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<mods:title id="964EC3A05ED6D3E03B2DC6706476BC23">Review of Northwestern African Buthacus, with description of Buthacus stockmanni sp. n. from Morocco and Western Sahara (Scorpiones, Buthidae)</mods:title>
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(
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Figures 1–2. Male holotype in dorsal (1) and ventral (2) views. Figures 3–4. Female paratype from the type locality in dorsal (3) and ventral (4) aspects. Scale bar: 10 mm." captionText-1="Figures 5–8: Buthacus stockmanni sp. n. Figures 5 and 7. Male holotype, carapace and tergites (5), coxosternal area and sternites III–V (7). Figures 6 and 8. Female paratype from the type locality, carapace and tergites (6), coxosternal area and sternites III–IV (8)." captionText-2="Figures 9–14: Buthacus stockmanni sp. n., metasoma and telson. Figures 9–11. Male holotype, lateral (9), ventral (10), and dorsal (11) views. Figures 12–14. Female paratype from the type locality, lateral (12), ventral (13), and dorsal (14) views. Scale bars: 10 mm." captionText-3="Figures 15–29: Buthacus stockmanni sp. n. Figures 15–27: pedipalp segments. Figures 15–16. Female paratype from the type locality, chela dorsal (15) and external (16). Figures 17–27. Male holotype, chela dorsal (17), external (18), and ventral (19). Patella dorsal (20), external (21) and ventral (22). Femur internal (23), dorsal (24), and ventral (25). Movable finger (26) and fixed finger (27) dentition. Trichobothrial pattern is indicated in Figures 17–21 and 23–24. Figures 28–29. Telson lateral, female paratype from the type locality (28) and male holotype (29)." captionText-4="Figures 30–33: Buthacus stockmanni sp. n., male holotype, left legs I–IV, retrolateral aspect." captionText-5="Figures 34–38: Buthacus stockmanni sp. n., right hemispermatophore, male holotype. Figure 34. Whole hemispermatophore, dorsal view. Figures 35–38. Capsule region and flagellum: dorsal (35), external (36), ventral (37) and internal (38) views. Abbreviations: bl, basal lobe; el, external lobe, il, internal lobe; ml, median lobe. Scale bars: 1 mm (34), 500 µm (35–38)." captionText-6="Figures 39–40: Buthacus stockmanni sp. n., in vivo habitus, male (39) holotype and female (40) paratype from the type locality." captionText-7="Figures 41–42: Buthacus stockmanni sp. n., type locality (41) and locality Morocco, North of Msied, 28.04383°N 10.85038°W. (42)." captionText-8="Figures 43–44: Buthacus stockmanni sp. n., mitotic metaphase (43) and ideogram (44) (y axis - % of the chromosome length of the diploid set, lines indicate min.-max. values) of paratype male from the type locality. Arrowheads indicate extra large chromosomes. Scale bar = 10 µm." captionText-9="Figures 45–51: Figure 45. Buthacus stockmanni sp. n., male paratype from locality Morocco, North of Msied, 28.04383°N 10.85038°W, pedipalp chela external. Figures 46–48. Buthacus occidentalis, male holotype of Buthacus ehrenbergi, pedipalp chela external (46) and dorsal (47), and movable finger dentition (48). Figures 49–51. Buthacus ziegleri, male from locality Morocco, Merzouga env., 31°02'59&quot;N 04°00'16&quot;W, pedipalp chela external (49) and dorsal (50), and movable finger dentition (51)." pageId="7" pageNumber="6">Figs. 1–45</figureCitation>
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<tableCitation id="DF350BF69472FFF6FDF6FABC9623FAC3" box="[539,619,1318,1341]" captionStart="Table 1" captionStartId="12.[192,257,1420,1443]" captionText="Table 1: Comparative measurements of adults of Buthacus stockmanni sp. n. Abbreviations: length (L), width (W, in carapace it corresponds to posterior width), depth (D)." pageId="7" pageNumber="6">Table 1</tableCitation>
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<taxonomicName id="55B745CE9472FFF6FF2DFA3B956BFA46" authorityName=": Simon" authorityYear="1910" box="[192,291,1441,1464]" class="Arachnida" family="Buthidae" genus="Buthacus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="6" phylum="Arthropoda" rank="genus">
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sp. Stockmann et al., 2016: 9.
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<figureCitation id="0A8C22C89472FFF6FDDDF90C96D1F953" box="[560,665,1686,1709]" captionStart="Figures 1–4" captionStartId="4.[192,278,1684,1707]" captionTargetBox="[205,1430,210,1642]" captionTargetId="figure-17@4.[192,1439,192,1655]" captionTargetPageId="4" captionText="Figures 1–4: Buthacus stockmanni sp. n. Figures 1–2. Male holotype in dorsal (1) and ventral (2) views. Figures 3–4. Female paratype from the type locality in dorsal (3) and ventral (4) aspects. Scale bar: 10 mm." pageId="7" pageNumber="6">Figs. 1–2</figureCitation>
, 
<figureCitation id="0A8C22C89472FFF6FD45F90C969EF953" box="[680,726,1686,1709]" captionStart="Figures 5–8" captionStartId="5.[192,278,1276,1299]" captionTargetBox="[191,1438,191,1246]" captionTargetId="figure-6@5.[192,1438,192,1245]" captionTargetPageId="5" captionText="Figures 5–8: Buthacus stockmanni sp. n. Figures 5 and 7. Male holotype, carapace and tergites (5), coxosternal area and sternites III–V (7). Figures 6 and 8. Female paratype from the type locality, carapace and tergites (6), coxosternal area and sternites III–IV (8)." pageId="7" pageNumber="6">5, 7</figureCitation>
, 
<figureCitation id="0A8C22C89472FFF6FD08F90C9753F953" box="[741,795,1686,1709]" captionStart="Figures 9–14" captionStartId="6.[192,278,1287,1310]" captionTargetBox="[202,1429,202,1243]" captionTargetId="figure-17@6.[192,1437,192,1257]" captionTargetPageId="6" captionText="Figures 9–14: Buthacus stockmanni sp. n., metasoma and telson. Figures 9–11. Male holotype, lateral (9), ventral (10), and dorsal (11) views. Figures 12–14. Female paratype from the type locality, lateral (12), ventral (13), and dorsal (14) views. Scale bars: 10 mm." pageId="7" pageNumber="6">9–11</figureCitation>
, 
<figureCitation id="0A8C22C89472FFF6FF2DF92F951CF932" box="[192,340,1717,1740]" captionStart-0="Figures 15–29" captionStart-1="Figures 30–33" captionStartId-0="7.[192,278,1113,1136]" captionStartId-1="8.[192,269,1225,1246]" captionTargetBox-0="[192,1433,202,1078]" captionTargetBox-1="[194,1424,204,1193]" captionTargetId-0="figure-6@7.[192,1437,192,1083]" captionTargetId-1="figure-17@8.[192,1438,192,1196]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionText-0="Figures 15–29: Buthacus stockmanni sp. n. Figures 15–27: pedipalp segments. Figures 15–16. Female paratype from the type locality, chela dorsal (15) and external (16). Figures 17–27. Male holotype, chela dorsal (17), external (18), and ventral (19). Patella dorsal (20), external (21) and ventral (22). Femur internal (23), dorsal (24), and ventral (25). Movable finger (26) and fixed finger (27) dentition. Trichobothrial pattern is indicated in Figures 17–21 and 23–24. Figures 28–29. Telson lateral, female paratype from the type locality (28) and male holotype (29)." captionText-1="Figures 30–33: Buthacus stockmanni sp. n., male holotype, left legs I–IV, retrolateral aspect." pageId="7" pageNumber="6">17–27, 29–33</figureCitation>
, 
<figureCitation id="0A8C22C89472FFF6FE8CF92F9533F932" box="[353,379,1717,1740]" captionStart="Figures 39–40" captionStartId="10.[192,278,1790,1813]" captionTargetBox="[191,1440,215,1761]" captionTargetId="figure-17@10.[192,1439,216,1760]" captionTargetPageId="10" captionText="Figures 39–40: Buthacus stockmanni sp. n., in vivo habitus, male (39) holotype and female (40) paratype from the type locality." pageId="7" pageNumber="6">39</figureCitation>
) 
<specimenCount id="84B1F5C49472FFF6FE61F92F95E5F932" box="[396,429,1717,1740]" pageId="7" pageNumber="6" type="male">1♂</specimenCount>
<specimenCount id="84B1F5C49472FFF6FE40F92F9586F932" box="[429,462,1717,1740]" pageId="7" pageNumber="6" type="female">1♀</specimenCount>
(
<typeStatus id="4D0C80EF9472FFF6FE33F92F960FF932" box="[478,583,1717,1740]" pageId="7" pageNumber="6" type="paratype">paratypes</typeStatus>
, 
<figureCitation id="0A8C22C89472FFF6FDBFF92F96FEF932" box="[594,694,1717,1740]" captionStart="Figures 1–4" captionStartId="4.[192,278,1684,1707]" captionTargetBox="[205,1430,210,1642]" captionTargetId="figure-17@4.[192,1439,192,1655]" captionTargetPageId="4" captionText="Figures 1–4: Buthacus stockmanni sp. n. Figures 1–2. Male holotype in dorsal (1) and ventral (2) views. Figures 3–4. Female paratype from the type locality in dorsal (3) and ventral (4) aspects. Scale bar: 10 mm." pageId="7" pageNumber="6">Figs. 3–4</figureCitation>
, 
<figureCitation id="0A8C22C89472FFF6FD29F92F96A6F932" box="[708,750,1717,1740]" captionStart="Figures 5–8" captionStartId="5.[192,278,1276,1299]" captionTargetBox="[191,1438,191,1246]" captionTargetId="figure-6@5.[192,1438,192,1245]" captionTargetPageId="5" captionText="Figures 5–8: Buthacus stockmanni sp. n. Figures 5 and 7. Male holotype, carapace and tergites (5), coxosternal area and sternites III–V (7). Figures 6 and 8. Female paratype from the type locality, carapace and tergites (6), coxosternal area and sternites III–IV (8)." pageId="7" pageNumber="6">6, 8</figureCitation>
, 
<figureCitation id="0A8C22C89472FFF6FD17F92F9493F915" captionStart="Figures 9–14" captionStartId="6.[192,278,1287,1310]" captionTargetBox="[202,1429,202,1243]" captionTargetId="figure-17@6.[192,1437,192,1257]" captionTargetPageId="6" captionText="Figures 9–14: Buthacus stockmanni sp. n., metasoma and telson. Figures 9–11. Male holotype, lateral (9), ventral (10), and dorsal (11) views. Figures 12–14. Female paratype from the type locality, lateral (12), ventral (13), and dorsal (14) views. Scale bars: 10 mm." pageId="7" pageNumber="6">12– 14</figureCitation>
, 
<figureCitation id="0A8C22C89472FFF6FF00F94E9528F915" box="[237,352,1748,1771]" captionStart="Figures 15–29" captionStartId="7.[192,278,1113,1136]" captionTargetBox="[192,1433,202,1078]" captionTargetId="figure-6@7.[192,1437,192,1083]" captionTargetPageId="7" captionText="Figures 15–29: Buthacus stockmanni sp. n. Figures 15–27: pedipalp segments. Figures 15–16. Female paratype from the type locality, chela dorsal (15) and external (16). Figures 17–27. Male holotype, chela dorsal (17), external (18), and ventral (19). Patella dorsal (20), external (21) and ventral (22). Femur internal (23), dorsal (24), and ventral (25). Movable finger (26) and fixed finger (27) dentition. Trichobothrial pattern is indicated in Figures 17–21 and 23–24. Figures 28–29. Telson lateral, female paratype from the type locality (28) and male holotype (29)." pageId="7" pageNumber="6">15–16, 28</figureCitation>
, 
<figureCitation id="0A8C22C89472FFF6FE9DF94E95FBF915" box="[368,435,1748,1771]" captionStart="Figures 34–38" captionStartId="9.[192,278,1038,1061]" captionTargetBox="[200,1429,178,1002]" captionTargetId="figure-6@9.[192,1437,168,1008]" captionTargetPageId="9" captionText="Figures 34–38: Buthacus stockmanni sp. n., right hemispermatophore, male holotype. Figure 34. Whole hemispermatophore, dorsal view. Figures 35–38. Capsule region and flagellum: dorsal (35), external (36), ventral (37) and internal (38) views. Abbreviations: bl, basal lobe; el, external lobe, il, internal lobe; ml, median lobe. Scale bars: 1 mm (34), 500 µm (35–38)." pageId="7" pageNumber="6">34–38</figureCitation>
, 
<figureCitation id="0A8C22C89472FFF6FE2BF94E95A9F915" box="[454,481,1748,1771]" captionStart="Figures 39–40" captionStartId="10.[192,278,1790,1813]" captionTargetBox="[191,1440,215,1761]" captionTargetId="figure-17@10.[192,1439,216,1760]" captionTargetPageId="10" captionText="Figures 39–40: Buthacus stockmanni sp. n., in vivo habitus, male (39) holotype and female (40) paratype from the type locality." pageId="7" pageNumber="6">40</figureCitation>
, 
<figureCitation id="0A8C22C89472FFF6FE1EF94E967EF915" box="[499,566,1748,1771]" captionStart="Figures 43–44" captionStartId="12.[192,278,658,681]" captionTargetBox="[191,1437,191,628]" captionTargetId="figure-17@12.[192,1437,192,627]" captionTargetPageId="12" captionText="Figures 43–44: Buthacus stockmanni sp. n., mitotic metaphase (43) and ideogram (44) (y axis - % of the chromosome length of the diploid set, lines indicate min.-max. values) of paratype male from the type locality. Arrowheads indicate extra large chromosomes. Scale bar = 10 µm." pageId="7" pageNumber="6">43–44</figureCitation>
), leg. 
<collectorName id="3F425B9B9472FFF6FD68F94E976AF915" box="[645,802,1748,1771]" pageId="7" pageNumber="6">Omar Hassan.</collectorName>
</materialsCitation>
<materialsCitation id="22DF34109472FFF6FF2DF96897DFFAC3" collectingDate="2011-05-08" collectingDateMax="2011-05-10" collectingDateMin="2011-05-08" collectionCode="FKCP" collectorName="P. Kabatek &amp; Saquia el Hamra N &amp; P. Kabatek. All" country="Western Sahara" elevation="123" latitude="26.85" location="Laayoune" longLatPrecision="1239" longitude="-11.933333" pageId="7" pageNumber="6" specimenCount="8" specimenCount-female="4" specimenCount-male="4" typeStatus="paratype">
<emphasis id="A0C3E25F9472FFF6FF2DF9689530F8F7" bold="true" box="[192,376,1778,1801]" pageId="7" pageNumber="6">
<collectingCountry id="EAA07EDD9472FFF6FF2DF9689530F8F7" box="[192,376,1778,1801]" name="Western Sahara" pageId="7" pageNumber="6">Western Sahara</collectingCountry>
</emphasis>
, 
<quantity id="554F93A89472FFF6FE6BF9689582F8F7" box="[390,458,1778,1801]" metricMagnitude="4" metricUnit="m" metricValue="2.8" pageId="7" pageNumber="6" unit="km" value="28.0">28 km</quantity>
E 
<location id="976868969472FFF6FE05F968961CF8F7" LSID="urn:lsid:plazi:treatment:1A1E8F5B9472FFFCFEB5FA9C90C3F988:976868969472FFF6FE05F968961CF8F7" box="[488,596,1778,1801]" country="Western Sahara" latitude="26.85" longLatPrecision="1239" longitude="-11.933333" name="Laayoune" pageId="7" pageNumber="6">Laayoune</location>
, 
<geoCoordinate id="F783588A9472FFF6FDB3F96896FEF8F7" box="[606,694,1778,1801]" degrees="27" direction="north" minutes="13" orientation="latitude" pageId="7" pageNumber="6" precision="925" value="27.216667">27°13'N</geoCoordinate>
<geoCoordinate id="F783588A9472FFF6FD50F9689756F8F7" box="[701,798,1778,1801]" degrees="12" direction="west" minutes="54" orientation="longitude" pageId="7" pageNumber="6" precision="925" value="-12.9">12°54'W</geoCoordinate>
, 
<elevation id="199AD97E9472FFF6FF2DF88B950AF8D6" box="[192,322,1809,1832]" metricMagnitude="2" metricUnit="m" metricValue="1.23" pageId="7" pageNumber="6" unit="m" value="123.0">
<quantity id="554F93A89472FFF6FF2DF88B954FF8D6" box="[192,263,1809,1832]" metricMagnitude="2" metricUnit="m" metricValue="1.23" pageId="7" pageNumber="6" unit="m" value="123.0">123 m</quantity>
a.s.l.
</elevation>
, 
<date id="E609188D9472FFF6FEBFF88B958BF8D6" box="[338,451,1809,1832]" pageId="7" pageNumber="6" value="2011-05-10">
<collectingDate id="F64DE1659472FFF6FEBFF88B958BF8D6" box="[338,451,1809,1832]" pageId="7" pageNumber="6" value="2011-05-10">10.V.2011</collectingDate>
</date>
, 
<specimenCount id="84B1F5C49472FFF6FE3EF88B95BDF8D6" box="[467,501,1809,1832]" pageId="7" pageNumber="6" type="female">2♀</specimenCount>
<specimenCount id="84B1F5C49472FFF6FE18F88B965EF8D6" box="[501,534,1809,1832]" pageId="7" pageNumber="6" type="male">3♂</specimenCount>
juvs. (
<typeStatus id="4D0C80EF9472FFF6FDB7F88B968EF8D6" box="[602,710,1809,1832]" pageId="7" pageNumber="6" type="paratype">paratypes</typeStatus>
), leg. 
<collectorName id="3F425B9B9472FFF6FCE1F88B9555F8B9" pageId="7" pageNumber="6">P. Kabátek</collectorName>
; 
<collectorName id="3F425B9B9472FFF6FEC7F8AA95B6F8B9" box="[298,510,1840,1863]" pageId="7" pageNumber="6">Saquia el Hamra N</collectorName>
Smara, 
<geoCoordinate id="F783588A9472FFF6FDB1F8AA96FBF8B9" box="[604,691,1840,1863]" degrees="26" direction="north" minutes="51" orientation="latitude" pageId="7" pageNumber="6" precision="925" value="26.85">26°51'N</geoCoordinate>
<geoCoordinate id="F783588A9472FFF6FD50F8AA9756F8B9" box="[701,798,1840,1863]" degrees="11" direction="west" minutes="56" orientation="longitude" pageId="7" pageNumber="6" precision="925" value="-11.933333">11°56'W</geoCoordinate>
, 
<elevation id="199AD97E9472FFF6FF2DF8D49500F89B" box="[192,328,1870,1893]" metricMagnitude="2" metricUnit="m" metricValue="1.4" pageId="7" pageNumber="6" unit="m" value="140.0">
<quantity id="554F93A89472FFF6FF2DF8D49542F89B" box="[192,266,1870,1893]" metricMagnitude="2" metricUnit="m" metricValue="1.4" pageId="7" pageNumber="6" unit="m" value="140.0">140 m</quantity>
a.s.l.
</elevation>
, 
<date id="E609188D9472FFF6FEB0F8D495F7F89B" box="[349,447,1870,1893]" pageId="7" pageNumber="6" value="2011-05-08">
<collectingDate id="F64DE1659472FFF6FEB0F8D495F7F89B" box="[349,447,1870,1893]" pageId="7" pageNumber="6" value="2011-05-08">8.V.2011</collectingDate>
</date>
, 
<specimenCount id="84B1F5C49472FFF6FE39F8D495BDF89B" box="[468,501,1870,1893]" pageId="7" pageNumber="6" type="female">2♀</specimenCount>
<specimenCount id="84B1F5C49472FFF6FE18F8D4965EF89B" box="[501,534,1870,1893]" pageId="7" pageNumber="6" type="male">1♂</specimenCount>
juv. (
<typeStatus id="4D0C80EF9472FFF6FDBEF8D496F7F89B" box="[595,703,1870,1893]" pageId="7" pageNumber="6" type="paratype">paratypes</typeStatus>
), leg. 
<collectorName id="3F425B9B9472FFF6FCE1F8D49786FAE1" box="[780,974,1288,1893]" pageId="7" pageNumber="6">P. Kabátek. All</collectorName>
types are in the first authors collection (
<collectionCode id="F4A6A6889472FFF6FCABFABC97C7FAC3" box="[838,911,1318,1341]" pageId="7" pageNumber="6">FKCP</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="DAAD6DC69472FFF6FCD3FAFE9142FA67" pageId="7" pageNumber="6" type="etymology">
<paragraph id="92083E4D9472FFF6FCD3FAFE9142FA67" blockId="7.[830,1440,1380,1433]" pageId="7" pageNumber="6">
ETYMOLOGY. It is a pleasure to name this species after a scorpiologist Mark Stockmann (
<collectingCountry id="EAA07EDD9472FFF6FB7AFA1890B2FA67" box="[1175,1274,1410,1433]" name="Germany" pageId="7" pageNumber="6">Germany</collectingCountry>
).
</paragraph>
</subSubSection>
<subSubSection id="DAAD6DC69472FFF9FCD3FA5A9627F9C0" lastPageId="8" lastPageNumber="7" pageId="7" pageNumber="6" type="diagnosis">
<paragraph id="92083E4D9472FFF9FCD3FA5A9627F9C0" blockId="7.[830,1441,1472,1893]" lastBlockId="8.[192,803,1299,1598]" lastPageId="8" lastPageNumber="7" pageId="7" pageNumber="6">
DIAGNOSIS. Total length 
<quantity id="554F93A89472FFF6FBB7FA5A909AFA29" box="[1114,1234,1472,1495]" metricMagnitude="-2" metricUnit="m" metricValue="5.35" metricValueMax="5.9" metricValueMin="4.8" pageId="7" pageNumber="6" unit="mm" value="53.5" valueMax="59.0" valueMin="48.0">48–59 mm</quantity>
; carapace densely granulated with only anterior median carinae developed; anterior margin of carapace straight; pectine teeth 
<quantity id="554F93A89472FFF6FAB0FA67971CF9CD" metricMagnitude="-1" metricUnit="m" metricValue="6.604" metricValueMax="7.366" metricValueMin="5.842" pageId="7" pageNumber="6" unit="in" value="26.0" valueMax="29.0" valueMin="23.0">23–29 in</quantity>
males and 
<quantity id="554F93A89472FFF6FC22F9869067F9CD" box="[975,1071,1564,1587]" metricMagnitude="-1" metricUnit="m" metricValue="4.826" metricValueMax="5.588" metricValueMin="4.064" pageId="7" pageNumber="6" unit="in" value="19.0" valueMax="22.0" valueMin="16.0">16–22 in</quantity>
females; sternites III––VI lacking carinae; sternite VII with four smooth carinae; metasomal segments I–IV intercarinal surfaces smooth or almost smooth; metasomal segment V of both sexes ventraly granulated; metasomal segment I bears 10 carinae, segments II–IV bear 8 carinae (lateromedial carinae on segments II–III indicated by several granules on posterior part only); ventrolateral carinae of metasomal segment V bear several lobate granules in their posterior half; dorsal and lateral surfaces of segment V smooth, without granules in both sexes; all metasomal segments sparsely setose; metasomal segment V with ca. 35 long setae in both sexes; telson with long curved aculeus, longer than vesicle in both sexes; legs I–III with tarsal bristle combs composed of long, thin setae; tibial spurs present on legs III–IV, longer on leg IV and moderate on leg III; movable and fixed fingers of pedipalp with 9 rows of granules, with external and internal accessory granules (
<taxonomicName id="55B745CE947DFFF9FEDFFA739653F9FE" baseAuthorityName="Ehrenberg" baseAuthorityYear="1829" box="[306,539,1513,1536]" class="Arachnida" family="Buthidae" genus="Buthacus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="leptochelys">
<emphasis id="A0C3E25F947DFFF9FEDFFA739653F9FE" box="[306,539,1513,1536]" italics="true" pageId="8" pageNumber="7">Buthacus leptochelys</emphasis>
</taxonomicName>
complex). Males with broader pedipalp chelae than females, fingers twisted proximally in males, straight in females.
</paragraph>
</subSubSection>
<caption id="C6C86EC5947DFFF9FF2DFB53900DFB20" box="[192,1093,1225,1246]" pageId="8" pageNumber="7" startId="8.[192,269,1225,1246]" targetBox="[194,1424,204,1193]" targetPageId="8">
<paragraph id="92083E4D947DFFF9FF2DFB53900DFB20" blockId="8.[192,1093,1225,1246]" box="[192,1093,1225,1246]" pageId="8" pageNumber="7">
<emphasis id="A0C3E25F947DFFF9FF2DFB53951FFB20" bold="true" box="[192,343,1225,1246]" pageId="8" pageNumber="7">Figures 30–33:</emphasis>
<taxonomicName id="55B745CE947DFFF9FEB0FB539662FB20" authority="Kovařík &amp; Lowe &amp; Šťáhlavský, 2016" authorityName="Kovařík &amp; Lowe &amp; Šťáhlavský" authorityYear="2016" box="[349,554,1225,1246]" class="Arachnida" family="Buthidae" genus="Buthacus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="stockmanni" status="sp. nov.">
<emphasis id="A0C3E25F947DFFF9FEB0FB539662FB20" box="[349,554,1225,1246]" italics="true" pageId="8" pageNumber="7">Buthacus stockmanni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBF05F24947DFFF9FDDDFB53962EFB20" box="[560,614,1225,1246]" pageId="8" pageNumber="7" rank="species">
<emphasis id="A0C3E25F947DFFF9FDDDFB53960FFB20" bold="true" box="[560,583,1225,1246]" pageId="8" pageNumber="7">sp</emphasis>
. 
<emphasis id="A0C3E25F947DFFF9FDBEFB539628FB20" bold="true" box="[595,608,1225,1246]" pageId="8" pageNumber="7">n</emphasis>
.
</taxonomicNameLabel>
, male holotype, left legs I–IV, retrolateral aspect.
</paragraph>
</caption>
<subSubSection id="DAAD6DC6947DFFFCFF2DF9FE90C3F988" lastPageId="13" lastPageNumber="12" pageId="8" pageNumber="7" type="description">
<paragraph id="92083E4D947DFFF9FF2DF9FE969DF8CD" blockId="8.[192,809,1636,1904]" pageId="8" pageNumber="7">
DESCRIPTION. Adult males are 
<quantity id="554F93A8947DFFF9FDF5F9FE96C6F985" box="[536,654,1636,1659]" metricMagnitude="-2" metricUnit="m" metricValue="5.1" metricValueMax="5.4" metricValueMin="4.8" pageId="8" pageNumber="7" unit="mm" value="51.0" valueMax="54.0" valueMin="48.0">48–54 mm</quantity>
long and the adult females are 
<quantity id="554F93A8947DFFF9FE6FF91995BDF964" box="[386,501,1667,1690]" metricMagnitude="-2" metricUnit="m" metricValue="5.55" metricValueMax="5.9" metricValueMin="5.2" pageId="8" pageNumber="7" unit="mm" value="55.5" valueMax="59.0" valueMin="52.0">52–59 mm</quantity>
long. The habitus is shown in 
<figureCitation id="0A8C22C8947DFFF9FF0EF93B9505F946" box="[227,333,1697,1720]" captionStart="Figures 1–4" captionStartId="4.[192,278,1684,1707]" captionTargetBox="[205,1430,210,1642]" captionTargetId="figure-17@4.[192,1439,192,1655]" captionTargetPageId="4" captionText="Figures 1–4: Buthacus stockmanni sp. n. Figures 1–2. Male holotype in dorsal (1) and ventral (2) views. Figures 3–4. Female paratype from the type locality in dorsal (3) and ventral (4) aspects. Scale bar: 10 mm." pageId="8" pageNumber="7">Figs. 1–4</figureCitation>
. For position and distribution of trichobothria of pedipalps see 
<figureCitation id="0A8C22C8947DFFF9FE0AF95A9625F929" box="[487,621,1728,1751]" captionStart="Figures 15–29" captionStartId="7.[192,278,1113,1136]" captionTargetBox="[192,1433,202,1078]" captionTargetId="figure-6@7.[192,1437,192,1083]" captionTargetPageId="7" captionText="Figures 15–29: Buthacus stockmanni sp. n. Figures 15–27: pedipalp segments. Figures 15–16. Female paratype from the type locality, chela dorsal (15) and external (16). Figures 17–27. Male holotype, chela dorsal (17), external (18), and ventral (19). Patella dorsal (20), external (21) and ventral (22). Femur internal (23), dorsal (24), and ventral (25). Movable finger (26) and fixed finger (27) dentition. Trichobothrial pattern is indicated in Figures 17–21 and 23–24. Figures 28–29. Telson lateral, female paratype from the type locality (28) and male holotype (29)." pageId="8" pageNumber="7">Figs. 17–24</figureCitation>
. 
<emphasis id="A0C3E25F947DFFF9FD6EF95A9548F908" italics="true" pageId="8" pageNumber="7">Sexual dimorphism</emphasis>
: males have longer pectines than females (
<figureCitation id="0A8C22C8947DFFF9FD26F9459485F8EA" captionStart="Figures 5–8" captionStartId="5.[192,278,1276,1299]" captionTargetBox="[191,1438,191,1246]" captionTargetId="figure-6@5.[192,1438,192,1245]" captionTargetPageId="5" captionText="Figures 5–8: Buthacus stockmanni sp. n. Figures 5 and 7. Male holotype, carapace and tergites (5), coxosternal area and sternites III–V (7). Figures 6 and 8. Female paratype from the type locality, carapace and tergites (6), coxosternal area and sternites III–IV (8)." pageId="8" pageNumber="7">Figs. 7– 8</figureCitation>
) and broader pedipalp chelae than females, with fingers twisted proximally (
<figureCitation id="0A8C22C8947DFFF9FE04F886962FF8CD" box="[489,615,1820,1843]" captionStart="Figures 15–29" captionStartId="7.[192,278,1113,1136]" captionTargetBox="[192,1433,202,1078]" captionTargetId="figure-6@7.[192,1437,192,1083]" captionTargetPageId="7" captionText="Figures 15–29: Buthacus stockmanni sp. n. Figures 15–27: pedipalp segments. Figures 15–16. Female paratype from the type locality, chela dorsal (15) and external (16). Figures 17–27. Male holotype, chela dorsal (17), external (18), and ventral (19). Patella dorsal (20), external (21) and ventral (22). Femur internal (23), dorsal (24), and ventral (25). Movable finger (26) and fixed finger (27) dentition. Trichobothrial pattern is indicated in Figures 17–21 and 23–24. Figures 28–29. Telson lateral, female paratype from the type locality (28) and male holotype (29)." pageId="8" pageNumber="7">Figs. 15–19</figureCitation>
, 
<tableCitation id="DF350BF6947DFFF9FD99F886968DF8CD" box="[628,709,1820,1843]" captionStart="Table 1" captionStartId="12.[192,257,1420,1443]" captionText="Table 1: Comparative measurements of adults of Buthacus stockmanni sp. n. Abbreviations: length (L), width (W, in carapace it corresponds to posterior width), depth (D)." pageId="8" pageNumber="7">Table 1</tableCitation>
).
</paragraph>
<paragraph id="92083E4D947DFFF9FF2DF8A19760F88E" blockId="8.[192,809,1636,1904]" pageId="8" pageNumber="7">
<emphasis id="A0C3E25F947DFFF9FF2DF8A19573F8AC" bold="true" box="[192,315,1851,1874]" pageId="8" pageNumber="7">Coloration</emphasis>
(
<figureCitation id="0A8C22C8947DFFF9FEA0F8A195FEF8AC" box="[333,438,1851,1874]" captionStart-0="Figures 1–4" captionStart-1="Figures 5–8" captionStartId-0="4.[192,278,1684,1707]" captionStartId-1="5.[192,278,1276,1299]" captionTargetBox-0="[205,1430,210,1642]" captionTargetBox-1="[191,1438,191,1246]" captionTargetId-0="figure-17@4.[192,1439,192,1655]" captionTargetId-1="figure-6@5.[192,1438,192,1245]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionText-0="Figures 1–4: Buthacus stockmanni sp. n. Figures 1–2. Male holotype in dorsal (1) and ventral (2) views. Figures 3–4. Female paratype from the type locality in dorsal (3) and ventral (4) aspects. Scale bar: 10 mm." captionText-1="Figures 5–8: Buthacus stockmanni sp. n. Figures 5 and 7. Male holotype, carapace and tergites (5), coxosternal area and sternites III–V (7). Figures 6 and 8. Female paratype from the type locality, carapace and tergites (6), coxosternal area and sternites III–IV (8)." pageId="8" pageNumber="7">Figs. 1–8</figureCitation>
). Basic color is yellow to orange with darker carapace and mesosoma with characteristic
</paragraph>
<paragraph id="92083E4D947DFFF9FCD3FA8991E9FA78" blockId="8.[830,1448,1299,1904]" pageId="8" pageNumber="7">
orange to brown area on the anterior part of carapace. The chelicerae are yellow without reticulation; dentition is reddish to black. 
<emphasis id="A0C3E25F947DFFF9FCD3FAF597E4FA78" bold="true" box="[830,940,1391,1414]" pageId="8" pageNumber="7">Carapace</emphasis>
(
<figureCitation id="0A8C22C8947DFFF9FC52FAF59062FA78" box="[959,1066,1391,1414]" captionStart="Figures 5–8" captionStartId="5.[192,278,1276,1299]" captionTargetBox="[191,1438,191,1246]" captionTargetId="figure-6@5.[192,1438,192,1245]" captionTargetPageId="5" captionText="Figures 5–8: Buthacus stockmanni sp. n. Figures 5 and 7. Male holotype, carapace and tergites (5), coxosternal area and sternites III–V (7). Figures 6 and 8. Female paratype from the type locality, carapace and tergites (6), coxosternal area and sternites III–IV (8)." pageId="8" pageNumber="7">Figs. 5–6</figureCitation>
). The surface is granulated. The
</paragraph>
<paragraph id="92083E4D947DFFF9FCD3FA1791E8F9E1" blockId="8.[830,1448,1299,1904]" pageId="8" pageNumber="7">
anterior margin is straight and bears eight to ten macrosetae. Anterior median carinae coarsely granular. There are 5 lateral eyes on each side (3 larger, 2 smaller). 
<emphasis id="A0C3E25F947DFFF9FCD3F99297FEF9E1" bold="true" box="[830,950,1544,1567]" pageId="8" pageNumber="7">Mesosoma</emphasis>
(
<figureCitation id="0A8C22C8947DFFF9FC2BF9929079F9E1" box="[966,1073,1544,1567]" captionStart="Figures 5–8" captionStartId="5.[192,278,1276,1299]" captionTargetBox="[191,1438,191,1246]" captionTargetId="figure-6@5.[192,1438,192,1245]" captionTargetPageId="5" captionText="Figures 5–8: Buthacus stockmanni sp. n. Figures 5 and 7. Male holotype, carapace and tergites (5), coxosternal area and sternites III–V (7). Figures 6 and 8. Female paratype from the type locality, carapace and tergites (6), coxosternal area and sternites III–IV (8)." pageId="8" pageNumber="7">Figs. 5–8</figureCitation>
). The tergites bear three coarsely
</paragraph>
<paragraph id="92083E4D947DFFF9FCD3F9BD91E8F88E" blockId="8.[830,1448,1299,1904]" pageId="8" pageNumber="7">granular carinae, of which the lateral pair on the tergites I–II could be inconspicuous. All tergites with dense coarse and fine granulation. The pectinal tooth count is 23–29 (1 × 23, 5 × 24, 5 × 25, 1 × 26, 3 × 27, 2 × 28, 1 × 29) in males and 16–22 (2 × 16, 1 × 17, 2 × 18, 4 × 19, 1 × 20, 4 × 22) in females. The marginal tips of the pectines extend to the end of sternite IV in females, and to the anterior half of sternite V in males. The pectines have 3 marginal lamellae and 7–9 middle lamellae. The lamellae and fulcra bear numerous dark setae. Sternites III–VI lack carinae, and surfaces are smooth. Sternite</paragraph>
<caption id="C6C86EC5947CFFF8FF2DFB949107FBA8" pageId="9" pageNumber="8" startId="9.[192,278,1038,1061]" targetBox="[200,1429,178,1002]" targetPageId="9">
<paragraph id="92083E4D947CFFF8FF2DFB949107FBA8" blockId="9.[192,1441,1038,1110]" pageId="9" pageNumber="8">
<emphasis id="A0C3E25F947CFFF8FF2DFB949521FBDB" bold="true" box="[192,361,1038,1061]" pageId="9" pageNumber="8">Figures 34–38:</emphasis>
<taxonomicName id="55B745CE947CFFF8FE9CFB959677FBDA" authority="Kovařík &amp; Lowe &amp; Šťáhlavský, 2016" authorityName="Kovařík &amp; Lowe &amp; Šťáhlavský" authorityYear="2016" box="[369,575,1039,1060]" class="Arachnida" family="Buthidae" genus="Buthacus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="stockmanni" status="sp. nov.">
<emphasis id="A0C3E25F947CFFF8FE9CFB959677FBDA" box="[369,575,1039,1060]" italics="true" pageId="9" pageNumber="8">Buthacus stockmanni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBF05F24947CFFF8FDABFB959635FBDA" box="[582,637,1039,1060]" pageId="9" pageNumber="8" rank="species">
<emphasis id="A0C3E25F947CFFF8FDABFB959615FBDA" bold="true" box="[582,605,1039,1060]" pageId="9" pageNumber="8">sp</emphasis>
. 
<emphasis id="A0C3E25F947CFFF8FD87FB95963FFBDA" bold="true" box="[618,631,1039,1060]" pageId="9" pageNumber="8">n</emphasis>
.
</taxonomicNameLabel>
, right hemispermatophore, male holotype. 
<emphasis id="A0C3E25F947CFFF8FBC8FB9590C5FBDA" bold="true" box="[1061,1165,1039,1060]" pageId="9" pageNumber="8">Figure 34.</emphasis>
Whole hemispermatophore, dorsal view. 
<emphasis id="A0C3E25F947CFFF8FEA9FBB29596FBC3" bold="true" box="[324,478,1064,1085]" pageId="9" pageNumber="8">Figures 35–38.</emphasis>
Capsule region and flagellum: dorsal (35), external (36), ventral (37) and internal (38) views. Abbreviations: 
<emphasis id="A0C3E25F947CFFF8FEB8FBDB9520FBA8" box="[341,360,1089,1110]" italics="true" pageId="9" pageNumber="8">bl</emphasis>
, basal lobe; 
<emphasis id="A0C3E25F947CFFF8FE0CFBDB95BAFBA8" box="[481,498,1089,1110]" italics="true" pageId="9" pageNumber="8">el</emphasis>
, external lobe, 
<emphasis id="A0C3E25F947CFFF8FD6BFBDB96DBFBA8" box="[646,659,1089,1110]" italics="true" pageId="9" pageNumber="8">il</emphasis>
, internal lobe; 
<emphasis id="A0C3E25F947CFFF8FCC8FBDB9775FBA8" box="[805,829,1089,1110]" italics="true" pageId="9" pageNumber="8">ml</emphasis>
, median lobe. Scale bars: 1 mm (34), 500 µm (35–38).
</paragraph>
</caption>
<paragraph id="92083E4D947CFFF8FF2DFB1196B0FB3F" blockId="9.[192,802,1163,1891]" pageId="9" pageNumber="8">VII has two pairs of smooth carinae. All sternites bear many long macrosetae on their surfaces and margins.</paragraph>
<paragraph id="92083E4D947CFFF8FF2DFB529542F990" blockId="9.[192,802,1163,1891]" pageId="9" pageNumber="8">
<emphasis id="A0C3E25F947CFFF8FF2DFB5295E4FB21" bold="true" box="[192,428,1224,1247]" pageId="9" pageNumber="8">Hemispermatophore</emphasis>
(
<figureCitation id="0A8C22C8947CFFF8FE53FB529609FB21" box="[446,577,1224,1247]" captionStart="Figures 34–38" captionStartId="9.[192,278,1038,1061]" captionTargetBox="[200,1429,178,1002]" captionTargetId="figure-6@9.[192,1437,168,1008]" captionTargetPageId="9" captionText="Figures 34–38: Buthacus stockmanni sp. n., right hemispermatophore, male holotype. Figure 34. Whole hemispermatophore, dorsal view. Figures 35–38. Capsule region and flagellum: dorsal (35), external (36), ventral (37) and internal (38) views. Abbreviations: bl, basal lobe; el, external lobe, il, internal lobe; ml, median lobe. Scale bars: 1 mm (34), 500 µm (35–38)." pageId="9" pageNumber="8">Figs. 34–38</figureCitation>
). Flagelliform, relatively stout, trunk 3.7 times length of capsule region. Flagellum well separated from capsule lobes, pars recta long, 0.77 times length of trunk, tapered, proximal half broader, compressed, with narrow fin running along internal margin; pars reflecta long, narrow, hyaline, 0.94 times length of trunk. Capsule region with 4 lobes arranged in 3 +1 configuration typical of the &quot; 
<taxonomicName id="55B745CE947CFFF8FD24FA059751FA48" box="[713,793,1439,1462]" class="Arachnida" family="Buthidae" genus="Buthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="8" phylum="Arthropoda" rank="genus">Buthus</taxonomicName>
&quot; group (
<bibRefCitation id="F62643BC947CFFF8FEFDFA2495F5FA2B" author="FET" box="[272,445,1470,1493]" pageId="9" pageNumber="8" pagination="1 - 40" refId="ref7158" refString="FET, V., M. E. SOLEGLAD &amp; G. LOWE. 2005. A new trichobothrial character for the high - level systematics of Buthoidea (Scorpiones: Buthida). Euscorpius, 23: 1 - 40." type="journal article" year="2005">Fet et al., 2005</bibRefCitation>
; 
<bibRefCitation id="F62643BC947CFFF8FE3DFA2496FEFA2B" author="KOVARIK" box="[464,694,1470,1493]" pageId="9" pageNumber="8" pagination="1 - 133" refId="ref7481" refString="KOVARIK, F., G. LOWE, K. B. RANAWANA, D. HOFEREK, V. A. SANJEEWA JAYARATHNE, J. PLISKOVA &amp; F. STAHLAVSKY. 2016. Scorpions of Sri Lanka (Arachnida, Scorpiones: Buthidae, Chaerilidae, Scorpionidae) with description of four new species of the genera Charmus Karsch, 1879 and Reddyanus Vachon, 1972 stat. n .. Euscorpius, 220: 1 - 133." type="journal article" year="2016">Kovařík et al., 2016</bibRefCitation>
): external lobe largest, forming tapered lamina with blunt apex; median lobe smallest, acuminate, with weak carina on internal margin; internal lobe intermediate in size, laminate; basal lobe a small, rounded knob with weak distal flange.
</paragraph>
<paragraph id="92083E4D947CFFF8FF2DF9EC97D3F9CF" blockId="9.[192,802,1163,1891]" lastBlockId="9.[830,1441,1163,1891]" pageId="9" pageNumber="8">
<emphasis id="A0C3E25F947CFFF8FF2DF9EC95F2F973" bold="true" box="[192,442,1654,1677]" pageId="9" pageNumber="8">Metasoma and telson</emphasis>
(
<figureCitation id="0A8C22C8947CFFF8FE26F9EC9609F973" box="[459,577,1654,1677]" captionStart="Figures 9–14" captionStartId="6.[192,278,1287,1310]" captionTargetBox="[202,1429,202,1243]" captionTargetId="figure-17@6.[192,1437,192,1257]" captionTargetPageId="6" captionText="Figures 9–14: Buthacus stockmanni sp. n., metasoma and telson. Figures 9–11. Male holotype, lateral (9), ventral (10), and dorsal (11) views. Figures 12–14. Female paratype from the type locality, lateral (12), ventral (13), and dorsal (14) views. Scale bars: 10 mm." pageId="9" pageNumber="8">Figs. 9–14</figureCitation>
). Metasoma I bears 10 carinae, the ventromedial pair being obsolete. Metasoma II–III bear 8 carinae. Median lateral carinae are indicated by three to six granules on posterior part only. Ventromedial and ventrolateral carinae on metasoma II–III are granulated, with larger granules posteriorly. Metasoma IV bears 8 carinae from which both ventromedial and dorsal carinae are obsolete and only ventrolateral carinae are present with several granules. Metasoma V in both sexes has only ventrolateral carinae, which in posterior halves bear several lobate granules. Granules on the ventral surface of segment V form a median carina in both sexes. Intercarinal surfaces of segments I–IV are almost smooth in both sexes. Dorsal and lateral surfaces of segment V are smooth, without granules and carinae in both sexes. The anal arch consists of three or four lobes in both sexes. All segments are sparsely setose; segment V has ca. 35 long setae in both sexes. The telson with long curved aculeus, longer than vesicle in both sexes. The surface of the telson is smooth, sparsely hirsute, without a subaculear tubercle.
</paragraph>
<paragraph id="92083E4D947CFFFDFCD3F9A294B0F97C" blockId="9.[830,1441,1163,1891]" lastBlockId="12.[192,809,1520,1881]" lastPageId="12" lastPageNumber="11" pageId="9" pageNumber="8">
<emphasis id="A0C3E25F947CFFF8FCD3F9A297E5F9B1" bold="true" box="[830,941,1592,1615]" pageId="9" pageNumber="8">Pedipalps</emphasis>
(
<figureCitation id="0A8C22C8947CFFF8FC24F9A2901EF9B1" box="[969,1110,1592,1615]" captionStart="Figures 15–29" captionStartId="7.[192,278,1113,1136]" captionTargetBox="[192,1433,202,1078]" captionTargetId="figure-6@7.[192,1437,192,1083]" captionTargetPageId="7" captionText="Figures 15–29: Buthacus stockmanni sp. n. Figures 15–27: pedipalp segments. Figures 15–16. Female paratype from the type locality, chela dorsal (15) and external (16). Figures 17–27. Male holotype, chela dorsal (17), external (18), and ventral (19). Patella dorsal (20), external (21) and ventral (22). Femur internal (23), dorsal (24), and ventral (25). Movable finger (26) and fixed finger (27) dentition. Trichobothrial pattern is indicated in Figures 17–21 and 23–24. Figures 28–29. Telson lateral, female paratype from the type locality (28) and male holotype (29)." pageId="9" pageNumber="8">Figs. 15–27</figureCitation>
, 
<figureCitation id="0A8C22C8947CFFF8FB82F9A290C2F9B1" box="[1135,1162,1592,1615]" captionStart="Figures 45–51" captionStartId="13.[192,278,873,896]" captionTargetBox="[202,1437,176,841]" captionTargetId="figure-6@13.[192,1437,168,844]" captionTargetPageId="13" captionText="Figures 45–51: Figure 45. Buthacus stockmanni sp. n., male paratype from locality Morocco, North of Msied, 28.04383°N 10.85038°W, pedipalp chela external. Figures 46–48. Buthacus occidentalis, male holotype of Buthacus ehrenbergi, pedipalp chela external (46) and dorsal (47), and movable finger dentition (48). Figures 49–51. Buthacus ziegleri, male from locality Morocco, Merzouga env., 31°02'59&quot;N 04°00'16&quot;W, pedipalp chela external (49) and dorsal (50), and movable finger dentition (51)." pageId="9" pageNumber="8">45</figureCitation>
). The femur is finely granulated and bears three to five carinae; the ventroexternal carina is incomplete or absent, the other carinae are granular. The patella is smooth, with seven smooth and usually obsolete carinae in both sexes. The chela is smooth, with only incomplete smooth carinae indicated in females. The movable and fixed fingers are proximally twisted in males (
<figureCitation id="0A8C22C8947CFFF8FB7DF89590ADF8D8" box="[1168,1253,1807,1830]" captionStart="Figures 45–51" captionStartId="13.[192,278,873,896]" captionTargetBox="[202,1437,176,841]" captionTargetId="figure-6@13.[192,1437,168,844]" captionTargetPageId="13" captionText="Figures 45–51: Figure 45. Buthacus stockmanni sp. n., male paratype from locality Morocco, North of Msied, 28.04383°N 10.85038°W, pedipalp chela external. Figures 46–48. Buthacus occidentalis, male holotype of Buthacus ehrenbergi, pedipalp chela external (46) and dorsal (47), and movable finger dentition (48). Figures 49–51. Buthacus ziegleri, male from locality Morocco, Merzouga env., 31°02'59&quot;N 04°00'16&quot;W, pedipalp chela external (49) and dorsal (50), and movable finger dentition (51)." pageId="9" pageNumber="8">Fig. 45</figureCitation>
) and straight in females (
<figureCitation id="0A8C22C8947CFFF8FC4EF8B497BCF8BB" box="[931,1012,1838,1861]" captionStart="Figures 15–29" captionStartId="7.[192,278,1113,1136]" captionTargetBox="[192,1433,202,1078]" captionTargetId="figure-6@7.[192,1437,192,1083]" captionTargetPageId="7" captionText="Figures 15–29: Buthacus stockmanni sp. n. Figures 15–27: pedipalp segments. Figures 15–16. Female paratype from the type locality, chela dorsal (15) and external (16). Figures 17–27. Male holotype, chela dorsal (17), external (18), and ventral (19). Patella dorsal (20), external (21) and ventral (22). Femur internal (23), dorsal (24), and ventral (25). Movable finger (26) and fixed finger (27) dentition. Trichobothrial pattern is indicated in Figures 17–21 and 23–24. Figures 28–29. Telson lateral, female paratype from the type locality (28) and male holotype (29)." pageId="9" pageNumber="8">Fig. 16</figureCitation>
). All pedipalp segments including the trochanter are sparsely hirsute, with long, dark macro- setae in both sexes. The dentate margin of the movable and fixed fingers (
<figureCitation id="0A8C22C89479FFFDFE63F995965BF9D8" box="[398,531,1551,1574]" captionStart="Figures 15–29" captionStartId="7.[192,278,1113,1136]" captionTargetBox="[192,1433,202,1078]" captionTargetId="figure-6@7.[192,1437,192,1083]" captionTargetPageId="7" captionText="Figures 15–29: Buthacus stockmanni sp. n. Figures 15–27: pedipalp segments. Figures 15–16. Female paratype from the type locality, chela dorsal (15) and external (16). Figures 17–27. Male holotype, chela dorsal (17), external (18), and ventral (19). Patella dorsal (20), external (21) and ventral (22). Femur internal (23), dorsal (24), and ventral (25). Movable finger (26) and fixed finger (27) dentition. Trichobothrial pattern is indicated in Figures 17–21 and 23–24. Figures 28–29. Telson lateral, female paratype from the type locality (28) and male holotype (29)." pageId="12" pageNumber="11">Figs. 26–27</figureCitation>
) has nine rows of granules, each with one external and one internal granule, and 5 terminal granules (4 terminal and one basal terminal).
</paragraph>
<caption id="C6C86EC5947FFFFBFF2DF9649547F8D0" pageId="10" pageNumber="9" startId="10.[192,278,1790,1813]" targetBox="[191,1440,215,1761]" targetPageId="10">
<paragraph id="92083E4D947FFFFBFF2DF9649547F8D0" blockId="10.[192,1440,1790,1838]" pageId="10" pageNumber="9">
<emphasis id="A0C3E25F947FFFFBFF2DF964952CF8EB" bold="true" box="[192,356,1790,1813]" pageId="10" pageNumber="9">Figures 39–40</emphasis>
: 
<taxonomicName id="55B745CE947FFFFBFE98F89A960EF8EB" authority="Kovařík &amp; Lowe &amp; Šťáhlavský, 2016" authorityName="Kovařík &amp; Lowe &amp; Šťáhlavský" authorityYear="2016" box="[373,582,1792,1813]" class="Arachnida" family="Buthidae" genus="Buthacus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="stockmanni" status="sp. nov.">
<emphasis id="A0C3E25F947FFFFBFE98F89A960EF8EB" box="[373,582,1792,1813]" italics="true" pageId="10" pageNumber="9">Buthacus stockmanni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBF05F24947FFFFBFDBCF89A96C2F8EB" box="[593,650,1792,1813]" pageId="10" pageNumber="9" rank="species">
<emphasis id="A0C3E25F947FFFFBFDBCF89A9620F8EB" bold="true" box="[593,616,1792,1813]" pageId="10" pageNumber="9">sp</emphasis>
. 
<emphasis id="A0C3E25F947FFFFBFD95F89A96CDF8EB" bold="true" box="[632,645,1792,1813]" pageId="10" pageNumber="9">n</emphasis>
.
</taxonomicNameLabel>
, in vivo habitus, male (39) holotype and female (40) paratype from the type locality.
</paragraph>
</caption>
<caption id="C6C86EC5947EFFFAFF2DF944953DF8F0" pageId="11" pageNumber="10" startId="11.[192,278,1758,1781]" targetBox="[191,1440,199,1729]" targetPageId="11">
<paragraph id="92083E4D947EFFFAFF2DF944953DF8F0" blockId="11.[192,1441,1758,1806]" pageId="11" pageNumber="10">
<emphasis id="A0C3E25F947EFFFAFF2DF9449520F90B" bold="true" box="[192,360,1758,1781]" pageId="11" pageNumber="10">Figures 41–42</emphasis>
: 
<taxonomicName id="55B745CE947EFFFAFE93F97A961CF90B" authority="Kovařík &amp; Lowe &amp; Šťáhlavský, 2016" authorityName="Kovařík &amp; Lowe &amp; Šťáhlavský" authorityYear="2016" box="[382,596,1760,1781]" class="Arachnida" family="Buthidae" genus="Buthacus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="stockmanni" status="sp. nov.">
<emphasis id="A0C3E25F947EFFFAFE93F97A961CF90B" box="[382,596,1760,1781]" italics="true" pageId="11" pageNumber="10">Buthacus stockmanni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBF05F24947EFFFAFD8EF97A96E9F90B" box="[611,673,1760,1781]" pageId="11" pageNumber="10" rank="species">
<emphasis id="A0C3E25F947EFFFAFD8EF97A9632F90B" bold="true" box="[611,634,1760,1781]" pageId="11" pageNumber="10">sp</emphasis>
. 
<emphasis id="A0C3E25F947EFFFAFD63F97A96D3F90B" bold="true" box="[654,667,1760,1781]" pageId="11" pageNumber="10">n</emphasis>
.
</taxonomicNameLabel>
, type locality (41) and locality Morocco, North of Msied, 28.04383°N 10.85038°W. (42).
</paragraph>
</caption>
<caption id="C6C86EC59479FFFDFF2DFD089640FD24" pageId="12" pageNumber="11" startId="12.[192,278,658,681]" targetBox="[191,1437,191,628]" targetPageId="12">
<paragraph id="92083E4D9479FFFDFF2DFD089640FD24" blockId="12.[192,1440,658,730]" pageId="12" pageNumber="11">
<emphasis id="A0C3E25F9479FFFDFF2DFD089521FD57" bold="true" box="[192,361,658,681]" pageId="12" pageNumber="11">Figures 43–44:</emphasis>
<taxonomicName id="55B745CE9479FFFDFE9CFD099676FD56" authority="Kovařík &amp; Lowe &amp; Šťáhlavský, 2016" authorityName="Kovařík &amp; Lowe &amp; Šťáhlavský" authorityYear="2016" box="[369,574,659,680]" class="Arachnida" family="Buthidae" genus="Buthacus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="stockmanni" status="sp. nov.">
<emphasis id="A0C3E25F9479FFFDFE9CFD099676FD56" box="[369,574,659,680]" italics="true" pageId="12" pageNumber="11">Buthacus stockmanni</emphasis>
</taxonomicName>
<emphasis id="A0C3E25F9479FFFDFDABFD099635FD56" bold="true" box="[582,637,659,680]" pageId="12" pageNumber="11">
<taxonomicNameLabel id="BBF05F249479FFFDFDABFD099635FD56" box="[582,637,659,680]" pageId="12" pageNumber="11" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, mitotic metaphase (43) and ideogram (44) (y axis - % of the chromosome length of the diploid set, lines indicate min.-max. values) of paratype male from the type locality. Arrowheads indicate extra large chromosomes. Scale bar = 10 µm.
</paragraph>
</caption>
<paragraph id="92083E4D9479FFFDFCF8FCB8909EFA94" pageId="12" pageNumber="11">
<table id="E0B7CCED9479000EFEF7FCB8916DFA94" box="[282,1317,802,1386]" gridcols="4" gridrows="19" pageId="12" pageNumber="11">
<tr id="2C873C0F9479000EFEF7FCB8916DFCC7" box="[282,1317,802,825]" gridrow="0" pageId="12" pageNumber="11" rowspan-0="1" rowspan-1="1">
<th id="6F5655739479000EFCF8FCB897A7FCC7" box="[789,1007,802,825]" gridcol="2" gridrow="0" pageId="12" pageNumber="11">
<taxonomicName id="55B745CE9479FFFDFCF8FCB897E4FCC7" authority="Kovařík &amp; Lowe &amp; Šťáhlavský, 2016" authorityName="Kovařík &amp; Lowe &amp; Šťáhlavský" authorityYear="2016" box="[789,940,802,825]" class="Arachnida" family="Buthidae" genus="Buthacus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="stockmanni" status="sp. nov.">
<emphasis id="A0C3E25F9479FFFDFCF8FCB897E4FCC7" box="[789,940,802,825]" italics="true" pageId="12" pageNumber="11">B. stockmanni</emphasis>
</taxonomicName>
<emphasis id="A0C3E25F9479FFFDFC5FFCB897A7FCC7" bold="true" box="[946,1007,802,825]" pageId="12" pageNumber="11">
<taxonomicNameLabel id="BBF05F249479FFFDFC5FFCB897A7FCC7" box="[946,1007,802,825]" pageId="12" pageNumber="11" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</th>
<th id="6F5655739479000EFBA6FCB8916DFCC7" box="[1099,1317,802,825]" gridcol="3" gridrow="0" pageId="12" pageNumber="11">
<taxonomicName id="55B745CE9479FFFDFBA6FCB890AAFCC7" authority="Kovařík &amp; Lowe &amp; Šťáhlavský, 2016" authorityName="Kovařík &amp; Lowe &amp; Šťáhlavský" authorityYear="2016" box="[1099,1250,802,825]" class="Arachnida" family="Buthidae" genus="Buthacus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="stockmanni" status="sp. nov.">
<emphasis id="A0C3E25F9479FFFDFBA6FCB890AAFCC7" box="[1099,1250,802,825]" italics="true" pageId="12" pageNumber="11">B. stockmanni</emphasis>
</taxonomicName>
<emphasis id="A0C3E25F9479FFFDFB04FCB8916DFCC7" bold="true" box="[1257,1317,802,825]" pageId="12" pageNumber="11">
<taxonomicNameLabel id="BBF05F249479FFFDFB04FCB8916DFCC7" box="[1257,1317,802,825]" pageId="12" pageNumber="11" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</th>
</tr>
<tr id="2C873C0F9479000EFEF7FCDB916DFCA6" box="[282,1317,833,856]" gridrow="1" pageId="12" pageNumber="11" rowspan-1="1">
<th id="6F5655739479000EFEF7FCDB964EFCA6" box="[282,518,833,856]" gridcol="0" gridrow="1" pageId="12" pageNumber="11">
<emphasis id="A0C3E25F9479FFFDFED5FCDB964EFCA6" bold="true" box="[312,518,833,856]" pageId="12" pageNumber="11">Dimensions (MM)</emphasis>
</th>
<td id="6F5655739479000EFCF8FCDB97A7FCA6" box="[789,1007,833,856]" gridcol="2" gridrow="1" pageId="12" pageNumber="11">
♂ 
<emphasis id="A0C3E25F9479FFFDFCB3FCDB9788FCA6" bold="true" box="[862,960,833,856]" pageId="12" pageNumber="11">holotype</emphasis>
</td>
<td id="6F5655739479000EFBA6FCDB916DFCA6" box="[1099,1317,833,856]" gridcol="3" gridrow="1" pageId="12" pageNumber="11">
<emphasis id="A0C3E25F9479FFFDFB95FCDB90B0FCA6" bold="true" box="[1144,1272,833,856]" pageId="12" pageNumber="11">♀ paratype</emphasis>
</td>
</tr>
<tr id="2C873C0F9479000EFEF7FCFE916DFC85" box="[282,1317,868,891]" gridrow="2" pageId="12" pageNumber="11">
<th id="6F5655739479000EFEF7FCFE964EFC85" box="[282,518,868,891]" gridcol="0" gridrow="2" pageId="12" pageNumber="11">Carapace</th>
<td id="6F5655739479000EFDB0FCFE968BFC85" box="[605,707,868,891]" gridcol="1" gridrow="2" pageId="12" pageNumber="11">L / W</td>
<td id="6F5655739479000EFCF8FCFE97A7FC85" box="[789,1007,868,891]" gridcol="2" gridrow="2" pageId="12" pageNumber="11">5.65 / 6.35</td>
<td id="6F5655739479000EFBA6FCFE916DFC85" box="[1099,1317,868,891]" gridcol="3" gridrow="2" pageId="12" pageNumber="11">6.25 / 7.15</td>
</tr>
<tr id="2C873C0F9479000EFEF7FC18916DFC67" box="[282,1317,898,921]" gridrow="3" pageId="12" pageNumber="11">
<th id="6F5655739479000EFEF7FC18964EFC67" box="[282,518,898,921]" gridcol="0" gridrow="3" pageId="12" pageNumber="11">Mesosoma</th>
<td id="6F5655739479000EFDB0FC18968BFC67" box="[605,707,898,921]" gridcol="1" gridrow="3" pageId="12" pageNumber="11">L</td>
<td id="6F5655739479000EFCF8FC1897A7FC67" box="[789,1007,898,921]" gridcol="2" gridrow="3" pageId="12" pageNumber="11">12.40</td>
<td id="6F5655739479000EFBA6FC18916DFC67" box="[1099,1317,898,921]" gridcol="3" gridrow="3" pageId="12" pageNumber="11">13.40</td>
</tr>
<tr id="2C873C0F9479000EFEF7FC3B916DFC46" box="[282,1317,929,952]" gridrow="4" pageId="12" pageNumber="11">
<th id="6F5655739479000EFEF7FC3B964EFC46" box="[282,518,929,952]" gridcol="0" gridrow="4" pageId="12" pageNumber="11">Tergite VII</th>
<td id="6F5655739479000EFDB0FC3B968BFC46" box="[605,707,929,952]" gridcol="1" gridrow="4" pageId="12" pageNumber="11">L / W</td>
<td id="6F5655739479000EFCF8FC3B97A7FC46" box="[789,1007,929,952]" gridcol="2" gridrow="4" pageId="12" pageNumber="11">3.55 / 5.55</td>
<td id="6F5655739479000EFBA6FC3B916DFC46" box="[1099,1317,929,952]" gridcol="3" gridrow="4" pageId="12" pageNumber="11">3.95 / 6.95</td>
</tr>
<tr id="2C873C0F9479000EFEF7FC5A916DFC29" box="[282,1317,960,983]" gridrow="5" pageId="12" pageNumber="11">
<th id="6F5655739479000EFEF7FC5A964EFC29" box="[282,518,960,983]" gridcol="0" gridrow="5" pageId="12" pageNumber="11">Metasoma et telson</th>
<td id="6F5655739479000EFDB0FC5A968BFC29" box="[605,707,960,983]" gridcol="1" gridrow="5" pageId="12" pageNumber="11">L</td>
<td id="6F5655739479000EFCF8FC5A97A7FC29" box="[789,1007,960,983]" gridcol="2" gridrow="5" pageId="12" pageNumber="11">35.37</td>
<td id="6F5655739479000EFBA6FC5A916DFC29" box="[1099,1317,960,983]" gridcol="3" gridrow="5" pageId="12" pageNumber="11">37.88</td>
</tr>
<tr id="2C873C0F9479000EFEF7FC44916DFC0B" box="[282,1317,990,1013]" gridrow="6" pageId="12" pageNumber="11">
<th id="6F5655739479000EFEF7FC44964EFC0B" box="[282,518,990,1013]" gridcol="0" gridrow="6" pageId="12" pageNumber="11">Segment I</th>
<td id="6F5655739479000EFDB0FC44968BFC0B" box="[605,707,990,1013]" gridcol="1" gridrow="6" pageId="12" pageNumber="11">L / W / D</td>
<td id="6F5655739479000EFCF8FC4497A7FC0B" box="[789,1007,990,1013]" gridcol="2" gridrow="6" pageId="12" pageNumber="11">4.70 / 3.65 / 3.05</td>
<td id="6F5655739479000EFBA6FC44916DFC0B" box="[1099,1317,990,1013]" gridcol="3" gridrow="6" pageId="12" pageNumber="11">5.03 / 3.85 / 3.35</td>
</tr>
<tr id="2C873C0F9479000EFEF7FC67916DFBEA" box="[282,1317,1021,1044]" gridrow="7" pageId="12" pageNumber="11">
<th id="6F5655739479000EFEF7FC67964EFBEA" box="[282,518,1021,1044]" gridcol="0" gridrow="7" pageId="12" pageNumber="11">Segment II</th>
<td id="6F5655739479000EFDB0FC67968BFBEA" box="[605,707,1021,1044]" gridcol="1" gridrow="7" pageId="12" pageNumber="11">L / W / D</td>
<td id="6F5655739479000EFCF8FC6797A7FBEA" box="[789,1007,1021,1044]" gridcol="2" gridrow="7" pageId="12" pageNumber="11">5.65 / 3.55 / 3.05</td>
<td id="6F5655739479000EFBA6FC67916DFBEA" box="[1099,1317,1021,1044]" gridcol="3" gridrow="7" pageId="12" pageNumber="11">5.95 / 3.63 / 3.15</td>
</tr>
<tr id="2C873C0F9479000EFEF7FB86916DFBCD" box="[282,1317,1052,1075]" gridrow="8" pageId="12" pageNumber="11">
<th id="6F5655739479000EFEF7FB86964EFBCD" box="[282,518,1052,1075]" gridcol="0" gridrow="8" pageId="12" pageNumber="11">Segment III</th>
<td id="6F5655739479000EFDB0FB86968BFBCD" box="[605,707,1052,1075]" gridcol="1" gridrow="8" pageId="12" pageNumber="11">L / W / D</td>
<td id="6F5655739479000EFCF8FB8697A7FBCD" box="[789,1007,1052,1075]" gridcol="2" gridrow="8" pageId="12" pageNumber="11">5.67 / 3.30 / 2.85</td>
<td id="6F5655739479000EFBA6FB86916DFBCD" box="[1099,1317,1052,1075]" gridcol="3" gridrow="8" pageId="12" pageNumber="11">6.15 / 3.40 / 3.05</td>
</tr>
<tr id="2C873C0F9479000EFEF7FBA0916DFBAF" box="[282,1317,1082,1105]" gridrow="9" pageId="12" pageNumber="11">
<th id="6F5655739479000EFEF7FBA0964EFBAF" box="[282,518,1082,1105]" gridcol="0" gridrow="9" pageId="12" pageNumber="11">Segment IV</th>
<td id="6F5655739479000EFDB0FBA0968BFBAF" box="[605,707,1082,1105]" gridcol="1" gridrow="9" pageId="12" pageNumber="11">L / W / D</td>
<td id="6F5655739479000EFCF8FBA097A7FBAF" box="[789,1007,1082,1105]" gridcol="2" gridrow="9" pageId="12" pageNumber="11">6.20 / 2.80 / 2.55</td>
<td id="6F5655739479000EFBA6FBA0916DFBAF" box="[1099,1317,1082,1105]" gridcol="3" gridrow="9" pageId="12" pageNumber="11">6.60 / 2.95 / 2.75</td>
</tr>
<tr id="2C873C0F9479000EFEF7FBC3916DFB8E" box="[282,1317,1113,1136]" gridrow="10" pageId="12" pageNumber="11">
<th id="6F5655739479000EFEF7FBC3964EFB8E" box="[282,518,1113,1136]" gridcol="0" gridrow="10" pageId="12" pageNumber="11">Segment V</th>
<td id="6F5655739479000EFDB0FBC3968BFB8E" box="[605,707,1113,1136]" gridcol="1" gridrow="10" pageId="12" pageNumber="11">L / W / D</td>
<td id="6F5655739479000EFCF8FBC397A7FB8E" box="[789,1007,1113,1136]" gridcol="2" gridrow="10" pageId="12" pageNumber="11">7.05 / 2.75 / 2.45</td>
<td id="6F5655739479000EFBA6FBC3916DFB8E" box="[1099,1317,1113,1136]" gridcol="3" gridrow="10" pageId="12" pageNumber="11">7.35 / 2.90 / 2.70</td>
</tr>
<tr id="2C873C0F9479000EFEF7FBE2916DFB71" box="[282,1317,1144,1167]" gridrow="11" pageId="12" pageNumber="11">
<th id="6F5655739479000EFEF7FBE2964EFB71" box="[282,518,1144,1167]" gridcol="0" gridrow="11" pageId="12" pageNumber="11">Telson</th>
<td id="6F5655739479000EFDB0FBE2968BFB71" box="[605,707,1144,1167]" gridcol="1" gridrow="11" pageId="12" pageNumber="11">L / W / D</td>
<td id="6F5655739479000EFCF8FBE297A7FB71" box="[789,1007,1144,1167]" gridcol="2" gridrow="11" pageId="12" pageNumber="11">6.10 / 1.95 / 1.85</td>
<td id="6F5655739479000EFBA6FBE2916DFB71" box="[1099,1317,1144,1167]" gridcol="3" gridrow="11" pageId="12" pageNumber="11">6.80 / 2.25 / 2.03</td>
</tr>
<tr id="2C873C0F9479000EFEF7FB0C916DFB53" box="[282,1317,1174,1197]" gridrow="12" pageId="12" pageNumber="11">
<th id="6F5655739479000EFEF7FB0C964EFB53" box="[282,518,1174,1197]" gridcol="0" gridrow="12" pageId="12" pageNumber="11">Pedipalp</th>
<td id="6F5655739479000EFDB0FB0C968BFB53" box="[605,707,1174,1197]" gridcol="1" gridrow="12" pageId="12" pageNumber="11">L</td>
<td id="6F5655739479000EFCF8FB0C97A7FB53" box="[789,1007,1174,1197]" gridcol="2" gridrow="12" pageId="12" pageNumber="11">20.3</td>
<td id="6F5655739479000EFBA6FB0C916DFB53" box="[1099,1317,1174,1197]" gridcol="3" gridrow="12" pageId="12" pageNumber="11">20.61</td>
</tr>
<tr id="2C873C0F9479000EFEF7FB2F916DFB32" box="[282,1317,1205,1228]" gridrow="13" pageId="12" pageNumber="11">
<th id="6F5655739479000EFEF7FB2F964EFB32" box="[282,518,1205,1228]" gridcol="0" gridrow="13" pageId="12" pageNumber="11">Femur</th>
<td id="6F5655739479000EFDB0FB2F968BFB32" box="[605,707,1205,1228]" gridcol="1" gridrow="13" pageId="12" pageNumber="11">L / W</td>
<td id="6F5655739479000EFCF8FB2F97A7FB32" box="[789,1007,1205,1228]" gridcol="2" gridrow="13" pageId="12" pageNumber="11">5.10 / 1.55</td>
<td id="6F5655739479000EFBA6FB2F916DFB32" box="[1099,1317,1205,1228]" gridcol="3" gridrow="13" pageId="12" pageNumber="11">5.20 / 1.65</td>
</tr>
<tr id="2C873C0F9479000EFEF7FB4E916DFB15" box="[282,1317,1236,1259]" gridrow="14" pageId="12" pageNumber="11">
<th id="6F5655739479000EFEF7FB4E964EFB15" box="[282,518,1236,1259]" gridcol="0" gridrow="14" pageId="12" pageNumber="11">Patella</th>
<td id="6F5655739479000EFDB0FB4E968BFB15" box="[605,707,1236,1259]" gridcol="1" gridrow="14" pageId="12" pageNumber="11">L / W</td>
<td id="6F5655739479000EFCF8FB4E97A7FB15" box="[789,1007,1236,1259]" gridcol="2" gridrow="14" pageId="12" pageNumber="11">6.20 / 2.20</td>
<td id="6F5655739479000EFBA6FB4E916DFB15" box="[1099,1317,1236,1259]" gridcol="3" gridrow="14" pageId="12" pageNumber="11">6.40 / 2.20</td>
</tr>
<tr id="2C873C0F9479000EFEF7FB68916DFAF7" box="[282,1317,1266,1289]" gridrow="15" pageId="12" pageNumber="11">
<th id="6F5655739479000EFEF7FB68964EFAF7" box="[282,518,1266,1289]" gridcol="0" gridrow="15" pageId="12" pageNumber="11">Chela</th>
<td id="6F5655739479000EFDB0FB68968BFAF7" box="[605,707,1266,1289]" gridcol="1" gridrow="15" pageId="12" pageNumber="11">L</td>
<td id="6F5655739479000EFCF8FB6897A7FAF7" box="[789,1007,1266,1289]" gridcol="2" gridrow="15" pageId="12" pageNumber="11">9.0</td>
<td id="6F5655739479000EFBA6FB68916DFAF7" box="[1099,1317,1266,1289]" gridcol="3" gridrow="15" pageId="12" pageNumber="11">9.01</td>
</tr>
<tr id="2C873C0F9479000EFEF7FA8B916DFAD6" box="[282,1317,1297,1320]" gridrow="16" pageId="12" pageNumber="11">
<th id="6F5655739479000EFEF7FA8B964EFAD6" box="[282,518,1297,1320]" gridcol="0" gridrow="16" pageId="12" pageNumber="11">Manus</th>
<td id="6F5655739479000EFDB0FA8B968BFAD6" box="[605,707,1297,1320]" gridcol="1" gridrow="16" pageId="12" pageNumber="11">L / W / D</td>
<td id="6F5655739479000EFCF8FA8B97A7FAD6" box="[789,1007,1297,1320]" gridcol="2" gridrow="16" pageId="12" pageNumber="11">4.05 / 2.90 / 2.75</td>
<td id="6F5655739479000EFBA6FA8B916DFAD6" box="[1099,1317,1297,1320]" gridcol="3" gridrow="16" pageId="12" pageNumber="11">3.16 / 1.85 / 1.93</td>
</tr>
<tr id="2C873C0F9479000EFEF7FAAA916DFAB9" box="[282,1317,1328,1351]" gridrow="17" pageId="12" pageNumber="11">
<th id="6F5655739479000EFEF7FAAA964EFAB9" box="[282,518,1328,1351]" gridcol="0" gridrow="17" pageId="12" pageNumber="11">Movable finger</th>
<td id="6F5655739479000EFDB0FAAA968BFAB9" box="[605,707,1328,1351]" gridcol="1" gridrow="17" pageId="12" pageNumber="11">L</td>
<td id="6F5655739479000EFCF8FAAA97A7FAB9" box="[789,1007,1328,1351]" gridcol="2" gridrow="17" pageId="12" pageNumber="11">4.95</td>
<td id="6F5655739479000EFBA6FAAA916DFAB9" box="[1099,1317,1328,1351]" gridcol="3" gridrow="17" pageId="12" pageNumber="11">5.85</td>
</tr>
<tr id="2C873C0F9479000EFEF7FAC9916DFA94" box="[282,1317,1363,1386]" gridrow="18" pageId="12" pageNumber="11">
<th id="6F5655739479000EFEF7FAC9964EFA94" box="[282,518,1363,1386]" gridcol="0" gridrow="18" pageId="12" pageNumber="11">Total</th>
<td id="6F5655739479000EFDB0FAC9968BFA94" box="[605,707,1363,1386]" gridcol="1" gridrow="18" pageId="12" pageNumber="11">L</td>
<td id="6F5655739479000EFCF8FAC997A7FA94" box="[789,1007,1363,1386]" gridcol="2" gridrow="18" pageId="12" pageNumber="11">53.42</td>
<td id="6F5655739479000EFBA6FAC9916DFA94" box="[1099,1317,1363,1386]" gridcol="3" gridrow="18" pageId="12" pageNumber="11">57.53</td>
</tr>
</table>
</paragraph>
<caption id="C6C86EC59479FFFDFF2DFA169627FA45" pageId="12" pageNumber="11" startId="12.[192,257,1420,1443]">
<paragraph id="92083E4D9479FFFDFF2DFA169627FA45" blockId="12.[192,1440,1420,1467]" pageId="12" pageNumber="11">
<emphasis id="A0C3E25F9479FFFDFF2DFA169557FA5D" bold="true" box="[192,287,1420,1443]" pageId="12" pageNumber="11">Table 1:</emphasis>
Comparative measurements of adults of 
<taxonomicName id="55B745CE9479FFFDFD58FA1497CBFA5D" authority="Kovařík &amp; Lowe &amp; Šťáhlavský, 2016" authorityName="Kovařík &amp; Lowe &amp; Šťáhlavský" authorityYear="2016" box="[693,899,1422,1443]" class="Arachnida" family="Buthidae" genus="Buthacus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="stockmanni" status="sp. nov.">
<emphasis id="A0C3E25F9479FFFDFD58FA1497CBFA5D" box="[693,899,1422,1443]" italics="true" pageId="12" pageNumber="11">Buthacus stockmanni</emphasis>
</taxonomicName>
<emphasis id="A0C3E25F9479FFFDFC64FA149788FA5D" bold="true" box="[905,960,1422,1443]" pageId="12" pageNumber="11">
<taxonomicNameLabel id="BBF05F249479FFFDFC64FA149788FA5D" box="[905,960,1422,1443]" pageId="12" pageNumber="11" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
Abbreviations: length (L), width (W, in carapace it corresponds to posterior width), depth (D).
</paragraph>
</caption>
<paragraph id="92083E4D9479FFFDFF2DF910904FF9D8" blockId="12.[192,809,1520,1881]" lastBlockId="12.[830,1447,1520,1881]" pageId="12" pageNumber="11">
<emphasis id="A0C3E25F9479FFFDFF2DF91094BDF95F" bold="true" box="[192,245,1674,1697]" pageId="12" pageNumber="11">Legs</emphasis>
(
<figureCitation id="0A8C22C89479FFFDFEE4F91095C7F95F" box="[265,399,1674,1697]" captionStart="Figures 30–33" captionStartId="8.[192,269,1225,1246]" captionTargetBox="[194,1424,204,1193]" captionTargetId="figure-17@8.[192,1438,192,1196]" captionTargetPageId="8" captionText="Figures 30–33: Buthacus stockmanni sp. n., male holotype, left legs I–IV, retrolateral aspect." pageId="12" pageNumber="11">Figs. 30–33</figureCitation>
). The tarsomeres bear two rows of macrosetae on the ventral surface and numerous macrosetae on the other surfaces, which on legs I–III form bristle combs. The macrosetae are thin in both sexes. The femur and patella may bear indications of four to six carinae, which however are usually obsolete. The femur bears only solitary macrosetae. The tibial spur on leg IV is strong and longer than on the leg III where tibial spur is rather moderate.
</paragraph>
<paragraph id="92083E4D9479FFFDFCD3F9B49030F9BB" blockId="12.[830,1447,1520,1881]" box="[830,1144,1582,1605]" pageId="12" pageNumber="11">
<emphasis id="A0C3E25F9479FFFDFCD3F9B497ADF9BB" bold="true" box="[830,997,1582,1605]" pageId="12" pageNumber="11">Measurements</emphasis>
. See 
<tableCitation id="DF350BF69479FFFDFBCDF9B4903AF9BB" box="[1056,1138,1582,1605]" captionStart="Table 1" captionStartId="12.[192,257,1420,1443]" captionText="Table 1: Comparative measurements of adults of Buthacus stockmanni sp. n. Abbreviations: length (L), width (W, in carapace it corresponds to posterior width), depth (D)." pageId="12" pageNumber="11">Table 1</tableCitation>
.
</paragraph>
<paragraph id="92083E4D9479FFFCFCD3F9D694A3FB93" blockId="12.[830,1447,1520,1881]" lastBlockId="13.[192,802,1049,1133]" lastPageId="13" lastPageNumber="12" pageId="12" pageNumber="11">
<emphasis id="A0C3E25F9479FFFDFCD3F9D697F0F99D" bold="true" box="[830,952,1612,1635]" pageId="12" pageNumber="11">Karyotype</emphasis>
(
<figureCitation id="0A8C22C89479FFFDFC3EF9D69017F99D" box="[979,1119,1612,1635]" captionStart="Figures 43–44" captionStartId="12.[192,278,658,681]" captionTargetBox="[191,1437,191,628]" captionTargetId="figure-17@12.[192,1437,192,627]" captionTargetPageId="12" captionText="Figures 43–44: Buthacus stockmanni sp. n., mitotic metaphase (43) and ideogram (44) (y axis - % of the chromosome length of the diploid set, lines indicate min.-max. values) of paratype male from the type locality. Arrowheads indicate extra large chromosomes. Scale bar = 10 µm." pageId="12" pageNumber="11">Figs. 43–44</figureCitation>
). We analyzed one male 
<typeStatus id="4D0C80EF9479FFFDFCD3F9F197D2F97C" box="[830,922,1643,1666]" pageId="12" pageNumber="11" type="paratype">paratype</typeStatus>
from the type locality using standard cytogenetic methods (e.g. Kovařík et al., 2009). The diploid complement of this specimen is composed of 20 chromosomes (
<figureCitation id="0A8C22C89479FFFDFC04F95D900AF920" box="[1001,1090,1735,1758]" captionStart="Figures 43–44" captionStartId="12.[192,278,658,681]" captionTargetBox="[191,1437,191,628]" captionTargetId="figure-17@12.[192,1437,192,627]" captionTargetPageId="12" captionText="Figures 43–44: Buthacus stockmanni sp. n., mitotic metaphase (43) and ideogram (44) (y axis - % of the chromosome length of the diploid set, lines indicate min.-max. values) of paratype male from the type locality. Arrowheads indicate extra large chromosomes. Scale bar = 10 µm." pageId="12" pageNumber="11">Fig. 43</figureCitation>
). The chromosomes are holocentric and the meiosis is achiasmatic. Both characters are typical for the scorpions from the family 
<taxonomicName id="55B745CE9479FFFDFAADF89E91E9F8E5" authorityName="C. L. Koch" authorityYear="1837" box="[1344,1441,1796,1819]" class="Arachnida" family="Buthidae" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="11" phylum="Arthropoda" rank="family">Buthidae</taxonomicName>
(e.g. Mattos et al., 2013). The chromosomes of the first pair are significantly longer (13.41 % of the diploid set) than the remaining chromosomes that gradually decrease from 5.84 % to 2.69 % of the diploid set (
<figureCitation id="0A8C22C89478FFFCFD15FBAD9493FB93" captionStart="Figures 43–44" captionStartId="12.[192,278,658,681]" captionTargetBox="[191,1437,191,628]" captionTargetId="figure-17@12.[192,1437,192,627]" captionTargetPageId="12" captionText="Figures 43–44: Buthacus stockmanni sp. n., mitotic metaphase (43) and ideogram (44) (y axis - % of the chromosome length of the diploid set, lines indicate min.-max. values) of paratype male from the type locality. Arrowheads indicate extra large chromosomes. Scale bar = 10 µm." pageId="13" pageNumber="12">Fig. 44</figureCitation>
).
</paragraph>
<caption id="C6C86EC59478FFFCFF2DFCF394A7FC1A" pageId="13" pageNumber="12" startId="13.[192,278,873,896]" targetBox="[202,1437,176,841]" targetPageId="13">
<paragraph id="92083E4D9478FFFCFF2DFCF394A7FC1A" blockId="13.[192,1441,873,996]" pageId="13" pageNumber="12">
<emphasis id="A0C3E25F9478FFFCFF2DFCF3952AFC7E" bold="true" box="[192,354,873,896]" pageId="13" pageNumber="12">Figures 45–51</emphasis>
: 
<emphasis id="A0C3E25F9478FFFCFE9FFCF1959EFC7E" bold="true" box="[370,470,875,896]" pageId="13" pageNumber="12">Figure 45</emphasis>
. 
<taxonomicName id="55B745CE9478FFFCFE08FCF196FCFC7E" authority="Kovařík &amp; Lowe &amp; Šťáhlavský, 2016" authorityName="Kovařík &amp; Lowe &amp; Šťáhlavský" authorityYear="2016" box="[485,692,875,896]" class="Arachnida" family="Buthidae" genus="Buthacus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="13" pageNumber="12" phylum="Arthropoda" rank="species" species="stockmanni" status="sp. nov.">
<emphasis id="A0C3E25F9478FFFCFE08FCF196FCFC7E" box="[485,692,875,896]" italics="true" pageId="13" pageNumber="12">Buthacus stockmanni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBF05F249478FFFCFD50FCF196BCFC7E" box="[701,756,875,896]" pageId="13" pageNumber="12" rank="species">
<emphasis id="A0C3E25F9478FFFCFD50FCF1969CFC7E" bold="true" box="[701,724,875,896]" pageId="13" pageNumber="12">sp</emphasis>
. 
<emphasis id="A0C3E25F9478FFFCFD0FFCF196A7FC7E" bold="true" box="[738,751,875,896]" pageId="13" pageNumber="12">n</emphasis>
.
</taxonomicNameLabel>
, male paratype from locality Morocco, North of Msied, 28.04383°N 10.85038°W, pedipalp chela external. 
<emphasis id="A0C3E25F9478FFFCFDD1FC1E9686FC67" bold="true" box="[572,718,900,921]" pageId="13" pageNumber="12">Figures 46–48</emphasis>
. 
<taxonomicName id="55B745CE9478FFFCFD30FC1E97FBFC67" authorityName=": Lourenco" authorityYear="2000" box="[733,947,900,921]" class="Arachnida" family="Buthidae" genus="Buthacus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="13" pageNumber="12" phylum="Arthropoda" rank="species" species="occidentalis">
<emphasis id="A0C3E25F9478FFFCFD30FC1E97FBFC67" box="[733,947,900,921]" italics="true" pageId="13" pageNumber="12">Buthacus occidentalis</emphasis>
</taxonomicName>
, male holotype of 
<taxonomicName id="55B745CE9478FFFCFB9EFC1E9176FC67" authorityName="Kovarik" authorityYear="2005" box="[1139,1342,900,921]" class="Arachnida" family="Buthidae" genus="Buthacus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="13" pageNumber="12" phylum="Arthropoda" rank="species" species="ehrenbergi">
<emphasis id="A0C3E25F9478FFFCFB9EFC1E9176FC67" box="[1139,1342,900,921]" italics="true" pageId="13" pageNumber="12">Buthacus ehrenbergi</emphasis>
</taxonomicName>
, pedipalp chela external (46) and dorsal (47), and movable finger dentition (48). 
<emphasis id="A0C3E25F9478FFFCFC61FC079056FC4C" bold="true" box="[908,1054,925,946]" pageId="13" pageNumber="12">Figures 49–51</emphasis>
. 
<taxonomicName id="55B745CE9478FFFCFBC3FC079091FC4C" authorityName="Lourenco" authorityYear="2000" box="[1070,1241,925,946]" class="Arachnida" family="Buthidae" genus="Buthacus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="13" pageNumber="12" phylum="Arthropoda" rank="species" species="ziegleri">
<emphasis id="A0C3E25F9478FFFCFBC3FC079091FC4C" box="[1070,1241,925,946]" italics="true" pageId="13" pageNumber="12">Buthacus ziegleri</emphasis>
</taxonomicName>
, male from locality Morocco, Merzouga env., 31°02'59&quot;N 04°00'16&quot;W, pedipalp chela external (49) and dorsal (50), and movable finger dentition (51).
</paragraph>
</caption>
<paragraph id="92083E4D9478FFFCFF2DFB09951DF8EE" blockId="13.[192,802,1171,1808]" pageId="13" pageNumber="12">
AFFINITIES. The described features distinguish 
<taxonomicName id="55B745CE9478FFFCFD2FFB09954BFB37" authority="Kovařík &amp; Lowe &amp; Šťáhlavský, 2016" authorityName="Kovařík &amp; Lowe &amp; Šťáhlavský" authorityYear="2016" class="Arachnida" family="Buthidae" genus="Buthacus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="13" pageNumber="12" phylum="Arthropoda" rank="species" species="stockmanni" status="sp. nov.">
<emphasis id="A0C3E25F9478FFFCFD2FFB09954BFB37" italics="true" pageId="13" pageNumber="12">B. stockmanni</emphasis>
</taxonomicName>
<emphasis id="A0C3E25F9478FFFCFEFDFB28951AFB37" bold="true" box="[272,338,1202,1225]" pageId="13" pageNumber="12">
<taxonomicNameLabel id="BBF05F249478FFFCFEFDFB28951AFB37" box="[272,338,1202,1225]" pageId="13" pageNumber="12" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
from all other species of the genus. 
<taxonomicName id="55B745CE9478FFFCFCE6FB289572FB16" authority="Kovařík &amp; Lowe &amp; Šťáhlavský, 2016" authorityName="Kovařík &amp; Lowe &amp; Šťáhlavský" authorityYear="2016" class="Arachnida" family="Buthidae" genus="Buthacus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="13" pageNumber="12" phylum="Arthropoda" rank="species" species="stockmanni" status="sp. nov.">
<emphasis id="A0C3E25F9478FFFCFCE6FB289572FB16" italics="true" pageId="13" pageNumber="12">B. stockmanni</emphasis>
</taxonomicName>
<emphasis id="A0C3E25F9478FFFCFEA8FB4B95CDFB16" bold="true" box="[325,389,1233,1256]" pageId="13" pageNumber="12">
<taxonomicNameLabel id="BBF05F249478FFFCFEA8FB4B95CDFB16" box="[325,389,1233,1256]" pageId="13" pageNumber="12" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
is a member of the 
<taxonomicName id="55B745CE9478FFFCFD98FB4B954BFAF8" baseAuthorityName="Ehrenberg" baseAuthorityYear="1829" class="Arachnida" family="Buthidae" genus="Buthacus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="13" pageNumber="12" phylum="Arthropoda" rank="species" species="leptochelys">
<emphasis id="A0C3E25F9478FFFCFD98FB4B954BFAF8" italics="true" pageId="13" pageNumber="12">Buthacus leptochelys</emphasis>
</taxonomicName>
complex (see generic diagnosis on the top) which is represented in 
<collectingRegion id="5073F0AF9478FFFCFE7FFA94966BFADB" box="[402,547,1294,1317]" country="Sri Lanka" name="North Western" pageId="13" pageNumber="12">Northwestern</collectingRegion>
Africa by two other species 
<taxonomicName id="55B745CE9478FFFCFEF6FAB79622FABA" authority="Vachon, 1953" authorityName="Vachon" authorityYear="1953" box="[283,618,1325,1348]" class="Arachnida" family="Buthidae" genus="Buthacus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="13" pageNumber="12" phylum="Arthropoda" rank="species" species="occidentalis">
<emphasis id="A0C3E25F9478FFFCFEF6FAB79589FABA" box="[283,449,1325,1348]" italics="true" pageId="13" pageNumber="12">B. occidentalis</emphasis>
Vachon, 1953
</taxonomicName>
and 
<taxonomicName id="55B745CE9478FFFCFD40FAB7953CFA9C" authority="Lourenco, 2000" authorityName="Lourenco" authorityYear="2000" class="Arachnida" family="Buthidae" genus="Buthacus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="13" pageNumber="12" phylum="Arthropoda" rank="species" species="ziegleri">
<emphasis id="A0C3E25F9478FFFCFD40FAB7976AFABA" box="[685,802,1325,1348]" italics="true" pageId="13" pageNumber="12">B. ziegleri</emphasis>
<bibRefCitation id="F62643BC9478FFFCFF2DFAD1953CFA9C" author="LOURENCO" box="[192,372,1355,1378]" pageId="13" pageNumber="12" refId="ref7746" refString="LOURENCO, W. R. 2000. A new species of Buthacus Birula from Morocco (Arachnida: Scorpiones:" type="book" year="2000">Lourenço, 2000</bibRefCitation>
</taxonomicName>
. Both of these species are separable from the new species geographically (see 
<figureCitation id="0A8C22C89478FFFCFD75FAF096A1FA7F" box="[664,745,1386,1409]" captionStart="Figure 63" captionStartId="17.[192,268,1362,1385]" captionTargetBox="[192,1402,192,1332]" captionTargetId="figure-6@17.[192,1402,192,1332]" captionTargetPageId="17" captionText="Figure 63: Map showing confirmed distribution of species of Buthacus leptochelys complex in Northwestern Africa." pageId="13" pageNumber="12">Fig. 63</figureCitation>
) and morphologically. Males of 
<taxonomicName id="55B745CE9478FFFCFE08FA139634FA5E" authority="Kovařík &amp; Lowe &amp; Šťáhlavský, 2016" authorityName="Kovařík &amp; Lowe &amp; Šťáhlavský" authorityYear="2016" box="[485,636,1417,1440]" class="Arachnida" family="Buthidae" genus="Buthacus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="13" pageNumber="12" phylum="Arthropoda" rank="species" species="stockmanni" status="sp. nov.">
<emphasis id="A0C3E25F9478FFFCFE08FA139634FA5E" box="[485,636,1417,1440]" italics="true" pageId="13" pageNumber="12">B. stockmanni</emphasis>
</taxonomicName>
<emphasis id="A0C3E25F9478FFFCFD69FA139688FA5E" bold="true" box="[644,704,1417,1440]" pageId="13" pageNumber="12">
<taxonomicNameLabel id="BBF05F249478FFFCFD69FA139688FA5E" box="[644,704,1417,1440]" pageId="13" pageNumber="12" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
have the fingers of the pedipalp chela strongly twisted proximally (
<figureCitation id="0A8C22C89478FFFCFF25FA5C955FFA23" box="[200,279,1478,1501]" captionStart="Figures 45–51" captionStartId="13.[192,278,873,896]" captionTargetBox="[202,1437,176,841]" captionTargetId="figure-6@13.[192,1437,168,844]" captionTargetPageId="13" captionText="Figures 45–51: Figure 45. Buthacus stockmanni sp. n., male paratype from locality Morocco, North of Msied, 28.04383°N 10.85038°W, pedipalp chela external. Figures 46–48. Buthacus occidentalis, male holotype of Buthacus ehrenbergi, pedipalp chela external (46) and dorsal (47), and movable finger dentition (48). Figures 49–51. Buthacus ziegleri, male from locality Morocco, Merzouga env., 31°02'59&quot;N 04°00'16&quot;W, pedipalp chela external (49) and dorsal (50), and movable finger dentition (51)." pageId="13" pageNumber="12">Fig. 45</figureCitation>
), whereas males of both 
<taxonomicName id="55B745CE9478FFFCFDDDFA5C969AFA23" authorityName="Vachon" authorityYear="1953" box="[560,722,1478,1501]" class="Arachnida" family="Buthidae" genus="Buthacus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="13" pageNumber="12" phylum="Arthropoda" rank="species" species="occidentalis">
<emphasis id="A0C3E25F9478FFFCFDDDFA5C969AFA23" box="[560,722,1478,1501]" italics="true" pageId="13" pageNumber="12">B. occidentalis</emphasis>
</taxonomicName>
and 
<taxonomicName id="55B745CE9478FFFCFCE6FA5C9558FA02" authorityName="Lourenco" authorityYear="2000" class="Arachnida" family="Buthidae" genus="Buthacus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="13" pageNumber="12" phylum="Arthropoda" rank="species" species="ziegleri">
<emphasis id="A0C3E25F9478FFFCFCE6FA5C9558FA02" italics="true" pageId="13" pageNumber="12">B. ziegleri</emphasis>
</taxonomicName>
have the fingers straight or almost straight (
<figureCitation id="0A8C22C89478FFFCFD00FA7F957AF9E4" captionStart="Figures 45–51" captionStartId="13.[192,278,873,896]" captionTargetBox="[202,1437,176,841]" captionTargetId="figure-6@13.[192,1437,168,844]" captionTargetPageId="13" captionText="Figures 45–51: Figure 45. Buthacus stockmanni sp. n., male paratype from locality Morocco, North of Msied, 28.04383°N 10.85038°W, pedipalp chela external. Figures 46–48. Buthacus occidentalis, male holotype of Buthacus ehrenbergi, pedipalp chela external (46) and dorsal (47), and movable finger dentition (48). Figures 49–51. Buthacus ziegleri, male from locality Morocco, Merzouga env., 31°02'59&quot;N 04°00'16&quot;W, pedipalp chela external (49) and dorsal (50), and movable finger dentition (51)." pageId="13" pageNumber="12">Figs. 46 and 49</figureCitation>
). In the past 
<taxonomicName id="55B745CE9478FFFCFE23F9999620F9E4" authority="Kovařík &amp; Lowe &amp; Šťáhlavský, 2016" authorityName="Kovařík &amp; Lowe &amp; Šťáhlavský" authorityYear="2016" box="[462,616,1539,1562]" class="Arachnida" family="Buthidae" genus="Buthacus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="13" pageNumber="12" phylum="Arthropoda" rank="species" species="stockmanni" status="sp. nov.">
<emphasis id="A0C3E25F9478FFFCFE23F9999620F9E4" box="[462,616,1539,1562]" italics="true" pageId="13" pageNumber="12">B. stockmanni</emphasis>
</taxonomicName>
<emphasis id="A0C3E25F9478FFFCFD9EF99996FCF9E4" bold="true" box="[627,692,1539,1562]" pageId="13" pageNumber="12">
<taxonomicNameLabel id="BBF05F249478FFFCFD9EF99996FCF9E4" box="[627,692,1539,1562]" pageId="13" pageNumber="12" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
was confused with 
<taxonomicName id="55B745CE9478FFFCFEAEF9B895A0F9C7" authorityName="Vachon" authorityYear="1953" box="[323,488,1570,1593]" class="Arachnida" family="Buthidae" genus="Buthacus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="13" pageNumber="12" phylum="Arthropoda" rank="species" species="occidentalis">
<emphasis id="A0C3E25F9478FFFCFEAEF9B895A0F9C7" box="[323,488,1570,1593]" italics="true" pageId="13" pageNumber="12">B. occidentalis</emphasis>
</taxonomicName>
(see comments in 
<taxonomicName id="55B745CE9478FFFCFD24F9B8955DF9A6" authorityName="Vachon" authorityYear="1953" class="Arachnida" family="Buthidae" genus="Buthacus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="13" pageNumber="12" phylum="Arthropoda" rank="species" species="occidentalis">
<emphasis id="A0C3E25F9478FFFCFD24F9B8955DF9A6" italics="true" pageId="13" pageNumber="12">B. occidentalis</emphasis>
</taxonomicName>
section and fig. 
<quantity id="554F93A89478FFFCFE33F9DB9652F9A6" box="[478,538,1601,1624]" metricMagnitude="-1" metricUnit="m" metricValue="5.842" pageId="13" pageNumber="12" unit="in" value="23.0">23 in</quantity>
Lourenço, 2001: 267). These two species can also be differentiated by their tibial spurs which are long in 
<taxonomicName id="55B745CE9478FFFCFDE4F9E496E8F96B" authority="Kovařík &amp; Lowe &amp; Šťáhlavský, 2016" authorityName="Kovařík &amp; Lowe &amp; Šťáhlavský" authorityYear="2016" box="[521,672,1662,1685]" class="Arachnida" family="Buthidae" genus="Buthacus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="13" pageNumber="12" phylum="Arthropoda" rank="species" species="stockmanni" status="sp. nov.">
<emphasis id="A0C3E25F9478FFFCFDE4F9E496E8F96B" box="[521,672,1662,1685]" italics="true" pageId="13" pageNumber="12">B. stockmanni</emphasis>
</taxonomicName>
<emphasis id="A0C3E25F9478FFFCFD47F9E496A0F96B" bold="true" box="[682,744,1662,1685]" pageId="13" pageNumber="12">
<taxonomicNameLabel id="BBF05F249478FFFCFD47F9E496A0F96B" box="[682,744,1662,1685]" pageId="13" pageNumber="12" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="0A8C22C89478FFFCFD1AF9E49493F94A" captionStart="Figures 30–33" captionStartId="8.[192,269,1225,1246]" captionTargetBox="[194,1424,204,1193]" captionTargetId="figure-17@8.[192,1438,192,1196]" captionTargetPageId="8" captionText="Figures 30–33: Buthacus stockmanni sp. n., male holotype, left legs I–IV, retrolateral aspect." pageId="13" pageNumber="12">Fig. 33</figureCitation>
) versus moderate in 
<taxonomicName id="55B745CE9478FFFCFE2AF9079621F94A" authorityName="Vachon" authorityYear="1953" box="[455,617,1693,1716]" class="Arachnida" family="Buthidae" genus="Buthacus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="13" pageNumber="12" phylum="Arthropoda" rank="species" species="occidentalis">
<emphasis id="A0C3E25F9478FFFCFE2AF9079621F94A" box="[455,617,1693,1716]" italics="true" pageId="13" pageNumber="12">B. occidentalis</emphasis>
</taxonomicName>
(
<figureCitation id="0A8C22C89478FFFCFD97F9079682F94A" box="[634,714,1693,1716]" captionStart="Figures 52–58" captionStartId="14.[192,278,1331,1354]" captionTargetBox="[191,1437,191,1301]" captionTargetId="figure-17@14.[192,1437,192,1300]" captionTargetPageId="14" captionText="Figures 52–58: Figures 52–55. Buthacus occidentalis, male holotype of Buthacus ehrenbergi, telson lateral (52), carapace and tergites I–IV (53), and retrolateral aspect of leg IV (54) and leg III (55). Figures 56–58. Buthacus ziegleri, male from locality Morocco, Merzouga env., 31°02'59&quot;N 04°00'16&quot;W, telson lateral (56) and retrolateral aspect of leg IV (57) and leg III (58)." pageId="13" pageNumber="12">Fig. 54</figureCitation>
) on leg IV; and moderate in 
<taxonomicName id="55B745CE9478FFFCFE4AF9219676F92C" authority="Kovařík &amp; Lowe &amp; Šťáhlavský, 2016" authorityName="Kovařík &amp; Lowe &amp; Šťáhlavský" authorityYear="2016" box="[423,574,1723,1746]" class="Arachnida" family="Buthidae" genus="Buthacus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="13" pageNumber="12" phylum="Arthropoda" rank="species" species="stockmanni" status="sp. nov.">
<emphasis id="A0C3E25F9478FFFCFE4AF9219676F92C" box="[423,574,1723,1746]" italics="true" pageId="13" pageNumber="12">B. stockmanni</emphasis>
</taxonomicName>
<emphasis id="A0C3E25F9478FFFCFDA4F92196CEF92C" bold="true" box="[585,646,1723,1746]" pageId="13" pageNumber="12">
<taxonomicNameLabel id="BBF05F249478FFFCFDA4F92196CEF92C" box="[585,646,1723,1746]" pageId="13" pageNumber="12" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="0A8C22C89478FFFCFD7AF92196ADF92C" box="[663,741,1723,1746]" captionStart="Figures 30–33" captionStartId="8.[192,269,1225,1246]" captionTargetBox="[194,1424,204,1193]" captionTargetId="figure-17@8.[192,1438,192,1196]" captionTargetPageId="8" captionText="Figures 30–33: Buthacus stockmanni sp. n., male holotype, left legs I–IV, retrolateral aspect." pageId="13" pageNumber="12">Fig. 32</figureCitation>
) versus extremely reduced or absent in 
<taxonomicName id="55B745CE9478FFFCFDAAF94096AEF90F" authorityName="Vachon" authorityYear="1953" box="[583,742,1754,1777]" class="Arachnida" family="Buthidae" genus="Buthacus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="13" pageNumber="12" phylum="Arthropoda" rank="species" species="occidentalis">
<emphasis id="A0C3E25F9478FFFCFDAAF94096AEF90F" box="[583,742,1754,1777]" italics="true" pageId="13" pageNumber="12">B. occidentalis</emphasis>
</taxonomicName>
(
<figureCitation id="0A8C22C89478FFFCFD1BF9409493F8EE" captionStart="Figures 52–58" captionStartId="14.[192,278,1331,1354]" captionTargetBox="[191,1437,191,1301]" captionTargetId="figure-17@14.[192,1437,192,1300]" captionTargetPageId="14" captionText="Figures 52–58: Figures 52–55. Buthacus occidentalis, male holotype of Buthacus ehrenbergi, telson lateral (52), carapace and tergites I–IV (53), and retrolateral aspect of leg IV (54) and leg III (55). Figures 56–58. Buthacus ziegleri, male from locality Morocco, Merzouga env., 31°02'59&quot;N 04°00'16&quot;W, telson lateral (56) and retrolateral aspect of leg IV (57) and leg III (58)." pageId="13" pageNumber="12">Fig. 55</figureCitation>
) on leg III.
</paragraph>
<paragraph id="92083E4D9478FFFCFF2DF8AC9769F892" blockId="13.[192,802,1846,1900]" pageId="13" pageNumber="12">
<materialsCitation id="22DF34109478FFFCFF2DF8AC9769F892" collectionCode="LOCALITIES, AND, LIFE, STRATEGY" country="Morocco" location="North of Zag" pageId="13" pageNumber="12" specimenCount="1">
COMMENTS ON 
<collectionCode id="F4A6A6889478FFFCFE98F8A096AAF8B3" box="[373,738,1850,1869]" pageId="13" pageNumber="12">LOCALITIES AND LIFE STRATEGY</collectionCode>
. The 
<typeStatus id="4D0C80EF9478FFFCFF2DF8CF94A6F892" box="[192,238,1877,1900]" pageId="13" pageNumber="12">type</typeStatus>
locality (
<figureCitation id="0A8C22C89478FFFCFEBEF8CF95E8F892" box="[339,416,1877,1900]" captionStart="Figures 41–42" captionStartId="11.[192,278,1758,1781]" captionTargetBox="[191,1440,199,1729]" captionTargetId="figure-6@11.[192,1439,200,1728]" captionTargetPageId="11" captionText="Figures 41–42: Buthacus stockmanni sp. n., type locality (41) and locality Morocco, North of Msied, 28.04383°N 10.85038°W. (42)." pageId="13" pageNumber="12">Fig. 41</figureCitation>
, 
<collectingCountry id="EAA07EDD9478FFFCFE43F8CF9646F892" box="[430,526,1877,1900]" name="Morocco" pageId="13" pageNumber="12">Morocco</collectingCountry>
, 
<location id="976868969478FFFCFDF0F8CF96EFF892" LSID="urn:lsid:plazi:treatment:1A1E8F5B9472FFFCFEB5FA9C90C3F988:976868969478FFFCFDF0F8CF96EFF892" box="[541,679,1877,1900]" country="Morocco" name="North of Zag" pageId="13" pageNumber="12">North of Zag</location>
) is a sandy
</materialsCitation>
</paragraph>
<paragraph id="92083E4D9478FFFCFCD2FB8390C3F988" blockId="13.[830,1441,1049,1654]" pageId="13" pageNumber="12">
flat area surrounded by rock-strewn sand dunes with small shrubs and even some solitary trees. 
<taxonomicName id="55B745CE9478FFFCFAD0FBAD97F0FB93" authority="Kovařík &amp; Lowe &amp; Šťáhlavský, 2016" authorityName="Kovařík &amp; Lowe &amp; Šťáhlavský" authorityYear="2016" class="Arachnida" family="Buthidae" genus="Buthacus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="13" pageNumber="12" phylum="Arthropoda" rank="species" species="stockmanni" status="sp. nov.">
<emphasis id="A0C3E25F9478FFFCFAD0FBAD97F0FB93" italics="true" pageId="13" pageNumber="12">Buthacus stockmanni</emphasis>
</taxonomicName>
<emphasis id="A0C3E25F9478FFFCFC2FFBCC904AFB93" bold="true" box="[962,1026,1110,1133]" pageId="13" pageNumber="12">
<taxonomicNameLabel id="BBF05F249478FFFCFC2FFBCC904AFB93" box="[962,1026,1110,1133]" pageId="13" pageNumber="12" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
occurs on the sandy flat. During the day it is possible to find a few small juveniles rarely under rocks, but the adults were collected by UV lamps at night mostly sitting at the bases of shrubs. Males were more active at night than females, and were more often found running in open terrain. In addition, at this locality 
<taxonomicName id="55B745CE9478FFFCFCD2FA9490BDFADB" authority="(Audouin, 1825)" baseAuthorityName="Audouin" baseAuthorityYear="1825" box="[831,1269,1294,1317]" class="Arachnida" family="Buthidae" genus="Androctonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="13" pageNumber="12" phylum="Arthropoda" rank="species" species="amoreuxi">
<emphasis id="A0C3E25F9478FFFCFCD2FA94907EFADB" box="[831,1078,1294,1317]" italics="true" pageId="13" pageNumber="12">Androctonus amoreuxi</emphasis>
(Audouin, 1825)
</taxonomicName>
, 
<taxonomicName id="55B745CE9478FFFCFAE8FA949010FABA" authority="Lourenco, 2003" authorityName="Lourenco" authorityYear="2003" class="Arachnida" family="Buthidae" genus="Buthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="13" pageNumber="12" phylum="Arthropoda" rank="species" species="mariefranceae">
<emphasis id="A0C3E25F9478FFFCFAE8FA9497D3FABA" italics="true" pageId="13" pageNumber="12">Buthus mariefranceae</emphasis>
Lourenço, 2003
</taxonomicName>
, 
<taxonomicName id="55B745CE9478FFFCFB85FAB797F4FA9C" authority="Vachon, 1950" authorityName="Vachon" authorityYear="1950" class="Arachnida" family="Buthidae" genus="Compsobuthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="13" pageNumber="12" phylum="Arthropoda" rank="species" species="berlandi">
<emphasis id="A0C3E25F9478FFFCFB85FAB79125FABA" box="[1128,1389,1325,1348]" italics="true" pageId="13" pageNumber="12">Compsobuthus berlandi</emphasis>
Vachon, 1950
</taxonomicName>
and 
<taxonomicName id="55B745CE9478FFFCFBEDFAD191E8FA9C" authority="Vachon, 1950" authorityName="Vachon" authorityYear="1950" box="[1024,1440,1355,1378]" class="Arachnida" family="Buthidae" genus="Lissothus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="13" pageNumber="12" phylum="Arthropoda" rank="species" species="occidentalis">
<emphasis id="A0C3E25F9478FFFCFBEDFAD190BCFA9C" box="[1024,1268,1355,1378]" italics="true" pageId="13" pageNumber="12">Lissothus occidentalis</emphasis>
Vachon, 1950
</taxonomicName>
were recorded. Another locality (
<figureCitation id="0A8C22C89478FFFCFB27FAF0916CFA7F" box="[1226,1316,1386,1409]" captionStart="Figures 41–42" captionStartId="11.[192,278,1758,1781]" captionTargetBox="[191,1440,199,1729]" captionTargetId="figure-6@11.[192,1439,200,1728]" captionTargetPageId="11" captionText="Figures 41–42: Buthacus stockmanni sp. n., type locality (41) and locality Morocco, North of Msied, 28.04383°N 10.85038°W. (42)." pageId="13" pageNumber="12">Fig. 42</figureCitation>
, 
<collectingCountry id="EAA07EDD9478FFFCFAD4FAF091D4FA7F" box="[1337,1436,1386,1409]" name="Morocco" pageId="13" pageNumber="12">Morocco</collectingCountry>
, North of Msied) is a sandy flat area with just a few rocks. Rocky Queds (Wadis) were not far away. The whole area has a sparse cover of shrubs. In addition, at this locality 
<taxonomicName id="55B745CE9478FFFCFC21FA7F908DFA02" baseAuthorityName="Audouin" baseAuthorityYear="1825" box="[972,1221,1509,1532]" class="Arachnida" family="Buthidae" genus="Androctonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="13" pageNumber="12" phylum="Arthropoda" rank="species" species="amoreuxi">
<emphasis id="A0C3E25F9478FFFCFC21FA7F908DFA02" box="[972,1221,1509,1532]" italics="true" pageId="13" pageNumber="12">Androctonus amoreuxi</emphasis>
</taxonomicName>
was commonly recorded. Near the hill on the horizon in 
<figureCitation id="0A8C22C89478FFFCFB1AF999910EF9E4" box="[1271,1350,1539,1562]" captionStart="Figures 41–42" captionStartId="11.[192,278,1758,1781]" captionTargetBox="[191,1440,199,1729]" captionTargetId="figure-6@11.[192,1439,200,1728]" captionTargetPageId="11" captionText="Figures 41–42: Buthacus stockmanni sp. n., type locality (41) and locality Morocco, North of Msied, 28.04383°N 10.85038°W. (42)." pageId="13" pageNumber="12">Fig. 42</figureCitation>
, neither 
<taxonomicName id="55B745CE9478FFFCFCD3F9B8906DF9C7" authority="Kovařík &amp; Lowe &amp; Šťáhlavský, 2016" authorityName="Kovařík &amp; Lowe &amp; Šťáhlavský" authorityYear="2016" box="[830,1061,1570,1593]" class="Arachnida" family="Buthidae" genus="Buthacus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="13" pageNumber="12" phylum="Arthropoda" rank="species" species="stockmanni" status="sp. nov.">
<emphasis id="A0C3E25F9478FFFCFCD3F9B8906DF9C7" box="[830,1061,1570,1593]" italics="true" pageId="13" pageNumber="12">Buthacus stockmanni</emphasis>
</taxonomicName>
<emphasis id="A0C3E25F9478FFFCFBDDF9B89038F9C7" bold="true" box="[1072,1136,1570,1593]" pageId="13" pageNumber="12">
<taxonomicNameLabel id="BBF05F249478FFFCFBDDF9B89038F9C7" box="[1072,1136,1570,1593]" pageId="13" pageNumber="12" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
nor 
<taxonomicName id="55B745CE9478FFFCFB44F9B891E8F9C7" baseAuthorityName="Audouin" baseAuthorityYear="1825" box="[1193,1440,1570,1593]" class="Arachnida" family="Buthidae" genus="Androctonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="13" pageNumber="12" phylum="Arthropoda" rank="species" species="amoreuxi">
<emphasis id="A0C3E25F9478FFFCFB44F9B891E8F9C7" box="[1193,1440,1570,1593]" italics="true" pageId="13" pageNumber="12">Androctonus amoreuxi</emphasis>
</taxonomicName>
were found, but 
<taxonomicName id="55B745CE9478FFFCFBE8F9DB9123F9A6" authority="Turiel, 2014" authorityName="Turiel" authorityYear="2014" box="[1029,1387,1601,1624]" class="Arachnida" family="Buthidae" genus="Hottentotta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="13" pageNumber="12" phylum="Arthropoda" rank="species" species="sousai">
<emphasis id="A0C3E25F9478FFFCFBE8F9DB9099F9A6" box="[1029,1233,1601,1624]" italics="true" pageId="13" pageNumber="12">Hottentotta sousai</emphasis>
Turiel, 2014
</taxonomicName>
and 
<taxonomicName id="55B745CE9478FFFCFCD3F9C59789F988" box="[830,961,1631,1654]" class="Arachnida" family="Buthidae" genus="Orthochirus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="13" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="A0C3E25F9478FFFCFCD3F9C59789F988" box="[830,961,1631,1654]" italics="true" pageId="13" pageNumber="12">Orthochirus</emphasis>
</taxonomicName>
sp. were recorded.
</paragraph>
</subSubSection>
</treatment>
</document>