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<mods:detail id="3AB510A6CB471A3FADBD5BCB9AB49331" type="volume">
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<mods:number id="C0161A1F663700945C4A74B859955BE2">4117</mods:number>
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</mods:detail>
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<mods:detail id="08BA40F4767A5113994B5B5C3D4A71CC" type="issue">
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<mods:number id="7E302B58A85B123B68B845981A67110D">4</mods:number>
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<mods:start id="DBF7477C4623D26CC7D9B682EA0C6226">451</mods:start>
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<mods:end id="DFC3B2A74FDFD4099223A4DB9C5B2B67">475</mods:end>
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<mods:classification id="FC76CE72F5A0F9375AE89A83D4A65A70">journal article</mods:classification>
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<mods:identifier id="9BE38BBF41DB77F44B48C1AE9924EA57" type="CLB-Dataset">38943</mods:identifier>
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<mods:identifier id="F7C9576AB2CB9A20A25A6AAA2A997E29" type="DOI">10.11646/zootaxa.4117.4.1</mods:identifier>
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<mods:identifier id="2F9C671E0100196824419D6F9DF3F82F" type="GBIF-Dataset">3ac9c002-f8fd-430b-8989-ff203416a264</mods:identifier>
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<mods:identifier id="D369395EC5FD16DD27F43D65E2CF0194" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="22B14E5CF4E851AA47203D210E7AAAF7" type="Zenodo-Dep">271659</mods:identifier>
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<mods:identifier id="717671719C7C7C97DB14960A5C550EC2" type="ZooBank">E52D10EF-B462-4F18-9C41-A7366170DFF7</mods:identifier>
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</mods:mods>
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<treatment id="397087F0FFF40207FF69EC63FB49FB2F" ID-DOI="http://doi.org/10.5281/zenodo.6090711" ID-GBIF-Taxon="120735637" ID-Zenodo-Dep="6090711" LSID="urn:lsid:plazi:treatment:397087F0FFF40207FF69EC63FB49FB2F" httpUri="http://treatment.plazi.org/id/397087F0FFF40207FF69EC63FB49FB2F" lastPageNumber="465" pageId="14" pageNumber="465">
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<subSubSection id="F9C3656DFFF40207FF69EC63FD96FE8C" box="[151,628,296,322]" pageId="14" pageNumber="465" type="nomenclature">
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<paragraph id="B16636E6FFF40207FF69EC63FD96FE8C" blockId="14.[151,628,296,322]" box="[151,628,296,322]" pageId="14" pageNumber="465">
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<heading id="EA2E818AFFF40207FF69EC63FD96FE8C" bold="true" box="[151,628,296,322]" fontSize="11" level="1" pageId="14" pageNumber="465" reason="1">
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<emphasis id="83ADEAF4FFF40207FF69EC63FD96FE8C" bold="true" box="[151,628,296,322]" pageId="14" pageNumber="465">
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Family
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<taxonomicName id="76D94D65FFF40207FF08EC63FD96FE8C" authority="Bonaparte, 1835" authorityName="Bonaparte" authorityYear="1835" box="[246,628,296,322]" class="Elasmobranchii" family="Rhinobatidae" kingdom="Animalia" order="Rajiformes" pageId="14" pageNumber="465" phylum="Chordata" rank="family">Rhinobatidae Bonaparte, 1835</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="F9C3656DFFF40207FF69EC25FD0EFE4A" box="[151,748,366,388]" pageId="14" pageNumber="465" type="materials_examined">
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<paragraph id="B16636E6FFF40207FF69EC25FD0EFE4A" blockId="14.[151,748,366,388]" box="[151,748,366,388]" pageId="14" pageNumber="465">
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<emphasis id="83ADEAF4FFF40207FF69EC25FEF4FE4A" bold="true" box="[151,278,366,388]" pageId="14" pageNumber="465">
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<typeStatus id="6E628844FFF40207FF69EC25FF2EFE4A" box="[151,204,366,388]" pageId="14" pageNumber="465">Type</typeStatus>
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genus.
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</emphasis>
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<taxonomicName id="76D94D65FFF40207FEE2EC25FDF3FE4A" authority="Linck, 1790" authorityName="Linck" authorityYear="1790" box="[284,529,366,388]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhinobatos" kingdom="Animalia" order="Rajiformes" pageId="14" pageNumber="465" phylum="Chordata" rank="genus">
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<emphasis id="83ADEAF4FFF40207FEE2EC25FE72FE4A" box="[284,400,366,388]" italics="true" pageId="14" pageNumber="465">Rhinobatos</emphasis>
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Linck, 1790
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</taxonomicName>
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(inferred from stem).
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</paragraph>
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</subSubSection>
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<subSubSection id="F9C3656DFFF40207FF69ECFBFBDDFC53" pageId="14" pageNumber="465" type="description">
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<paragraph id="B16636E6FFF40207FF69ECFBFBDDFC53" blockId="14.[151,1437,432,1250]" pageId="14" pageNumber="465">
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<emphasis id="83ADEAF4FFF40207FF69ECFBFECBFE07" bold="true" box="[151,297,432,457]" pageId="14" pageNumber="465">Description.</emphasis>
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Small to large rays guitarfishes (adults attaining
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<quantity id="76219B03FFF40207FCB0ECFAFC6CFE07" box="[846,910,433,457]" metricMagnitude="0" metricUnit="m" metricValue="1.7" pageId="14" pageNumber="465" unit="m" value="1.7">1.7 m</quantity>
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TL) with a flattened, wedge- or shovel-shaped disc with a strongly depressed trunk. Snout often elongated and its tip narrowly pointed to broadly rounded. Eyes and spiracles medium to large; spiracles with 1–2 variably developed folds. Nostrils short and very oblique (with fewer than 62 lamellae), nasal curtain absent; anterior nasal flaps often very broad (almost joined mesially), inserted close to inner edge of nostril, or insertions extending slightly into internasal space (to level of nostril inner corner). Mouth profile straight to lower jaw weakly convex. Skin usually covered with fine denticles (sometimes partly naked); small thorns and thornlets variably developed in row along dorsal midline of body, in small patches near eyes, and on shoulder and snout. Pelvic fins long-based, positioned laterally and posteriorly to the disc. Tail not especially robust, depressed, with well-developed lateral skin folds extending along its ventrolateral margin; folds almost united on caudal-fin base near origin of ventral lobe of caudal fin. Two dorsal fins well separated, first positioned well to slightly behind rear tips of pelvic fins. Caudal fin small, lacking an obvious ventral lobe. Dorsal coloration plain (usually greyish or brownish) or with strong pattern of lines, bars, spots and/or blotches; edges of cranium and rostral cartilage not usually sharply demarcated with snout; ventral surface usually white, black blotch often present on snout. Three genera:
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<taxonomicName id="76D94D65FFF40207FDC0EECEFD07FC50" box="[574,741,901,926]" class="Elasmobranchii" family="Rhinobatidae" genus="Acroteriobatus" kingdom="Animalia" order="Rajiformes" pageId="14" pageNumber="465" phylum="Chordata" rank="genus">
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<emphasis id="83ADEAF4FFF40207FDC0EECEFD07FC50" box="[574,741,901,926]" italics="true" pageId="14" pageNumber="465">Acroteriobatus</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="76D94D65FFF40207FD0AEECEFC91FC50" box="[756,883,901,926]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhinobatos" kingdom="Animalia" order="Rajiformes" pageId="14" pageNumber="465" phylum="Chordata" rank="genus">
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<emphasis id="83ADEAF4FFF40207FD0AEECEFC91FC50" box="[756,883,901,926]" italics="true" pageId="14" pageNumber="465">Rhinobatos</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="76D94D65FFF40207FC57EECEFBDAFC50" box="[937,1080,901,926]" class="Elasmobranchii" family="Rhinobatidae" genus="Pseudobatos" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rajiformes" pageId="14" pageNumber="470" phylum="Chordata" rank="genus" status="gen. nov.">
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<emphasis id="83ADEAF4FFF40207FC57EECEFBDAFC50" box="[937,1080,901,926]" italics="true" pageId="14" pageNumber="465">Pseudobatos</emphasis>
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="F9C3656DFFF40207FF39EEE2FB49FB2F" pageId="14" pageNumber="465" type="discussion">
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<paragraph id="B16636E6FFF40207FF39EEE2FB49FB2F" blockId="14.[151,1437,432,1250]" pageId="14" pageNumber="465">
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<emphasis id="83ADEAF4FFF40207FF39EEE2FED9FC0F" bold="true" box="[199,315,937,961]" pageId="14" pageNumber="465">Remarks.</emphasis>
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<taxonomicName id="76D94D65FFF40207FEB9EEE2FE34FC0C" box="[327,470,937,962]" class="Elasmobranchii" family="Rhinobatidae" genus="Pseudobatos" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rajiformes" pageId="14" pageNumber="470" phylum="Chordata" rank="genus" status="gen. nov.">
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<emphasis id="83ADEAF4FFF40207FEB9EEE2FE34FC0C" box="[327,470,937,962]" italics="true" pageId="14" pageNumber="465">Pseudobatos</emphasis>
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</taxonomicName>
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undoubtedly represents a valid genus-level taxon but is not fully resolved as its familial assignment needs further investigation based on its apparent polyphyly with other genera based on molecular data. Relatively poorly developed nasal flaps in members of this amphi-American group is shared with
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<taxonomicName id="76D94D65FFF40207FF69E95EFE7BFBE0" box="[151,409,1045,1070]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhinobatos" kingdom="Animalia" order="Rajiformes" pageId="14" pageNumber="465" phylum="Chordata" rank="species" species="rhinobatos">
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<emphasis id="83ADEAF4FFF40207FF69E95EFE7BFBE0" box="[151,409,1045,1070]" italics="true" pageId="14" pageNumber="465">Rhinobatos rhinobatos</emphasis>
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</taxonomicName>
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, the
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<typeStatus id="6E628844FFF40207FE2BE95DFDE7FBE0" box="[469,517,1046,1070]" pageId="14" pageNumber="465">type</typeStatus>
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species of the genus (
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<collectingCountry id="C9CE7676FFF40207FCF9E95EFCD8FBE0" box="[775,826,1045,1070]" name="India" pageId="14" pageNumber="465">Indo</collectingCountry>
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–Pacific
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<taxonomicName id="76D94D65FFF40207FC5CE95EFBC0FBE0" box="[930,1058,1045,1070]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhinobatos" kingdom="Animalia" order="Rajiformes" pageId="14" pageNumber="465" phylum="Chordata" rank="genus">
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<emphasis id="83ADEAF4FFF40207FC5CE95EFBC0FBE0" box="[930,1058,1045,1070]" italics="true" pageId="14" pageNumber="465">Rhinobatos</emphasis>
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</taxonomicName>
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typically have larger flaps, most strongly expressed by the posterior nasal flap, which is broad and can extend almost to the innermost corner of the nostril).
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<taxonomicName id="76D94D65FFF40207FF08E916FE97FBB8" box="[246,373,1117,1142]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhinobatos" kingdom="Animalia" order="Rajiformes" pageId="14" pageNumber="465" phylum="Chordata" rank="genus">
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<emphasis id="83ADEAF4FFF40207FF08E916FE97FBB8" box="[246,373,1117,1142]" italics="true" pageId="14" pageNumber="465">Rhinobatos</emphasis>
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</taxonomicName>
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is sister to
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<taxonomicName id="76D94D65FFF40207FE06E916FD7DFBB8" box="[504,671,1117,1142]" class="Elasmobranchii" family="Rhinobatidae" genus="Acroteriobatus" kingdom="Animalia" order="Rajiformes" pageId="14" pageNumber="465" phylum="Chordata" rank="genus">
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<emphasis id="83ADEAF4FFF40207FE06E916FD7DFBB8" box="[504,671,1117,1142]" italics="true" pageId="14" pageNumber="465">Acroteriobatus</emphasis>
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</taxonomicName>
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but paraphyletic to the
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<collectingCountry id="C9CE7676FFF40207FC52E916FC3CFBB8" box="[940,990,1117,1142]" name="India" pageId="14" pageNumber="465">Indo</collectingCountry>
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–Pacific
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<emphasis id="83ADEAF4FFF40207FBBCE916FB2BFBB8" box="[1090,1225,1117,1142]" italics="true" pageId="14" pageNumber="465">
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<taxonomicName id="76D94D65FFF40207FBBCE916FB27FBB8" box="[1090,1221,1117,1142]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhinobatos" kingdom="Animalia" order="Rajiformes" pageId="14" pageNumber="465" phylum="Chordata" rank="genus">Rhinobatos</taxonomicName>
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.
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</emphasis>
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The anterior nasal flaps of
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<taxonomicName id="76D94D65FFF40207FF0FE9CAFE7AFB54" box="[241,408,1153,1178]" class="Elasmobranchii" family="Rhinobatidae" genus="Acroteriobatus" kingdom="Animalia" order="Rajiformes" pageId="14" pageNumber="465" phylum="Chordata" rank="genus">
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<emphasis id="83ADEAF4FFF40207FF0FE9CAFE7AFB54" box="[241,408,1153,1178]" italics="true" pageId="14" pageNumber="465">Acroteriobatus</emphasis>
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</taxonomicName>
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, which are almost joined in the medial region of the internasal space and well anterior of the innermost corners of the nostrils, are among the most well-developed of rhinopristiforms. The trygonorrhinid genus
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<taxonomicName id="76D94D65FFF40207FF21E982FE99FB2C" box="[223,379,1225,1250]" class="Elasmobranchii" family="Rhinobatidae" genus="Aptychotrema" kingdom="Animalia" order="Rajiformes" pageId="14" pageNumber="465" phylum="Chordata" rank="genus">
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<emphasis id="83ADEAF4FFF40207FF21E982FE99FB2C" box="[223,379,1225,1250]" italics="true" pageId="14" pageNumber="465">Aptychotrema</emphasis>
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</taxonomicName>
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has anterior nasal flaps inserted more posteriorly in the internasal space.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |