793 lines
99 KiB
XML
793 lines
99 KiB
XML
<document id="C18C1ADAAD947E556D51C0648360214D" ID-CLB-Dataset="57778" ID-DOI="10.11646/zootaxa.5319.2.6" ID-GBIF-Dataset="49b790fb-5325-46da-aa80-d36d2e1fcc1a" ID-ISSN="1175-5326" ID-Zenodo-Dep="8182539" ID-ZooBank="065A967F-2832-4BDF-890A-2EA852CACD74" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="valdenar" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="valdenar" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1690278797849" checkinUser="plazi" docAuthor="Chaves, Gerardo, Ryan, Mason J., Bolaños, Federico, Márquez, Cruz, Köhler, Gunther & Poe, Steven" docDate="2023" docId="09619A08FFA4190D8FECF9D69A64FEA5" docLanguage="en" docName="zootaxa.5319.2.6.pdf" docOrigin="Zootaxa 5319 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5319.2.6" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Anolis riparius Chaves & Ryan & Bolaños & Márquez & Köhler & Poe 2023, new species" docType="treatment" docVersion="6" lastPageNumber="259" masterDocId="F558E270FFAC19068F7BFFD09E22FFE4" masterDocTitle="Two new species of semiaquatic Anolis (Squamata: Dactyloidae) from Costa Rica" masterLastPageNumber="262" masterPageNumber="249" pageNumber="257" updateTime="1699546558579" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="1665BAE51C323C12D71026CC07040012">Two new species of semiaquatic Anolis (Squamata: Dactyloidae) from Costa Rica</mods:title>
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<mods:namePart id="FA4878EE1A37EC7FD91B91522C5F2489">Chaves, Gerardo</mods:namePart>
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<mods:affiliation id="F32EB180130AA948E43AB491D5A1DA1D">Centro de Investigación en Biodiversidad y Ecología Tropical (Museo de Zoología) de la Universidad de Costa Rica & cachi 13 @ gmail. com; https: // orcid. org / 0000 - 0002 - 4301 - 6569</mods:affiliation>
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<mods:nameIdentifier id="04E32E3FAADB28A46B80AA6BEF6C249C" type="email">cachi13@gmail.com</mods:nameIdentifier>
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<mods:namePart id="46E099DE1C1CC748A8BAB1A570C541F5">Ryan, Mason J.</mods:namePart>
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<mods:affiliation id="A6CF97F1E6816F07EFF3CFA3FCF8B00C">Department of Biology, University of New Mexico, Albuquerque, NM, 87131, USA & Arizona Game and Fish Department, 5000 W Carefree Hwy, Phoenix, Arizona 85086</mods:affiliation>
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<mods:namePart id="546E2ED83EDD8F17E6B0C8A0F85A0DDE">Bolaños, Federico</mods:namePart>
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<mods:affiliation id="043007CF5F39F3D783905574D915F696">Centro de Investigación en Biodiversidad y Ecología Tropical (Museo de Zoología) de la Universidad de Costa Rica & Escuela de Biología, Universidad de Costa Rica, San Pedro, Costa Rica & federico. bolanos @ ucr. ac. cr; https: // orcid. org / 0000 - 0002 - 7935 - 6418</mods:affiliation>
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<mods:nameIdentifier id="1A9A5749A6A11822DE9AB82C2D7AEA7B" type="email">federico.bolanos@ucr.ac.cr</mods:nameIdentifier>
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<mods:namePart id="DD10AC63261156380744B626BB64BDBC">Márquez, Cruz</mods:namePart>
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<mods:affiliation id="0403414A76798B6B88DA2A36DFB82B55">Direccion, Conservacion, Restauracion y Desarrollo Sustentable de Ecosistemas Insulares; Parque Nacional Galapagos, Ecuador cruzkallw @ gmail. com; https: // orcid. org / 0009 - 0003 - 2220 - 7379</mods:affiliation>
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<mods:nameIdentifier id="2337C5DDFCC37626F0E4FB2D0C427F5F" type="email">cruzkallw@gmail.com</mods:nameIdentifier>
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<mods:namePart id="F8C402AE5CB76230E31FBF3F7E3F83C9">Köhler, Gunther</mods:namePart>
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<mods:affiliation id="D5660449B54042F284F9107A20FED6B9">Senckenberg Forschungsinstitut und Naturmuseum, Senckenberganlage 25, 60325 Frankfurt a. M., Germany gkoehler @ senckenberg. de; https: // orcid. org / 0000 - 0002 - 2563 - 5331</mods:affiliation>
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<mods:nameIdentifier id="4DE22B0D63000F75CD6C90C54AAA649E" type="email">gkoehler@senckenberg.de</mods:nameIdentifier>
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<mods:namePart id="EE94DF5889C475727957662674372C3E">Poe, Steven</mods:namePart>
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<mods:affiliation id="D65C6DCE6DF33F81624F0EF2501901F1">Department of Biology, University of New Mexico, Albuquerque, NM, 87131, USA</mods:affiliation>
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<mods:title id="7212150BCA5984511FFB43519E586C0E">Zootaxa</mods:title>
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<mods:date id="3051FC847F74E7A6980A43B207C612B2">2023</mods:date>
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<mods:number id="62404FFDB8712383E2A7677736918DE4">2023-07-25</mods:number>
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<mods:number id="FAB2C7CB1828804B126CEEA9D583AAAD">5319</mods:number>
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<mods:url id="F286DD89DD1893F16271F5AE13B6DEB6">http://dx.doi.org/10.11646/zootaxa.5319.2.6</mods:url>
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<treatment id="09619A08FFA4190D8FECF9D69A64FEA5" ID-DOI="http://doi.org/10.5281/zenodo.8184547" ID-GBIF-Taxon="212419560" ID-Zenodo-Dep="8184547" LSID="urn:lsid:plazi:treatment:09619A08FFA4190D8FECF9D69A64FEA5" httpUri="http://treatment.plazi.org/id/09619A08FFA4190D8FECF9D69A64FEA5" lastPageId="11" lastPageNumber="259" pageId="8" pageNumber="257">
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<taxonomicName id="46C8509DFFA4190E8FECF9D69F66F9C5" ID-CoL="675PJ" authority="Chaves & Ryan & Bolaños & Márquez & Köhler & Poe, 2023" authorityName="Chaves & Ryan & Bolaños & Márquez & Köhler & Poe" authorityYear="2023" box="[151,324,1542,1569]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="8" pageNumber="257" phylum="Chordata" rank="species" species="riparius" status="sp. nov.">
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<emphasis id="B3BCF70CFFA4190E8FECF9D69F66F9C5" bold="true" box="[151,324,1542,1569]" italics="true" pageId="8" pageNumber="257">Anolis riparius</emphasis>
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</taxonomicName>
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,
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<subSubSection id="C9D27895FFA4190F8FECF99F9A3EFE49" lastPageId="9" lastPageNumber="258" pageId="8" pageNumber="257" type="materials_examined">
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<paragraph id="81772B1EFFA4190E8FECF99F9EF2F955" blockId="8.[151,1437,1614,2037]" pageId="8" pageNumber="257">
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<materialsCitation id="31A02143FFA4190E8FECF99F9EF2F955" ID-GBIF-Occurrence="4164459315" collectingDate="2009-09-07" country="Costa Rica" county="Perez Zeledon" elevation="880" latitude="9.3139" location="Douglas Robinson" longLatPrecision="1" longitude="-83.7722" municipality="Km" pageId="8" pageNumber="257" specimenCount="1" stateProvince="San Jose" typeStatus="holotype">
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<emphasis id="B3BCF70CFFA4190E8FECF99F9EDDF98D" bold="true" box="[151,255,1615,1641]" pageId="8" pageNumber="257">
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<typeStatus id="5E7395BCFFA4190E8FECF99F9EDDF98D" box="[151,255,1615,1641]" pageId="8" pageNumber="257" type="holotype">Holotype</typeStatus>
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</emphasis>
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. UCR 5579 (
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<figureCitation id="19F3379BFFA4190E8EF4F99F9FEBF98D" box="[399,457,1615,1641]" captionStart="FIGURE 6" captionStartId="9.[152,255,1398,1422]" captionTargetBox="[151,1436,453,1375]" captionTargetId="figure-172@9.[151,1436,453,1375]" captionTargetPageId="9" captionText="FIGURE 6. A) Dorsal body and B) dewlap views of Anolis riparius in life. C) Dorsal view of head and D) Small middorsal scales at the base of the tail of paratype (UCR 5579) of A. riparius between breakpoint (a) and the first enlarge middorsal caudal scale (b). Photos A and B by David Laurencio and C and D by Gerardo Chaves." figureDoi="http://doi.org/10.5281/zenodo.8182551" httpUri="https://zenodo.org/record/8182551/files/figure.png" pageId="8" pageNumber="257">Fig 6</figureCitation>
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) collected on
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<date id="F5760DDEFFA4190E8D1BF99F9D01F98D" box="[608,803,1615,1641]" pageId="8" pageNumber="257" value="2009-09-07">
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<collectingDate id="E532F436FFA4190E8D1BF99F9D01F98D" box="[608,803,1615,1641]" pageId="8" pageNumber="257" value="2009-09-07">7 September 2009</collectingDate>
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</date>
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by
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and his students; an adult male from
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<location id="84177DC5FFA4190E8FECF9A39F15F969" LSID="urn:lsid:plazi:treatment:09619A08FFA4190D8FECF9D69A64FEA5:84177DC5FFA4190E8FECF9A39F15F969" box="[151,311,1651,1677]" country="Costa Rica" county="Perez Zeledon" latitude="9.3139" longLatPrecision="1" longitude="-83.7722" municipality="Km" name="Pacuarito river" pageId="8" pageNumber="257" stateProvince="San Jose">Pacuarito river</location>
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,
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<location id="84177DC5FFA4190E8E3AF9A39F84F969" LSID="urn:lsid:plazi:treatment:09619A08FFA4190D8FECF9D69A64FEA5:84177DC5FFA4190E8E3AF9A39F84F969" box="[321,422,1651,1677]" country="Costa Rica" county="Perez Zeledon" latitude="9.3139" longLatPrecision="1" longitude="-83.7722" municipality="Km" name="Pacuarito" pageId="8" pageNumber="257" stateProvince="San Jose">Pacuarito</location>
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10
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<collectingMunicipality id="6113B164FFA4190E8EB5F9A39FD5F969" box="[462,503,1651,1677]" pageId="8" pageNumber="257">Km</collectingMunicipality>
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S-SW of
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<collectingCounty id="68165392FFA4190E8D19F9A39CDCF969" box="[610,766,1651,1677]" pageId="8" pageNumber="257">Pérez Zeledón</collectingCounty>
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,
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, -
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,
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<quantity id="463086FBFFA4190E8B5BF9A39AA6F969" box="[1056,1156,1651,1677]" metricMagnitude="2" metricUnit="m" metricValue="8.8" pageId="8" pageNumber="257" unit="m" value="880.0">
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<elevation id="0AE5CC2DFFA4190E8B5BF9A39AA6F969" box="[1056,1156,1651,1677]" metricMagnitude="2" metricUnit="m" metricValue="8.8" pageId="8" pageNumber="257" unit="m" value="880.0">880 masl</elevation>
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</quantity>
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,
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<collectingRegion id="430CE5FCFFA4190E8BF4F9A39B76F969" box="[1167,1364,1651,1677]" country="Costa Rica" name="San Jose" pageId="8" pageNumber="257">San José Province</collectingRegion>
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,
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<collectingCountry id="F9DF6B8EFFA4190E8A24F9A39EEEF955" name="Costa Rica" pageId="8" pageNumber="257">Costa Rica</collectingCountry>
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.
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</materialsCitation>
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</paragraph>
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<paragraph id="81772B1EFFA4190F8FBCF96B9A3EFE49" blockId="8.[151,1437,1614,2037]" lastBlockId="9.[151,1436,150,429]" lastPageId="9" lastPageNumber="258" pageId="8" pageNumber="257">
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<materialsCitation id="31A02143FFA4190E8FBCF96B9B32F91D" ID-GBIF-Occurrence="4164459304" collectingDate="2002-10-09" collectionCode="SMF" collectorName="Axel Flaschendrager" country="Costa Rica" latitude="9.31056" location="Adult" longitude="-83.775" pageId="8" pageNumber="257" specimenCode="SMF 82183-82184" specimenCount="1" stateProvince="San Jose" typeStatus="paratype">
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<emphasis id="B3BCF70CFFA4190E8FBCF96B9F18F931" bold="true" box="[199,314,1723,1749]" pageId="8" pageNumber="257">
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<typeStatus id="5E7395BCFFA4190E8FBCF96B9F18F931" box="[199,314,1723,1749]" pageId="8" pageNumber="257" type="paratype">Paratypes</typeStatus>
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</emphasis>
|
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(all from
|
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<collectingCountry id="F9DF6B8EFFA4190E8ED8F96B9C39F931" box="[419,539,1723,1749]" name="Costa Rica" pageId="8" pageNumber="257">Costa Rica</collectingCountry>
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:
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<collectingRegion id="430CE5FCFFA4190E8D5FF96B9CC5F931" box="[548,743,1723,1749]" country="Costa Rica" name="San Jose" pageId="8" pageNumber="257">San José Province</collectingRegion>
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).
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<emphasis id="B3BCF70CFFA4190E8D81F96A9DA6F931" bold="true" box="[762,900,1722,1749]" pageId="8" pageNumber="257">
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<location id="84177DC5FFA4190E8D81F96A9D1EF930" LSID="urn:lsid:plazi:treatment:09619A08FFA4190D8FECF9D69A64FEA5:84177DC5FFA4190E8D81F96A9D1EF930" box="[762,828,1722,1748]" country="Costa Rica" latitude="9.31056" name="Adult" pageId="8" pageNumber="257" stateProvince="San Jose">Adult</location>
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males
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||
</emphasis>
|
||
:
|
||
<specimenCode id="D16E8365FFA4190E8CEAF96B9A42F931" box="[913,1120,1723,1749]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="8" pageNumber="257">SMF 82183–82184</specimenCode>
|
||
(collected
|
||
<date id="F5760DDEFFA4190E8BADF96B9B59F931" box="[1238,1403,1723,1749]" pageId="8" pageNumber="257" value="2002-10-09">
|
||
<collectingDate id="E532F436FFA4190E8BADF96B9B59F931" box="[1238,1403,1723,1749]" pageId="8" pageNumber="257" value="2002-10-09">9 October 2002</collectingDate>
|
||
</date>
|
||
by
|
||
<collectorName id="2C3D4EC8FFA4190E8FECF90F9F55F91D" box="[151,375,1759,1785]" pageId="8" pageNumber="257">Axel Fläschendräger</collectorName>
|
||
) from road
|
||
<locationDeviation id="90C5323CFFA4190E8E8CF90F9D43F91D" box="[503,865,1759,1785]" firstLocation="Platanillo" pageId="8" pageNumber="257" secondLocation="San Isidro">between Platanillo and San Isidro</locationDeviation>
|
||
,
|
||
<geoCoordinate id="E4FC4DD9FFA4190E8C16F90E9DC3F91D" box="[877,993,1758,1785]" degrees="9.31056" direction="north" orientation="latitude" pageId="8" pageNumber="257" precision="1" value="9.31056">9.31056 N</geoCoordinate>
|
||
; -
|
||
<geoCoordinate id="E4FC4DD9FFA4190E8C8FF90E9A59F91D" box="[1012,1147,1758,1785]" degrees="83.77500" direction="west" orientation="longitude" pageId="8" pageNumber="257" precision="1" value="-83.775">83.77500 W</geoCoordinate>
|
||
, ca.
|
||
<quantity id="463086FBFFA4190E8BD0F90E9B32F91D" box="[1195,1296,1758,1785]" metricMagnitude="2" metricUnit="m" metricValue="8.5" pageId="8" pageNumber="257" unit="m" value="850.0">850 masl</quantity>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="31A02143FFA4190E8A60F90F9B39F8F9" ID-GBIF-Occurrence="4164459309" collectingDate="2010-02-15" collectionCode="SMF" collectorName="Gunther Kohler" country="Costa Rica" elevation="1425" latitude="9.35789" location="1 km N Cedral" longLatPrecision="1" longitude="-83.56094" pageId="8" pageNumber="257" specimenCode="SMF 92428" specimenCount="1" stateProvince="San Jose" typeStatus="paratype">
|
||
<specimenCode id="D16E8365FFA4190E8A60F90F9BBEF91D" box="[1307,1436,1759,1785]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="8" pageNumber="257">SMF 92428</specimenCode>
|
||
(collected
|
||
<date id="F5760DDEFFA4190E8E7CF8D29FEAF8F9" box="[263,456,1794,1821]" pageId="8" pageNumber="257" value="2010-02-15">
|
||
<collectingDate id="E532F436FFA4190E8E7CF8D29FEAF8F9" box="[263,456,1794,1821]" pageId="8" pageNumber="257" value="2010-02-15">15 February 2010</collectingDate>
|
||
</date>
|
||
by
|
||
<collectorName id="2C3D4EC8FFA4190E8E8AF8D39CBEF8F9" box="[497,668,1795,1821]" pageId="8" pageNumber="257">Gunther Köhler</collectorName>
|
||
) from
|
||
<location id="84177DC5FFA4190E8D9DF8D39DA5F8F9" LSID="urn:lsid:plazi:treatment:09619A08FFA4190D8FECF9D69A64FEA5:84177DC5FFA4190E8D9DF8D39DA5F8F9" box="[742,903,1795,1821]" country="Costa Rica" latitude="9.35789" longLatPrecision="1" longitude="-83.56094" name="1 km N Cedral" pageId="8" pageNumber="257" stateProvince="San Jose">
|
||
<quantity id="463086FBFFA4190E8D9DF8D39D3CF8F9" box="[742,798,1795,1821]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="8" pageNumber="257" unit="km" value="1.0">1 km</quantity>
|
||
N Cedral
|
||
</location>
|
||
<geoCoordinate id="E4FC4DD9FFA4190E8CF4F8D29A26F8F9" box="[911,1028,1794,1821]" degrees="9.35789" direction="north" orientation="latitude" pageId="8" pageNumber="257" precision="1" value="9.35789">9.35789 N</geoCoordinate>
|
||
, -
|
||
<geoCoordinate id="E4FC4DD9FFA4190E8B6DF8D29ABFF8F9" box="[1046,1181,1794,1821]" degrees="83.56094" direction="west" orientation="longitude" pageId="8" pageNumber="257" precision="1" value="-83.56094">83.56094 W</geoCoordinate>
|
||
,
|
||
<quantity id="463086FBFFA4190E8BD3F8D29B39F8F9" box="[1192,1307,1794,1821]" metricMagnitude="3" metricUnit="m" metricValue="1.425" pageId="8" pageNumber="257" unit="m" value="1425.0">
|
||
<elevation id="0AE5CC2DFFA4190E8BD3F8D29B39F8F9" box="[1192,1307,1794,1821]" metricMagnitude="3" metricUnit="m" metricValue="1.425" pageId="8" pageNumber="257" unit="m" value="1425.0">1425 masl</elevation>
|
||
</quantity>
|
||
</materialsCitation>
|
||
; UCR 7893 (collected
|
||
<date id="F5760DDEFFA4190E8E7CF8F79F90F8A5" box="[263,434,1831,1857]" pageId="8" pageNumber="257" value="1979-08-24">24 August 1979</date>
|
||
by Douglas Robinson) from I.85 Km NE Alfombra,
|
||
<geoCoordinate id="E4FC4DD9FFA4190E8C85F8F79A4BF8A5" box="[1022,1129,1831,1857]" degrees="9.3167" direction="north" orientation="latitude" pageId="8" pageNumber="257" precision="5" value="9.3167">9.3167 N</geoCoordinate>
|
||
, -
|
||
<geoCoordinate id="E4FC4DD9FFA4190E8B07F8F79AD9F8A5" box="[1148,1275,1831,1857]" degrees="83.7722" direction="west" orientation="longitude" pageId="8" pageNumber="257" precision="5" value="-83.7722">83.7722 W</geoCoordinate>
|
||
, ca.
|
||
<quantity id="463086FBFFA4190E8A55F8F79BB5F8A5" box="[1326,1431,1831,1857]" metricMagnitude="2" metricUnit="m" metricValue="9.3" pageId="8" pageNumber="257" unit="m" value="930.0">930 masl</quantity>
|
||
; MCZ-R 18171 (collected
|
||
<date id="F5760DDEFFA4190E8EBBF89A9CB8F881" box="[448,666,1866,1893]" pageId="8" pageNumber="257" value="2007-12-15">15 December 2007</date>
|
||
by Steve Poe and Mason J. Ryan) from Alfombra, 12 Km S-SW of Pérez Zeledón (
|
||
<geoCoordinate id="E4FC4DD9FFA4190E8E3AF8BF9F95F86D" box="[321,439,1903,1929]" degrees="9.31670" direction="north" orientation="latitude" pageId="8" pageNumber="257" precision="1" value="9.3167">9.31670 N</geoCoordinate>
|
||
, -
|
||
<geoCoordinate id="E4FC4DD9FFA4190E8EB1F8BF9C73F86D" box="[458,593,1903,1929]" degrees="83.77220" direction="west" orientation="longitude" pageId="8" pageNumber="257" precision="1" value="-83.7722">83.77220 W</geoCoordinate>
|
||
,
|
||
<quantity id="463086FBFFA4190E8D26F8BF9CE1F86D" box="[605,707,1903,1929]" metricMagnitude="2" metricUnit="m" metricValue="9.3" pageId="8" pageNumber="257" unit="m" value="930.0">930 masl</quantity>
|
||
);
|
||
<materialsCitation id="31A02143FFA4190E8DACF8BF9A8CF849" ID-GBIF-Occurrence="4164459305" collectingDate="2009-01-10" collectionCode="MSB" country="Costa Rica" elevation="930" latitude="9.3167" longLatPrecision="1" longitude="-83.7722" pageId="8" pageNumber="257" specimenCode="MSB 95201" specimenCount="1" stateProvince="San Jose" typeStatus="paratype">
|
||
<specimenCode id="D16E8365FFA4190E8DACF8BF9D79F86D" box="[727,859,1902,1929]" collectionCode="MSB" country="USA" httpUri="http://grbio.org/cool/km06-gtbn" name="Museum of Southwestern Biology" pageId="8" pageNumber="257" type="Museum">MSB 95201</specimenCode>
|
||
(collected on
|
||
<date id="F5760DDEFFA4190E8C8DF8BF9A8BF86D" box="[1014,1193,1903,1929]" pageId="8" pageNumber="257" value="2009-01-10">
|
||
<collectingDate id="E532F436FFA4190E8C8DF8BF9A8BF86D" box="[1014,1193,1903,1929]" pageId="8" pageNumber="257" value="2009-01-10">10 January 2009</collectingDate>
|
||
</date>
|
||
by
|
||
<collectorName id="2C3D4EC8FFA4190E8BA8F8BF9B3EF86D" box="[1235,1308,1903,1929]" pageId="8" pageNumber="257">Mason</collectorName>
|
||
J
|
||
<collectorName id="2C3D4EC8FFA4190E8A4DF8BF9B4CF86D" box="[1334,1390,1903,1929]" pageId="8" pageNumber="257">Ryan</collectorName>
|
||
and
|
||
<collectorName id="2C3D4EC8FFA4190E8FECF8439F29F849" box="[151,267,1939,1965]" pageId="8" pageNumber="257">Ian Latella</collectorName>
|
||
) from
|
||
<collectorName id="2C3D4EC8FFA4190E8E28F8439F9CF849" box="[339,446,1939,1965]" pageId="8" pageNumber="257">Alfombra</collectorName>
|
||
, 12
|
||
<collectorName id="2C3D4EC8FFA4190E8E90F8439C36F849" box="[491,532,1939,1965]" pageId="8" pageNumber="257">Km</collectorName>
|
||
S-SW of
|
||
<collectorName id="2C3D4EC8FFA4190E8D04F8439D3BF849" box="[639,793,1939,1965]" pageId="8" pageNumber="257">Pérez Zeledón</collectorName>
|
||
(
|
||
<geoCoordinate id="E4FC4DD9FFA4190E8C5CF8439DBFF849" box="[807,925,1939,1965]" degrees="9.31670" direction="north" orientation="latitude" pageId="8" pageNumber="257" precision="1" value="9.3167">9.31670 N</geoCoordinate>
|
||
, -
|
||
<geoCoordinate id="E4FC4DD9FFA4190E8CD4F8439A14F849" box="[943,1078,1939,1965]" degrees="83.77220" direction="west" orientation="longitude" pageId="8" pageNumber="257" precision="1" value="-83.7722">83.77220 W</geoCoordinate>
|
||
,
|
||
<quantity id="463086FBFFA4190E8B3BF8439A85F849" box="[1088,1191,1939,1965]" metricMagnitude="2" metricUnit="m" metricValue="9.3" pageId="8" pageNumber="257" unit="m" value="930.0">
|
||
<elevation id="0AE5CC2DFFA4190E8B3BF8439A85F849" box="[1088,1191,1939,1965]" metricMagnitude="2" metricUnit="m" metricValue="9.3" pageId="8" pageNumber="257" unit="m" value="930.0">930 masl</elevation>
|
||
</quantity>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="31A02143FFA4190E8BC3F8429EF4F811" ID-GBIF-Occurrence="4164459310" collectingDate="2002-10-09" collectionCode="SMF" collectorName="Axel Flaschendrager" country="Costa Rica" elevation="850" location="Adult" pageId="8" pageNumber="257" specimenCode="SMF 82185-82186" specimenCount="1" stateProvince="San Jose" typeStatus="paratype">
|
||
<emphasis id="B3BCF70CFFA4190E8BC3F8429B75F848" bold="true" box="[1208,1367,1938,1964]" pageId="8" pageNumber="257">
|
||
<location id="84177DC5FFA4190E8BC3F8429AD8F848" LSID="urn:lsid:plazi:treatment:09619A08FFA4190D8FECF9D69A64FEA5:84177DC5FFA4190E8BC3F8429AD8F848" box="[1208,1274,1938,1964]" country="Costa Rica" name="Adult" pageId="8" pageNumber="257" stateProvince="San Jose">Adult</location>
|
||
females
|
||
</emphasis>
|
||
:
|
||
<specimenCode id="D16E8365FFA4190E8A1EF8439F09F835" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="8" pageNumber="257">SMF 82185–82186</specimenCode>
|
||
(collected
|
||
<date id="F5760DDEFFA4190E8ED8F8679C69F835" box="[419,587,1975,2001]" pageId="8" pageNumber="257" value="2002-10-09">
|
||
<collectingDate id="E532F436FFA4190E8ED8F8679C69F835" box="[419,587,1975,2001]" pageId="8" pageNumber="257" value="2002-10-09">9 October 2002</collectingDate>
|
||
</date>
|
||
by
|
||
<collectorName id="2C3D4EC8FFA4190E8D08F8679D71F835" box="[627,851,1975,2001]" pageId="8" pageNumber="257">Axel Fläschendräger</collectorName>
|
||
) from road
|
||
<locationDeviation id="90C5323CFFA4190E8CAFF8679B1CF835" box="[980,1342,1975,2001]" firstLocation="Platanillo" pageId="8" pageNumber="257" secondLocation="San Isidro">between Platanillo and San Isidro</locationDeviation>
|
||
(ca.
|
||
<quantity id="463086FBFFA4190E8A08F8669EECF811" metricMagnitude="2" metricUnit="m" metricValue="8.5" pageId="8" pageNumber="257" unit="m" value="850.0">
|
||
<elevation id="0AE5CC2DFFA4190E8A08F8669EECF811" metricMagnitude="2" metricUnit="m" metricValue="8.5" pageId="8" pageNumber="257" unit="m" value="850.0">850 masl</elevation>
|
||
</quantity>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="31A02143FFA4190E8F9AF80B9B79F811" ID-GBIF-Occurrence="4164459318" box="[225,1371,2010,2037]" collectingDate="2002-10-09" collectionCode="SMF" collectorName="Axel Flaschendrager" country="Costa Rica" elevation="850" location="Platanillo" pageId="8" pageNumber="257" specimenCode="SMF 83093-83094" specimenCount="1" stateProvince="San Jose" typeStatus="paratype">
|
||
<specimenCode id="D16E8365FFA4190E8F9AF80B9F93F811" box="[225,433,2011,2037]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="8" pageNumber="257">SMF 83093–83094</specimenCode>
|
||
(collected
|
||
<date id="F5760DDEFFA4190E8D53F80B9CEDF811" box="[552,719,2011,2037]" pageId="8" pageNumber="257" value="2002-10-09">
|
||
<collectingDate id="E532F436FFA4190E8D53F80B9CEDF811" box="[552,719,2011,2037]" pageId="8" pageNumber="257" value="2002-10-09">9 October 2002</collectingDate>
|
||
</date>
|
||
by
|
||
<collectorName id="2C3D4EC8FFA4190E8D8DF80B9DF4F811" box="[758,982,2011,2037]" pageId="8" pageNumber="257">Axel Fläschendräger</collectorName>
|
||
) from near
|
||
<location id="84177DC5FFA4190E8B28F80B9A9BF811" LSID="urn:lsid:plazi:treatment:09619A08FFA4190D8FECF9D69A64FEA5:84177DC5FFA4190E8B28F80B9A9BF811" box="[1107,1209,2011,2037]" country="Costa Rica" name="Platanillo" pageId="8" pageNumber="257" stateProvince="San Jose">Platanillo</location>
|
||
(ca.
|
||
<quantity id="463086FBFFA4190E8B96F80A9B71F811" box="[1261,1363,2010,2037]" metricMagnitude="2" metricUnit="m" metricValue="8.5" pageId="8" pageNumber="257" unit="m" value="850.0">
|
||
<elevation id="0AE5CC2DFFA4190E8B96F80A9B71F811" box="[1261,1363,2010,2037]" metricMagnitude="2" metricUnit="m" metricValue="8.5" pageId="8" pageNumber="257" unit="m" value="850.0">850 masl</elevation>
|
||
</quantity>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="31A02143FFA4190F8A1DF80B9B78FF55" ID-GBIF-Occurrence="4164459303" collectingDate="2010-02-15" collectionCode="SMF" collectorName="Gunther Kohler" country="Costa Rica" elevation="1425" lastPageId="9" lastPageNumber="258" latitude="9.35789" location="1 km N Cedral" longLatPrecision="1" longitude="-83.56094" pageId="8" pageNumber="257" specimenCode="SMF 92429" specimenCount="1" stateProvince="San Jose" typeStatus="paratype">
|
||
<specimenCode id="D16E8365FFA4190F8A1DF80B9EF8FF55" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lastPageId="9" lastPageNumber="258" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="8" pageNumber="257">SMF 92429</specimenCode>
|
||
(collected
|
||
<date id="F5760DDEFFA5190F8E34FF469C2EFF55" box="[335,524,150,177]" pageId="9" pageNumber="258" value="2010-02-15">
|
||
<collectingDate id="E532F436FFA5190F8E34FF469C2EFF55" box="[335,524,150,177]" pageId="9" pageNumber="258" value="2010-02-15">15 February 2010</collectingDate>
|
||
</date>
|
||
by
|
||
<collectorName id="2C3D4EC8FFA5190F8D49FF479CFEFF55" box="[562,732,151,177]" pageId="9" pageNumber="258">Gunther Köhler</collectorName>
|
||
) from
|
||
<location id="84177DC5FFA5190F8C59FF479D9CFF55" LSID="urn:lsid:plazi:treatment:09619A08FFA4190D8FECF9D69A64FEA5:84177DC5FFA5190F8C59FF479D9CFF55" box="[802,958,151,177]" country="Costa Rica" latitude="9.35789" longLatPrecision="1" longitude="-83.56094" name="1 km N Cedral" pageId="9" pageNumber="258" stateProvince="San Jose">
|
||
<quantity id="463086FBFFA5190F8C59FF479D7BFF55" box="[802,857,151,177]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="9" pageNumber="258" unit="km" value="1.0">1 km</quantity>
|
||
N Cedral
|
||
</location>
|
||
(
|
||
<geoCoordinate id="E4FC4DD9FFA5190F8CB7FF469A62FF55" box="[972,1088,150,177]" degrees="9.35789" direction="north" orientation="latitude" pageId="9" pageNumber="258" precision="1" value="9.35789">9.35789 N</geoCoordinate>
|
||
, -
|
||
<geoCoordinate id="E4FC4DD9FFA5190F8B2AFF469AF5FF55" box="[1105,1239,150,177]" degrees="83,56094" direction="west" orientation="longitude" pageId="9" pageNumber="258" precision="1" value="-83.56094">83,56094 W</geoCoordinate>
|
||
,
|
||
<quantity id="463086FBFFA5190F8B9BFF469B71FF55" box="[1248,1363,150,177]" metricMagnitude="3" metricUnit="m" metricValue="1.425" pageId="9" pageNumber="258" unit="m" value="1425.0">
|
||
<elevation id="0AE5CC2DFFA5190F8B9BFF469B71FF55" box="[1248,1363,150,177]" metricMagnitude="3" metricUnit="m" metricValue="1.425" pageId="9" pageNumber="258" unit="m" value="1425.0">1425 masl</elevation>
|
||
</quantity>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="31A02143FFA5190F8A1FFF479FBDFF1D" ID-GBIF-Occurrence="4164459308" collectingDate="2001-03-03" collectionCode="SMF" country="Costa Rica" elevation="800" latitude="9.2293" location="1 km N Cedral" longLatPrecision="1" longitude="-83.642" pageId="9" pageNumber="258" specimenCount="1" stateProvince="San Jose" typeStatus="holotype">
|
||
UCR 5580 (same data of the
|
||
<typeStatus id="5E7395BCFFA5190F8EEBFF6B9FD0FF31" box="[400,498,187,213]" pageId="9" pageNumber="258" type="holotype">holotype</typeStatus>
|
||
); UCR 15959 (collected
|
||
<date id="F5760DDEFFA5190F8D87FF6B9DB0FF31" box="[764,914,187,213]" pageId="9" pageNumber="258" value="2001-03-03">
|
||
<collectingDate id="E532F436FFA5190F8D87FF6B9DB0FF31" box="[764,914,187,213]" pageId="9" pageNumber="258" value="2001-03-03">3 March 2001</collectingDate>
|
||
</date>
|
||
by
|
||
<collectorName id="2C3D4EC8FFA5190F8CC1FF6B9A21FF31" box="[954,1027,187,213]" pageId="9" pageNumber="258">Mason</collectorName>
|
||
J
|
||
<collectorName id="2C3D4EC8FFA5190F8B60FF6B9A76FF31" box="[1051,1108,187,213]" pageId="9" pageNumber="258">Ryan</collectorName>
|
||
) from
|
||
<collectorName id="2C3D4EC8FFA5190F8BE7FF6B9B36FF31" box="[1180,1300,187,213]" pageId="9" pageNumber="258">Mollejones</collectorName>
|
||
(
|
||
<geoCoordinate id="E4FC4DD9FFA5190F8A59FF6B9BBAFF31" box="[1314,1432,187,213]" degrees="9.22930" direction="north" orientation="latitude" pageId="9" pageNumber="258" precision="1" value="9.2293">9.22930 N</geoCoordinate>
|
||
, -
|
||
<geoCoordinate id="E4FC4DD9FFA5190F8FE5FF0F9F07FF1D" box="[158,293,223,249]" degrees="83.64200" direction="west" orientation="longitude" pageId="9" pageNumber="258" precision="1" value="-83.642">83.64200 W</geoCoordinate>
|
||
,
|
||
<quantity id="463086FBFFA5190F8E4BFF0F9FB5FF1D" box="[304,407,223,249]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="9" pageNumber="258" unit="m" value="800.0">
|
||
<elevation id="0AE5CC2DFFA5190F8E4BFF0F9FB5FF1D" box="[304,407,223,249]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="9" pageNumber="258" unit="m" value="800.0">800 masl</elevation>
|
||
</quantity>
|
||
)
|
||
</materialsCitation>
|
||
; UCR 16162 (collected
|
||
<date id="F5760DDEFFA5190F8DDFFF0F9D01FF1D" box="[676,803,223,249]" pageId="9" pageNumber="258" value="2001-07-02">2 July 2001</date>
|
||
by Mason J Ryan) from Alfombra, 12 Km S-SW of Pérez Zeledón (
|
||
<geoCoordinate id="E4FC4DD9FFA5190F8F86FED39F50FEF9" box="[253,370,259,285]" degrees="9.31670" direction="north" orientation="latitude" pageId="9" pageNumber="258" precision="1" value="9.3167">9.31670 N</geoCoordinate>
|
||
, -
|
||
<geoCoordinate id="E4FC4DD9FFA5190F8EF8FED39C28FEF9" box="[387,522,259,285]" degrees="83.77220" direction="west" orientation="longitude" pageId="9" pageNumber="258" precision="1" value="-83.7722">83.77220 W</geoCoordinate>
|
||
,
|
||
<quantity id="463086FBFFA5190F8D6FFED39C58FEF9" box="[532,634,259,285]" metricMagnitude="2" metricUnit="m" metricValue="9.3" pageId="9" pageNumber="258" unit="m" value="930.0">930 masl</quantity>
|
||
);
|
||
<materialsCitation id="31A02143FFA5190F8DF7FED39A58FEA5" ID-GBIF-Occurrence="4164459302" collectingDate="2007-12-15" collectionCode="MCZ" country="Costa Rica" elevation="930" latitude="9.3167" longLatPrecision="1" longitude="-83.7722" pageId="9" pageNumber="258" specimenCount="1" stateProvince="San Jose" typeStatus="paratype">
|
||
<collectionCode id="E7D9B3DBFFA5190F8DF7FED39CE5FEF9" box="[652,711,259,285]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="9" pageNumber="258" type="Museum">MCZ</collectionCode>
|
||
R-186143 (collected
|
||
<date id="F5760DDEFFA5190F8CD4FED29A5EFEF9" box="[943,1148,258,285]" pageId="9" pageNumber="258" value="2007-12-15">
|
||
<collectingDate id="E532F436FFA5190F8CD4FED29A5EFEF9" box="[943,1148,258,285]" pageId="9" pageNumber="258" value="2007-12-15">15 December 2007</collectingDate>
|
||
</date>
|
||
by
|
||
<collectorName id="2C3D4EC8FFA5190F8BD8FED39B2FFEF9" box="[1187,1293,259,285]" pageId="9" pageNumber="258">Steve Poe</collectorName>
|
||
and
|
||
<collectorName id="2C3D4EC8FFA5190F8A3AFED39BA8FEF9" box="[1345,1418,259,285]" pageId="9" pageNumber="258">Mason</collectorName>
|
||
J
|
||
<collectorName id="2C3D4EC8FFA5190F8FECFEF79EF2FEA5" box="[151,208,295,321]" pageId="9" pageNumber="258">Ryan</collectorName>
|
||
) from
|
||
<collectorName id="2C3D4EC8FFA5190F8E62FEF79FA6FEA5" box="[281,388,295,321]" pageId="9" pageNumber="258">Alfombra</collectorName>
|
||
, 12
|
||
<collectorName id="2C3D4EC8FFA5190F8EC9FEF79FF9FEA5" box="[434,475,295,321]" pageId="9" pageNumber="258">Km</collectorName>
|
||
S-SW of
|
||
<collectorName id="2C3D4EC8FFA5190F8D3CFEF79CC0FEA5" box="[583,738,295,321]" pageId="9" pageNumber="258">Pérez Zeledón</collectorName>
|
||
(
|
||
<geoCoordinate id="E4FC4DD9FFA5190F8D8BFEF79D45FEA5" box="[752,871,295,321]" degrees="9.31670" direction="north" orientation="latitude" pageId="9" pageNumber="258" precision="1" value="9.3167">9.31670 N</geoCoordinate>
|
||
, -
|
||
<geoCoordinate id="E4FC4DD9FFA5190F8C02FEF79A22FEA5" box="[889,1024,295,321]" degrees="83.77220" direction="west" orientation="longitude" pageId="9" pageNumber="258" precision="1" value="-83.7722">83.77220 W</geoCoordinate>
|
||
,
|
||
<quantity id="463086FBFFA5190F8B70FEF79A50FEA5" box="[1035,1138,295,321]" metricMagnitude="2" metricUnit="m" metricValue="9.3" pageId="9" pageNumber="258" unit="m" value="930.0">
|
||
<elevation id="0AE5CC2DFFA5190F8B70FEF79A50FEA5" box="[1035,1138,295,321]" metricMagnitude="2" metricUnit="m" metricValue="9.3" pageId="9" pageNumber="258" unit="m" value="930.0">930 masl</elevation>
|
||
</quantity>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="31A02143FFA5190F8BFEFEF79F0EFE6D" ID-GBIF-Occurrence="4164459319" collectingDate="2007-12-15" collectionCode="MSB" country="Costa Rica" elevation="930" latitude="9.3167" longLatPrecision="1" longitude="-83.7722" pageId="9" pageNumber="258" specimenCode="MSB 95200" specimenCount="1" stateProvince="San Jose" typeStatus="paratype">
|
||
<specimenCode id="D16E8365FFA5190F8BFEFEF79B2BFEA5" box="[1157,1289,294,321]" collectionCode="MSB" country="USA" httpUri="http://grbio.org/cool/km06-gtbn" name="Museum of Southwestern Biology" pageId="9" pageNumber="258" type="Museum">MSB 95200</specimenCode>
|
||
(collected
|
||
<date id="F5760DDEFFA5190F8AFAFEF69F60FE81" pageId="9" pageNumber="258" value="2007-12-15">
|
||
<collectingDate id="E532F436FFA5190F8AFAFEF69F60FE81" pageId="9" pageNumber="258" value="2007-12-15">15 December 2007</collectingDate>
|
||
</date>
|
||
by
|
||
<collectorName id="2C3D4EC8FFA5190F8E13FE9B9FF3FE81" box="[360,465,331,357]" pageId="9" pageNumber="258">Steve Poe</collectorName>
|
||
and
|
||
<collectorName id="2C3D4EC8FFA5190F8D7FFE9B9C6FFE81" box="[516,589,331,357]" pageId="9" pageNumber="258">Mason</collectorName>
|
||
J
|
||
<collectorName id="2C3D4EC8FFA5190F8D1FFE9B9CBFFE81" box="[612,669,331,357]" pageId="9" pageNumber="258">Ryan</collectorName>
|
||
) from
|
||
<collectorName id="2C3D4EC8FFA5190F8D98FE9B9D6CFE81" box="[739,846,331,357]" pageId="9" pageNumber="258">Alfombra</collectorName>
|
||
, 12
|
||
<collectorName id="2C3D4EC8FFA5190F8C02FE9B9D80FE81" box="[889,930,331,357]" pageId="9" pageNumber="258">Km</collectorName>
|
||
S-SW of
|
||
<collectorName id="2C3D4EC8FFA5190F8B71FE9B9A81FE81" box="[1034,1187,331,357]" pageId="9" pageNumber="258">Pérez Zeledón</collectorName>
|
||
(
|
||
<geoCoordinate id="E4FC4DD9FFA5190F8BCBFE9B9B06FE81" box="[1200,1316,331,357]" degrees="9.31670" direction="north" orientation="latitude" pageId="9" pageNumber="258" precision="1" value="9.3167">9.31670 N</geoCoordinate>
|
||
, -
|
||
<geoCoordinate id="E4FC4DD9FFA5190F8A4EFE9B9E93FE6D" degrees="83.77220" direction="west" orientation="longitude" pageId="9" pageNumber="258" precision="1" value="-83.7722">83.77220 W</geoCoordinate>
|
||
,
|
||
<quantity id="463086FBFFA5190F8FC6FEBF9F06FE6D" box="[189,292,367,393]" metricMagnitude="2" metricUnit="m" metricValue="9.3" pageId="9" pageNumber="258" unit="m" value="930.0">
|
||
<elevation id="0AE5CC2DFFA5190F8FC6FEBF9F06FE6D" box="[189,292,367,393]" metricMagnitude="2" metricUnit="m" metricValue="9.3" pageId="9" pageNumber="258" unit="m" value="930.0">930 masl</elevation>
|
||
</quantity>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="31A02143FFA5190F8E4CFEBF9FBAFE49" ID-GBIF-Occurrence="4164459314" collectingDate="2002-03-15" collectorName="Franklin Aguilar" country="Costa Rica" elevation="900" latitude="9.5013" location="Fila Piedras Blancas area" longLatPrecision="1" longitude="-83.9128" pageId="9" pageNumber="258" specimenCount="1" stateProvince="San Jose" typeStatus="paratype">
|
||
UCR 16638 (collected
|
||
<date id="F5760DDEFFA5190F8D49FEBE9CFBFE6D" box="[562,729,366,393]" pageId="9" pageNumber="258" value="2002-03-15">
|
||
<collectingDate id="E532F436FFA5190F8D49FEBE9CFBFE6D" box="[562,729,366,393]" pageId="9" pageNumber="258" value="2002-03-15">15 March 2002</collectingDate>
|
||
</date>
|
||
by
|
||
<collectorName id="2C3D4EC8FFA5190F8C78FEBF9D95FE6D" box="[771,951,367,393]" pageId="9" pageNumber="258">Franklin Aguilar</collectorName>
|
||
) from
|
||
<location id="84177DC5FFA5190F8B79FEBF9B30FE6D" LSID="urn:lsid:plazi:treatment:09619A08FFA4190D8FECF9D69A64FEA5:84177DC5FFA5190F8B79FEBF9B30FE6D" box="[1026,1298,367,393]" country="Costa Rica" latitude="9.5013" longLatPrecision="1" longitude="-83.9128" name="Fila Piedras Blancas area" pageId="9" pageNumber="258" stateProvince="San Jose">Fila Piedras Blancas area</location>
|
||
(
|
||
<geoCoordinate id="E4FC4DD9FFA5190F8A5AFEBE9BB5FE6D" box="[1313,1431,366,393]" degrees="9.50130" direction="north" orientation="latitude" pageId="9" pageNumber="258" precision="1" value="9.5013">9.50130 N</geoCoordinate>
|
||
, -
|
||
<geoCoordinate id="E4FC4DD9FFA5190F8FE5FE439F06FE49" box="[158,292,403,429]" degrees="83.91280" direction="west" orientation="longitude" pageId="9" pageNumber="258" precision="1" value="-83.9128">83.91280 W</geoCoordinate>
|
||
,
|
||
<quantity id="463086FBFFA5190F8E54FE439FB3FE49" box="[303,401,403,429]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="9" pageNumber="258" unit="m" value="900.0">
|
||
<elevation id="0AE5CC2DFFA5190F8E54FE439FB3FE49" box="[303,401,403,429]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="9" pageNumber="258" unit="m" value="900.0">900 masl</elevation>
|
||
</quantity>
|
||
)
|
||
</materialsCitation>
|
||
<materialsCitation id="31A02143FFA5190F8EE4FE439A35FE49" ID-GBIF-Occurrence="4164459316" box="[415,1047,403,429]" collectorName="Zona Protectora Los Santos" country="Costa Rica" location="Rio Savegre" pageId="9" pageNumber="258" specimenCount="1" stateProvince="San Jose" typeStatus="paratype">
|
||
in the
|
||
<collectorName id="2C3D4EC8FFA5190F8E9AFE439D2BFE49" box="[481,777,403,429]" pageId="9" pageNumber="258">Zona Protectora Los Santos</collectorName>
|
||
in the
|
||
<location id="84177DC5FFA5190F8C29FE439DF7FE49" LSID="urn:lsid:plazi:treatment:09619A08FFA4190D8FECF9D69A64FEA5:84177DC5FFA5190F8C29FE439DF7FE49" box="[850,981,403,429]" country="Costa Rica" name="Rio Savegre" pageId="9" pageNumber="258" stateProvince="San Jose">Río Savegre</location>
|
||
basin
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="D5B77B96FFA5190F8FE3FAA69D9DFA33" ID-DOI="http://doi.org/10.5281/zenodo.8182551" ID-Zenodo-Dep="8182551" httpUri="https://zenodo.org/record/8182551/files/figure.png" pageId="9" pageNumber="258" startId="9.[152,255,1398,1422]" targetBox="[151,1436,453,1375]" targetPageId="9" targetType="figure">
|
||
<paragraph id="81772B1EFFA5190F8FE3FAA69D9DFA33" blockId="9.[151,1436,1398,1495]" pageId="9" pageNumber="258">
|
||
<emphasis id="B3BCF70CFFA5190F8FE3FAA69F3BFA6B" bold="true" box="[152,281,1398,1423]" pageId="9" pageNumber="258">FIGURE 6.</emphasis>
|
||
A) Dorsal body and B) dewlap views of
|
||
<taxonomicName id="46C8509DFFA5190F8DB8FAA79D7FFA6A" authorityName="Chaves & Ryan & Bolaños & Márquez & Köhler & Poe" authorityYear="2023" box="[707,861,1399,1423]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="9" pageNumber="258" phylum="Chordata" rank="species" species="riparius">
|
||
<emphasis id="B3BCF70CFFA5190F8DB8FAA79D7FFA6A" box="[707,861,1399,1423]" italics="true" pageId="9" pageNumber="258">Anolis riparius</emphasis>
|
||
</taxonomicName>
|
||
in life. C) Dorsal view of head and D) Small middorsal scales at the base of the tail of paratype (UCR 5579) of
|
||
<taxonomicName id="46C8509DFFA5190F8DB9FA4B9D12FA56" authorityName="Chaves & Ryan & Bolaños & Márquez & Köhler & Poe" authorityYear="2023" box="[706,816,1435,1458]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="9" pageNumber="258" phylum="Chordata" rank="species" species="riparius">
|
||
<emphasis id="B3BCF70CFFA5190F8DB9FA4B9D12FA56" box="[706,816,1435,1458]" italics="true" pageId="9" pageNumber="258">A. riparius</emphasis>
|
||
</taxonomicName>
|
||
between breakpoint (a) and the first enlarge middorsal caudal scale (b). Photos A and B by David Laurencio and C and D by Gerardo Chaves.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C9D27895FFA5190F8FBCF9F99D6FF82B" pageId="9" pageNumber="258" type="diagnosis">
|
||
<paragraph id="81772B1EFFA5190F8FBCF9F99D6FF82B" blockId="9.[151,1437,1577,2035]" pageId="9" pageNumber="258">
|
||
<emphasis id="B3BCF70CFFA5190F8FBCF9F99F17F9A7" bold="true" box="[199,309,1577,1603]" pageId="9" pageNumber="258">Diagnosis</emphasis>
|
||
.
|
||
<taxonomicName id="46C8509DFFA5190F8E38F9F99FC7F9A7" authority="Chaves & Ryan & Bolaños & Márquez & Köhler & Poe, 2023" authorityName="Chaves & Ryan & Bolaños & Márquez & Köhler & Poe" authorityYear="2023" box="[323,485,1577,1603]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="9" pageNumber="258" phylum="Chordata" rank="species" species="riparius" status="sp. nov.">
|
||
<emphasis id="B3BCF70CFFA5190F8E38F9F99FC7F9A7" box="[323,485,1577,1603]" italics="true" pageId="9" pageNumber="258">Anolis riparius</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B3BCF70CFFA5190F8E96F9F99C66F9A7" bold="true" box="[493,580,1577,1603]" pageId="9" pageNumber="258">
|
||
<taxonomicNameLabel id="A88F4A77FFA5190F8E96F9F99C66F9A7" box="[493,580,1577,1603]" pageId="9" pageNumber="258" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="46C8509DFFA5190F8D02F9F99CDDF9A7" authorityName="Taylor" authorityYear="1956" box="[633,767,1577,1603]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="9" pageNumber="258" phylum="Chordata" rank="species" species="aquaticus">
|
||
<emphasis id="B3BCF70CFFA5190F8D02F9F99CDDF9A7" box="[633,767,1577,1603]" italics="true" pageId="9" pageNumber="258">A. aquaticus</emphasis>
|
||
</taxonomicName>
|
||
are the only semiaquatic Middle American anoles that have a large orange-red dewlap with yellow in males (
|
||
<figureCitation id="19F3379BFFA5190F8DEBF99D9CC7F983" box="[656,741,1613,1639]" captionStart="FIGURE 6" captionStartId="9.[152,255,1398,1422]" captionTargetBox="[151,1436,453,1375]" captionTargetId="figure-172@9.[151,1436,453,1375]" captionTargetPageId="9" captionText="FIGURE 6. A) Dorsal body and B) dewlap views of Anolis riparius in life. C) Dorsal view of head and D) Small middorsal scales at the base of the tail of paratype (UCR 5579) of A. riparius between breakpoint (a) and the first enlarge middorsal caudal scale (b). Photos A and B by David Laurencio and C and D by Gerardo Chaves." figureDoi="http://doi.org/10.5281/zenodo.8182551" httpUri="https://zenodo.org/record/8182551/files/figure.png" pageId="9" pageNumber="258">Fig. 6B</figureCitation>
|
||
).
|
||
<taxonomicName id="46C8509DFFA5190F8D8CF99D9DBAF983" authority="Chaves & Ryan & Bolaños & Márquez & Köhler & Poe, 2023" authorityName="Chaves & Ryan & Bolaños & Márquez & Köhler & Poe" authorityYear="2023" box="[759,920,1613,1639]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="9" pageNumber="258" phylum="Chordata" rank="species" species="riparius" status="sp. nov.">
|
||
<emphasis id="B3BCF70CFFA5190F8D8CF99D9DBAF983" box="[759,920,1613,1639]" italics="true" pageId="9" pageNumber="258">Anolis riparius</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B3BCF70CFFA5190F8CE4F99D9DD7F983" bold="true" box="[927,1013,1613,1639]" pageId="9" pageNumber="258">
|
||
<taxonomicNameLabel id="A88F4A77FFA5190F8CE4F99D9DD7F983" box="[927,1013,1613,1639]" pageId="9" pageNumber="258" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs from
|
||
<taxonomicName id="46C8509DFFA5190F8BF9F99D9B25F983" authorityName="Taylor" authorityYear="1956" box="[1154,1287,1613,1639]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="9" pageNumber="258" phylum="Chordata" rank="species" species="aquaticus">
|
||
<emphasis id="B3BCF70CFFA5190F8BF9F99D9B25F983" box="[1154,1287,1613,1639]" italics="true" pageId="9" pageNumber="258">A. aquaticus</emphasis>
|
||
</taxonomicName>
|
||
in possessing smaller dorsal scales of the head and body (e.g., 15–19 scales across the snout between the second canthals in
|
||
<taxonomicName id="46C8509DFFA5190F8A52F9A19BBEF96F" authority="Chaves & Ryan & Bolaños & Márquez & Köhler & Poe, 2023" authorityName="Chaves & Ryan & Bolaños & Márquez & Köhler & Poe" authorityYear="2023" box="[1321,1436,1649,1675]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="9" pageNumber="258" phylum="Chordata" rank="species" species="riparius" status="sp. nov.">
|
||
<emphasis id="B3BCF70CFFA5190F8A52F9A19BBEF96F" box="[1321,1436,1649,1675]" italics="true" pageId="9" pageNumber="258">A. riparius</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B3BCF70CFFA5190F8FECF9459ECDF94B" bold="true" box="[151,239,1685,1711]" pageId="9" pageNumber="258">
|
||
<taxonomicNameLabel id="A88F4A77FFA5190F8FECF9459ECDF94B" box="[151,239,1685,1711]" pageId="9" pageNumber="258" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
versus
|
||
<quantity id="463086FBFFA5190F8E39F9459FB7F94B" box="[322,405,1685,1711]" metricMagnitude="-1" metricUnit="m" metricValue="2.667" metricValueMax="3.556" metricValueMin="1.778" pageId="9" pageNumber="258" unit="in" value="10.5" valueMax="14.0" valueMin="7.0">7–14 in</quantity>
|
||
<taxonomicName id="46C8509DFFA5190F8EE6F9459C06F94B" authorityName="Taylor" authorityYear="1956" box="[413,548,1685,1711]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="9" pageNumber="258" phylum="Chordata" rank="species" species="aquaticus">
|
||
<emphasis id="B3BCF70CFFA5190F8EE6F9459C06F94B" box="[413,548,1685,1711]" italics="true" pageId="9" pageNumber="258">A. aquaticus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<figureCitation id="19F3379BFFA5190F8D48F9459CA5F954" box="[563,647,1685,1712]" captionStart="FIGURE 5" captionStartId="8.[152,255,1364,1388]" captionTargetBox="[260,1328,381,1341]" captionTargetId="figure-113@8.[260,1328,381,1341]" captionTargetPageId="8" captionText="FIGURE 5. A) Lateral and dewlap view of Anolis aquaticus in life. B) Dorsal view of head and C) Small middorsal scales in the base of the tail of A. aquaticus (UCR 5880) locate between breakpoint (a) and the first enlarge middorsal caudal scale (b). Photo A by Steven Poe and B and C by Gerardo Chaves." figureDoi="http://doi.org/10.5281/zenodo.8182549" httpUri="https://zenodo.org/record/8182549/files/figure.png" pageId="9" pageNumber="258">Fig. 5B</figureCitation>
|
||
), and is phylogenetically distinct (see below).
|
||
<taxonomicName id="46C8509DFFA5190F8BFAF9459B01F94B" authority="Chaves & Ryan & Bolaños & Márquez & Köhler & Poe, 2023" authorityName="Chaves & Ryan & Bolaños & Márquez & Köhler & Poe" authorityYear="2023" box="[1153,1315,1685,1711]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="9" pageNumber="258" phylum="Chordata" rank="species" species="riparius" status="sp. nov.">
|
||
<emphasis id="B3BCF70CFFA5190F8BFAF9459B01F94B" box="[1153,1315,1685,1711]" italics="true" pageId="9" pageNumber="258">Anolis riparius</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B3BCF70CFFA5190F8A50F9459BA1F94B" bold="true" box="[1323,1411,1685,1711]" pageId="9" pageNumber="258">
|
||
<taxonomicNameLabel id="A88F4A77FFA5190F8A50F9459BA1F94B" box="[1323,1411,1685,1711]" pageId="9" pageNumber="258" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is distinguished from the other Middle American semiaquatic anoles as follows: from
|
||
<taxonomicName id="46C8509DFFA5190F8B74F9699AB1F937" authority="Chaves & Ryan & Bolaños & Márquez & Köhler & Poe, 2023" authorityName="Chaves & Ryan & Bolaños & Márquez & Köhler & Poe" authorityYear="2023" box="[1039,1171,1721,1747]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="9" pageNumber="258" phylum="Chordata" rank="species" species="robinsoni" status="sp. nov.">
|
||
<emphasis id="B3BCF70CFFA5190F8B74F9699AB1F937" box="[1039,1171,1721,1747]" italics="true" pageId="9" pageNumber="258">A. robinsoni</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B3BCF70CFFA5190F8BE3F9699ACFF937" bold="true" box="[1176,1261,1721,1747]" pageId="9" pageNumber="258">
|
||
<taxonomicNameLabel id="A88F4A77FFA5190F8BE3F9699ACFF937" box="[1176,1261,1721,1747]" pageId="9" pageNumber="258" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
by male dewlap color (chocolate brown with ill-defined brick red horizontal stripes in
|
||
<taxonomicName id="46C8509DFFA5190F8CE2F90D9A3DF913" authority="Chaves & Ryan & Bolaños & Márquez & Köhler & Poe, 2023" authorityName="Chaves & Ryan & Bolaños & Márquez & Köhler & Poe" authorityYear="2023" box="[921,1055,1757,1783]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="9" pageNumber="258" phylum="Chordata" rank="species" species="robinsoni" status="sp. nov.">
|
||
<emphasis id="B3BCF70CFFA5190F8CE2F90D9A3DF913" box="[921,1055,1757,1783]" italics="true" pageId="9" pageNumber="258">A. robinsoni</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B3BCF70CFFA5190F8B53F90D9AA2F913" bold="true" box="[1064,1152,1757,1783]" pageId="9" pageNumber="258">
|
||
<taxonomicNameLabel id="A88F4A77FFA5190F8B53F90D9AA2F913" box="[1064,1152,1757,1783]" pageId="9" pageNumber="258" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
); from
|
||
<taxonomicName id="46C8509DFFA5190F8BADF90D9B6EF913" authorityName="Cope" authorityYear="1861" box="[1238,1356,1757,1783]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="9" pageNumber="258" phylum="Chordata" rank="species" species="lionotus">
|
||
<emphasis id="B3BCF70CFFA5190F8BADF90D9B6EF913" box="[1238,1356,1757,1783]" italics="true" pageId="9" pageNumber="258">A. lionotus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="46C8509DFFA5190F8AFEF90D9F29F8FF" authorityName="Cope" authorityYear="1862" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="9" pageNumber="258" phylum="Chordata" rank="species" species="poecilopus">
|
||
<emphasis id="B3BCF70CFFA5190F8AFEF90D9F29F8FF" italics="true" pageId="9" pageNumber="258">A. poecilopus</emphasis>
|
||
</taxonomicName>
|
||
by male dewlap color (solid yellow-orange in
|
||
<taxonomicName id="46C8509DFFA5190F8C6BF8D19DA4F8FF" authorityName="Cope" authorityYear="1861" box="[784,902,1793,1819]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="9" pageNumber="258" phylum="Chordata" rank="species" species="lionotus">
|
||
<emphasis id="B3BCF70CFFA5190F8C6BF8D19DA4F8FF" box="[784,902,1793,1819]" italics="true" pageId="9" pageNumber="258">A. lionotus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="46C8509DFFA5190F8CC5F8D19A70F8FF" authorityName="Cope" authorityYear="1862" box="[958,1106,1793,1819]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="9" pageNumber="258" phylum="Chordata" rank="species" species="poecilopus">
|
||
<emphasis id="B3BCF70CFFA5190F8CC5F8D19A70F8FF" box="[958,1106,1793,1819]" italics="true" pageId="9" pageNumber="258">A. poecilopus</emphasis>
|
||
</taxonomicName>
|
||
), lack of postcloacal scales in males (usually present in
|
||
<taxonomicName id="46C8509DFFA5190F8ED8F8F59C34F8DB" authorityName="Cope" authorityYear="1861" box="[419,534,1829,1855]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="9" pageNumber="258" phylum="Chordata" rank="species" species="lionotus">
|
||
<emphasis id="B3BCF70CFFA5190F8ED8F8F59C34F8DB" box="[419,534,1829,1855]" italics="true" pageId="9" pageNumber="258">A. lionotus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="46C8509DFFA5190F8D33F8F59CFAF8DB" authorityName="Cope" authorityYear="1862" box="[584,728,1829,1855]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="9" pageNumber="258" phylum="Chordata" rank="species" species="poecilopus">
|
||
<emphasis id="B3BCF70CFFA5190F8D33F8F59CFAF8DB" box="[584,728,1829,1855]" italics="true" pageId="9" pageNumber="258">A. poecilopus</emphasis>
|
||
</taxonomicName>
|
||
), and possession of zero to three enlarged rows of middorsal scales (a broadband of 10–24 enlarged middorsal scale rows in
|
||
<taxonomicName id="46C8509DFFA5190F8D81F8999DAEF887" authorityName="Cope" authorityYear="1862" box="[762,908,1865,1891]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="9" pageNumber="258" phylum="Chordata" rank="species" species="poecilopus">
|
||
<emphasis id="B3BCF70CFFA5190F8D81F8999DAEF887" box="[762,908,1865,1891]" italics="true" pageId="9" pageNumber="258">A. poecilopus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="46C8509DFFA5190F8CBAF8999A17F887" authorityName="Cope" authorityYear="1861" box="[961,1077,1865,1891]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="9" pageNumber="258" phylum="Chordata" rank="species" species="lionotus">
|
||
<emphasis id="B3BCF70CFFA5190F8CBAF8999A17F887" box="[961,1077,1865,1891]" italics="true" pageId="9" pageNumber="258">A. lionotus</emphasis>
|
||
</taxonomicName>
|
||
); from
|
||
<taxonomicName id="46C8509DFFA5190F8BFCF8999AD6F887" baseAuthorityName="Schmidt" baseAuthorityYear="1939" box="[1159,1268,1865,1891]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="9" pageNumber="258" phylum="Chordata" rank="species" species="barkeri">
|
||
<emphasis id="B3BCF70CFFA5190F8BFCF8999AD6F887" box="[1159,1268,1865,1891]" italics="true" pageId="9" pageNumber="258">A. barkeri</emphasis>
|
||
</taxonomicName>
|
||
by smaller size (maximum SVL
|
||
<quantity id="463086FBFFA5190F8E31F8BD9FB6F863" box="[330,404,1901,1927]" metricMagnitude="-2" metricUnit="m" metricValue="7.3" pageId="9" pageNumber="258" unit="mm" value="73.0">73 mm</quantity>
|
||
in
|
||
<taxonomicName id="46C8509DFFA5190F8ECFF8BD9C04F863" authority="Chaves & Ryan & Bolaños & Márquez & Köhler & Poe, 2023" authorityName="Chaves & Ryan & Bolaños & Márquez & Köhler & Poe" authorityYear="2023" box="[436,550,1901,1927]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="9" pageNumber="258" phylum="Chordata" rank="species" species="riparius" status="sp. nov.">
|
||
<emphasis id="B3BCF70CFFA5190F8ECFF8BD9C04F863" box="[436,550,1901,1927]" italics="true" pageId="9" pageNumber="258">A. riparius</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B3BCF70CFFA5190F8D50F8BD9C5DF863" bold="true" box="[555,639,1901,1927]" pageId="9" pageNumber="258">
|
||
<taxonomicNameLabel id="A88F4A77FFA5190F8D50F8BD9C5DF863" box="[555,639,1901,1927]" pageId="9" pageNumber="258" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
,
|
||
<quantity id="463086FBFFA5190F8DF1F8BD9CF6F863" box="[650,724,1901,1927]" metricMagnitude="-2" metricUnit="m" metricValue="9.1" pageId="9" pageNumber="258" unit="mm" value="91.0">91 mm</quantity>
|
||
in
|
||
<taxonomicName id="46C8509DFFA5190F8D88F8BD9D7CF863" baseAuthorityName="Schmidt" baseAuthorityYear="1939" box="[755,862,1901,1927]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="9" pageNumber="258" phylum="Chordata" rank="species" species="barkeri">
|
||
<emphasis id="B3BCF70CFFA5190F8D88F8BD9D7CF863" box="[755,862,1901,1927]" italics="true" pageId="9" pageNumber="258">A. barkeri</emphasis>
|
||
</taxonomicName>
|
||
), presence of distinctly expanded toe pads (ratio width of expanded pads / width of distal phalanx
|
||
<quantity id="463086FBFFA5190F8D1BF8419CE9F84F" box="[608,715,1937,1963]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" metricValueMax="5.588" metricValueMin="4.572" pageId="9" pageNumber="258" unit="in" value="2.0" valueMax="2.2" valueMin="1.8">1.8–2.2 in</quantity>
|
||
<taxonomicName id="46C8509DFFA5190F8DAAF8419D66F84F" authority="Chaves & Ryan & Bolaños & Márquez & Köhler & Poe, 2023" authorityName="Chaves & Ryan & Bolaños & Márquez & Köhler & Poe" authorityYear="2023" box="[721,836,1937,1963]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="9" pageNumber="258" phylum="Chordata" rank="species" species="riparius" status="sp. nov.">
|
||
<emphasis id="B3BCF70CFFA5190F8DAAF8419D66F84F" box="[721,836,1937,1963]" italics="true" pageId="9" pageNumber="258">A. riparius</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B3BCF70CFFA5190F8C31F8419D82F84F" bold="true" box="[842,928,1937,1963]" pageId="9" pageNumber="258">
|
||
<taxonomicNameLabel id="A88F4A77FFA5190F8C31F8419D82F84F" box="[842,928,1937,1963]" pageId="9" pageNumber="258" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
versus
|
||
<quantity id="463086FBFFA5190F8C8BF8419A79F84F" box="[1008,1115,1937,1963]" metricMagnitude="-2" metricUnit="m" metricValue="3.81" metricValueMax="4.064" metricValueMin="3.5559999999999996" pageId="9" pageNumber="258" unit="in" value="1.5" valueMax="1.6" valueMin="1.4">1.4–1.6 in</quantity>
|
||
<taxonomicName id="46C8509DFFA5190F8B1AF8419AECF84F" baseAuthorityName="Schmidt" baseAuthorityYear="1939" box="[1121,1230,1937,1963]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="9" pageNumber="258" phylum="Chordata" rank="species" species="barkeri">
|
||
<emphasis id="B3BCF70CFFA5190F8B1AF8419AECF84F" box="[1121,1230,1937,1963]" italics="true" pageId="9" pageNumber="258">A. barkeri</emphasis>
|
||
</taxonomicName>
|
||
), and presence of a single row of middorsal caudal scales (double row in
|
||
<taxonomicName id="46C8509DFFA5190F8DAAF8659D1FF82B" baseAuthorityName="Schmidt" baseAuthorityYear="1939" box="[721,829,1973,1999]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="9" pageNumber="258" phylum="Chordata" rank="species" species="barkeri">
|
||
<emphasis id="B3BCF70CFFA5190F8DAAF8659D1FF82B" box="[721,829,1973,1999]" italics="true" pageId="9" pageNumber="258">A. barkeri</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C9D27895FFA5190C8FBCF8099AAFFB06" lastPageId="10" lastPageNumber="259" pageId="9" pageNumber="258" type="description">
|
||
<paragraph id="81772B1EFFA5190C8FBCF8099FC9FCD1" blockId="9.[151,1437,1577,2035]" lastBlockId="10.[151,1437,150,2038]" lastPageId="10" lastPageNumber="259" pageId="9" pageNumber="258">
|
||
<emphasis id="B3BCF70CFFA5190F8FBCF8099C57F816" bold="true" box="[199,629,2008,2035]" pageId="9" pageNumber="258">
|
||
Description of the
|
||
<typeStatus id="5E7395BCFFA5190F8EE0F8099FDFF817" box="[411,509,2009,2035]" pageId="9" pageNumber="258" type="holotype">holotype</typeStatus>
|
||
(
|
||
<figureCitation id="19F3379BFFA5190F8D70F8089C4FF816" box="[523,621,2008,2034]" captionStart="FIGURE 6" captionStartId="9.[152,255,1398,1422]" captionTargetBox="[151,1436,453,1375]" captionTargetId="figure-172@9.[151,1436,453,1375]" captionTargetPageId="9" captionText="FIGURE 6. A) Dorsal body and B) dewlap views of Anolis riparius in life. C) Dorsal view of head and D) Small middorsal scales at the base of the tail of paratype (UCR 5579) of A. riparius between breakpoint (a) and the first enlarge middorsal caudal scale (b). Photos A and B by David Laurencio and C and D by Gerardo Chaves." figureDoi="http://doi.org/10.5281/zenodo.8182551" httpUri="https://zenodo.org/record/8182551/files/figure.png" pageId="9" pageNumber="258">Figure 6</figureCitation>
|
||
)
|
||
</emphasis>
|
||
. Adult male as indicated by dewlap and everted hemipenes SVL
|
||
<quantity id="463086FBFFA5190F8A4FF8099BBAF817" box="[1332,1432,2009,2035]" metricMagnitude="-2" metricUnit="m" metricValue="7.33" pageId="9" pageNumber="258" unit="mm" value="73.3">73.3 mm</quantity>
|
||
; HL
|
||
<quantity id="463086FBFFA6190C8FB9FF479F05FF55" box="[194,295,151,177]" metricMagnitude="-2" metricUnit="m" metricValue="2.01" pageId="10" pageNumber="259" unit="mm" value="20.1">20.1 mm</quantity>
|
||
; HL/SVL 0.27; HW
|
||
<quantity id="463086FBFFA6190C8D71FF479C4DFF55" box="[522,623,151,177]" metricMagnitude="-2" metricUnit="m" metricValue="1.0699999999999998" pageId="10" pageNumber="259" unit="mm" value="10.7">10.7 mm</quantity>
|
||
; EOH
|
||
<quantity id="463086FBFFA6190C8DC1FF479D3CFF55" box="[698,798,151,177]" metricMagnitude="-3" metricUnit="m" metricValue="1.64" pageId="10" pageNumber="259" unit="mm" value="1.64">1.64 mm</quantity>
|
||
; IL
|
||
<quantity id="463086FBFFA6190C8C31FF469D8CFF55" box="[842,942,150,177]" metricMagnitude="-3" metricUnit="m" metricValue="1.45" pageId="10" pageNumber="259" unit="mm" value="1.45">1.45 mm</quantity>
|
||
; IL/EOH.88; SLS-I 0.1; AGL
|
||
<quantity id="463086FBFFA6190C8A7BFF479B50FF55" box="[1280,1394,151,177]" metricMagnitude="-2" metricUnit="m" metricValue="3.126" pageId="10" pageNumber="259" unit="mm" value="31.26">31.26 mm</quantity>
|
||
; FL
|
||
<quantity id="463086FBFFA6190C8FECFF6B9F25FF31" box="[151,263,187,213]" metricMagnitude="-2" metricUnit="m" metricValue="1.834" pageId="10" pageNumber="259" unit="mm" value="18.34">18.34 mm</quantity>
|
||
; T4L
|
||
<quantity id="463086FBFFA6190C8E38FF6A9F91FF31" box="[323,435,186,213]" metricMagnitude="-2" metricUnit="m" metricValue="1.595" pageId="10" pageNumber="259" unit="mm" value="15.95">15.95 mm</quantity>
|
||
; T4W
|
||
<quantity id="463086FBFFA6190C8E83FF6A9C79FF31" box="[504,603,186,213]" metricMagnitude="-3" metricUnit="m" metricValue="1.54" pageId="10" pageNumber="259" unit="mm" value="1.54">1.54 mm</quantity>
|
||
; TL 135.00 mm, tail complete; TL/SVL 1.84; TH
|
||
<quantity id="463086FBFFA6190C8B11FF6A9AF7FF31" box="[1130,1237,186,213]" metricMagnitude="-3" metricUnit="m" metricValue="5.994" pageId="10" pageNumber="259" unit="mm" value="5.994">5.994 mm</quantity>
|
||
and TW
|
||
<quantity id="463086FBFFA6190C8A4CFF6B9BBBFF31" box="[1335,1433,187,213]" metricMagnitude="-3" metricUnit="m" metricValue="3.42" pageId="10" pageNumber="259" unit="mm" value="3.42">3.42 mm</quantity>
|
||
. 18 scales across the snout between second canthals; two scales separate the nasal opening from rostral scale; ten postrostral scales between supralabials; 12 supralabial scales from rostral scale to level of middle of eye; four scales separating supraorbital semicircles; 2 rows of scales separate subocular scales from supralabials; one slightly elongate superciliary scale; nine scales separating interparietal scale from supraorbital semicircles; four enlarged scales in supraocular disk; eight loreal scales in column just anterior to eye; 12 postmental scales posteriorly in contact with mental scale between infralabials; 15 scales in the loreal region; 13 rows of single scales in the dewlap (21–48 scales per row, rows somewhat irregular); seven enlarge middorsal scale row; 16 dorsal scales in 5% of SVL; 12 ventral scales in 5% of SVL; 220 scales around midbody; 15 expanded lamellae under fourth toe; 23 small middorsal caudal scales before the first large caudal scale (
|
||
<figureCitation id="19F3379BFFA6190C8FA1FE2D9F0CFDF3" box="[218,302,509,535]" captionStart="FIGURE 6" captionStartId="9.[152,255,1398,1422]" captionTargetBox="[151,1436,453,1375]" captionTargetId="figure-172@9.[151,1436,453,1375]" captionTargetPageId="9" captionText="FIGURE 6. A) Dorsal body and B) dewlap views of Anolis riparius in life. C) Dorsal view of head and D) Small middorsal scales at the base of the tail of paratype (UCR 5579) of A. riparius between breakpoint (a) and the first enlarge middorsal caudal scale (b). Photos A and B by David Laurencio and C and D by Gerardo Chaves." figureDoi="http://doi.org/10.5281/zenodo.8182551" httpUri="https://zenodo.org/record/8182551/files/figure.png" pageId="10" pageNumber="259">Fig. 6D</figureCitation>
|
||
). Preoccipital, enlarged infralabial, tail crest, middorsal, postcloacal scales absent; supraocular disc presents with small scales. Frontal region of head concave; scales in supraocular disc unicarinate; rostral scale with very weak cleft and overlaps mental scale; ear opening vertically ovoid; dorsal edge of ear lacks ornamentation; mental scale completely divided posteriorly and with its posterior edge convex; dewlap extends well on to chest with anterior insertion at level of anterior portion of the eye and without protruding scales at distal edge; dorsal scales strongly keeled and uniform in size; ventral scales in diagonal rows keeled and larger than dorsal scales; lateral scales homogeneous; tail is compressed and triangular in cross-section with the base taller than wide; middorsal caudal scales in single row and its size is less than three times than the adjacent scales (
|
||
<figureCitation id="19F3379BFFA6190C8D64FD279C57FCF5" box="[543,629,759,785]" captionStart="FIGURE 6" captionStartId="9.[152,255,1398,1422]" captionTargetBox="[151,1436,453,1375]" captionTargetId="figure-172@9.[151,1436,453,1375]" captionTargetPageId="9" captionText="FIGURE 6. A) Dorsal body and B) dewlap views of Anolis riparius in life. C) Dorsal view of head and D) Small middorsal scales at the base of the tail of paratype (UCR 5579) of A. riparius between breakpoint (a) and the first enlarge middorsal caudal scale (b). Photos A and B by David Laurencio and C and D by Gerardo Chaves." figureDoi="http://doi.org/10.5281/zenodo.8182551" httpUri="https://zenodo.org/record/8182551/files/figure.png" pageId="10" pageNumber="259">Fig. 6D</figureCitation>
|
||
); the longest toe of adpressed limb reaches anterior to eye and, supradigital scales keeled and multicarinate.
|
||
</paragraph>
|
||
<paragraph id="81772B1EFFA6190C8FBCFCEF9A82FC0C" blockId="10.[151,1437,150,2038]" pageId="10" pageNumber="259">
|
||
<emphasis id="B3BCF70CFFA6190C8FBCFCEF9F46FCBD" bold="true" box="[199,356,831,857]" pageId="10" pageNumber="259">Color in Life</emphasis>
|
||
(based on field notes and color photographs;
|
||
<figureCitation id="19F3379BFFA6190C8C14FCEF9A01FCBD" box="[879,1059,831,857]" captionStart="FIGURE 6" captionStartId="9.[152,255,1398,1422]" captionTargetBox="[151,1436,453,1375]" captionTargetId="figure-172@9.[151,1436,453,1375]" captionTargetPageId="9" captionText="FIGURE 6. A) Dorsal body and B) dewlap views of Anolis riparius in life. C) Dorsal view of head and D) Small middorsal scales at the base of the tail of paratype (UCR 5579) of A. riparius between breakpoint (a) and the first enlarge middorsal caudal scale (b). Photos A and B by David Laurencio and C and D by Gerardo Chaves." figureDoi="http://doi.org/10.5281/zenodo.8182551" httpUri="https://zenodo.org/record/8182551/files/figure.png" pageId="10" pageNumber="259">Fig. 6A and 6B</figureCitation>
|
||
). Dewlap yellow with orange-red horizontal streaks. Dorsal ground color olive to chestnut brown with transverse olive-green bars across dorsum down flanks extending to tail. A dirty cream-colored line extends posteriorly from the labial scales down the side of the body. Scattered small greenish spots are present on the dorsum. The dorsum and tail are marked with olive-green transverse bars. The limbs are marked with pale green to yellowish transverse bars with small punctuations.
|
||
</paragraph>
|
||
<paragraph id="81772B1EFFA6190C8FBCFC219F5DFB93" blockId="10.[151,1437,150,2038]" pageId="10" pageNumber="259">
|
||
<emphasis id="B3BCF70CFFA6190C8FBCFC219F10FBEF" bold="true" box="[199,306,1009,1035]" pageId="10" pageNumber="259">Variation</emphasis>
|
||
. Size of reproductive males (69.6 ± 5.3,
|
||
<quantity id="463086FBFFA6190C8D89FC219DB0FBE8" box="[754,914,1009,1036]" metricMagnitude="-2" metricUnit="m" metricValue="6.955" metricValueMax="7.33" metricValueMin="6.58" pageId="10" pageNumber="259" unit="mm" value="69.55" valueMax="73.3" valueMin="65.8">65.8–73.3 mm</quantity>
|
||
) was larger than females (58.0 ± 6.0,
|
||
<quantity id="463086FBFFA6190C8A4AFC219EE4FBCB" metricMagnitude="-2" metricUnit="m" metricValue="5.709999999999999" metricValueMax="6.51" metricValueMin="4.91" pageId="10" pageNumber="259" unit="mm" value="57.099999999999994" valueMax="65.1" valueMin="49.1">49.1–65.1 mm</quantity>
|
||
).
|
||
<tableCitation id="CC4A1EA5FFA6190C8FACFBC59F04FBCB" box="[215,294,1045,1071]" captionStart="TABLE 1" captionStartId="4.[151,239,512,536]" captionTargetPageId="4" captionText="TABLE 1. Selected measurements and scale characters of Anolis aquaticus, A. riparius, and A. robinsoni. Mean value" httpUri="http://table.plazi.org/id/D5B77B96FFA819028FECFDD09BBEFDFC" pageId="10" pageNumber="259" tableUuid="D5B77B96FFA819028FECFDD09BBEFDFC">Table 1</tableCitation>
|
||
shows variation in the lepidosis characters of
|
||
<taxonomicName id="46C8509DFFA6190C8C6FFBC59D97FBCB" authority="Chaves & Ryan & Bolaños & Márquez & Köhler & Poe, 2023" authorityName="Chaves & Ryan & Bolaños & Márquez & Köhler & Poe" authorityYear="2023" box="[788,949,1045,1071]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="10" pageNumber="259" phylum="Chordata" rank="species" species="riparius" status="sp. nov.">
|
||
<emphasis id="B3BCF70CFFA6190C8C6FFBC59D97FBCB" box="[788,949,1045,1071]" italics="true" pageId="10" pageNumber="259">Anolis riparius</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B3BCF70CFFA6190C8CC0FBC59A33FBCB" bold="true" box="[955,1041,1045,1071]" pageId="10" pageNumber="259">
|
||
<taxonomicNameLabel id="A88F4A77FFA6190C8CC0FBC59A33FBCB" box="[955,1041,1045,1071]" pageId="10" pageNumber="259" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
. Individuals exhibited little variation in coloration. Although some individuals were dark, the amount and distribution of the light bands and dots were similar among all specimens.
|
||
</paragraph>
|
||
<paragraph id="81772B1EFFA6190C8FBCFB509AAFFB06" blockId="10.[151,1437,150,2038]" pageId="10" pageNumber="259">
|
||
<emphasis id="B3BCF70CFFA6190C8FBCFB509FF8FB7F" bold="true" box="[199,474,1152,1179]" pageId="10" pageNumber="259">Hemipenis morphology.</emphasis>
|
||
The completely everted hemipenis of SMF 92428 (
|
||
<figureCitation id="19F3379BFFA6190C8B73FB509A78FB7F" box="[1032,1114,1152,1179]" captionStart="FIGURE 7" captionStartId="11.[152,255,1295,1319]" captionTargetBox="[356,1232,345,1266]" captionTargetId="figure-125@11.[356,1232,345,1272]" captionTargetPageId="11" captionText="FIGURE 7. Hemipenes of A. robinsoni (A), A. riparius (B), and Anolis aquaticus (C). The notable differences in appearance are due to preservation artifacts (i.e., the hemipenes of A. riparius and A. robinsoni are not completely everted). Scale bar = 1 mm for all photographs. Photos by Gunther Köhler." figureDoi="http://doi.org/10.5281/zenodo.8182555" httpUri="https://zenodo.org/record/8182555/files/figure.png" pageId="10" pageNumber="259">Fig. 7B</figureCitation>
|
||
) is a stout bilobed organ. The sulcus spermaticus is bordered by well-developed sulcal lips and opens into two broad concave areas, one on each lobe. A small asulcate ridge is present. The apex is strongly calyculate, truncus with transverse folds.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C9D27895FFA6190C8FBCFB3C9FFFFACD" pageId="10" pageNumber="259" type="etymology">
|
||
<paragraph id="81772B1EFFA6190C8FBCFB3C9FFFFACD" blockId="10.[151,1437,150,2038]" pageId="10" pageNumber="259">
|
||
<emphasis id="B3BCF70CFFA6190C8FBCFB3C9F61FAE2" bold="true" box="[199,323,1260,1286]" pageId="10" pageNumber="259">Etymology</emphasis>
|
||
. The specific epithet is an adjective taken directly from Latin (
|
||
<taxonomicName id="46C8509DFFA6190C8C84FB3C9A77FAE1" authorityName="Chaves & Ryan & Bolaños & Márquez & Köhler & Poe" authorityYear="2023" box="[1023,1109,1260,1285]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="10" pageNumber="259" phylum="Chordata" rank="species" species="riparius">
|
||
<emphasis id="B3BCF70CFFA6190C8C84FB3C9A77FAE1" box="[1023,1109,1260,1285]" italics="true" pageId="10" pageNumber="259">riparius</emphasis>
|
||
</taxonomicName>
|
||
= of stream banks) about the riparian habitat of this species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C9D27895FFA6190C8FBCFAE39F27FA71" pageId="10" pageNumber="259" type="distribution">
|
||
<paragraph id="81772B1EFFA6190C8FBCFAE39F27FA71" blockId="10.[151,1437,150,2038]" pageId="10" pageNumber="259">
|
||
<emphasis id="B3BCF70CFFA6190C8FBCFAE39F71FAA9" bold="true" box="[199,339,1331,1357]" pageId="10" pageNumber="259">Distribution</emphasis>
|
||
.
|
||
<taxonomicName id="46C8509DFFA6190C8E24FAE39FDDFAA9" authority="Chaves & Ryan & Bolaños & Márquez & Köhler & Poe, 2023" authorityName="Chaves & Ryan & Bolaños & Márquez & Köhler & Poe" authorityYear="2023" box="[351,511,1331,1357]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="10" pageNumber="259" phylum="Chordata" rank="species" species="riparius" status="sp. nov.">
|
||
<emphasis id="B3BCF70CFFA6190C8E24FAE39FDDFAA9" box="[351,511,1331,1357]" italics="true" pageId="10" pageNumber="259">Anolis riparius</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B3BCF70CFFA6190C8D7FFAE39C7BFAA9" bold="true" box="[516,601,1331,1357]" pageId="10" pageNumber="259">
|
||
<taxonomicNameLabel id="A88F4A77FFA6190C8D7FFAE39C7BFAA9" box="[516,601,1331,1357]" pageId="10" pageNumber="259" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is known from streams in the riparian gallery forests of the Pacuar, Guabo, and Savegre river basins (
|
||
<figureCitation id="19F3379BFFA6190C8EF9FA879FE3FA95" box="[386,449,1367,1393]" captionStart="FIGURE 1" captionStartId="2.[152,255,1647,1671]" captionTargetBox="[151,1436,453,1623]" captionTargetId="figure-155@2.[151,1436,453,1623]" captionTargetPageId="2" captionText="FIGURE 1. Map of Costa Rica showing collecting sites for Anolis aquaticus group. Caribbean individuals appear to be A. aquaticus, but we consider their status uncertain on geographic grounds." figureDoi="http://doi.org/10.5281/zenodo.8182541" httpUri="https://zenodo.org/record/8182541/files/figure.png" pageId="10" pageNumber="259">Fig. 1</figureCitation>
|
||
) from
|
||
<quantity id="463086FBFFA6190C8D6BFA879CF2FA95" box="[528,720,1367,1394]" metricMagnitude="2" metricUnit="m" metricValue="7.75" metricValueMax="14.5" metricValueMin="1.0" pageId="10" pageNumber="259" unit="m" value="775.0" valueMax="1450.0" valueMin="100.0">100 to 1450 masl</quantity>
|
||
. This species inhabits lowland and premontane tropical wet forest life zones.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C9D27895FFA6190D8FBCFA4E9A64FEA5" lastPageId="11" lastPageNumber="260" pageId="10" pageNumber="259" type="biology_ecology">
|
||
<paragraph id="81772B1EFFA6190C8FBCFA4E9C4DF8FA" blockId="10.[151,1437,150,2038]" pageId="10" pageNumber="259">
|
||
<emphasis id="B3BCF70CFFA6190C8FBCFA4E9F00FA5C" bold="true" box="[199,290,1438,1464]" pageId="10" pageNumber="259">Ecology</emphasis>
|
||
. Little is known about the ecology and life history of
|
||
<taxonomicName id="46C8509DFFA6190C8C02FA4E9A3EFA5C" authority="Chaves & Ryan & Bolaños & Márquez & Köhler & Poe, 2023" authorityName="Chaves & Ryan & Bolaños & Márquez & Köhler & Poe" authorityYear="2023" box="[889,1052,1438,1464]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="10" pageNumber="259" phylum="Chordata" rank="species" species="riparius" status="sp. nov.">
|
||
<emphasis id="B3BCF70CFFA6190C8C02FA4E9A3EFA5C" box="[889,1052,1438,1464]" italics="true" pageId="10" pageNumber="259">Anolis riparius</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B3BCF70CFFA6190C8B5FFA4E9A5EFA5C" bold="true" box="[1060,1148,1438,1464]" pageId="10" pageNumber="259">
|
||
<taxonomicNameLabel id="A88F4A77FFA6190C8B5FFA4E9A5EFA5C" box="[1060,1148,1438,1464]" pageId="10" pageNumber="259" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
but it seems to share lifehistory traits with
|
||
<taxonomicName id="46C8509DFFA6190C8E25FA129FC7FA38" authorityName="Taylor" authorityYear="1956" box="[350,485,1474,1500]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="10" pageNumber="259" phylum="Chordata" rank="species" species="aquaticus">
|
||
<emphasis id="B3BCF70CFFA6190C8E25FA129FC7FA38" box="[350,485,1474,1500]" italics="true" pageId="10" pageNumber="259">A. aquaticus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="46C8509DFFA6190C8D67FA129C83FA38" authority="Chaves & Ryan & Bolaños & Márquez & Köhler & Poe, 2023" authorityName="Chaves & Ryan & Bolaños & Márquez & Köhler & Poe" authorityYear="2023" box="[540,673,1474,1500]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="10" pageNumber="259" phylum="Chordata" rank="species" species="robinsoni" status="sp. nov.">
|
||
<emphasis id="B3BCF70CFFA6190C8D67FA129C83FA38" box="[540,673,1474,1500]" italics="true" pageId="10" pageNumber="259">A. robinsoni</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B3BCF70CFFA6190C8DD3FA129CDDFA38" bold="true" box="[680,767,1474,1500]" pageId="10" pageNumber="259">
|
||
<taxonomicNameLabel id="A88F4A77FFA6190C8DD3FA129CDDFA38" box="[680,767,1474,1500]" pageId="10" pageNumber="259" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
. What follows is based mainly on field observations by MJR from Alfombra and additional unnamed streams of the Rio Guabo watershed. This species may be common in small order and headwater streams but absent from some other streams within a watershed. During the day this species may be found perched on rocks and boulders along streams and can jump into the water or crawl into interstitial cracks between rocks and boulders when pursued. At night these lizards are found sleeping on moss mats, fern fronds, and other large leafy riparian vegetation, especially near the splash zone around boulders. We have also found juveniles and adults sleeping on vertical rock walls and boulders in and next to waterfalls. We have observed these lizards sleeping from 0.5 to 2.0 meters above the ground. At Alfombra on
|
||
<date id="F5760DDEFFA6190C8D76F96C9CFBF932" box="[525,729,1724,1750]" pageId="10" pageNumber="259" value="2007-12-21">21 December 2007</date>
|
||
MJR and SP found a congregation of seven gravid females on the same fern-covered boulder. One meter from the sleeping females was a cluster of 6 loosely aggregated eggs that were stuck to the moss on the same large boulder.
|
||
</paragraph>
|
||
<paragraph id="81772B1EFFA6190D8FBCF8F79A64FEA5" blockId="10.[151,1437,150,2038]" lastBlockId="11.[151,1437,150,321]" lastPageId="11" lastPageNumber="260" pageId="10" pageNumber="259">
|
||
<emphasis id="B3BCF70CFFA6190C8FBCF8F79DD3F8A6" bold="true" box="[199,1009,1831,1858]" pageId="10" pageNumber="259">
|
||
Morphological comparisons of the two new species and
|
||
<taxonomicName id="46C8509DFFA6190C8C42F8F79DD3F8A6" authorityName="Taylor" authorityYear="1956" box="[825,1009,1831,1858]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="10" pageNumber="259" phylum="Chordata" rank="species" species="aquaticus">
|
||
<emphasis id="B3BCF70CFFA6190C8C42F8F79DD3F8A6" bold="true" box="[825,1009,1831,1858]" italics="true" pageId="10" pageNumber="259">Anolis aquaticus</emphasis>
|
||
</taxonomicName>
|
||
</emphasis>
|
||
.
|
||
<taxonomicName id="46C8509DFFA6190C8C86F8F89A8DF8A5" authorityName="Taylor" authorityYear="1956" box="[1021,1199,1832,1858]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="10" pageNumber="259" phylum="Chordata" rank="species" species="aquaticus">
|
||
<emphasis id="B3BCF70CFFA6190C8C86F8F89A8DF8A5" box="[1021,1199,1832,1858]" italics="true" pageId="10" pageNumber="259">Anolis aquaticus</emphasis>
|
||
</taxonomicName>
|
||
is the smallest species in this complex. It has fewer scales for all traits except for the number of scales between the suboculars and supralabials (
|
||
<tableCitation id="CC4A1EA5FFA6190C8FE5F8A09ECCF86E" box="[158,238,1904,1930]" captionStart="TABLE 1" captionStartId="4.[151,239,512,536]" captionTargetPageId="4" captionText="TABLE 1. Selected measurements and scale characters of Anolis aquaticus, A. riparius, and A. robinsoni. Mean value" httpUri="http://table.plazi.org/id/D5B77B96FFA819028FECFDD09BBEFDFC" pageId="10" pageNumber="259" tableUuid="D5B77B96FFA819028FECFDD09BBEFDFC">Table 1</tableCitation>
|
||
<emphasis id="B3BCF70CFFA6190C8F94F8A09F94F86D" box="[239,438,1904,1930]" italics="true" pageId="10" pageNumber="259">
|
||
).
|
||
<taxonomicName id="46C8509DFFA6190C8E7FF8A09F94F86D" authorityName="Taylor" authorityYear="1956" box="[260,438,1904,1930]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="10" pageNumber="259" phylum="Chordata" rank="species" species="aquaticus">Anolis aquaticus</taxonomicName>
|
||
</emphasis>
|
||
has relatively larger dorsal scales, whereas
|
||
<taxonomicName id="46C8509DFFA6190C8CFEF8A09A2AF86E" authority="Chaves & Ryan & Bolaños & Márquez & Köhler & Poe, 2023" authorityName="Chaves & Ryan & Bolaños & Márquez & Köhler & Poe" authorityYear="2023" box="[901,1032,1904,1930]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="10" pageNumber="259" phylum="Chordata" rank="species" species="robinsoni" status="sp. nov.">
|
||
<emphasis id="B3BCF70CFFA6190C8CFEF8A09A2AF86E" box="[901,1032,1904,1930]" italics="true" pageId="10" pageNumber="259">A. robinsoni</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B3BCF70CFFA6190C8B76F8A09A41F86E" bold="true" box="[1037,1123,1904,1930]" pageId="10" pageNumber="259">
|
||
<taxonomicNameLabel id="A88F4A77FFA6190C8B76F8A09A41F86E" box="[1037,1123,1904,1930]" pageId="10" pageNumber="259" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="46C8509DFFA6190C8BEEF8A09B2AF86D" authority="Chaves & Ryan & Bolaños & Márquez & Köhler & Poe, 2023" authorityName="Chaves & Ryan & Bolaños & Márquez & Köhler & Poe" authorityYear="2023" box="[1173,1288,1904,1930]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="10" pageNumber="259" phylum="Chordata" rank="species" species="riparius" status="sp. nov.">
|
||
<emphasis id="B3BCF70CFFA6190C8BEEF8A09B2AF86D" box="[1173,1288,1904,1930]" italics="true" pageId="10" pageNumber="259">A. riparius</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B3BCF70CFFA6190C8A75F8A09B46F86E" bold="true" box="[1294,1380,1904,1930]" pageId="10" pageNumber="259">
|
||
<taxonomicNameLabel id="A88F4A77FFA6190C8A75F8A09B46F86E" box="[1294,1380,1904,1930]" pageId="10" pageNumber="259" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
have small, granular dorsal scales that result in a higher number of total of scales around the body (more than 160, whereas
|
||
<taxonomicName id="46C8509DFFA6190C8AFEF8449EDDF835" authorityName="Taylor" authorityYear="1956" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="10" pageNumber="259" phylum="Chordata" rank="species" species="aquaticus">
|
||
<emphasis id="B3BCF70CFFA6190C8AFEF8449EDDF835" italics="true" pageId="10" pageNumber="259">A. aquaticus</emphasis>
|
||
</taxonomicName>
|
||
has 160 or fewer). The number of scales between the second canthals is close to half in number in
|
||
<taxonomicName id="46C8509DFFA6190C8A6CF8689BBEF835" authorityName="Taylor" authorityYear="1956" box="[1303,1436,1976,2002]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="10" pageNumber="259" phylum="Chordata" rank="species" species="aquaticus">
|
||
<emphasis id="B3BCF70CFFA6190C8A6CF8689BBEF835" box="[1303,1436,1976,2002]" italics="true" pageId="10" pageNumber="259">A. aquaticus</emphasis>
|
||
</taxonomicName>
|
||
relative to the other species (
|
||
<tableCitation id="CC4A1EA5FFA6190C8EBFF80C9C36F812" box="[452,532,2012,2038]" captionStart="TABLE 1" captionStartId="4.[151,239,512,536]" captionTargetPageId="4" captionText="TABLE 1. Selected measurements and scale characters of Anolis aquaticus, A. riparius, and A. robinsoni. Mean value" httpUri="http://table.plazi.org/id/D5B77B96FFA819028FECFDD09BBEFDFC" pageId="10" pageNumber="259" tableUuid="D5B77B96FFA819028FECFDD09BBEFDFC">Table 1</tableCitation>
|
||
,
|
||
<figureCitation id="19F3379BFFA6190C8D5AF80C9C51F812" box="[545,627,2012,2038]" captionStart="FIGURE 4" captionStartId="6.[152,255,1472,1496]" captionTargetBox="[157,1429,489,1449]" captionTargetId="figure-181@6.[157,1429,489,1449]" captionTargetPageId="6" captionText="FIGURE 4. A) Anolis robinsoni in life; B) Male dewlap of Anolis robinsoni (SMF 92437) in life. C) Dorsal view of head and D) Small middorsal scales in the base of the tail of paratype (UCR 2464) of A. robinsoni locate between breakpoint (a) and the first enlarge middorsal caudal scale (b). Photo A by Robert Puschendorf, B by Gunther Köhler, and C and D by Gerardo Chaves." figureDoi="http://doi.org/10.5281/zenodo.8182547" httpUri="https://zenodo.org/record/8182547/files/figure.png" pageId="10" pageNumber="259">Fig. 4B</figureCitation>
|
||
,
|
||
<figureCitation id="19F3379BFFA6190C8D06F80B9CBCF812" box="[637,670,2011,2038]" captionStart="FIGURE 5" captionStartId="8.[152,255,1364,1388]" captionTargetBox="[260,1328,381,1341]" captionTargetId="figure-113@8.[260,1328,381,1341]" captionTargetPageId="8" captionText="FIGURE 5. A) Lateral and dewlap view of Anolis aquaticus in life. B) Dorsal view of head and C) Small middorsal scales in the base of the tail of A. aquaticus (UCR 5880) locate between breakpoint (a) and the first enlarge middorsal caudal scale (b). Photo A by Steven Poe and B and C by Gerardo Chaves." figureDoi="http://doi.org/10.5281/zenodo.8182549" httpUri="https://zenodo.org/record/8182549/files/figure.png" pageId="10" pageNumber="259">5B</figureCitation>
|
||
,
|
||
<figureCitation id="19F3379BFFA6190C8DD2F80C9CE8F812" box="[681,714,2012,2038]" captionStart="FIGURE 6" captionStartId="9.[152,255,1398,1422]" captionTargetBox="[151,1436,453,1375]" captionTargetId="figure-172@9.[151,1436,453,1375]" captionTargetPageId="9" captionText="FIGURE 6. A) Dorsal body and B) dewlap views of Anolis riparius in life. C) Dorsal view of head and D) Small middorsal scales at the base of the tail of paratype (UCR 5579) of A. riparius between breakpoint (a) and the first enlarge middorsal caudal scale (b). Photos A and B by David Laurencio and C and D by Gerardo Chaves." figureDoi="http://doi.org/10.5281/zenodo.8182551" httpUri="https://zenodo.org/record/8182551/files/figure.png" pageId="10" pageNumber="259">6B</figureCitation>
|
||
), indicating the larger size of the scales in
|
||
<taxonomicName id="46C8509DFFA6190C8BF2F80C9B2CF811" authorityName="Taylor" authorityYear="1956" box="[1161,1294,2012,2038]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="10" pageNumber="259" phylum="Chordata" rank="species" species="aquaticus">
|
||
<emphasis id="B3BCF70CFFA6190C8BF2F80C9B2CF811" box="[1161,1294,2012,2038]" italics="true" pageId="10" pageNumber="259">A. aquaticus</emphasis>
|
||
</taxonomicName>
|
||
. The number of small middorsal caudal scale varies between
|
||
<quantity id="463086FBFFA7190D8DEAFF479CD7FF55" box="[657,757,151,177]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" metricValueMax="2.54" metricValueMin="0.0" pageId="11" pageNumber="260" unit="in" value="5.0" valueMax="10.0" valueMin="0.0">0 to 10 in</quantity>
|
||
<taxonomicName id="46C8509DFFA7190D8D80FF479DA2FF54" authorityName="Taylor" authorityYear="1956" box="[763,896,151,177]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="11" pageNumber="260" phylum="Chordata" rank="species" species="aquaticus">
|
||
<emphasis id="B3BCF70CFFA7190D8D80FF479DA2FF54" box="[763,896,151,177]" italics="true" pageId="11" pageNumber="260">A. aquaticus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="19F3379BFFA7190D8CF5FF479DFDFF55" box="[910,991,150,177]" captionStart="FIGURE 5" captionStartId="8.[152,255,1364,1388]" captionTargetBox="[260,1328,381,1341]" captionTargetId="figure-113@8.[260,1328,381,1341]" captionTargetPageId="8" captionText="FIGURE 5. A) Lateral and dewlap view of Anolis aquaticus in life. B) Dorsal view of head and C) Small middorsal scales in the base of the tail of A. aquaticus (UCR 5880) locate between breakpoint (a) and the first enlarge middorsal caudal scale (b). Photo A by Steven Poe and B and C by Gerardo Chaves." figureDoi="http://doi.org/10.5281/zenodo.8182549" httpUri="https://zenodo.org/record/8182549/files/figure.png" pageId="11" pageNumber="260">Fig. 5C</figureCitation>
|
||
) while the other species has more than 20 small middorsal caudal scales (
|
||
<figureCitation id="19F3379BFFA7190D8E99FF6B9CAEFF31" box="[482,652,187,213]" captionStart="FIGURE 4" captionStartId="6.[152,255,1472,1496]" captionTargetBox="[157,1429,489,1449]" captionTargetId="figure-181@6.[157,1429,489,1449]" captionTargetPageId="6" captionText="FIGURE 4. A) Anolis robinsoni in life; B) Male dewlap of Anolis robinsoni (SMF 92437) in life. C) Dorsal view of head and D) Small middorsal scales in the base of the tail of paratype (UCR 2464) of A. robinsoni locate between breakpoint (a) and the first enlarge middorsal caudal scale (b). Photo A by Robert Puschendorf, B by Gunther Köhler, and C and D by Gerardo Chaves." figureDoi="http://doi.org/10.5281/zenodo.8182547" httpUri="https://zenodo.org/record/8182547/files/figure.png" pageId="11" pageNumber="260">Fig. 4D and 4D</figureCitation>
|
||
). However, the most striking feature separating one of the three species of this group is the color of the male dewlap. The red and yellow dewlap of
|
||
<taxonomicName id="46C8509DFFA7190D8CE9FF0F9A34FF1C" authorityName="Taylor" authorityYear="1956" box="[914,1046,223,249]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="11" pageNumber="260" phylum="Chordata" rank="species" species="aquaticus">
|
||
<emphasis id="B3BCF70CFFA7190D8CE9FF0F9A34FF1C" box="[914,1046,223,249]" italics="true" pageId="11" pageNumber="260">A. aquaticus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="46C8509DFFA7190D8B3CFF0F9A98FF1C" authority="Chaves & Ryan & Bolaños & Márquez & Köhler & Poe, 2023" authorityName="Chaves & Ryan & Bolaños & Márquez & Köhler & Poe" authorityYear="2023" box="[1095,1210,223,249]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="11" pageNumber="260" phylum="Chordata" rank="species" species="riparius" status="sp. nov.">
|
||
<emphasis id="B3BCF70CFFA7190D8B3CFF0F9A98FF1C" box="[1095,1210,223,249]" italics="true" pageId="11" pageNumber="260">A. riparius</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B3BCF70CFFA7190D8BC4FF0F9B31FF1D" bold="true" box="[1215,1299,223,249]" pageId="11" pageNumber="260">
|
||
<taxonomicNameLabel id="A88F4A77FFA7190D8BC4FF0F9B31FF1D" box="[1215,1299,223,249]" pageId="11" pageNumber="260" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="19F3379BFFA7190D8A5BFF0F9B52FF1D" box="[1312,1392,222,249]" captionStart="FIGURE 5" captionStartId="8.[152,255,1364,1388]" captionTargetBox="[260,1328,381,1341]" captionTargetId="figure-113@8.[260,1328,381,1341]" captionTargetPageId="8" captionText="FIGURE 5. A) Lateral and dewlap view of Anolis aquaticus in life. B) Dorsal view of head and C) Small middorsal scales in the base of the tail of A. aquaticus (UCR 5880) locate between breakpoint (a) and the first enlarge middorsal caudal scale (b). Photo A by Steven Poe and B and C by Gerardo Chaves." figureDoi="http://doi.org/10.5281/zenodo.8182549" httpUri="https://zenodo.org/record/8182549/files/figure.png" pageId="11" pageNumber="260">Fig. 5A</figureCitation>
|
||
and
|
||
<figureCitation id="19F3379BFFA7190D8FECFED39E95FEF9" box="[151,183,259,285]" captionStart="FIGURE 6" captionStartId="9.[152,255,1398,1422]" captionTargetBox="[151,1436,453,1375]" captionTargetId="figure-172@9.[151,1436,453,1375]" captionTargetPageId="9" captionText="FIGURE 6. A) Dorsal body and B) dewlap views of Anolis riparius in life. C) Dorsal view of head and D) Small middorsal scales at the base of the tail of paratype (UCR 5579) of A. riparius between breakpoint (a) and the first enlarge middorsal caudal scale (b). Photos A and B by David Laurencio and C and D by Gerardo Chaves." figureDoi="http://doi.org/10.5281/zenodo.8182551" httpUri="https://zenodo.org/record/8182551/files/figure.png" pageId="11" pageNumber="260">6B</figureCitation>
|
||
) contrasts with the dark brown dewlap of
|
||
<taxonomicName id="46C8509DFFA7190D8D0DFED39CDAFEF9" authority="Chaves & Ryan & Bolaños & Márquez & Köhler & Poe, 2023" authorityName="Chaves & Ryan & Bolaños & Márquez & Köhler & Poe" authorityYear="2023" box="[630,760,259,285]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="11" pageNumber="260" phylum="Chordata" rank="species" species="robinsoni" status="sp. nov.">
|
||
<emphasis id="B3BCF70CFFA7190D8D0DFED39CDAFEF9" box="[630,760,259,285]" italics="true" pageId="11" pageNumber="260">A. robinsoni</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B3BCF70CFFA7190D8D85FED39D71FEF9" bold="true" box="[766,851,259,285]" pageId="11" pageNumber="260">
|
||
<taxonomicNameLabel id="A88F4A77FFA7190D8D85FED39D71FEF9" box="[766,851,259,285]" pageId="11" pageNumber="260" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="19F3379BFFA7190D8C1AFED39D96FEF9" box="[865,948,259,285]" captionStart="FIGURE 4" captionStartId="6.[152,255,1472,1496]" captionTargetBox="[157,1429,489,1449]" captionTargetId="figure-181@6.[157,1429,489,1449]" captionTargetPageId="6" captionText="FIGURE 4. A) Anolis robinsoni in life; B) Male dewlap of Anolis robinsoni (SMF 92437) in life. C) Dorsal view of head and D) Small middorsal scales in the base of the tail of paratype (UCR 2464) of A. robinsoni locate between breakpoint (a) and the first enlarge middorsal caudal scale (b). Photo A by Robert Puschendorf, B by Gunther Köhler, and C and D by Gerardo Chaves." figureDoi="http://doi.org/10.5281/zenodo.8182547" httpUri="https://zenodo.org/record/8182547/files/figure.png" pageId="11" pageNumber="260">Fig. 4B</figureCitation>
|
||
). We did not find any difference in hemipenes morphology among
|
||
<emphasis id="B3BCF70CFFA7190D8E0AFEF79C54FEA4" box="[369,630,295,321]" italics="true" pageId="11" pageNumber="260">
|
||
<taxonomicName id="46C8509DFFA7190D8E0AFEF79FDAFEA4" authorityName="Taylor" authorityYear="1956" box="[369,504,295,321]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="11" pageNumber="260" phylum="Chordata" rank="species" species="aquaticus">A. aquaticus</taxonomicName>
|
||
,
|
||
<taxonomicName id="46C8509DFFA7190D8D79FEF79C54FEA4" authority="Chaves & Ryan & Bolaños & Márquez & Köhler & Poe, 2023" authorityName="Chaves & Ryan & Bolaños & Márquez & Köhler & Poe" authorityYear="2023" box="[514,630,295,321]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="11" pageNumber="260" phylum="Chordata" rank="species" species="riparius" status="sp. nov.">A. riparius</taxonomicName>
|
||
</emphasis>
|
||
<emphasis id="B3BCF70CFFA7190D8D07FEF79CF3FEA5" bold="true" box="[636,721,295,321]" pageId="11" pageNumber="260">
|
||
<taxonomicNameLabel id="A88F4A77FFA7190D8D07FEF79CF3FEA5" box="[636,721,295,321]" pageId="11" pageNumber="260" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, and
|
||
<taxonomicName id="46C8509DFFA7190D8C70FEF79DACFEA5" authority="Chaves & Ryan & Bolaños & Márquez & Köhler & Poe, 2023" authorityName="Chaves & Ryan & Bolaños & Márquez & Köhler & Poe" authorityYear="2023" box="[779,910,295,321]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="11" pageNumber="260" phylum="Chordata" rank="species" species="robinsoni" status="sp. nov.">
|
||
<emphasis id="B3BCF70CFFA7190D8C70FEF79DACFEA5" box="[779,910,295,321]" italics="true" pageId="11" pageNumber="260">A. robinsoni</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B3BCF70CFFA7190D8CEFFEF79DC8FEA5" bold="true" box="[916,1002,295,321]" pageId="11" pageNumber="260">
|
||
<taxonomicNameLabel id="A88F4A77FFA7190D8CEFFEF79DC8FEA5" box="[916,1002,295,321]" pageId="11" pageNumber="260" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="19F3379BFFA7190D8C83FEF79A1BFEA5" box="[1016,1081,295,321]" captionStart="FIGURE 7" captionStartId="11.[152,255,1295,1319]" captionTargetBox="[356,1232,345,1266]" captionTargetId="figure-125@11.[356,1232,345,1272]" captionTargetPageId="11" captionText="FIGURE 7. Hemipenes of A. robinsoni (A), A. riparius (B), and Anolis aquaticus (C). The notable differences in appearance are due to preservation artifacts (i.e., the hemipenes of A. riparius and A. robinsoni are not completely everted). Scale bar = 1 mm for all photographs. Photos by Gunther Köhler." figureDoi="http://doi.org/10.5281/zenodo.8182555" httpUri="https://zenodo.org/record/8182555/files/figure.png" pageId="11" pageNumber="260">Fig. 7</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |