treatments-xml/data/39/70/87/397087F0FFEC021FFF69EF27FD69FA58.xml
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<document id="66CA5CA77118FF516D379D347E5B4F08" ID="10.11646/zootaxa.4117.4.1" ID-CLB-Dataset="38943" ID-DOI="10.11646/zootaxa.4117.4.1" ID-GBIF-Dataset="3ac9c002-f8fd-430b-8989-ff203416a264" ID-ISSN="1175-5326" ID-Zenodo-Dep="271659" ID-ZooBank="E52D10EF-B462-4F18-9C41-A7366170DFF7" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1464847700924" checkinUser="plazi" docAuthor="Last, Peter R., Séret, Bernard &amp; Naylor, Gavin J. P." docDate="2016" docId="397087F0FFEC021FFF69EF27FD69FA58" docLanguage="en" docName="zootaxa.4117.4.1.pdf" docOrigin="Zootaxa 4117 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pristidae Bonaparte 1835" docType="treatment" docVersion="8" lastPageNumber="473" masterDocId="C549FF88FFFA0209FFFEED4BFFE2FFCE" masterDocTitle="A new species of guitarfish, Rhinobatos borneensis sp. nov. with a redefinition of the family-level classification in the order Rhinopristiformes (Chondrichthyes: Batoidea)" masterLastPageNumber="475" masterPageNumber="451" pageNumber="473" updateTime="1698662679303" updateUser="ExternalLinkService">
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<mods:title id="9C0CA6309797084F58EDAB55093A52DA">A new species of guitarfish, Rhinobatos borneensis sp. nov. with a redefinition of the family-level classification in the order Rhinopristiformes (Chondrichthyes: Batoidea)</mods:title>
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<mods:namePart id="B02CB04B8FA0B8E53E83A941623A81AE">Last, Peter R.</mods:namePart>
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<mods:namePart id="BFA9604CDF01D7AACDD398DF006B41BD">Séret, Bernard</mods:namePart>
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<mods:namePart id="468F7B8307776DFCFE2645ED473FC112">Naylor, Gavin J. P.</mods:namePart>
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<mods:date id="CAAC24A9F480355DD73A1CF51E36FA1F">2016</mods:date>
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<treatment id="397087F0FFEC021FFF69EF27FD69FA58" ID-DOI="http://doi.org/10.5281/zenodo.6090737" ID-GBIF-Taxon="120735632" ID-Zenodo-Dep="6090737" LSID="urn:lsid:plazi:treatment:397087F0FFEC021FFF69EF27FD69FA58" httpUri="http://treatment.plazi.org/id/397087F0FFEC021FFF69EF27FD69FA58" lastPageNumber="473" pageId="22" pageNumber="473">
<subSubSection id="F9C3656DFFEC021FFF69EF27FDD9FD48" box="[151,571,620,646]" pageId="22" pageNumber="473" type="nomenclature">
<paragraph id="B16636E6FFEC021FFF69EF27FDD9FD48" blockId="22.[151,571,620,646]" box="[151,571,620,646]" pageId="22" pageNumber="473">
<heading id="EA2E818AFFEC021FFF69EF27FDD9FD48" bold="true" box="[151,571,620,646]" fontSize="11" level="1" pageId="22" pageNumber="473" reason="1">
<emphasis id="83ADEAF4FFEC021FFF69EF27FDD9FD48" bold="true" box="[151,571,620,646]" pageId="22" pageNumber="473">
Family
<taxonomicName id="76D94D65FFEC021FFF08EF27FDD9FD48" authority="Bonaparte, 1835" authorityName="Bonaparte" authorityYear="1835" box="[246,571,620,646]" class="Elasmobranchii" family="Pristidae" kingdom="Animalia" order="Pristiformes" pageId="22" pageNumber="473" phylum="Chordata" rank="family">Pristidae Bonaparte, 1835</taxonomicName>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="F9C3656DFFEC021FFF69EFF9FD5EFD06" box="[151,700,690,712]" pageId="22" pageNumber="473" type="materials_examined">
<paragraph id="B16636E6FFEC021FFF69EFF9FD5EFD06" blockId="22.[151,700,690,712]" box="[151,700,690,712]" pageId="22" pageNumber="473">
<emphasis id="83ADEAF4FFEC021FFF69EFF9FEF4FD06" bold="true" box="[151,278,690,712]" pageId="22" pageNumber="473">
<typeStatus id="6E628844FFEC021FFF69EFF9FF2EFD06" box="[151,204,690,712]" pageId="22" pageNumber="473">Type</typeStatus>
genus.
</emphasis>
<taxonomicName id="76D94D65FFEC021FFEE2EFF8FE02FD06" authority="Linck, 1790" authorityName="Linck" authorityYear="1790" box="[284,480,690,712]" class="Elasmobranchii" family="Pristidae" genus="Pristis" kingdom="Animalia" order="Pristiformes" pageId="22" pageNumber="473" phylum="Chordata" rank="genus">
<emphasis id="83ADEAF4FFEC021FFEE2EFF8FEBCFD06" box="[284,350,691,712]" italics="true" pageId="22" pageNumber="473">Pristis</emphasis>
Linck, 1790
</taxonomicName>
(inferred from stem).
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<subSubSection id="F9C3656DFFEC021FFF69EFBFFC75FB57" pageId="22" pageNumber="473" type="description">
<paragraph id="B16636E6FFEC021FFF69EFBFFC75FB57" blockId="22.[151,1437,756,1430]" pageId="22" pageNumber="473">
<emphasis id="83ADEAF4FFEC021FFF69EFBFFECBFCC3" bold="true" box="[151,297,756,781]" pageId="22" pageNumber="473">Description.</emphasis>
Large to very large rays (adults up to
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TL) with snout greatly extended to form a hard, flattened blade armed along each edge with a row of sharp, robust, hypertrophied tooth-like denticles. Pectoral fins relatively small, separate, not fused to head or forming an obvious disc. Head slightly flattened, projecting well forward of pectoral fins. Eyes positioned very close to sides of head; well removed from spiracles. Spiracles lacking folds. Mouth broad, lacking oral grooves. Nostrils variably developed, short to relatively elongate, well separated, located well anterior to mouth and posterior to toothed part of rostrum; mouth broad and transverse. Tail elongate, more or less subcylindrical anteriorly; lateral skin folds weak anteriorly but more expanded posteriorly to form a welldefined keel; keels extending well onto caudal fin (their posterior extremities not almost joined adjacent origin of ventral lobe of fin). Two dorsal fins similar in shape, rather tall, with first located fully or partly above pelvic fins; pectoral fins well separated from pelvic fins. Pelvic fins short-based, subtriangular. Caudal fin variably developed, its ventral lobe weak or short (
<taxonomicName id="76D94D65FFEC021FFE0FE915FDD8FBBB" box="[497,570,1118,1141]" class="Elasmobranchii" family="Pristidae" genus="Pristis" kingdom="Animalia" order="Pristiformes" pageId="22" pageNumber="473" phylum="Chordata" rank="genus">
<emphasis id="83ADEAF4FFEC021FFE0FE915FDD8FBBB" box="[497,570,1118,1141]" italics="true" pageId="22" pageNumber="473">Pristis</emphasis>
</taxonomicName>
), or present as a long lobe (
<taxonomicName id="76D94D65FFEC021FFC86E915FBE6FBBB" box="[888,1028,1118,1141]" class="Elasmobranchii" family="Pristidae" genus="Anoxypristis" kingdom="Animalia" order="Pristiformes" pageId="22" pageNumber="473" phylum="Chordata" rank="genus">
<emphasis id="83ADEAF4FFEC021FFC86E915FBE6FBBB" box="[888,1028,1118,1141]" italics="true" pageId="22" pageNumber="473">Anoxypristis</emphasis>
</taxonomicName>
). Dorsal coloration plain yellowish, brownish, greyish or greenish. Two genera:
<emphasis id="83ADEAF4FFEC021FFD7DE9C9FC72FB57" box="[643,912,1153,1178]" italics="true" pageId="22" pageNumber="473">
<taxonomicName id="76D94D65FFEC021FFD7DE9C9FCEDFB57" box="[643,783,1154,1177]" class="Elasmobranchii" family="Pristidae" genus="Anoxypristis" kingdom="Animalia" order="Pristiformes" pageId="22" pageNumber="473" phylum="Chordata" rank="genus">Anoxypristis</taxonomicName>
and
<taxonomicName id="76D94D65FFEC021FFCB9E9C9FC72FB57" box="[839,912,1154,1177]" class="Elasmobranchii" family="Pristidae" genus="Pristis" kingdom="Animalia" order="Pristiformes" pageId="22" pageNumber="473" phylum="Chordata" rank="genus">Pristis</taxonomicName>
</emphasis>
.
</paragraph>
</subSubSection>
<subSubSection id="F9C3656DFFEC021FFF39E9EEFD69FA58" pageId="22" pageNumber="473" type="discussion">
<paragraph id="B16636E6FFEC021FFF39E9EEFD69FA58" blockId="22.[151,1437,756,1430]" pageId="22" pageNumber="473">
<emphasis id="83ADEAF4FFEC021FFF39E9EEFED9FB73" bold="true" box="[199,315,1189,1213]" pageId="22" pageNumber="473">Remarks.</emphasis>
No other batoid has a heavily modified, blade-like snout armed with enlarged tooth-like denticles. The shark family
<taxonomicName id="76D94D65FFEC021FFE96E982FDF1FB2C" box="[360,531,1225,1250]" class="Elasmobranchii" family="Pristiophoridae" kingdom="Animalia" order="Pristiophoriformes" pageId="22" pageNumber="473" phylum="Chordata" rank="family">Pristiophoridae</taxonomicName>
also has a modified saw-like snout but otherwise possesses characteristics of sharks (e.g. gill slits lateral rather than ventral on head). In most recent morphology-based classifications, pristids have been placed alone in the order
<taxonomicName id="76D94D65FFEC021FFDBEE85AFD2BFAE4" box="[576,713,1297,1322]" class="Elasmobranchii" kingdom="Animalia" order="Pristiformes" pageId="22" pageNumber="473" phylum="Chordata" rank="order">Pristiformes</taxonomicName>
(e.g.
<bibRefCitation id="D5484B17FFEC021FFCF1E85AFBC0FAE4" author="McEachran" box="[783,1058,1297,1322]" pageId="22" pageNumber="473" refString="McEachran, J. D., Dunn, K. A. &amp; Miyake, T. (1996) Interrelationships of the batoid fishes (Chondrichthyes: Batoidea). In: Stiassny, M. J., Parenti, L. R. &amp; Johnson, G. D. (Eds.), Interrelationships of Fishes. Academic Press, London, pp. 63 - 82. http: // dx. doi. org / 10.1016 / b 978 - 012670950 - 6 / 50005 - 9" type="book chapter" year="1996">
McEachran
<emphasis id="83ADEAF4FFEC021FFC64E859FC37FAE4" box="[922,981,1297,1322]" italics="true" pageId="22" pageNumber="473">et al.</emphasis>
, 1996
</bibRefCitation>
; van der
<bibRefCitation id="D5484B17FFEC021FFB6EE859FA80FAE4" author="Laan" box="[1168,1378,1297,1322]" pageId="22" pageNumber="473" refString="Laan, R. van der, Eschmeyer, W. N. &amp; Fricke, R. (2014) Family-group names of recent fishes. Zootaxa, 3882 (1), 1 - 230. http: // dx. doi. org / 10.11646 / zootaxa. 3882.1.1" type="journal article" year="2014">
Laan,
<emphasis id="83ADEAF4FFEC021FFB27E859FAF6FAE4" box="[1241,1300,1297,1322]" italics="true" pageId="22" pageNumber="473">et al.</emphasis>
, 2014
</bibRefCitation>
), but recurrent recovery of the group as being monophyletic within the Rhinopristiformes based on molecular data is compelling evidence. Apart from their snout morphology, they share many key characters with other rhinopristiform families, notably the rhinids.
</paragraph>
</subSubSection>
</treatment>
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