treatments-xml/data/12/38/87/123887EBFFE4FF83FF436746FD7FC98E.xml

<document id="D5BDBD55FF92F25A1AD1CF8476FC1C9C" ID-DOI="10.5281/zenodo.5434686" ID-ISSN="0253-116X" ID-Zenodo-Dep="5434686" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630433216301" checkinUser="valdenar" docAuthor="A. S, D. W, A, W &amp; H, E" docDate="2006" docId="123887EBFFE4FF83FF436746FD7FC98E" docLanguage="en" docName="LinzerbiolBeitr.2006.38.1.991-1008.pdf" docOrigin="Linzer biologische Beiträge 38 (1)" docStyle="DocumentStyle:24E5EB772D355DE2EE23B1E8D07853E0.9:LinzerbiolBeitr.2006-.journal_article" docStyleId="24E5EB772D355DE2EE23B1E8D07853E0" docStyleName="LinzerbiolBeitr.2006-.journal_article" docStyleVersion="9" docTitle="Parapenetretus (Propenetretus) A.S &amp; D.W &amp; A &amp; H 2006, subgen. nov." docType="treatment" docVersion="4" lastPageNumber="998" masterDocId="EE01FF93FFE1FF84FFD46455FFC4CB15" masterDocTitle="New taxa of the tribe Patrobini (Coleoptera, Carabidae) from the East Himalayas" masterLastPageNumber="1008" masterPageNumber="991" pageNumber="996" updateTime="1699269285600" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="E5E44CDC473E13477BCB2597817DB36C">New taxa of the tribe Patrobini (Coleoptera, Carabidae) from the East Himalayas</mods:title>
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<mods:namePart id="E687FEBC41AF16DF2467F28F931206F6">H, E</mods:namePart>
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<mods:title id="25619CFF95CC458CF6E245FB7CF1F6A6">Linzer biologische Beiträge</mods:title>
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<mods:date id="9DCC8EBA82B370955629A3D8A4706690">2006</mods:date>
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<mods:number id="E49CB023007633D670E4CA8E6CE840DA">2006-07-21</mods:number>
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<mods:number id="00F6F7167D4F940CD1177247197755CC">38</mods:number>
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<subSubSection id="D28B6576FFE4FF81FF436746FE72C83D" box="[151,438,787,808]" pageId="5" pageNumber="996" type="nomenclature">
<paragraph id="9A2E36FDFFE4FF81FF436746FE72C83D" blockId="5.[151,438,787,808]" box="[151,438,787,808]" pageId="5" pageNumber="996">
<heading id="C1668191FFE4FF81FF436746FE72C83D" box="[151,438,787,808]" fontSize="8" level="2" pageId="5" pageNumber="996" reason="2">
<emphasis id="A8E5EAEFFFE4FF81FF436746FE72C83D" bold="true" box="[151,438,787,808]" pageId="5" pageNumber="996">
<taxonomicName id="5D914D7EFFE4FF81FF436746FEEDC83D" ID-CoL="8MKLR" authority="A.S &amp; D.W &amp; A &amp; H, 2006" authorityName="A.S &amp; D.W &amp; A &amp; H" authorityYear="2006" box="[151,297,787,808]" class="Insecta" family="Carabidae" genus="Parapenetretus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="996" phylum="Arthropoda" rank="subGenus" status="subgen. nov." subGenus="Propenetretus">
<emphasis id="A8E5EAEFFFE4FF81FF436746FEEDC83D" bold="true" box="[151,297,787,808]" italics="true" pageId="5" pageNumber="996">Propenetretus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B3D65794FFE4FF81FEE46746FE72C83D" box="[304,438,787,808]" pageId="5" pageNumber="996" rank="subGenus">subgen. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="D28B6576FFE4FF83FF43676CFD7FC98E" lastPageId="7" lastPageNumber="998" pageId="5" pageNumber="996" type="materials_examined">
<paragraph id="9A2E36FDFFE4FF81FF43676CFCE3C858" blockId="5.[151,807,824,845]" box="[151,807,824,845]" pageId="5" pageNumber="996">
<typeStatus id="452A885FFFE4FF81FF43676CFF02C858" box="[151,198,825,845]" pageId="5" pageNumber="996">Type</typeStatus>
species: 
<taxonomicName id="5D914D7EFFE4FF81FECD676DFE39C858" authority="A.S &amp; D.W &amp; A &amp; H, 2006" authorityName="A.S &amp; D.W &amp; A &amp; H" authorityYear="2006" box="[281,509,824,845]" class="Insecta" family="Carabidae" genus="Parapenetretus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="996" phylum="Arthropoda" rank="species" species="selaensis" status="sp. nov.">
<emphasis id="A8E5EAEFFFE4FF81FECD676DFE39C858" box="[281,509,824,845]" italics="true" pageId="5" pageNumber="996">Parapenetretus selaensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B3D65794FFE4FF81FDD6676CFDA5C858" box="[514,609,825,845]" pageId="5" pageNumber="996" rank="species">spec. nov.</taxonomicNameLabel>
, herewith designated.
</paragraph>
<paragraph id="9A2E36FDFFE4FF82FF436709FD41C93E" blockId="5.[151,1059,860,1584]" lastBlockId="6.[151,1059,211,555]" lastPageId="6" lastPageNumber="997" pageId="5" pageNumber="996">
D i a g n o s t i c f e a t u r e s:Initsmainfeaturesagreeswithspeciesofthegenus 
<taxonomicName id="5D914D7EFFE4FF81FF43672FFEF7C89A" box="[151,307,890,911]" class="Insecta" family="Carabidae" genus="Parapenetretus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="996" phylum="Arthropoda" rank="genus">
<emphasis id="A8E5EAEFFFE4FF81FF43672FFEF7C89A" box="[151,307,890,911]" italics="true" pageId="5" pageNumber="996">Parapenetretus</emphasis>
</taxonomicName>
, being the most similar habitually to 
<taxonomicName id="5D914D7EFFE4FF81FD67672FFEE5C8BE" authority="ZAMOTAJLOV" authorityName="ZAMOTAJLOV" class="Insecta" family="Carabidae" genus="Parapenetretus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="996" phylum="Arthropoda" rank="species" species="shimianensis" subGenus="Ambigopenetretus">
<emphasis id="A8E5EAEFFFE4FF81FD67672FFD0DC89A" box="[691,713,890,911]" italics="true" pageId="5" pageNumber="996">P.</emphasis>
(
<emphasis id="A8E5EAEFFFE4FF81FD0C672CFC54C89B" box="[728,912,889,910]" italics="true" pageId="5" pageNumber="996">Ambigopenetretus</emphasis>
) 
<emphasis id="A8E5EAEFFFE4FF81FC74672CFBE7C89B" box="[928,1059,889,910]" italics="true" pageId="5" pageNumber="996">shimianensis</emphasis>
ZAMOTAJLOV
</taxonomicName>
from Sichuan. However, 
<taxonomicName id="5D914D7EFFE4FF81FDF967C2FD78C8B9" authority="A.S &amp; D.W &amp; A &amp; H, 2006" authorityName="A.S &amp; D.W &amp; A &amp; H" authorityYear="2006" box="[557,700,919,940]" class="Insecta" family="Carabidae" genus="Parapenetretus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="996" phylum="Arthropoda" rank="subGenus" status="subgen. nov." subGenus="Propenetretus">
<emphasis id="A8E5EAEFFFE4FF81FDF967C2FD78C8B9" box="[557,700,919,940]" italics="true" pageId="5" pageNumber="996">Propenetretus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B3D65794FFE4FF81FD1067C3FC83C8B9" box="[708,839,918,940]" pageId="5" pageNumber="996" rank="subGenus">subgen. nov.</taxonomicNameLabel>
is easily distinguishable from 
<taxonomicName id="5D914D7EFFE4FF81FEDF67E1FE63C8DC" box="[267,423,948,969]" class="Insecta" family="Carabidae" genus="Parapenetretus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="996" phylum="Arthropoda" rank="genus">
<emphasis id="A8E5EAEFFFE4FF81FEDF67E1FE63C8DC" box="[267,423,948,969]" italics="true" pageId="5" pageNumber="996">Parapenetretus</emphasis>
</taxonomicName>
(s. str.), 
<emphasis id="A8E5EAEFFFE4FF81FDC767E1FD01C8DC" box="[531,709,948,969]" italics="true" pageId="5" pageNumber="996">Butanopenetretus</emphasis>
, the bulk of 
<emphasis id="A8E5EAEFFFE4FF81FC8A67E1FBDEC8DC" box="[862,1050,948,969]" italics="true" pageId="5" pageNumber="996">Robustopenetretus</emphasis>
- species, and 
<emphasis id="A8E5EAEFFFE4FF81FECE6787FE60C8F3" box="[282,420,977,999]" italics="true" pageId="5" pageNumber="996">P. reticulatus</emphasis>
ZAMOTAJLOV by presence of only 2 (very seldom 3) supraorbital setae on each side of head. In this respect it resembles 
<emphasis id="A8E5EAEFFFE4FF81FCCD67BBFC15CF16" box="[793,977,1006,1027]" italics="true" pageId="5" pageNumber="996">Ambigopenetretus</emphasis>
-species and some species of 
<emphasis id="A8E5EAEFFFE4FF81FEA4605EFDE8CF35" box="[368,556,1035,1056]" italics="true" pageId="5" pageNumber="996">Robustopenetretus</emphasis>
. From the first subgenus it differs in polysetose lateral margin of pronotum, longer mesepisternum (
<figureCitation id="02AA2A78FFE4FF81FD62607CFD35CF2A" box="[694,753,1065,1087]" captionStart="Figs 3-8" captionStartId="12.[151,191,1258,1280]" captionTargetBox="[175,1039,242,1234]" captionTargetId="figure-83@12.[151,1057,235,1248]" captionTargetPageId="12" captionText="Figs 3-8: Patrobini, details of external morphology (3-4 - mentum, dorsal view, 5-6 - lateral part of thorax, lateroventral view, 7-8 - female tergite 8, dorsal view, ac2 - middle coxal cavity, epm2 - mesepimeron, epm3 - metepimeron, eps2 - mesepisternum, eps3 - metepisternum, st2 - mesosternum, st3 - metasternum). Indopatrobus bashtai spec. nov., paratype, (3, 5, 7). Parapenetretus selaensis spec. nov., paratype, (4, 6, 8). Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5434690" httpUri="https://zenodo.org/record/5434690/files/figure.png" pageId="5" pageNumber="996">fig. 6</figureCitation>
), absence of ventral setae of tarsomere 5, different shape of female tergite 8 (
<figureCitation id="02AA2A78FFE4FF81FD5C6013FD05CF49" box="[648,705,1094,1116]" captionStart="Figs 3-8" captionStartId="12.[151,191,1258,1280]" captionTargetBox="[175,1039,242,1234]" captionTargetId="figure-83@12.[151,1057,235,1248]" captionTargetPageId="12" captionText="Figs 3-8: Patrobini, details of external morphology (3-4 - mentum, dorsal view, 5-6 - lateral part of thorax, lateroventral view, 7-8 - female tergite 8, dorsal view, ac2 - middle coxal cavity, epm2 - mesepimeron, epm3 - metepimeron, eps2 - mesepisternum, eps3 - metepisternum, st2 - mesosternum, st3 - metasternum). Indopatrobus bashtai spec. nov., paratype, (3, 5, 7). Parapenetretus selaensis spec. nov., paratype, (4, 6, 8). Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5434690" httpUri="https://zenodo.org/record/5434690/files/figure.png" pageId="5" pageNumber="996">fig. 8</figureCitation>
), male and female genitalia. From the latter it differs, first of all, in its more slender body, head with much less developed puncturation and rugosity, and longer mesepisternum. From all hitherto described subgenera and groups of 
<taxonomicName id="5D914D7EFFE4FF81FEBD60CAFDC1CFA1" box="[361,517,1183,1204]" class="Insecta" family="Carabidae" genus="Parapenetretus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="996" phylum="Arthropoda" rank="genus">
<emphasis id="A8E5EAEFFFE4FF81FEBD60CAFDC1CFA1" box="[361,517,1183,1204]" italics="true" pageId="5" pageNumber="996">Parapenetretus</emphasis>
</taxonomicName>
, 
<taxonomicName id="5D914D7EFFE4FF81FDC260CAFD61CFA1" authority="A.S &amp; D.W &amp; A &amp; H, 2006" authorityName="A.S &amp; D.W &amp; A &amp; H" authorityYear="2006" box="[534,677,1183,1204]" class="Insecta" family="Carabidae" genus="Parapenetretus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="996" phylum="Arthropoda" rank="subGenus" status="subgen. nov." subGenus="Propenetretus">
<emphasis id="A8E5EAEFFFE4FF81FDC260CAFD61CFA1" box="[534,677,1183,1204]" italics="true" pageId="5" pageNumber="996">Propenetretus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B3D65794FFE4FF81FD7B60CBFCF0CFA6" box="[687,820,1182,1204]" pageId="5" pageNumber="996" rank="subGenus">subgen. nov.</taxonomicNameLabel>
differs in less cordate pronotum and more elongate, possessing lateral keels, apical lamella of aedeagus (only 
<emphasis id="A8E5EAEFFFE4FF81FF43608CFEE7CFF8" box="[151,291,1240,1262]" italics="true" pageId="5" pageNumber="996">P. reticulatus</emphasis>
ZAMOTAJLOV known upon female being unstudied in the latter respect) (
<figureCitation id="02AA2A78FFE4FF81FF4A60A0FEDDCE1E" box="[158,281,1269,1291]" captionStart="Figs 9-18" captionStartId="13.[151,191,1493,1515]" captionTargetBox="[203,992,200,1479]" captionTargetId="figure-76@13.[192,1008,192,1483]" captionTargetPageId="13" captionText="Figs 9-18: Patrobini, male genitalia (9, 14 - aedeagus, left lateral view, 10, 15 - aedeagus, dorsal view, 11, 16 - left paramere, left lateral view, 12, 17 - right paramere, right lateral view, 13, 18 - ventrite 9, dorsal view). Indopatrobus bashtai spec. nov., paratype, (9-13). Parapenetretus selaensis spec. nov., paratype, (14-18). Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5434692" httpUri="https://zenodo.org/record/5434692/files/figure.png" pageId="5" pageNumber="996">figs 14, 15</figureCitation>
). In habitus strongly resembles also &quot; 
<taxonomicName id="5D914D7EFFE4FF81FD6260A3FCDBCE1E" authorityName="HABU &amp; BABA" authorityYear="1960" box="[694,799,1270,1291]" class="Insecta" family="Carabidae" genus="Apatrobus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="996" phylum="Arthropoda" rank="genus">
<emphasis id="A8E5EAEFFFE4FF81FD6260A3FCDBCE1E" box="[694,799,1270,1291]" italics="true" pageId="5" pageNumber="996">Apatrobus</emphasis>
</taxonomicName>
&quot; 
<emphasis id="A8E5EAEFFFE4FF81FCE160A3FC6CCE1E" box="[821,936,1270,1291]" italics="true" pageId="5" pageNumber="996">sikkimensis</emphasis>
(
<collectionCode id="FC80AE38FFE4FF81FC6860A0FBC0CE1F" box="[956,1028,1269,1291]" pageId="5" pageNumber="996">DEUVE</collectionCode>
&amp; 
<collectionCode id="FC80AE38FFE4FF81FF436146FF29CE32" box="[151,237,1299,1320]" pageId="5" pageNumber="996">LEDOUX</collectionCode>
), however, larger in average, body more slender, hind supraorbital seta located closer to neck-constriction, pore puncture larger, lateral margin of pronotum polysetose (in &quot; 
<taxonomicName id="5D914D7EFFE4FF81FF1D611BFEF6CE76" authorityName="HABU &amp; BABA" authorityYear="1960" box="[201,306,1358,1379]" class="Insecta" family="Carabidae" genus="Apatrobus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="996" phylum="Arthropoda" rank="genus">
<emphasis id="A8E5EAEFFFE4FF81FF1D611BFEF6CE76" box="[201,306,1358,1379]" italics="true" pageId="5" pageNumber="996">Apatrobus</emphasis>
</taxonomicName>
&quot; 
<emphasis id="A8E5EAEFFFE4FF81FE9D611BFE78CE76" box="[329,444,1358,1379]" italics="true" pageId="5" pageNumber="996">sikkimensis</emphasis>
bears single seta), marginal series of elytra not forming distinct groups (in &quot; 
<taxonomicName id="5D914D7EFFE4FF81FEB2613EFE0BCE95" authorityName="HABU &amp; BABA" authorityYear="1960" box="[358,463,1387,1408]" class="Insecta" family="Carabidae" genus="Apatrobus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="996" phylum="Arthropoda" rank="genus">
<emphasis id="A8E5EAEFFFE4FF81FEB2613EFE0BCE95" box="[358,463,1387,1408]" italics="true" pageId="5" pageNumber="996">Apatrobus</emphasis>
</taxonomicName>
&quot; 
<emphasis id="A8E5EAEFFFE4FF81FE36613EFD91CE95" box="[482,597,1387,1408]" italics="true" pageId="5" pageNumber="996">sikkimensis</emphasis>
composed of 3 more or less distinctly separated groups, formula of series being 2 + 1 + 6), shape of apical lamella of aedeagus (see 
<figureCitation id="02AA2A78FFE4FF81FF4361F0FEC9CEAE" box="[151,269,1445,1467]" captionStart="Figs 9-18" captionStartId="13.[151,191,1493,1515]" captionTargetBox="[203,992,200,1479]" captionTargetId="figure-76@13.[192,1008,192,1483]" captionTargetPageId="13" captionText="Figs 9-18: Patrobini, male genitalia (9, 14 - aedeagus, left lateral view, 10, 15 - aedeagus, dorsal view, 11, 16 - left paramere, left lateral view, 12, 17 - right paramere, right lateral view, 13, 18 - ventrite 9, dorsal view). Indopatrobus bashtai spec. nov., paratype, (9-13). Parapenetretus selaensis spec. nov., paratype, (14-18). Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5434692" httpUri="https://zenodo.org/record/5434692/files/figure.png" pageId="5" pageNumber="996">figs 14, 15</figureCitation>
) different, elongate, with lateral keels (in &quot; 
<taxonomicName id="5D914D7EFFE4FF81FD3661F0FC8FCEAF" authorityName="HABU &amp; BABA" authorityYear="1960" box="[738,843,1445,1466]" class="Insecta" family="Carabidae" genus="Apatrobus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="996" phylum="Arthropoda" rank="genus">
<emphasis id="A8E5EAEFFFE4FF81FD3661F0FC8FCEAF" box="[738,843,1445,1466]" italics="true" pageId="5" pageNumber="996">Apatrobus</emphasis>
</taxonomicName>
&quot; 
<emphasis id="A8E5EAEFFFE4FF81FCB561F0FC10CEAF" box="[865,980,1445,1466]" italics="true" pageId="5" pageNumber="996">sikkimensis</emphasis>
guttershaped and open dorsally). Aedeagus resembles the most closely that of 
<taxonomicName id="5D914D7EFFE4FF81FC426196FE6ECEE0" authority="(ZAMOTAJLOV)" baseAuthorityName="ZAMOTAJLOV" class="Insecta" family="Carabidae" genus="Parapatrobus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="996" phylum="Arthropoda" rank="species" species="brancuccii">
<emphasis id="A8E5EAEFFFE4FF81FC426196FEC7CEE0" italics="true" pageId="5" pageNumber="996">Parapatrobus brancuccii</emphasis>
(ZAMOTAJLOV)
</taxonomicName>
, however, smaller, apical protuberance much longer, apical lamella broader (viewed laterally), not twisted leftward (viewed dorsally), proximal copulatory piece smaller, of different structure, apical projections of parameres shorter in average. Being manifestly different in general appearance, it is distinguishable also in numerous important details of external morphology: lateral margin of pronotum polysetose, suture separating mesepisternum and mesosternum (
<figureCitation id="02AA2A78FFE7FF82FD3A655BFCECCA31" box="[750,808,270,292]" captionStart="Figs 3-8" captionStartId="12.[151,191,1258,1280]" captionTargetBox="[175,1039,242,1234]" captionTargetId="figure-83@12.[151,1057,235,1248]" captionTargetPageId="12" captionText="Figs 3-8: Patrobini, details of external morphology (3-4 - mentum, dorsal view, 5-6 - lateral part of thorax, lateroventral view, 7-8 - female tergite 8, dorsal view, ac2 - middle coxal cavity, epm2 - mesepimeron, epm3 - metepimeron, eps2 - mesepisternum, eps3 - metepisternum, st2 - mesosternum, st3 - metasternum). Indopatrobus bashtai spec. nov., paratype, (3, 5, 7). Parapenetretus selaensis spec. nov., paratype, (4, 6, 8). Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5434690" httpUri="https://zenodo.org/record/5434690/files/figure.png" pageId="6" pageNumber="997">fig. 6</figureCitation>
) joins lateral margin of metathorax, mesepimeron of different shape, only slightly broadened inward, mesepisternum longer, front margin of metathorax near middle coxal cavities valliculiform, male protarsomere 2 smaller than protarsomere 3 than in 
<taxonomicName id="5D914D7EFFE7FF82FD356533FC25CA6E" box="[737,993,358,379]" class="Insecta" family="Carabidae" genus="Parapatrobus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="997" phylum="Arthropoda" rank="species" species="brancuccii">
<emphasis id="A8E5EAEFFFE7FF82FD356533FC25CA6E" box="[737,993,358,379]" italics="true" pageId="6" pageNumber="997">Parapatrobus brancuccii</emphasis>
</taxonomicName>
, latter nearly longitudinal, anterolateral apophyse of female tergite 8 indistinct (
<figureCitation id="02AA2A78FFE7FF82FC5E65D6FC07CA8C" box="[906,963,387,409]" captionStart="Figs 3-8" captionStartId="12.[151,191,1258,1280]" captionTargetBox="[175,1039,242,1234]" captionTargetId="figure-83@12.[151,1057,235,1248]" captionTargetPageId="12" captionText="Figs 3-8: Patrobini, details of external morphology (3-4 - mentum, dorsal view, 5-6 - lateral part of thorax, lateroventral view, 7-8 - female tergite 8, dorsal view, ac2 - middle coxal cavity, epm2 - mesepimeron, epm3 - metepimeron, eps2 - mesepisternum, eps3 - metepisternum, st2 - mesosternum, st3 - metasternum). Indopatrobus bashtai spec. nov., paratype, (3, 5, 7). Parapenetretus selaensis spec. nov., paratype, (4, 6, 8). Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5434690" httpUri="https://zenodo.org/record/5434690/files/figure.png" pageId="6" pageNumber="997">fig. 8</figureCitation>
), base of tergite moderately exceeding epitergite, etc. Elongate, complicated bursal sclerite and presence of distal sclerite of female reproductive tract outwardly resemble the 
<emphasis id="A8E5EAEFFFE7FF82FBD865EBFF32CAE5" italics="true" pageId="6" pageNumber="997">P. hikisanus</emphasis>
-group and the 
<taxonomicName id="5D914D7EFFE7FF82FE76658EFD90CAE5" box="[418,596,475,496]" class="Insecta" family="Carabidae" genus="Parapatrobus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="997" phylum="Arthropoda" rank="species" species="hayachinensis">
<emphasis id="A8E5EAEFFFE7FF82FE76658EFD90CAE5" box="[418,596,475,496]" italics="true" pageId="6" pageNumber="997">P. hayachinensis</emphasis>
</taxonomicName>
-group of 
<taxonomicName id="5D914D7EFFE7FF82FD11658EFCEACAE5" authorityName="HABU &amp; BABA" authorityYear="1960" box="[709,814,475,496]" class="Insecta" family="Carabidae" genus="Apatrobus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="1008" phylum="Arthropoda" rank="genus">
<emphasis id="A8E5EAEFFFE7FF82FD11658EFCEACAE5" box="[709,814,475,496]" italics="true" pageId="6" pageNumber="997">Apatrobus</emphasis>
</taxonomicName>
from 
<collectingCountry id="E286766DFFE7FF82FCAD658EFC71CAE5" box="[889,949,475,496]" name="Japan" pageId="6" pageNumber="997">Japan</collectingCountry>
, however, 
<taxonomicName id="5D914D7EFFE7FF82FF4365ACFEE2C91B" authority="A.S &amp; D.W &amp; A &amp; H, 2006" authorityName="A.S &amp; D.W &amp; A &amp; H" authorityYear="2006" box="[151,294,505,526]" class="Insecta" family="Carabidae" genus="Parapenetretus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="997" phylum="Arthropoda" rank="subGenus" status="subgen. nov." subGenus="Propenetretus">
<emphasis id="A8E5EAEFFFE7FF82FF4365ACFEE2C91B" box="[151,294,505,526]" italics="true" pageId="6" pageNumber="997">Propenetretus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B3D65794FFE7FF82FEFA65ADFE74C91B" box="[302,432,504,526]" pageId="6" pageNumber="997" rank="subGenus">subgen. nov.</taxonomicNameLabel>
differs from them in the same features as other subgenera of 
<taxonomicName id="5D914D7EFFE7FF82FF436643FEF7C93E" box="[151,307,534,555]" class="Insecta" family="Carabidae" genus="Parapenetretus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="997" phylum="Arthropoda" rank="genus">
<emphasis id="A8E5EAEFFFE7FF82FF436643FEF7C93E" box="[151,307,534,555]" italics="true" pageId="6" pageNumber="997">Parapenetretus</emphasis>
</taxonomicName>
(see key by 
<bibRefCitation id="FE004B0CFFE7FF82FE676643FDBEC93E" author="ZAMOTAJLOV A" box="[435,634,533,555]" pageId="6" pageNumber="997" pagination="1 - 145" refId="ref6582" refString="ZAMOTAJLOV A. S. (2002): Inferring phylogenetic system of the carabid subfamily Patrobinae (Coleoptera, Carabidae). - St-Petersburg: Zoological Institute of Russian Academy of Sciences (Meetings in memory N. A. Cholodkovsky; Iss. 55): 1 - 145. [In Russian]." type="journal article" year="2002">ZAMOTAJLOV 2002</bibRefCitation>
).
</paragraph>
<paragraph id="9A2E36FDFFE7FF82FF43666EFC1BC97B" blockId="6.[151,1058,570,622]" pageId="6" pageNumber="997">D e s c r i p t i o n: Body of medium size. Integument fully pigmented, dark, nearly monochromatic, without metallic lustre. Rather slender, with elongate appendages.</paragraph>
<paragraph id="9A2E36FDFFE7FF82FF436628FC19C830" blockId="6.[151,1059,637,805]" pageId="6" pageNumber="997">
Head normal; tempora well-developed, long; mandibles normal; antennomere 1 with single anterodorsal seta (exceptionally with a few minute additional setae), antennomere 2 with a corona of setae apically; apical palpomere of maxillar palpi broadest in the middle; mentum (
<figureCitation id="02AA2A78FFE7FF82FE9A6680FE4CC9FE" box="[334,392,725,747]" captionStart="Figs 3-8" captionStartId="12.[151,191,1258,1280]" captionTargetBox="[175,1039,242,1234]" captionTargetId="figure-83@12.[151,1057,235,1248]" captionTargetPageId="12" captionText="Figs 3-8: Patrobini, details of external morphology (3-4 - mentum, dorsal view, 5-6 - lateral part of thorax, lateroventral view, 7-8 - female tergite 8, dorsal view, ac2 - middle coxal cavity, epm2 - mesepimeron, epm3 - metepimeron, eps2 - mesepisternum, eps3 - metepisternum, st2 - mesosternum, st3 - metasternum). Indopatrobus bashtai spec. nov., paratype, (3, 5, 7). Parapenetretus selaensis spec. nov., paratype, (4, 6, 8). Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5434690" httpUri="https://zenodo.org/record/5434690/files/figure.png" pageId="6" pageNumber="997">fig. 4</figureCitation>
) with moderately deep furrows, mentum tooth with 2 setae subapically; submentum with 2 setiferous pores on each side; only 2 (exceptionally 3) supraorbital setae present, hind one situated beside neck-constriction; tempora glabrous.
</paragraph>
<paragraph id="9A2E36FDFFE7FF82FF436760FCDDCF47" blockId="6.[151,1059,821,1106]" pageId="6" pageNumber="997">
Disk of pronotum glabrous; lateral margin before its middle polysetose (usually with 2 seta); median line simple; prothorax glabrous; suture separating mesepisternum and mesosternum (
<figureCitation id="02AA2A78FFE7FF82FEF9673AFEA1C890" box="[301,357,879,901]" captionStart="Figs 3-8" captionStartId="12.[151,191,1258,1280]" captionTargetBox="[175,1039,242,1234]" captionTargetId="figure-83@12.[151,1057,235,1248]" captionTargetPageId="12" captionText="Figs 3-8: Patrobini, details of external morphology (3-4 - mentum, dorsal view, 5-6 - lateral part of thorax, lateroventral view, 7-8 - female tergite 8, dorsal view, ac2 - middle coxal cavity, epm2 - mesepimeron, epm3 - metepimeron, eps2 - mesepisternum, eps3 - metepisternum, st2 - mesosternum, st3 - metasternum). Indopatrobus bashtai spec. nov., paratype, (3, 5, 7). Parapenetretus selaensis spec. nov., paratype, (4, 6, 8). Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5434690" httpUri="https://zenodo.org/record/5434690/files/figure.png" pageId="6" pageNumber="997">fig. 6</figureCitation>
) joins lateral margin of metasternum; mesepimeron narrow, slightly broadened laterally and hardly broadened inward, without median process penetrating between meso- and metathorax, disjointed from middle coxal cavity by rather broad area of meso- and metasternum; metepisternum rather long and narrow; front margin of metasternum near middle coxal cavities valliculiform (see 
<figureCitation id="02AA2A78FFE7FF82FD3667B1FCD9C8EF" box="[738,797,996,1018]" captionStart="Figs 3-8" captionStartId="12.[151,191,1258,1280]" captionTargetBox="[175,1039,242,1234]" captionTargetId="figure-83@12.[151,1057,235,1248]" captionTargetPageId="12" captionText="Figs 3-8: Patrobini, details of external morphology (3-4 - mentum, dorsal view, 5-6 - lateral part of thorax, lateroventral view, 7-8 - female tergite 8, dorsal view, ac2 - middle coxal cavity, epm2 - mesepimeron, epm3 - metepimeron, eps2 - mesepisternum, eps3 - metepisternum, st2 - mesosternum, st3 - metasternum). Indopatrobus bashtai spec. nov., paratype, (3, 5, 7). Parapenetretus selaensis spec. nov., paratype, (4, 6, 8). Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5434690" httpUri="https://zenodo.org/record/5434690/files/figure.png" pageId="6" pageNumber="997">fig. 6</figureCitation>
); inner side of profemur without tubercle; metatrochanter normal; male protarsomere 2 distinctly larger than protarsomere 3, latter longitudinal, protarsomere 4 strongly emarginate apically, bilobed; tarsi glabrous on upper surface; tarsomere 5 glabrous ventrally.
</paragraph>
<paragraph id="9A2E36FDFFE7FF82FF436034FC98CF80" blockId="6.[151,1059,1121,1173]" pageId="6" pageNumber="997">Elytra elongate, somewhat depressed; scutellar pore puncture present; elytral striae welldeveloped; setae arranged into discal series present only on interval 3.</paragraph>
<paragraph id="9A2E36FDFFE7FF82FF4360F1FEE8CE07" blockId="6.[151,1059,1188,1298]" pageId="6" pageNumber="997">
Anterolateral apophyse of female tergite 8 indistinct (
<figureCitation id="02AA2A78FFE7FF82FD1060F1FD3ACFAF" box="[708,766,1188,1210]" captionStart="Figs 3-8" captionStartId="12.[151,191,1258,1280]" captionTargetBox="[175,1039,242,1234]" captionTargetId="figure-83@12.[151,1057,235,1248]" captionTargetPageId="12" captionText="Figs 3-8: Patrobini, details of external morphology (3-4 - mentum, dorsal view, 5-6 - lateral part of thorax, lateroventral view, 7-8 - female tergite 8, dorsal view, ac2 - middle coxal cavity, epm2 - mesepimeron, epm3 - metepimeron, eps2 - mesepisternum, eps3 - metepisternum, st2 - mesosternum, st3 - metasternum). Indopatrobus bashtai spec. nov., paratype, (3, 5, 7). Parapenetretus selaensis spec. nov., paratype, (4, 6, 8). Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5434690" httpUri="https://zenodo.org/record/5434690/files/figure.png" pageId="6" pageNumber="997">fig. 8</figureCitation>
), base of tergite moderately exceeding epitergite, longitudinal keels absent, transversal ones long, median sclerotization absent, both basal and apical longitudinal depigmentation absent; abdominal sternites glabrous.
</paragraph>
<paragraph id="9A2E36FDFFE7FF82FF436174FF2ECE9A" blockId="6.[151,1059,1313,1423]" pageId="6" pageNumber="997">
Apical lamella of aedeagus moderately elongate, with lateral keels (
<figureCitation id="02AA2A78FFE7FF82FCB36174FC1ACE22" box="[871,990,1313,1335]" captionStart="Figs 9-18" captionStartId="13.[151,191,1493,1515]" captionTargetBox="[203,992,200,1479]" captionTargetId="figure-76@13.[192,1008,192,1483]" captionTargetPageId="13" captionText="Figs 9-18: Patrobini, male genitalia (9, 14 - aedeagus, left lateral view, 10, 15 - aedeagus, dorsal view, 11, 16 - left paramere, left lateral view, 12, 17 - right paramere, right lateral view, 13, 18 - ventrite 9, dorsal view). Indopatrobus bashtai spec. nov., paratype, (9-13). Parapenetretus selaensis spec. nov., paratype, (14-18). Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5434692" httpUri="https://zenodo.org/record/5434692/files/figure.png" pageId="6" pageNumber="997">figs 14, 15</figureCitation>
); only proximal scleriotization of endophallus present; basal phlagellum absent; parameres fully developed, apical projections rather short, somewhat gradually tapering apically and pointed.
</paragraph>
<paragraph id="9A2E36FDFFE7FF82FF4361CBFD7BCEC4" blockId="6.[151,1058,1438,1489]" pageId="6" pageNumber="997">
Distal sclerite of female reproductive tract present (
<figureCitation id="02AA2A78FFE7FF82FD0A61CBFCEACEA1" box="[734,814,1438,1460]" captionStart="Figs 19-20" captionStartId="14.[151,191,757,779]" captionTargetBox="[168,1047,227,744]" captionTargetId="figure-76@14.[151,1056,204,748]" captionTargetPageId="14" captionText="Figs 19-20: Patrobini, reproductive tract (bc - bursa copulatrix, bs - bursal sclerite, sg - spermathecal gland, sp - spermatheca). Indopatrobus bashtai spec. nov., paratype, (19). Parapenetretus selaensis spec. nov., paratype, (20). Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5434694" httpUri="https://zenodo.org/record/5434694/files/figure.png" pageId="6" pageNumber="997">fig. 20</figureCitation>
); bursal sclerite welldeveloped, asymmetric; stylus with 1 seta subapically.
</paragraph>
<paragraph id="9A2E36FDFFE7FF83FF4361B4FCCDCA6E" blockId="6.[151,1059,1505,1585]" lastBlockId="7.[151,1059,211,379]" lastPageId="7" lastPageNumber="998" pageId="6" pageNumber="997">
<collectionCode id="FC80AE38FFE7FF82FF4361B4FF6CCEE3" box="[151,168,1505,1526]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="6" pageNumber="997">R</collectionCode>
e m a r k s: Judging from the bulk of characters known, 
<taxonomicName id="5D914D7EFFE7FF82FCDC61B7FC53CEE2" authority="A.S &amp; D.W &amp; A &amp; H, 2006" authorityName="A.S &amp; D.W &amp; A &amp; H" authorityYear="2006" box="[776,919,1506,1527]" class="Insecta" family="Carabidae" genus="Parapenetretus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="997" phylum="Arthropoda" rank="subGenus" status="subgen. nov." subGenus="Propenetretus">
<emphasis id="A8E5EAEFFFE7FF82FCDC61B7FC53CEE2" box="[776,919,1506,1527]" italics="true" pageId="6" pageNumber="997">Propenetretus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B3D65794FFE7FF82FC7461B4FBE7CEE3" box="[928,1059,1505,1527]" pageId="6" pageNumber="997" rank="subGenus">subgen. nov.</taxonomicNameLabel>
must be undoubtedly placed close to other taxa of the genus 
<taxonomicName id="5D914D7EFFE7FF82FCC961AAFC7DCD01" box="[797,953,1535,1556]" class="Insecta" family="Carabidae" genus="Parapenetretus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="997" phylum="Arthropoda" rank="genus">
<emphasis id="A8E5EAEFFFE7FF82FCC961AAFC7DCD01" box="[797,953,1535,1556]" italics="true" pageId="6" pageNumber="997">Parapenetretus</emphasis>
</taxonomicName>
. Cladistic analysis, based on the detruncated matrix comprising taxa with all characters known, proves its affiliation with the other groups of 
<taxonomicName id="5D914D7EFFE6FF83FD506481FCE4CBFC" box="[644,800,212,233]" class="Insecta" family="Carabidae" genus="Parapenetretus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="998" phylum="Arthropoda" rank="genus">
<emphasis id="A8E5EAEFFFE6FF83FD506481FCE4CBFC" box="[644,800,212,233]" italics="true" pageId="7" pageNumber="998">Parapenetretus</emphasis>
</taxonomicName>
(
<figureCitation id="02AA2A78FFE6FF83FCE56486FCBDCBFC" box="[817,889,211,233]" captionStart="Fig" captionStartId="14.[151,184,1530,1552]" captionTargetBox="[155,661,840,1517]" captionTargetId="figure-77@14.[151,661,840,1520]" captionTargetPageId="14" captionText="Fig. 21: Simplified majority rule consensus cladogram of the tribe Patrobini, including Indopatrobus gen. nov. and Propenetretus subgen. nov. Length=297, CI=0,29, RI=0,71. Numbers are the percentage of coincidence." figureDoi="http://doi.org/10.5281/zenodo.5434696" httpUri="https://zenodo.org/record/5434696/files/figure.png" pageId="7" pageNumber="998">fig. 21</figureCitation>
) (tree remained unresolved as concerns relationships of 
<emphasis id="A8E5EAEFFFE6FF83FD9264A4FD3ACA13" box="[582,766,241,262]" italics="true" pageId="7" pageNumber="998">Ambigopenetretus</emphasis>
and 
<taxonomicName id="5D914D7EFFE6FF83FCEE64A4FC0DCA13" authority="A.S &amp; D.W &amp; A &amp; H, 2006" authorityName="A.S &amp; D.W &amp; A &amp; H" authorityYear="2006" box="[826,969,241,262]" class="Insecta" family="Carabidae" genus="Parapenetretus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="998" phylum="Arthropoda" rank="subGenus" status="subgen. nov." subGenus="Propenetretus">
<emphasis id="A8E5EAEFFFE6FF83FCEE64A4FC0DCA13" box="[826,969,241,262]" italics="true" pageId="7" pageNumber="998">Propenetretus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B3D65794FFE6FF83FC0064A5FF03CA36" pageId="7" pageNumber="998" rank="subGenus">subgen. nov.</taxonomicNameLabel>
). This monophyletic group of 
<taxonomicName id="5D914D7EFFE6FF83FE2B655AFD5FCA31" box="[511,667,271,292]" class="Insecta" family="Carabidae" genus="Parapenetretus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="998" phylum="Arthropoda" rank="genus">
<emphasis id="A8E5EAEFFFE6FF83FE2B655AFD5FCA31" box="[511,667,271,292]" italics="true" pageId="7" pageNumber="998">Parapenetretus</emphasis>
</taxonomicName>
taxa is observed at 100% of all resulting equally parsimonious trees. Analysis involving all groups (including units with ambiguity) (
<figureCitation id="02AA2A78FFE6FF83FEC2651DFE9FCA4B" box="[278,347,328,350]" captionStart="Fig" captionStartId="15.[151,184,1372,1394]" captionTargetBox="[162,783,1008,1356]" captionTargetId="figure-112@15.[151,791,1007,1363]" captionTargetPageId="15" captionText="Fig. 23: Reconstruction of possible phylogeny of the genus Parapenetretus on one of the most parsimonious trees of Patrobini including units with ambiguity. Length=342, CI=0,26, RI=0,66. White circles indicate homoplasies, black circles indicate aut- or synapomorphies, character numbers indicated above lines, character states below lines (numbers expansion s. ZAMOTAJLOV 2002)." figureDoi="http://doi.org/10.5281/zenodo.5434700" httpUri="https://zenodo.org/record/5434700/files/figure.png" pageId="7" pageNumber="998">fig. 23</figureCitation>
) reveals that 
<taxonomicName id="5D914D7EFFE6FF83FE3D651CFDBCCA4B" authority="A.S &amp; D.W &amp; A &amp; H, 2006" authorityName="A.S &amp; D.W &amp; A &amp; H" authorityYear="2006" box="[489,632,329,350]" class="Insecta" family="Carabidae" genus="Parapenetretus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="998" phylum="Arthropoda" rank="subGenus" status="subgen. nov." subGenus="Propenetretus">
<emphasis id="A8E5EAEFFFE6FF83FE3D651CFDBCCA4B" box="[489,632,329,350]" italics="true" pageId="7" pageNumber="998">Propenetretus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B3D65794FFE6FF83FD54651DFCC0CA4B" box="[640,772,328,350]" pageId="7" pageNumber="998" rank="subGenus">subgen. nov.</taxonomicNameLabel>
represents an adelphotaxon of the entire complex of the known groups of 
<taxonomicName id="5D914D7EFFE6FF83FDBC6533FCC0CA6E" box="[616,772,358,379]" class="Insecta" family="Carabidae" genus="Parapenetretus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="998" phylum="Arthropoda" rank="genus">
<emphasis id="A8E5EAEFFFE6FF83FDBC6533FCC0CA6E" box="[616,772,358,379]" italics="true" pageId="7" pageNumber="998">Parapenetretus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="9A2E36FDFFE6FF83FF4365DEFD9ECAED" blockId="7.[151,1059,395,505]" pageId="7" pageNumber="998">
Similarity dendrograms obtained by different clustering methods (
<collectionCode id="FC80AE38FFE6FF83FC9965DEFC69CAB4" box="[845,941,395,417]" pageId="7" pageNumber="998">UPGMA</collectionCode>
, 
<collectionCode id="FC80AE38FFE6FF83FC6F65DEFBDBCAB4" box="[955,1055,395,417]" pageId="7" pageNumber="998">WPGMA</collectionCode>
, single linkage, complete linkage, etc., example see 
<figureCitation id="02AA2A78FFE6FF83FD7065FDFD2CCAAB" box="[676,744,424,446]" captionStart="Fig" captionStartId="16.[151,184,1497,1519]" captionTargetBox="[151,690,212,1488]" captionTargetId="figure-23@16.[151,690,204,1488]" captionTargetPageId="16" captionText="Fig. 24: Similarity of several Patrobini units on the dendrogram of the tribe constructed using UPGMA method. Scale indicates similarity indices." figureDoi="http://doi.org/10.5281/zenodo.5434702" httpUri="https://zenodo.org/record/5434702/files/figure.png" pageId="7" pageNumber="998">fig. 24</figureCitation>
) possess a stable congregation comprising all groups of 
<taxonomicName id="5D914D7EFFE6FF83FE766593FDFACACE" box="[418,574,454,475]" class="Insecta" family="Carabidae" genus="Parapenetretus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="998" phylum="Arthropoda" rank="genus">
<emphasis id="A8E5EAEFFFE6FF83FE766593FDFACACE" box="[418,574,454,475]" italics="true" pageId="7" pageNumber="998">Parapenetretus</emphasis>
</taxonomicName>
and 
<taxonomicName id="5D914D7EFFE6FF83FDA16593FCC0CACE" authority="A.S &amp; D.W &amp; A &amp; H, 2006" authorityName="A.S &amp; D.W &amp; A &amp; H" authorityYear="2006" box="[629,772,454,475]" class="Insecta" family="Carabidae" genus="Parapenetretus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="998" phylum="Arthropoda" rank="subGenus" status="subgen. nov." subGenus="Propenetretus">
<emphasis id="A8E5EAEFFFE6FF83FDA16593FCC0CACE" box="[629,772,454,475]" italics="true" pageId="7" pageNumber="998">Propenetretus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B3D65794FFE6FF83FCDA6590FC50CACE" box="[782,916,453,475]" pageId="7" pageNumber="998" rank="subGenus">subgen. nov.</taxonomicNameLabel>
, the latter always occupies a position basal to other taxa.
</paragraph>
<paragraph id="9A2E36FDFFE6FF83FF43665DFC8CC94D" blockId="7.[151,1059,520,600]" pageId="7" pageNumber="998">
Thus, basing on both phylogenetic and phenetic analysis, in spite of some pronounced habitual differences, above described taxon must be attributed to the genus 
<taxonomicName id="5D914D7EFFE6FF83FF436616FEF7C94D" box="[151,307,579,600]" class="Insecta" family="Carabidae" genus="Parapenetretus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="998" phylum="Arthropoda" rank="genus">
<emphasis id="A8E5EAEFFFE6FF83FF436616FEF7C94D" box="[151,307,579,600]" italics="true" pageId="7" pageNumber="998">Parapenetretus</emphasis>
</taxonomicName>
, though deserves separation as a different subgenus.
</paragraph>
<paragraph id="9A2E36FDFFE6FF83FF43663DFD7FC98E" blockId="7.[151,1059,616,667]" pageId="7" pageNumber="998">
E t y m o l o g y: The subgeneric epithet refers to the phylogenetic position of this subgenus, basal to other subgenera of 
<taxonomicName id="5D914D7EFFE6FF83FDCD66D3FD71C98E" box="[537,693,646,667]" class="Insecta" family="Carabidae" genus="Parapenetretus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="998" phylum="Arthropoda" rank="genus">
<emphasis id="A8E5EAEFFFE6FF83FDCD66D3FD71C98E" box="[537,693,646,667]" italics="true" pageId="7" pageNumber="998">Parapenetretus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>