303 lines
37 KiB
XML
303 lines
37 KiB
XML
<document id="F8F9A3862D4F63073B78D0D91B307A96" ID-CLB-Dataset="32590" ID-DOI="10.11646/zootaxa.4299.4.1" ID-GBIF-Dataset="e9fcac10-f2bf-4f24-b015-b8c0f18faa14" ID-ISSN="1175-5326" ID-Zenodo-Dep="836971" ID-ZooBank="DEB133FB-9D5A-4F17-A718-568ACA9BA9D7" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1501579068187" checkinUser="plazi" docAuthor="Sirenko, Boris I. & Saito, Hiroshi" docDate="2017" docId="2C2987BBDD49FFAFFF10354DFBFDD2A2" docLanguage="en" docName="zootaxa.4299.4.1.pdf" docOrigin="Zootaxa 4299 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Acanthochitona biformis Nierstrasz 1905" docType="treatment" docVersion="8" lastPageNumber="480" masterDocId="D010FFC3DD52FFB2FF87362AFFCBD52F" masterDocTitle="New species of chitons of the superfamily Cryptoplacoidea (Mollusca: Polyplacophora) from Vietnamese waters" masterLastPageNumber="506" masterPageNumber="451" pageNumber="478" updateTime="1698466057706" updateUser="ExternalLinkService">
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<mods:mods id="5E8DDF76D2FF6F06E5C8E7BB53E7DCF2" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="C6E3E82D189B19C5893B82A94083B0F4">
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<mods:title id="5449C75D49662569F3F9E1E3A4F0B863">New species of chitons of the superfamily Cryptoplacoidea (Mollusca: Polyplacophora) from Vietnamese waters</mods:title>
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<mods:name id="2425D41F3C6185BCA7BD90B67675A3E1" type="personal">
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<mods:namePart id="806D129CFE12F1EAE64C1FCF60626D49">Sirenko, Boris I.</mods:namePart>
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<mods:name id="25BE0EA9A963A884C898C963CACF771F" type="personal">
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<mods:namePart id="F8CEBB5F44C5969E73B149BB8ACEA80D">Saito, Hiroshi</mods:namePart>
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<mods:typeOfResource id="F7CE239E9E5B351346F4906F80D06620">text</mods:typeOfResource>
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<mods:title id="54D45117AAE9F6470A615A22A926906B">Zootaxa</mods:title>
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<mods:part id="11F7672F36363B79248C5F910E8618D2">
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<mods:date id="B15BAA93B77A59B96711F28A3811407E">2017</mods:date>
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<mods:number id="2ECF49E662962FD10E0D131E04B8FB41">4299</mods:number>
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<mods:identifier id="94FE502884F8C2A755CD566109CB74F4" type="DOI">10.11646/zootaxa.4299.4.1</mods:identifier>
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<treatment id="2C2987BBDD49FFAFFF10354DFBFDD2A2" ID-DOI="http://doi.org/10.5281/zenodo.5998847" ID-GBIF-Taxon="132617936" ID-Zenodo-Dep="5998847" LSID="urn:lsid:plazi:treatment:2C2987BBDD49FFAFFF10354DFBFDD2A2" httpUri="http://treatment.plazi.org/id/2C2987BBDD49FFAFFF10354DFBFDD2A2" lastPageId="29" lastPageNumber="480" pageId="27" pageNumber="478">
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<subSubSection id="EC9A6526DD49FFA9FF10354DFE9AD68C" pageId="27" pageNumber="478" type="nomenclature">
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<paragraph id="A43F36ADDD49FFA9FF10354DFD62D6AD" blockId="27.[151,681,871,931]" box="[151,681,871,898]" pageId="27" pageNumber="478">
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<heading id="FF7781C1DD49FFA9FF10354DFD62D6AD" bold="true" box="[151,681,871,898]" fontSize="11" level="1" pageId="27" pageNumber="478" reason="1">
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<taxonomicName id="63804D2EDD49FFA9FF10354DFD62D6AD" ID-CoL="64DSH" authority="Nierstrasz, 1905" authorityName="Nierstrasz" authorityYear="1905" box="[151,681,871,898]" class="Polyplacophora" family="Acanthochitonidae" genus="Acanthochitona" kingdom="Animalia" order="Chitonida" pageId="27" pageNumber="478" phylum="Mollusca" rank="species" species="biformis">
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<emphasis id="96F4EABFDD49FFA9FF10354DFD62D6AD" bold="true" box="[151,681,871,898]" pageId="27" pageNumber="478">
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<emphasis id="96F4EABFDD49FFA9FF10354DFE0ED6AE" bold="true" box="[151,453,871,897]" italics="true" pageId="27" pageNumber="478">Acanthochitona biformis</emphasis>
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||
(
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<bibRefCitation id="C0114B5CDD49FFA9FE533542FD6AD6AD" author="Nierstrasz" box="[468,673,872,898]" pageId="27" pageNumber="478" refString="Nierstrasz, H. F. (1905) Die Chitonen der Siboga-Expedition. - Siboga Expeditie, 48, 1 - 112, pls 1 - 8." type="journal article" year="1905">Nierstrasz, 1905</bibRefCitation>
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||
)
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||
</emphasis>
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||
</taxonomicName>
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||
</heading>
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||
</paragraph>
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||
<paragraph id="A43F36ADDD49FFA9FF1035A0FE9AD68C" blockId="27.[151,681,871,931]" box="[151,337,906,931]" pageId="27" pageNumber="478">
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<figureCitation id="3CBB2A28DD49FFA9FF1035A0FEDCD68C" box="[151,279,906,931]" captionStart-0="FIGURE 24" captionStart-1="FIGURE 25" captionStart-2="FIGURE 26" captionStartId-0="28.[151,249,1621,1643]" captionStartId-1="29.[151,249,1156,1178]" captionStartId-2="30.[151,249,1934,1956]" captionTargetBox-0="[202,1386,186,1579]" captionTargetBox-1="[220,1374,186,1127]" captionTargetBox-2="[496,1084,209,1859]" captionTargetId-0="figure@28.[151,1436,147,1636]" captionTargetId-1="figure@29.[174,1426,139,1157]" captionTargetId-2="figure@30.[481,1100,193,1907]" captionTargetPageId-0="28" captionTargetPageId-1="29" captionTargetPageId-2="30" captionText-0="FIGURE 24. Acanthochitona biformis, Southern Vietnam, Cau Id., BL 11 mm: A. Valve I, dorsal view; B. Valve V, dorsal view; C. Valve VIII, dorsal view; D. Valve IV, ventral view; E, F. Valve V, detail of tegmentum in lateropleural area; G. Valve V, rostral view: H. Valve VIII, lateral view." captionText-1="FIGURE 25. Acanthochitona biformis, Southern Vietnam, Cau Id., BL 11 mm: A. Valve V, detail of tegmentum in jugal area; B, C. Tufts, dorsal needles and spicules and ventral spicules; D. Radula." captionText-2="FIGURE 26. Acanthochitona biformis, Southern Vietnam, Cau Id., BL 11 mm: A. Needles of tufts; B. Dorsal needles and spicules; C. Marginal needle; D. Ventral spicules; E. Head of major lateral tooth of radula; F. Central and first lateral teeth of radula; G. Pustule of lateropleural areas of tegmentum with aesthete pores. Scale bars: 100 µm." httpUri-0="https://zenodo.org/record/837017/files/figure.png" httpUri-1="https://zenodo.org/record/837019/files/figure.png" httpUri-2="https://zenodo.org/record/837021/files/figure.png" pageId="27" pageNumber="478">Figs 24–26</figureCitation>
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,
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<figureCitation id="3CBB2A28DD49FFA9FEA535A1FEF4D68C" box="[290,319,907,931]" captionStart="FIGURE 44" captionStartId="50.[151,249,1837,1859]" captionTargetBox="[168,1412,193,1804]" captionTargetId="figure@50.[168,1412,193,1804]" captionTargetPageId="50" captionText="FIGURE 44. Images of species of genus Acanthochitona and Cryptoplax: A. A. achates, near Da Nang, intertidal, on barnacles, BL 15 mm; B. A. biformis, Southern Vietnam, Cau Id., 13 – 17 m, on old shells, BL 11 mm; C. A condaoi n. sp., Gulf of Thailand, Vang Id., 9 – 11 m, on old shells, paratype, BL 6 mm; D. A. lanae n. sp., near Da Nang, intertidal, on barnacles, paratype, BL 12 mm; E. A. ostreaphila n. sp., Nha Trang Bay, Tre Island, Dambay Inlet, intertidal, mangrove community, on oysters, paratype, BL 12 mm; F. A. ostreaphila n. sp., Gulf of Tongking, Cat Ba Island, 0.5 m, on oysters, paratype, BL 5.5 mm; G. Cryptoplax burrowi, South China Sea, Macclesfield Bank, 21 m, BL 29 mm." httpUri="https://zenodo.org/record/837057/files/figure.png" pageId="27" pageNumber="478">44</figureCitation>
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B
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="EC9A6526DD49FFA9FF1035F8FBE1D128" pageId="27" pageNumber="478" type="reference_group">
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||
<paragraph id="A43F36ADDD49FFA9FF1035F8FC57D6C7" blockId="27.[151,1066,978,1031]" box="[151,924,978,1000]" pageId="27" pageNumber="478">
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||
<treatmentCitationGroup id="84901183DD49FFA9FF1035F8FC57D6C7" box="[151,924,978,1000]" pageId="27" pageNumber="478">
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||
<taxonomicName id="63804D2EDD49FFA9FF1035F8FD9DD6C7" ID-CoL="64CDN" authority="Nierstrasz, 1905: 55" authorityName="Nierstrasz" authorityPageNumber="55" authorityYear="1905" box="[151,598,978,1000]" class="Polyplacophora" family="Acanthochitonidae" genus="Acanthochites" higherTaxonomySource="CoL" kingdom="Animalia" order="Chitonida" pageId="27" pageNumber="478" phylum="Mollusca" rank="species" species="biformis">
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||
<emphasis id="96F4EABFDD49FFA9FF1035F8FE4AD6C7" box="[151,385,978,1000]" italics="true" pageId="27" pageNumber="478">Acanthochites biformis</emphasis>
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||
<treatmentCitation id="252110BCDD49FFA9FE0035F8FD9DD6C7" author="Nierstrasz" box="[391,598,978,1000]" page="55" pageId="27" pageNumber="478" year="1905">
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<bibRefCitation id="C0114B5CDD49FFA9FE0035F8FDFAD6C7" author="Nierstrasz" box="[391,561,978,1000]" pageId="27" pageNumber="478" refString="Nierstrasz, H. F. (1905) Die Chitonen der Siboga-Expedition. - Siboga Expeditie, 48, 1 - 112, pls 1 - 8." type="journal article" year="1905">Nierstrasz, 1905</bibRefCitation>
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||
: 55
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||
</treatmentCitation>
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||
</taxonomicName>
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||
, pl. 1, fig.1, pl. 4, figs 104–108.
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||
</treatmentCitationGroup>
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||
</paragraph>
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||
<paragraph id="A43F36ADDD49FFA9FF1035DBFBE1D128" blockId="27.[151,1066,978,1031]" box="[151,1066,1009,1031]" pageId="27" pageNumber="478">
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||
<treatmentCitationGroup id="84901183DD49FFA9FF1035DBFBE1D128" box="[151,1066,1009,1031]" pageId="27" pageNumber="478">
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||
<taxonomicName id="63804D2EDD49FFA9FF1035DBFCEFD128" ID-CoL="64DSH" authority="Kaas & Van Belle, 1980: 16" authorityName="Kaas & Van Belle" authorityPageNumber="16" authorityYear="1980" box="[151,804,1009,1031]" class="Polyplacophora" family="Acanthochitonidae" genus="Acanthochitona" kingdom="Animalia" order="Chitonida" pageId="27" pageNumber="478" phylum="Mollusca" rank="species" species="biformis">
|
||
<emphasis id="96F4EABFDD49FFA9FF1035DBFE5AD128" box="[151,401,1009,1031]" italics="true" pageId="27" pageNumber="478">Acanthochitona biformis</emphasis>
|
||
:
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<treatmentCitation id="252110BCDD49FFA9FE1835D8FD76D128" author="Kaas" box="[415,701,1009,1031]" page="16" pageId="27" pageNumber="478" year="1980">
|
||
<bibRefCitation id="C0114B5CDD49FFA9FE1835D8FD53D128" author="Kaas" box="[415,664,1009,1031]" pageId="27" pageNumber="478" refString="Kaas, P. & Van Belle, R. A. (1980) Catalogue of living chitons. Dr. W. Backhuys, Publisher, Rotterdam, vi + 144 pp." type="book" year="1980">Kaas & Van Belle, 1980</bibRefCitation>
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||
: 16
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||
</treatmentCitation>
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||
, 1998: 32
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||
</taxonomicName>
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||
;
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||
<treatmentCitation id="252110BCDD49FFA9FCA935DBFC2FD128" author="Saito" box="[814,996,1009,1031]" page="125" pageId="27" pageNumber="478" year="2006">
|
||
<bibRefCitation id="C0114B5CDD49FFA9FCA935DBFC79D128" author="Saito" box="[814,946,1009,1031]" pageId="27" pageNumber="478" refString="Saito, H. (2006 b) Shallow water chitons (Mollusca: Polyplacophora) from Puerto Galera, Mindoro Island, the Philippines. Memoirs of the National Science Museum, 44, 119 - 133." type="journal article" year="2006" yearSuffix="b">Saito, 2006b</bibRefCitation>
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||
: 125
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</treatmentCitation>
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, fig. 3.
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||
</treatmentCitationGroup>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="EC9A6526DD49FFA9FF10321EFD33D1F2" pageId="27" pageNumber="478" type="materials_examined">
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||
<paragraph id="A43F36ADDD49FFA9FF10321EFD55D162" blockId="27.[151,1436,1076,1893]" box="[151,670,1076,1101]" pageId="27" pageNumber="478">
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||
<emphasis id="96F4EABFDD49FFA9FF10321EFE8ED162" bold="true" box="[151,325,1076,1101]" pageId="27" pageNumber="478">Type material.</emphasis>
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Syntypes, ZMA Moll 138563.
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</paragraph>
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<paragraph id="A43F36ADDD49FFA9FF40327DFD7CD1BA" blockId="27.[151,1436,1076,1893]" pageId="27" pageNumber="478">
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<materialsCitation id="14E83CF0DD49FFA9FF40327DFD7CD1BA" ID-GBIF-Occurrence="1585189399" country="Indonesia" county="Banda Island" location="Not" municipality="Sanguisiapo Island" pageId="27" pageNumber="478" specimenCount="1" typeStatus="holotype">
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||
<emphasis id="96F4EABFDD49FFA9FF40327DFEADD15F" bold="true" box="[199,358,1111,1136]" pageId="27" pageNumber="478">
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||
<typeStatus id="7B3B880FDD49FFA9FF40327DFEC9D15F" box="[199,258,1111,1136]" pageId="27" pageNumber="478">Type</typeStatus>
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||
locality.
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||
</emphasis>
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||
<location id="A15F6076DD49FFA9FEEA3272FE5CD140" LSID="urn:lsid:plazi:treatment:2C2987BBDD49FFAFFF10354DFBFDD2A2:A15F6076DD49FFA9FEEA3272FE5CD140" box="[365,407,1112,1135]" country="Indonesia" county="Banda Island" municipality="Sanguisiapo Island" name="Not" pageId="27" pageNumber="478">Not</location>
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designated, original lots were mentioned from near
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||
<location id="A15F6076DD49FFA9FC603272FBF9D140" LSID="urn:lsid:plazi:treatment:2C2987BBDD49FFAFFF10354DFBFDD2A2:A15F6076DD49FFA9FC603272FBF9D140" box="[999,1074,1112,1135]" country="Indonesia" county="Banda Island" municipality="Sanguisiapo Island" name="Kupan" pageId="27" pageNumber="478">Kupan</location>
|
||
and
|
||
<location id="A15F6076DD49FFA9FBED3272FA53D15F" LSID="urn:lsid:plazi:treatment:2C2987BBDD49FFAFFF10354DFBFDD2A2:A15F6076DD49FFA9FBED3272FA53D15F" box="[1130,1432,1111,1136]" country="Indonesia" county="Banda Island" municipality="Sanguisiapo Island" name="Nusa Besi of Timor Island" pageId="27" pageNumber="478">Nusa Besi of Timor Island</location>
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||
,
|
||
<collectingMunicipality id="445BACD7DD49FFA9FF103256FEA0D1BA" box="[151,363,1148,1173]" pageId="27" pageNumber="478">Sanguisiapo Island</collectingMunicipality>
|
||
and
|
||
<collectingCounty id="4D5E4E21DD49FFA9FE253256FDFCD1BA" box="[418,567,1148,1173]" pageId="27" pageNumber="478">Banda Island</collectingCounty>
|
||
,
|
||
<collectingCountry id="DC97763DDD49FFA9FDC43256FD7FD1BA" box="[579,692,1148,1173]" name="Indonesia" pageId="27" pageNumber="478">Indonesia</collectingCountry>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="A43F36ADDD49FFA9FF4032B5FD33D1F2" blockId="27.[151,1436,1076,1893]" pageId="27" pageNumber="478">
|
||
<materialsCitation id="14E83CF0DD49FFA9FF4032B5FD33D1F2" ID-GBIF-Occurrence="1585189406" collectingDate="2013-12-05" collectionCode="SCUBA" collectorName="Cau Id. & B. Sirenko." country="Vietnam" elevation="15" latitude="11.22045" location="Material" longLatPrecision="1" longitude="108.829765" pageId="27" pageNumber="478" specimenCount="1">
|
||
<emphasis id="96F4EABFDD49FFA9FF4032B5FE7DD197" bold="true" box="[199,438,1183,1208]" pageId="27" pageNumber="478">
|
||
<location id="A15F6076DD49FFA9FF4032B5FEE4D197" LSID="urn:lsid:plazi:treatment:2C2987BBDD49FFAFFF10354DFBFDD2A2:A15F6076DD49FFA9FF4032B5FEE4D197" box="[199,303,1183,1208]" country="Vietnam" latitude="11.22045" longLatPrecision="1" longitude="108.829765" name="Material" pageId="27" pageNumber="478">Material</location>
|
||
examined.
|
||
</emphasis>
|
||
Southern
|
||
<collectingCountry id="DC97763DDD49FFA9FDB432B5FD5ED197" box="[563,661,1183,1208]" name="Vietnam" pageId="27" pageNumber="478">Vietnam</collectingCountry>
|
||
,
|
||
<collectorName id="0975537BDD49FFA9FD22328AFD34D197" box="[677,767,1183,1208]" pageId="27" pageNumber="478">Cau Id.</collectorName>
|
||
,
|
||
<geoCoordinate id="C1B4506ADD49FFA9FC97328AFC68D197" box="[784,931,1184,1208]" direction="north" orientation="latitude" pageId="27" pageNumber="478" precision="1" value="11.22045">11°13.227´N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="C1B4506ADD49FFA9FC3332B5FB98D197" box="[948,1107,1183,1208]" direction="east" orientation="longitude" pageId="27" pageNumber="478" precision="1" value="108.829765">108°49.786´E</geoCoordinate>
|
||
,
|
||
<elevation id="2FADD19EDD49FFA9FBE3328AFB05D198" box="[1124,1230,1184,1208]" metricMagnitude="1" metricUnit="m" metricValue="1.5" metricValueMax="1.7" metricValueMin="1.3" pageId="27" pageNumber="478" unit="m" value="15.0" valueMax="17.0" valueMin="13.0">
|
||
<quantity id="63789B48DD49FFA9FBE3328AFB05D198" box="[1124,1230,1184,1208]" metricMagnitude="1" metricUnit="m" metricValue="1.5" metricValueMax="1.7" metricValueMin="1.3" pageId="27" pageNumber="478" unit="m" value="15.0" valueMax="17.0" valueMin="13.0">13–17 m</quantity>
|
||
</elevation>
|
||
,
|
||
<collectionCode id="C291AE68DD49FFA9FB58328AFAF4D197" box="[1247,1343,1184,1208]" pageId="27" pageNumber="478">SCUBA</collectionCode>
|
||
, on old shells, 1 spm, BL
|
||
<quantity id="63789B48DD49FFA9FEE632EEFE7AD1F3" box="[353,433,1220,1244]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" pageId="27" pageNumber="478" unit="mm" value="11.0">11 mm</quantity>
|
||
,
|
||
<collectingDate id="C07AE985DD49FFA9FE3A32EEFDF6D1F2" box="[445,573,1220,1245]" pageId="27" pageNumber="478" value="2013-12-05">
|
||
<date id="D03E106DDD49FFA9FE3A32EEFDF6D1F2" box="[445,573,1220,1245]" pageId="27" pageNumber="478" value="2013-12-05">12.05.2013</date>
|
||
</collectingDate>
|
||
, leg.
|
||
<collectorName id="0975537BDD49FFA9FDFE32EFFD33D1F2" box="[633,760,1220,1245]" pageId="27" pageNumber="478">B. Sirenko.</collectorName>
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="EC9A6526DD49FFAEFF4032CDFDEAD2B8" lastPageId="28" lastPageNumber="479" pageId="27" pageNumber="478" type="description">
|
||
<paragraph id="A43F36ADDD49FFA9FF4032CDFD35D00A" blockId="27.[151,1436,1076,1893]" pageId="27" pageNumber="478">
|
||
<emphasis id="96F4EABFDD49FFA9FF4032CDFE93D02F" bold="true" box="[199,344,1255,1280]" pageId="27" pageNumber="478">Description.</emphasis>
|
||
Animal small, oval, with small, moderately elevated, round backed valves. Color of tegmentum white with brownish flecks; jugum with yellowish tint.
|
||
</paragraph>
|
||
<paragraph id="A43F36ADDD49FFA9FF403305FCB9D09A" blockId="27.[151,1436,1076,1893]" pageId="27" pageNumber="478">Head valve semicircular with five obsolete radial undulations; posterior margin almost straight; anterior slope convex. Intermediate valves roughly oval to fan-shaped in outline, beaked, round-backed; lateral areas hardly separable; side slopes slightly convex; jugum narrow wedge-shape, smooth. Tail valve small, with central mucro; width of tegmentum slightly wider than half width of head valve.</paragraph>
|
||
<paragraph id="A43F36ADDD49FFA9FF4033EAFD9AD36A" blockId="27.[151,1436,1076,1893]" pageId="27" pageNumber="478">Tegmentum sculptured with droplet-shaped or oval, weakly concave pustules arranged in quincunx order; prepustular slope very narrow, sunken in slit. Each pustule with one macraesthete pore or occasionally two pores near center. No aesthete pore on tegmental plain. Jugum with fine growth lines, but practically smooth; micraesthete pores sparsely distributed.</paragraph>
|
||
<paragraph id="A43F36ADDD49FFA9FF403065FD46D39F" blockId="27.[151,1436,1076,1893]" pageId="27" pageNumber="478">Apophyses rounded, widely apart from each other in intermediate valves, truncated in tail valve. Insertion plate rather short, smooth on surface except for tail valve, which has rough surface with eight minute denticles at the edge in studied specimen. Slit formula 5/1/2.</paragraph>
|
||
<paragraph id="A43F36ADDD49FFA9FF403096FCF4D24A" blockId="27.[151,1436,1076,1893]" pageId="27" pageNumber="478">
|
||
Girdle rather wide, about
|
||
<quantity id="63789B48DD49FFA9FE733096FD86D3FB" box="[500,589,1724,1748]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="27" pageNumber="478" unit="mm" value="2.8">2.8 mm</quantity>
|
||
near valve V (width
|
||
<quantity id="63789B48DD49FFA9FCCD3096FC6ED3FB" box="[842,933,1724,1748]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="27" pageNumber="478" unit="mm" value="3.3">3.3 mm</quantity>
|
||
) in studied specimen. Dorsal side of girdle densely covered with minute, pointed spicules, 110–125 µm
|
||
<date id="D03E106DDD49FFA9FCB830CBFC51D3D7" box="[831,922,1759,1784]" pageId="27" pageNumber="478" value="2010-10-16">x 10–16</date>
|
||
µm, intermingling with thin needles attaining 218 µm
|
||
<date id="D03E106DDD49FFA9FF72312FFEDCD233" box="[245,279,1796,1820]" pageId="27" pageNumber="478">x 9</date>
|
||
µm. Sutural tufts prominent, consisting of up to 40 long, slightly curved, smooth or occasionally finely ribbed needles, 770–1200 µm
|
||
<date id="D03E106DDD49FFA9FE773103FDE9D26F" box="[496,546,1832,1856]" pageId="27" pageNumber="478" value="1940-10">x 40</date>
|
||
–42 µm. Marginal needles long, slightly curved, finely ribbed, up to 413 µm
|
||
<date id="D03E106DDD49FFA9FA093103FF78D24B" pageId="27" pageNumber="478" value="1927-10">x 27</date>
|
||
µm. Ventral spicules minute, flat, 66–80 µm
|
||
<date id="D03E106DDD49FFA9FD373167FCC1D24A" box="[688,778,1868,1893]" pageId="27" pageNumber="478" value="2014-10-15">x 14–15</date>
|
||
µm.
|
||
</paragraph>
|
||
<caption id="F0FF6625DD4EFFAEFF10307FFDFBD387" httpUri="https://zenodo.org/record/837017/files/figure.png" pageId="28" pageNumber="479" targetBox="[202,1386,186,1579]" targetPageId="28">
|
||
<paragraph id="A43F36ADDD4EFFAEFF10307FFDFBD387" blockId="28.[151,1436,1621,1704]" pageId="28" pageNumber="479">
|
||
<emphasis id="96F4EABFDD4EFFAEFF10307FFEEAD344" bold="true" box="[151,289,1621,1643]" pageId="28" pageNumber="479">FIGURE 24.</emphasis>
|
||
<taxonomicName id="63804D2EDD4EFFAEFEAD307FFDECD344" authorityName="Kaas & Van Belle" authorityYear="1980" box="[298,551,1621,1643]" class="Polyplacophora" family="Acanthochitonidae" genus="Acanthochitona" kingdom="Animalia" order="Chitonida" pageId="28" pageNumber="479" phylum="Mollusca" rank="species" species="biformis">
|
||
<emphasis id="96F4EABFDD4EFFAEFEAD307FFDECD344" box="[298,551,1621,1643]" italics="true" pageId="28" pageNumber="479">Acanthochitona biformis</emphasis>
|
||
</taxonomicName>
|
||
, Southern Vietnam, Cau Id., BL 11 mm: A. Valve I, dorsal view; B. Valve V, dorsal view; C. Valve VIII, dorsal view; D. Valve IV, ventral view; E, F. Valve V, detail of tegmentum in lateropleural area; G. Valve V, rostral view: H. Valve VIII, lateral view.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="A43F36ADDD4EFFAEFF4930C4FDEAD2B8" blockId="28.[151,1437,1774,2015]" pageId="28" pageNumber="479">
|
||
Gills extending from valve VII to valve V, composed of 10 ctenidia on each side in studied specimen. Radula of studied specimen
|
||
<quantity id="63789B48DD4EFFAEFD923138FDA6D205" box="[533,621,1810,1835]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="28" pageNumber="479" unit="mm" value="3.5">3.5 mm</quantity>
|
||
with 25 rows of mature teeth. Central tooth small, spatulate in outline, thin, sharply keeled in basal portion. First lateral (centro-lateral) tooth with thick, nodulous antero-dorsal corner. Major lateral tooth with tricuspid head; denticles blunt at tip; central one longer than others. Major uncinal tooth with long blade of moderate width.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="EC9A6526DD4EFFAEFF403188FDD5D2F1" pageId="28" pageNumber="479" type="distribution">
|
||
<paragraph id="A43F36ADDD4EFFAEFF403188FDD5D2F1" blockId="28.[151,1437,1774,2015]" pageId="28" pageNumber="479">
|
||
<emphasis id="96F4EABFDD4EFFAEFF403188FEABD294" bold="true" box="[199,352,1954,1979]" pageId="28" pageNumber="479">Distribution.</emphasis>
|
||
Timor and Banda Islands,
|
||
<collectingCountry id="DC97763DDD4EFFAEFD183188FCC4D294" box="[671,783,1954,1979]" name="Indonesia" pageId="28" pageNumber="479">Indonesia</collectingCountry>
|
||
; southern
|
||
<collectingRegion id="6644F84FDD4EFFAEFC0E3188FC76D294" box="[905,957,1954,1979]" country="Philippines" name="Sulu" pageId="28" pageNumber="479">Sulu</collectingRegion>
|
||
Islands and Mindoro Island,
|
||
<collectingCountry id="DC97763DDD4EFFAEFA913188FA53D294" box="[1302,1432,1954,1979]" name="Philippines" pageId="28" pageNumber="479">Philippines</collectingCountry>
|
||
; southern
|
||
<collectingCountry id="DC97763DDD4EFFAEFF7931ECFEABD2F0" box="[254,352,1990,2015]" name="Vietnam" pageId="28" pageNumber="479">Vietnam</collectingCountry>
|
||
, intertidal–
|
||
<specimenCount id="B286FD24DD4EFFAEFE6631ECFDD5D2F1" box="[481,542,1990,2015]" pageId="28" pageNumber="479" type="generic">
|
||
<quantity id="63789B48DD4EFFAEFE6631ECFDD1D2F1" box="[481,538,1990,2015]" metricMagnitude="1" metricUnit="m" metricValue="1.2" pageId="28" pageNumber="479" unit="m" value="12.0">12 m</quantity>
|
||
.
|
||
</specimenCount>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="F0FF6625DD4FFFAFFF1032AEFCBAD193" httpUri="https://zenodo.org/record/837019/files/figure.png" pageId="29" pageNumber="480" targetBox="[220,1374,186,1127]" targetPageId="29">
|
||
<paragraph id="A43F36ADDD4FFFAFFF1032AEFCBAD193" blockId="29.[151,1436,1156,1212]" pageId="29" pageNumber="480">
|
||
<emphasis id="96F4EABFDD4FFFAFFF1032AEFED4D1B5" bold="true" box="[151,287,1156,1178]" pageId="29" pageNumber="480">FIGURE 25.</emphasis>
|
||
<taxonomicName id="63804D2EDD4FFFAFFEA132AEFDEAD1B5" authorityName="Kaas & Van Belle" authorityYear="1980" box="[294,545,1156,1178]" class="Polyplacophora" family="Acanthochitonidae" genus="Acanthochitona" kingdom="Animalia" order="Chitonida" pageId="29" pageNumber="480" phylum="Mollusca" rank="species" species="biformis">
|
||
<emphasis id="96F4EABFDD4FFFAFFEA132AEFDEAD1B5" box="[294,545,1156,1178]" italics="true" pageId="29" pageNumber="480">Acanthochitona biformis</emphasis>
|
||
</taxonomicName>
|
||
, Southern Vietnam, Cau Id., BL 11 mm: A. Valve V, detail of tegmentum in jugal area; B, C. Tufts, dorsal needles and spicules and ventral spicules; D. Radula.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="EC9A6526DD4FFFAFFF4032C6FD81D341" pageId="29" pageNumber="480" type="discussion">
|
||
<paragraph id="A43F36ADDD4FFFAFFF4032C6FD81D341" blockId="29.[151,1437,1260,1934]" pageId="29" pageNumber="480">
|
||
<emphasis id="96F4EABFDD4FFFAFFF4032C6FEF0D02A" bold="true" box="[199,315,1260,1285]" pageId="29" pageNumber="480">Remarks.</emphasis>
|
||
<bibRefCitation id="C0114B5CDD4FFFAFFEC232C6FDC7D02A" author="Nierstrasz" box="[325,524,1260,1285]" pageId="29" pageNumber="480" refString="Nierstrasz, H. F. (1905) Die Chitonen der Siboga-Expedition. - Siboga Expeditie, 48, 1 - 112, pls 1 - 8." type="journal article" year="1905">Nierstrasz (1905)</bibRefCitation>
|
||
remarked that there are two variations within the present species, which have almost same morphology in the valves and same color (tint) in the sutural tufts, but differences in the color and the width of the valves. One of the two color variations referred by Nierstrasz has white valves with a yellowish jugum and dark reddish flecks as has been reported by
|
||
<bibRefCitation id="C0114B5CDD4FFFAFFD2D3373FC88D05D" author="Saito" box="[682,835,1369,1394]" pageId="29" pageNumber="480" refString="Saito, H. (2006 b) Shallow water chitons (Mollusca: Polyplacophora) from Puerto Galera, Mindoro Island, the Philippines. Memoirs of the National Science Museum, 44, 119 - 133." type="journal article" year="2006" yearSuffix="b">Saito (2006b)</bibRefCitation>
|
||
in a specimen from Mindoro Island,
|
||
<collectingCountry id="DC97763DDD4FFFAFFB583373FAAAD05D" box="[1247,1377,1369,1394]" name="Philippines" pageId="29" pageNumber="480">Philippines</collectingCountry>
|
||
. This specimen has a little wider valves than the present material but other features match well in both specimens. Another variation reported by Nierstrasz has uniformly orange-yellow valves which are slightly narrower than those of the white variation. Examination of the orange-yellow specimen among the
|
||
<typeStatus id="7B3B880FDD4FFFAFFBE733ECFB09D0F1" box="[1120,1218,1478,1502]" pageId="29" pageNumber="480" type="syntype">syntypes</typeStatus>
|
||
(from Sanguisiapo Island, Siboga station 93) confirmed the same morphology among those variations. As similar differences are known also from some
|
||
<taxonomicName id="63804D2EDD4FFFAFFE2E3027FD91D309" authorityName="Gray" authorityYear="1821" box="[425,602,1549,1574]" class="Polyplacophora" family="Acanthochitonidae" genus="Acanthochitona" kingdom="Animalia" order="Chitonida" pageId="29" pageNumber="480" phylum="Mollusca" rank="genus">
|
||
<emphasis id="96F4EABFDD4FFFAFFE2E3027FD91D309" box="[425,602,1549,1574]" italics="true" pageId="29" pageNumber="480">Acanthochitona</emphasis>
|
||
</taxonomicName>
|
||
species, such as
|
||
<taxonomicName id="63804D2EDD4FFFAFFCA33027FB2DD309" authority="(Pennant, 1777) (Kaas 1985)" authorityName="Pennant" authorityYear="1777" baseAuthorityName="Pennant" baseAuthorityYear="1777" box="[804,1254,1549,1574]" class="Polyplacophora" family="Acanthochitonidae" genus="Acanthochitona" kingdom="Animalia" order="Chitonida" pageId="29" pageNumber="480" phylum="Mollusca" rank="species" species="crinita">
|
||
<emphasis id="96F4EABFDD4FFFAFFCA33027FC5AD30A" box="[804,913,1549,1573]" italics="true" pageId="29" pageNumber="480">A. crinita</emphasis>
|
||
(Pennant, 1777) (
|
||
<bibRefCitation id="C0114B5CDD4FFFAFFBE53027FB15D309" author="Kaas" box="[1122,1246,1549,1574]" pageId="29" pageNumber="480" refString="Kaas, P. (1985) The genus Acanthochitona Gray, 1821 (Mollusca, Polyplacophora) in the north-eastern Atlantic Ocean and in the Mediterranean Sea, designation of neotype of A. fascicularis (L., 1767) and of A. crinita (Pennant, 1777). Bulletin de Museum national d'Histoire Naturelle, Paris, 4 e Serie, 7, Section A, 3, 579 - 609." type="journal article" year="1985">Kaas 1985</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="63804D2EDD4FFFAFFB703027FE93D365" authority="Linnaeus, 1767" authorityName="Linnaeus" authorityYear="1767" class="Polyplacophora" family="Chitonidae" genus="Chiton" kingdom="Animalia" order="Chitonida" pageId="29" pageNumber="480" phylum="Mollusca" rank="species" species="fascicularis">
|
||
<emphasis id="96F4EABFDD4FFFAFFB703027FA57D309" box="[1271,1436,1549,1574]" italics="true" pageId="29" pageNumber="480">A. fascicularis</emphasis>
|
||
(Linnaeus, 1767)
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="C0114B5CDD4FFFAFFEEF301BFD70D366" author="Leloup" box="[360,699,1585,1610]" pageId="29" pageNumber="480" refString="Leloup, E. (1941) A propos de quelques Acanthochitons peu connus ou nouveaux II. - Region Atlantique. Bulletin du Musee royal d'Histoire naturelle de Belgique, 17 (43), 1 - 16, pl. 1." type="journal article" year="1941">
|
||
Leloup 1941: as
|
||
<taxonomicName id="63804D2EDD4FFFAFFDAD3018FD70D366" authorityName="Risso" authorityYear="1826" box="[554,699,1586,1609]" class="Polyplacophora" family="Acanthochitonidae" genus="Acanthochitona" kingdom="Animalia" order="Chitonida" pageId="29" pageNumber="480" phylum="Mollusca" rank="species" species="communis">
|
||
<emphasis id="96F4EABFDD4FFFAFFDAD3018FD70D366" box="[554,699,1586,1609]" italics="true" pageId="29" pageNumber="480">A. communis</emphasis>
|
||
</taxonomicName>
|
||
</bibRefCitation>
|
||
) and
|
||
<taxonomicName id="63804D2EDD4FFFAFFD783018FAF3D366" authority="(Sowerby, 1832) (Watters 1990)" authorityName="Sowerby I" authorityYear="1832" baseAuthorityName="Sowerby" baseAuthorityYear="1832" box="[767,1336,1585,1610]" class="Polyplacophora" family="Acanthochitonidae" genus="Acanthochitona" kingdom="Animalia" order="Chitonida" pageId="29" pageNumber="480" phylum="Mollusca" rank="species" species="hirudiniformis">
|
||
<emphasis id="96F4EABFDD4FFFAFFD783018FC0AD365" box="[767,961,1585,1610]" italics="true" pageId="29" pageNumber="480">A. hirudiniformis</emphasis>
|
||
(Sowerby, 1832) (
|
||
<bibRefCitation id="C0114B5CDD4FFFAFFB12301BFAFAD366" author="Watters" box="[1173,1329,1585,1609]" pageId="29" pageNumber="480" refString="Watters. G. T. (1990) A review of the recent eastern Pacific Acanthochitoninae (Mollusca: Polyplacophora: Cryptoplacidae) with the description of a new genus, Americhiton. The Feliger, 33, 241 - 271." type="journal article" year="1990">Watters 1990</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, we also regard them as intraspecific variations.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="EC9A6526DD4FFFAFFF403053FBFDD2A2" pageId="29" pageNumber="480" type="reference_group">
|
||
<paragraph id="A43F36ADDD4FFFAFFF403053FB3BD3D1" blockId="29.[151,1437,1260,1934]" pageId="29" pageNumber="480">
|
||
<taxonomicName id="63804D2EDD4FFFAFFF403053FE14D3BD" ID-CoL="64DSH" authorityName="Kaas & Van Belle" authorityYear="1980" box="[199,479,1657,1682]" class="Polyplacophora" family="Acanthochitonidae" genus="Acanthochitona" kingdom="Animalia" order="Chitonida" pageId="29" pageNumber="480" phylum="Mollusca" rank="species" species="biformis">
|
||
<emphasis id="96F4EABFDD4FFFAFFF403053FE14D3BD" box="[199,479,1657,1682]" italics="true" pageId="29" pageNumber="480">Acanthochitona biformis</emphasis>
|
||
</taxonomicName>
|
||
differs from its Indo-West Pacific congeners by having wide valves with a narrow jugum and densely packed oval pustules. The morphology of the pustules and radula show similarity to those of
|
||
<taxonomicName id="63804D2EDD4FFFAFFF1030EBFD6ED3F5" ID-CoL="64DTL" authority="Nierstrasz, 1905" authorityName="Nierstrasz" authorityYear="1905" box="[151,677,1729,1754]" class="Polyplacophora" family="Acanthochitonidae" genus="Acanthochitona" kingdom="Animalia" order="Chitonida" pageId="29" pageNumber="480" phylum="Mollusca" rank="species" species="intermedia">
|
||
<emphasis id="96F4EABFDD4FFFAFFF1030EBFE00D3F5" box="[151,459,1729,1754]" italics="true" pageId="29" pageNumber="480">Acanthochitona intermedia</emphasis>
|
||
(
|
||
<bibRefCitation id="C0114B5CDD4FFFAFFE5A30EBFD57D3F5" author="Nierstrasz" box="[477,668,1729,1754]" pageId="29" pageNumber="480" refString="Nierstrasz, H. F. (1905) Die Chitonen der Siboga-Expedition. - Siboga Expeditie, 48, 1 - 112, pls 1 - 8." type="journal article" year="1905">Nierstrasz, 1905</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, but
|
||
<taxonomicName id="63804D2EDD4FFFAFFD6530E8FCAAD3F5" ID-CoL="64DSH" authorityName="Kaas & Van Belle" authorityYear="1980" box="[738,865,1729,1754]" class="Polyplacophora" family="Acanthochitonidae" genus="Acanthochitona" kingdom="Animalia" order="Chitonida" pageId="29" pageNumber="480" phylum="Mollusca" rank="species" species="biformis">
|
||
<emphasis id="96F4EABFDD4FFFAFFD6530E8FCAAD3F5" box="[738,865,1729,1754]" italics="true" pageId="29" pageNumber="480">A. biformis</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="63804D2EDD4FFFAFFC7930E8FB51D3F5" ID-CoL="64DTL" authorityName="Nierstrasz" authorityYear="1905" box="[1022,1178,1729,1754]" class="Polyplacophora" family="Acanthochitonidae" genus="Acanthochitona" kingdom="Animalia" order="Chitonida" pageId="29" pageNumber="480" phylum="Mollusca" rank="species" species="intermedia">
|
||
<emphasis id="96F4EABFDD4FFFAFFC7930E8FB51D3F5" box="[1022,1178,1729,1754]" italics="true" pageId="29" pageNumber="480">A. intermedia</emphasis>
|
||
</taxonomicName>
|
||
by having a narrower jugum and more round pustules with a much narrower prepustular slope which is sunken in the slit.
|
||
</paragraph>
|
||
<paragraph id="A43F36ADDD4FFFAFFF403123FBFDD2A2" blockId="29.[151,1437,1260,1934]" pageId="29" pageNumber="480">
|
||
The present species superficially resembles
|
||
<taxonomicName id="63804D2EDD4FFFAFFD3D3123FB48D20E" ID-CoL="64DSQ" authority="Leloup, 1933" authorityName="Leloup" authorityYear="1933" box="[698,1155,1801,1826]" class="Polyplacophora" family="Acanthochitonidae" genus="Acanthochitona" kingdom="Animalia" order="Chitonida" pageId="29" pageNumber="480" phylum="Mollusca" rank="species" species="leopoldi">
|
||
<emphasis id="96F4EABFDD4FFFAFFD3D3123FC1BD20D" box="[698,976,1801,1826]" italics="true" pageId="29" pageNumber="480">Acanthochitona leopoldi</emphasis>
|
||
(
|
||
<bibRefCitation id="C0114B5CDD4FFFAFFC583123FBB1D20E" author="Leloup" box="[991,1146,1801,1826]" pageId="29" pageNumber="480" refString="Leloup, E. (1933) Resultats scientifiques du voyage aux Indes Orientales neerlandaises de LL. AA. RR. le Prince et la Princesse Leopold de Belgique. Brachiopodes et Amphineures. Memoires du Musee royal d'histoire naturelle de Belgique, 2 (13), 5 - 33, pls. 1 - 2." type="journal article" year="1933">Leloup, 1933</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, but differs in shape and in the arrangement of the pustules (oval to droplet-shape and arranged in quincunx order in the former, while narrow droplet-shape and arranged in comarginally in the latter. Also
|
||
<taxonomicName id="63804D2EDD4FFFAFFC363178FBE6D245" ID-CoL="64DSQ" authorityName="Leloup" authorityYear="1933" box="[945,1069,1873,1898]" class="Polyplacophora" family="Acanthochitonidae" genus="Acanthochitona" kingdom="Animalia" order="Chitonida" pageId="29" pageNumber="480" phylum="Mollusca" rank="species" species="leopoldi">
|
||
<emphasis id="96F4EABFDD4FFFAFFC363178FBE6D245" box="[945,1069,1873,1898]" italics="true" pageId="29" pageNumber="480">A. leopoldi</emphasis>
|
||
</taxonomicName>
|
||
is more spiculous in the sutures between the valves. This is the first record of this species from Vietnamese waters.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |