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<mods:title id="2B5B13011929E2901F0AA50FFEE776BF">The arboreal snail genus Amphidromus Albers, 1850 (Eupulmonata, Camaenidae) of Southeast Asia: 1. Molecular systematics of some Vietnamese species and related species from Cambodia, Indonesia, and Laos</mods:title>
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<figureCitation id="A1C6260C1FE46B76810A70CA51136F87" captionStart="Figure 30" captionStartId="F30" captionText="Figure 30. Shells of Amphidromus spp A-F Amphidromus roseolabiatus Fulton, 1896 A lectotype of &quot; Amphidromus roseolabiatus &quot; (NHMUK 19601462) B paralectotype of &quot; Amphidromus roseolabiatus &quot; (NHMUK 19601463) C holotype of &quot; Amphidromus phuonglinhae &quot; (MNHN-IM- 2000 - 33200) D-F specimens from Kampong Siem, Kampong Cham, Cambodia (NMNS- 8764 - 254, NMNS- 8764 - 258, NMNS- 8764 - 260) G-L Amphidromus pankowskianus Thach, 2020 G holotype (NHMUK 20200213) H-J specimens from Lak Sao, Khamkeut, Bolikhamsai, Laos (NMNS- 8764 - 154, NMNS- 8764 - 170, NMNS- 8764 - 192) K, L specimens from Khammouan Province, Laos, near Minh Hoa District, Quang Binh Province, Vietnam (NMNS- 8764 - 150, NMNS- 8764 - 151). Credit: M. Caballer (C), K. Webb (G)." figureDoi="10.3897/zookeys.1196.112146.figure30" httpUri="https://binary.pensoft.net/fig/1007848" pageId="0" pageNumber="15">Figs 30A-F</figureCitation>
<figureCitation id="00D17811170600F849174EC726F23777" captionStart="Figure 31" captionStartId="F31" captionText="Figure 31. Mitochondrial COI haplotype minimum spanning networks of Amphidromus roseolabiatus Fulton, 1896 and Amphidromus pankowskianus Thach, 2020. The size of each circle corresponds to the frequency of that haplotype, also shown as the number in that circle. The bars on the branches indicate the number of mutational steps between haplotypes. Specimen codes correspond to those in Table 1." figureDoi="10.3897/zookeys.1196.112146.figure31" httpUri="https://binary.pensoft.net/fig/1007849" pageId="0" pageNumber="15">, 31</figureCitation>
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Fulton, 1896: 89, pl. 6, fig. 8. Type locality: Siam [Thailand]. 
<bibRefCitation id="E0125CD6DD5CA94A64D0E6806B5AE910" author="Pilsbry, HA" journalOrPublisher="Academy of Natural Sciences of Philadelphia, Philadelphia" pageId="0" pageNumber="15" publicationUrl="https://www.biodiversitylibrary.org/item/16708" refId="B48" refString="Pilsbry, HA, 1900. Manual of Conchology, Second Series, Vol. 13. Academy of Natural Sciences of Philadelphia, Philadelphia, https://www.biodiversitylibrary.org/item/16708" title="Manual of Conchology, Second Series, Vol. 13." url="https://www.biodiversitylibrary.org/item/16708" year="1900">Pilsbry 1900</bibRefCitation>
: 188, pl. 60, fig. 36. 
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: 407. 
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: 527, 655. 
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: 42. 
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: 51. 
<bibRefCitation id="26BA4808974B012FBE361176B1872A6A" DOI="https://doi.org/10.3897/zookeys.492.8641" author="Sutcharit, C" journalOrPublisher="ZooKeys" pageId="0" pageNumber="15" pagination="49 - 105" refId="B68" refString="Sutcharit, C, Ablett, J, Tongkerd, P, Naggs, F, Panha, S, 2015. Illustrated type catalogue of Amphidromus Albers, 1850 in the Natural History Museum, London, and descriptions of two new species. ZooKeys 492: 49 - 105, DOI: https://doi.org/10.3897/zookeys.492.8641" title="Illustrated type catalogue of Amphidromus Albers, 1850 in the Natural History Museum, London, and descriptions of two new species." url="https://doi.org/10.3897/zookeys.492.8641" volume="492" year="2015">Sutcharit et al. 2015</bibRefCitation>
: 88, fig. 13j, k. 
<bibRefCitation id="7F0418243537438BDDAC98864D3190A8" DOI="https://doi.org/10.3897/zookeys.834.28800" author="Inkhavilay, K" journalOrPublisher="ZooKeys" pageId="0" pageNumber="15" pagination="1 - 166" refId="B27" refString="Inkhavilay, K, Sutcharit, C, Bantaowong, U, Chanabun, R, Siriwut, W, Srisonchai, R, Pholyotha, A, Jirapatrasilp, P, Panha, S, 2019. Annotated checklist of the terrestrial molluscs from Laos (Mollusca, Gastropoda). ZooKeys 834: 1 - 166, DOI: https://doi.org/10.3897/zookeys.834.28800" title="Annotated checklist of the terrestrial molluscs from Laos (Mollusca, Gastropoda)." url="https://doi.org/10.3897/zookeys.834.28800" volume="834" year="2019">Inkhavilay et al. 2019</bibRefCitation>
: 94, figs 45d-f, 58a. 
<bibRefCitation id="B5B49028EEE15196740D947E6F8C3889" DOI="https://doi.org/10.12657/folmal.028.002" author="Pall-Gergely, B" journalOrPublisher="Folia Malacologica" pageId="0" pageNumber="15" pagination="35 - 76" refId="B45" refString="Pall-Gergely, B, Hunyadi, A, Auffenberg, K, 2020. Taxonomic vandalism in malacology: Comments on molluscan taxa recently described by N. N. Thach and colleagues (2014-2019). Folia Malacologica 28 (1): 35 - 76, DOI: https://doi.org/10.12657/folmal.028.002" title="Taxonomic vandalism in malacology: Comments on molluscan taxa recently described by N. N. Thach and colleagues (2014 - 2019)." url="https://doi.org/10.12657/folmal.028.002" volume="28" year="2020">
<normalizedToken id="6751FF90BF4CBEF79377B2E23A7FCD8F" originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
et al. 2020
</bibRefCitation>
: 54. 
<bibRefCitation id="9DC9B4CF8F45EFCC289067E13C625734" DOI="https://doi.org/10.54173/F524359" author="Thach, NN" journalOrPublisher="The Festivus" pageId="0" pageNumber="15" pagination="359 - 362" refId="B76" refString="Thach, NN, 2020b. Remarks on Amphidromus reflexilabris Schepman, 1892, Amphidromus roseolabiatus Fulton, 1896, Amphidromus anhduongae Thach, 2020 and correction of errors in &quot;New Shells of South Asia, Volume 2&quot;. The Festivus 52 (4): 359 - 362, DOI: https://doi.org/10.54173/F524359" title="Remarks on Amphidromus reflexilabris Schepman, 1892, Amphidromus roseolabiatus Fulton, 1896, Amphidromus anhduongae Thach, 2020 and correction of errors in &quot; New Shells of South Asia, Volume 2 &quot;." url="https://doi.org/10.54173/F524359" volume="52" year="2020 b">Thach 2020b</bibRefCitation>
: 360, fig. 7.
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<taxonomicName id="07EC5EA94F2586764FAB76F854DE2E96" authorityName="Albers" authorityYear="1850" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Amphidromus" order="Stylommatophora" pageId="0" pageNumber="15" phylum="Mollusca" rank="genus">Amphidromus</taxonomicName>
<taxonomicName id="7A620B5A807A3D8AB2D6E57BF635136B" authorityName="Fulton" authorityYear="1896" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Amphidromus (Amphidromus) roseolabiatus" order="Stylommatophora" pageId="0" pageNumber="15" phylum="Mollusca" rank="species" species="roseolabiatus" subGenus="Amphidromus">Amphidromus (Amphidromus) roseolabiatus</taxonomicName>
. 
<bibRefCitation id="80FEBD56C7DCA05A04553290967EFF77" DOI="https://doi.org/10.5852/ejt.2017.330" author="Inkhavilay, K" journalOrPublisher="European Journal of Taxonomy" pageId="0" pageNumber="15" pagination="1 - 40" refId="B26" refString="Inkhavilay, K, Sutcharit, C, Panha, S, 2017. Taxonomic review of the tree snail genus Amphidromus Albers, 1850 (Pulmonata: Camaenidae) in Laos, with the description of two new species. European Journal of Taxonomy 330 (330): 1 - 40, DOI: https://doi.org/10.5852/ejt.2017.330" title="Taxonomic review of the tree snail genus Amphidromus Albers, 1850 (Pulmonata: Camaenidae) in Laos, with the description of two new species." url="https://doi.org/10.5852/ejt.2017.330" volume="330" year="2017">Inkhavilay et al. 2017</bibRefCitation>
: 3, 6, 9, 10, figs 2a, b, 3a, b, 4a-f, 6a, b, 7a-c.
</paragraph>
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<taxonomicName id="0672DC8066F394E03E2A8E1AA394E95C" authorityName="Thach" authorityYear="2017" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Amphidromus phuonglinhae" order="Stylommatophora" pageId="0" pageNumber="15" phylum="Mollusca" rank="species" species="phuonglinhae">Amphidromus phuonglinhae</taxonomicName>
Thach, 2017: 45, pl. 46, figs 581-584. Type locality: Bo Trach District, Quang Binh Province, Central Vietnam. 
<bibRefCitation id="A3C5B7ACEFA2046DB13243155E90D225" author="Thach, NN" journalOrPublisher="48 HRBooks Company, Akron, Ohio, USA" pageId="0" pageNumber="15" refId="B77" refString="Thach, NN, 2021. New shells of South Asia and Taiwan, China, Tanzania. Seashells*Freshwater*Land snails. With 116 new species and subspecies and rejected synonyms, accepted species. 48HRBooks Company, Akron, Ohio, USA" title="New shells of South Asia and Taiwan, China, Tanzania. Seashells * Freshwater * Land snails. With 116 new species and subspecies and rejected synonyms, accepted species." year="2021">Thach 2021</bibRefCitation>
: 76.
</paragraph>
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<paragraph id="7E056B41830A7CE5D7536475E433853E" pageId="0" pageNumber="15">Material examined.</paragraph>
<paragraph id="8ED8736C750BF2CDC43ADB7B1ABF3EED" pageId="0" pageNumber="15">
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: 
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, 
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of &quot; 
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<emphasis id="EF35C153E60A4E1DF4095B621B0D78BC" italics="true" pageId="0" pageNumber="15">Amphidromus roseolabiatus</emphasis>
</taxonomicName>
&quot;, NHMUK 19601462 (Fig. 
<figureCitation id="52F8833751E502AFD16E64D116AA34E1" captionStart="Figure 30" captionStartId="F30" captionText="Figure 30. Shells of Amphidromus spp A-F Amphidromus roseolabiatus Fulton, 1896 A lectotype of &quot; Amphidromus roseolabiatus &quot; (NHMUK 19601462) B paralectotype of &quot; Amphidromus roseolabiatus &quot; (NHMUK 19601463) C holotype of &quot; Amphidromus phuonglinhae &quot; (MNHN-IM- 2000 - 33200) D-F specimens from Kampong Siem, Kampong Cham, Cambodia (NMNS- 8764 - 254, NMNS- 8764 - 258, NMNS- 8764 - 260) G-L Amphidromus pankowskianus Thach, 2020 G holotype (NHMUK 20200213) H-J specimens from Lak Sao, Khamkeut, Bolikhamsai, Laos (NMNS- 8764 - 154, NMNS- 8764 - 170, NMNS- 8764 - 192) K, L specimens from Khammouan Province, Laos, near Minh Hoa District, Quang Binh Province, Vietnam (NMNS- 8764 - 150, NMNS- 8764 - 151). Credit: M. Caballer (C), K. Webb (G)." figureDoi="10.3897/zookeys.1196.112146.figure30" httpUri="https://binary.pensoft.net/fig/1007848" pageId="0" pageNumber="15">30A</figureCitation>
); 1S, 
<typeStatus id="27BF4633929FDF93C7A0A17B2ED8147C" type="paralectotype">paralectotype</typeStatus>
of &quot; 
<taxonomicName id="7F3FD3D16C2FF6028F52A4E992613698" authorityName="Fulton" authorityYear="1896" class="Gastropoda" family="Camaenidae" genus="Amphidromus" kingdom="Animalia" lsidName="Amphidromus roseolabiatus" order="Stylommatophora" pageId="0" pageNumber="15" phylum="Mollusca" rank="species" species="roseolabiatus">
<emphasis id="EB063D945D1D8652CB26AE093E7DF216" italics="true" pageId="0" pageNumber="15">Amphidromus roseolabiatus</emphasis>
</taxonomicName>
&quot;, NHMUK 19601463 (Fig. 
<figureCitation id="26D9FFFCCED1A6445C5EB337D2D2CE61" captionStart="Figure 30" captionStartId="F30" captionText="Figure 30. Shells of Amphidromus spp A-F Amphidromus roseolabiatus Fulton, 1896 A lectotype of &quot; Amphidromus roseolabiatus &quot; (NHMUK 19601462) B paralectotype of &quot; Amphidromus roseolabiatus &quot; (NHMUK 19601463) C holotype of &quot; Amphidromus phuonglinhae &quot; (MNHN-IM- 2000 - 33200) D-F specimens from Kampong Siem, Kampong Cham, Cambodia (NMNS- 8764 - 254, NMNS- 8764 - 258, NMNS- 8764 - 260) G-L Amphidromus pankowskianus Thach, 2020 G holotype (NHMUK 20200213) H-J specimens from Lak Sao, Khamkeut, Bolikhamsai, Laos (NMNS- 8764 - 154, NMNS- 8764 - 170, NMNS- 8764 - 192) K, L specimens from Khammouan Province, Laos, near Minh Hoa District, Quang Binh Province, Vietnam (NMNS- 8764 - 150, NMNS- 8764 - 151). Credit: M. Caballer (C), K. Webb (G)." figureDoi="10.3897/zookeys.1196.112146.figure30" httpUri="https://binary.pensoft.net/fig/1007848" pageId="0" pageNumber="15">30B</figureCitation>
)
</materialsCitation>
.
</paragraph>
<paragraph id="4F3A544030BBC1444176FEBF175E0AF7" pageId="0" pageNumber="15">
<materialsCitation id="F6D0BD4B466AC2542947F51B4682BF6E" collectingDate="2024-01-01" collectingDateMax="2024-12-31" collectingDateMin="2024-01-01" country="Vietnam" location="Sinistral" specimenCount="1" typeStatus="holotype">
<collectingCountry id="B7D65F815D45200F857268E1D74F89F1" name="Vietnam">Vietnam</collectingCountry>
: 
<location id="5F7214AC749EE20B4F3411C8495B415C" LSID="urn:lsid:plazi:treatment:EE0304285DA751C6BCD91642F7D44CBB:5F7214AC749EE20B4F3411C8495B415C" country="Vietnam" name="Sinistral">Sinistral</location>
, 
<emphasis id="EC5688A778204DE1A97D6051EB5FD7E0" bold="true" italics="true" pageId="0" pageNumber="15">
<typeStatus id="2C534CA82EF58F71AA68BD1F3603910A" type="holotype">holotype</typeStatus>
</emphasis>
of &quot; 
<taxonomicName id="6B69ABE5A3A393F7673CFC4F0EE1631B" authorityName="Thach" authorityYear="2017" class="Gastropoda" family="Camaenidae" genus="Amphidromus" kingdom="Animalia" lsidName="Amphidromus phuonglinhae" order="Stylommatophora" pageId="0" pageNumber="15" phylum="Mollusca" rank="species" species="phuonglinhae">
<emphasis id="39FC4113EB00D8AC590F15559B97201A" italics="true" pageId="0" pageNumber="15">Amphidromus phuonglinhae</emphasis>
</taxonomicName>
&quot;, MNHN-IM-2000-33200 (Fig. 
<figureCitation id="21BC79CDDEC32E1C03C8487F75688569" captionStart="Figure 30" captionStartId="F30" captionText="Figure 30. Shells of Amphidromus spp A-F Amphidromus roseolabiatus Fulton, 1896 A lectotype of &quot; Amphidromus roseolabiatus &quot; (NHMUK 19601462) B paralectotype of &quot; Amphidromus roseolabiatus &quot; (NHMUK 19601463) C holotype of &quot; Amphidromus phuonglinhae &quot; (MNHN-IM- 2000 - 33200) D-F specimens from Kampong Siem, Kampong Cham, Cambodia (NMNS- 8764 - 254, NMNS- 8764 - 258, NMNS- 8764 - 260) G-L Amphidromus pankowskianus Thach, 2020 G holotype (NHMUK 20200213) H-J specimens from Lak Sao, Khamkeut, Bolikhamsai, Laos (NMNS- 8764 - 154, NMNS- 8764 - 170, NMNS- 8764 - 192) K, L specimens from Khammouan Province, Laos, near Minh Hoa District, Quang Binh Province, Vietnam (NMNS- 8764 - 150, NMNS- 8764 - 151). Credit: M. Caballer (C), K. Webb (G)." figureDoi="10.3897/zookeys.1196.112146.figure30" httpUri="https://binary.pensoft.net/fig/1007848" pageId="0" pageNumber="15">30C</figureCitation>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="531EBF8ADCA30107567D0ABF92865A51" pageId="0" pageNumber="15" type="materials_examined">
<paragraph id="307C4795E661989CE49CCC2A204C2EAA" pageId="0" pageNumber="15">Other material examined.</paragraph>
<paragraph id="5537C15C155E3DAE89CCA3CEC3C54811" pageId="0" pageNumber="15">
<collectingCountry id="45AA1BC0C07C3DCE24799F3CAA116C3C" name="Cambodia">Cambodia</collectingCountry>
: 4D + 6S specimens, Kampong Siem District, 
<collectingRegion id="4F0DADD9A1EC70D60928CC09F8678AD5" country="Cambodia" name="Kampong Cham">Kampong Cham Province</collectingRegion>
, NMNS-8764-254- NMNS-8764-263 (Fig. 
<figureCitation id="433AF7333A86FD741133457580B1E609" captionStart="Figure 30" captionStartId="F30" captionText="Figure 30. Shells of Amphidromus spp A-F Amphidromus roseolabiatus Fulton, 1896 A lectotype of &quot; Amphidromus roseolabiatus &quot; (NHMUK 19601462) B paralectotype of &quot; Amphidromus roseolabiatus &quot; (NHMUK 19601463) C holotype of &quot; Amphidromus phuonglinhae &quot; (MNHN-IM- 2000 - 33200) D-F specimens from Kampong Siem, Kampong Cham, Cambodia (NMNS- 8764 - 254, NMNS- 8764 - 258, NMNS- 8764 - 260) G-L Amphidromus pankowskianus Thach, 2020 G holotype (NHMUK 20200213) H-J specimens from Lak Sao, Khamkeut, Bolikhamsai, Laos (NMNS- 8764 - 154, NMNS- 8764 - 170, NMNS- 8764 - 192) K, L specimens from Khammouan Province, Laos, near Minh Hoa District, Quang Binh Province, Vietnam (NMNS- 8764 - 150, NMNS- 8764 - 151). Credit: M. Caballer (C), K. Webb (G)." figureDoi="10.3897/zookeys.1196.112146.figure30" httpUri="https://binary.pensoft.net/fig/1007848" pageId="0" pageNumber="15">30D-F</figureCitation>
).
</paragraph>
<caption id="85B91166596F034DFAE9007979049BBF" doi="10.3897/zookeys.1196.112146.figure30" httpUri="https://binary.pensoft.net/fig/1007848" pageId="0" pageNumber="15" start="Figure 30" startId="F30">
<paragraph id="FA0BDAE1F970530AD002F9595BE5F5B9" pageId="0" pageNumber="15">
<emphasis id="9AB4229724F5D8DCCA94EAE3F015236F" bold="true" pageId="0" pageNumber="15">Figure 30.</emphasis>
Shells of 
<taxonomicName id="4118CDEDDFCEC0B6F371438846CCE65E" authorityName="Albers" authorityYear="1850" class="Gastropoda" family="Camaenidae" genus="Amphidromus" kingdom="Animalia" lsidName="Amphidromus" order="Stylommatophora" pageId="0" pageNumber="15" phylum="Mollusca" rank="genus">
<emphasis id="23FD9C8A59E82B2217C00465E5AFB421" italics="true" pageId="0" pageNumber="15">Amphidromus</emphasis>
</taxonomicName>
spp 
<emphasis id="D51F6A05B5C92A60DEF0FB86A9D7C7ED" bold="true" pageId="0" pageNumber="15">A-F</emphasis>
<taxonomicName id="402E076AB8BA665D144327F01C8B604D" authorityName="Fulton" authorityYear="1896" class="Gastropoda" family="Camaenidae" genus="Amphidromus" kingdom="Animalia" lsidName="Amphidromus roseolabiatus" order="Stylommatophora" pageId="0" pageNumber="15" phylum="Mollusca" rank="species" species="roseolabiatus">
<emphasis id="D386028064A09558C05F24B3C176A21F" italics="true" pageId="0" pageNumber="15">Amphidromus roseolabiatus</emphasis>
</taxonomicName>
Fulton, 1896 
<emphasis id="85DF7357685C2546382E41C7BBED4D90" bold="true" pageId="0" pageNumber="15">A</emphasis>
<typeStatus id="00E2B7FD2D0010A5C5B4D5B106DFFE99" type="lectotype">lectotype</typeStatus>
of &quot; 
<taxonomicName id="B3C285128116E0CA1D7A2EC8837473FC" authorityName="Fulton" authorityYear="1896" class="Gastropoda" family="Camaenidae" genus="Amphidromus" kingdom="Animalia" lsidName="Amphidromus roseolabiatus" order="Stylommatophora" pageId="0" pageNumber="15" phylum="Mollusca" rank="species" species="roseolabiatus">
<emphasis id="E1112C39A48A8469EFB30C6B282F86B7" italics="true" pageId="0" pageNumber="15">Amphidromus roseolabiatus</emphasis>
</taxonomicName>
&quot; (NHMUK 19601462) 
<emphasis id="4B2AABCC280E82D0C3B1AF1C13C4ED02" bold="true" pageId="0" pageNumber="15">B</emphasis>
<typeStatus id="2CD2A74D380D7F528A918A69FF1FB5CD" type="paralectotype">paralectotype</typeStatus>
of &quot; 
<taxonomicName id="349720BE6B1681526FEA9437F2A386EE" authorityName="Fulton" authorityYear="1896" class="Gastropoda" family="Camaenidae" genus="Amphidromus" kingdom="Animalia" lsidName="Amphidromus roseolabiatus" order="Stylommatophora" pageId="0" pageNumber="15" phylum="Mollusca" rank="species" species="roseolabiatus">
<emphasis id="7F3F9D9273CBE188FFF146820C8477EB" italics="true" pageId="0" pageNumber="15">Amphidromus roseolabiatus</emphasis>
</taxonomicName>
&quot; (NHMUK 19601463) 
<emphasis id="8588F7F500C75882E2D565035258E954" bold="true" pageId="0" pageNumber="15">C</emphasis>
<typeStatus id="CA461A3CE3DA5437E9ACBDF8F0347D2E" type="holotype">holotype</typeStatus>
of &quot; 
<taxonomicName id="E44D30A52787EE106C3E3C4DDF4A7EF5" authorityName="Thach" authorityYear="2017" class="Gastropoda" family="Camaenidae" genus="Amphidromus" kingdom="Animalia" lsidName="Amphidromus phuonglinhae" order="Stylommatophora" pageId="0" pageNumber="15" phylum="Mollusca" rank="species" species="phuonglinhae">
<emphasis id="B3CA407F65B99F77A55B1A53394C475A" italics="true" pageId="0" pageNumber="15">Amphidromus phuonglinhae</emphasis>
</taxonomicName>
&quot; (MNHN-IM-2000-33200) 
<emphasis id="697A8A330EB702EA41A87B4D52E3035B" bold="true" pageId="0" pageNumber="15">D-F</emphasis>
specimens from Kampong Siem, 
<collectingRegion id="05A99F2B50F17B141792D075751616C8" country="Cambodia" name="Kampong Cham">Kampong Cham</collectingRegion>
, 
<collectingCountry id="60508913A63C236D2A310BE7AF80A01D" name="Cambodia">Cambodia</collectingCountry>
(NMNS-8764-254, NMNS-8764-258, NMNS-8764-260) 
<emphasis id="5F1964AC91687FB3FAF67E3336A39A1E" bold="true" pageId="0" pageNumber="15">G-L</emphasis>
<taxonomicName id="D372A67F6ABB55AFDC433C15CDE81901" authorityName="Thach" authorityYear="2020" class="Gastropoda" family="Camaenidae" genus="Amphidromus" kingdom="Animalia" lsidName="Amphidromus pankowskianus" order="Stylommatophora" pageId="0" pageNumber="15" phylum="Mollusca" rank="species" species="pankowskianus">
<emphasis id="1C115C8D83EFF65486E617B96E2ED7B2" italics="true" pageId="0" pageNumber="15">Amphidromus pankowskianus</emphasis>
</taxonomicName>
Thach, 2020 
<emphasis id="7C201735D3A02ECA11C69270222B0292" bold="true" pageId="0" pageNumber="15">G</emphasis>
<typeStatus id="CF0B5FB0C32E7D91046589B36DD6343D" type="holotype">holotype</typeStatus>
(NHMUK 20200213) 
<emphasis id="51D1DA54C73540DADBA60FE0EB110569" bold="true" pageId="0" pageNumber="15">H-J</emphasis>
specimens from Lak Sao, Khamkeut, 
<collectingRegion id="F0C8C24CEBF525C81A4A7C65D897E308" country="Laos" name="Bolikhamxai">Bolikhamsai</collectingRegion>
, 
<collectingCountry id="AD46548D02DCD8D0B114BFA2C38C9ED3" name="Laos">Laos</collectingCountry>
(NMNS-8764-154, NMNS-8764-170, NMNS-8764-192) 
<emphasis id="3092A8167C1902CBE15FE385088C7B62" bold="true" pageId="0" pageNumber="15">K, L</emphasis>
specimens from 
<collectingRegion id="3FF32076770902DEE08986128063258A" country="Laos" name="Khammouan">Khammouan Province</collectingRegion>
, 
<collectingCountry id="5AE3EB413014A896D8D6247A7183641A" name="Laos">Laos</collectingCountry>
, near Minh Hoa District, 
<collectingRegion id="D265EDBAA6469B9C029A065E2CD8DBC6" country="Vietnam" name="Quang Binh">Quang Binh Province</collectingRegion>
, 
<collectingCountry id="F7C2EAD017BE847DE27CAEBC025A848A" name="Vietnam">Vietnam</collectingCountry>
(NMNS-8764-150, NMNS-8764-151). Credit: M. Caballer (
<emphasis id="E341B031B2C3C4A52CC2606ED7A66279" bold="true" pageId="0" pageNumber="15">C</emphasis>
), K. Webb (
<emphasis id="EB1AB0AEB7EBF891FE9E7082CF71463F" bold="true" pageId="0" pageNumber="15">G</emphasis>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="199F03BB3CBEEAA3BC8A867BCBDF2D49" pageId="0" pageNumber="15" type="diagnosis">
<paragraph id="447B0DC1A5ADAF7383589CB3F54D0542" pageId="0" pageNumber="15">Diagnosis.</paragraph>
<paragraph id="91AD80A6D375616A2F85DA4DA068EB2A" pageId="0" pageNumber="15">Shell medium and chirally dimorphic. Parietal callus transparent; lip and columella purplish pink. Genitalia with appendix.</paragraph>
</subSubSection>
<subSubSection id="929CDC717C4B0A02A9775F85787D0699" pageId="0" pageNumber="15" type="differential diagnosis">
<paragraph id="E4D4B0A8453C417F8767492DC471F038" pageId="0" pageNumber="15">Differential diagnosis.</paragraph>
<paragraph id="57876526EE3DA4CFA32D32B09CE2C4A6" pageId="0" pageNumber="15">
This species is very closely similar to 
<taxonomicName id="A8A7D36D08A2E55C2BB0289E879BEB71" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. pankowskianus" pageId="0" pageNumber="15" phylum="Mollusca" rank="species" species="pankowskianus">
<emphasis id="A528BC3F0D72BE8B166A5D0A78361684" italics="true" pageId="0" pageNumber="15">A. pankowskianus</emphasis>
</taxonomicName>
in terms of shell morphology and colour pattern. However, this species lacks a dark radial band behind the reflected lip which is also visible in the inner side of the shell, and a dark spiral band below periphery, both of which are present in 
<taxonomicName id="6376D072165D043E7AEB4987F428099C" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. pankowskianus" pageId="0" pageNumber="15" phylum="Mollusca" rank="species" species="pankowskianus">
<emphasis id="DB462FC3ADDA08671580489BB3DF378F" italics="true" pageId="0" pageNumber="15">A. pankowskianus</emphasis>
</taxonomicName>
. 
<taxonomicName id="3E4F09959E974821E902DA9A30D4A118" authorityName="Fulton" authorityYear="1896" class="Gastropoda" family="Camaenidae" genus="Amphidromus" kingdom="Animalia" lsidName="Amphidromus roseolabiatus" order="Stylommatophora" pageId="0" pageNumber="15" phylum="Mollusca" rank="species" species="roseolabiatus">
<emphasis id="A6D9AE2E499B959F4CF573DB5E19DBF5" italics="true" pageId="0" pageNumber="15">Amphidromus roseolabiatus</emphasis>
</taxonomicName>
differs from the similar species 
<taxonomicName id="0F57A6A001382FD184F841DA62E3F01B" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. madelineae" pageId="0" pageNumber="15" phylum="Mollusca" rank="species" species="madelineae">
<emphasis id="E5848BDCEA520F4F05CFF0541086FDDD" italics="true" pageId="0" pageNumber="15">A. madelineae</emphasis>
</taxonomicName>
and 
<taxonomicName id="6E5E3479651851E8DCEE330162709B36" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. haematostoma" pageId="0" pageNumber="15" phylum="Mollusca" rank="species" species="haematostoma">
<emphasis id="C64AF6B654283E4F2F987B0B75B82AC9" italics="true" pageId="0" pageNumber="15">A. haematostoma</emphasis>
</taxonomicName>
in having a chirally dimorphic shell, and genitalia with a very long appendix, while both 
<taxonomicName id="ECB9158DD1259C49A10D7C8403B02A6A" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. madelineae" pageId="0" pageNumber="15" phylum="Mollusca" rank="species" species="madelineae">
<emphasis id="6126AB56E4BEFD649F141A46F4321F6C" italics="true" pageId="0" pageNumber="15">A. madelineae</emphasis>
</taxonomicName>
and 
<taxonomicName id="9E0E6F397E7ED898A782219F29331B40" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. haematostoma" pageId="0" pageNumber="15" phylum="Mollusca" rank="species" species="haematostoma">
<emphasis id="7F9787358B05D0818DF1685B02675888" italics="true" pageId="0" pageNumber="15">A. haematostoma</emphasis>
</taxonomicName>
are exclusively sinistral, and the genitalia lacks an appendix. 
<taxonomicName id="72271436E39BF5B5D16941651F2307EB" authorityName="Fulton" authorityYear="1896" class="Gastropoda" family="Camaenidae" genus="Amphidromus" kingdom="Animalia" lsidName="Amphidromus roseolabiatus" order="Stylommatophora" pageId="0" pageNumber="15" phylum="Mollusca" rank="species" species="roseolabiatus">
<emphasis id="163577D65824A0FDDB7D0B76A3C018C3" italics="true" pageId="0" pageNumber="15">Amphidromus roseolabiatus</emphasis>
</taxonomicName>
also differs from both 
<taxonomicName id="DFC8EC00F86B963550C14ABFBF8C635A" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. smithi" pageId="0" pageNumber="15" phylum="Mollusca" rank="species" species="smithi">
<emphasis id="EF4446EB295E1ECF8E4FDA648E9D6385" italics="true" pageId="0" pageNumber="15">A. smithi</emphasis>
</taxonomicName>
Fulton, 1896 and 
<taxonomicName id="607B5744337863DDCF60FD80DF9B4A56" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. ventrosulus" pageId="0" pageNumber="15" phylum="Mollusca" rank="species" species="ventrosulus">
<emphasis id="0643C385EAEB46C717042E607B546CCF" italics="true" pageId="0" pageNumber="15">A. ventrosulus</emphasis>
</taxonomicName>
<normalizedToken id="065EFF4EEF748CF38E79E2B9EE221E44" originalValue="Möllendorff">Moellendorff</normalizedToken>
, 1900 from Vietnam (
<bibRefCitation id="147FC83BF3E38DC8D3EAE1D81C37FE18" author="Zilch, A" journalOrPublisher="Archiv fuer Molluskenkunde" pageId="0" pageNumber="15" pagination="131 - 140" refId="B83" refString="Zilch, A, 1953. Die Typen und Typoide des Natur-Museums Senckenberg, 10: Mollusca, Pleurodontidae (1). Archiv fuer Molluskenkunde 82: 131 - 140" title="Die Typen und Typoide des Natur-Museums Senckenberg, 10: Mollusca, Pleurodontidae (1)." volume="82" year="1953">Zilch 1953</bibRefCitation>
; 
<bibRefCitation id="3D681145C00BA81DF6E2B86985DDE99A" DOI="https://doi.org/10.3897/zookeys.492.8641" author="Sutcharit, C" journalOrPublisher="ZooKeys" pageId="0" pageNumber="15" pagination="49 - 105" refId="B68" refString="Sutcharit, C, Ablett, J, Tongkerd, P, Naggs, F, Panha, S, 2015. Illustrated type catalogue of Amphidromus Albers, 1850 in the Natural History Museum, London, and descriptions of two new species. ZooKeys 492: 49 - 105, DOI: https://doi.org/10.3897/zookeys.492.8641" title="Illustrated type catalogue of Amphidromus Albers, 1850 in the Natural History Museum, London, and descriptions of two new species." url="https://doi.org/10.3897/zookeys.492.8641" volume="492" year="2015">Sutcharit et al. 2015</bibRefCitation>
) in having a chirally dimorphic shell, a purplish pink lip and fine green streaks. In contrast, 
<taxonomicName id="F291FF543D96B438250F8C6369A14F7C" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. smithi" pageId="0" pageNumber="15" phylum="Mollusca" rank="species" species="smithi">
<emphasis id="36D920C504E785F55808F6C43586DC36" italics="true" pageId="0" pageNumber="15">A. smithi</emphasis>
</taxonomicName>
has a sinistral shell, a dark red to brownish lip with dark spot on the apex, and 
<taxonomicName id="0AD02ABB04D64C363EF18D3EFF88946E" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. ventrosulus" pageId="0" pageNumber="15" phylum="Mollusca" rank="species" species="ventrosulus">
<emphasis id="7E8CC7FFC837C3437F6CEBDF900C3B75" italics="true" pageId="0" pageNumber="15">A. ventrosulus</emphasis>
</taxonomicName>
has a sinistral shell, uniform green colour, elongate spire and more depressed suture. The molecular phylogeny in this study reveals that 
<taxonomicName id="06F6E01DCD9A59A799AEC6EF995F7D4E" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. roseolabiatus" pageId="0" pageNumber="15" phylum="Mollusca" rank="species" species="roseolabiatus">
<emphasis id="7B4C8058083EA332B0F6887186356C4F" italics="true" pageId="0" pageNumber="15">A. roseolabiatus</emphasis>
</taxonomicName>
is a distinct clade from its sister 
<taxonomicName id="282D3AA6E7FBD8BF581DC9BF86C808E8" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. pankowskianus" pageId="0" pageNumber="15" phylum="Mollusca" rank="species" species="pankowskianus">
<emphasis id="8826EAF8399BF082E570BA4A39403DB7" italics="true" pageId="0" pageNumber="15">A. pankowskianus</emphasis>
</taxonomicName>
(Fig. 
<figureCitation id="5CE8E728D69ED26B00588FDB93CFB9CA" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Bayesian phylogeny of Amphidromus spp. based on mitochondrial COI and 16 S genes. Nodal support values are given as SH-aLRT / aBayes / ultra-fast bootstrap (IQ-TREE, ML) / posterior probability (MrBayes, BI). An asterisk on each branch indicates a clade with all well-supported values (SH-aLRT ≥ 80 %, aBayes ≥ 0.95, BS ≥ 95 %, PP ≥ 0.95). Species numbers correspond to those in Fig. 1 and Tables 1, 2." figureDoi="10.3897/zookeys.1196.112146.figure2" httpUri="https://binary.pensoft.net/fig/1007820" pageId="0" pageNumber="15">2</figureCitation>
). The COI and 16S 
<emphasis id="AB4DB95A15C4915FD8FF04F13C4E0D50" italics="true" pageId="0" pageNumber="15">p</emphasis>
-distances between 
<taxonomicName id="FB5A2CAB02B648FA3DABD6579F3BC723" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. roseolabiatus" pageId="0" pageNumber="15" phylum="Mollusca" rank="species" species="roseolabiatus">
<emphasis id="764735472B8733499A577E9A3DEAD2E1" italics="true" pageId="0" pageNumber="15">A. roseolabiatus</emphasis>
</taxonomicName>
and 
<taxonomicName id="F706EB4C4F38401E9D02817E5287CD7C" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. pankowskianus" pageId="0" pageNumber="15" phylum="Mollusca" rank="species" species="pankowskianus">
<emphasis id="746372FA3B41C1F11975512BD4529BAF" italics="true" pageId="0" pageNumber="15">A. pankowskianus</emphasis>
</taxonomicName>
are 13.02% and 6.14%, respectively (Table 
<tableCitation id="8F5CC55A4C404CFFE8D9225E592ED902" captionStart="Table 2" captionStartId="T2" captionText="Table 2. Percentage of uncorrected pairwise interspecific distances for the partial COI (above the diagonal) and 16 S rRNA (below the diagonal) gene fragments among the Amphidromus species in this study. Intraspecific distances for COI / 16 S rRNA are shown on the diagonal (bold). Species numbers correspond to those in Figs 1, 2, and Table 1. The numbers of sequences used to calculate the distances of each respective gene fragment are given as n = COI, 16 S in the first column." httpUri="http://table.plazi.org/id/A1B4F960F999761C3840BD02D5BE2608" pageId="0" pageNumber="15" tableUuid="A1B4F960F999761C3840BD02D5BE2608">2</tableCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="AAD5810565FBC8806FB4F3A0D0683E5F" pageId="0" pageNumber="15" type="description">
<paragraph id="126213FE406C173D331FBFFF2E57BCF8" pageId="0" pageNumber="15">Description.</paragraph>
<paragraph id="9F52DC50768108F821C8E1B4D6CCC5F3" pageId="0" pageNumber="15">
<emphasis id="2B7DD916B045761BD16FC2B866EC069F" bold="true" italics="true" pageId="0" pageNumber="15">Shell</emphasis>
medium (height 33.1-38.6 mm, width 19.2-21.6 mm), chirally dimorphic, elongate to ovate conical, rather thin and glossy. Spire conical; apex acute, light brown and without black spot on tip. Whorls 6-7 nearly smooth; suture wide and depressed; last whorl rounded. Periostracum usually deciduous to yellowish green radial streaks, more conspicuous on last whorl and faded in earlier whorls. Last whorl with thin, whitish subsutural band; with or without reddish brown spiral band on periphery but usually not reaching apertural lip; varix absent. Parietal callus thin and transparent. Aperture ovate to elongate; peristome expanded and not reflected; lip usually purplish pink. Columella straight, thickened, purplish pink. Umbilicus narrowly opened.
</paragraph>
</subSubSection>
<subSubSection id="193F614F3ECD6191E77D2EC4F6C95567" pageId="0" pageNumber="15" type="haplotype network">
<paragraph id="521D8F816642945A99ACF5C9393A6501" pageId="0" pageNumber="15">Haplotype network.</paragraph>
<paragraph id="7901F6DE43DE5D06181762097F055267" pageId="0" pageNumber="15">
There was one COI haplotype of 
<taxonomicName id="C1BA4DF0B9714A1189175625707E95CF" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. roseolabiatus" pageId="0" pageNumber="15" phylum="Mollusca" rank="species" species="roseolabiatus">
<emphasis id="28EE5DD7AB00698BBAF131C1C6523792" italics="true" pageId="0" pageNumber="15">A. roseolabiatus</emphasis>
</taxonomicName>
in this study (Fig. 
<figureCitation id="C10FDF38781943567E05CFFB70973F8C" captionStart="Figure 31" captionStartId="F31" captionText="Figure 31. Mitochondrial COI haplotype minimum spanning networks of Amphidromus roseolabiatus Fulton, 1896 and Amphidromus pankowskianus Thach, 2020. The size of each circle corresponds to the frequency of that haplotype, also shown as the number in that circle. The bars on the branches indicate the number of mutational steps between haplotypes. Specimen codes correspond to those in Table 1." figureDoi="10.3897/zookeys.1196.112146.figure31" httpUri="https://binary.pensoft.net/fig/1007849" pageId="0" pageNumber="15">31</figureCitation>
).
</paragraph>
<caption id="4E79855581B49AA85629BDBED7B107E5" doi="10.3897/zookeys.1196.112146.figure31" httpUri="https://binary.pensoft.net/fig/1007849" pageId="0" pageNumber="15" start="Figure 31" startId="F31">
<paragraph id="3697CBF0F8F55F0A6FBA5374611569AA" pageId="0" pageNumber="15">
<emphasis id="5D36CD51BE532E485215EB22EC76B4B5" bold="true" pageId="0" pageNumber="15">Figure 31.</emphasis>
Mitochondrial COI haplotype minimum spanning networks of 
<taxonomicName id="8AD8A4276E56A357E34721AF3721BB0F" authorityName="Fulton" authorityYear="1896" class="Gastropoda" family="Camaenidae" genus="Amphidromus" kingdom="Animalia" lsidName="Amphidromus roseolabiatus" order="Stylommatophora" pageId="0" pageNumber="15" phylum="Mollusca" rank="species" species="roseolabiatus">
<emphasis id="6BA2300FE6628A95CE6F46B9B36E3698" italics="true" pageId="0" pageNumber="15">Amphidromus roseolabiatus</emphasis>
</taxonomicName>
Fulton, 1896 and 
<taxonomicName id="DAD1B6E0F758B819FBDFFAC4F2A43E10" authorityName="Thach" authorityYear="2020" class="Gastropoda" family="Camaenidae" genus="Amphidromus" kingdom="Animalia" lsidName="Amphidromus pankowskianus" order="Stylommatophora" pageId="0" pageNumber="15" phylum="Mollusca" rank="species" species="pankowskianus">
<emphasis id="410E3517A45615DA2ADC2B06BA2380C3" italics="true" pageId="0" pageNumber="15">Amphidromus pankowskianus</emphasis>
</taxonomicName>
Thach, 2020. The size of each circle corresponds to the frequency of that haplotype, also shown as the number in that circle. The bars on the branches indicate the number of mutational steps between haplotypes. Specimen codes correspond to those in Table 
<tableCitation id="DAFD8F0497E72093F40F866395C1385F" captionStart="Table 1" captionStartId="T1" captionText="Table 1. List of specimens used in this study with species name, locality details, voucher and GenBank accession numbers. Species numbers correspond to those in Figs 1, 2, and Table 2." httpUri="http://table.plazi.org/id/536020F98423605B1D469F287C9C42E1" pageId="0" pageNumber="15" tableUuid="536020F98423605B1D469F287C9C42E1">1</tableCitation>
.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="5E34D985FBE46B77831C25DD7699F112" pageId="0" pageNumber="15" type="distribution">
<paragraph id="BB85ACF13DC998E742D9C2A13239F925" pageId="0" pageNumber="15">Distribution.</paragraph>
<paragraph id="2CF3034E847E59135DB4AFF2C4CA6F61" pageId="0" pageNumber="15">
This species is found in Khammouan Province, Laos, Kampong Cham Province, Cambodia, and Quang Binh Province, Vietnam. The distribution of this species in Bolikhamxay, Laos according to 
<bibRefCitation id="0F33D1ECB29F2983C2CCDACBF1513A1E" DOI="https://doi.org/10.5852/ejt.2017.330" author="Inkhavilay, K" journalOrPublisher="European Journal of Taxonomy" pageId="0" pageNumber="15" pagination="1 - 40" refId="B26" refString="Inkhavilay, K, Sutcharit, C, Panha, S, 2017. Taxonomic review of the tree snail genus Amphidromus Albers, 1850 (Pulmonata: Camaenidae) in Laos, with the description of two new species. European Journal of Taxonomy 330 (330): 1 - 40, DOI: https://doi.org/10.5852/ejt.2017.330" title="Taxonomic review of the tree snail genus Amphidromus Albers, 1850 (Pulmonata: Camaenidae) in Laos, with the description of two new species." url="https://doi.org/10.5852/ejt.2017.330" volume="330" year="2017">Inkhavilay et al. (2017)</bibRefCitation>
is dubious (see below).
</paragraph>
</subSubSection>
<subSubSection id="EB2AD145E4702C51563FB2ADC371D228" pageId="0" pageNumber="15" type="remarks">
<paragraph id="85B3233971308AB627646E60B7A699CE" pageId="0" pageNumber="15">Remarks.</paragraph>
<paragraph id="AE0AFE893D847CDA585E18134FAB3FAF" pageId="0" pageNumber="15">
<bibRefCitation id="7B5BB9B77EC2963B9ECAF3BD0F1D4628" DOI="https://doi.org/10.5852/ejt.2017.330" author="Inkhavilay, K" journalOrPublisher="European Journal of Taxonomy" pageId="0" pageNumber="15" pagination="1 - 40" refId="B26" refString="Inkhavilay, K, Sutcharit, C, Panha, S, 2017. Taxonomic review of the tree snail genus Amphidromus Albers, 1850 (Pulmonata: Camaenidae) in Laos, with the description of two new species. European Journal of Taxonomy 330 (330): 1 - 40, DOI: https://doi.org/10.5852/ejt.2017.330" title="Taxonomic review of the tree snail genus Amphidromus Albers, 1850 (Pulmonata: Camaenidae) in Laos, with the description of two new species." url="https://doi.org/10.5852/ejt.2017.330" volume="330" year="2017">Inkhavilay et al. (2017)</bibRefCitation>
also included the white-lipped morph from Bolikhamxay, Laos in 
<taxonomicName id="53F8D455A69BB81982DBD6DEF9C707BC" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. roseolabiatus" pageId="0" pageNumber="15" phylum="Mollusca" rank="species" species="roseolabiatus">
<emphasis id="6A5DEAC16958E300C961A1B59A98496C" italics="true" pageId="0" pageNumber="15">A. roseolabiatus</emphasis>
</taxonomicName>
. However, in this study we only incorporated the typical purplish pink-lipped morph in the phylogenetic analyses. Therefore, the identification of the white-lipped morph from Bolikhamxay, Laos as 
<taxonomicName id="F2375A98C5B7414FF99E55473354134E" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. roseolabiatus" pageId="0" pageNumber="15" phylum="Mollusca" rank="species" species="roseolabiatus">
<emphasis id="BC14AEB63B48AF0B8BAAF9920AF2C85B" italics="true" pageId="0" pageNumber="15">A. roseolabiatus</emphasis>
</taxonomicName>
or another distinct species remains to be further investigated.
</paragraph>
</subSubSection>
</treatment>
</document>