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<document ID-CLB-Dataset="9572" ID-DOI="http://dx.doi.org/10.3897/zookeys.994.56810" ID-GBIF-Dataset="8639369d-0b63-4725-aa0c-aed78224bbe5" ID-PMC="PMC7686225" ID-Pensoft-Pub="1313-2970-994-1" ID-Pensoft-UUID="11A23FB601685F7ABFCC123875393447" ID-PubMed="33273881" ID-ZooBank="DB1879C919044320BE674D9F863F6775" ModsDocID="1313-2970-994-1" checkinTime="1605693934925" checkinUser="pensoft" docAuthor="Ng, Peter K. L." docDate="2020" docId="253627CA174B5C69B8B6231C187A0709" docLanguage="en" docName="ZooKeys 994: 1-34" docOrigin="ZooKeys 994" docSource="http://dx.doi.org/10.3897/zookeys.994.56810" docTitle="Johora booliati Ng 2020, sp. nov." docType="treatment" docUuid="8A3C9173-A7A5-4559-92D7-A548453C6688" docUuidSource="ZooBank" docVersion="6" id="11A23FB601685F7ABFCC123875393447" lastPageNumber="1" masterDocId="11A23FB601685F7ABFCC123875393447" masterDocTitle="Revision of the freshwater crabs of the Johora tahanensis (Bott, 1966) species group (Crustacea, Brachyura, Potamidae), with a key to the genus" masterLastPageNumber="34" masterPageNumber="1" pageNumber="1" updateTime="1732749455418" updateUser="ExternalLinkService">
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<mods:title id="586C735944B49C9673C55273B1B81CC3">Revision of the freshwater crabs of the Johora tahanensis (Bott, 1966) species group (Crustacea, Brachyura, Potamidae), with a key to the genus</mods:title>
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<mods:name id="D122C1B52EAAE0915F033DBB404F7600" type="personal">
<mods:role id="F9A5556CA1952BBAFE2CA0244776D201">
<mods:roleTerm id="2C98C552857EC1EA9339BF9120E2641F">Author</mods:roleTerm>
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<mods:namePart id="9E1A9D571205E6C1948A43231FFD6BF3">Ng, Peter K. L.</mods:namePart>
<mods:affiliation id="4B1B85AEDD2A8FFA98DCF1B17195E509">Lee Kong Chian Natural History Museum, National University of Singapore, 2 Conservatory Drive, Singapore 117377, Singapore</mods:affiliation>
<mods:nameIdentifier id="F80CD11B0DE6AC4D6CAC369E7A663D2F" type="orcid">https://orcid.org/0000-0001-5946-0608</mods:nameIdentifier>
<mods:nameIdentifier id="632CC7AA7F25DE4A465E8CEF87104C59" type="email">peterng@nus.edu.sg</mods:nameIdentifier>
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<mods:title id="6A555D0B85D619F43C05880C60FF4F29">ZooKeys</mods:title>
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<mods:part id="2F0D1E9ECCFE7760ADDA75EE1F5D9888">
<mods:date id="048C59710345F6F4BA379D551FB43600">2020</mods:date>
<mods:detail id="16A495CB0A27294F3ACAC8E8678ED2A3" type="volume">
<mods:number id="5FF9992886FE95EC991904224ADAD54E">994</mods:number>
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<mods:url id="8B0666955A561A348B2A3339CAD2A11D">http://dx.doi.org/10.3897/zookeys.994.56810</mods:url>
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<mods:classification id="B52889ED72A7521FD7D0835F8970938E">journal article</mods:classification>
<mods:identifier id="79AC1CC40003D05C5A992FF486232F92" type="DOI">http://dx.doi.org/10.3897/zookeys.994.56810</mods:identifier>
<mods:identifier id="FAF529B874FEE3D3E3F6E12149B7CE0B" type="Pensoft-Pub">1313-2970-994-1</mods:identifier>
<mods:identifier id="BFFF86047A0165650F63EB772EF51241" type="ZooBank">DB1879C919044320BE674D9F863F6775</mods:identifier>
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<treatment id="253627CA174B5C69B8B6231C187A0709" ID-GBIF-Taxon="169094345" LSID="urn:lsid:zoobank.org:act:8A3C9173-A7A5-4559-92D7-A548453C6688" httpUri="http://treatment.plazi.org/id/253627CA174B5C69B8B6231C187A0709" lastPageNumber="1" pageId="0" pageNumber="1" scope_family="Potamidae" scope_infraOrder="Brachyura">
<subSubSection id="6C0E3E19E5C8C65BB9EAAF23C418AAC9" pageId="0" pageNumber="1" type="nomenclature">
<paragraph id="990A4FC7D861E3242AFADE1A87818E88" pageId="0" pageNumber="1">
<taxonomicName id="C7F20C93E1FD59582117C2C8E912DF4A" ID-CoL="84GXG" LSID="253627CA-174B-5C69-B8B6-231C187A0709" authority="Ng, 2020" authorityName="Ng" authorityYear="2020" class="Malacostraca" family="Potamidae" genus="Johora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Johora booliati" order="Decapoda" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="booliati" status="sp. nov.">Johora booliati</taxonomicName>
<taxonomicNameLabel id="056AA6BC78B2657ED2B24689878D5985" pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
<figureCitation id="D950420F53A6058947B519DBB18B25EE" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Overall dorsal view A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249), Terengganu D J. hoiseni Ng &amp; Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure1" httpUri="https://binary.pensoft.net/fig/476312" pageId="0" pageNumber="1">Figures 1F</figureCitation>
<figureCitation id="E3A39C642547E679D85A3828786AAF59" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dorsal view of carapace A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249) Terengganu D J. hoiseni Ng &amp; Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure2" httpUri="https://binary.pensoft.net/fig/476313" pageId="0" pageNumber="1">, 2F</figureCitation>
<figureCitation id="1D23BDE5B29B3C39094079FD840174E7" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Frontal view of cephalothorax A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249), Terengganu D J. hoiseni Ng &amp; Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure3" httpUri="https://binary.pensoft.net/fig/476314" pageId="0" pageNumber="1">, 3F</figureCitation>
<figureCitation id="A091D18B9ABC8C414F8066735389CECD" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Left third maxilliped A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249), Terengganu D J. hoiseni Ng &amp; Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure4" httpUri="https://binary.pensoft.net/fig/476315" pageId="0" pageNumber="1">, 4F</figureCitation>
<figureCitation id="EA284283D919140A2401C433E8E46736" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Anterior thoracic sternum and pleon A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249), Terengganu D J. hoiseni Ng &amp; Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure5" httpUri="https://binary.pensoft.net/fig/476316" pageId="0" pageNumber="1">, 5F</figureCitation>
<figureCitation id="7F245CAFDE2D8B891E627E185065F78A" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. A-D Johora thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand E-I J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang A, E left G 1 (ventral view) B, F distal part of left G 1 (ventral view) C, H distal part of left G 1 (dorsal view) G distal part of left G 1 (mesial view) D, I left G 2 (ventral view). Scale bas: 1.0 mm (A, D, E, I); 0.5 mm (B, C, F-H)." figureDoi="10.3897/zookeys.994.56810.figure10" httpUri="https://binary.pensoft.net/fig/476321" pageId="0" pageNumber="1">, 10E-I</figureCitation>
<figureCitation id="80B40749EFACA0FE1889AF7C7CD1F987" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Overall dorsal view A Johora intermedia Ng, 1986, female (17.9 x 14.7 mm) (ZRC 2001.1002), Selangor B J. tahanensis (Bott, 1966), female (24.4 x 21.0 mm) (ZRC 1984.6765), Pahang C J. thoi Ng, 1990, female (31.7 x 25.5 mm) (ZRC 2001.1167), Terengganu D J. hoiseni Ng &amp; Takeda, 1992, paratype female (22.1 x 19.0 mm) (ZRC 1984.6675), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype female (21.0 x 17.4 mm) (ZRC 2006.0053), Thailand F J. booliati sp. nov., paratype female (40.4 x 34.4 mm) (ZRC 1995.270), Pahang G J. erici sp. nov., paratype female (32.8 x 26.9 mm) (ZRC 1995.268), Perak H J. michaeli sp. nov., paratype female (29.9 x 25.0 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure12" httpUri="https://binary.pensoft.net/fig/476323" pageId="0" pageNumber="1">, 12F</figureCitation>
<figureCitation id="E3255CCED8703197A5465C7F7A81C5BE" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Female pleon A Johora intermedia Ng, 1986, female (17.9 x 14.7 mm) (ZRC 2001.1002), Selangor B J. tahanensis (Bott, 1966), female (24.4 x 21.0 mm) (ZRC 1984.6765), Pahang C J. thoi Ng, 1990, female (31.7 x 25.5 mm) (ZRC 2001.1167), Terengganu D J. hoiseni Ng &amp; Takeda, 1992, paratype female (22.1 x 19.0 mm) (ZRC 1984.6675), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype female (21.0 x 17.4 mm) (ZRC 2006.0053), Thailand F J. booliati sp. nov., paratype female (40.4 x 34.4 mm) (ZRC 1995.270), Pahang G J. erici sp. nov., paratype female (32.8 x 26.9 mm) (ZRC 1995.268), Perak H J. michaeli sp. nov., paratype female (29.9 x 25.0 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure13" httpUri="https://binary.pensoft.net/fig/476324" pageId="0" pageNumber="1">, 13F</figureCitation>
<figureCitation id="9F3957A8EFB970A5558DD2DFAFCE6B5D" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Sternopleonal cavity and vulvae A Johora intermedia Ng, 1986, female (17.9 x 14.7 mm) (ZRC 2001.1002), Selangor B J. tahanensis (Bott, 1966), female (24.4 x 21.0 mm) (ZRC 1984.6765), Pahang C J. thoi Ng, 1990, female (31.7 x 25.5 mm) (ZRC 2001.1167), Terengganu D J. hoiseni Ng &amp; Takeda, 1992, paratype female (22.1 x 19.0 mm) (ZRC 1984.6675), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype female (21.0 x 17.4 mm) (ZRC 2006.0053), Thailand F J. booliati sp. nov., paratype female (40.4 x 34.4 mm) (ZRC 1995.270), Pahang G J. erici sp. nov., paratype female (32.8 x 26.9 mm) (ZRC 1995.268), Perak H J. michaeli sp. nov., paratype female (29.9 x 25.0 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure14" httpUri="https://binary.pensoft.net/fig/476325" pageId="0" pageNumber="1">, 14F</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="38E287492680E56A6133F034BC2A6773" pageId="0" pageNumber="1" type="reference_group">
<paragraph id="63DEC5A618E865BA493892988AA343F6" pageId="0" pageNumber="1">
<taxonomicName id="63F131481920FA00F305491957A3114A" authorityName="Bott" authorityYear="1966" class="Malacostraca" family="Potamidae" genus="Johora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Johora tahanensis" order="Decapoda" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="tahanensis">Johora tahanensis</taxonomicName>
- Takeda &amp; Ng, 1992: 107 (part); Ng 1995: 249 (part) (not
<taxonomicName id="E44633A785C206D58AB3DFDB1A8682BA" class="Malacostraca" family="Potamidae" genus="Stoliczia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stoliczia (Johora) johorensis subsp. tahanensis" order="Decapoda" pageId="0" pageNumber="1" phylum="Arthropoda" rank="subSpecies" species="johorensis" subGenus="Johora" subSpecies="tahanensis">Stoliczia (Johora) johorensis tahanensis</taxonomicName>
Bott, 1966).
</paragraph>
</subSubSection>
<subSubSection id="60EC6151B3EDAB645AA85AF4D5A31A30" pageId="0" pageNumber="1" type="materials_examined">
<paragraph id="D5C893233DBEECF800FF3E08311DBA13" pageId="0" pageNumber="1">Material examined.</paragraph>
<paragraph id="8A11DD3BC7782615C5BA3B88B69F184F" pageId="0" pageNumber="1">
<emphasis id="D844C060A2AF9FAE58C9B4BC6A1C44AE" bold="true" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
: male (34.6
<normalizedToken id="23FFD3075242B3C14783CCE8E4B21507" originalValue="×">x</normalizedToken>
30.0 mm) (ZRC 2020.0072), stream next to house, under rocks, in old farmland and secondary forest, Bukit Tinggi village, west side of northbound Karak Highway, ca. 480 m a.s.l., Bentong District, Pahang,
<geoCoordinate id="8E3363F6153D98B842FB8FCCF20B120F" degrees="3" direction="north" minutes="22" orientation="latitude" precision="1" seconds="3.0396" value="3.367511">3°22'3.0396&quot;N</geoCoordinate>
,
<geoCoordinate id="4ED49A70390F54214C190DC90BF5F3C8" degrees="101" direction="east" minutes="48" orientation="longitude" precision="1" seconds="50.994" value="101.81417">101°48'50.994&quot;E</geoCoordinate>
, coll. July 2016. Paratype: 1 female (40.4
<normalizedToken id="6E23BA00601D474583FCBC04E5516F7C" originalValue="×">x</normalizedToken>
34.4 mm) (ZRC 1995.270), in shallow stream with leaf litter, The Gap,
<normalizedToken id="25E7FB6511BBBE647E37AB8E2D08102D" originalValue="Fraser’s">Fraser's</normalizedToken>
Hill, 853 m a.s.l., Pahang, ca.
<geoCoordinate id="4D606E5AE7D8C2096974B14A8D491D83" degrees="3" direction="north" minutes="41" orientation="latitude" precision="15" seconds="29" value="3.6913888">3°41'29&quot;N</geoCoordinate>
,
<geoCoordinate id="2BC9DC49FE8072FDB733F9952B6BE821" degrees="101" direction="east" minutes="44" orientation="longitude" precision="15" seconds="56" value="101.748886">101°44'56&quot;E</geoCoordinate>
, coll. Lim KKP, 1 June 1990; 1 juvenile male (10.5
<normalizedToken id="04EF927501615538C97C3730FE7F9133" originalValue="×">x</normalizedToken>
9.1 mm) (ZRC 2020.0364), under rocks, first stream on road towards Raub, descending from
<normalizedToken id="3B9D6EC81A08FC3223A6211464FF94C3" originalValue="Fraser’s">Fraser's</normalizedToken>
Hill, Pahang, coll. Lai JCY, 6 December 2015. All locations in Peninsular Malaysia.
</paragraph>
</subSubSection>
<subSubSection id="DC3663242B1C28ADE45A3A1E5804DD1B" pageId="0" pageNumber="1" type="diagnosis">
<paragraph id="4ED1BC9EEAF3754F08A957956E81DB33" pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph id="87C370A22E36A612EBA4187F784DEEDB" pageId="0" pageNumber="1">
Adult carapace width to length ratio 1.15-1.17 (Figs
<figureCitation id="39D928D3FEFC93A192BFB3CF57594937" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Overall dorsal view A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249), Terengganu D J. hoiseni Ng &amp; Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure1" httpUri="https://binary.pensoft.net/fig/476312" pageId="0" pageNumber="1">1F</figureCitation>
,
<figureCitation id="D4912506856F008F285C1ED5C740609F" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dorsal view of carapace A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249) Terengganu D J. hoiseni Ng &amp; Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure2" httpUri="https://binary.pensoft.net/fig/476313" pageId="0" pageNumber="1">2F</figureCitation>
,
<figureCitation id="1BF2AAEA3AB48C10FE94EF5D979F650D" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Overall dorsal view A Johora intermedia Ng, 1986, female (17.9 x 14.7 mm) (ZRC 2001.1002), Selangor B J. tahanensis (Bott, 1966), female (24.4 x 21.0 mm) (ZRC 1984.6765), Pahang C J. thoi Ng, 1990, female (31.7 x 25.5 mm) (ZRC 2001.1167), Terengganu D J. hoiseni Ng &amp; Takeda, 1992, paratype female (22.1 x 19.0 mm) (ZRC 1984.6675), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype female (21.0 x 17.4 mm) (ZRC 2006.0053), Thailand F J. booliati sp. nov., paratype female (40.4 x 34.4 mm) (ZRC 1995.270), Pahang G J. erici sp. nov., paratype female (32.8 x 26.9 mm) (ZRC 1995.268), Perak H J. michaeli sp. nov., paratype female (29.9 x 25.0 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure12" httpUri="https://binary.pensoft.net/fig/476323" pageId="0" pageNumber="1">12F</figureCitation>
); dorsal surface gently convex in frontal view, not inflated (Fig.
<figureCitation id="510338188717B074EE0ABF08569F2E07" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Frontal view of cephalothorax A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249), Terengganu D J. hoiseni Ng &amp; Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure3" httpUri="https://binary.pensoft.net/fig/476314" pageId="0" pageNumber="1">3F</figureCitation>
); frontal margin almost straight (Fig.
<figureCitation id="A417704708382B66E3ADB2B26B2C9E2C" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dorsal view of carapace A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249) Terengganu D J. hoiseni Ng &amp; Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure2" httpUri="https://binary.pensoft.net/fig/476313" pageId="0" pageNumber="1">2F</figureCitation>
); suborbital, pterygostomial and sub-branchial regions rugose, pterygostomial region covered with dense setae (Fig.
<figureCitation id="D7688594CB4B2C3A317A910461731ABC" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Frontal view of cephalothorax A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249), Terengganu D J. hoiseni Ng &amp; Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure3" httpUri="https://binary.pensoft.net/fig/476314" pageId="0" pageNumber="1">3F</figureCitation>
); epigastric cristae distinct, distinctly anterior to sharp postorbital cristae, postorbital cristae with lateral edges low, joining lateral margin through oblique striae (Fig.
<figureCitation id="8DBDE50B7EB22AC1DFA8637717975FC9" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dorsal view of carapace A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249) Terengganu D J. hoiseni Ng &amp; Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure2" httpUri="https://binary.pensoft.net/fig/476313" pageId="0" pageNumber="1">2F</figureCitation>
); external orbital tooth separated from epibranchial tooth by distinct cleft, epibranchial tooth sharp, distinct (Fig.
<figureCitation id="BE1545E7FD7B576CCCC0F5588BCFABEB" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dorsal view of carapace A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249) Terengganu D J. hoiseni Ng &amp; Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure2" httpUri="https://binary.pensoft.net/fig/476313" pageId="0" pageNumber="1">2F</figureCitation>
); anterolateral margin gently convex (Fig.
<figureCitation id="2A91D6399E922BB8E494A4B34847F89E" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dorsal view of carapace A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249) Terengganu D J. hoiseni Ng &amp; Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure2" httpUri="https://binary.pensoft.net/fig/476313" pageId="0" pageNumber="1">2F</figureCitation>
); posterolateral margin gently sinuous to almost straight, gently converging towards sinuous posterior carapace margin with shallow median indentation (Fig.
<figureCitation id="861423E860A4C0341718E11ACC7FB6A5" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dorsal view of carapace A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249) Terengganu D J. hoiseni Ng &amp; Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure2" httpUri="https://binary.pensoft.net/fig/476313" pageId="0" pageNumber="1">2F</figureCitation>
); posterior margin of epistome with triangular median triangle, lateral margin obliquely sloping (Fig.
<figureCitation id="29578D47630C3392019164B3DF01501F" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Frontal view of cephalothorax A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249), Terengganu D J. hoiseni Ng &amp; Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure3" httpUri="https://binary.pensoft.net/fig/476314" pageId="0" pageNumber="1">3F</figureCitation>
); outer surfaces of third maxillipeds with dense, long stiff setae; ischium subrectangular, with shallow median oblique groove (Figs
<figureCitation id="D56978971B479C8F6D95AC1065AC8421" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Frontal view of cephalothorax A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249), Terengganu D J. hoiseni Ng &amp; Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure3" httpUri="https://binary.pensoft.net/fig/476314" pageId="0" pageNumber="1">3F</figureCitation>
,
<figureCitation id="06D64E4B5DC0B6B1E2D371AE3DE58A0B" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Left third maxilliped A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249), Terengganu D J. hoiseni Ng &amp; Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure4" httpUri="https://binary.pensoft.net/fig/476315" pageId="0" pageNumber="1">4F</figureCitation>
); ambulatory legs not elongate, length to width ratio of merus of fourth ambulatory leg 2.7-2.8 (Figs
<figureCitation id="C9EB35E1A005B82289A7FA1D9176CCE3" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Overall dorsal view A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249), Terengganu D J. hoiseni Ng &amp; Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure1" httpUri="https://binary.pensoft.net/fig/476312" pageId="0" pageNumber="1">1F</figureCitation>
,
<figureCitation id="D5D9E199D197851A85E8CE3A4FE3F96B" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Overall dorsal view A Johora intermedia Ng, 1986, female (17.9 x 14.7 mm) (ZRC 2001.1002), Selangor B J. tahanensis (Bott, 1966), female (24.4 x 21.0 mm) (ZRC 1984.6765), Pahang C J. thoi Ng, 1990, female (31.7 x 25.5 mm) (ZRC 2001.1167), Terengganu D J. hoiseni Ng &amp; Takeda, 1992, paratype female (22.1 x 19.0 mm) (ZRC 1984.6675), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype female (21.0 x 17.4 mm) (ZRC 2006.0053), Thailand F J. booliati sp. nov., paratype female (40.4 x 34.4 mm) (ZRC 1995.270), Pahang G J. erici sp. nov., paratype female (32.8 x 26.9 mm) (ZRC 1995.268), Perak H J. michaeli sp. nov., paratype female (29.9 x 25.0 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure12" httpUri="https://binary.pensoft.net/fig/476323" pageId="0" pageNumber="1">12F</figureCitation>
); G1 subterminal segment with broad proximal part, tapering relatively suddenly to slender distal part, with low shelf-like structure on outer margin; terminal segment gently curved outwards (from median part of sternum), less than half length of subterminal segment, surfaces with numerous short setae (Fig.
<figureCitation id="F1AB4969C7FEC3B4C27418F2C7820BC8" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. A-D Johora thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand E-I J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang A, E left G 1 (ventral view) B, F distal part of left G 1 (ventral view) C, H distal part of left G 1 (dorsal view) G distal part of left G 1 (mesial view) D, I left G 2 (ventral view). Scale bas: 1.0 mm (A, D, E, I); 0.5 mm (B, C, F-H)." figureDoi="10.3897/zookeys.994.56810.figure10" httpUri="https://binary.pensoft.net/fig/476321" pageId="0" pageNumber="1">10E-H</figureCitation>
); G2 slightly longer than G1, distal segment long, about half length of basal segment (Fig.
<figureCitation id="4B4711F081D4291ADF643E7CD4A203E6" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. A-D Johora thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand E-I J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang A, E left G 1 (ventral view) B, F distal part of left G 1 (ventral view) C, H distal part of left G 1 (dorsal view) G distal part of left G 1 (mesial view) D, I left G 2 (ventral view). Scale bas: 1.0 mm (A, D, E, I); 0.5 mm (B, C, F-H)." figureDoi="10.3897/zookeys.994.56810.figure10" httpUri="https://binary.pensoft.net/fig/476321" pageId="0" pageNumber="1">10I</figureCitation>
). Female pleon ovate; somite 3 less wide than somite 4, somites 4-6 progressively narrower; telson semi-circular (Fig.
<figureCitation id="75FB805073DAB74E96036B01C07DE9F3" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Female pleon A Johora intermedia Ng, 1986, female (17.9 x 14.7 mm) (ZRC 2001.1002), Selangor B J. tahanensis (Bott, 1966), female (24.4 x 21.0 mm) (ZRC 1984.6765), Pahang C J. thoi Ng, 1990, female (31.7 x 25.5 mm) (ZRC 2001.1167), Terengganu D J. hoiseni Ng &amp; Takeda, 1992, paratype female (22.1 x 19.0 mm) (ZRC 1984.6675), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype female (21.0 x 17.4 mm) (ZRC 2006.0053), Thailand F J. booliati sp. nov., paratype female (40.4 x 34.4 mm) (ZRC 1995.270), Pahang G J. erici sp. nov., paratype female (32.8 x 26.9 mm) (ZRC 1995.268), Perak H J. michaeli sp. nov., paratype female (29.9 x 25.0 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure13" httpUri="https://binary.pensoft.net/fig/476324" pageId="0" pageNumber="1">13F</figureCitation>
). Vulvae large, on anterior half of sternite 6, slightly pushing into suture with sternite 5, lateral sternal vulvar cover semi-circular (Fig.
<figureCitation id="3B4B4826AA60891F6ED0C71DB4DAD68E" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Sternopleonal cavity and vulvae A Johora intermedia Ng, 1986, female (17.9 x 14.7 mm) (ZRC 2001.1002), Selangor B J. tahanensis (Bott, 1966), female (24.4 x 21.0 mm) (ZRC 1984.6765), Pahang C J. thoi Ng, 1990, female (31.7 x 25.5 mm) (ZRC 2001.1167), Terengganu D J. hoiseni Ng &amp; Takeda, 1992, paratype female (22.1 x 19.0 mm) (ZRC 1984.6675), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype female (21.0 x 17.4 mm) (ZRC 2006.0053), Thailand F J. booliati sp. nov., paratype female (40.4 x 34.4 mm) (ZRC 1995.270), Pahang G J. erici sp. nov., paratype female (32.8 x 26.9 mm) (ZRC 1995.268), Perak H J. michaeli sp. nov., paratype female (29.9 x 25.0 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure14" httpUri="https://binary.pensoft.net/fig/476325" pageId="0" pageNumber="1">14F</figureCitation>
).
</paragraph>
<caption id="F491AD5AD63FDDA6FDF9170964270CF9" doi="10.3897/zookeys.994.56810.figure12" httpUri="https://binary.pensoft.net/fig/476323" pageId="0" pageNumber="1" start="Figure 12" startId="F12">
<paragraph id="45FB350AC88B37FA173FB8B69C8BC4A3" pageId="0" pageNumber="1">
<emphasis id="4DC6502C5CBE32291DCB80F6D1B98DE8" bold="true" pageId="0" pageNumber="1">Figure 12.</emphasis>
Overall dorsal view
<emphasis id="269D7CAB89404C54088D14E89547ABD3" bold="true" pageId="0" pageNumber="1">A</emphasis>
<taxonomicName id="40286D70F1DFB937890F0966C5D50FC6" baseAuthorityName="Ng" baseAuthorityYear="1986" class="Malacostraca" family="Potamidae" genus="Johora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Johora intermedia" order="Decapoda" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="intermedia">
<emphasis id="07296548EC0919C154A9A1DCC7538940" italics="true" pageId="0" pageNumber="1">Johora intermedia</emphasis>
</taxonomicName>
Ng, 1986, female (17.9
<normalizedToken id="BD54C5F105D7E8A164C23DD2644D562F" originalValue="×">x</normalizedToken>
14.7 mm) (ZRC 2001.1002), Selangor
<emphasis id="B9EE1BC06591F5B5B80FA87B0E277218" bold="true" pageId="0" pageNumber="1">B</emphasis>
<taxonomicName id="2C263AF17FB65A6B13079CFD73E3F99C" lsidName="J. tahanensis" pageId="0" pageNumber="1" rank="species" species="tahanensis">
<emphasis id="C7BDAEBD400700200B0E31D6A46EE1EA" italics="true" pageId="0" pageNumber="1">J. tahanensis</emphasis>
</taxonomicName>
(Bott, 1966), female (24.4
<normalizedToken id="80958342FE9BC107E7A60669F22F0279" originalValue="×">x</normalizedToken>
21.0 mm) (ZRC 1984.6765), Pahang
<emphasis id="E1BD8D65568CF469FC6237E8E8C0C483" bold="true" pageId="0" pageNumber="1">C</emphasis>
<taxonomicName id="F6609DE701BE0AEC43071E5307DDA7F3" lsidName="J. thoi" pageId="0" pageNumber="1" rank="species" species="thoi">
<emphasis id="2E836947D87539FE8CB7CF8A3B541E2C" italics="true" pageId="0" pageNumber="1">J. thoi</emphasis>
</taxonomicName>
Ng, 1990, female (31.7
<normalizedToken id="D96BD3A48C5F01723F7F7B5CBC36B6F3" originalValue="×">x</normalizedToken>
25.5 mm) (ZRC 2001.1167), Terengganu
<emphasis id="249E3D498B302B9A7DFAABC0BED6518A" bold="true" pageId="0" pageNumber="1">D</emphasis>
<taxonomicName id="D0723681FA96BCB802AD9E4FB2B7A336" lsidName="J. hoiseni" pageId="0" pageNumber="1" rank="species" species="hoiseni">
<emphasis id="9FF87E6594C03BFB6D76DAB9B8C82EAB" italics="true" pageId="0" pageNumber="1">J. hoiseni</emphasis>
</taxonomicName>
Ng &amp; Takeda, 1992, paratype female (22.1
<normalizedToken id="A9E2D7AE7552BA60B7CF9444A7CEAE7F" originalValue="×">x</normalizedToken>
19.0 mm) (ZRC 1984.6675), Kelantan
<emphasis id="02F51834C990AD2FCC1BC0E3092797F7" bold="true" pageId="0" pageNumber="1">E</emphasis>
<taxonomicName id="8361378D1EDD4AA5A6D6CF07E8A56A6F" lsidName="J. thaiana" pageId="0" pageNumber="1" rank="species" species="thaiana">
<emphasis id="80F840AFB96F72FEBBC9D16D9E11B3A1" italics="true" pageId="0" pageNumber="1">J. thaiana</emphasis>
</taxonomicName>
Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype female (21.0
<normalizedToken id="217EDA778196DAF4AD55CF6A0DA2D850" originalValue="×">x</normalizedToken>
17.4 mm) (ZRC 2006.0053), Thailand
<emphasis id="394EDABC3E20764DE4806E295D9BED05" bold="true" pageId="0" pageNumber="1">F</emphasis>
<taxonomicName id="107CDB743125BEEE631A679ED695B666" lsidName="J. booliati" pageId="0" pageNumber="1" rank="species" species="booliati">
<emphasis id="4E7CE2E21E77F5AB07758A25E7FA0511" italics="true" pageId="0" pageNumber="1">J. booliati</emphasis>
</taxonomicName>
sp. nov., paratype female (40.4
<normalizedToken id="29FA07CBBCC1CD7447D1953CECD1CD58" originalValue="×">x</normalizedToken>
34.4 mm) (ZRC 1995.270), Pahang
<emphasis id="79B18632226D0EB1AF3C914944EFA56A" bold="true" pageId="0" pageNumber="1">G</emphasis>
<taxonomicName id="CB103EB2BD5117777C993EDB8265C4D5" lsidName="J. erici" pageId="0" pageNumber="1" rank="species" species="erici">
<emphasis id="9D99CDA49062D9F6585033D24443A265" italics="true" pageId="0" pageNumber="1">J. erici</emphasis>
</taxonomicName>
sp. nov., paratype female (32.8
<normalizedToken id="6CB380C18E0F4B5A7DB50BA7B59BDBBF" originalValue="×">x</normalizedToken>
26.9 mm) (ZRC 1995.268), Perak
<emphasis id="9AF3F4D0409993F32AEE22947F3D2217" bold="true" pageId="0" pageNumber="1">H</emphasis>
<taxonomicName id="8A60EF588D1E8B6620B39AA809990EF0" lsidName="J. michaeli" pageId="0" pageNumber="1" rank="species" species="michaeli">
<emphasis id="2F0624500FC101428181E697F7306769" italics="true" pageId="0" pageNumber="1">J. michaeli</emphasis>
</taxonomicName>
sp. nov., paratype female (29.9
<normalizedToken id="1190D9D857A3F4799417A5B5D66CAA03" originalValue="×">x</normalizedToken>
25.0 mm) (ZRC 2010.0047), Terengganu.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="EA21CA74E800380E46917778D200A39B" pageId="0" pageNumber="1" type="etymology">
<paragraph id="568D554CD1BB338B66CFA06E7E010EE7" pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph id="109A555A8EDAE3DFD56F0ACB05CA00BB" pageId="0" pageNumber="1">
The species is named after an old friend and mentor, the late Dr Lim Boo Liat (1926-2020), one of
<normalizedToken id="F55182104C4662F37E9033F5E970C5A9" originalValue="Malaysia’s">Malaysia's</normalizedToken>
most eminent biologists and naturalists. In a remarkable career spanning 70 years, he has contributed immensely to biodiversity and conservation science in Malaysia; more importantly, he has inspired, mentored, and trained many generations of biologists, many who now lead science in Malaysia and Singapore.
</paragraph>
</subSubSection>
<subSubSection id="C28C72211A44ABCF7561A67042C2AD60" pageId="0" pageNumber="1" type="remarks">
<paragraph id="3BC5AFE2E70476CCF7380DF14F626F72" pageId="0" pageNumber="1">Remarks.</paragraph>
<paragraph id="FF87AB73BEE960D12F894BF4857B116A" pageId="0" pageNumber="1">
The large adult size of
<taxonomicName id="48B153B90E8AB00912A8E59F7931A0E1" lsidName="J. booliati" pageId="0" pageNumber="1" rank="species" species="booliati">
<emphasis id="141886BC63A7108B7ACB811C2E97DD4E" italics="true" pageId="0" pageNumber="1">J. booliati</emphasis>
</taxonomicName>
sp. nov. and
<taxonomicName id="61580C971B04B3F0F5EBF9290F88C82C" lsidName="J. erici" pageId="0" pageNumber="1" rank="species" species="erici">
<emphasis id="3603B74DFC2C63FE2BAA63618AA1547E" italics="true" pageId="0" pageNumber="1">J. erici</emphasis>
</taxonomicName>
sp. nov. (carapace width in excess of 25 mm) with the pronounced epibranchial teeth allies them with
<taxonomicName id="A88BC144FB135250F0649145DDE17F13" lsidName="J. tahanensis" pageId="0" pageNumber="1" rank="species" species="tahanensis">
<emphasis id="5E357913764FE1F2734F51DD97044D0C" italics="true" pageId="0" pageNumber="1">J. tahanensis</emphasis>
</taxonomicName>
s. str.,
<taxonomicName id="67AE305E769D9DB98D599F37EA0FC77E" lsidName="J. hoiseni" pageId="0" pageNumber="1" rank="species" species="hoiseni">
<emphasis id="BCFFEB0498F034EA5A33C2F3FCDF3BCC" italics="true" pageId="0" pageNumber="1">J. hoiseni</emphasis>
</taxonomicName>
,
<taxonomicName id="EC38BBC93853DA02DA727F9F1881D90B" lsidName="J. thaiana" pageId="0" pageNumber="1" rank="species" species="thaiana">
<emphasis id="2C9AD5F892F0017A2529BD36F700290A" italics="true" pageId="0" pageNumber="1">J. thaiana</emphasis>
</taxonomicName>
,
<taxonomicName id="C36BF2DF90FC1EC66514B5D09A3229B3" lsidName="J. thoi" pageId="0" pageNumber="1" rank="species" species="thoi">
<emphasis id="D7A62A9A073D0282A971027ED48B0EEB" italics="true" pageId="0" pageNumber="1">J. thoi</emphasis>
</taxonomicName>
,
<taxonomicName id="5B101DA7A7C162EDECBEB81D97906F6B" lsidName="J. tiomanensis" pageId="0" pageNumber="1" rank="species" species="tiomanensis">
<emphasis id="A4A36059E5D86E75B5D32BF918A410D0" italics="true" pageId="0" pageNumber="1">J. tiomanensis</emphasis>
</taxonomicName>
, and
<taxonomicName id="873148FF977471274A5E4D2D98671AC5" lsidName="J. counsilmani" pageId="0" pageNumber="1" rank="species" species="counsilmani">
<emphasis id="E66A355C91E6259610BC373BB71D0BB6" italics="true" pageId="0" pageNumber="1">J. counsilmani</emphasis>
</taxonomicName>
(the last two being endemic to Pulau Tioman, Peninsular Malaysia).
</paragraph>
<paragraph id="D1AA4F74D4B3293F8B550454EE1AFF99" pageId="0" pageNumber="1">
<taxonomicName id="1BB791BE9AD72EDF089136765BD5D66D" authorityName="Ng" authorityYear="2020" class="Malacostraca" family="Potamidae" genus="Johora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Johora booliati" order="Decapoda" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="booliati">
<emphasis id="6CD053D8F8DB45750CAFFA4E28CACA91" italics="true" pageId="0" pageNumber="1">Johora booliati</emphasis>
</taxonomicName>
and
<taxonomicName id="C73366A70D9A36B3EB18CFD97144B978" lsidName="J. erici" pageId="0" pageNumber="1" rank="species" species="erici">
<emphasis id="51A67D5F59FD53A4499102C53AFF3885" italics="true" pageId="0" pageNumber="1">J. erici</emphasis>
</taxonomicName>
resemble
<taxonomicName id="91E6BCACAB95AA026B8C73869E65BBBD" lsidName="J. tahanensis" pageId="0" pageNumber="1" rank="species" species="tahanensis">
<emphasis id="D0D8F434226836D424657822C6671ED6" italics="true" pageId="0" pageNumber="1">J. tahanensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4F347B4FB444B2060546657129511D89" lsidName="J. hoiseni" pageId="0" pageNumber="1" rank="species" species="hoiseni">
<emphasis id="F4B6E69C647B49F88963DB91236796E1" italics="true" pageId="0" pageNumber="1">J. hoiseni</emphasis>
</taxonomicName>
, and
<taxonomicName id="32D9CE5FE3DFE1170035B1BA74D661C2" lsidName="J. thaiana" pageId="0" pageNumber="1" rank="species" species="thaiana">
<emphasis id="B57193AF1BB4A95A9DF8D02BA45870CA" italics="true" pageId="0" pageNumber="1">J. thaiana</emphasis>
</taxonomicName>
in the carapace shape and general features. Their G1 structures, however, are different. In
<taxonomicName id="31E50C63FDDA2B6DA774C83F699DD0C8" lsidName="J. booliati" pageId="0" pageNumber="1" rank="species" species="booliati">
<emphasis id="5CD4DD8F57F61D19661F5E49EB67041F" italics="true" pageId="0" pageNumber="1">J. booliati</emphasis>
</taxonomicName>
and
<taxonomicName id="68B5BE14647968E3D4E1492ABC561455" lsidName="J. erici" pageId="0" pageNumber="1" rank="species" species="erici">
<emphasis id="4F05DC98D5D56CAE1EBB47D87D8AF979" italics="true" pageId="0" pageNumber="1">J. erici</emphasis>
</taxonomicName>
, the G1 subterminal segment is proportionately more elongate and the terminal segment is relatively shorter (Figs
<figureCitation id="B836F38F83DE9B6FCCEEEDA612628D20" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. A-D Johora thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand E-I J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang A, E left G 1 (ventral view) B, F distal part of left G 1 (ventral view) C, H distal part of left G 1 (dorsal view) G distal part of left G 1 (mesial view) D, I left G 2 (ventral view). Scale bas: 1.0 mm (A, D, E, I); 0.5 mm (B, C, F-H)." figureDoi="10.3897/zookeys.994.56810.figure10" httpUri="https://binary.pensoft.net/fig/476321" pageId="0" pageNumber="1">10E-H</figureCitation>
,
<figureCitation id="05EA46C83D50DFB751BF828690DF6C65" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. A-D Johora erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak E-H J. erici sp. nov., male (41.2 x 33.8 mm) (ZRC 1995.0269), Kelantan A, E left G 1 (ventral view) B, F distal part of left G 1 (ventral view) C, G distal part of left G 1 (dorsal view) D, H left G 2 (ventral view). Scale bars: 1.0 mm (A, D, E, H); 0.5 mm (B, C, F, G)." figureDoi="10.3897/zookeys.994.56810.figure11" httpUri="https://binary.pensoft.net/fig/476322" pageId="0" pageNumber="1">11A-C, E-G</figureCitation>
), and significantly, the terminal segment has only scattered short setae (versus subterminal segment shorter with the terminal segment longer and the surface of the latter more densely covered with short setae in
<taxonomicName id="C26151AD371698C494722232D62058C8" lsidName="J. tahanensis" pageId="0" pageNumber="1" rank="species" species="tahanensis">
<emphasis id="126D2F5E563E7E6C43498A4EC1AD69CA" italics="true" pageId="0" pageNumber="1">J. tahanensis</emphasis>
</taxonomicName>
(Fig.
<figureCitation id="7FF5A685929B7321D5B30CD1A7EA20DD" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. A-D Johora tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang E-H J. hoiseni Ng &amp; Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan A, E left G 1 (ventral view) B, F distal part of left G 1 (ventral view) C, G distal part of left G 1 (dorsal view) D, H left G 2 (ventral view) E-H after Ng and Takeda (1992: fig. 3 B, D-F). Scale bars: 1.0 mm (A, D, E, H); 0.5 mm (B, C, F, G)." figureDoi="10.3897/zookeys.994.56810.figure8" httpUri="https://binary.pensoft.net/fig/476319" pageId="0" pageNumber="1">8A-C</figureCitation>
). Both
<taxonomicName id="DE1768AEBF4A991AD70E62A12A9641A1" lsidName="J. hoiseni" pageId="0" pageNumber="1" rank="species" species="hoiseni">
<emphasis id="7CC72A280D83F408DEA4F673EA09CEE7" italics="true" pageId="0" pageNumber="1">J. hoiseni</emphasis>
</taxonomicName>
and
<taxonomicName id="7D0BC440C947788A16B50BC33FD4B320" lsidName="J. thaiana" pageId="0" pageNumber="1" rank="species" species="thaiana">
<emphasis id="9FC84231B64D2489D41C67AC895565EB" italics="true" pageId="0" pageNumber="1">J. thaiana</emphasis>
</taxonomicName>
have relatively straight G1 terminal segments which are prominently setose (Figs
<figureCitation id="F39E90428D21648A9ED54FC827A7B809" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. A-D Johora tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang E-H J. hoiseni Ng &amp; Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan A, E left G 1 (ventral view) B, F distal part of left G 1 (ventral view) C, G distal part of left G 1 (dorsal view) D, H left G 2 (ventral view) E-H after Ng and Takeda (1992: fig. 3 B, D-F). Scale bars: 1.0 mm (A, D, E, H); 0.5 mm (B, C, F, G)." figureDoi="10.3897/zookeys.994.56810.figure8" httpUri="https://binary.pensoft.net/fig/476319" pageId="0" pageNumber="1">8E-G</figureCitation>
,
<figureCitation id="F5F9E1F982393032A519E6B3465D3877" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. A-D Johora thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand E-I J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang A, E left G 1 (ventral view) B, F distal part of left G 1 (ventral view) C, H distal part of left G 1 (dorsal view) G distal part of left G 1 (mesial view) D, I left G 2 (ventral view). Scale bas: 1.0 mm (A, D, E, I); 0.5 mm (B, C, F-H)." figureDoi="10.3897/zookeys.994.56810.figure10" httpUri="https://binary.pensoft.net/fig/476321" pageId="0" pageNumber="1">10A-C</figureCitation>
).
</paragraph>
<paragraph id="A9D32373FD74DB9AD86306E92B63E929" pageId="0" pageNumber="1">
<taxonomicName id="AA100BD82E50E47BBEC2C2048CF0910F" authorityName="Ng" authorityYear="2020" class="Malacostraca" family="Potamidae" genus="Johora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Johora booliati" order="Decapoda" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="booliati">
<emphasis id="1782F7B189793456ACEAB408C90E5C7A" italics="true" pageId="0" pageNumber="1">Johora booliati</emphasis>
</taxonomicName>
and
<taxonomicName id="54ED3B6052D6773BADE546002817DCC6" lsidName="J. erici" pageId="0" pageNumber="1" rank="species" species="erici">
<emphasis id="024D3FAF5C0E8863D5EEB2B812F0D907" italics="true" pageId="0" pageNumber="1">J. erici</emphasis>
</taxonomicName>
can be distinguished from
<taxonomicName id="A181A3307967875B996803D1FEA56E50" lsidName="J. tiomanensis" pageId="0" pageNumber="1" rank="species" species="tiomanensis">
<emphasis id="7AD3E8EFD7F6580E2026B23891687F80" italics="true" pageId="0" pageNumber="1">J. tiomanensis</emphasis>
</taxonomicName>
and
<taxonomicName id="59149571DC73D55CAFB86115B96C208C" lsidName="J. counsilmani" pageId="0" pageNumber="1" rank="species" species="counsilmani">
<emphasis id="34FCE809D45FE9BA5CA6578C47A2CC20" italics="true" pageId="0" pageNumber="1">J. counsilmani</emphasis>
</taxonomicName>
by their carapace being relatively more quadrate in shape (width to length ratio less than 1.2) and the adult carapace is distinctly wider than long (width to length ratio 1.3-1.4) in
<taxonomicName id="265FE31A35377EF783A08F024E37CF78" lsidName="J. tiomanensis" pageId="0" pageNumber="1" rank="species" species="tiomanensis">
<emphasis id="974B080B104289540B218A34899CE28F" italics="true" pageId="0" pageNumber="1">J. tiomanensis</emphasis>
</taxonomicName>
and
<taxonomicName id="A00CEDDACBC4CF5B04628979C66CAA96" lsidName="J. counsilmani" pageId="0" pageNumber="1" rank="species" species="counsilmani">
<emphasis id="181CB82AB334DA5A10C22B007F90BE05" italics="true" pageId="0" pageNumber="1">J. counsilmani</emphasis>
</taxonomicName>
. In addition, the epigastric cristae of
<taxonomicName id="182B9B0E5407586F8AD0071BA2BE6048" lsidName="J. booliati" pageId="0" pageNumber="1" rank="species" species="booliati">
<emphasis id="BFDC9A73FB2C52D8E4099BE0A65E1A6D" italics="true" pageId="0" pageNumber="1">J. booliati</emphasis>
</taxonomicName>
and
<taxonomicName id="B0AEA7B317D7D5157B69DA575BD1A237" lsidName="J. erici" pageId="0" pageNumber="1" rank="species" species="erici">
<emphasis id="5045537D97505F6FAD9E98B8CCF99686" italics="true" pageId="0" pageNumber="1">J. erici</emphasis>
</taxonomicName>
are distinctly anterior to the postorbital cristae and separated by a gap with the entire postorbital cristae sharp (Figs
<figureCitation id="90931B1E2FBCB051DE33E1310ECD8F08" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dorsal view of carapace A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249) Terengganu D J. hoiseni Ng &amp; Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure2" httpUri="https://binary.pensoft.net/fig/476313" pageId="0" pageNumber="1">2F, G</figureCitation>
,
<figureCitation id="34E4DD56EDFB546DFD6E437D7FE782B1" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Johora erici sp. nov., male (41.2 x 33.8 mm) (ZRC 1995.0269), Kelantan A overall dorsal view B dorsal view of carapace C frontal view of cephalothorax D anterior thoracic sternum and pleon." figureDoi="10.3897/zookeys.994.56810.figure7" httpUri="https://binary.pensoft.net/fig/476318" pageId="0" pageNumber="1">7B</figureCitation>
) while in
<taxonomicName id="A1FE2A0C4470A04B441C640DACF83107" lsidName="J. tiomanensis" pageId="0" pageNumber="1" rank="species" species="tiomanensis">
<emphasis id="A120F252D58B816D2E9DCFF662AA6293" italics="true" pageId="0" pageNumber="1">J. tiomanensis</emphasis>
</taxonomicName>
and
<taxonomicName id="D8C6EE54DBDFBFC0364572C50D76EAB1" lsidName="J. counsilmani" pageId="0" pageNumber="1" rank="species" species="counsilmani">
<emphasis id="C68DE0839613F2D806EDF78AD60D9585" italics="true" pageId="0" pageNumber="1">J. counsilmani</emphasis>
</taxonomicName>
, the epigastric cristae are only slightly anterior of the postorbital cristae and separated only by striae (cf.
<bibRefCitation id="7BBB2B9EDF10518D4BB9583A39977AA3" author="Ng, PKL" journalOrPublisher="Malayan Nature Journal" pageId="0" pageNumber="1" pagination="167 - 174" refId="B20" refString="Ng, PKL, Tan, LWH, 1984. A new freshwater crab, Stoliczia (Johora) tiomanensis n. sp. (Decapoda: Brachyura: Potamidae) from Pulau Tioman, West Malaysia. Malayan Nature Journal 37 (3): 167 - 174" title="A new freshwater crab, Stoliczia (Johora) tiomanensis n. sp. (Decapoda: Brachyura: Potamidae) from Pulau Tioman, West Malaysia." volume="37" year="1984">Ng and Tan 1984</bibRefCitation>
: figs 1A, 2A;
<bibRefCitation id="AEC3B9B31BB91C5614B88DA7340531F1" author="Ng, PKL" journalOrPublisher="Zoologische Mededelingen" pageId="0" pageNumber="1" pagination="149 - 162" refId="B8" refString="Ng, PKL, 1985. Freshwater decapod crustaceans from Pulau Tioman, West Malaysia. Zoologische Mededelingen 59 (14): 149 - 162" title="Freshwater decapod crustaceans from Pulau Tioman, West Malaysia." volume="59" year="1985">Ng 1985</bibRefCitation>
: fig. 3). Most significantly, the G1 terminal segments of
<taxonomicName id="8822041A3433F1D62F570D45DBF2A153" lsidName="J. booliati" pageId="0" pageNumber="1" rank="species" species="booliati">
<emphasis id="D8D792FF19E1E1A59026FFB83BBE595B" italics="true" pageId="0" pageNumber="1">J. booliati</emphasis>
</taxonomicName>
and
<taxonomicName id="BB082DAE58BF89F757A1D74F51297C89" lsidName="J. erici" pageId="0" pageNumber="1" rank="species" species="erici">
<emphasis id="937B445E107FEC9D78FA2E628AC72788" italics="true" pageId="0" pageNumber="1">J. erici</emphasis>
</taxonomicName>
are proportionately stouter and less curved (Figs
<figureCitation id="BEF84A60C1125FDDAF75FD3863D7EEC6" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. A-D Johora thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand E-I J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang A, E left G 1 (ventral view) B, F distal part of left G 1 (ventral view) C, H distal part of left G 1 (dorsal view) G distal part of left G 1 (mesial view) D, I left G 2 (ventral view). Scale bas: 1.0 mm (A, D, E, I); 0.5 mm (B, C, F-H)." figureDoi="10.3897/zookeys.994.56810.figure10" httpUri="https://binary.pensoft.net/fig/476321" pageId="0" pageNumber="1">10E-H</figureCitation>
,
<figureCitation id="5836EF9ED8123FFE05C007833920160A" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. A-D Johora erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak E-H J. erici sp. nov., male (41.2 x 33.8 mm) (ZRC 1995.0269), Kelantan A, E left G 1 (ventral view) B, F distal part of left G 1 (ventral view) C, G distal part of left G 1 (dorsal view) D, H left G 2 (ventral view). Scale bars: 1.0 mm (A, D, E, H); 0.5 mm (B, C, F, G)." figureDoi="10.3897/zookeys.994.56810.figure11" httpUri="https://binary.pensoft.net/fig/476322" pageId="0" pageNumber="1">11A-C, E-G</figureCitation>
); in
<taxonomicName id="F11FD3F731E4B07DE01FCB20CF1B3A94" lsidName="J. tiomanensis" pageId="0" pageNumber="1" rank="species" species="tiomanensis">
<emphasis id="B9F88F1297C2006D1C11AE30B21D36C6" italics="true" pageId="0" pageNumber="1">J. tiomanensis</emphasis>
</taxonomicName>
and
<taxonomicName id="B36024168A28EC9996742DA5E47DC152" lsidName="J. counsilmani" pageId="0" pageNumber="1" rank="species" species="counsilmani">
<emphasis id="5A5161F8CC2C01C9C7674A84F700E709" italics="true" pageId="0" pageNumber="1">J. counsilmani</emphasis>
</taxonomicName>
, it is distinctly slenderer, more elongate and strongly curved (cf.
<bibRefCitation id="2903A4866F77ABF1B15392A3605A4520" author="Ng, PKL" journalOrPublisher="Malayan Nature Journal" pageId="0" pageNumber="1" pagination="167 - 174" refId="B20" refString="Ng, PKL, Tan, LWH, 1984. A new freshwater crab, Stoliczia (Johora) tiomanensis n. sp. (Decapoda: Brachyura: Potamidae) from Pulau Tioman, West Malaysia. Malayan Nature Journal 37 (3): 167 - 174" title="A new freshwater crab, Stoliczia (Johora) tiomanensis n. sp. (Decapoda: Brachyura: Potamidae) from Pulau Tioman, West Malaysia." volume="37" year="1984">Ng and Tan 1984</bibRefCitation>
: fig. 3a-c;
<bibRefCitation id="79C4AA51793D72E37373CCF4E258F999" author="Ng, PKL" journalOrPublisher="Zoologische Mededelingen" pageId="0" pageNumber="1" pagination="149 - 162" refId="B8" refString="Ng, PKL, 1985. Freshwater decapod crustaceans from Pulau Tioman, West Malaysia. Zoologische Mededelingen 59 (14): 149 - 162" title="Freshwater decapod crustaceans from Pulau Tioman, West Malaysia." volume="59" year="1985">Ng 1985</bibRefCitation>
: figs 2a, c, e, 4a-c).
<taxonomicName id="6E0BFC8CFB98859236C3C7D1625979F9" authorityName="Ng" authorityYear="2020" class="Malacostraca" family="Potamidae" genus="Johora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Johora booliati" order="Decapoda" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="booliati">
<emphasis id="A97AB802E69699EB68E2D02EFE692C57" italics="true" pageId="0" pageNumber="1">Johora booliati</emphasis>
</taxonomicName>
and
<taxonomicName id="86543197D727348A63AB5E89601C7086" lsidName="J. erici" pageId="0" pageNumber="1" rank="species" species="erici">
<emphasis id="94C548A7ACA75F1EE76273F5F05C8646" italics="true" pageId="0" pageNumber="1">J. erici</emphasis>
</taxonomicName>
differ from
<taxonomicName id="F9EF0D564C6DA63B844C4C3CF952D3DA" lsidName="J. thoi" pageId="0" pageNumber="1" rank="species" species="thoi">
<emphasis id="00DB1C39DDDF803E3B905ABE503ACFF3" italics="true" pageId="0" pageNumber="1">J. thoi</emphasis>
</taxonomicName>
in possessing slightly shorter ambulatory legs (length to width ratio of merus of fourth leg 2.7-2.8 versus 3.0-3.2), the lateral carapace surfaces are relatively more rugose, and the distinctly shorter and gently curved G1 terminal segment (Figs
<figureCitation id="6EA92A0A9D12BD63C69A33CD38093C70" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. A-D Johora thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand E-I J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang A, E left G 1 (ventral view) B, F distal part of left G 1 (ventral view) C, H distal part of left G 1 (dorsal view) G distal part of left G 1 (mesial view) D, I left G 2 (ventral view). Scale bas: 1.0 mm (A, D, E, I); 0.5 mm (B, C, F-H)." figureDoi="10.3897/zookeys.994.56810.figure10" httpUri="https://binary.pensoft.net/fig/476321" pageId="0" pageNumber="1">10E-H</figureCitation>
,
<figureCitation id="828F67FA772EBD654916709035D1DEB5" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. A-D Johora erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak E-H J. erici sp. nov., male (41.2 x 33.8 mm) (ZRC 1995.0269), Kelantan A, E left G 1 (ventral view) B, F distal part of left G 1 (ventral view) C, G distal part of left G 1 (dorsal view) D, H left G 2 (ventral view). Scale bars: 1.0 mm (A, D, E, H); 0.5 mm (B, C, F, G)." figureDoi="10.3897/zookeys.994.56810.figure11" httpUri="https://binary.pensoft.net/fig/476322" pageId="0" pageNumber="1">11A-C, E-G</figureCitation>
); in
<taxonomicName id="11A51A7BC4655020CABBC7F2FDE60658" lsidName="J. thoi" pageId="0" pageNumber="1" rank="species" species="thoi">
<emphasis id="39F8E9E677BB93A62D65D03650490F93" italics="true" pageId="0" pageNumber="1">J. thoi</emphasis>
</taxonomicName>
, this structure is diagnostic, being very elongate and straight (Fig.
<figureCitation id="8CF52D06EBA13E7745436B7AFADB13CF" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. A-D Johora thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249), Terengganu E-H J. thoi Ng, 1990, male (21.1 x 17.7 mm) (ZRC 2001.1167), Terengganu I-L J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2020.0361), Terengganu A, E, I left G 1 (ventral view) B, F, J distal part of left G 1 (ventral view) C, G, K distal part of left G 1 (dorsal view) D, H, L left G 2 (ventral view) A-D after Ng (1990: fig. 2 A, C-E). Scale bars: 1.0 mm (A, D, E, H, I, L); 0.5 mm (B, C, F, G, J, K)." figureDoi="10.3897/zookeys.994.56810.figure9" httpUri="https://binary.pensoft.net/fig/476320" pageId="0" pageNumber="1">9A-C, E-G</figureCitation>
).
</paragraph>
<paragraph id="AF1652BE3EA73E4730AD664C5B63FDDC" pageId="0" pageNumber="1">
The G1 structure of
<taxonomicName id="79908A2F313DC6167EE67264DC9914A7" lsidName="J. booliati" pageId="0" pageNumber="1" rank="species" species="booliati">
<emphasis id="57897F5EDFAA427ED20886ACF839F215" italics="true" pageId="0" pageNumber="1">J. booliati</emphasis>
</taxonomicName>
closely resembles that of
<taxonomicName id="EFA3D8743A4945CE3C24C415DC227406" lsidName="J. erici" pageId="0" pageNumber="1" rank="species" species="erici">
<emphasis id="058EBD395709900A6A69915EEC8D42B5" italics="true" pageId="0" pageNumber="1">J. erici</emphasis>
</taxonomicName>
, with the terminal segments similar in shape and proportions. The G1 subterminal segments of the two species, however, differ in form, with that of
<taxonomicName id="0E1819EE935CA2E9852CA5EF2DC9B340" lsidName="J. booliati" pageId="0" pageNumber="1" rank="species" species="booliati">
<emphasis id="64A6A06EECEBC3CC565130BABE89F059" italics="true" pageId="0" pageNumber="1">J. booliati</emphasis>
</taxonomicName>
forming a shelf-like structure along the outer margin (Fig.
<figureCitation id="0332A32C6C35604E12ABC84AC9955133" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. A-D Johora thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand E-I J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang A, E left G 1 (ventral view) B, F distal part of left G 1 (ventral view) C, H distal part of left G 1 (dorsal view) G distal part of left G 1 (mesial view) D, I left G 2 (ventral view). Scale bas: 1.0 mm (A, D, E, I); 0.5 mm (B, C, F-H)." figureDoi="10.3897/zookeys.994.56810.figure10" httpUri="https://binary.pensoft.net/fig/476321" pageId="0" pageNumber="1">10E</figureCitation>
) while in
<taxonomicName id="280EE73087F1D7A9518C169FCF6EFC42" lsidName="J. erici" pageId="0" pageNumber="1" rank="species" species="erici">
<emphasis id="AEC831CD8FAEDB65E361FFEDEBF13B26" italics="true" pageId="0" pageNumber="1">J. erici</emphasis>
</taxonomicName>
, the tapering of the segment is gradual, and no shelf is visible (Fig.
<figureCitation id="03068CE77C1241DED8A8241893E739B4" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. A-D Johora erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak E-H J. erici sp. nov., male (41.2 x 33.8 mm) (ZRC 1995.0269), Kelantan A, E left G 1 (ventral view) B, F distal part of left G 1 (ventral view) C, G distal part of left G 1 (dorsal view) D, H left G 2 (ventral view). Scale bars: 1.0 mm (A, D, E, H); 0.5 mm (B, C, F, G)." figureDoi="10.3897/zookeys.994.56810.figure11" httpUri="https://binary.pensoft.net/fig/476322" pageId="0" pageNumber="1">11A, E</figureCitation>
). The carapaces of the two species are different in adult males and females. In
<taxonomicName id="9E9E9C18B9EA6882D1D292E4C9FDEB76" lsidName="J. booliati" pageId="0" pageNumber="1" rank="species" species="booliati">
<emphasis id="D4C4278F1C3830E99DE482021AA2250B" italics="true" pageId="0" pageNumber="1">J. booliati</emphasis>
</taxonomicName>
, the carapace has a more quadrate form because the anterolateral margin is only gently convex, even in the largest male and the posterolateral margin is straighter and gently converging, with the posterior carapace margin medially indented (even in females) (Fig.
<figureCitation id="4685385D5A43C44BF455483088C45DCB" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dorsal view of carapace A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249) Terengganu D J. hoiseni Ng &amp; Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure2" httpUri="https://binary.pensoft.net/fig/476313" pageId="0" pageNumber="1">2F</figureCitation>
). In
<taxonomicName id="6963B892EAD74077BF2F432C706DD404" lsidName="J. erici" pageId="0" pageNumber="1" rank="species" species="erici">
<emphasis id="74832385BD1B4AEF7B3389EC73902500" italics="true" pageId="0" pageNumber="1">J. erici</emphasis>
</taxonomicName>
, the carapace appears more ovate, with the anterolateral margin distinctly convex, and the posterolateral margin is medially concave and strongly converging, and the posterior carapace margin is entire (Figs
<figureCitation id="7302C837C6481B3E89F2D6A2CC0CBD31" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dorsal view of carapace A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249) Terengganu D J. hoiseni Ng &amp; Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure2" httpUri="https://binary.pensoft.net/fig/476313" pageId="0" pageNumber="1">2G</figureCitation>
,
<figureCitation id="E0792EF62518F2BEB44AF07C145CDF03" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Johora erici sp. nov., male (41.2 x 33.8 mm) (ZRC 1995.0269), Kelantan A overall dorsal view B dorsal view of carapace C frontal view of cephalothorax D anterior thoracic sternum and pleon." figureDoi="10.3897/zookeys.994.56810.figure7" httpUri="https://binary.pensoft.net/fig/476318" pageId="0" pageNumber="1">7B</figureCitation>
).
</paragraph>
<paragraph id="DE24BB5592DC8E0EFF5665D0C315D884" pageId="0" pageNumber="1">
The G1 structure of
<taxonomicName id="E6BAEF00986406B1792B1EF708935E48" lsidName="J. booliati" pageId="0" pageNumber="1" rank="species" species="booliati">
<emphasis id="72A9BEA4EB1966970337AE25FA8AD32E" italics="true" pageId="0" pageNumber="1">J. booliati</emphasis>
</taxonomicName>
and
<taxonomicName id="82C369754B0C01AE96CFC71CA007E9C5" lsidName="J. erici" pageId="0" pageNumber="1" rank="species" species="erici">
<emphasis id="06A19AA862B5DE9FBA817080EE8F5F5A" italics="true" pageId="0" pageNumber="1">J. erici</emphasis>
</taxonomicName>
is superficially similar to that of
<taxonomicName id="7AAD2165B6118D89833DA21BA53C984C" lsidName="J. murphyi" pageId="0" pageNumber="1" rank="species" species="murphyi">
<emphasis id="A7438193394A2906614A3274F7183CD0" italics="true" pageId="0" pageNumber="1">J. murphyi</emphasis>
</taxonomicName>
from southern Peninsular Malaysia, notably in the shape of the G1 terminal and subterminal segments (
<bibRefCitation id="5CF24B698B22B22F1A14F83376F999B8" author="Ng, PKL" journalOrPublisher="Journal of the Singapore National Academy of Science" pageId="0" pageNumber="1" pagination="36 - 44" refId="B10" refString="Ng, PKL, 1986b. Preliminary descriptions of 17 new freshwater crabs of the genera Geosesarma, Parathelphusa, Johora and Stoliczia (Crustacea Decapoda, Brachyura) from South East Asia. Journal of the Singapore National Academy of Science 15: 36 - 44" title="Preliminary descriptions of 17 new freshwater crabs of the genera Geosesarma, Parathelphusa, Johora and Stoliczia (Crustacea Decapoda, Brachyura) from South East Asia." volume="15" year="1986 b">Ng 1986b</bibRefCitation>
: fig. 14a, b;
<bibRefCitation id="2172654569FE873F85DC53677E1CB28A" author="Ng, PKL" journalOrPublisher="Malayan Nature Journal" pageId="0" pageNumber="1" pagination="13 - 44" refId="B11" refString="Ng, PKL, 1987. A revision of the Malayan freshwater crabs of the genus Johora Bott, 1966 stat. nov. (Decapoda: Brachyura: Potamidae). Malayan Nature Journal 41: 13 - 44" title="A revision of the Malayan freshwater crabs of the genus Johora Bott, 1966 stat. nov. (Decapoda: Brachyura: Potamidae)." volume="41" year="1987">1987</bibRefCitation>
: fig. 6D, E;
<bibRefCitation id="D3F3BDAC3606F8E54A84A10F8D43C4A9" author="Ng, PKL" journalOrPublisher="Department of Zoology, National University of Singapore, Shinglee Press" pageId="0" pageNumber="1" refId="B12" refString="Ng, PKL, 1988. The Freshwater Crabs of Peninsular Malaysia and Singapore. Department of Zoology, National University of Singapore, Shinglee Press" title="The Freshwater Crabs of Peninsular Malaysia and Singapore." year="1988">1988</bibRefCitation>
: fig. 16D, E), but differs in having the distal part of the terminal segment more rounded and less sharply tapering (Figs
<figureCitation id="4D9F99AB4204C7A60D29F4FAE79C3CB7" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. A-D Johora thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand E-I J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang A, E left G 1 (ventral view) B, F distal part of left G 1 (ventral view) C, H distal part of left G 1 (dorsal view) G distal part of left G 1 (mesial view) D, I left G 2 (ventral view). Scale bas: 1.0 mm (A, D, E, I); 0.5 mm (B, C, F-H)." figureDoi="10.3897/zookeys.994.56810.figure10" httpUri="https://binary.pensoft.net/fig/476321" pageId="0" pageNumber="1">10E-H</figureCitation>
,
<figureCitation id="6F04C4D23D34382392A138B11382A050" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. A-D Johora erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak E-H J. erici sp. nov., male (41.2 x 33.8 mm) (ZRC 1995.0269), Kelantan A, E left G 1 (ventral view) B, F distal part of left G 1 (ventral view) C, G distal part of left G 1 (dorsal view) D, H left G 2 (ventral view). Scale bars: 1.0 mm (A, D, E, H); 0.5 mm (B, C, F, G)." figureDoi="10.3897/zookeys.994.56810.figure11" httpUri="https://binary.pensoft.net/fig/476322" pageId="0" pageNumber="1">11A-C, E-G</figureCitation>
). The G1 of
<taxonomicName id="39348FD46132C796FFF6D124314BF544" lsidName="J. booliati" pageId="0" pageNumber="1" rank="species" species="booliati">
<emphasis id="8D7A55A3B5A77F3C36B5E16B77D46310" italics="true" pageId="0" pageNumber="1">J. booliati</emphasis>
</taxonomicName>
is also similar to
<taxonomicName id="55A5A9559ECFBCE6E4239170C5468810" lsidName="J. intermedia" pageId="0" pageNumber="1" rank="species" species="intermedia">
<emphasis id="C18CF39C38B21A1CFD0EF921B7FE213C" italics="true" pageId="0" pageNumber="1">J. intermedia</emphasis>
</taxonomicName>
, which has a wide distribution in central Peninsular Malaysia. The G1 terminal segment of
<taxonomicName id="B136B640275514A4C2536D78E41488FA" lsidName="J. booliati" pageId="0" pageNumber="1" rank="species" species="booliati">
<emphasis id="DACE504A831DFA6DE30062A00AC83518" italics="true" pageId="0" pageNumber="1">J. booliati</emphasis>
</taxonomicName>
and
<taxonomicName id="DDB8DF5C644EC06B9FABFD7C16AA1F9F" lsidName="J. erici" pageId="0" pageNumber="1" rank="species" species="erici">
<emphasis id="C3BC5D3741CDB3703E1926B00E23D43A" italics="true" pageId="0" pageNumber="1">J. erici</emphasis>
</taxonomicName>
, however, is relatively stouter and less curved, and the subterminal segment narrows along the distal third to form a neck-like structure with the terminal segment (Figs
<figureCitation id="EC23DDCE781544DCACECE55D39E857AB" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. A-D Johora thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand E-I J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang A, E left G 1 (ventral view) B, F distal part of left G 1 (ventral view) C, H distal part of left G 1 (dorsal view) G distal part of left G 1 (mesial view) D, I left G 2 (ventral view). Scale bas: 1.0 mm (A, D, E, I); 0.5 mm (B, C, F-H)." figureDoi="10.3897/zookeys.994.56810.figure10" httpUri="https://binary.pensoft.net/fig/476321" pageId="0" pageNumber="1">10E-H</figureCitation>
,
<figureCitation id="740FB7D6111DD9A41209BD4DF77ADF6B" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. A-D Johora erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak E-H J. erici sp. nov., male (41.2 x 33.8 mm) (ZRC 1995.0269), Kelantan A, E left G 1 (ventral view) B, F distal part of left G 1 (ventral view) C, G distal part of left G 1 (dorsal view) D, H left G 2 (ventral view). Scale bars: 1.0 mm (A, D, E, H); 0.5 mm (B, C, F, G)." figureDoi="10.3897/zookeys.994.56810.figure11" httpUri="https://binary.pensoft.net/fig/476322" pageId="0" pageNumber="1">11A-C, E-G</figureCitation>
) (versus the G1 terminal segment is more curved and tapers to a slender tip with only the distalmost part of the subterminal segment distinctly narrowed to form a broad cleft in
<taxonomicName id="D3BD46648A4D041D3677B5BFD67CFB32" lsidName="J. intermedia" pageId="0" pageNumber="1" rank="species" species="intermedia">
<emphasis id="DB3FB9DD41A67A78BF69EF391FEA743A" italics="true" pageId="0" pageNumber="1">J. intermedia</emphasis>
</taxonomicName>
, cf.
<bibRefCitation id="8F474064054CCC0A9B050AD4C6EFEA3C" author="Ng, PKL" journalOrPublisher="Malayan Nature Journal" pageId="0" pageNumber="1" pagination="13 - 44" refId="B11" refString="Ng, PKL, 1987. A revision of the Malayan freshwater crabs of the genus Johora Bott, 1966 stat. nov. (Decapoda: Brachyura: Potamidae). Malayan Nature Journal 41: 13 - 44" title="A revision of the Malayan freshwater crabs of the genus Johora Bott, 1966 stat. nov. (Decapoda: Brachyura: Potamidae)." volume="41" year="1987">Ng 1987</bibRefCitation>
: figs 3A-D, G-J, 4A-D, F, G, I, J;
<bibRefCitation id="EABD4DE9102333A9CDC7E27F79DD05DF" author="Ng, PKL" journalOrPublisher="Department of Zoology, National University of Singapore, Shinglee Press" pageId="0" pageNumber="1" refId="B12" refString="Ng, PKL, 1988. The Freshwater Crabs of Peninsular Malaysia and Singapore. Department of Zoology, National University of Singapore, Shinglee Press" title="The Freshwater Crabs of Peninsular Malaysia and Singapore." year="1988">1988</bibRefCitation>
: fig. 15D-F;
<bibRefCitation id="C7FFF0CABD22B3AAF1E8256C67A61313" author="Ng, PKL" journalOrPublisher="Bulletin of the National Science Museum, Tokyo, Series A" pageId="0" pageNumber="1" pagination="103 - 116" refId="B19" refString="Ng, PKL, Takeda, M, 1992. On some freshwater crabs (Crustacea, Brachyura, Potamidae, Parathelphusidae and Grapsidae) from Peninsular Malaysia. Bulletin of the National Science Museum, Tokyo, Series A 18 (3): 103 - 116" title="On some freshwater crabs (Crustacea, Brachyura, Potamidae, Parathelphusidae and Grapsidae) from Peninsular Malaysia." volume="18" year="1992">Ng and Takeda 1992</bibRefCitation>
: fig. 1A-E). It is also noteworthy that both
<taxonomicName id="AF803F26FF6981B3C91E32ADB2A09ADA" lsidName="J. murphyi" pageId="0" pageNumber="1" rank="species" species="murphyi">
<emphasis id="D7A830C3C0C1CC269D056D4AC472B30F" italics="true" pageId="0" pageNumber="1">J. murphyi</emphasis>
</taxonomicName>
and
<taxonomicName id="9E53A4F809A2CA504F3A08653387A769" lsidName="J. intermedia" pageId="0" pageNumber="1" rank="species" species="intermedia">
<emphasis id="E735F4D570B5DF5C28A4C24451A6F804" italics="true" pageId="0" pageNumber="1">J. intermedia</emphasis>
</taxonomicName>
are smaller species than
<taxonomicName id="A53ED1751FD9357A14FDD0D00FB2A487" lsidName="J. booliati" pageId="0" pageNumber="1" rank="species" species="booliati">
<emphasis id="62A251E3E75156E08FFEF54980FEF7A4" italics="true" pageId="0" pageNumber="1">J. booliati</emphasis>
</taxonomicName>
and
<taxonomicName id="78502BEB113910CDE5444808FA62497A" lsidName="J. erici" pageId="0" pageNumber="1" rank="species" species="erici">
<emphasis id="6BB0BF8B0C0BDC43D2539D416E8FDD6C" italics="true" pageId="0" pageNumber="1">J. erici</emphasis>
</taxonomicName>
, with adults not reaching carapace widths of 25 mm.
</paragraph>
<paragraph id="A39306FC0DF2C9EDA1B5E2915D69CE77" pageId="0" pageNumber="1">
<bibRefCitation id="FEA01B517E388E290440E68028B37899" author="Ng, PKL" journalOrPublisher="Bulletin of the National Science Museum, Tokyo, Series A" pageId="0" pageNumber="1" pagination="103 - 116" refId="B19" refString="Ng, PKL, Takeda, M, 1992. On some freshwater crabs (Crustacea, Brachyura, Potamidae, Parathelphusidae and Grapsidae) from Peninsular Malaysia. Bulletin of the National Science Museum, Tokyo, Series A 18 (3): 103 - 116" title="On some freshwater crabs (Crustacea, Brachyura, Potamidae, Parathelphusidae and Grapsidae) from Peninsular Malaysia." volume="18" year="1992">Ng and Takeda (1992</bibRefCitation>
: 107) had specimens from
<normalizedToken id="1A824EBAD2892C069BC67A893C1C4B99" originalValue="Fraser’s">Fraser's</normalizedToken>
Hill as well as adjacent areas (Sungei Gumut, Peretak, Selangor,
<geoCoordinate id="F09B90FF47A2AEC63E1560A3C1FD1BD6" degrees="3" direction="north" minutes="36" orientation="latitude" precision="1" seconds="53.3" value="3.6148055">3°36'53.3&quot;N</geoCoordinate>
,
<geoCoordinate id="AA1951F9277040DF7B36D99356BE04F0" degrees="101" direction="east" minutes="44" orientation="longitude" precision="1" seconds="40.4" value="101.74455">101°44'40.4&quot;E</geoCoordinate>
; Sungei Sum [probably Sungei Sum Sum], near Genting Highlands, Pahang,
<geoCoordinate id="54498DE0B73506FB90DDCC013D8D8B91" degrees="3" direction="north" minutes="20" orientation="latitude" precision="1" seconds="42.2" value="3.3450556">3°20'42.2&quot;N</geoCoordinate>
,
<geoCoordinate id="FA5D7CE2FE7457E7E64E414CA15546B9" degrees="101" direction="east" minutes="51" orientation="longitude" precision="1" seconds="12.0" value="101.85333">101°51'12.0&quot;E</geoCoordinate>
; Ulu Teranum, Teras, Pahang,
<geoCoordinate id="E130851FEEF51F44E617C923E1BE6D41" degrees="3" direction="north" minutes="44" orientation="latitude" precision="1" seconds="12.6" value="3.7368333">3°44'12.6&quot;N</geoCoordinate>
,
<geoCoordinate id="C5E42B9D6841D8E25B6432EB88CC79B2" degrees="101" direction="east" minutes="47" orientation="longitude" precision="1" seconds="29.5" value="101.79153">101°47'29.5&quot;E</geoCoordinate>
) which they identified as &quot;
<taxonomicName id="FC04A8963D08620F00C3834D5A5A30E9" lsidName="J. tahanensis" pageId="0" pageNumber="1" rank="species" species="tahanensis">
<emphasis id="DFCBDF34A3A932F1DA8F1711834520A8" italics="true" pageId="0" pageNumber="1">J. tahanensis</emphasis>
</taxonomicName>
&quot;. On the basis of geography, they are probably all
<taxonomicName id="29B96A53526F8C693BED4240DC5E171E" lsidName="J. booliati" pageId="0" pageNumber="1" rank="species" species="booliati">
<emphasis id="566BC9B172C2104E3556DE14F4EA773C" italics="true" pageId="0" pageNumber="1">J. booliati</emphasis>
</taxonomicName>
as defined here.
</paragraph>
</subSubSection>
<subSubSection id="C7905A5E762944749B5A716B564A0C1E" pageId="0" pageNumber="1" type="distribution">
<paragraph id="1EC9CE34BA2E8D38D72CAC7FE7CF7A76" pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph id="120F3C4AAE6C68EA95312E35D0192A0E" pageId="0" pageNumber="1">
<taxonomicName id="9F3EBA4303F844E1371D419B769335AC" authorityName="Ng" authorityYear="2020" class="Malacostraca" family="Potamidae" genus="Johora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Johora booliati" order="Decapoda" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="booliati">
<emphasis id="74F828522FF705F3B20A0AD38A0A1530" italics="true" pageId="0" pageNumber="1">Johora booliati</emphasis>
</taxonomicName>
sp. nov. is known from highland streams in the central highlands of Pahang (Genting Highlands, Bukit Tinggi and
<normalizedToken id="CA32527822EE2C178FEB6C4063DB140B" originalValue="Fraser’s">Fraser's</normalizedToken>
Hill) (Fig.
<figureCitation id="9DB242360031159D6E8023F50A2F06D2" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Distribution of species of the Johora tahanensis species group 1 J. tahanensis (Bott, 1966) 2 J. thoi Ng, 1990 3 J. hoiseni Ng &amp; Takeda, 1992 4 J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005 5 J. booliati sp. nov. 6 J. erici sp. nov. 7 J. michaeli sp. nov. Map data 2020 Google." figureDoi="10.3897/zookeys.994.56810.figure15" httpUri="https://binary.pensoft.net/fig/476326" pageId="0" pageNumber="1">15</figureCitation>
). The distribution of
<taxonomicName id="4C70F35F1D03529D8A755CD4560EA508" lsidName="J. booliati" pageId="0" pageNumber="1" rank="species" species="booliati">
<emphasis id="F1E518C83CCE71A0D5F4F3DAC11D563B" italics="true" pageId="0" pageNumber="1">J. booliati</emphasis>
</taxonomicName>
overlaps with those of
<taxonomicName id="C5E9E1CA9ECDD6579C565ADE9322C6AC" lsidName="J. intermedia" pageId="0" pageNumber="1" rank="species" species="intermedia">
<emphasis id="62D969A957D5718B7C24CD1F001B7A79" italics="true" pageId="0" pageNumber="1">J. intermedia</emphasis>
</taxonomicName>
and
<taxonomicName id="E4F861FDF3E7DCD231B17BC3A700A48B" lsidName="J. gapensis" pageId="0" pageNumber="1" rank="species" species="gapensis">
<emphasis id="CFD003E6BBCDD78D0DFC0E3ABC3ED0CF" italics="true" pageId="0" pageNumber="1">J. gapensis</emphasis>
</taxonomicName>
, and we can expect the taxa to be found together. One juvenile male of
<taxonomicName id="54F98E0ADB0283D9FB19956BFE227AE0" lsidName="J. booliati" pageId="0" pageNumber="1" rank="species" species="booliati">
<emphasis id="4157A1B8651790E38DEDC0DFB96379C3" italics="true" pageId="0" pageNumber="1">J. booliati</emphasis>
</taxonomicName>
(10.5
<normalizedToken id="D5EC641A9D299446EEAB9E555722D47F" originalValue="×">x</normalizedToken>
9.1 mm, ZRC 2020.0364) was in fact collected with two adult males of
<taxonomicName id="60C6BE91C3419B03156EF0E5E3C8BCA5" lsidName="J. intermedia" pageId="0" pageNumber="1" rank="species" species="intermedia">
<emphasis id="A052418018BB9C5F6FA575BF103E315C" italics="true" pageId="0" pageNumber="1">J. intermedia</emphasis>
</taxonomicName>
at
<normalizedToken id="572886945F28C87AFC9C94405B960ECE" originalValue="Fraser’s">Fraser's</normalizedToken>
Hill (ZRC 2016.003).
</paragraph>
</subSubSection>
<subSubSection id="B07167BBE4798AAC802A739465015180" pageId="0" pageNumber="1" type="conservation">
<paragraph id="026B58F183862A391B3689B723E54036" pageId="0" pageNumber="1">Conservation.</paragraph>
<paragraph id="97AE1E0214FF4DAE1E040C1F999C2E83" pageId="0" pageNumber="1">
The conservation status for
<taxonomicName id="CFAC41F26BCB478CE2F2D736BB05AD58" lsidName="J. booliati" pageId="0" pageNumber="1" rank="species" species="booliati">
<emphasis id="18D2E9A4A45282301C94A2E23157E36D" italics="true" pageId="0" pageNumber="1">J. booliati</emphasis>
</taxonomicName>
is not known as its actual distribution is not known. For the moment, it is known only from a 30 km stretch of hills along the Central Highlands of Peninsular Malaysia, in the area of
<normalizedToken id="3D6C13F49CF818865A28E0050315A61E" originalValue="Fraser’s">Fraser's</normalizedToken>
Hill, Genting Highlands and Bukit Tinggi. This area is not protected, and in this context, the species should be categorised as vulnerable for the time being (see
<bibRefCitation id="94D323C3EDFCF8980F28CD74320777C7" DOI="https://doi.org/10.1016/j.biocon.2009.02.038" author="Cumberlidge, N" journalOrPublisher="Biological Conservation" pageId="0" pageNumber="1" pagination="1665 - 1673" refId="B3" refString="Cumberlidge, N, Ng, PKL, Yeo, DCJ, Magalhaes, C, Campos, MR, Alvarez, F, Naruse, T, Daniels, SR, Esser, LJ, Attipoe, FYK, Clotilde-Ba, F-L, Darwall, W, Mcivor, A, Collen, B, Ram, M, 2009. Freshwater crabs and the biodiversity crisis: importance, threats, status, and conservation challenges. Biological Conservation 142: 1665 - 1673, DOI: https://doi.org/10.1016/j.biocon.2009.02.038" title="Freshwater crabs and the biodiversity crisis: importance, threats, status, and conservation challenges." url="https://doi.org/10.1016/j.biocon.2009.02.038" volume="142" year="2009">Cumberlidge et al. 2009</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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