312 lines
32 KiB
XML
312 lines
32 KiB
XML
<document id="0A64318F16AA86FA80BEB2CEB4413114" ID-DOI="10.3897/dez.71.134268" ID-ZooBank="A1638EDA-A566-4837-A558-AA2707FBD0EB" ID-publisher-id="134268" URI-arpha="174753C4-D831-5CF5-A5C0-E6529D4F5A1C" URI-zoobank="http://zoobank.org/A1638EDA-A566-4837-A558-AA2707FBD0EB" article-type="research-article" checkinTime="1732027755009" checkinUser="pensoft" docAuthor="Liebherr, James K., Roig-Juñent, Sergio & Will, Kipling W." docDate="2024" docId="14FE6E48D8265989B1B8B4B003008764" docLanguage="en" docName="Deutsche Entomologische Zeitschrift 71(2): 319-338" docOrigin="Deutsche Entomologische Zeitschrift 71 (2)" docSource="https://dez.pensoft.net/article/134268/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Amaroxenus embersoni Liebherr & Roig-Juñent & Will 2024, sp. nov." docType="treatment" docUuid="45DA3BFE-373C-47A0-B736-B8406053EDAD" docUuidSource="ZooBank" docVersion="1" dtd-version="3.0" lastPageNumber="338" masterDocId="174753C4D8315CF5A5C0E6529D4F5A1C" masterDocTitle="Phylogenetic analysis of the circum-Antarctic Subfamily Migadopinae (Coleoptera, Carabidae) and assessment of the trans-Tasman Amarotypus clade" masterLastPageNumber="338" masterPageNumber="319" pageNumber="319" updateTime="1732027817041" updateUser="pensoft">
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<mods:title id="38F4A9ADC9C6CFD1EBADD2DA7A420A61">Phylogenetic analysis of the circum-Antarctic Subfamily Migadopinae (Coleoptera, Carabidae) and assessment of the trans-Tasman Amarotypus clade</mods:title>
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<mods:name id="C6CC6E0F1BFF1BAB525176F112160032" type="personal">
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<mods:namePart id="33CCCB0E2B2D90F045AD8C4AECD45CF3">Liebherr, James K.</mods:namePart>
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<mods:affiliation id="E7AFF02F5B5466C1FCE38CDA4DE03CEC">Cornell University Insect Collection, Ithaca, NY 14853 - 2601 USA</mods:affiliation>
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</mods:name>
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<mods:name id="E0E20ECA509230FF13EE0D6AACD7EDE7" type="personal">
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<mods:roleTerm id="C1E32A148334F604A8B633BB8C3F160E">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="D72236801D56C1B2A4BAB4EF0A51CD4C">Roig-Juñent, Sergio</mods:namePart>
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<mods:nameIdentifier id="232EE7306F3648F1CA2FD4D25CEDE5A3" type="ORCID">0000-0002-3619-5351</mods:nameIdentifier>
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<mods:affiliation id="A963AFF54122EC3037D8A3F889752024">Laboratorio de Entomología, Instituto Argentina de Investigaciones de las Zonas Áridas (IADIZA, CCT – CONICET, Mendoza), 5500 Mendoza, Argentina</mods:affiliation>
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<mods:namePart id="33A79AB84D5A53FCA8EDFF90F99B267C">Will, Kipling W.</mods:namePart>
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<mods:nameIdentifier id="790B69247EB6EF56E1DAAC39D2E64F66" type="ORCID">0000-0002-7056-9011</mods:nameIdentifier>
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<mods:affiliation id="AF5211A4FC96A48D1CA0866D5FEE2A23">Essig Museum of Entomology, University of California, Berkeley, CA, 94720 - 3112 USA</mods:affiliation>
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<mods:title id="1D6795537D059A983EA61D53E950F055">Deutsche Entomologische Zeitschrift</mods:title>
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<mods:part id="23E8A07DD88CD984EAD24033E981F7F4">
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<mods:date id="C586ECF4DDFEFC40251A08041CBC327C">2024</mods:date>
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<mods:number id="E7C078611A6AE4EF1DF081DDCAED7717">2024-11-19</mods:number>
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<mods:number id="DCE446103DE2FF7C20603AF741DD3FDB">71</mods:number>
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<treatment id="14FE6E48D8265989B1B8B4B003008764" ID-arpha="14FE6E48-D826-5989-B1B8-B4B003008764" ID-zoobank="https://zoobank.org/45DA3BFE-373C-47A0-B736-B8406053EDAD" LSID="urn:lsid:zoobank.org:act:45DA3BFE-373C-47A0-B736-B8406053EDAD" LSID-ZBK="urn:lsid:zoobank.org:act:45DA3BFE-373C-47A0-B736-B8406053EDAD" httpUri="http://treatment.plazi.org/id/14FE6E48D8265989B1B8B4B003008764">
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<subSubSection id="CB1248DA2C4C4F901337DA2FF6598FD4" type="nomenclature">
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||
<paragraph id="D6D1F7AC1642AE47F7279B9596057D2A">
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||
<taxonomicName id="A5AD26F4026072A4F39C919C5C0B174E" authority="Liebherr & Will" authorityName="Liebherr & Roig-Juñent & Will" authorityYear="2024" class="Insecta" family="Carabidae" genus="Amaroxenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="embersoni" status="sp. nov.">
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<emphasis id="9114274878BBCCF6DE07C27F9C103CA9" italics="true">Amaroxenus embersoni</emphasis>
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Liebherr & Will
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</taxonomicName>
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<taxonomicNameLabel id="9355F963C5B99A7E58E5F7072E63C848" rank="species">sp. nov.</taxonomicNameLabel>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="SECID0ENTBG" type="Types">
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||
<paragraph id="F2FE2E2AD59F22452D6DFD40325DC997">
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||
<heading id="8915C62407520643CED560905F373655" reason="title">Types.</heading>
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||
</paragraph>
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||
<paragraph id="B273C7FF76D7082B0809CF285AF27B17">
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||
<emphasis id="88BB177C571E15717CE25E616A567BBB" bold="true" italics="true">
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<typeStatus id="BAF5DEA7C699A2902917DB868EA1105A" type="holotype">Holotype</typeStatus>
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</emphasis>
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male (
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<named-content id="3B76AC952F7B91C373AF5DA8063C2481" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/lincoln-university-entomology-research-museum" xlink_title="Lincoln University Entomology Research Collection">
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<collectionCode id="C94B1B3648C9724044EAC0009E879D49" country="New Zealand" httpUri="http://grbio.org/cool/n5zm-bhpt" name="Lincoln University Entomology Research Museum" type="">LUNZ</collectionCode>
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</named-content>
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) mislabeled:
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<emphasis id="2BA6C3F36AC5D62B53C7EFDD3EEA4940" italics="true">
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Arthurs Pass Nat. / Pk, NC
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<quantity id="97754CEAEA424686E132515049B28F0D" metricMagnitude="3" metricUnit="m" metricValue="1.65" unit="m" value="1650.0">1650 m</quantity>
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/ Mt. Philistine /
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<date id="AA61D2EF64035F54C0DAB96FE7F63111" value="1988-01-01">1 - i- 1988</date>
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P. Syrett /
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<collectionCode id="020D1DB1A67A5C2467FC4122DE3C2E4D" country="Chile" name="Departamento de Geologia, Universidad de Chile">R</collectionCode>
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. M. Emberson // under rock / in fine damp / gravel
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</emphasis>
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//
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<typeStatus id="684150230B297AAD183D8BBBF9A247CA" type="holotype">HOLOTYPE</typeStatus>
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<specimenCount id="ACEB853D356DF851B9FAB43F0F3449B5" count="1" type="male">♂</specimenCount>
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/
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<taxonomicName id="46AA8D3ED27D109C95FBC433EFD5AD66" class="Insecta" family="Carabidae" genus="Amaroxenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
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<emphasis id="A593AC7E794B023F1C592DD92FDD6305" italics="true">Amaroxenus</emphasis>
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</taxonomicName>
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/
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<taxonomicName id="1A05C68E14205CE957133F7F905A5557" authorityName="Liebherr & Roig-Juñent & Will" authorityYear="2024" class="Insecta" family="Carabidae" genus="Amaroxenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="embersoni">
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<emphasis id="FBE7C29B434935794E329559E48A5962" italics="true">embersoni</emphasis>
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</taxonomicName>
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/ Liebherr and Will 2023 (black-bordered red label). Based on consultation with Emberson field notes (P. Syrett pers. comm.) the
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<typeStatus id="A784372909BDE9D9A52FE02E43407BBD" type="holotype">holotype</typeStatus>
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was actually collected at: Arthur’s Pass N. P., Mt. Aicken,
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<quantity id="258EBBCB9686D8D9DB1BC7409F75DBD6" metricMagnitude="3" metricUnit="m" metricValue="1.75" unit="m" value="1750.0">1750 m</quantity>
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,
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<date id="57DEA64201A769919B5E0545062335F9" value="1988-01-01">1 - i- 1988</date>
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(Fig.
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<figureCitation id="8CADFD8E1E93129F9B0EE84C53E15C92" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Type locality of Amaroxenus embersoni showing Professor Rowan Emberson collecting the type series (photo courtesy Pol Syrett)." figureDoi="10.3897/dez.71.134268.figure7" httpUri="https://binary.pensoft.net/fig/1177808">7</figureCitation>
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).
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<emphasis id="3DC0D8D91187E9A15417F9450B353908" bold="true" italics="true">
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<typeStatus id="A7A7923676508BA0C54DBBA23215AD9F" type="paratype">Paratypes</typeStatus>
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</emphasis>
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: same incorrect data label as
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<typeStatus id="52D168B0ED10565A6243D39538637A59" type="holotype">holotype</typeStatus>
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||
(
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<named-content id="0A95149B93FFF22D9081C256EA058F7C" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/lincoln-university-entomology-research-museum" xlink_title="Lincoln University Entomology Research Collection">
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||
<collectionCode id="AB1D91D5842C923E8A774007666B7294" country="New Zealand" httpUri="http://grbio.org/cool/n5zm-bhpt" name="Lincoln University Entomology Research Museum" type="">LUNZ</collectionCode>
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||
</named-content>
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,
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<specimenCount id="83016E56FA829A14F53D015D524A3CCA" count="1" type="male">1 ♂</specimenCount>
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);
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<collectingCountry id="FC8841AB4478D64FAE1291BB4F0DEE9E" name="New Zealand">New Zealand</collectingCountry>
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: Arthur’s Pass N. P., Mt. Aicken,
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<quantity id="00DD360EDB0BC341F5C2B98DCD6E4A81" metricMagnitude="3" metricUnit="m" metricValue="1.75" unit="m" value="1750.0">1750 m</quantity>
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||
,
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<date id="3198CACC66CAC8C155172FA377FB3DF1" value="1988-01-01">1 - i- 1988</date>
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,
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<collectionCode id="98001BA46A8287B18266F22E879EFB86" country="Chile" name="Departamento de Geologia, Universidad de Chile">R</collectionCode>
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. M. Emberson and P. Syrett, under rock sparse fell field vegetation (
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||
<named-content id="F5928D6EA1677E1FA521FF19FB2ACC5B" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/lincoln-university-entomology-research-museum" xlink_title="Lincoln University Entomology Research Collection">
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||
<collectionCode id="D7558F8DB771376678949B97EADFBE27" country="New Zealand" httpUri="http://grbio.org/cool/n5zm-bhpt" name="Lincoln University Entomology Research Museum" type="">LUNZ</collectionCode>
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||
</named-content>
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,
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<specimenCount id="FC67E8E951B1F8A75F7083940D5BC38A" count="1" type="female">1 ♀</specimenCount>
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) [as noted above, these data are appropriate for all
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<specimenCount id="51A98B93852711E526823784357B4CB0" count="3" type="generic">three specimens</specimenCount>
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of
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<taxonomicName id="C7A125190BA891102ECCB4AEFF2524BD" authorityName="Liebherr & Roig-Juñent & Will" authorityYear="2024" class="Insecta" family="Carabidae" genus="Amaroxenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="embersoni">
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<emphasis id="F2574423042C2D3E439A177EA389B321" italics="true">A. embersoni</emphasis>
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</taxonomicName>
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].
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</paragraph>
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</subSubSection>
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<subSubSection id="SECID0EHVBG" type="diagnosis">
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<paragraph id="56FE2DEC2B1E64BAE13DC10F9D1B6522">
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<heading id="0059F5996DCBEAF3CA8C479A81A9A0F5" reason="title">Diagnosis.</heading>
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</paragraph>
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<paragraph id="97C7F34CBD8B648F2DB4351949ADF6BC">
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A broad-bodied species, with transverse pronotum,
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<abbrev id="ABBRID0ENVBG" xlink_title="maximal pronotal width">MPW</abbrev>
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/
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<abbrev id="ABBRID0ERVBG" xlink_title="pronotal length measured along the midline">
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<collectionCode id="4D6CDD4D86BF398EA14969989AB298D5" country="Czech Republic" name="Západoceské muzeum v Plzni" type="Museum">PL</collectionCode>
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</abbrev>
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= 1.42, and broadly ovate elytra,
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<abbrev id="ABBRID0EVVBG" xlink_title="maximal elytral width">MEW</abbrev>
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/
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<abbrev id="ABBRID0EZVBG" xlink_title="elytral length measured from the base of the scutellum to the apex of the left elytron">EL</abbrev>
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= 0.76 (Fig.
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<figureCitation id="91D90BF941AC9532B8372121A0D0A67F" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Representative taxa of the Amarotypus clade, dorsal view. A. Amarotypus edwardsii; B. Amaroxenus embersoni; C. Amaroxenus marrisi; D. Amarophilus otagoensis; E. Migadopiella convexipennis; F. Migadopiella octoguttata." figureDoi="10.3897/dez.71.134268.figure2" httpUri="https://binary.pensoft.net/fig/1177803">2 B</figureCitation>
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); elytra moderately convex but with disc flat; elytral striae punctate in basal ¾ of length, intervals moderately convex on disc; standardized body length
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<quantity id="EB354262A02FA1254DB212A3BD5043F5" metricMagnitude="-3" metricUnit="m" metricValue="7.949999999999999" metricValueMax="8.1" metricValueMin="7.8" unit="mm" value="7.949999999999999" valueMax="8.1" valueMin="7.8">7.8–8.1 mm</quantity>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="SECID0EBWBG" type="description">
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<paragraph id="D6D5DE41035D2942D1F7D0747B6F344E">
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<heading id="9034EF8733EFA7160B5848D07E33ABD7" reason="title">Description.</heading>
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</paragraph>
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<paragraph id="F62B152DED07B258B0C3E388B81F6480">
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||
<emphasis id="89852FE19B769B1F8CE32AA7E802B2F9" bold="true">
|
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<emphasis id="C5EDE78C08BC746BD865C026F9A5A8DD" bold="true" italics="true">Head</emphasis>
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.
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</emphasis>
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Frons broad, frontal grooves broad and shallow depressions isolated from clypeus, lined with shallow oblique wrinkles on mesal surface; eyes slightly convex, horizontal diameter intersecting ~ 20 ommatidia, ocular ratio = 1.28.
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<emphasis id="CFB2F284A69464644BFCDBA6AFAB9AA1" bold="true" italics="true">Prothorax</emphasis>
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broadly convex dorsally, middle of disc flat, lateral margins distinctly sinuate anteriad slightly acute hind angles,
|
||
<abbrev id="ABBRID0EPWBG" xlink_title="maximal pronotal width">MPW</abbrev>
|
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/
|
||
<abbrev id="ABBRID0ETWBG" xlink_title="basal pronotal width measured between the basal angles">BPW</abbrev>
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||
= 1.05; basal margin trisinuate, medial margin extended slightly posteriad line intersecting hind angles; base indistinctly, narrowly margined, marginal bead narrowest posteriad laterobasal depression; median base smooth, with shallow ovoid depression at midline; laterobasal depressions broad, defined mesally by irregular declivity, surface dimpled laterad declivity and depression curved upward to broadly meet the narrow lateral margin; median impression very fine, intersecting ~ 12 fine, transverse impressions that represent little more than irregularities in microsculpture; lateral marginal bead narrowly upraised, lateral marginal depression very narrow and immediately abutting convex disc; anterior margin with distinct, broad lateral bead laterally, but margin nearly smooth medially, margin traceable only in certain orientations of light source; front angles acute, inner margin adhering to lateral surface of head,
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<abbrev id="ABBRID0EXWBG" xlink_title="apical pronotal width measured at the front angles">APW</abbrev>
|
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/
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<abbrev id="ABBRID0E2WBG" xlink_title="basal pronotal width measured between the basal angles">BPW</abbrev>
|
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= 0.74; prosternum smooth medially, lateral reaches slightly undulated; posterior margin of prosternal process extended as adze-like projection, its ventral surface bearing a deep longitudinal declivity; proepisternum / proepimeron juncture lined with ~ 7 rugose longitudinal depressions.
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<emphasis id="FAB8C843B7A9D302CDBD4BC9010604E8" bold="true" italics="true">Elytra</emphasis>
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narrow basally,
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<abbrev id="ABBRID0ECXBG" xlink_title="width across the humeral angles">HuW</abbrev>
|
||
/
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||
<abbrev id="ABBRID0EGXBG" xlink_title="maximal elytral width">MEW</abbrev>
|
||
= 0.59, humeri evenly expanded posteriad narrowly rounded humeral angle defined by basal and lateral margins; elongate parascutellar stria joined basally to stria 2, the fused trunk curved laterally parallel to sutural stria; parascutellar stria free apically, terminated at 0.70 × elytral length; elytral stria 2 extended nearly to apex, finely punctate even apically, striae 3–6 progressively shorter on apex, stria 7 fused both basally and apically to stria 8, both striae distinctly punctate anteriad posterior juncture, stria 8 irregularly punctate posteriad juncture; stria 9 — i. e. lateral marginal depression — punctate nearly to juncture of striae 7 and 8; elytral interval 3 bearing two very fine setae in obscure depressions set at and posteriad elytral mid-length, the impressions situated just mesad stria and associated with slight deviations in the strial orientation; 11–13 lateral elytral setae situated just laterad stria 8.
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<emphasis id="844B36CD7B87C57C5578A277A9C70BEF" bold="true" italics="true">Pterothorax</emphasis>
|
||
foreshortened, mesepisternum broadly punctate, with ~ 20 punctures in posterior 2 / 3 of length; mesosternum smooth, with fine median crest aligned with adze-like prosternal projection; mesepimeron a narrow longitudinal strap bordering posterior margin of mesepisternum, both sclerites reaching disjunct mesothoracic coxal cavity; metepisternum irregularly quadrate, anterior and posterior margins parallel, medial margin concave along juncture with mesosternum; lateral reaches of metasternum punctate, ~ 8 punctures over surface.
|
||
<emphasis id="98E9EDE66E58416A252EB6DEE734A0D6" bold="true" italics="true">Abdomen</emphasis>
|
||
finely punctate basally on first visible ventrite; lateral reaches of ventrites 2–3 longitudinally wrinkled, ventrites 4–6 very finely punctulate over surface; apical ventrite of
|
||
<specimenCount id="58649DFAB72990C95B3D152CF1112E2C" count="1" type="male">one male</specimenCount>
|
||
and the female specimen with one seta each side of midline, second male with two setae on right, and one on left side of the ventrite.
|
||
<emphasis id="12D672D40D1D624224A1A107DEC0BB25" bold="true" italics="true">Legs</emphasis>
|
||
with expanded tarsomeres on pro- and mesothoracic legs; males with protarsomere 2 w / l = 1.33, mesotarsomere 2 w / l = 1.04, and females with protarsomere 2 w / l = 1.37, mesotarsomere 2 w / l = 0.81; male pro- and mesotarsomeres 1–4 both with ventral surfaces clothed with laterally expanded setae, those on protarsomeres 1–3 broadly expanded laterally, squamose; female protarsomeres 1–4 and mesotarsomeres 2–4 clothed with dense, apical fields of thick, silky, presumably flexible setae.
|
||
</paragraph>
|
||
<paragraph id="C22609752EAACCB3DA231D0BA082AA9C">
|
||
<emphasis id="35C3D3D7E78AD4B05BF9D17C26CEF9E3" bold="true">Male genitalia</emphasis>
|
||
(Fig.
|
||
<figureCitation id="B14ACFC46035CD9690AFDF988D3AF3AC" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Male aedeagal complex. A – C. Amaroxenus embersoni. A. Median lobe and parameres, dextral view; B. Median lobe and parameres, sinistral view; C. Metatergite, ventral view. D – F. Amaroxenus marrisi: D. Median lobe and parameres, dextral view; E. Median lobe and parameres, sinistral view; F. Median lobe and parameres, ventral view." figureDoi="10.3897/dez.71.134268.figure6" httpUri="https://binary.pensoft.net/fig/1177807">6 A – C</figureCitation>
|
||
). Aedeagal median lobe robust, broad dorsoventrally and bilaterally from base to apically rounded apex (Fig.
|
||
<figureCitation id="DC812409E1AAF11B8B748179DAF9FC95" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Male aedeagal complex. A – C. Amaroxenus embersoni. A. Median lobe and parameres, dextral view; B. Median lobe and parameres, sinistral view; C. Metatergite, ventral view. D – F. Amaroxenus marrisi: D. Median lobe and parameres, dextral view; E. Median lobe and parameres, sinistral view; F. Median lobe and parameres, ventral view." figureDoi="10.3897/dez.71.134268.figure6" httpUri="https://binary.pensoft.net/fig/1177807">6 A, B</figureCitation>
|
||
), median lobe basal bulb closed, bearing an apically divergent sagittal crest; median lobe sclerotized basally, ostium opening apically on left side; right paramere broadly conchoid, parallel sided with narrowly rounded ventral apex, ventral margin lined with 3 dense rows of long setae; left paramere conchoid, parallel sided, glabrous, acuminate apicoventrally (Fig.
|
||
<figureCitation id="8B9627C53800DC43770A5C3E28B010E2" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Male aedeagal complex. A – C. Amaroxenus embersoni. A. Median lobe and parameres, dextral view; B. Median lobe and parameres, sinistral view; C. Metatergite, ventral view. D – F. Amaroxenus marrisi: D. Median lobe and parameres, dextral view; E. Median lobe and parameres, sinistral view; F. Median lobe and parameres, ventral view." figureDoi="10.3897/dez.71.134268.figure6" httpUri="https://binary.pensoft.net/fig/1177807">6 B</figureCitation>
|
||
); aedeagal internal sac with heavily sclerotized fields including a dorsal flagellum and a dense ventral spicular field (Fig.
|
||
<figureCitation id="FB5165C87D6EC7202A2BF2F9D6D7D520" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Male aedeagal complex. A – C. Amaroxenus embersoni. A. Median lobe and parameres, dextral view; B. Median lobe and parameres, sinistral view; C. Metatergite, ventral view. D – F. Amaroxenus marrisi: D. Median lobe and parameres, dextral view; E. Median lobe and parameres, sinistral view; F. Median lobe and parameres, ventral view." figureDoi="10.3897/dez.71.134268.figure6" httpUri="https://binary.pensoft.net/fig/1177807">6 A</figureCitation>
|
||
); mediotergite VIII broadly rounded distally, sclerotized apex broadened relative to tubular lateral margin (Fig.
|
||
<figureCitation id="E83FDCAA6C5B91C7819989F557D8E47A" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Male aedeagal complex. A – C. Amaroxenus embersoni. A. Median lobe and parameres, dextral view; B. Median lobe and parameres, sinistral view; C. Metatergite, ventral view. D – F. Amaroxenus marrisi: D. Median lobe and parameres, dextral view; E. Median lobe and parameres, sinistral view; F. Median lobe and parameres, ventral view." figureDoi="10.3897/dez.71.134268.figure6" httpUri="https://binary.pensoft.net/fig/1177807">6 C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0ELYBG" type="Female reproductive tract">
|
||
<paragraph id="A85FF20314CC2D2F664DABC62BE45455">
|
||
<heading id="1413AD6A9B07DF1B0788D917CA062B09" reason="title">Female reproductive tract.</heading>
|
||
</paragraph>
|
||
<paragraph id="3E6EDFAB25BD24A634F657B9B8F5A090">
|
||
Gonocoxite narrow, elongate, unipartite, articulated basally with heavily sclerotized median boss along anterior margin of laterotergite IX (Fig.
|
||
<figureCitation id="9105BBF87429B8728770B8F3389FFDB0" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Female left gonocoxite, ventral view. A. Amarotypus edwardsii; B. Amaroxenus embersoni; C. Amaroxenus marrisi; D. Migadopiella octoguttata; E. Amarophilus otagoensis." figureDoi="10.3897/dez.71.134268.figure3" httpUri="https://binary.pensoft.net/fig/1177804">3 B</figureCitation>
|
||
); two nematiform setae in apical sensorial pit; ventroapical surface of gonocoxite lined with campaniform sensoria, with several small trichoid sensilla in median half of coxite; bursa copulatrix circular (when compressed on microslide), with common oviduct-bursal juncture on ventral surface, and cristate “ helminthoid sclerite ” near juncture (Fig.
|
||
<figureCitation id="56FB3E16B19BF80504471A483223371B" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Female reproductive tract, ventral view, scale bar = 0.5 mm to right of each figure. A. Stichonotus piceus; B. Amarotypus edwardsii; C. Migadopiella octoguttata; D. Amaroxenus embersoni; E. Amaroxenus marrisi; F. Amarophilus otagoensis. Abbreviations for anatomical structures include: bc, bursa copulatrix; bd, bursal diverticulum; co, common oviduct; dbs, distal bursal sclerite; gc, gonocoxa; ls, ligular sclerite; lt, laterotergite." figureDoi="10.3897/dez.71.134268.figure4" httpUri="https://binary.pensoft.net/fig/1177805">4 B</figureCitation>
|
||
); ventroapical surface of bursa broadly, moderately sclerotized, resulting in a discrete plate just distad the bursal-oviduct juncture.
|
||
</paragraph>
|
||
<caption id="EC24B9D01E92D6FF3C801C808EE3CF03" ID-DOI="10.3897/dez.71.134268.figure4" ID-arpha="B75D0D10-EC15-5AB6-96B2-099962665324" httpUri="https://binary.pensoft.net/fig/1177805" startId="F4">
|
||
<paragraph id="FBF2FBA0119BE53DC6E7360600B94E75">
|
||
<label id="12E235460CAF5E9061F2411725B59066">Figure 4.</label>
|
||
</paragraph>
|
||
<paragraph id="4842BD323B20EBE5064532DFFEC15517">
|
||
Female reproductive tract, ventral view, scale bar = 0.5 mm to right of each figure.
|
||
<emphasis id="0CFEBDF6E899D49CECAC3A0BC91EAADE" bold="true">A.</emphasis>
|
||
<taxonomicName id="1F485D61C52B1C4C88B3DB7775CB4E4C" authorityName="Sloane" authorityYear="1915" class="Insecta" family="Carabidae" genus="Stichonotus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="piceus">
|
||
<emphasis id="29AF2B8A760B1C74175DEE4E1098B530" italics="true">Stichonotus piceus</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<emphasis id="734D3A02F7A3EC5814D0087A82FBE3DE" bold="true">B.</emphasis>
|
||
<taxonomicName id="29CB112349EA3263B2DBB5548E00FC94" authorityName="Bates" authorityYear="1872" class="Insecta" family="Carabidae" genus="Amarotypus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="edwardsii">
|
||
<emphasis id="70DCEE1355A5128B81DBF1CE9F850CBD" italics="true">Amarotypus edwardsii</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<emphasis id="7462B46302AF8C7A6ED8843572D99D54" bold="true">C.</emphasis>
|
||
<taxonomicName id="6440E25F63F9EED34018A9EC0EDB8E7F" authorityName="Baehr" authorityYear="2009" class="Insecta" family="Carabidae" genus="Migadopiella" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="octoguttata">
|
||
<emphasis id="AF0912B4D5EC23F5692F0B7F9DBBE545" italics="true">Migadopiella octoguttata</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<emphasis id="F192E2661382CDB0CF102E7F4F840B2B" bold="true">D.</emphasis>
|
||
<taxonomicName id="98D234E5D1BF44E3581BA6D00E7CCE18" authorityName="Liebherr & Roig-Juñent & Will" authorityYear="2024" class="Insecta" family="Carabidae" genus="Amaroxenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="embersoni">
|
||
<emphasis id="0E52271BC69EB1AEEF6EAAA9F0993587" italics="true">Amaroxenus embersoni</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<emphasis id="6664D70AFF92B35A4F9ABDBAE68F425E" bold="true">E.</emphasis>
|
||
<taxonomicName id="54E6174BAB8120AD8BBDDEE9FFE3EC37" authorityName="Liebherr & Will" authorityYear="2024" class="Insecta" family="Carabidae" genus="Amaroxenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="marrisi">
|
||
<emphasis id="7C52C75BFBAA3347BDEF2737AE20EB19" italics="true">Amaroxenus marrisi</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<emphasis id="C60608A0C54C6B8929D17F57479FFB80" bold="true">F.</emphasis>
|
||
<taxonomicName id="8A8E94B5A1515A0ECB2787B0682E8B34" class="Insecta" family="Carabidae" genus="Amarophilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="otagoensis">
|
||
<emphasis id="E3245B68EF120D9E36550FEEF8309873" italics="true">Amarophilus otagoensis</emphasis>
|
||
</taxonomicName>
|
||
. Abbreviations for anatomical structures include: bc, bursa copulatrix; bd, bursal diverticulum; co, common oviduct; dbs, distal bursal sclerite; gc, gonocoxa; ls, ligular sclerite; lt, laterotergite.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0EU2BG" type="etymology">
|
||
<paragraph id="5666A72EA7D48EF41A742C14E62088B3">
|
||
<heading id="63091820E7364D980A24DD05028A150F" reason="title">Etymology.</heading>
|
||
</paragraph>
|
||
<paragraph id="1DD6C80B18F92FCA4207BF28DD0F147B">
|
||
We take great pleasure in naming this species for Professor Rowan M. Emberson (Fig.
|
||
<figureCitation id="C95D782E9EC5A94BB8008EE77841616E" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Type locality of Amaroxenus embersoni showing Professor Rowan Emberson collecting the type series (photo courtesy Pol Syrett)." figureDoi="10.3897/dez.71.134268.figure7" httpUri="https://binary.pensoft.net/fig/1177808">7</figureCitation>
|
||
), late of Lincoln University. He collected the
|
||
<specimenCount id="F47F0CA08E6C4E2F9ABF13F35AA71938" count="3" type="generic" typeStatus="type">three type</specimenCount>
|
||
specimens during a hike, accompanied by Pauline Syrett, to the summit of Mt. Aicken in the Southern Alps of
|
||
<collectingCountry id="BED403A0B60775CE4439983F77A42972" name="New Zealand">New Zealand</collectingCountry>
|
||
, December, 1987. The species epithet recognizes Professor Emberson’s dedication to the study of New Zealand’s insects, including discovery of this high montane
|
||
<collectingCountry id="0337C2D6F8B87D3E9CB43F67F47CB36C" name="New Zealand">New Zealand</collectingCountry>
|
||
insect species comprising animals that live in extreme situations far beyond the limits of most entomological exploration.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0E52BG" type="distribution">
|
||
<paragraph id="3DDF59B533701A40D11C5E37A7CD1B23">
|
||
<heading id="E5A1F5012E1A8F19D13D2D4F849427E3" reason="title">Distribution.</heading>
|
||
</paragraph>
|
||
<paragraph id="7EE0A70679D80724996B7FE6BC7C311D">
|
||
Mt. Aicken is located at
|
||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EH3BG" specific-use="{"type":"Point","coordinates":[171.595000,-42.930833]}">
|
||
<geoCoordinate id="549D6474A2F8FD413D36710287164B66" degrees="42" direction="south" minutes="55.85" orientation="latitude" precision="9" value="-42.930832">42 ° 55.85 ' S</geoCoordinate>
|
||
, 171 ° 35.70 ′ Е
|
||
</named-content>
|
||
, and has a summit elevation of
|
||
<quantity id="E4C0EBE31FA022257F110418E2724377" metricMagnitude="3" metricUnit="m" metricValue="1.859" unit="m" value="1859.0">1859 m</quantity>
|
||
(
|
||
<ext-link id="B6C5E9858758D2AA2A2BB1CF09E7B411" ext-link-type="uri" xlink_href="https://climbnz.org.nz/nz/si/arthurs-pass/mt-aicken" xlink_type="simple">https://climbnz.org.nz/nz/si/arthurs-pass/mt-aicken</ext-link>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0ER3BG" type="habitat">
|
||
<paragraph id="4F8CB095FBEA441DCD0EE69EE3495F6E">
|
||
<heading id="C76AB136A36333A7131B811AF543A939" reason="title">Habitat.</heading>
|
||
</paragraph>
|
||
<paragraph id="9DD0DA143CCAA43199CEED4B0DB2D3A0">
|
||
This is a species of high montane grassland and exposed glacial till. At elevations of
|
||
<quantity id="E6058152D861FF50AB632E8DE1501FE7" metricMagnitude="3" metricUnit="m" metricValue="1.7" metricValueMax="1.75" metricValueMin="1.65" unit="m" value="1700.0" valueMax="1750.0" valueMin="1650.0">1650–1750 m</quantity>
|
||
in the Southern Alps, these habitats are strictly alpine in character. The
|
||
<specimenCount id="4A3E1CB7FFC3625A9857EF28462C04A6" count="3" type="generic">three specimens</specimenCount>
|
||
were collected under rocks at the edge of receding snowfields (Fig.
|
||
<figureCitation id="E0F45AFCD1B9028A04CE12520E70F0AA" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Type locality of Amaroxenus embersoni showing Professor Rowan Emberson collecting the type series (photo courtesy Pol Syrett)." figureDoi="10.3897/dez.71.134268.figure7" httpUri="https://binary.pensoft.net/fig/1177808">7</figureCitation>
|
||
)
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |