252 lines
30 KiB
XML
252 lines
30 KiB
XML
<document id="299732A590471AE6B68D866F5762BC8C" ID-DOI="10.15298/rusentj.30.4.06" ID-ISSN="0132-8069" ID-Zenodo-Dep="13178304" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722593993438" checkinUser="felipe" docAuthor="Serrano, J." docDate="2021" docId="25357A69FFBD9F23FC32F9BE0A275798" docLanguage="en" docName="RussEntomolJ.30.4.430-447.pdf" docOrigin="Russian Entomological Journal 30 (4)" docSource="http://dx.doi.org/10.15298/rusentj.30.4.06" docStyle="DocumentStyle:0D5BAC5AA0C130B3809AF578A601D837.3:RussEntomolJ.2019-.journal_article" docStyleId="0D5BAC5AA0C130B3809AF578A601D837" docStyleName="RussEntomolJ.2019-.journal_article" docStyleVersion="3" docTitle="Orthomus lacouri" docType="treatment" docVersion="2" lastPageNumber="441" masterDocId="D90C0211FFB49F28FFA5FF850B595403" masterDocTitle="The genus Orthomus Chaudoir, 1838 in the Iberian Peninsula and Morocco" masterLastPageNumber="447" masterPageNumber="430" pageNumber="439" updateTime="1722633854025" updateUser="ExternalLinkService">
|
||
<mods:mods id="93509AAD748EB6F5B5DC763C96429E25" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="CD7854FD2EF0A8F401C2F5815128AABA">
|
||
<mods:title id="16016A3AA9C389DD1379C0C1D12DDC82">The genus Orthomus Chaudoir, 1838 in the Iberian Peninsula and Morocco</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="73C65DB5E93D35D1E505655C8BAD7BE4" type="personal">
|
||
<mods:role id="95C2D337B08C0DFE29ADD50692555D46">
|
||
<mods:roleTerm id="3AFEA2F0F898FD2792DEE68268EBCCF9">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="05B9FF89DA3C37FD5B2F405FDEE13C00">Serrano, J.</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="9CA1EDF12823DFCF157539C988EE81F1">text</mods:typeOfResource>
|
||
<mods:relatedItem id="C5341165969C433DCA1FF4B2FD373121" type="host">
|
||
<mods:titleInfo id="B290B9BDFA2175F4C14753E202BC0BD8">
|
||
<mods:title id="1C4E3511D75B5DA5F42476B67B1B2250">Russian Entomological Journal</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="D133A13339C81FBA04E41394EBE182A7">
|
||
<mods:date id="61D0DC9488771078C804616A32EDAD12">2021</mods:date>
|
||
<mods:detail id="2F5CC195DA69E18CDEAF3FB01FA62FF8" type="pubDate">
|
||
<mods:number id="A35D8A591E7DD3459DC9F3C4610DE8C3">2021-12-31</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="94F4168C450F18EA819F21B076E26789" type="volume">
|
||
<mods:number id="E8DBCFCC7BAB75E8BA659D8A1A65DCE4">30</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="49671497757056DBF2065216E176CA95" type="issue">
|
||
<mods:number id="6C2664E6AD054E9F47DDC065B51268C3">4</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="BADB06F3874FECECA3628B73126E5302" unit="page">
|
||
<mods:start id="5E047DAE403A5936EAF41A29F438996A">430</mods:start>
|
||
<mods:end id="9720F5F8953CF9C36027992AAFBDC362">447</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:location id="115DC2AC337DB4415F5F8D5107F3AAF2">
|
||
<mods:url id="4F711AE909B802450493B6E136AA58C4">http://dx.doi.org/10.15298/rusentj.30.4.06</mods:url>
|
||
</mods:location>
|
||
<mods:classification id="A5B99156C03D2973C37A3CA8B26F321A">journal article</mods:classification>
|
||
<mods:identifier id="3FEE2633C3BD6455D58AF5AF05D3DB62" type="DOI">10.15298/rusentj.30.4.06</mods:identifier>
|
||
<mods:identifier id="89FF696E81B9ADFF7C822F3123272B3F" type="ISSN">0132-8069</mods:identifier>
|
||
<mods:identifier id="F3E3DD29AFB030217C6E7BF33DE26B1B" type="Zenodo-Dep">13178304</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="25357A69FFBD9F23FC32F9BE0A275798" LSID="urn:lsid:plazi:treatment:25357A69FFBD9F23FC32F9BE0A275798" httpUri="http://treatment.plazi.org/id/25357A69FFBD9F23FC32F9BE0A275798" lastPageId="11" lastPageNumber="441" pageId="9" pageNumber="439">
|
||
<subSubSection id="E58698F4FFBD9F21FC32F9BE0FCB5273" pageId="9" pageNumber="439" type="nomenclature">
|
||
<paragraph id="AD23CB7FFFBD9F21FC32F9BE0E525250" blockId="9.[812,1399,1595,2057]" box="[919,1291,1595,1619]" pageId="9" pageNumber="439">
|
||
<heading id="F66B7C13FFBD9F21FC32F9BE0E525250" box="[919,1291,1595,1619]" centered="true" fontSize="10" level="2" pageId="9" pageNumber="439" reason="8">
|
||
<taxonomicName id="6A9CB0FCFFBD9F21FC32F9BE0E525250" ID-CoL="6T92C" authority="(Antoine, 1941)" baseAuthorityName="Antoine" baseAuthorityYear="1941" box="[919,1291,1595,1619]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="439" phylum="Arthropoda" rank="species" species="lacouri">
|
||
<emphasis id="9FE8176DFFBD9F21FC32F9BE0F0E5250" box="[919,1111,1595,1619]" italics="true" pageId="9" pageNumber="439">Orthomus lacouri</emphasis>
|
||
(Antoine, 1941)
|
||
</taxonomicName>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="AD23CB7FFFBD9F21FBB7F9DC0FCB5273" blockId="9.[812,1399,1595,2057]" box="[1042,1170,1624,1649]" pageId="9" pageNumber="439">Figs 56–64.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="E58698F4FFBD9F23FCF7F9F80A275798" lastPageId="11" lastPageNumber="441" pageId="9" pageNumber="439" type="description">
|
||
<paragraph id="AD23CB7FFFBD9F21FCF7F9F80F3B5C0A" blockId="9.[812,1399,1595,2057]" pageId="9" pageNumber="439">
|
||
According to
|
||
<bibRefCitation id="C90DB68EFFBD9F21FC41F9F80FAD5290" author="Pupier R. & Coulon J." box="[996,1268,1661,1683]" pageId="9" pageNumber="439" refId="ref17156" refString="Pupier R., Coulon J. 2013. Etude des organes genitaux des femelles et considerations sur la taxonomie de deux groupes d'Orthomus" type="book" year="2013">Pupier and Coulon [2013]</bibRefCitation>
|
||
this species includes different geographic races that are distinguished by morphological features, and thus deserve their ranking as subspecies. Three of them are found in NE
|
||
<collectingCountry id="D58B8BEFFFBD9F21FB49F94B0E1F52E0" box="[1260,1350,1742,1763]" name="Morocco" pageId="9" pageNumber="439">Morocco</collectingCountry>
|
||
,
|
||
<taxonomicName id="6A9CB0FCFFBD9F21FAF6F94B0FEC52FE" authority="(Antoine, 1941)" baseAuthorityName="Antoine" baseAuthorityYear="1941" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="439" phylum="Arthropoda" rank="subSpecies" species="lacouri" subSpecies="lacouri">
|
||
<emphasis id="9FE8176DFFBD9F21FAF6F94B0F5752FE" italics="true" pageId="9" pageNumber="439">Orthomus lacouri lacouri</emphasis>
|
||
(Antoine, 1941)
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="6A9CB0FCFFBD9F21FB64F96C0F57531B" authority="Pupier et Coulon, 2013" authorityName="Pupier et Coulon" authorityYear="2013" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="439" phylum="Arthropoda" rank="subSpecies" species="lacouri" subSpecies="haroldi">
|
||
<emphasis id="9FE8176DFFBD9F21FB64F96C0E2F52FE" box="[1217,1398,1768,1790]" italics="true" pageId="9" pageNumber="439">O. lacouri haroldi</emphasis>
|
||
<bibRefCitation id="C90DB68EFFBD9F21FC89F8860F57531B" author="Pupier R. & Coulon J." box="[812,1038,1795,1817]" pageId="9" pageNumber="439" refId="ref17156" refString="Pupier R., Coulon J. 2013. Etude des organes genitaux des femelles et considerations sur la taxonomie de deux groupes d'Orthomus" type="book" year="2013">Pupier et Coulon, 2013</bibRefCitation>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="6A9CB0FCFFBD9F21FBE5F8860E2A531A" authority="Mateu, 1955" authorityName="Mateu" authorityYear="1955" box="[1088,1395,1795,1817]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="439" phylum="Arthropoda" rank="subSpecies" species="lacouri" subSpecies="kocheri">
|
||
<emphasis id="9FE8176DFFBD9F21FBE5F8860FA9531B" box="[1088,1264,1795,1816]" italics="true" pageId="9" pageNumber="439">O. lacouri kocheri</emphasis>
|
||
<bibRefCitation id="C90DB68EFFBD9F21FB52F8860E2A531A" author="Mateu J." box="[1271,1395,1795,1817]" pageId="9" pageNumber="439" pagination="53 - 85" refId="ref17020" refString="Mateu J. 1955. Notas sobre los Orthomus Chaudoir (segunda nota) // Eos. Vol. 31. P. 53 - 85." type="book chapter" year="1955">Mateu, 1955</bibRefCitation>
|
||
</taxonomicName>
|
||
. The fourth subspecies,
|
||
<taxonomicName id="6A9CB0FCFFBD9F21FBB3F89B0E005330" authority="Jeanne, 1988" authorityName="Jeanne" authorityYear="1988" box="[1046,1369,1821,1843]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="439" phylum="Arthropoda" rank="subSpecies" species="lacouri" subSpecies="pupieri">
|
||
<emphasis id="9FE8176DFFBD9F21FBB3F89B0F955330" box="[1046,1228,1821,1843]" italics="true" pageId="9" pageNumber="439">O. lacouri pupieri</emphasis>
|
||
<bibRefCitation id="C90DB68EFFBD9F21FB70F89B0E005330" author="Jeanne C." box="[1237,1369,1822,1843]" pageId="9" pageNumber="439" pagination="69 - 87" refId="ref16833" refString="Jeanne C. 1988. Carabiques noveaux ou remarquables (9 eme note) // Bull. Soc. linn. Bordeaux. Vol. 16. P. 69 - 87." type="journal article" year="1988">Jeanne, 1988</bibRefCitation>
|
||
</taxonomicName>
|
||
, is proper of NW
|
||
<collectingCountry id="D58B8BEFFFBD9F21FC67F8BD0F57534D" box="[962,1038,1848,1870]" name="Algeria" pageId="9" pageNumber="439">Algeria</collectingCountry>
|
||
. Here I follow the taxa descriptions published by
|
||
<bibRefCitation id="C90DB68EFFBD9F21FC15F8D60FF7536B" author="Pupier R. & Coulon J." box="[944,1198,1875,1897]" pageId="9" pageNumber="439" refId="ref17156" refString="Pupier R., Coulon J. 2013. Etude des organes genitaux des femelles et considerations sur la taxonomie de deux groupes d'Orthomus" type="book" year="2013">Pupier and Coulon [2013]</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="C90DB68EFFBD9F21FB7BF8D608365380" author="Chavanon G. & Mabrouki Y. & Taybi F. A." pageId="9" pageNumber="439" pagination="173 - 186" refId="ref16408" refString="Chavanon G., Mabrouki Y., Taybi F. A. 2016. Sur la repartition des populations d' Orthomus lacouri (Antoine, 1941) (Coleoptera, Carabidae, Pterostichinae) dans les Beni Snassen (Maroc Oriental) // Bull. mens. Soc. linn. Lyon. Vol. 85. Nos 5 - 6. P. 173 - 186." type="book chapter" year="2016">Chavanon et al. [2016]</bibRefCitation>
|
||
, together with personal observations of material of this species deposited in the MNCN Madrid and DZAFUM. Male genitalia are the key characters that distinguish the subspecies, which share in common most external characters. Thus, a general description follows and peculiarities of the subspecies are indicated below.
|
||
</paragraph>
|
||
<paragraph id="AD23CB7FFFBE9F22FF47FEB9090255BF" blockId="10.[189,776,315,657]" pageId="10" pageNumber="440">
|
||
Dorsum from black to piceous brown, appendages brown. Body length 10.5–11.3 mm. Eye of moderate size and convex, tempora oblique. Males shiny, females shiny or somewhat dull with visible microreticulation (
|
||
<emphasis id="9FE8176DFFBE9F22FDE8FE09099F55A3" box="[589,710,395,417]" italics="true" pageId="10" pageNumber="440">
|
||
O. l.
|
||
<taxonomicName id="6A9CB0FCFFBE9F22FD25FE0E099F55A3" baseAuthorityName="Antoine" baseAuthorityYear="1941" box="[640,710,395,416]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="440" phylum="Arthropoda" rank="species" species="lacouri">lacouri</taxonomicName>
|
||
</emphasis>
|
||
); both sexes of
|
||
<emphasis id="9FE8176DFFBE9F22FEB6FE230AD455B8" box="[275,397,422,443]" italics="true" pageId="10" pageNumber="440">O. l. kocheri</emphasis>
|
||
show a dull dorsum.
|
||
</paragraph>
|
||
<paragraph id="AD23CB7FFFBE9F22FF47FE4409B05692" blockId="10.[189,776,315,657]" pageId="10" pageNumber="440">
|
||
Pronotum shape trapezoid, anterior angle briefly protruding ahead except for
|
||
<taxonomicName id="6A9CB0FCFFBE9F22FE21FE59096855F3" authorityName="Pupier et Coulon" authorityYear="2013" box="[388,561,475,497]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="440" phylum="Arthropoda" rank="subSpecies" species="lacouri" subSpecies="haroldi">
|
||
<emphasis id="9FE8176DFFBE9F22FE21FE59096855F3" box="[388,561,475,497]" italics="true" pageId="10" pageNumber="440">O. lacouri haroldi</emphasis>
|
||
</taxonomicName>
|
||
, anterior basis moderately concave; side arcuate, more narrowed forward, slightly narrowed backward and showing maximum width near posterior basis (Fig. 56); posterior angle slightly obtuse and without tooth; basis no or slightly (
|
||
<emphasis id="9FE8176DFFBE9F22FE6BFDC309125658" box="[462,587,582,603]" italics="true" pageId="10" pageNumber="440">O. l. haroldi</emphasis>
|
||
) emarginate; inner fovea linear and marked, outer fovea poorly impressed; basis with points varying between scarce and moderately dense.
|
||
</paragraph>
|
||
<paragraph id="AD23CB7FFFBE9F22FCF7FEBE0E785584" blockId="10.[812,1399,315,657]" pageId="10" pageNumber="440">Elytra parallel, humeral tooth vestigial, striae punctured with variable intensity; discal pores on stria 3; 8th interval almost vertically tumbling down to elytron margin.</paragraph>
|
||
<paragraph id="AD23CB7FFFBE9F22FCF7FE0E0E16560F" blockId="10.[812,1399,315,657]" pageId="10" pageNumber="440">
|
||
Prosternal process truncate; metepisterna long, smooth and narrowed backward (Fig. 57); ventrites almost smooth, with fine punctures except for
|
||
<taxonomicName id="6A9CB0FCFFBE9F22FBF6FE440E5A55D5" authorityName="Mateu" authorityYear="1955" box="[1107,1283,449,470]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="440" phylum="Arthropoda" rank="subSpecies" species="lacouri" subSpecies="kocheri">
|
||
<emphasis id="9FE8176DFFBE9F22FBF6FE440E5A55D5" box="[1107,1283,449,470]" italics="true" pageId="10" pageNumber="440">O. lacouri kocheri</emphasis>
|
||
</taxonomicName>
|
||
, that shows well-marked punctures; male mesotibia distally dilated, metatibia with marked crenulation in inner side of distal half.
|
||
</paragraph>
|
||
<paragraph id="AD23CB7FFFBE9F22FCF7FD940F275658" blockId="10.[812,1399,315,657]" pageId="10" pageNumber="440">
|
||
Female genitalia (Fig. 58) with a short spermatic conduct followed by a progressively widened spermatheca of moderate length [
|
||
<bibRefCitation id="C90DB68EFFBE9F22FC3FFDC30F2A5658" author="Pupier R. & Coulon J." box="[922,1139,582,604]" pageId="10" pageNumber="440" refId="ref17156" refString="Pupier R., Coulon J. 2013. Etude des organes genitaux des femelles et considerations sur la taxonomie de deux groupes d'Orthomus" type="book" year="2013">Pupier, Coulon, 2013</bibRefCitation>
|
||
].
|
||
</paragraph>
|
||
<paragraph id="AD23CB7FFFBE9F22FCF7FDE40E2F5692" blockId="10.[812,1399,315,657]" pageId="10" pageNumber="440">
|
||
According to the detailed study of
|
||
<bibRefCitation id="C90DB68EFFBE9F22FB6AFDE408295692" author="Chavanon G. & Mabrouki Y. & Taybi F. A." pageId="10" pageNumber="440" pagination="173 - 186" refId="ref16408" refString="Chavanon G., Mabrouki Y., Taybi F. A. 2016. Sur la repartition des populations d' Orthomus lacouri (Antoine, 1941) (Coleoptera, Carabidae, Pterostichinae) dans les Beni Snassen (Maroc Oriental) // Bull. mens. Soc. linn. Lyon. Vol. 85. Nos 5 - 6. P. 173 - 186." type="book chapter" year="2016">Chavanon et al. [2016]</bibRefCitation>
|
||
, each subspecies has a “typical” shape of the median
|
||
</paragraph>
|
||
<paragraph id="AD23CB7FFFBE9F22FE93F9420E1952E2" blockId="10.[310,649,1735,1761]" box="[310,1344,1735,1761]" lastBlockId="10.[862,1344,1735,1761]" pageId="10" pageNumber="440">
|
||
<emphasis id="9FE8176DFFBE9F22FE93F9420E1952E2" bold="true" box="[310,1344,1735,1761]" pageId="10" pageNumber="440">57 56 62 63 64 58</emphasis>
|
||
</paragraph>
|
||
<paragraph id="AD23CB7FFFBE9F22FF47F9740E03537F" blockId="10.[189,1399,1777,2064]" pageId="10" pageNumber="440">
|
||
Figs 51–64.
|
||
<taxonomicName id="6A9CB0FCFFBE9F22FEEAF9740A955306" box="[335,460,1777,1797]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="440" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="9FE8176DFFBE9F22FEEAF9740AFC5307" box="[335,421,1777,1796]" italics="true" pageId="10" pageNumber="440">Orthomus</emphasis>
|
||
spp
|
||
</taxonomicName>
|
||
: 51–55 —
|
||
<taxonomicName id="6A9CB0FCFFBE9F22FD88F97409C65307" baseAuthorityName="Lucas" baseAuthorityYear="1846" box="[557,671,1777,1796]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="440" phylum="Arthropoda" rank="species" species="abacoides">
|
||
<emphasis id="9FE8176DFFBE9F22FD88F97409C65307" box="[557,671,1777,1796]" italics="true" pageId="10" pageNumber="440">O. abacoides</emphasis>
|
||
</taxonomicName>
|
||
; 56–64 —
|
||
<taxonomicName id="6A9CB0FCFFBE9F22FCA6F97408055307" baseAuthorityName="Antoine" baseAuthorityYear="1941" box="[771,860,1777,1796]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="440" phylum="Arthropoda" rank="species" species="lacouri">
|
||
<emphasis id="9FE8176DFFBE9F22FCA6F97408055307" box="[771,860,1777,1796]" italics="true" pageId="10" pageNumber="440">O. lacouri</emphasis>
|
||
</taxonomicName>
|
||
(56–57, 63 —
|
||
<emphasis id="9FE8176DFFBE9F22FC46F9740F095307" box="[995,1104,1777,1796]" italics="true" pageId="10" pageNumber="440">O. l. kocheri</emphasis>
|
||
(56 — female); 58–60, 62, 64 —
|
||
<emphasis id="9FE8176DFFBE9F22FF18F88C0A73531F" box="[189,298,1801,1820]" italics="true" pageId="10" pageNumber="440">O. l. haroldi</emphasis>
|
||
; 61 —
|
||
<emphasis id="9FE8176DFFBE9F22FECAF88C0A82531F" box="[367,475,1801,1820]" italics="true" pageId="10" pageNumber="440">O. l. pupieri</emphasis>
|
||
); 51, 56 — habitus; 57 — metepisterna; 52 — punctures of abdominal segments; 53, 59 — median lobe of aedeagus; 54, 61–64 — apex of median lobe of aedeagus; 55, 58 — genitalia of female; 51, 54, 56, 61–64 — dorsal view; 57–58, 60 — right lateral view; 53, 59 — left lateral view; 52 — ventral view; 51 — reservoir El Fakia,
|
||
<collectingCountry id="D58B8BEFFFBE9F22FC7CF8BC0F44534F" box="[985,1053,1849,1868]" name="Algeria" pageId="10" pageNumber="440">Algerie</collectingCountry>
|
||
; 55, 61 — Sidi Daho ez Zahir,
|
||
<collectingCountry id="D58B8BEFFFBE9F22FA95F8BC0E2D534F" box="[1328,1396,1849,1868]" name="Algeria" pageId="10" pageNumber="440">Algerie</collectingCountry>
|
||
; 58 — Jebel Tikermine,
|
||
<collectingCountry id="D58B8BEFFFBE9F22FE2FF8D70A825366" box="[394,475,1874,1893]" name="Morocco" pageId="10" pageNumber="440">Morocco</collectingCountry>
|
||
; 62 — Tannezatte,
|
||
<collectingCountry id="D58B8BEFFFBE9F22FD21F8D7098C5366" box="[644,725,1874,1893]" name="Morocco" pageId="10" pageNumber="440">Morocco</collectingCountry>
|
||
; 63 — Farkhana, Nador,
|
||
<collectingCountry id="D58B8BEFFFBE9F22FC14F8D70F5B5366" box="[945,1026,1874,1893]" name="Morocco" pageId="10" pageNumber="440">Morocco</collectingCountry>
|
||
; 64 — Sidi Bouhouria, Berkane,
|
||
<collectingCountry id="D58B8BEFFFBE9F22FA86F8D70E2D5366" box="[1315,1396,1874,1893]" name="Morocco" pageId="10" pageNumber="440">Morocco</collectingCountry>
|
||
; 53–55, 58–61 — after
|
||
<bibRefCitation id="C90DB68EFFBE9F22FE23F8EC0915537F" author="Pupier R. & Coulon J." box="[390,588,1897,1916]" pageId="10" pageNumber="440" refId="ref17156" refString="Pupier R., Coulon J. 2013. Etude des organes genitaux des femelles et considerations sur la taxonomie de deux groupes d'Orthomus" type="book" year="2013">Pupier, Coulon [2013]</bibRefCitation>
|
||
; 62–64 — after
|
||
<bibRefCitation id="C90DB68EFFBE9F22FD7BF8EC08F3537F" author="Chavanon G. & Mabrouki Y. & Taybi F. A." box="[734,938,1897,1917]" pageId="10" pageNumber="440" pagination="173 - 186" refId="ref16408" refString="Chavanon G., Mabrouki Y., Taybi F. A. 2016. Sur la repartition des populations d' Orthomus lacouri (Antoine, 1941) (Coleoptera, Carabidae, Pterostichinae) dans les Beni Snassen (Maroc Oriental) // Bull. mens. Soc. linn. Lyon. Vol. 85. Nos 5 - 6. P. 173 - 186." type="book chapter" year="2016">Chavanon et al. [2016]</bibRefCitation>
|
||
. Scale bars: 1.5 mm (Fig. 51); 1.6 mm (Fig. 56).
|
||
</paragraph>
|
||
<paragraph id="AD23CB7FFFBE9F22FF47F8000E695C13" blockId="10.[189,1399,1777,2064]" pageId="10" pageNumber="440">
|
||
Figs 51–64.
|
||
<taxonomicName id="6A9CB0FCFFBE9F22FEE8F8000A9E539B" box="[333,455,1925,1944]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="440" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="9FE8176DFFBE9F22FEE8F8000AF8539B" box="[333,417,1925,1944]" italics="true" pageId="10" pageNumber="440">Orthomus</emphasis>
|
||
spp
|
||
</taxonomicName>
|
||
: 51–55 —
|
||
<taxonomicName id="6A9CB0FCFFBE9F22FD82F80009CE539B" baseAuthorityName="Lucas" baseAuthorityYear="1846" box="[551,663,1925,1944]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="440" phylum="Arthropoda" rank="species" species="abacoides">
|
||
<emphasis id="9FE8176DFFBE9F22FD82F80009CE539B" box="[551,663,1925,1944]" italics="true" pageId="10" pageNumber="440">O. abacoides</emphasis>
|
||
</taxonomicName>
|
||
; 56–64 —
|
||
<taxonomicName id="6A9CB0FCFFBE9F22FD5CF8000808539B" baseAuthorityName="Antoine" baseAuthorityYear="1941" box="[761,849,1925,1944]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="440" phylum="Arthropoda" rank="species" species="lacouri">
|
||
<emphasis id="9FE8176DFFBE9F22FD5CF8000808539B" box="[761,849,1925,1944]" italics="true" pageId="10" pageNumber="440">O. lacouri</emphasis>
|
||
</taxonomicName>
|
||
(56–57, 63 —
|
||
<emphasis id="9FE8176DFFBE9F22FC70F8000F66539B" box="[981,1087,1925,1944]" italics="true" pageId="10" pageNumber="440">O. l. kocheri</emphasis>
|
||
(56 — самка); 58–60, 62, 64 —
|
||
<emphasis id="9FE8176DFFBE9F22FAC7F8000A5353B3" italics="true" pageId="10" pageNumber="440">O. l. haroldi</emphasis>
|
||
; 61 —
|
||
<emphasis id="9FE8176DFFBE9F22FEE3F8180AF253B3" box="[326,427,1949,1968]" italics="true" pageId="10" pageNumber="440">O. l. pupieri</emphasis>
|
||
); 51, 56 — внеШний вид; 57 — Эпистерн Заднегруди; 52 — пунктировка сегментов брюШка; 53, 59 — среднЯЯ долЯ Эдеагуса; 54, 61–64 — верШина средней доли Эдеагуса; 55, 58 — гениталии самки; 51, 54, 56, 61–64 — сверху; 57–58, 60 — сбоку справа; 53, 59 — сбоку слева; 52 — сниЗу; 51 — водохранилиЩе Эль- ФакиЯ,
|
||
<collectingCountry id="D58B8BEFFFBE9F22FC35F848089653E3" box="[912,975,1997,2016]" name="Algeria" pageId="10" pageNumber="440">Алжир</collectingCountry>
|
||
; 55, 61 — Сиди Дахо ЭЗ Захир,
|
||
<collectingCountry id="D58B8BEFFFBE9F22FB5CF8480E6153E3" box="[1273,1336,1997,2016]" name="Algeria" pageId="10" pageNumber="440">Алжир</collectingCountry>
|
||
; 58 — Джебель Тикермин,
|
||
<collectingCountry id="D58B8BEFFFBE9F22FEDFF8600A9553FB" box="[378,460,2021,2040]" name="Morocco" pageId="10" pageNumber="440">Марокко</collectingCountry>
|
||
; 62 — ТаннеЗатт,
|
||
<collectingCountry id="D58B8BEFFFBE9F22FDD0F860099E53FB" box="[629,711,2021,2040]" name="Morocco" pageId="10" pageNumber="440">Марокко</collectingCountry>
|
||
; 63 — Фархана, Надор,
|
||
<collectingCountry id="D58B8BEFFFBE9F22FC0DF86008A353FB" box="[936,1018,2021,2040]" name="Morocco" pageId="10" pageNumber="440">Марокко</collectingCountry>
|
||
; 64 — Сиди БухуриЯ, Беркане,
|
||
<collectingCountry id="D58B8BEFFFBE9F22FABBF8600E2953FB" box="[1310,1392,2021,2040]" name="Morocco" pageId="10" pageNumber="440">Марокко</collectingCountry>
|
||
; 53–55, 58–61 — по
|
||
<bibRefCitation id="C90DB68EFFBE9F22FED0F87809625C13" author="Pupier R. & Coulon J." box="[373,571,2045,2064]" pageId="10" pageNumber="440" refId="ref17156" refString="Pupier R., Coulon J. 2013. Etude des organes genitaux des femelles et considerations sur la taxonomie de deux groupes d'Orthomus" type="book" year="2013">Pupier, Coulon [2013]</bibRefCitation>
|
||
; 62–64 — по
|
||
<bibRefCitation id="C90DB68EFFBE9F22FD19F87808D15C13" author="Chavanon G. & Mabrouki Y. & Taybi F. A." box="[700,904,2045,2064]" pageId="10" pageNumber="440" pagination="173 - 186" refId="ref16408" refString="Chavanon G., Mabrouki Y., Taybi F. A. 2016. Sur la repartition des populations d' Orthomus lacouri (Antoine, 1941) (Coleoptera, Carabidae, Pterostichinae) dans les Beni Snassen (Maroc Oriental) // Bull. mens. Soc. linn. Lyon. Vol. 85. Nos 5 - 6. P. 173 - 186." type="book chapter" year="2016">Chavanon et al. [2016]</bibRefCitation>
|
||
. МасШтаб: 1,5 мм (Рис. 51); 1,6 мм (Рис. 56).
|
||
</paragraph>
|
||
<paragraph id="AD23CB7FFFBF9F23FF18FEBE0A5F55D4" blockId="11.[189,775,315,923]" pageId="11" pageNumber="441">lobe of aedeagus, but this shape may vary notably depending on the degree of population gene flow, that results in “hybrid” shapes intermediate between those of “typical” exemplars. That paper should be consulted to check the variability; here I shall refer to the so-called “typical” shapes.</paragraph>
|
||
<paragraph id="AD23CB7FFFBF9F23FF47FE590AC856E2" blockId="11.[189,775,315,923]" pageId="11" pageNumber="441">
|
||
In lateral view the distal half of the median lobe of aedeagus is almost straight in
|
||
<emphasis id="9FE8176DFFBF9F23FE4DFE73093C5608" box="[488,613,502,523]" italics="true" pageId="11" pageNumber="441">O. l. kocheri</emphasis>
|
||
, sinuate in
|
||
<emphasis id="9FE8176DFFBF9F23FD7FFE730A5D5625" italics="true" pageId="11" pageNumber="441">O. l. haroldi</emphasis>
|
||
and
|
||
<emphasis id="9FE8176DFFBF9F23FE94FD940AFF5625" box="[305,422,529,550]" italics="true" pageId="11" pageNumber="441">
|
||
O. l.
|
||
<taxonomicName id="6A9CB0FCFFBF9F23FEC5FD940AFF5625" baseAuthorityName="Antoine" baseAuthorityYear="1941" box="[352,422,529,550]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="441" phylum="Arthropoda" rank="species" species="lacouri">lacouri</taxonomicName>
|
||
</emphasis>
|
||
, and is ventrally bent in
|
||
<emphasis id="9FE8176DFFBF9F23FD37FD94085E5625" box="[658,775,529,550]" italics="true" pageId="11" pageNumber="441">O. l. pupieri</emphasis>
|
||
[
|
||
<bibRefCitation id="C90DB68EFFBF9F23FF67FDAE0AC35643" author="Pupier R. & Coulon J." box="[194,410,555,577]" pageId="11" pageNumber="441" refId="ref17156" refString="Pupier R., Coulon J. 2013. Etude des organes genitaux des femelles et considerations sur la taxonomie de deux groupes d'Orthomus" type="book" year="2013">Pupier, Coulon, 2013</bibRefCitation>
|
||
] (Fig. 59). In dorsal view, the apical shaft is truncate and slightly sinuate in
|
||
<emphasis id="9FE8176DFFBF9F23FD93FDC309F55658" box="[566,684,582,603]" italics="true" pageId="11" pageNumber="441">
|
||
O. l.
|
||
<taxonomicName id="6A9CB0FCFFBF9F23FDC3FDC309F55658" baseAuthorityName="Antoine" baseAuthorityYear="1941" box="[614,684,582,603]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="441" phylum="Arthropoda" rank="species" species="lacouri">lacouri</taxonomicName>
|
||
</emphasis>
|
||
(Fig. 64) and truncate with a clear tooth in
|
||
<emphasis id="9FE8176DFFBF9F23FDA2FDE409265675" box="[519,639,609,630]" italics="true" pageId="11" pageNumber="441">O. l. pupieri</emphasis>
|
||
(Fig. 61); the other two subspecies show a rounded apex, sinuate to the left, being wider in
|
||
<emphasis id="9FE8176DFFBF9F23FEFEFD130A8456A8" box="[347,477,662,683]" italics="true" pageId="11" pageNumber="441">O. l. kocheri</emphasis>
|
||
[
|
||
<bibRefCitation id="C90DB68EFFBF9F23FE49FD13099056A8" author="Pupier R. & Coulon J." box="[492,713,662,684]" pageId="11" pageNumber="441" refId="ref17156" refString="Pupier R., Coulon J. 2013. Etude des organes genitaux des femelles et considerations sur la taxonomie de deux groupes d'Orthomus" type="book" year="2013">Pupier, Coulon, 2013</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="C90DB68EFFBF9F23FD7DFD130A2D56C5" author="Chavanon G. & Mabrouki Y. & Taybi F. A." pageId="11" pageNumber="441" pagination="173 - 186" refId="ref16408" refString="Chavanon G., Mabrouki Y., Taybi F. A. 2016. Sur la repartition des populations d' Orthomus lacouri (Antoine, 1941) (Coleoptera, Carabidae, Pterostichinae) dans les Beni Snassen (Maroc Oriental) // Bull. mens. Soc. linn. Lyon. Vol. 85. Nos 5 - 6. P. 173 - 186." type="book chapter" year="2016">Chavanon et al., 2016</bibRefCitation>
|
||
] and slender and more elongate in
|
||
<emphasis id="9FE8176DFFBF9F23FD7CFD340A5D56E3" italics="true" pageId="11" pageNumber="441">O. l. haroldi</emphasis>
|
||
(Figs 62–63).
|
||
</paragraph>
|
||
<paragraph id="AD23CB7FFFBF9F23FF47FD630A275798" blockId="11.[189,775,315,923]" pageId="11" pageNumber="441">
|
||
According to the map provided by
|
||
<bibRefCitation id="C90DB68EFFBF9F23FD88FD63085E56FF" author="Chavanon G. & Mabrouki Y. & Taybi F. A." box="[557,775,742,764]" pageId="11" pageNumber="441" pagination="173 - 186" refId="ref16408" refString="Chavanon G., Mabrouki Y., Taybi F. A. 2016. Sur la repartition des populations d' Orthomus lacouri (Antoine, 1941) (Coleoptera, Carabidae, Pterostichinae) dans les Beni Snassen (Maroc Oriental) // Bull. mens. Soc. linn. Lyon. Vol. 85. Nos 5 - 6. P. 173 - 186." type="book chapter" year="2016">Chavanon et al. [2016]</bibRefCitation>
|
||
the subspecies
|
||
<emphasis id="9FE8176DFFBF9F23FEF4FC840A975715" box="[337,462,769,790]" italics="true" pageId="11" pageNumber="441">O. l. kocheri</emphasis>
|
||
is found west of River Muluya whereas
|
||
<emphasis id="9FE8176DFFBF9F23FEB6FC990AD55733" box="[275,396,795,817]" italics="true" pageId="11" pageNumber="441">
|
||
O. l.
|
||
<taxonomicName id="6A9CB0FCFFBF9F23FEE3FC9E0AD55733" baseAuthorityName="Antoine" baseAuthorityYear="1941" box="[326,396,795,816]" class="Insecta" family="Carabidae" genus="Orthomus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="441" phylum="Arthropoda" rank="species" species="lacouri">lacouri</taxonomicName>
|
||
</emphasis>
|
||
is found in the surroundings of Oujda and the Algerian border, to the west near Berkane, El Aioun and Guerfane;
|
||
<emphasis id="9FE8176DFFBF9F23FEEFFCD40A995765" box="[330,448,849,870]" italics="true" pageId="11" pageNumber="441">O. l. haroldi</emphasis>
|
||
is found in-between the other two subspecies. Hybrids between these subspecies are found in many contact areas.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |