633 lines
91 KiB
XML
633 lines
91 KiB
XML
<document id="BD05D2569EED30CE93A97A6120663CA9" ID-CLB-Dataset="20273" ID-DOI="10.11646/zootaxa.5115.1.9" ID-GBIF-Dataset="183d6d80-cdfb-4ec4-9500-17cb2e89f635" ID-ISSN="1175-5326" ID-Zenodo-Dep="6346178" ID-ZooBank="5FF52BDB-1928-4C4A-ACA2-47B5B869A611" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1646989739340" checkinUser="plazi" docAuthor="Starkevich, Pavel, Podėnas, Sigitas & Sivell, Duncan" docDate="2022" docId="03B38A2C5A1F265B29BAFC88FB049A5C" docLanguage="en" docName="zootaxa.5115.1.9.pdf" docOrigin="Zootaxa 5115 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tipula (Vestiplex) levisoni Starkevich & Podenas 2022, sp. nov." docType="treatment" docUuid="7D95A140-6F3B-4577-8862-F64D35DBDD8F" docUuidSource="ZooBank" docVersion="7" lastPageNumber="137" masterDocId="FF8AF2545A1D265D292DFF92FF859F25" masterDocTitle="Tipula (Vestiplex) levisoni sp. nov., a new cranefly (Diptera, Tipulidae) from the Western Himalayas" masterLastPageNumber="145" masterPageNumber="131" pageNumber="133" updateTime="1699317865228" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="4592E225C15C0B87E1E7E1FA70A954EA">Tipula (Vestiplex) levisoni sp. nov., a new cranefly (Diptera, Tipulidae) from the Western Himalayas</mods:title>
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<mods:namePart id="544AAA21491DFCF28E39CAB335399BFB">Starkevich, Pavel</mods:namePart>
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<mods:nameIdentifier id="512DBE748DE632827B34025608C2D205" type="ORCID">0000-0002-0746-0398</mods:nameIdentifier>
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<mods:affiliation id="F6E769B10FC5843E3CFF0F90AA42B0E8">Nature Research Centre, Akademijos str. 2, LT- 08412 Vilnius, Lithuania</mods:affiliation>
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<mods:nameIdentifier id="5DBCCA7D75E2AFE40F01453267527217" type="email">pavel.starkevic@gmail.com</mods:nameIdentifier>
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<mods:namePart id="6D6062B2C5E7761145BC194E6FEC0AB7">Podėnas, Sigitas</mods:namePart>
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<mods:affiliation id="0CF3B1C2CCFECAF014218BE0C4C6036D">Nature Research Centre, Akademijos str. 2, LT- 08412 Vilnius, Lithuania & Life Sciences Centre of Vilnius University, Sauletekio str. 7, LT- 10257 Vilnius, Lithuania</mods:affiliation>
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<mods:namePart id="4C8E59E73A31F199F284C0CC45EE09EE">Sivell, Duncan</mods:namePart>
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<mods:affiliation id="F46D20AAECB2D5C3E81E418D12895B13">Natural History Museum, Cromwell Road, London, SW 7 5 BD, United Kingdom</mods:affiliation>
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<mods:title id="D413A657A42668A73669643A34D38182">Zootaxa</mods:title>
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<mods:date id="1155F40FC799E4E2F45174AAA48CCD4D">2022</mods:date>
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<mods:number id="4B618DD55793C362C22BF872F8CC8E43">2022-03-11</mods:number>
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<treatment id="03B38A2C5A1F265B29BAFC88FB049A5C" ID-DOI="http://doi.org/10.5281/zenodo.6347071" ID-GBIF-Taxon="193941080" ID-Zenodo-Dep="6347071" LSID="urn:lsid:zoobank.org:act:7D95A140-6F3B-4577-8862-F64D35DBDD8F" httpUri="http://treatment.plazi.org/id/03B38A2C5A1F265B29BAFC88FB049A5C" lastPageId="6" lastPageNumber="137" pageId="2" pageNumber="133">
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<subSubSection id="C30068B15A1F265F29BAFC88FCD19C10" box="[151,852,794,821]" pageId="2" pageNumber="133" type="nomenclature">
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<paragraph id="8BA53B3A5A1F265F29BAFC88FCD19C10" blockId="2.[151,852,794,857]" box="[151,852,794,821]" pageId="2" pageNumber="133">
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<heading id="D0ED8C565A1F265F29BAFC88FCD19C10" bold="true" box="[151,852,794,821]" fontSize="11" level="1" pageId="2" pageNumber="133" reason="1">
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<emphasis id="B96EE7285A1F265F29BAFC88FCD19C10" bold="true" box="[151,852,794,821]" pageId="2" pageNumber="133">
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<taxonomicName id="4C1A40B95A1F265F29BAFC88FD6B9C10" authority="Starkevich & Podenas" authorityName="Starkevich & Podenas" authorityYear="2022" box="[151,750,794,821]" class="Insecta" family="Tipulidae" genus="Tipula" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="133" phylum="Arthropoda" rank="species" species="levisoni" status="sp. nov." subGenus="Vestiplex">
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<emphasis id="B96EE7285A1F265F29BAFC88FF639C10" bold="true" box="[151,230,794,821]" italics="true" pageId="2" pageNumber="133">Tipula</emphasis>
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(
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||
<emphasis id="B96EE7285A1F265F29DAFC88FEE69C10" bold="true" box="[247,355,794,821]" italics="true" pageId="2" pageNumber="133">Vestiplex</emphasis>
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)
|
||
<emphasis id="B96EE7285A1F265F2859FC88FE549C10" bold="true" box="[372,465,794,821]" italics="true" pageId="2" pageNumber="133">levisoni</emphasis>
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Starkevich & Podenas
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||
</taxonomicName>
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||
<taxonomicNameLabel id="A25D5A535A1F265F2BDBFC89FCD19C10" box="[758,852,795,821]" pageId="2" pageNumber="133" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C30068B15A1F265F29BAFCACFE9C9C7C" box="[151,281,830,857]" pageId="2" pageNumber="133" type="description">
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<paragraph id="8BA53B3A5A1F265F29BAFCACFE9C9C7C" blockId="2.[151,852,794,857]" box="[151,281,830,857]" pageId="2" pageNumber="133">
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(
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<figureCitation id="132127BF5A1F265F29B2FCACFE949C7C" box="[159,273,830,857]" captionStart-0="FIGURES 1–4" captionStart-1="FIGURES 5–14" captionStart-2="FIGURES 15–20" captionStartId-0="3.[151,264,1821,1845]" captionStartId-1="4.[151,264,1794,1818]" captionStartId-2="5.[151,264,1883,1907]" captionTargetBox-0="[160,1423,781,1791]" captionTargetBox-1="[255,1322,189,1764]" captionTargetBox-2="[173,1415,193,1844]" captionTargetId-0="figure-261@3.[151,1435,774,1794]" captionTargetId-1="figure-26@4.[245,1343,181,1770]" captionTargetId-2="figure-20@5.[163,1425,181,1859]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionText-0="FIGURES 1–4. Tipula (Vestiplex) levisoni sp. nov. 1, holotype male with larval exuvium, lateral view; 2, head and thorax, holotype, dorsal view; 3, female, paratype, dorsal view; 4, wing, male, paratype. Scale of 1, 3, 4 = 0.5 mm; 2 = 1 mm." captionText-1="FIGURES 5–14. Male terminalia of Tipula (Vestiplex) levisoni sp. nov. 5, hypopygium, dorsal view; 6, hypopygium, lateral view; 7, epandrium, dorsal view; 8, left gonocoxite, lateral view; 9, left outer gonostylus; 10, left inner gonostylus, lateral view; 11, aedeagal guide, lateral view; 12, gonocoxal fragment, dorsal view; 13, sperm pump, dorsal view; 14, distal part of intromittent organ, lateral view. Abbreviations: aia, anterior immovable apodeme; ca, compressor apodeme; ep, epandrium; gcx, gonocoxite; ig, inner gonostylus; ls, lateral sclerite of gonocoxal fragment; ms, medial sclerite of gonocoxal fragment; mt, mid-dorsal tooth of inner gonostylus; og, outer gonostylus; pia, posterior immovable apodeme; pt, posterolateral tooth of epandrium. Scale a: 5, 6 = 0.5 mm; b: 7–13 = 0.5 mm; c: 14 = 0.25 mm." captionText-2="FIGURES 15–20. Female terminalia of Tipula (Vestiplex) levisoni sp. nov. 15, ovipositor, left lateral view; 16, sternite 8 with hypovalvae, ventral view; 17, sternite 9, dorsal view; 18, furca, dorsal view; 19, bursa copulatrix, dorsal view; 20, spermatheca. Abbreviations: bp sd, basal part of spermathecal duct; cds, cul-de-sac of bursa copulatrix; c, cerci; h, hypovalvae; s8, sternite 8; t10, tergite 10. Scale a: 15 = 1 mm; b: 16–20 = 0.5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.6346180" figureDoi-1="http://doi.org/10.5281/zenodo.6346182" figureDoi-2="http://doi.org/10.5281/zenodo.6346184" httpUri-0="https://zenodo.org/record/6346180/files/figure.png" httpUri-1="https://zenodo.org/record/6346182/files/figure.png" httpUri-2="https://zenodo.org/record/6346184/files/figure.png" pageId="2" pageNumber="133">Figs 1–20</figureCitation>
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||
)
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C30068B15A1F265F29BAFC15FEB89B14" pageId="2" pageNumber="133" type="materials_examined">
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<paragraph id="8BA53B3A5A1F265F29BAFC15FEB89B14" blockId="2.[151,1437,902,2009]" pageId="2" pageNumber="133">
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<emphasis id="B96EE7285A1F265F29BAFC15FECD9C85" bold="true" box="[151,328,902,929]" pageId="2" pageNumber="133">Type material.</emphasis>
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||
<materialsCitation id="3B7231675A1F265F287EFC14FC6F9CE0" ID-GBIF-Occurrence="3699968305" collectingDate="1931-06-29" collectionCode="BMNH" collectorName="Fletcher & No. PS" country="India" location="Kashmir" municipality="Gulmarg" pageId="2" pageNumber="133" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
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||
<emphasis id="B96EE7285A1F265F287EFC14FE779C85" bold="true" box="[339,498,902,928]" pageId="2" pageNumber="133">
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<typeStatus id="54A185985A1F265F287EFC14FE779C85" box="[339,498,902,928]" pageId="2" pageNumber="133" type="holotype">HOLOTYPE</typeStatus>
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||
</emphasis>
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||
<specimenCount id="9D1CF0B35A1F265F28D1FC15FD969C84" box="[508,531,903,929]" pageId="2" pageNumber="133" type="male">♂</specimenCount>
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||
, labelled:
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||
<emphasis id="B96EE7285A1F265F2BA0FC14FD5A9C85" bold="true" box="[653,735,902,928]" pageId="2" pageNumber="133">
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<collectingCountry id="F30D7BAA5A1F265F2BA0FC14FD5A9C85" box="[653,735,902,928]" name="India" pageId="2" pageNumber="133">INDIA</collectingCountry>
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</emphasis>
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||
,
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||
<location id="8EC56DE15A1F265F2BDCFC14FCD69C85" LSID="urn:lsid:plazi:treatment:03B38A2C5A1F265B29BAFC88FB049A5C:8EC56DE15A1F265F2BDCFC14FCD69C85" box="[753,851,902,928]" country="India" municipality="Gulmarg" name="Kashmir" pageId="2" pageNumber="133">Kashmir</location>
|
||
,
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||
<collectingMunicipality id="6BC1A1405A1F265F2A4FFC14FC4D9C85" box="[866,968,902,928]" pageId="2" pageNumber="133">Gulmarg</collectingMunicipality>
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,
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<date id="FFA41DFA5A1F265F2AFBFC14FBD59C85" box="[982,1104,902,928]" pageId="2" pageNumber="133" value="1931-06-29">
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<collectingDate id="EFE0E4125A1F265F2AFBFC14FBD59C85" box="[982,1104,902,928]" pageId="2" pageNumber="133" value="1931-06-29">29.vi.1931</collectingDate>
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</date>
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,
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<collectorName id="26EF5EEC5A1F265F2D72FC14FB3E9C85" box="[1119,1211,902,928]" pageId="2" pageNumber="133">Fletcher</collectorName>
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coll., B.M.1932.13 [larval exuvium on same pin, male genitalia slide
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||
<collectorName id="26EF5EEC5A1F265F2BFDFC39FCA59CE0" box="[720,800,938,965]" pageId="2" pageNumber="133">No. PS</collectorName>
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0375m] (
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||
<collectionCode id="ED0BA3FF5A1F265F2AAAFC39FC649CE0" box="[903,993,939,965]" pageId="2" pageNumber="133">BMNH</collectionCode>
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||
)
|
||
</materialsCitation>
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||
.
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||
<materialsCitation id="3B7231675A1F265F2AD8FC38FCF39CCC" ID-GBIF-Occurrence="3699968303" collectingDate="1931-07-26" collectionCode="BMNH" collectorName="No. PS" country="India" location="Kashmir" municipality="Gulmarg" pageId="2" pageNumber="133" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
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<emphasis id="B96EE7285A1F265F2AD8FC38FB789CE1" bold="true" box="[1013,1277,938,965]" pageId="2" pageNumber="133">
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<typeStatus id="54A185985A1F265F2AD8FC38FB1C9CE1" box="[1013,1177,938,964]" pageId="2" pageNumber="133" type="paratype">PARATYPES</typeStatus>
|
||
:
|
||
<collectingCountry id="F30D7BAA5A1F265F2D86FC38FB789CE1" box="[1195,1277,938,964]" name="India" pageId="2" pageNumber="133">INDIA</collectingCountry>
|
||
</emphasis>
|
||
, same data as holotype,
|
||
<date id="FFA41DFA5A1F265F282BFC5CFE039CCD" box="[262,390,974,1000]" pageId="2" pageNumber="133" value="1931-07-26">
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<collectingDate id="EFE0E4125A1F265F282BFC5CFE039CCD" box="[262,390,974,1000]" pageId="2" pageNumber="133" value="1931-07-26">26.vii.1931</collectingDate>
|
||
</date>
|
||
[genitalia slide
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||
<collectorName id="26EF5EEC5A1F265F2B1AFC5DFD009CCC" box="[567,645,974,1001]" pageId="2" pageNumber="133">No. PS</collectorName>
|
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0376f] (
|
||
<specimenCount id="9D1CF0B35A1F265F2BCCFC5DFC8D9CCC" box="[737,776,975,1001]" pageId="2" pageNumber="133" type="female">1 ♀</specimenCount>
|
||
,
|
||
<collectionCode id="ED0BA3FF5A1F265F2A38FC5DFCEB9CCC" box="[789,878,975,1001]" pageId="2" pageNumber="133">BMNH</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B7231675A1F265F2AADFC5DFA1C9CCC" ID-GBIF-Occurrence="3699968310" box="[896,1433,974,1001]" collectingDate="1931-07-09" collectionCode="BMNH" collectorName="Fletcher & No. PS" country="India" location="Kashmir" municipality="Gulmarg" pageId="2" pageNumber="133" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
|
||
same data as holotype,
|
||
<date id="FFA41DFA5A1F265F2DACFC5CFB779CCD" box="[1153,1266,974,1000]" pageId="2" pageNumber="133" value="1931-07-09">
|
||
<collectingDate id="EFE0E4125A1F265F2DACFC5CFB779CCD" box="[1153,1266,974,1000]" pageId="2" pageNumber="133" value="1931-07-09">9.vii.1931</collectingDate>
|
||
</date>
|
||
(
|
||
<specimenCount id="9D1CF0B35A1F265F2C29FC5DFAAE9CCC" box="[1284,1323,975,1001]" pageId="2" pageNumber="133" type="female">1 ♀</specimenCount>
|
||
,
|
||
<collectionCode id="ED0BA3FF5A1F265F2C15FC5DFA149CCC" box="[1336,1425,975,1001]" pageId="2" pageNumber="133">BMNH</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B7231675A1F265F29BAFC60FEBF9B14" ID-GBIF-Occurrence="3699968304" collectingDate="1953-07-05" collectionCode="BMNH" collectorName="J. B. Tyson & W. Nepal Expedition" country="Nepal" elevation="3962" location="Baitadi" municipality="Tinkar Khola" pageId="2" pageNumber="133" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
|
||
<emphasis id="B96EE7285A1F265F29BAFC60FF719B29" bold="true" box="[151,244,1010,1036]" pageId="2" pageNumber="133">
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<collectingCountry id="F30D7BAA5A1F265F29BAFC60FF719B29" box="[151,244,1010,1036]" name="Nepal" pageId="2" pageNumber="133">NEPAL</collectingCountry>
|
||
</emphasis>
|
||
,
|
||
<location id="8EC56DE15A1F265F282EFC60FED29B29" LSID="urn:lsid:plazi:treatment:03B38A2C5A1F265B29BAFC88FB049A5C:8EC56DE15A1F265F282EFC60FED29B29" box="[259,343,1010,1036]" country="Nepal" municipality="Tinkar Khola" name="Baitadi" pageId="2" pageNumber="133">Baitadi</location>
|
||
,
|
||
<collectingMunicipality id="6BC1A1405A1F265F284FFC60FE7E9B29" box="[354,507,1010,1036]" pageId="2" pageNumber="133">Tinkar Khola</collectingMunicipality>
|
||
,
|
||
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|
||
<elevation id="0037DC095A1F265F2B25FC61FDF09B29" box="[520,629,1010,1037]" metricMagnitude="3" metricUnit="m" metricValue="3.9624" pageId="2" pageNumber="133" unit="ft" value="13000.0">13,000 ft.</elevation>
|
||
</quantity>
|
||
[
|
||
<quantity id="4CE296DF5A1F265F2BAEFC60FD659B28" box="[643,736,1010,1037]" metricMagnitude="3" metricUnit="m" metricValue="3.9619999999999997" pageId="2" pageNumber="133" unit="m" value="3962.0">
|
||
<elevation id="0037DC095A1F265F2BAEFC60FD659B28" box="[643,736,1010,1037]" metricMagnitude="3" metricUnit="m" metricValue="3.9619999999999997" pageId="2" pageNumber="133" unit="m" value="3962.0">3962 m</elevation>
|
||
</quantity>
|
||
],
|
||
<date id="FFA41DFA5A1F265F2BD8FC60FCEF9B28" box="[757,874,1010,1037]" pageId="2" pageNumber="133" value="1953-07-05">
|
||
<collectingDate id="EFE0E4125A1F265F2BD8FC60FCEF9B28" box="[757,874,1010,1037]" pageId="2" pageNumber="133" value="1953-07-05">5.vii.1953</collectingDate>
|
||
</date>
|
||
.
|
||
<collectorName id="26EF5EEC5A1F265F2A58FC61FC759B28" box="[885,1008,1011,1037]" pageId="2" pageNumber="133">J.B. Tyson</collectorName>
|
||
;
|
||
<collectorName id="26EF5EEC5A1F265F2AD1FC61FB6E9B29" box="[1020,1259,1010,1037]" pageId="2" pageNumber="133">W. Nepal Expedition</collectorName>
|
||
, B.M.1953-592 (
|
||
<specimenCount id="9D1CF0B35A1F265F298FFB85FF489B14" box="[162,205,1047,1073]" pageId="2" pageNumber="133" type="male">1 ♂</specimenCount>
|
||
,
|
||
<collectionCode id="ED0BA3FF5A1F265F29F4FB85FEB79B14" box="[217,306,1047,1073]" pageId="2" pageNumber="133">BMNH</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30068B15A1F265F29EAFBA8FD7F9A08" pageId="2" pageNumber="133" type="diagnosis">
|
||
<paragraph id="8BA53B3A5A1F265F29EAFBA8FD7F9A08" blockId="2.[151,1437,902,2009]" pageId="2" pageNumber="133">
|
||
<emphasis id="B96EE7285A1F265F29EAFBA8FEC79B71" bold="true" box="[199,322,1082,1108]" pageId="2" pageNumber="133">Diagnosis.</emphasis>
|
||
<taxonomicName id="4C1A40B95A1F265F2861FBA8FDA39B71" authority="Starkevich & Podėnas & Sivell, 2022" authorityName="Starkevich & Podėnas & Sivell" authorityYear="2022" box="[332,550,1082,1109]" class="Insecta" family="Tipulidae" genus="Tipula" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="133" phylum="Arthropoda" rank="species" species="levisoni" status="sp. nov." subGenus="Vestiplex">
|
||
<emphasis id="B96EE7285A1F265F2861FBA8FE119B71" box="[332,404,1082,1108]" italics="true" pageId="2" pageNumber="133">Tipula</emphasis>
|
||
(
|
||
<emphasis id="B96EE7285A1F265F288AFBA9FE399B71" box="[423,444,1083,1108]" italics="true" pageId="2" pageNumber="133">V.</emphasis>
|
||
)
|
||
<emphasis id="B96EE7285A1F265F28E2FBA8FDA39B71" box="[463,550,1082,1108]" italics="true" pageId="2" pageNumber="133">levisoni</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B96EE7285A1F265F2B1CFBA9FD0B9B70" bold="true" box="[561,654,1083,1109]" pageId="2" pageNumber="133">
|
||
<taxonomicNameLabel id="A25D5A535A1F265F2B1CFBA9FD0B9B70" box="[561,654,1083,1109]" pageId="2" pageNumber="133" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
can be recognized by its brownish-yellow body, elongated antenna which reaches the base of the abdomen and the epandrium forming a large, crescent in outline, saucer-shaped plate with deep posterior U-shaped notch bordered by denticles on either side. The gonocoxite is unarmed with a rectangular ventromesal part. Females are large with an elongated abdomen and a body length longer than
|
||
<quantity id="4CE296DF5A1F265F2DDEFB35FAC09BE4" box="[1267,1349,1191,1217]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="2" pageNumber="133" unit="mm" value="30.0">30 mm</quantity>
|
||
. Cercus slender with upturned tip and ventral margin distally with small, distinct serrations. The hypovalva comprises a pale, elongated, slender filament. The median incision between the hypovalvae is deeper than the posterior margin of sternite 8, with the lateral incision deep and narrow.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30068B15A1F265B29EAFAA4FD059E88" lastPageId="6" lastPageNumber="137" pageId="2" pageNumber="133" type="description">
|
||
<paragraph id="8BA53B3A5A1F265F29EAFAA4FE089A50" blockId="2.[151,1437,902,2009]" pageId="2" pageNumber="133">
|
||
<emphasis id="B96EE7285A1F265F29EAFAA4FE279A74" bold="true" box="[199,418,1334,1361]" pageId="2" pageNumber="133">Description. Male.</emphasis>
|
||
Body length 18.9–21.0 mm, wing length
|
||
<quantity id="4CE296DF5A1F265F2A46FAA4FB889A74" box="[875,1037,1334,1361]" metricMagnitude="-2" metricUnit="m" metricValue="2.09" metricValueMax="2.12" metricValueMin="2.06" pageId="2" pageNumber="133" unit="mm" value="20.9" valueMax="21.2" valueMin="20.6">20.6–21.2 mm</quantity>
|
||
(N=2). General body colour brownish yellow (
|
||
<figureCitation id="132127BF5A1F265F2837FAC8FE059A50" box="[282,384,1370,1397]" captionStart="FIGURES 1–4" captionStartId="3.[151,264,1821,1845]" captionTargetBox="[160,1423,781,1791]" captionTargetId="figure-261@3.[151,1435,774,1794]" captionTargetPageId="3" captionText="FIGURES 1–4. Tipula (Vestiplex) levisoni sp. nov. 1, holotype male with larval exuvium, lateral view; 2, head and thorax, holotype, dorsal view; 3, female, paratype, dorsal view; 4, wing, male, paratype. Scale of 1, 3, 4 = 0.5 mm; 2 = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6346180" httpUri="https://zenodo.org/record/6346180/files/figure.png" pageId="2" pageNumber="133">Figs 1, 2</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA53B3A5A1F265F29EAFAECFCB99955" blockId="2.[151,1437,902,2009]" pageId="2" pageNumber="133">
|
||
<emphasis id="B96EE7285A1F265F29EAFAECFE819ABD" box="[199,260,1406,1432]" italics="true" pageId="2" pageNumber="133">Head</emphasis>
|
||
. Vertex and occiput grey with narrow brown median line (
|
||
<figureCitation id="132127BF5A1F265F2A80FAECFC709ABC" box="[941,1013,1406,1433]" captionStart="FIGURES 1–4" captionStartId="3.[151,264,1821,1845]" captionTargetBox="[160,1423,781,1791]" captionTargetId="figure-261@3.[151,1435,774,1794]" captionTargetPageId="3" captionText="FIGURES 1–4. Tipula (Vestiplex) levisoni sp. nov. 1, holotype male with larval exuvium, lateral view; 2, head and thorax, holotype, dorsal view; 3, female, paratype, dorsal view; 4, wing, male, paratype. Scale of 1, 3, 4 = 0.5 mm; 2 = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6346180" httpUri="https://zenodo.org/record/6346180/files/figure.png" pageId="2" pageNumber="133">Fig. 2</figureCitation>
|
||
). Rostrum brownish yellow, dorsally thinly dusted with grey pruinescence. Nasus distinct. Antenna 13-segmented, when bent backward almost reaching base of abdomen. Scape and pedicel yellow, first flagellar segment yellow, brown at tip, remaining flagellomeres brown. Each flagellomere, except first, with basal enlargement with small incision and narrowing apex; below narrowing, with additional narrowing at apex, making each segment slightly sinuous. Apical flagellomere small. Longest verticils approximately same length as their corresponding segments. Palpus with basal segment pale, second segment pale with brown surface, last segments dark brown.
|
||
</paragraph>
|
||
<paragraph id="8BA53B3A5A1F265F29EAF9E8FDC698B5" blockId="2.[151,1437,902,2009]" pageId="2" pageNumber="133">
|
||
<emphasis id="B96EE7285A1F265F29EAF9E8FE9D99B1" box="[199,280,1658,1684]" italics="true" pageId="2" pageNumber="133">Thorax</emphasis>
|
||
. Pronotum yellowish brown with brown median line. Prescutum and presutural scutum with four greyish brown stripes narrowly bordered with brown (
|
||
<figureCitation id="132127BF5A1F265F2B90F90CFC84999C" box="[701,769,1694,1721]" captionStart="FIGURES 1–4" captionStartId="3.[151,264,1821,1845]" captionTargetBox="[160,1423,781,1791]" captionTargetId="figure-261@3.[151,1435,774,1794]" captionTargetPageId="3" captionText="FIGURES 1–4. Tipula (Vestiplex) levisoni sp. nov. 1, holotype male with larval exuvium, lateral view; 2, head and thorax, holotype, dorsal view; 3, female, paratype, dorsal view; 4, wing, male, paratype. Scale of 1, 3, 4 = 0.5 mm; 2 = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6346180" httpUri="https://zenodo.org/record/6346180/files/figure.png" pageId="2" pageNumber="133">Fig. 2</figureCitation>
|
||
). Medial pair of stripes fused in basal half, distally narrowly separated. Interspaces yellowish grey, anteriorly with small brown dots. Postsutural scutum with lobes each having greenish-brown nearly oval spot bordered with brown. Scutellum yellowish brown, mediotergite brown, thinly dusted, both sclerites with brown medial line. Pleura yellowish brown, thinly dusted with grey pruinescence. Coxae yellow, thinly dusted, trochanter yellow, femora and tibiae yellow, narrowly darkened at tip; first tarsal segment yellow, darkened at tip, remaining tarsal segments dark brown. Tarsal claw without tooth. Wing patterned with brown (
|
||
<figureCitation id="132127BF5A1F265F298DF8E4FF6098B4" box="[160,229,1910,1937]" captionStart="FIGURES 1–4" captionStartId="3.[151,264,1821,1845]" captionTargetBox="[160,1423,781,1791]" captionTargetId="figure-261@3.[151,1435,774,1794]" captionTargetPageId="3" captionText="FIGURES 1–4. Tipula (Vestiplex) levisoni sp. nov. 1, holotype male with larval exuvium, lateral view; 2, head and thorax, holotype, dorsal view; 3, female, paratype, dorsal view; 4, wing, male, paratype. Scale of 1, 3, 4 = 0.5 mm; 2 = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6346180" httpUri="https://zenodo.org/record/6346180/files/figure.png" pageId="2" pageNumber="133">Fig. 4</figureCitation>
|
||
). Halter pale with brown knob.
|
||
</paragraph>
|
||
<paragraph id="8BA53B3A5A1F265F29EAF808FA0C9891" blockId="2.[151,1437,902,2009]" box="[199,1417,1946,1973]" pageId="2" pageNumber="133">
|
||
<emphasis id="B96EE7285A1F265F29EAF808FEB49891" box="[199,305,1946,1972]" italics="true" pageId="2" pageNumber="133">Abdomen</emphasis>
|
||
. Brownish yellow, terminal segments brown. Tergites with broad brown median and lateral stripes.
|
||
</paragraph>
|
||
<paragraph id="8BA53B3A5A1F265E29EAF82DFE859D8C" blockId="2.[151,1437,902,2009]" lastBlockId="3.[151,1437,150,753]" lastPageId="3" lastPageNumber="134" pageId="2" pageNumber="133">
|
||
<emphasis id="B96EE7285A1F265F29EAF82DFED398FD" box="[199,342,1983,2008]" italics="true" pageId="2" pageNumber="133">Hypopygium</emphasis>
|
||
(
|
||
<figureCitation id="132127BF5A1F265F284BF82CFE5E98FC" box="[358,475,1982,2009]" captionStart="FIGURES 5–14" captionStartId="4.[151,264,1794,1818]" captionTargetBox="[255,1322,189,1764]" captionTargetId="figure-26@4.[245,1343,181,1770]" captionTargetPageId="4" captionText="FIGURES 5–14. Male terminalia of Tipula (Vestiplex) levisoni sp. nov. 5, hypopygium, dorsal view; 6, hypopygium, lateral view; 7, epandrium, dorsal view; 8, left gonocoxite, lateral view; 9, left outer gonostylus; 10, left inner gonostylus, lateral view; 11, aedeagal guide, lateral view; 12, gonocoxal fragment, dorsal view; 13, sperm pump, dorsal view; 14, distal part of intromittent organ, lateral view. Abbreviations: aia, anterior immovable apodeme; ca, compressor apodeme; ep, epandrium; gcx, gonocoxite; ig, inner gonostylus; ls, lateral sclerite of gonocoxal fragment; ms, medial sclerite of gonocoxal fragment; mt, mid-dorsal tooth of inner gonostylus; og, outer gonostylus; pia, posterior immovable apodeme; pt, posterolateral tooth of epandrium. Scale a: 5, 6 = 0.5 mm; b: 7–13 = 0.5 mm; c: 14 = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.6346182" httpUri="https://zenodo.org/record/6346182/files/figure.png" pageId="2" pageNumber="133">Figs 5–14</figureCitation>
|
||
). Brown, broader than abdomen. Epandrium forming large, concave, sclerotised saucer-shaped plate, crescent-shaped in outline (
|
||
<figureCitation id="132127BF5A1E265E2BB5FF05FC849F94" box="[664,769,150,177]" captionStart="FIGURES 5–14" captionStartId="4.[151,264,1794,1818]" captionTargetBox="[255,1322,189,1764]" captionTargetId="figure-26@4.[245,1343,181,1770]" captionTargetPageId="4" captionText="FIGURES 5–14. Male terminalia of Tipula (Vestiplex) levisoni sp. nov. 5, hypopygium, dorsal view; 6, hypopygium, lateral view; 7, epandrium, dorsal view; 8, left gonocoxite, lateral view; 9, left outer gonostylus; 10, left inner gonostylus, lateral view; 11, aedeagal guide, lateral view; 12, gonocoxal fragment, dorsal view; 13, sperm pump, dorsal view; 14, distal part of intromittent organ, lateral view. Abbreviations: aia, anterior immovable apodeme; ca, compressor apodeme; ep, epandrium; gcx, gonocoxite; ig, inner gonostylus; ls, lateral sclerite of gonocoxal fragment; ms, medial sclerite of gonocoxal fragment; mt, mid-dorsal tooth of inner gonostylus; og, outer gonostylus; pia, posterior immovable apodeme; pt, posterolateral tooth of epandrium. Scale a: 5, 6 = 0.5 mm; b: 7–13 = 0.5 mm; c: 14 = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.6346182" httpUri="https://zenodo.org/record/6346182/files/figure.png" pageId="3" pageNumber="134">Figs 5–7</figureCitation>
|
||
). Main body of plate brown with blackened rim. Posterior margin of plate broadly emarginated medially with deep U-shaped notch, laterally bordered by black denticles on either side; rest of margin toothed. Lateral angle of plate broadly obtuse. Anterior and lateral portions of plate raised into narrow sclerotised border, posterolaterally terminating in large, black and obtuse teeth. Gonocoxite unarmed, ventromesal part rectangular in shape (
|
||
<figureCitation id="132127BF5A1E265E2B7FFEB5FD1D9E64" box="[594,664,295,321]" captionStart="FIGURES 5–14" captionStartId="4.[151,264,1794,1818]" captionTargetBox="[255,1322,189,1764]" captionTargetId="figure-26@4.[245,1343,181,1770]" captionTargetPageId="4" captionText="FIGURES 5–14. Male terminalia of Tipula (Vestiplex) levisoni sp. nov. 5, hypopygium, dorsal view; 6, hypopygium, lateral view; 7, epandrium, dorsal view; 8, left gonocoxite, lateral view; 9, left outer gonostylus; 10, left inner gonostylus, lateral view; 11, aedeagal guide, lateral view; 12, gonocoxal fragment, dorsal view; 13, sperm pump, dorsal view; 14, distal part of intromittent organ, lateral view. Abbreviations: aia, anterior immovable apodeme; ca, compressor apodeme; ep, epandrium; gcx, gonocoxite; ig, inner gonostylus; ls, lateral sclerite of gonocoxal fragment; ms, medial sclerite of gonocoxal fragment; mt, mid-dorsal tooth of inner gonostylus; og, outer gonostylus; pia, posterior immovable apodeme; pt, posterolateral tooth of epandrium. Scale a: 5, 6 = 0.5 mm; b: 7–13 = 0.5 mm; c: 14 = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.6346182" httpUri="https://zenodo.org/record/6346182/files/figure.png" pageId="3" pageNumber="134">Fig. 8</figureCitation>
|
||
). Outer gonostylus flattened, leaf-shaped (
|
||
<figureCitation id="132127BF5A1E265E2D51FEB5FB479E64" box="[1148,1218,295,321]" captionStart="FIGURES 5–14" captionStartId="4.[151,264,1794,1818]" captionTargetBox="[255,1322,189,1764]" captionTargetId="figure-26@4.[245,1343,181,1770]" captionTargetPageId="4" captionText="FIGURES 5–14. Male terminalia of Tipula (Vestiplex) levisoni sp. nov. 5, hypopygium, dorsal view; 6, hypopygium, lateral view; 7, epandrium, dorsal view; 8, left gonocoxite, lateral view; 9, left outer gonostylus; 10, left inner gonostylus, lateral view; 11, aedeagal guide, lateral view; 12, gonocoxal fragment, dorsal view; 13, sperm pump, dorsal view; 14, distal part of intromittent organ, lateral view. Abbreviations: aia, anterior immovable apodeme; ca, compressor apodeme; ep, epandrium; gcx, gonocoxite; ig, inner gonostylus; ls, lateral sclerite of gonocoxal fragment; ms, medial sclerite of gonocoxal fragment; mt, mid-dorsal tooth of inner gonostylus; og, outer gonostylus; pia, posterior immovable apodeme; pt, posterolateral tooth of epandrium. Scale a: 5, 6 = 0.5 mm; b: 7–13 = 0.5 mm; c: 14 = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.6346182" httpUri="https://zenodo.org/record/6346182/files/figure.png" pageId="3" pageNumber="134">Fig. 9</figureCitation>
|
||
). Inner gonostylus, large, curved, claw-shaped with single black, large mid-dorsal tooth. Beak extended into blackened, narrowed rostrum with indistinctly split tip (
|
||
<figureCitation id="132127BF5A1E265E2B08FEFDFDFE9EAC" box="[549,635,367,393]" captionStart="FIGURES 5–14" captionStartId="4.[151,264,1794,1818]" captionTargetBox="[255,1322,189,1764]" captionTargetId="figure-26@4.[245,1343,181,1770]" captionTargetPageId="4" captionText="FIGURES 5–14. Male terminalia of Tipula (Vestiplex) levisoni sp. nov. 5, hypopygium, dorsal view; 6, hypopygium, lateral view; 7, epandrium, dorsal view; 8, left gonocoxite, lateral view; 9, left outer gonostylus; 10, left inner gonostylus, lateral view; 11, aedeagal guide, lateral view; 12, gonocoxal fragment, dorsal view; 13, sperm pump, dorsal view; 14, distal part of intromittent organ, lateral view. Abbreviations: aia, anterior immovable apodeme; ca, compressor apodeme; ep, epandrium; gcx, gonocoxite; ig, inner gonostylus; ls, lateral sclerite of gonocoxal fragment; ms, medial sclerite of gonocoxal fragment; mt, mid-dorsal tooth of inner gonostylus; og, outer gonostylus; pia, posterior immovable apodeme; pt, posterolateral tooth of epandrium. Scale a: 5, 6 = 0.5 mm; b: 7–13 = 0.5 mm; c: 14 = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.6346182" httpUri="https://zenodo.org/record/6346182/files/figure.png" pageId="3" pageNumber="134">Fig. 10</figureCitation>
|
||
). Aedeagal guide inshape of narrow, elongated tube, base with dorsal flange narrow and elongated, membranous at base, ventral flange flattened; distal margin bent inward forming two microscopic hooks (
|
||
<figureCitation id="132127BF5A1E265E2857FE25FE499EF4" box="[378,460,439,465]" captionStart="FIGURES 5–14" captionStartId="4.[151,264,1794,1818]" captionTargetBox="[255,1322,189,1764]" captionTargetId="figure-26@4.[245,1343,181,1770]" captionTargetPageId="4" captionText="FIGURES 5–14. Male terminalia of Tipula (Vestiplex) levisoni sp. nov. 5, hypopygium, dorsal view; 6, hypopygium, lateral view; 7, epandrium, dorsal view; 8, left gonocoxite, lateral view; 9, left outer gonostylus; 10, left inner gonostylus, lateral view; 11, aedeagal guide, lateral view; 12, gonocoxal fragment, dorsal view; 13, sperm pump, dorsal view; 14, distal part of intromittent organ, lateral view. Abbreviations: aia, anterior immovable apodeme; ca, compressor apodeme; ep, epandrium; gcx, gonocoxite; ig, inner gonostylus; ls, lateral sclerite of gonocoxal fragment; ms, medial sclerite of gonocoxal fragment; mt, mid-dorsal tooth of inner gonostylus; og, outer gonostylus; pia, posterior immovable apodeme; pt, posterolateral tooth of epandrium. Scale a: 5, 6 = 0.5 mm; b: 7–13 = 0.5 mm; c: 14 = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.6346182" httpUri="https://zenodo.org/record/6346182/files/figure.png" pageId="3" pageNumber="134">Fig. 11</figureCitation>
|
||
). Gonocoxal fragment large with lateral and medial sclerites well-developed (
|
||
<figureCitation id="132127BF5A1E265E2C16FE25FA0A9EF4" box="[1339,1423,439,465]" captionStart="FIGURES 5–14" captionStartId="4.[151,264,1794,1818]" captionTargetBox="[255,1322,189,1764]" captionTargetId="figure-26@4.[245,1343,181,1770]" captionTargetPageId="4" captionText="FIGURES 5–14. Male terminalia of Tipula (Vestiplex) levisoni sp. nov. 5, hypopygium, dorsal view; 6, hypopygium, lateral view; 7, epandrium, dorsal view; 8, left gonocoxite, lateral view; 9, left outer gonostylus; 10, left inner gonostylus, lateral view; 11, aedeagal guide, lateral view; 12, gonocoxal fragment, dorsal view; 13, sperm pump, dorsal view; 14, distal part of intromittent organ, lateral view. Abbreviations: aia, anterior immovable apodeme; ca, compressor apodeme; ep, epandrium; gcx, gonocoxite; ig, inner gonostylus; ls, lateral sclerite of gonocoxal fragment; ms, medial sclerite of gonocoxal fragment; mt, mid-dorsal tooth of inner gonostylus; og, outer gonostylus; pia, posterior immovable apodeme; pt, posterolateral tooth of epandrium. Scale a: 5, 6 = 0.5 mm; b: 7–13 = 0.5 mm; c: 14 = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.6346182" httpUri="https://zenodo.org/record/6346182/files/figure.png" pageId="3" pageNumber="134">Fig. 12</figureCitation>
|
||
). Medial sclerites fused, anterior apodeme distinct, flattened and narrowed in middle; posterior part broadly flattened, expanded at base, distal parts flattened and arched. Lateral sclerites U-shaped. Sperm pump with small and flattened central vesicle (
|
||
<figureCitation id="132127BF5A1E265E2860FDB1FE269D18" box="[333,419,547,573]" captionStart="FIGURES 5–14" captionStartId="4.[151,264,1794,1818]" captionTargetBox="[255,1322,189,1764]" captionTargetId="figure-26@4.[245,1343,181,1770]" captionTargetPageId="4" captionText="FIGURES 5–14. Male terminalia of Tipula (Vestiplex) levisoni sp. nov. 5, hypopygium, dorsal view; 6, hypopygium, lateral view; 7, epandrium, dorsal view; 8, left gonocoxite, lateral view; 9, left outer gonostylus; 10, left inner gonostylus, lateral view; 11, aedeagal guide, lateral view; 12, gonocoxal fragment, dorsal view; 13, sperm pump, dorsal view; 14, distal part of intromittent organ, lateral view. Abbreviations: aia, anterior immovable apodeme; ca, compressor apodeme; ep, epandrium; gcx, gonocoxite; ig, inner gonostylus; ls, lateral sclerite of gonocoxal fragment; ms, medial sclerite of gonocoxal fragment; mt, mid-dorsal tooth of inner gonostylus; og, outer gonostylus; pia, posterior immovable apodeme; pt, posterolateral tooth of epandrium. Scale a: 5, 6 = 0.5 mm; b: 7–13 = 0.5 mm; c: 14 = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.6346182" httpUri="https://zenodo.org/record/6346182/files/figure.png" pageId="3" pageNumber="134">Fig. 13</figureCitation>
|
||
). Compressor apodeme with broad median emargination. Posterior immovable apodeme longer than compressor apodeme, large and extended laterally into broad plate, medially with darker ridge. Anterior immovable apodeme nearly triangular, narrowed at lateral margin. Aedeagus with distal part ventrally membranous (
|
||
<figureCitation id="132127BF5A1E265E298DFD1DFF769D8C" box="[160,243,655,681]" captionStart="FIGURES 5–14" captionStartId="4.[151,264,1794,1818]" captionTargetBox="[255,1322,189,1764]" captionTargetId="figure-26@4.[245,1343,181,1770]" captionTargetPageId="4" captionText="FIGURES 5–14. Male terminalia of Tipula (Vestiplex) levisoni sp. nov. 5, hypopygium, dorsal view; 6, hypopygium, lateral view; 7, epandrium, dorsal view; 8, left gonocoxite, lateral view; 9, left outer gonostylus; 10, left inner gonostylus, lateral view; 11, aedeagal guide, lateral view; 12, gonocoxal fragment, dorsal view; 13, sperm pump, dorsal view; 14, distal part of intromittent organ, lateral view. Abbreviations: aia, anterior immovable apodeme; ca, compressor apodeme; ep, epandrium; gcx, gonocoxite; ig, inner gonostylus; ls, lateral sclerite of gonocoxal fragment; ms, medial sclerite of gonocoxal fragment; mt, mid-dorsal tooth of inner gonostylus; og, outer gonostylus; pia, posterior immovable apodeme; pt, posterolateral tooth of epandrium. Scale a: 5, 6 = 0.5 mm; b: 7–13 = 0.5 mm; c: 14 = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.6346182" httpUri="https://zenodo.org/record/6346182/files/figure.png" pageId="3" pageNumber="134">Fig. 14</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA53B3A5A1E265E29EAFD21FE4A9DD4" blockId="3.[151,1437,150,753]" pageId="3" pageNumber="134">
|
||
<emphasis id="B96EE7285A1E265E29EAFD21FEA09DE8" bold="true" box="[199,293,691,717]" pageId="3" pageNumber="134">Female.</emphasis>
|
||
Body length
|
||
<quantity id="4CE296DF5A1E265E28EDFD21FDE39DE8" box="[448,614,691,717]" metricMagnitude="-2" metricUnit="m" metricValue="3.6" metricValueMax="3.82" metricValueMin="3.38" pageId="3" pageNumber="134" unit="mm" value="36.0" valueMax="38.2" valueMin="33.8">33.8–38.2 mm</quantity>
|
||
, wing length
|
||
<quantity id="4CE296DF5A1E265E2A2CFD20FC209DE8" box="[769,933,690,717]" metricMagnitude="-2" metricUnit="m" metricValue="2.665" metricValueMax="2.7399999999999998" metricValueMin="2.59" pageId="3" pageNumber="134" unit="mm" value="26.65" valueMax="27.4" valueMin="25.9">25.9–27.4 mm</quantity>
|
||
(N=2). Generally similar to male, but abdomen more elongate (
|
||
<figureCitation id="132127BF5A1E265E2850FD45FE449DD4" box="[381,449,727,753]" captionStart="FIGURES 1–4" captionStartId="3.[151,264,1821,1845]" captionTargetBox="[160,1423,781,1791]" captionTargetId="figure-261@3.[151,1435,774,1794]" captionTargetPageId="3" captionText="FIGURES 1–4. Tipula (Vestiplex) levisoni sp. nov. 1, holotype male with larval exuvium, lateral view; 2, head and thorax, holotype, dorsal view; 3, female, paratype, dorsal view; 4, wing, male, paratype. Scale of 1, 3, 4 = 0.5 mm; 2 = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6346180" httpUri="https://zenodo.org/record/6346180/files/figure.png" pageId="3" pageNumber="134">Fig. 3</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF656BB25A1E265E29BAF88FFAC7987C" ID-DOI="http://doi.org/10.5281/zenodo.6346180" ID-Zenodo-Dep="6346180" httpUri="https://zenodo.org/record/6346180/files/figure.png" pageId="3" pageNumber="134" startId="3.[151,264,1821,1845]" targetBox="[160,1423,781,1791]" targetPageId="3">
|
||
<paragraph id="8BA53B3A5A1E265E29BAF88FFAC7987C" blockId="3.[151,1436,1821,1882]" pageId="3" pageNumber="134">
|
||
<emphasis id="B96EE7285A1E265E29BAF88FFEB89810" bold="true" box="[151,317,1821,1845]" pageId="3" pageNumber="134">FIGURES 1–4.</emphasis>
|
||
<taxonomicName id="4C1A40B95A1E265E2868F88FFDD49810" authority="Starkevich & Podėnas & Sivell, 2022" authorityName="Starkevich & Podėnas & Sivell" authorityYear="2022" box="[325,593,1821,1845]" class="Insecta" family="Tipulidae" genus="Tipula" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="134" phylum="Arthropoda" rank="species" species="levisoni" status="sp. nov." subGenus="Vestiplex">
|
||
<emphasis id="B96EE7285A1E265E2868F88FFE039810" box="[325,390,1821,1845]" italics="true" pageId="3" pageNumber="134">Tipula</emphasis>
|
||
(
|
||
<emphasis id="B96EE7285A1E265E28BAF88FFE749810" box="[407,497,1821,1845]" italics="true" pageId="3" pageNumber="134">Vestiplex</emphasis>
|
||
)
|
||
<emphasis id="B96EE7285A1E265E2B2FF88FFDD49810" box="[514,593,1821,1845]" italics="true" pageId="3" pageNumber="134">levisoni</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B96EE7285A1E265E2B74F88FFD4D9810" bold="true" box="[601,712,1821,1845]" pageId="3" pageNumber="134">
|
||
<taxonomicNameLabel id="A25D5A535A1E265E2B74F88FFD299810" box="[601,684,1821,1845]" pageId="3" pageNumber="134" rank="species">sp. nov.</taxonomicNameLabel>
|
||
1,
|
||
</emphasis>
|
||
holotype male with larval exuvium, lateral view;
|
||
<emphasis id="B96EE7285A1E265E2DF9F88FFB629810" bold="true" box="[1236,1255,1821,1845]" pageId="3" pageNumber="134">2,</emphasis>
|
||
head and thorax, holotype, dorsal view;
|
||
<emphasis id="B96EE7285A1E265E2852F8D3FE17987C" bold="true" box="[383,402,1857,1881]" pageId="3" pageNumber="134">3,</emphasis>
|
||
female, paratype, dorsal view;
|
||
<emphasis id="B96EE7285A1E265E2BFCF8D3FD61987C" bold="true" box="[721,740,1857,1881]" pageId="3" pageNumber="134">4,</emphasis>
|
||
wing, male, paratype. Scale of 1, 3, 4 = 0.5 mm; 2 = 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF656BB25A19265929BAF890FD3998D7" ID-DOI="http://doi.org/10.5281/zenodo.6346182" ID-Zenodo-Dep="6346182" httpUri="https://zenodo.org/record/6346182/files/figure.png" pageId="4" pageNumber="135" startId="4.[151,264,1794,1818]" targetBox="[255,1322,189,1764]" targetPageId="4">
|
||
<paragraph id="8BA53B3A5A19265929BAF890FD3998D7" blockId="4.[151,1436,1794,2035]" pageId="4" pageNumber="135">
|
||
<emphasis id="B96EE7285A19265929BAF890FECC983F" bold="true" box="[151,329,1794,1818]" pageId="4" pageNumber="135">FIGURES 5–14.</emphasis>
|
||
Male terminalia of
|
||
<taxonomicName id="4C1A40B95A1926592B35F890FCA6983F" authority="Starkevich & Podėnas & Sivell, 2022" authorityName="Starkevich & Podėnas & Sivell" authorityYear="2022" box="[536,803,1794,1819]" class="Insecta" family="Tipulidae" genus="Tipula" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="135" phylum="Arthropoda" rank="species" species="levisoni" status="sp. nov." subGenus="Vestiplex">
|
||
<emphasis id="B96EE7285A1926592B35F890FDDC983F" box="[536,601,1794,1818]" italics="true" pageId="4" pageNumber="135">Tipula</emphasis>
|
||
(
|
||
<emphasis id="B96EE7285A1926592B44F890FD46983F" box="[617,707,1794,1818]" italics="true" pageId="4" pageNumber="135">Vestiplex</emphasis>
|
||
)
|
||
<emphasis id="B96EE7285A1926592BF9F890FCA6983F" box="[724,803,1794,1818]" italics="true" pageId="4" pageNumber="135">levisoni</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B96EE7285A1926592A07F891FC1D983E" bold="true" box="[810,920,1795,1819]" pageId="4" pageNumber="135">
|
||
<taxonomicNameLabel id="A25D5A535A1926592A07F891FCF9983E" box="[810,892,1795,1819]" pageId="4" pageNumber="135" rank="species">sp. nov.</taxonomicNameLabel>
|
||
5,
|
||
</emphasis>
|
||
hypopygium, dorsal view;
|
||
<emphasis id="B96EE7285A1926592D99F890FB42983F" bold="true" box="[1204,1223,1794,1818]" pageId="4" pageNumber="135">6,</emphasis>
|
||
hypopygium, lateral view;
|
||
<emphasis id="B96EE7285A19265929F8F8B5FF6D981A" bold="true" box="[213,232,1831,1855]" pageId="4" pageNumber="135">7,</emphasis>
|
||
epandrium, dorsal view;
|
||
<emphasis id="B96EE7285A19265928C5F8B4FE79981B" bold="true" box="[488,508,1830,1854]" pageId="4" pageNumber="135">8,</emphasis>
|
||
left gonocoxite, lateral view;
|
||
<emphasis id="B96EE7285A1926592A04F8B4FCB9981B" bold="true" box="[809,828,1830,1854]" pageId="4" pageNumber="135">9,</emphasis>
|
||
left outer gonostylus;
|
||
<emphasis id="B96EE7285A1926592D33F8B4FBB8981B" bold="true" box="[1054,1085,1830,1854]" pageId="4" pageNumber="135">10,</emphasis>
|
||
left inner gonostylus, lateral view;
|
||
<emphasis id="B96EE7285A19265929BAF8D8FF309847" bold="true" box="[151,181,1866,1890]" pageId="4" pageNumber="135">11,</emphasis>
|
||
aedeagal guide, lateral view;
|
||
<emphasis id="B96EE7285A19265928CEF8D8FD869847" bold="true" box="[483,515,1866,1890]" pageId="4" pageNumber="135">12,</emphasis>
|
||
gonocoxal fragment, dorsal view;
|
||
<emphasis id="B96EE7285A1926592A4EF8D8FC069847" bold="true" box="[867,899,1866,1890]" pageId="4" pageNumber="135">13,</emphasis>
|
||
sperm pump, dorsal view;
|
||
<emphasis id="B96EE7285A1926592DBBF8D8FB339847" bold="true" box="[1174,1206,1866,1890]" pageId="4" pageNumber="135">14,</emphasis>
|
||
distal part of intromittent organ, lateral view. Abbreviations: aia, anterior immovable apodeme; ca, compressor apodeme; ep, epandrium; gcx, gonocoxite; ig, inner gonostylus; ls, lateral sclerite of gonocoxal fragment; ms, medial sclerite of gonocoxal fragment; mt, mid-dorsal tooth of inner gonostylus; og, outer gonostylus; pia, posterior immovable apodeme; pt, posterolateral tooth of epandrium. Scale a: 5, 6 = 0.5 mm; b: 7–13 = 0.5 mm; c: 14 = 0.25 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF656BB25A18265829BAF8C9FD4998FA" ID-DOI="http://doi.org/10.5281/zenodo.6346184" ID-Zenodo-Dep="6346184" httpUri="https://zenodo.org/record/6346184/files/figure.png" pageId="5" pageNumber="136" startId="5.[151,264,1883,1907]" targetBox="[173,1415,193,1844]" targetPageId="5">
|
||
<paragraph id="8BA53B3A5A18265829BAF8C9FD4998FA" blockId="5.[151,1437,1883,2015]" pageId="5" pageNumber="136">
|
||
<emphasis id="B96EE7285A18265829BAF8C9FED09856" bold="true" box="[151,341,1883,1907]" pageId="5" pageNumber="136">FIGURES 15–20.</emphasis>
|
||
Female terminalia of
|
||
<taxonomicName id="4C1A40B95A1826582B1BF8C9FCBB9856" authority="Starkevich & Podėnas & Sivell, 2022" authorityName="Starkevich & Podėnas & Sivell" authorityYear="2022" box="[566,830,1883,1907]" class="Insecta" family="Tipulidae" genus="Tipula" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="136" phylum="Arthropoda" rank="species" species="levisoni" status="sp. nov." subGenus="Vestiplex">
|
||
<emphasis id="B96EE7285A1826582B1BF8C9FDF29856" box="[566,631,1883,1907]" italics="true" pageId="5" pageNumber="136">Tipula</emphasis>
|
||
(
|
||
<emphasis id="B96EE7285A1826582BABF8C9FD659856" box="[646,736,1883,1907]" italics="true" pageId="5" pageNumber="136">Vestiplex</emphasis>
|
||
)
|
||
<emphasis id="B96EE7285A1826582BC2F8C9FCBB9856" box="[751,830,1883,1907]" italics="true" pageId="5" pageNumber="136">levisoni</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B96EE7285A1826582A68F8C9FC399856" bold="true" box="[837,956,1883,1907]" pageId="5" pageNumber="136">
|
||
<taxonomicNameLabel id="A25D5A535A1826582A68F8C9FC139856" box="[837,918,1883,1907]" pageId="5" pageNumber="136" rank="species">sp. nov.</taxonomicNameLabel>
|
||
15,
|
||
</emphasis>
|
||
ovipositor, left lateral view;
|
||
<emphasis id="B96EE7285A1826582DC9F8C9FA869856" bold="true" box="[1252,1283,1883,1907]" pageId="5" pageNumber="136">16,</emphasis>
|
||
sternite 8 with hypovalvae, ventral view;
|
||
<emphasis id="B96EE7285A182658288FF8EDFE4798B2" bold="true" box="[418,450,1919,1943]" pageId="5" pageNumber="136">17,</emphasis>
|
||
sternite 9, dorsal view;
|
||
<emphasis id="B96EE7285A1826582B9EF8EDFD5798B2" bold="true" box="[691,722,1919,1943]" pageId="5" pageNumber="136">18,</emphasis>
|
||
furca, dorsal view;
|
||
<emphasis id="B96EE7285A1826582AB6F8EDFC3F98B2" bold="true" box="[923,954,1919,1943]" pageId="5" pageNumber="136">19,</emphasis>
|
||
bursa copulatrix, dorsal view;
|
||
<emphasis id="B96EE7285A1826582DDFF8EDFA9498B2" bold="true" box="[1266,1297,1919,1943]" pageId="5" pageNumber="136">20,</emphasis>
|
||
spermatheca. Abbreviations: bp sd, basal part of spermathecal duct; cds, cul-de-sac of bursa copulatrix; c, cerci; h, hypovalvae; s8, sternite 8; t10, tergite 10. Scale a: 15 = 1 mm; b: 16–20 = 0.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BA53B3A5A1B265B29EAFF05FD059E88" blockId="6.[151,1437,150,1401]" pageId="6" pageNumber="137">
|
||
<emphasis id="B96EE7285A1B265B29EAFF05FE1D9F94" box="[199,408,151,177]" italics="true" pageId="6" pageNumber="137">Female terminalia</emphasis>
|
||
(
|
||
<figureCitation id="132127BF5A1B265B288AFF05FDAF9F94" box="[423,554,150,177]" captionStart="FIGURES 15–20" captionStartId="5.[151,264,1883,1907]" captionTargetBox="[173,1415,193,1844]" captionTargetId="figure-20@5.[163,1425,181,1859]" captionTargetPageId="5" captionText="FIGURES 15–20. Female terminalia of Tipula (Vestiplex) levisoni sp. nov. 15, ovipositor, left lateral view; 16, sternite 8 with hypovalvae, ventral view; 17, sternite 9, dorsal view; 18, furca, dorsal view; 19, bursa copulatrix, dorsal view; 20, spermatheca. Abbreviations: bp sd, basal part of spermathecal duct; cds, cul-de-sac of bursa copulatrix; c, cerci; h, hypovalvae; s8, sternite 8; t10, tergite 10. Scale a: 15 = 1 mm; b: 16–20 = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6346184" httpUri="https://zenodo.org/record/6346184/files/figure.png" pageId="6" pageNumber="137">Figs 15–20</figureCitation>
|
||
). Tergite 10 shiny brown. Cercus brownish yellow, about as long as tergite 10, slender with up-turned tip (
|
||
<figureCitation id="132127BF5A1B265B28E8FF29FD9D9FF0" box="[453,536,186,213]" captionStart="FIGURES 15–20" captionStartId="5.[151,264,1883,1907]" captionTargetBox="[173,1415,193,1844]" captionTargetId="figure-20@5.[163,1425,181,1859]" captionTargetPageId="5" captionText="FIGURES 15–20. Female terminalia of Tipula (Vestiplex) levisoni sp. nov. 15, ovipositor, left lateral view; 16, sternite 8 with hypovalvae, ventral view; 17, sternite 9, dorsal view; 18, furca, dorsal view; 19, bursa copulatrix, dorsal view; 20, spermatheca. Abbreviations: bp sd, basal part of spermathecal duct; cds, cul-de-sac of bursa copulatrix; c, cerci; h, hypovalvae; s8, sternite 8; t10, tergite 10. Scale a: 15 = 1 mm; b: 16–20 = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6346184" httpUri="https://zenodo.org/record/6346184/files/figure.png" pageId="6" pageNumber="137">Fig. 15</figureCitation>
|
||
). Ventral margin distally with small, distinct serrations. Hypovalva pale, elongated, filamentous, with short trichia at tip (
|
||
<figureCitation id="132127BF5A1B265B2B7BFF4DFD2D9FDC" box="[598,680,222,249]" captionStart="FIGURES 15–20" captionStartId="5.[151,264,1883,1907]" captionTargetBox="[173,1415,193,1844]" captionTargetId="figure-20@5.[163,1425,181,1859]" captionTargetPageId="5" captionText="FIGURES 15–20. Female terminalia of Tipula (Vestiplex) levisoni sp. nov. 15, ovipositor, left lateral view; 16, sternite 8 with hypovalvae, ventral view; 17, sternite 9, dorsal view; 18, furca, dorsal view; 19, bursa copulatrix, dorsal view; 20, spermatheca. Abbreviations: bp sd, basal part of spermathecal duct; cds, cul-de-sac of bursa copulatrix; c, cerci; h, hypovalvae; s8, sternite 8; t10, tergite 10. Scale a: 15 = 1 mm; b: 16–20 = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6346184" httpUri="https://zenodo.org/record/6346184/files/figure.png" pageId="6" pageNumber="137">Fig. 15</figureCitation>
|
||
). Median incision between hypovalvae deeper than posterior margin of sternite 8. Lateral incision deep and narrow, about three times as deep as maximum width. Posterolateral angle of sternite 8 rounded. Sternite 9 with posterior part shovel-shaped, margin with lateral incisions, medially forming groove; anterior parts flattened (
|
||
<figureCitation id="132127BF5A1B265B28D1FED9FDCA9E40" box="[508,591,331,357]" captionStart="FIGURES 15–20" captionStartId="5.[151,264,1883,1907]" captionTargetBox="[173,1415,193,1844]" captionTargetId="figure-20@5.[163,1425,181,1859]" captionTargetPageId="5" captionText="FIGURES 15–20. Female terminalia of Tipula (Vestiplex) levisoni sp. nov. 15, ovipositor, left lateral view; 16, sternite 8 with hypovalvae, ventral view; 17, sternite 9, dorsal view; 18, furca, dorsal view; 19, bursa copulatrix, dorsal view; 20, spermatheca. Abbreviations: bp sd, basal part of spermathecal duct; cds, cul-de-sac of bursa copulatrix; c, cerci; h, hypovalvae; s8, sternite 8; t10, tergite 10. Scale a: 15 = 1 mm; b: 16–20 = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6346184" httpUri="https://zenodo.org/record/6346184/files/figure.png" pageId="6" pageNumber="137">Fig. 17</figureCitation>
|
||
). Furca flattened, base narrow (
|
||
<figureCitation id="132127BF5A1B265B2A81FED9FC7A9E40" box="[940,1023,331,357]" captionStart="FIGURES 15–20" captionStartId="5.[151,264,1883,1907]" captionTargetBox="[173,1415,193,1844]" captionTargetId="figure-20@5.[163,1425,181,1859]" captionTargetPageId="5" captionText="FIGURES 15–20. Female terminalia of Tipula (Vestiplex) levisoni sp. nov. 15, ovipositor, left lateral view; 16, sternite 8 with hypovalvae, ventral view; 17, sternite 9, dorsal view; 18, furca, dorsal view; 19, bursa copulatrix, dorsal view; 20, spermatheca. Abbreviations: bp sd, basal part of spermathecal duct; cds, cul-de-sac of bursa copulatrix; c, cerci; h, hypovalvae; s8, sternite 8; t10, tergite 10. Scale a: 15 = 1 mm; b: 16–20 = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6346184" httpUri="https://zenodo.org/record/6346184/files/figure.png" pageId="6" pageNumber="137">Fig. 18</figureCitation>
|
||
). Bursa copulatrix with spermathecal ducts sclerotised at the base, in shape of curved, slender, dark brown sticks (
|
||
<figureCitation id="132127BF5A1B265B2AC9FEFDFBB29EAC" box="[996,1079,367,393]" captionStart="FIGURES 15–20" captionStartId="5.[151,264,1883,1907]" captionTargetBox="[173,1415,193,1844]" captionTargetId="figure-20@5.[163,1425,181,1859]" captionTargetPageId="5" captionText="FIGURES 15–20. Female terminalia of Tipula (Vestiplex) levisoni sp. nov. 15, ovipositor, left lateral view; 16, sternite 8 with hypovalvae, ventral view; 17, sternite 9, dorsal view; 18, furca, dorsal view; 19, bursa copulatrix, dorsal view; 20, spermatheca. Abbreviations: bp sd, basal part of spermathecal duct; cds, cul-de-sac of bursa copulatrix; c, cerci; h, hypovalvae; s8, sternite 8; t10, tergite 10. Scale a: 15 = 1 mm; b: 16–20 = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6346184" httpUri="https://zenodo.org/record/6346184/files/figure.png" pageId="6" pageNumber="137">Fig. 19</figureCitation>
|
||
). Cul-de-sac of bursa copulatrix curved. Spermatheca pear-shaped (
|
||
<figureCitation id="132127BF5A1B265B2B32FE01FDF69E88" box="[543,627,403,429]" captionStart="FIGURES 15–20" captionStartId="5.[151,264,1883,1907]" captionTargetBox="[173,1415,193,1844]" captionTargetId="figure-20@5.[163,1425,181,1859]" captionTargetPageId="5" captionText="FIGURES 15–20. Female terminalia of Tipula (Vestiplex) levisoni sp. nov. 15, ovipositor, left lateral view; 16, sternite 8 with hypovalvae, ventral view; 17, sternite 9, dorsal view; 18, furca, dorsal view; 19, bursa copulatrix, dorsal view; 20, spermatheca. Abbreviations: bp sd, basal part of spermathecal duct; cds, cul-de-sac of bursa copulatrix; c, cerci; h, hypovalvae; s8, sternite 8; t10, tergite 10. Scale a: 15 = 1 mm; b: 16–20 = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6346184" httpUri="https://zenodo.org/record/6346184/files/figure.png" pageId="6" pageNumber="137">Fig. 20</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30068B15A1B265B29EAFE25FDFF9D3C" pageId="6" pageNumber="137" type="etymology">
|
||
<paragraph id="8BA53B3A5A1B265B29EAFE25FDFF9D3C" blockId="6.[151,1437,150,1401]" pageId="6" pageNumber="137">
|
||
<emphasis id="B96EE7285A1B265B29EAFE25FECB9EF4" bold="true" box="[199,334,439,465]" pageId="6" pageNumber="137">Etymology.</emphasis>
|
||
The new species is named after Levison James Wood (1982–), a British army officer, explorer and writer, best known for his remarkable expeditions in the Himalayas, Central America, the Arabian Peninsula and walking the entire length of the River Nile.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30068B15A1B265B29EAFDB0FBA89D1B" box="[199,1069,546,574]" pageId="6" pageNumber="137" type="distribution">
|
||
<paragraph id="8BA53B3A5A1B265B29EAFDB0FBA89D1B" blockId="6.[151,1437,150,1401]" box="[199,1069,546,574]" pageId="6" pageNumber="137">
|
||
<emphasis id="B96EE7285A1B265B29EAFDB0FEE49D19" bold="true" box="[199,353,546,572]" pageId="6" pageNumber="137">Distribution.</emphasis>
|
||
<collectingCountry id="F30D7BAA5A1B265B284AFDB1FE249D18" box="[359,417,547,573]" name="India" pageId="6" pageNumber="137">India</collectingCountry>
|
||
(
|
||
<collectingRegion id="49DEF5D85A1B265B2882FDB1FD1B9D1B" box="[431,670,547,574]" country="India" name="Jammu and Kashmir" pageId="6" pageNumber="137">Jammu and Kashmir</collectingRegion>
|
||
),
|
||
<collectingCountry id="F30D7BAA5A1B265B2B9DFDB1FD769D18" box="[688,755,547,573]" name="Nepal" pageId="6" pageNumber="137">Nepal</collectingCountry>
|
||
(Sudurpashchim Province).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30068B15A1B265B29EAFDD5FB049A5C" pageId="6" pageNumber="137" type="discussion">
|
||
<paragraph id="8BA53B3A5A1B265B29EAFDD5FAE29C33" blockId="6.[151,1437,150,1401]" pageId="6" pageNumber="137">
|
||
<emphasis id="B96EE7285A1B265B29EAFDD5FEBE9D44" bold="true" box="[199,315,583,609]" pageId="6" pageNumber="137">Remarks.</emphasis>
|
||
<taxonomicName id="4C1A40B95A1B265B286EFDD5FD9D9D44" authority="Starkevich & Podėnas & Sivell, 2022" authorityName="Starkevich & Podėnas & Sivell" authorityYear="2022" box="[323,536,583,609]" class="Insecta" family="Tipulidae" genus="Tipula" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="137" phylum="Arthropoda" rank="species" species="levisoni" status="sp. nov." subGenus="Vestiplex">
|
||
<emphasis id="B96EE7285A1B265B286EFDD5FE0E9D44" box="[323,395,583,609]" italics="true" pageId="6" pageNumber="137">Tipula</emphasis>
|
||
(
|
||
<emphasis id="B96EE7285A1B265B28B1FDD5FE349D45" box="[412,433,583,608]" italics="true" pageId="6" pageNumber="137">V.</emphasis>
|
||
)
|
||
<emphasis id="B96EE7285A1B265B28ECFDD5FD9D9D44" box="[449,536,583,609]" italics="true" pageId="6" pageNumber="137">levisoni</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B96EE7285A1B265B2B0DFDD5FDFE9D44" bold="true" box="[544,635,583,609]" pageId="6" pageNumber="137">
|
||
<taxonomicNameLabel id="A25D5A535A1B265B2B0DFDD5FDFE9D44" box="[544,635,583,609]" pageId="6" pageNumber="137" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is assigned here as a member of
|
||
<taxonomicName id="4C1A40B95A1B265B2AD9FDD5FB419D44" authorityName="Mik" authorityYear="1889" box="[1012,1220,583,609]" class="Insecta" family="Tipulidae" genus="Tipula" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="137" phylum="Arthropoda" rank="species" species="leucoprocta" subGenus="Vestiplex">
|
||
<emphasis id="B96EE7285A1B265B2AD9FDD5FB8C9D45" box="[1012,1033,583,608]" italics="true" pageId="6" pageNumber="137">T.</emphasis>
|
||
(
|
||
<emphasis id="B96EE7285A1B265B2D34FDD5FBAA9D45" box="[1049,1071,583,608]" italics="true" pageId="6" pageNumber="137">V.</emphasis>
|
||
)
|
||
<emphasis id="B96EE7285A1B265B2D12FDD5FB419D44" box="[1087,1220,583,609]" italics="true" pageId="6" pageNumber="137">leucoprocta</emphasis>
|
||
</taxonomicName>
|
||
species group proposed by
|
||
<bibRefCitation id="EF8B46CB5A1B265B2828FDF9FE489DA0" author="Savchenko, E. N." box="[261,461,619,645]" pageId="6" pageNumber="137" pagination="143 - 216" refId="ref8361" refString="Savchenko, E. N. (1960) A contribution to the taxonomy of crane-flies (Diptera, Tipulidae) of the subgenus Vestiplex Bezzi of the genus Tipula L. Horae Societatis Entomologicae Rossicae Unionis Soveticae, 47, 143 - 216. [in Russian]" type="journal article" year="1960">Savchenko (1960</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EF8B46CB5A1B265B28F4FDF9FD979DA3" author="Savchenko, E. N." box="[473,530,619,646]" pageId="6" pageNumber="137" pagination="1 - 503" refId="ref8410" refString="Savchenko, E. N. (1964) Crane-flies (Diptera, Tipulidae), Subfam. Tipulinae, Genus Tipula L., 2. Fauna USSR, New Series, 89, Nasekomye Dvukrylye [Insecta Diptera], 2 (4), 1 - 503. [in Russian]" type="journal article" year="1964">1964</bibRefCitation>
|
||
) and as discussed by
|
||
<bibRefCitation id="EF8B46CB5A1B265B2A26FDF9FB9E9DA0" author="Starkevich, P. & Podenas, S. & Gelhaus, J." box="[779,1051,619,645]" pageId="6" pageNumber="137" pagination="1 - 88" refId="ref8465" refString="Starkevich, P., Podenas, S. & Gelhaus, J. (2020) Taxonomic review of Tipula (Vestiplex Bezzi) crane flies (Diptera: Tipulidae) in Mongolia. Zootaxa, 4837 (1), 1 - 88. https: // doi. org / 10.11646 / zootaxa. 4837.1.1" type="journal article" year="2020">
|
||
Starkevich
|
||
<emphasis id="B96EE7285A1B265B2AA6FDF9FC3B9DA0" box="[907,958,619,645]" italics="true" pageId="6" pageNumber="137">et al</emphasis>
|
||
. (2020)
|
||
</bibRefCitation>
|
||
. Males of the
|
||
<taxonomicName id="4C1A40B95A1B265B2D93FDF9FAC69DA0" authorityName="Mik" authorityYear="1889" box="[1214,1347,619,645]" class="Insecta" family="Tipulidae" genus="Tipula" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="137" phylum="Arthropoda" rank="species" species="leucoprocta">
|
||
<emphasis id="B96EE7285A1B265B2D93FDF9FAC69DA0" box="[1214,1347,619,645]" italics="true" pageId="6" pageNumber="137">leucoprocta</emphasis>
|
||
</taxonomicName>
|
||
species group are distinguished by an unarmed gonocoxite, the epandrium having a large sclerotised plate with an extensive saucer with a raised border, posterolaterally terminating in teeth and a remarkably large gonocoxal fragment. In females, the bursa copulatrix and basal part of the spermathecal ducts are sclerotised. Two members of the
|
||
<taxonomicName id="4C1A40B95A1B265B2C35FD45FA189DD4" authorityName="Mik" authorityYear="1889" box="[1304,1437,727,753]" class="Insecta" family="Tipulidae" genus="Tipula" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="137" phylum="Arthropoda" rank="species" species="leucoprocta">
|
||
<emphasis id="B96EE7285A1B265B2C35FD45FA189DD4" box="[1304,1437,727,753]" italics="true" pageId="6" pageNumber="137">leucoprocta</emphasis>
|
||
</taxonomicName>
|
||
species group are known from the Western Himalayas,
|
||
<taxonomicName id="4C1A40B95A1B265B2A2FFD69FBAD9C33" authority="(Pakistan)" authorityName="Alexander" authorityYear="1961" baseAuthorityName="Pakistan" box="[770,1064,763,790]" class="Insecta" family="Tipulidae" genus="Tipula" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="137" phylum="Arthropoda" rank="species" species="tanycera" subGenus="Vestiplex">
|
||
<emphasis id="B96EE7285A1B265B2A2FFD69FC979C31" box="[770,786,763,788]" italics="true" pageId="6" pageNumber="137">T</emphasis>
|
||
. (
|
||
<emphasis id="B96EE7285A1B265B2A04FD69FCBB9C31" box="[809,830,763,788]" italics="true" pageId="6" pageNumber="137">V.</emphasis>
|
||
)
|
||
<emphasis id="B96EE7285A1B265B2A60FD69FC2A9C31" box="[845,943,763,788]" italics="true" pageId="6" pageNumber="137">tanycera</emphasis>
|
||
(
|
||
<collectingCountry id="F30D7BAA5A1B265B2A93FD69FBA59C33" box="[958,1056,763,790]" name="Pakistan" pageId="6" pageNumber="137">Pakistan</collectingCountry>
|
||
)
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C1A40B95A1B265B2D73FD69FAE69C33" authority="(India)" authorityName="Edwards" authorityYear="1927" baseAuthorityName="India" box="[1118,1379,763,790]" class="Insecta" family="Tipulidae" genus="Tipula" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="137" phylum="Arthropoda" rank="species" species="mitchelli" subGenus="Vestiplex">
|
||
<emphasis id="B96EE7285A1B265B2D73FD69FBEB9C31" box="[1118,1134,763,788]" italics="true" pageId="6" pageNumber="137">T</emphasis>
|
||
. (
|
||
<emphasis id="B96EE7285A1B265B2DA8FD69FB1E9C31" box="[1157,1179,763,788]" italics="true" pageId="6" pageNumber="137">V.</emphasis>
|
||
)
|
||
<emphasis id="B96EE7285A1B265B2D87FD69FA899C30" box="[1194,1292,763,789]" italics="true" pageId="6" pageNumber="137">mitchelli</emphasis>
|
||
(
|
||
<collectingCountry id="F30D7BAA5A1B265B2C36FD69FADF9C33" box="[1307,1370,763,790]" name="India" pageId="6" pageNumber="137">India</collectingCountry>
|
||
)
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BA53B3A5A1B265B29EAFC8DFECA9C80" blockId="6.[151,1437,150,1401]" pageId="6" pageNumber="137">
|
||
<taxonomicName id="4C1A40B95A1B265B29EAFC8DFE1B9C1C" authority="Starkevich & Podėnas & Sivell, 2022" authorityName="Starkevich & Podėnas & Sivell" authorityYear="2022" box="[199,414,799,825]" class="Insecta" family="Tipulidae" genus="Tipula" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="137" phylum="Arthropoda" rank="species" species="levisoni" status="sp. nov." subGenus="Vestiplex">
|
||
<emphasis id="B96EE7285A1B265B29EAFC8DFE8A9C1C" box="[199,271,799,825]" italics="true" pageId="6" pageNumber="137">Tipula</emphasis>
|
||
(
|
||
<emphasis id="B96EE7285A1B265B280CFC8DFEB39C1D" box="[289,310,799,824]" italics="true" pageId="6" pageNumber="137">V.</emphasis>
|
||
)
|
||
<emphasis id="B96EE7285A1B265B286AFC8DFE1B9C1C" box="[327,414,799,825]" italics="true" pageId="6" pageNumber="137">levisoni</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B96EE7285A1B265B288AFC8DFD879C1C" bold="true" box="[423,514,799,825]" pageId="6" pageNumber="137">
|
||
<taxonomicNameLabel id="A25D5A535A1B265B288AFC8DFD879C1C" box="[423,514,799,825]" pageId="6" pageNumber="137" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is closest to
|
||
<taxonomicName id="4C1A40B95A1B265B2BB4FC8DFCCF9C1D" authorityName="Alexander" authorityYear="1961" box="[665,842,799,825]" class="Insecta" family="Tipulidae" genus="Tipula" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="137" phylum="Arthropoda" rank="species" species="tanycera" subGenus="Vestiplex">
|
||
<emphasis id="B96EE7285A1B265B2BB4FC8DFD2C9C1D" box="[665,681,799,824]" italics="true" pageId="6" pageNumber="137">T</emphasis>
|
||
. (
|
||
<emphasis id="B96EE7285A1B265B2BECFC8DFD529C1D" box="[705,727,799,824]" italics="true" pageId="6" pageNumber="137">V.</emphasis>
|
||
)
|
||
<emphasis id="B96EE7285A1B265B2BC5FC8DFCCF9C1D" box="[744,842,799,824]" italics="true" pageId="6" pageNumber="137">tanycera</emphasis>
|
||
</taxonomicName>
|
||
based on body colouration and the shape of the hypopygium. Both species are characterized by a crescent-shaped epandrium, inner gonostylus with middorsal tooth, slender aedeagal guide and the shape of the gonocoxal fragment. Females are characterized by sternite 8 with deep lateral incisions.
|
||
</paragraph>
|
||
<paragraph id="8BA53B3A5A1B265B29EAFC3DFE299BE3" blockId="6.[151,1437,150,1401]" pageId="6" pageNumber="137">
|
||
Males can be separated by their antennae which reaches the base of the abdomen in
|
||
<taxonomicName id="4C1A40B95A1B265B2D73FC3DFB7E9CEC" authority="Starkevich & Podėnas & Sivell, 2022" authorityName="Starkevich & Podėnas & Sivell" authorityYear="2022" box="[1118,1275,943,969]" class="Insecta" family="Tipulidae" genus="Tipula" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="137" phylum="Arthropoda" rank="species" species="levisoni" status="sp. nov." subGenus="Vestiplex">
|
||
<emphasis id="B96EE7285A1B265B2D73FC3DFBF69CED" box="[1118,1139,943,968]" italics="true" pageId="6" pageNumber="137">T.</emphasis>
|
||
(
|
||
<emphasis id="B96EE7285A1B265B2DACFC3DFB139CED" box="[1153,1174,943,968]" italics="true" pageId="6" pageNumber="137">V.</emphasis>
|
||
)
|
||
<emphasis id="B96EE7285A1B265B2D89FC3DFB7E9CEC" box="[1188,1275,943,969]" italics="true" pageId="6" pageNumber="137">levisoni</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B96EE7285A1B265B2C2CFC3DFADC9CEC" bold="true" box="[1281,1369,943,969]" pageId="6" pageNumber="137">
|
||
<taxonomicNameLabel id="A25D5A535A1B265B2C2CFC3DFADC9CEC" box="[1281,1369,943,969]" pageId="6" pageNumber="137" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
while in
|
||
<taxonomicName id="4C1A40B95A1B265B2999FC41FEE59CC9" authorityName="Alexander" authorityYear="1961" box="[180,352,979,1005]" class="Insecta" family="Tipulidae" genus="Tipula" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="137" phylum="Arthropoda" rank="species" species="tanycera" subGenus="Vestiplex">
|
||
<emphasis id="B96EE7285A1B265B2999FC41FF4C9CC9" box="[180,201,979,1004]" italics="true" pageId="6" pageNumber="137">T.</emphasis>
|
||
(
|
||
<emphasis id="B96EE7285A1B265B29F4FC41FF6A9CC9" box="[217,239,979,1004]" italics="true" pageId="6" pageNumber="137">V.</emphasis>
|
||
)
|
||
<emphasis id="B96EE7285A1B265B29D3FC41FEE59CC9" box="[254,352,979,1004]" italics="true" pageId="6" pageNumber="137">tanycera</emphasis>
|
||
</taxonomicName>
|
||
the antenna is longer,almost reaching the middle of the abdomen. They can also be separated by the following details of the epandrium:
|
||
<taxonomicName id="4C1A40B95A1B265B2B7BFC65FD7D9B34" authority="Starkevich & Podėnas & Sivell, 2022" authorityName="Starkevich & Podėnas & Sivell" authorityYear="2022" box="[598,760,1015,1041]" class="Insecta" family="Tipulidae" genus="Tipula" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="137" phylum="Arthropoda" rank="species" species="levisoni" status="sp. nov." subGenus="Vestiplex">
|
||
<emphasis id="B96EE7285A1B265B2B7BFC65FDEE9B35" box="[598,619,1015,1040]" italics="true" pageId="6" pageNumber="137">T.</emphasis>
|
||
(
|
||
<emphasis id="B96EE7285A1B265B2B51FC65FD149B35" box="[636,657,1015,1040]" italics="true" pageId="6" pageNumber="137">V.</emphasis>
|
||
)
|
||
<emphasis id="B96EE7285A1B265B2B8CFC65FD7D9B34" box="[673,760,1015,1041]" italics="true" pageId="6" pageNumber="137">levisoni</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B96EE7285A1B265B2A2DFC65FCDF9B34" bold="true" box="[768,858,1015,1041]" pageId="6" pageNumber="137">
|
||
<taxonomicNameLabel id="A25D5A535A1B265B2A2DFC65FCDF9B34" box="[768,858,1015,1041]" pageId="6" pageNumber="137" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
has the posterior margin with a deep median notch and a large posterolateral tooth (
|
||
<figureCitation id="132127BF5A1B265B2B2CFB89FDC09B10" box="[513,581,1051,1077]" captionStart="FIGURES 5–14" captionStartId="4.[151,264,1794,1818]" captionTargetBox="[255,1322,189,1764]" captionTargetId="figure-26@4.[245,1343,181,1770]" captionTargetPageId="4" captionText="FIGURES 5–14. Male terminalia of Tipula (Vestiplex) levisoni sp. nov. 5, hypopygium, dorsal view; 6, hypopygium, lateral view; 7, epandrium, dorsal view; 8, left gonocoxite, lateral view; 9, left outer gonostylus; 10, left inner gonostylus, lateral view; 11, aedeagal guide, lateral view; 12, gonocoxal fragment, dorsal view; 13, sperm pump, dorsal view; 14, distal part of intromittent organ, lateral view. Abbreviations: aia, anterior immovable apodeme; ca, compressor apodeme; ep, epandrium; gcx, gonocoxite; ig, inner gonostylus; ls, lateral sclerite of gonocoxal fragment; ms, medial sclerite of gonocoxal fragment; mt, mid-dorsal tooth of inner gonostylus; og, outer gonostylus; pia, posterior immovable apodeme; pt, posterolateral tooth of epandrium. Scale a: 5, 6 = 0.5 mm; b: 7–13 = 0.5 mm; c: 14 = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.6346182" httpUri="https://zenodo.org/record/6346182/files/figure.png" pageId="6" pageNumber="137">Fig. 7</figureCitation>
|
||
), while in
|
||
<taxonomicName id="4C1A40B95A1B265B2B90FB89FCED9B11" authorityName="Alexander" authorityYear="1961" box="[701,872,1051,1077]" class="Insecta" family="Tipulidae" genus="Tipula" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="137" phylum="Arthropoda" rank="species" species="tanycera" subGenus="Vestiplex">
|
||
<emphasis id="B96EE7285A1B265B2B90FB89FD579B11" box="[701,722,1051,1076]" italics="true" pageId="6" pageNumber="137">T.</emphasis>
|
||
(
|
||
<emphasis id="B96EE7285A1B265B2BCCFB89FD729B11" box="[737,759,1051,1076]" italics="true" pageId="6" pageNumber="137">V.</emphasis>
|
||
)
|
||
<emphasis id="B96EE7285A1B265B2A2BFB89FCED9B11" box="[774,872,1051,1076]" italics="true" pageId="6" pageNumber="137">tanycera</emphasis>
|
||
</taxonomicName>
|
||
the median notch is shallower and has a distinctly smaller posterolateral tooth (
|
||
<figureCitation id="132127BF5A1B265B28F5FBADFDA99B7C" box="[472,556,1087,1113]" captionStart="FIGURES 41–50" captionStartId="11.[151,264,1865,1889]" captionTargetBox="[210,1377,191,1829]" captionTargetId="figure-20@11.[198,1389,181,1841]" captionTargetPageId="11" captionText="FIGURES 41–50. Male terminalia of Tipula (Vestiplex) tanycera. 41, hypopygium, dorsal view; 42, hypopygium, lateral view; 43, epandrium, dorsal view; 44, left gonocoxite, lateral view; 45, left outer gonostylus; 46, left inner gonostylus, lateral view; 47, aedeagal guide, lateral view; 48, gonocoxal fragment, dorsal view; 49, sperm pump, dorsal view; 50, distal part of intromittent organ, lateral view. Abbreviations: mt, mid-dorsal tooth of inner gonostylus; pt, posterolateral tooth of epandrium. Scale a: 41, 42 = 0.5 mm; b: 43–49 = 0.5 mm; c: 50 = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.6346194" httpUri="https://zenodo.org/record/6346194/files/figure.png" pageId="6" pageNumber="137">Fig. 43</figureCitation>
|
||
). The outer gonostylus of
|
||
<taxonomicName id="4C1A40B95A1B265B2A62FBADFC6A9B7C" authority="Starkevich & Podėnas & Sivell, 2022" authorityName="Starkevich & Podėnas & Sivell" authorityYear="2022" box="[847,1007,1087,1113]" class="Insecta" family="Tipulidae" genus="Tipula" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="137" phylum="Arthropoda" rank="species" species="levisoni" status="sp. nov." subGenus="Vestiplex">
|
||
<emphasis id="B96EE7285A1B265B2A62FBADFCE19B7D" box="[847,868,1087,1112]" italics="true" pageId="6" pageNumber="137">T.</emphasis>
|
||
(
|
||
<emphasis id="B96EE7285A1B265B2A59FBADFC0C9B7D" box="[884,905,1087,1112]" italics="true" pageId="6" pageNumber="137">V.</emphasis>
|
||
)
|
||
<emphasis id="B96EE7285A1B265B2AB5FBADFC6A9B7C" box="[920,1007,1087,1113]" italics="true" pageId="6" pageNumber="137">levisoni</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B96EE7285A1B265B2ADBFBADFBCA9B7C" bold="true" box="[1014,1103,1087,1113]" pageId="6" pageNumber="137">
|
||
<taxonomicNameLabel id="A25D5A535A1B265B2ADBFBADFBCA9B7C" box="[1014,1103,1087,1113]" pageId="6" pageNumber="137" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is flattened, leaf-shaped (
|
||
<figureCitation id="132127BF5A1B265B2C42FBADFF229B58" captionStart="FIGURES 5–14" captionStartId="4.[151,264,1794,1818]" captionTargetBox="[255,1322,189,1764]" captionTargetId="figure-26@4.[245,1343,181,1770]" captionTargetPageId="4" captionText="FIGURES 5–14. Male terminalia of Tipula (Vestiplex) levisoni sp. nov. 5, hypopygium, dorsal view; 6, hypopygium, lateral view; 7, epandrium, dorsal view; 8, left gonocoxite, lateral view; 9, left outer gonostylus; 10, left inner gonostylus, lateral view; 11, aedeagal guide, lateral view; 12, gonocoxal fragment, dorsal view; 13, sperm pump, dorsal view; 14, distal part of intromittent organ, lateral view. Abbreviations: aia, anterior immovable apodeme; ca, compressor apodeme; ep, epandrium; gcx, gonocoxite; ig, inner gonostylus; ls, lateral sclerite of gonocoxal fragment; ms, medial sclerite of gonocoxal fragment; mt, mid-dorsal tooth of inner gonostylus; og, outer gonostylus; pia, posterior immovable apodeme; pt, posterolateral tooth of epandrium. Scale a: 5, 6 = 0.5 mm; b: 7–13 = 0.5 mm; c: 14 = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.6346182" httpUri="https://zenodo.org/record/6346182/files/figure.png" pageId="6" pageNumber="137">Fig. 9</figureCitation>
|
||
), while in
|
||
<taxonomicName id="4C1A40B95A1B265B2832FBF1FE4E9B59" authorityName="Alexander" authorityYear="1961" box="[287,459,1123,1149]" class="Insecta" family="Tipulidae" genus="Tipula" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="137" phylum="Arthropoda" rank="species" species="tanycera" subGenus="Vestiplex">
|
||
<emphasis id="B96EE7285A1B265B2832FBF1FEB19B59" box="[287,308,1123,1148]" italics="true" pageId="6" pageNumber="137">T.</emphasis>
|
||
(
|
||
<emphasis id="B96EE7285A1B265B2869FBF1FEDF9B59" box="[324,346,1123,1148]" italics="true" pageId="6" pageNumber="137">V.</emphasis>
|
||
)
|
||
<emphasis id="B96EE7285A1B265B2844FBF1FE4E9B59" box="[361,459,1123,1148]" italics="true" pageId="6" pageNumber="137">tanycera</emphasis>
|
||
</taxonomicName>
|
||
it is nearly oval (
|
||
<figureCitation id="132127BF5A1B265B2BBDFBF1FD609B58" box="[656,741,1122,1149]" captionStart="FIGURES 41–50" captionStartId="11.[151,264,1865,1889]" captionTargetBox="[210,1377,191,1829]" captionTargetId="figure-20@11.[198,1389,181,1841]" captionTargetPageId="11" captionText="FIGURES 41–50. Male terminalia of Tipula (Vestiplex) tanycera. 41, hypopygium, dorsal view; 42, hypopygium, lateral view; 43, epandrium, dorsal view; 44, left gonocoxite, lateral view; 45, left outer gonostylus; 46, left inner gonostylus, lateral view; 47, aedeagal guide, lateral view; 48, gonocoxal fragment, dorsal view; 49, sperm pump, dorsal view; 50, distal part of intromittent organ, lateral view. Abbreviations: mt, mid-dorsal tooth of inner gonostylus; pt, posterolateral tooth of epandrium. Scale a: 41, 42 = 0.5 mm; b: 43–49 = 0.5 mm; c: 50 = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.6346194" httpUri="https://zenodo.org/record/6346194/files/figure.png" pageId="6" pageNumber="137">Fig. 45</figureCitation>
|
||
). The inner gonostylus bears a middorsal tooth with a broadly expanded base in
|
||
<taxonomicName id="4C1A40B95A1B265B2872FB15FD859B84" authority="Starkevich & Podėnas & Sivell, 2022" authorityName="Starkevich & Podėnas & Sivell" authorityYear="2022" box="[351,512,1159,1185]" class="Insecta" family="Tipulidae" genus="Tipula" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="137" phylum="Arthropoda" rank="species" species="levisoni" status="sp. nov." subGenus="Vestiplex">
|
||
<emphasis id="B96EE7285A1B265B2872FB15FEF09B85" box="[351,373,1159,1184]" italics="true" pageId="6" pageNumber="137">T.</emphasis>
|
||
(
|
||
<emphasis id="B96EE7285A1B265B28A9FB15FE1F9B85" box="[388,410,1159,1184]" italics="true" pageId="6" pageNumber="137">V.</emphasis>
|
||
)
|
||
<emphasis id="B96EE7285A1B265B2884FB15FD859B84" box="[425,512,1159,1185]" italics="true" pageId="6" pageNumber="137">levisoni</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B96EE7285A1B265B2B2AFB15FDE49B84" bold="true" box="[519,609,1159,1185]" pageId="6" pageNumber="137">
|
||
<taxonomicNameLabel id="A25D5A535A1B265B2B2AFB15FDE49B84" box="[519,609,1159,1185]" pageId="6" pageNumber="137" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="132127BF5A1B265B2B5CFB15FD419B84" box="[625,708,1159,1185]" captionStart="FIGURES 5–14" captionStartId="4.[151,264,1794,1818]" captionTargetBox="[255,1322,189,1764]" captionTargetId="figure-26@4.[245,1343,181,1770]" captionTargetPageId="4" captionText="FIGURES 5–14. Male terminalia of Tipula (Vestiplex) levisoni sp. nov. 5, hypopygium, dorsal view; 6, hypopygium, lateral view; 7, epandrium, dorsal view; 8, left gonocoxite, lateral view; 9, left outer gonostylus; 10, left inner gonostylus, lateral view; 11, aedeagal guide, lateral view; 12, gonocoxal fragment, dorsal view; 13, sperm pump, dorsal view; 14, distal part of intromittent organ, lateral view. Abbreviations: aia, anterior immovable apodeme; ca, compressor apodeme; ep, epandrium; gcx, gonocoxite; ig, inner gonostylus; ls, lateral sclerite of gonocoxal fragment; ms, medial sclerite of gonocoxal fragment; mt, mid-dorsal tooth of inner gonostylus; og, outer gonostylus; pia, posterior immovable apodeme; pt, posterolateral tooth of epandrium. Scale a: 5, 6 = 0.5 mm; b: 7–13 = 0.5 mm; c: 14 = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.6346182" httpUri="https://zenodo.org/record/6346182/files/figure.png" pageId="6" pageNumber="137">Fig. 10</figureCitation>
|
||
).
|
||
<taxonomicName id="4C1A40B95A1B265B2BF4FB15FC3D9B85" authorityName="Alexander" authorityYear="1961" box="[729,952,1159,1185]" class="Insecta" family="Tipulidae" genus="Tipula" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="137" phylum="Arthropoda" rank="species" species="tanycera" subGenus="Vestiplex">
|
||
<emphasis id="B96EE7285A1B265B2BF4FB15FCA49B84" box="[729,801,1159,1185]" italics="true" pageId="6" pageNumber="137">Tipula</emphasis>
|
||
(
|
||
<emphasis id="B96EE7285A1B265B2A1CFB15FCC39B85" box="[817,838,1159,1184]" italics="true" pageId="6" pageNumber="137">V.</emphasis>
|
||
)
|
||
<emphasis id="B96EE7285A1B265B2A7BFB15FC3D9B85" box="[854,952,1159,1184]" italics="true" pageId="6" pageNumber="137">tanycera</emphasis>
|
||
</taxonomicName>
|
||
has a middorsal tooth with a distinctly less expanded base (
|
||
<figureCitation id="132127BF5A1B265B2866FB39FE1A9BE3" box="[331,415,1195,1222]" captionStart="FIGURES 41–50" captionStartId="11.[151,264,1865,1889]" captionTargetBox="[210,1377,191,1829]" captionTargetId="figure-20@11.[198,1389,181,1841]" captionTargetPageId="11" captionText="FIGURES 41–50. Male terminalia of Tipula (Vestiplex) tanycera. 41, hypopygium, dorsal view; 42, hypopygium, lateral view; 43, epandrium, dorsal view; 44, left gonocoxite, lateral view; 45, left outer gonostylus; 46, left inner gonostylus, lateral view; 47, aedeagal guide, lateral view; 48, gonocoxal fragment, dorsal view; 49, sperm pump, dorsal view; 50, distal part of intromittent organ, lateral view. Abbreviations: mt, mid-dorsal tooth of inner gonostylus; pt, posterolateral tooth of epandrium. Scale a: 41, 42 = 0.5 mm; b: 43–49 = 0.5 mm; c: 50 = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.6346194" httpUri="https://zenodo.org/record/6346194/files/figure.png" pageId="6" pageNumber="137">Fig. 46</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA53B3A5A1B265B29EAFB5DFAB19A14" blockId="6.[151,1437,150,1401]" pageId="6" pageNumber="137">
|
||
Females can be separated by their body size which exceeds
|
||
<quantity id="4CE296DF5A1B265B2A73FB5DFC299BCC" box="[862,940,1231,1257]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="6" pageNumber="137" unit="mm" value="30.0">30 mm</quantity>
|
||
in
|
||
<taxonomicName id="4C1A40B95A1B265B2AE3FB5DFBE89BCC" authority="Starkevich & Podėnas & Sivell, 2022" authorityName="Starkevich & Podėnas & Sivell" authorityYear="2022" box="[974,1133,1231,1257]" class="Insecta" family="Tipulidae" genus="Tipula" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="137" phylum="Arthropoda" rank="species" species="levisoni" status="sp. nov." subGenus="Vestiplex">
|
||
<emphasis id="B96EE7285A1B265B2AE3FB5DFC669BCD" box="[974,995,1231,1256]" italics="true" pageId="6" pageNumber="137">T.</emphasis>
|
||
(
|
||
<emphasis id="B96EE7285A1B265B2ADFFB5DFB829BCD" box="[1010,1031,1231,1256]" italics="true" pageId="6" pageNumber="137">V.</emphasis>
|
||
)
|
||
<emphasis id="B96EE7285A1B265B2D3BFB5DFBE89BCC" box="[1046,1133,1231,1257]" italics="true" pageId="6" pageNumber="137">levisoni</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B96EE7285A1B265B2D5EFB5DFB499BCC" bold="true" box="[1139,1228,1231,1257]" pageId="6" pageNumber="137">
|
||
<taxonomicNameLabel id="A25D5A535A1B265B2D5EFB5DFB499BCC" box="[1139,1228,1231,1257]" pageId="6" pageNumber="137" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and is less than
|
||
<quantity id="4CE296DF5A1B265B2CADFB5DFF469A28" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="6" pageNumber="137" unit="mm" value="20.0">20 mm</quantity>
|
||
in
|
||
<taxonomicName id="4C1A40B95A1B265B29C8FB61FE159A29" authorityName="Alexander" authorityYear="1961" box="[229,400,1267,1293]" class="Insecta" family="Tipulidae" genus="Tipula" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="137" phylum="Arthropoda" rank="species" species="tanycera" subGenus="Vestiplex">
|
||
<emphasis id="B96EE7285A1B265B29C8FB61FF7F9A29" box="[229,250,1267,1292]" italics="true" pageId="6" pageNumber="137">T.</emphasis>
|
||
(
|
||
<emphasis id="B96EE7285A1B265B2827FB61FE9A9A29" box="[266,287,1267,1292]" italics="true" pageId="6" pageNumber="137">V.</emphasis>
|
||
)
|
||
<emphasis id="B96EE7285A1B265B2803FB61FE159A29" box="[302,400,1267,1292]" italics="true" pageId="6" pageNumber="137">tanycera</emphasis>
|
||
</taxonomicName>
|
||
. The cercus has small serrations and a nearly rounded tip in
|
||
<taxonomicName id="4C1A40B95A1B265B2D02FB61FB4B9A28" authority="Starkevich & Podėnas & Sivell, 2022" authorityName="Starkevich & Podėnas & Sivell" authorityYear="2022" box="[1071,1230,1267,1293]" class="Insecta" family="Tipulidae" genus="Tipula" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="137" phylum="Arthropoda" rank="species" species="levisoni" status="sp. nov." subGenus="Vestiplex">
|
||
<emphasis id="B96EE7285A1B265B2D02FB61FBC19A29" box="[1071,1092,1267,1292]" italics="true" pageId="6" pageNumber="137">T.</emphasis>
|
||
(
|
||
<emphasis id="B96EE7285A1B265B2D7EFB61FBED9A29" box="[1107,1128,1267,1292]" italics="true" pageId="6" pageNumber="137">V.</emphasis>
|
||
)
|
||
<emphasis id="B96EE7285A1B265B2D5AFB61FB4B9A28" box="[1143,1230,1267,1293]" italics="true" pageId="6" pageNumber="137">levisoni</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B96EE7285A1B265B2DF8FB61FAAB9A28" bold="true" box="[1237,1326,1267,1293]" pageId="6" pageNumber="137">
|
||
<taxonomicNameLabel id="A25D5A535A1B265B2DF8FB61FAAB9A28" box="[1237,1326,1267,1293]" pageId="6" pageNumber="137" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="132127BF5A1B265B2C11FB61FA0B9A28" box="[1340,1422,1266,1293]" captionStart="FIGURES 15–20" captionStartId="5.[151,264,1883,1907]" captionTargetBox="[173,1415,193,1844]" captionTargetId="figure-20@5.[163,1425,181,1859]" captionTargetPageId="5" captionText="FIGURES 15–20. Female terminalia of Tipula (Vestiplex) levisoni sp. nov. 15, ovipositor, left lateral view; 16, sternite 8 with hypovalvae, ventral view; 17, sternite 9, dorsal view; 18, furca, dorsal view; 19, bursa copulatrix, dorsal view; 20, spermatheca. Abbreviations: bp sd, basal part of spermathecal duct; cds, cul-de-sac of bursa copulatrix; c, cerci; h, hypovalvae; s8, sternite 8; t10, tergite 10. Scale a: 15 = 1 mm; b: 16–20 = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6346184" httpUri="https://zenodo.org/record/6346184/files/figure.png" pageId="6" pageNumber="137">Fig. 15</figureCitation>
|
||
), while in
|
||
<taxonomicName id="4C1A40B95A1B265B29D4FA85FE209A15" authorityName="Alexander" authorityYear="1961" box="[249,421,1303,1329]" class="Insecta" family="Tipulidae" genus="Tipula" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="137" phylum="Arthropoda" rank="species" species="tanycera" subGenus="Vestiplex">
|
||
<emphasis id="B96EE7285A1B265B29D4FA85FE8B9A15" box="[249,270,1303,1328]" italics="true" pageId="6" pageNumber="137">T.</emphasis>
|
||
(
|
||
<emphasis id="B96EE7285A1B265B2833FA85FEB69A15" box="[286,307,1303,1328]" italics="true" pageId="6" pageNumber="137">V.</emphasis>
|
||
)
|
||
<emphasis id="B96EE7285A1B265B286EFA85FE209A15" box="[323,421,1303,1328]" italics="true" pageId="6" pageNumber="137">tanycera</emphasis>
|
||
</taxonomicName>
|
||
the serrations are distinct on the distal end and the cercus tip is pointed (
|
||
<figureCitation id="132127BF5A1B265B2DF9FA85FAA29A14" box="[1236,1319,1302,1329]" captionStart="FIGURES 51–56" captionStartId="12.[151,264,1887,1911]" captionTargetBox="[177,1411,190,1849]" captionTargetId="figure-26@12.[166,1422,181,1864]" captionTargetPageId="12" captionText="FIGURES 51–56. Female terminalia of Tipula (Vestiplex) tanycera. 51, ovipositor, left lateral view; 52, sternite 8 with hypovalvae, ventral view; 53, sternite 9, dorsal view; 54, furca, dorsal view; 55, bursa copulatrix, dorsal view; 56, spermatheca. Abbreviation: cds, cul-de-sac of bursa copulatrix. Scale a: 50 = 1 mm; b: 51–54 = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6346196" httpUri="https://zenodo.org/record/6346196/files/figure.png" pageId="6" pageNumber="137">Fig. 51</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA53B3A5A1B265B29EAFAA9FB049A5C" blockId="6.[151,1437,150,1401]" pageId="6" pageNumber="137">
|
||
Another regional species related to
|
||
<taxonomicName id="4C1A40B95A1B265B2B7FFAA9FD749A70" authority="Starkevich & Podėnas & Sivell, 2022" authorityName="Starkevich & Podėnas & Sivell" authorityYear="2022" box="[594,753,1339,1365]" class="Insecta" family="Tipulidae" genus="Tipula" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="137" phylum="Arthropoda" rank="species" species="levisoni" status="sp. nov." subGenus="Vestiplex">
|
||
<emphasis id="B96EE7285A1B265B2B7FFAA9FDE29A71" box="[594,615,1339,1364]" italics="true" pageId="6" pageNumber="137">T.</emphasis>
|
||
(
|
||
<emphasis id="B96EE7285A1B265B2B5BFAA9FD0E9A71" box="[630,651,1339,1364]" italics="true" pageId="6" pageNumber="137">V.</emphasis>
|
||
)
|
||
<emphasis id="B96EE7285A1B265B2BB7FAA9FD749A70" box="[666,753,1339,1365]" italics="true" pageId="6" pageNumber="137">levisoni</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B96EE7285A1B265B2BD5FAA9FCD49A70" bold="true" box="[760,849,1339,1365]" pageId="6" pageNumber="137">
|
||
<taxonomicNameLabel id="A25D5A535A1B265B2BD5FAA9FCD49A70" box="[760,849,1339,1365]" pageId="6" pageNumber="137" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C1A40B95A1B265B2AABFAA9FBB59A71" authorityName="Alexander" authorityYear="1961" box="[902,1072,1339,1365]" class="Insecta" family="Tipulidae" genus="Tipula" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="137" phylum="Arthropoda" rank="species" species="tanycera" subGenus="Vestiplex">
|
||
<emphasis id="B96EE7285A1B265B2AABFAA9FC1E9A71" box="[902,923,1339,1364]" italics="true" pageId="6" pageNumber="137">T.</emphasis>
|
||
(
|
||
<emphasis id="B96EE7285A1B265B2A87FAA9FC3A9A71" box="[938,959,1339,1364]" italics="true" pageId="6" pageNumber="137">V.</emphasis>
|
||
)
|
||
<emphasis id="B96EE7285A1B265B2AE3FAA9FBB59A71" box="[974,1072,1339,1364]" italics="true" pageId="6" pageNumber="137">tanycera</emphasis>
|
||
</taxonomicName>
|
||
is
|
||
<taxonomicName id="4C1A40B95A1B265B2D7DFAA9FB799A70" authorityName="Edwards" authorityYear="1927" box="[1104,1276,1339,1365]" class="Insecta" family="Tipulidae" genus="Tipula" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="137" phylum="Arthropoda" rank="species" species="mitchelli" subGenus="Vestiplex">
|
||
<emphasis id="B96EE7285A1B265B2D7DFAA9FBE59A71" box="[1104,1120,1339,1364]" italics="true" pageId="6" pageNumber="137">T</emphasis>
|
||
. (
|
||
<emphasis id="B96EE7285A1B265B2D5BFAA9FB0E9A71" box="[1142,1163,1339,1364]" italics="true" pageId="6" pageNumber="137">V.</emphasis>
|
||
)
|
||
<emphasis id="B96EE7285A1B265B2DB7FAA9FB799A70" box="[1178,1276,1339,1365]" italics="true" pageId="6" pageNumber="137">mitchelli</emphasis>
|
||
</taxonomicName>
|
||
, which can be separated by a rectangular epandrium and the inner gonostylus lacking a middorsal tooth.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |