209 lines
29 KiB
XML
209 lines
29 KiB
XML
<document id="4B8544DCE1A33F747A421B2C0D36C570" ID-ISSN="1536-9307" ID-ZooBank="43AA8CDF-9C04-40E8-BE69-BDE1E436DA18" IM.materialsCitations_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1617910976782" checkinUser="carolina" docAuthor="Kovařík, František, Lowe, Graeme & Šťáhlavský, František" docDate="2016" docId="1A1E8F5B9476FFF2FE9BFCCC9076FACE" docLanguage="en" docName="Euscorpius.236.1-18.pdf" docOrigin="Euscorpius 236" docStyle="DocumentStyle:05AD4CDCECA6C90C60787CEF3EC4672A.3:Euscorpius.2002-2019.journal_article" docStyleId="05AD4CDCECA6C90C60787CEF3EC4672A" docStyleName="Euscorpius.2002-2019.journal_article" docStyleVersion="3" docTitle="Buthacus Birula 1908" docType="treatment" docVersion="3" lastPageNumber="2" masterDocId="E627F7239475FFF1FFEDFF9A9448FFFE" masterDocTitle="Review of Northwestern African Buthacus, with description of Buthacus stockmanni sp. n. from Morocco and Western Sahara (Scorpiones, Buthidae)" masterLastPageNumber="18" masterPageNumber="1" pageNumber="2" updateTime="1698902728836" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="88C4CBB7441347687EF7295F2C8D1A1C">Review of Northwestern African Buthacus, with description of Buthacus stockmanni sp. n. from Morocco and Western Sahara (Scorpiones, Buthidae)</mods:title>
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<mods:namePart id="15C27C150FA8C7C0D34816E71C5703CE">Kovařík, František</mods:namePart>
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<mods:namePart id="08D5FF80E5B43C5D806643291C3093DB">Lowe, Graeme</mods:namePart>
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<mods:namePart id="7E7B7EEE13B8C066755DB5004A200335">Šťáhlavský, František</mods:namePart>
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<mods:title id="B0CA0BC1559B89C69F24AAA7753F7D6E">Euscorpius</mods:title>
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<mods:date id="EB4C49A07D7B461E24E7809E25BAEF20">2016</mods:date>
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<mods:number id="BEB4F64D770F78819A7F96573AB1132A">236</mods:number>
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<treatment id="1A1E8F5B9476FFF2FE9BFCCC9076FACE" ID-DOI="http://doi.org/10.5281/zenodo.4673059" ID-Zenodo-Dep="4673059" LSID="urn:lsid:plazi:treatment:1A1E8F5B9476FFF2FE9BFCCC9076FACE" httpUri="http://treatment.plazi.org/id/1A1E8F5B9476FFF2FE9BFCCC9076FACE" lastPageNumber="2" pageId="3" pageNumber="2">
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<taxonomicName id="55B745CE9476FFF2FE9BFCCC9623FC90" ID-CoL="62HVF" authority="Birula, 1908" authorityName="Birula" authorityYear="1908" box="[374,619,854,878]" class="Arachnida" family="Buthidae" genus="Buthacus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="genus">
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<emphasis id="A0C3E25F9476FFF2FE9BFCCC9597FC90" bold="true" box="[374,479,854,878]" italics="true" pageId="3" pageNumber="2">Buthacus</emphasis>
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<bibRefCitation id="F62643BC9476FFF2FE0BFCCD9623FC90" author="BIRULA" box="[486,619,855,878]" pageId="3" pageNumber="2" pagination="121 - 152" refId="ref7027" refString="BIRULA, A. A. 1908. Ergebnisse der mit Subvention aus der Erbschaft Treitl unternommenen zoologischen Forschungsreise Dr. F. Werner`s nach dem agyptischen Sudan und Nord-Uganda. XIV. Scorpiones und Solifugae. Sitzungsberichte der Kaiserlich- Koniglichen Akademie der Wissenschaften, Wien, 117 / 2 (1): 121 - 152." type="journal article" year="1908">Birula, 1908</bibRefCitation>
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</taxonomicName>
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</heading>
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</paragraph>
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<subSubSection id="DAAD6DC69476FFF2FE6CFCEC9629FC73" box="[385,609,886,909]" pageId="3" pageNumber="2" type="description">
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<paragraph id="92083E4D9476FFF2FE6CFCEC9629FC73" blockId="3.[374,619,854,909]" box="[385,609,886,909]" pageId="3" pageNumber="2">
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(
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<figureCitation id="0A8C22C89476FFF2FE64FCEC95B2FC73" box="[393,506,886,909]" captionStart-0="Figures 1–4" captionStart-1="Figures 5–8" captionStart-10="Figures 52–58" captionStart-11="Figures 59–60" captionStart-12="Figure 63" captionStart-2="Figures 9–14" captionStart-3="Figures 15–29" captionStart-4="Figures 30–33" captionStart-5="Figures 34–38" captionStart-6="Figures 39–40" captionStart-7="Figures 41–42" captionStart-8="Figures 43–44" captionStart-9="Figures 45–51" captionStartId-0="4.[192,278,1684,1707]" captionStartId-1="5.[192,278,1276,1299]" captionStartId-10="14.[192,278,1331,1354]" captionStartId-11="15.[192,278,1614,1637]" captionStartId-12="17.[192,268,1362,1385]" captionStartId-2="6.[192,278,1287,1310]" captionStartId-3="7.[192,278,1113,1136]" captionStartId-4="8.[192,269,1225,1246]" captionStartId-5="9.[192,278,1038,1061]" captionStartId-6="10.[192,278,1790,1813]" captionStartId-7="11.[192,278,1758,1781]" captionStartId-8="12.[192,278,658,681]" captionStartId-9="13.[192,278,873,896]" captionTargetBox-0="[205,1430,210,1642]" captionTargetBox-1="[191,1438,191,1246]" captionTargetBox-10="[191,1437,191,1301]" captionTargetBox-11="[191,1440,203,1585]" captionTargetBox-12="[192,1402,192,1332]" captionTargetBox-2="[202,1429,202,1243]" captionTargetBox-3="[192,1433,202,1078]" captionTargetBox-4="[194,1424,204,1193]" captionTargetBox-5="[200,1429,178,1002]" captionTargetBox-6="[191,1440,215,1761]" captionTargetBox-7="[191,1440,199,1729]" captionTargetBox-8="[191,1437,191,628]" captionTargetBox-9="[202,1437,176,841]" captionTargetId-0="figure-17@4.[192,1439,192,1655]" captionTargetId-1="figure-6@5.[192,1438,192,1245]" captionTargetId-10="figure-17@14.[192,1437,192,1300]" captionTargetId-11="figure-6@15.[192,1439,204,1584]" captionTargetId-12="figure-6@17.[192,1402,192,1332]" captionTargetId-2="figure-17@6.[192,1437,192,1257]" captionTargetId-3="figure-6@7.[192,1437,192,1083]" captionTargetId-4="figure-17@8.[192,1438,192,1196]" captionTargetId-5="figure-6@9.[192,1437,168,1008]" captionTargetId-6="figure-17@10.[192,1439,216,1760]" captionTargetId-7="figure-6@11.[192,1439,200,1728]" captionTargetId-8="figure-17@12.[192,1437,192,627]" captionTargetId-9="figure-6@13.[192,1437,168,844]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-10="14" captionTargetPageId-11="15" captionTargetPageId-12="17" captionTargetPageId-2="6" captionTargetPageId-3="7" captionTargetPageId-4="8" captionTargetPageId-5="9" captionTargetPageId-6="10" captionTargetPageId-7="11" captionTargetPageId-8="12" captionTargetPageId-9="13" captionText-0="Figures 1–4: Buthacus stockmanni sp. n. Figures 1–2. Male holotype in dorsal (1) and ventral (2) views. Figures 3–4. Female paratype from the type locality in dorsal (3) and ventral (4) aspects. Scale bar: 10 mm." captionText-1="Figures 5–8: Buthacus stockmanni sp. n. Figures 5 and 7. Male holotype, carapace and tergites (5), coxosternal area and sternites III–V (7). Figures 6 and 8. Female paratype from the type locality, carapace and tergites (6), coxosternal area and sternites III–IV (8)." captionText-10="Figures 52–58: Figures 52–55. Buthacus occidentalis, male holotype of Buthacus ehrenbergi, telson lateral (52), carapace and tergites I–IV (53), and retrolateral aspect of leg IV (54) and leg III (55). Figures 56–58. Buthacus ziegleri, male from locality Morocco, Merzouga env., 31°02'59"N 04°00'16"W, telson lateral (56) and retrolateral aspect of leg IV (57) and leg III (58)." captionText-11="Figures 59–60: Buthacus ziegleri, in vivo habitus, male (59) and female (60) at the locality Morocco, Merzouga env., 31°02'59"N 04°00'16"W." captionText-12="Figure 63: Map showing confirmed distribution of species of Buthacus leptochelys complex in Northwestern Africa." captionText-2="Figures 9–14: Buthacus stockmanni sp. n., metasoma and telson. Figures 9–11. Male holotype, lateral (9), ventral (10), and dorsal (11) views. Figures 12–14. Female paratype from the type locality, lateral (12), ventral (13), and dorsal (14) views. Scale bars: 10 mm." captionText-3="Figures 15–29: Buthacus stockmanni sp. n. Figures 15–27: pedipalp segments. Figures 15–16. Female paratype from the type locality, chela dorsal (15) and external (16). Figures 17–27. Male holotype, chela dorsal (17), external (18), and ventral (19). Patella dorsal (20), external (21) and ventral (22). Femur internal (23), dorsal (24), and ventral (25). Movable finger (26) and fixed finger (27) dentition. Trichobothrial pattern is indicated in Figures 17–21 and 23–24. Figures 28–29. Telson lateral, female paratype from the type locality (28) and male holotype (29)." captionText-4="Figures 30–33: Buthacus stockmanni sp. n., male holotype, left legs I–IV, retrolateral aspect." captionText-5="Figures 34–38: Buthacus stockmanni sp. n., right hemispermatophore, male holotype. Figure 34. Whole hemispermatophore, dorsal view. Figures 35–38. Capsule region and flagellum: dorsal (35), external (36), ventral (37) and internal (38) views. Abbreviations: bl, basal lobe; el, external lobe, il, internal lobe; ml, median lobe. Scale bars: 1 mm (34), 500 µm (35–38)." captionText-6="Figures 39–40: Buthacus stockmanni sp. n., in vivo habitus, male (39) holotype and female (40) paratype from the type locality." captionText-7="Figures 41–42: Buthacus stockmanni sp. n., type locality (41) and locality Morocco, North of Msied, 28.04383°N 10.85038°W. (42)." captionText-8="Figures 43–44: Buthacus stockmanni sp. n., mitotic metaphase (43) and ideogram (44) (y axis - % of the chromosome length of the diploid set, lines indicate min.-max. values) of paratype male from the type locality. Arrowheads indicate extra large chromosomes. Scale bar = 10 µm." captionText-9="Figures 45–51: Figure 45. Buthacus stockmanni sp. n., male paratype from locality Morocco, North of Msied, 28.04383°N 10.85038°W, pedipalp chela external. Figures 46–48. Buthacus occidentalis, male holotype of Buthacus ehrenbergi, pedipalp chela external (46) and dorsal (47), and movable finger dentition (48). Figures 49–51. Buthacus ziegleri, male from locality Morocco, Merzouga env., 31°02'59"N 04°00'16"W, pedipalp chela external (49) and dorsal (50), and movable finger dentition (51)." pageId="3" pageNumber="2">Figs. 1–63</figureCitation>
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,
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<tableCitation id="DF350BF69476FFF2FDEAFCEC961FFC73" box="[519,599,886,909]" captionStart="Table 1" captionStartId="12.[192,257,1420,1443]" captionText="Table 1: Comparative measurements of adults of Buthacus stockmanni sp. n. Abbreviations: length (L), width (W, in carapace it corresponds to posterior width), depth (D)." pageId="3" pageNumber="2">Table 1</tableCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="DAAD6DC69476FFF2FF2DFC2996FCFBBB" pageId="3" pageNumber="2" type="reference_group">
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<paragraph id="92083E4D9476FFF2FF2DFC2996CCFC34" blockId="3.[192,802,947,1093]" box="[192,644,947,970]" pageId="3" pageNumber="2">
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<treatmentCitationGroup id="B2A719639476FFF2FF2DFC2996CCFC34" box="[192,644,947,970]" pageId="3" pageNumber="2">
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<treatmentCitation id="1316185C9476FFF2FF2DFC299636FC34" author="BIRULA" box="[192,638,947,970]" page="140" pageId="3" pageNumber="2" year="1908">
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<taxonomicName id="55B745CE9476FFF2FF2DFC299636FC34" authority="Birula, 1908: 139 - 140" authorityName="Birula" authorityPageNumber="139 - 140" authorityYear="1908" box="[192,638,947,970]" class="Arachnida" family="Buthidae" genus="Buthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="subGenus" subGenus="Buthacus">
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<emphasis id="A0C3E25F9476FFF2FF2DFC299542FC34" box="[192,266,947,970]" italics="true" pageId="3" pageNumber="2">Buthus</emphasis>
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(
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<emphasis id="A0C3E25F9476FFF2FEF7FC299535FC34" box="[282,381,947,970]" italics="true" pageId="3" pageNumber="2">Buthacus</emphasis>
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)
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<bibRefCitation id="F62643BC9476FFF2FE60FC299636FC34" author="BIRULA" box="[397,638,947,970]" pageId="3" pageNumber="2" pagination="121 - 152" refId="ref7027" refString="BIRULA, A. A. 1908. Ergebnisse der mit Subvention aus der Erbschaft Treitl unternommenen zoologischen Forschungsreise Dr. F. Werner`s nach dem agyptischen Sudan und Nord-Uganda. XIV. Scorpiones und Solifugae. Sitzungsberichte der Kaiserlich- Koniglichen Akademie der Wissenschaften, Wien, 117 / 2 (1): 121 - 152." type="journal article" year="1908">Birula, 1908: 139–140</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="92083E4D9476FFF2FF2DFC4896FCFBBB" blockId="3.[192,802,947,1093]" pageId="3" pageNumber="2">
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<taxonomicName id="55B745CE9476FFF2FF2DFC4895AEFC17" ID-CoL="62HVF" authority=": Simon, 1910: 74" authorityName=": Simon" authorityPageNumber="74" authorityYear="1910" box="[192,486,978,1001]" class="Arachnida" family="Buthidae" genus="Buthacus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="genus">
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<emphasis id="A0C3E25F9476FFF2FF2DFC48956BFC17" box="[192,291,978,1001]" italics="true" pageId="3" pageNumber="2">Buthacus</emphasis>
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: Simon, 1910: 74
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</taxonomicName>
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(in part); Fet & Lowe, 2000: 81–86 (in part, complete reference list until 2000);
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<bibRefCitation id="F62643BC9476FFF2FF1DFB95959FFBD8" author="KOVARIK" box="[240,471,1039,1062]" pageId="3" pageNumber="2" pagination="1 - 13" refId="ref7347" refString="KOVARIK, F. 2005. Taxonomic position of species of the genus Buthacus Birula, 1908 described by Ehrenberg and Lourenco, and description of a new species (Scorpiones: Buthidae). Euscorpius, 28: 1 - 13." type="journal article" year="2005">Kovařík, 2005: 1–10</bibRefCitation>
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, figs. 1–8 (in part); Kovařík, 2009: 30; Kovařík et al., 2013: 1–4, fig. 5.
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||
</paragraph>
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</subSubSection>
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<subSubSection id="DAAD6DC69476FFF2FF2DFBF19531FB5F" pageId="3" pageNumber="2" type="materials_examined">
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<paragraph id="92083E4D9476FFF2FF2DFBF19531FB5F" blockId="3.[192,801,1131,1185]" pageId="3" pageNumber="2">
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<materialsCitation id="22DF34109476FFF2FF2DFBF1953EFB5F" collectionCode="SPECIES" pageId="3" pageNumber="2" specimenCount="1" typeStatus="holotype">
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<typeStatus id="4D0C80EF9476FFF2FF2DFBF194B0FB7C" box="[192,248,1131,1154]" pageId="3" pageNumber="2">TYPE</typeStatus>
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<collectionCode id="F4A6A6889476FFF2FEE2FBF5952AFB7C" box="[271,354,1135,1154]" pageId="3" pageNumber="2">SPECIES</collectionCode>
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.
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<taxonomicName id="55B745CE9476FFF2FE6CFBF1953EFB5F" authority="Ehrenberg, 1829" authorityName="Ehrenberg" authorityYear="1829" class="Arachnida" family="Buthidae" genus="Androctonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="leptopchelys" subGenus="Leiurus">
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<emphasis id="A0C3E25F9476FFF2FE6CFBF19641FB7C" box="[385,521,1131,1154]" italics="true" pageId="3" pageNumber="2">Androctonus</emphasis>
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(
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<emphasis id="A0C3E25F9476FFF2FDC7FBF19633FB7C" box="[554,635,1131,1154]" italics="true" pageId="3" pageNumber="2">Leiurus</emphasis>
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)
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<emphasis id="A0C3E25F9476FFF2FD71FBF19769FB7C" box="[668,801,1131,1154]" italics="true" pageId="3" pageNumber="2">leptopchelys</emphasis>
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Ehrenberg, 1829
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</taxonomicName>
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="DAAD6DC69476FFF2FF2DFB5D9109FE2B" pageId="3" pageNumber="2" type="diagnosis">
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<paragraph id="92083E4D9476FFF2FF2DFB5D9109FE2B" blockId="3.[192,802,1223,1890]" lastBlockId="3.[830,1440,199,469]" pageId="3" pageNumber="2">
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DIAGNOSIS. Total length
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<quantity id="554F93A89476FFF2FE3BFB5D9605FB20" box="[470,589,1223,1246]" metricMagnitude="-2" metricUnit="m" metricValue="6.5" metricValueMax="9.0" metricValueMin="4.0" pageId="3" pageNumber="2" unit="mm" value="65.0" valueMax="90.0" valueMin="40.0">40–90 mm</quantity>
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(except
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<taxonomicName id="55B745CE9476FFF2FD42FB5D9511FB03" authority="Vachon, 1949" authorityName="Vachon" authorityYear="1949" class="Arachnida" family="Buthidae" genus="Buthacus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="villiersi">
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<emphasis id="A0C3E25F9476FFF2FD42FB5D9769FB20" box="[687,801,1223,1246]" italics="true" pageId="3" pageNumber="2">B. villiersi</emphasis>
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Vachon, 1949
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</taxonomicName>
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and
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<taxonomicName id="55B745CE9476FFF2FE79FB7C96E7FB03" authority="Lourenco, 2001" authorityName="Lourenco" authorityYear="2001" box="[404,687,1254,1277]" class="Arachnida" family="Buthidae" genus="Buthacus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="clevai">
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<emphasis id="A0C3E25F9476FFF2FE79FB7C95BCFB03" box="[404,500,1254,1277]" italics="true" pageId="3" pageNumber="2">B. clevai</emphasis>
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Lourenço, 2001
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</taxonomicName>
|
||
); carapace trapezoidal, in lateral view preocular area not distinctly inclined towards anterior margin, level with or higher than postocular area; surface of carapace conspicuously granular, with only anterior median carinae developed, anterior part of carapace glossy; ventral aspect of cheliceral fixed finger with two denticles; tergites with three carinae of which lateral pair on I and II inconspicuous; pectines with fulcra, hirsute; hemispermatophore flagelliform, capsule with 4 lobes, lobes separated from flagellum, basal lobe small, knob-like; sternum subtriangular; metasomal segments I–III with 8–10 carinae; metasoma II as wide as other metasomal segments; metasoma V with enlarged "lobate" dentition on ventrolateral carinae; telson without subaculear tubercle, with long curved aculeus, longer than vesicle (except
|
||
<taxonomicName id="55B745CE9476FFF2FEFAF94A962CF919" authority="Hendrixson, 2006" authorityName="Hendrixson" authorityYear="2006" box="[279,612,1744,1767]" class="Arachnida" family="Buthidae" genus="Buthacus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="buettikeri">
|
||
<emphasis id="A0C3E25F9476FFF2FEFAF94A95D4F919" box="[279,412,1744,1767]" italics="true" pageId="3" pageNumber="2">B. buettikeri</emphasis>
|
||
Hendrixson, 2006
|
||
</taxonomicName>
|
||
); all segments of metasoma hirsute, with long setae in both sexes, dentate margin of movable finger of pedipalp with 9–12 rows of granules, each row equipped with one internal accessory granule, and with (
|
||
<taxonomicName id="55B745CE9476FFF2FE75F8D1967BF89C" baseAuthorityName="Ehrenberg" baseAuthorityYear="1829" box="[408,563,1867,1890]" class="Arachnida" family="Buthidae" genus="Buthacus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="leptochelys">
|
||
<emphasis id="A0C3E25F9476FFF2FE75F8D1967BF89C" box="[408,563,1867,1890]" italics="true" pageId="3" pageNumber="2">B. leptochelys</emphasis>
|
||
</taxonomicName>
|
||
complex) or without (
|
||
<taxonomicName id="55B745CE9476FFF2FCAAFF5D9783FF20" baseAuthorityName="Simon" baseAuthorityYear="1885" box="[839,971,199,222]" class="Arachnida" family="Buthidae" genus="Buthacus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="arenicola">
|
||
<emphasis id="A0C3E25F9476FFF2FCAAFF5D9783FF20" box="[839,971,199,222]" italics="true" pageId="3" pageNumber="2">B. arenicola</emphasis>
|
||
</taxonomicName>
|
||
complex) one external accessory granule, 4 terminal and one basal terminal granules present; trichobothrial pattern orthobothriotaxic
|
||
<typeStatus id="4D0C80EF9476FFF2FB1FFE9E9168FEE5" box="[1266,1312,260,283]" pageId="3" pageNumber="2">type</typeStatus>
|
||
A; dorsal trichobothria of femur arranged in β- configuration; pedipalp patella with 7 external trichobothria; pedipalp femur with trichobothrium
|
||
<emphasis id="A0C3E25F9476FFF2FB9CFEF89036FE87" box="[1137,1150,354,377]" italics="true" pageId="3" pageNumber="2">d</emphasis>
|
||
<subScript id="0E333C089476FFF2FB92FEF790C0FE82" attach="left" box="[1151,1160,365,380]" fontSize="6" pageId="3" pageNumber="2">2</subScript>
|
||
on dorsal surface;
|
||
<emphasis id="A0C3E25F9476FFF2FA85FEF8913DFE87" box="[1384,1397,354,377]" italics="true" pageId="3" pageNumber="2">d</emphasis>
|
||
<subScript id="0E333C089476FFF2FA98FEF79136FE82" attach="left" box="[1397,1406,365,380]" fontSize="6" pageId="3" pageNumber="2">2</subScript>
|
||
of pedipalp patella present; patella trichobothrium
|
||
<emphasis id="A0C3E25F9476FFF2FAB2FE1B9124FE66" box="[1375,1388,385,408]" italics="true" pageId="3" pageNumber="2">d</emphasis>
|
||
<subScript id="0E333C089476FFF2FA81FE11913DFE64" attach="left" box="[1388,1397,395,410]" fontSize="6" pageId="3" pageNumber="2">3</subScript>
|
||
internal to dorsomedian carina; tibial spurs present on legs III–IV but could be reduced or absent on leg III.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="DAAD6DC69476FFF2FCD3FE669076FACE" pageId="3" pageNumber="2" type="discussion">
|
||
<paragraph id="92083E4D9476FFF2FCD3FE6690ABFD35" blockId="3.[830,1442,508,1328]" pageId="3" pageNumber="2">
|
||
COMMENTS. Many of the diagnostic characters applied to differentiate
|
||
<taxonomicName id="55B745CE9476FFF2FC05FD809004FDCF" authorityName=": Simon" authorityYear="1910" box="[1000,1100,538,561]" class="Arachnida" family="Buthidae" genus="Buthacus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="A0C3E25F9476FFF2FC05FD809004FDCF" box="[1000,1100,538,561]" italics="true" pageId="3" pageNumber="2">Buthacus</emphasis>
|
||
</taxonomicName>
|
||
from other buthid genera seem to be plesiomorphic, so the genus is not strongly supported by derived characters. Thus,
|
||
<taxonomicName id="55B745CE9476FFF2FB20FDC29178FD91" authorityName=": Simon" authorityYear="1910" box="[1229,1328,600,623]" class="Arachnida" family="Buthidae" genus="Buthacus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="A0C3E25F9476FFF2FB20FDC29178FD91" box="[1229,1328,600,623]" italics="true" pageId="3" pageNumber="2">Buthacus</emphasis>
|
||
</taxonomicName>
|
||
may be a paraphyletic assemblage of taxa containing several distinct lineages. It is beyond the scope of this paper to completely revise and divide
|
||
<taxonomicName id="55B745CE9476FFF2FB95FD2E9093FD35" authorityName=": Simon" authorityYear="1910" box="[1144,1243,692,715]" class="Arachnida" family="Buthidae" genus="Buthacus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="A0C3E25F9476FFF2FB95FD2E9093FD35" box="[1144,1243,692,715]" italics="true" pageId="3" pageNumber="2">Buthacus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="92083E4D9476FFF2FC83FD48904EFC03" blockId="3.[830,1442,508,1328]" pageId="3" pageNumber="2">
|
||
<taxonomicName id="55B745CE9476FFF2FC83FD489169FD17" authority="Zambre et Lourenco, 2010" authorityName="Zambre et Lourenco" authorityYear="2010" box="[878,1313,722,745]" class="Arachnida" family="Buthidae" genus="Buthacus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="agarwali">
|
||
<emphasis id="A0C3E25F9476FFF2FC83FD4897B8FD17" box="[878,1008,722,745]" italics="true" pageId="3" pageNumber="2">B. agarwali</emphasis>
|
||
Zambre et Lourenço, 2010
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry id="EAA07EDD9476FFF2FA84FD4891E8FD17" box="[1385,1440,722,745]" name="India" pageId="3" pageNumber="2">India</collectingCountry>
|
||
definitely does not belong in
|
||
<taxonomicName id="55B745CE9476FFF2FB90FD6B9097FCF6" authorityName=": Simon" authorityYear="1910" box="[1149,1247,753,776]" class="Arachnida" family="Buthidae" genus="Buthacus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="A0C3E25F9476FFF2FB90FD6B9097FCF6" box="[1149,1247,753,776]" italics="true" pageId="3" pageNumber="2">Buthacus</emphasis>
|
||
</taxonomicName>
|
||
because it differs in the characters cited in the diagnosis above as well as in other aspects of its morphology, mainly in shape of the pedipalp segments. The taxonomic position of
|
||
<taxonomicName id="55B745CE9476FFF2FA64FCD790D9FC7D" authority="Lourenco et Qi, 2007" authorityName="Lourenco et Qi" authorityYear="2007" class="Arachnida" family="Buthidae" genus="Buthacus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="maliensis">
|
||
<emphasis id="A0C3E25F9476FFF2FA64FCD797EBFC7D" italics="true" pageId="3" pageNumber="2">B. maliensis</emphasis>
|
||
Lourenço et Qi, 2007
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry id="EAA07EDD9476FFF2FB38FCF6914FFC7D" box="[1237,1287,876,899]" name="Serbia" pageId="3" pageNumber="2">Mali</collectingCountry>
|
||
is also doubtful. It has a short aculeus, the anterior part of the carapace is probably not glossy, the movable finger of pedipalp has nearly linear rows of granules, and the total length is 29.8 mm.
|
||
</paragraph>
|
||
<paragraph id="92083E4D9476FFF2FC83FB9F9076FACE" blockId="3.[830,1442,508,1328]" pageId="3" pageNumber="2">
|
||
It appears that the most reliable diagnostic character for
|
||
<taxonomicName id="55B745CE9476FFF2FC88FBBE9780FBC5" authorityName=": Simon" authorityYear="1910" box="[869,968,1060,1083]" class="Arachnida" family="Buthidae" genus="Buthacus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="A0C3E25F9476FFF2FC88FBBE9780FBC5" box="[869,968,1060,1083]" italics="true" pageId="3" pageNumber="2">Buthacus</emphasis>
|
||
</taxonomicName>
|
||
is the presence of 9–12 rows of granules on the movable finger. The only species for which this character is inapplicable is
|
||
<taxonomicName id="55B745CE9476FFF2FB8BFBFB904AFB69" authority="Lourenco et Leguin, 2009" authorityName="Lourenco et Leguin" authorityYear="2009" class="Arachnida" family="Buthidae" genus="Buthacus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="williamsi">
|
||
<emphasis id="A0C3E25F9476FFF2FB8BFBFB917AFB86" box="[1126,1330,1121,1144]" italics="true" pageId="3" pageNumber="2">Buthacus williamsi</emphasis>
|
||
Lourenço et Leguin, 2009
|
||
</taxonomicName>
|
||
from the
|
||
<collectingCountry id="EAA07EDD9476FFF2FB79FB1A91D4FB69" box="[1172,1436,1152,1175]" name="United Arab Emirates" pageId="3" pageNumber="2">United Arab Emirates</collectingCountry>
|
||
, described from three females that have 6 or 7 rows of granules on the movable finger. We are convinced, however, that this species does not belong to the genus
|
||
<taxonomicName id="55B745CE9476FFF2FCD3FB6097E9FAEF" authorityName=": Simon" authorityYear="1910" box="[830,929,1274,1297]" class="Arachnida" family="Buthidae" genus="Buthacus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="A0C3E25F9476FFF2FCD3FB6097E9FAEF" box="[830,929,1274,1297]" italics="true" pageId="3" pageNumber="2">Buthacus</emphasis>
|
||
</taxonomicName>
|
||
but probably to the genus
|
||
<taxonomicName id="55B745CE9476FFF2FB3EFB60907FFACE" authority="Levy, Amitai et Shulov, 1973" authorityName="Levy, Amitai et Shulov" authorityYear="1973" class="Arachnida" family="Buthidae" genus="Vachoniolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="A0C3E25F9476FFF2FB3EFB609110FAEF" box="[1235,1368,1274,1297]" italics="true" pageId="3" pageNumber="2">Vachoniolus</emphasis>
|
||
Levy, Amitai et Shulov, 1973
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |