520 lines
82 KiB
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520 lines
82 KiB
XML
<document ID-DOI="10.1080/00222933.2023.2192431" ID-GBIF-Dataset="64f4ca9e-4556-45ac-8a57-00fc3a16e837" ID-ISSN="1464-5262" ID-Zenodo-Dep="7859817" IM.bibliography_approvedBy="diego" IM.illustrations_approvedBy="diego" IM.materialsCitations_approvedBy="diego" IM.metadata_approvedBy="diego" IM.tables_approvedBy="diego" IM.taxonomicNames_approvedBy="diego" IM.treatments_approvedBy="diego" checkinTime="1682060567496" checkinUser="plazi" docAuthor="Cumberlidge, Neil, Daniels, Savel R., Soma, Julia B. & Leever, Ellen M." docDate="2023" docId="503B8E36676CB147FDA0A7AFE9555CB5" docLanguage="en" docName="JNATHIST.57.5-8.463-474.pdf" docOrigin="Journal of Natural History 57 (5 - 8)" docSource="http://dx.doi.org/10.1080/00222933.2023.2192431" docStyle="DocumentStyle:3C762F9783B768A90048B82ACD135A62.3:JNatHist.2017-.journal_article.0cover" docStyleId="3C762F9783B768A90048B82ACD135A62" docStyleName="JNatHist.2017-.journal_article.0cover" docStyleVersion="3" docTitle="Vichai Cumberlidge & Daniels & Soma & Leever 2023, gen. nov." docType="treatment" docVersion="3" lastPageNumber="470" masterDocId="AC02F64E676EB140FF8DA501ED46586F" masterDocTitle="Vichai cyanapelou gen. et sp. nov. (Crustacea: Deckeniidae: Hydrothelphusinae), a new genus and new species of freshwater crab from northern Madagascar" masterLastPageNumber="474" masterPageNumber="463" pageNumber="465" updateTime="1682358530964" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Vichai cyanapelou gen. et sp. nov. (Crustacea: Deckeniidae: Hydrothelphusinae), a new genus and new species of freshwater crab from northern Madagascar</mods:title>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Cumberlidge, Neil</mods:namePart>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Daniels, Savel R.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Soma, Julia B.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Leever, Ellen M.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Journal of Natural History</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2023</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2023-04-20</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>57</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>5 - 8</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>463</mods:start>
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<mods:end>474</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.1080/00222933.2023.2192431</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.1080/00222933.2023.2192431</mods:identifier>
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<mods:identifier type="GBIF-Dataset">64f4ca9e-4556-45ac-8a57-00fc3a16e837</mods:identifier>
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<mods:identifier type="ISSN">1464-5262</mods:identifier>
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<mods:identifier type="Zenodo-Dep">7859817</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.7859819" ID-Zenodo-Dep="7859819" LSID="urn:lsid:plazi:treatment:503B8E36676CB147FDA0A7AFE9555CB5" httpUri="http://treatment.plazi.org/id/503B8E36676CB147FDA0A7AFE9555CB5" lastPageId="7" lastPageNumber="470" pageId="2" pageNumber="465">
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<subSubSection box="[557,759,686,713]" pageId="2" pageNumber="465" type="nomenclature">
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<paragraph blockId="2.[557,759,686,713]" box="[557,759,686,713]" pageId="2" pageNumber="465">
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<heading box="[557,759,686,713]" centered="true" fontSize="11" level="2" pageId="2" pageNumber="465" reason="5">
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<taxonomicName authority="Cumberlidge & Daniels & Soma & Leever, 2023" authorityName="Cumberlidge & Daniels & Soma & Leever" authorityYear="2023" box="[557,636,686,713]" class="Malacostraca" family="Potamonautidae" genus="Vichai" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="465" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis bold="true" box="[557,636,686,713]" italics="true" pageId="2" pageNumber="465">Vichai</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[644,759,686,713]" pageId="2" pageNumber="465" rank="genus">gen. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="2" pageNumber="465" type="description">
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<paragraph blockId="2.[427,888,743,770]" box="[427,888,743,770]" pageId="2" pageNumber="465">
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(
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<figureCitation box="[436,543,743,770]" captionStart="Figure 1" captionStartId="3.[160,225,1754,1777]" captionTargetBox="[172,1144,142,1722]" captionTargetId="figure-3@3.[172,1144,142,1722]" captionTargetPageId="3" captionText="Figure 1. Vichai cyanalepou sp. nov., holotype, adult J CW 38.5 mm, FMNH 7589, from Marojejy National Park,Sava region,northernMadagascar,whole animal:(a) dorsal view;(b) cephalothorax,ventral view.Scale bar: a = 10.8 mm; b = 5.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7859821" httpUri="https://zenodo.org/record/7859821/files/figure.png" pageId="2" pageNumber="465">Figures 1</figureCitation>
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(a,b), 2(a–i), 3(a–d);
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<tableCitation box="[789,880,743,770]" captionStart="Table 1" captionStartId="1.[160,216,1316,1339]" captionTargetPageId="1" captionText="Table 1. Vichai cyanalepou sp. nov. male holotype (FMNH 7589). Length in mm of the five articles (ischium, merus, carpus, propodus, dactylus) of the cheliped (P1) and the four ambulatory legs (P2–5), and the leg length in relation to the carapace width (CW). The sum of the length of each article of the four ambulatory legs divided by the carapace width is also shown (Total of 5 articles/CW), as is the sum of the relative length of the four ambulatory legs (ΣP2–5/CW = 5.35), placing this species in the short ambulatory leg size class (short: 5.0–5.9; medium: 6.0–6.9; long:>7.0)." httpUri="http://table.plazi.org/id/8CED6FA8676FB141FF2DA025EE1C5DA2" pageId="2" pageNumber="465" tableUuid="8CED6FA8676FB141FF2DA025EE1C5DA2">Table 1</tableCitation>
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)
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</paragraph>
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<paragraph blockId="2.[160,1156,832,927]" box="[160,463,832,858]" pageId="2" pageNumber="465">
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<emphasis bold="true" box="[160,463,832,858]" italics="true" pageId="2" pageNumber="465">Nomenclatural statement</emphasis>
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</paragraph>
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<paragraph blockId="2.[160,1156,832,927]" pageId="2" pageNumber="465">
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A life science identifier (LSID) number was obtained for the new genus:
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<uri pageId="2" pageNumber="465">
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urn:lsid:zoobank. org:pub:
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<uuid box="[240,735,902,927]" pageId="2" pageNumber="465">46218584-43FE-4C05-9537-FFC81EACCFDB</uuid>
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</uri>
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</paragraph>
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<paragraph blockId="2.[160,690,972,1032]" box="[160,282,972,998]" pageId="2" pageNumber="465">
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<emphasis bold="true" box="[160,282,972,998]" italics="true" pageId="2" pageNumber="465">Synonymy</emphasis>
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</paragraph>
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<paragraph blockId="2.[160,690,972,1032]" box="[160,690,1007,1032]" pageId="2" pageNumber="465">
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′New genus E̍,
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<bibRefCitation author="Cumberlidge N & Soma JB & Leever EM & Daniels SR" box="[331,617,1007,1032]" pageId="2" pageNumber="465" pagination="584 - 599" refId="ref6294" refString="Cumberlidge N, Soma JB, Leever EM, Daniels SR. 2020. New lineages within the Malagasy freshwater crab fauna: agora n. gen. for Thelphusa goudoti H. Milne Edwards, 1853, and a phytotelmic new species of Malagasya Cumberlidge & Sternberg, 2002 (Brachyura: Potamonautidae: Deckeniidae). J Crust Biol. 40 (5): 584 - 599. doi: 10.1093 / jcbiol / ruaa 050." type="journal article" year="2020">Cumberlidge et al. (2020)</bibRefCitation>
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: fig. 1.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="2" pageNumber="465" type="type_taxon">
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<paragraph blockId="2.[160,946,1077,1138]" box="[160,307,1077,1103]" pageId="2" pageNumber="465">
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<emphasis bold="true" box="[160,307,1077,1103]" italics="true" pageId="2" pageNumber="465">
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<typeStatus box="[160,216,1077,1103]" pageId="2" pageNumber="465">Type</typeStatus>
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species
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</emphasis>
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</paragraph>
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<paragraph blockId="2.[160,946,1077,1138]" box="[160,946,1113,1138]" pageId="2" pageNumber="465">
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<taxonomicName authority="Cumberlidge & Daniels & Soma & Leever, 2023" authorityName="Cumberlidge & Daniels & Soma & Leever" authorityYear="2023" box="[160,358,1113,1138]" class="Malacostraca" family="Potamonautidae" genus="Vichai" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="465" phylum="Arthropoda" rank="species" species="cyanalepou" status="sp. nov.">
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<emphasis box="[160,358,1113,1138]" italics="true" pageId="2" pageNumber="465">Vichai cyanalepou</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[366,456,1113,1138]" pageId="2" pageNumber="465" rank="species">sp. nov.</taxonomicNameLabel>
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, by present designation, gender masculine.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="5" lastPageNumber="468" pageId="2" pageNumber="465" type="diagnosis">
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<paragraph blockId="2.[160,1156,1182,1728]" box="[160,276,1182,1208]" pageId="2" pageNumber="465">
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<emphasis bold="true" box="[160,276,1182,1208]" italics="true" pageId="2" pageNumber="465">Diagnosis</emphasis>
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</paragraph>
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<paragraph blockId="2.[160,1156,1182,1728]" lastBlockId="5.[160,1155,142,479]" lastPageId="5" lastPageNumber="468" pageId="2" pageNumber="465">
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Carapace moderate height (CH/FW 1.0); front broad (CW/FW = 3.1), deflexed; surface with faint carinae in anterolateral and posterlateral regions, otherwise smooth; anterolateral margin between exorbital tooth, epibranchial tooth concave, granulated,lacking intermediate tooth; epibranchial tooth reduced to small granule, close to exorbital tooth, positioned in line with postorbital margin; lateral margin strongly convex, lined with granules, posterior end curving inward, not continuous with posterolateral margin; postfrontal crest faint, incomplete, not traversing entire carapace, epigastric crests faint, in advanced position, in line with postorbital margin, postorbital crests faint, ending before meeting epibranchial teeth; semicircular, cervical sulci deep, cervical sulcus ending before meeting postorbital crest (
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<figureCitation box="[230,318,1530,1555]" captionStart="Figure 1" captionStartId="3.[160,225,1754,1777]" captionTargetBox="[172,1144,142,1722]" captionTargetId="figure-3@3.[172,1144,142,1722]" captionTargetPageId="3" captionText="Figure 1. Vichai cyanalepou sp. nov., holotype, adult J CW 38.5 mm, FMNH 7589, from Marojejy National Park,Sava region,northernMadagascar,whole animal:(a) dorsal view;(b) cephalothorax,ventral view.Scale bar: a = 10.8 mm; b = 5.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7859821" httpUri="https://zenodo.org/record/7859821/files/figure.png" pageId="2" pageNumber="465">Figure 1</figureCitation>
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(a)). Suborbital, subhepatic regions of branchiostegite with faint granules, pterygostomial region smooth except for granules along epimeral (longitudinal) sulcus; vertical sulcus on branchiostegite curved, granular, running downward from base of epibranchial tooth to epimeral sulcus (
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<figureCitation box="[553,639,1634,1659]" captionStart="Figure 1" captionStartId="3.[160,225,1754,1777]" captionTargetBox="[172,1144,142,1722]" captionTargetId="figure-3@3.[172,1144,142,1722]" captionTargetPageId="3" captionText="Figure 1. Vichai cyanalepou sp. nov., holotype, adult J CW 38.5 mm, FMNH 7589, from Marojejy National Park,Sava region,northernMadagascar,whole animal:(a) dorsal view;(b) cephalothorax,ventral view.Scale bar: a = 10.8 mm; b = 5.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7859821" httpUri="https://zenodo.org/record/7859821/files/figure.png" pageId="2" pageNumber="465">Figure 1</figureCitation>
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(b), 2(a)). Epistomial tooth triangular, deflexed, edges with faint granules, epistomial margins smooth (
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<figureCitation box="[752,839,1669,1694]" captionStart="Figure 2" captionStartId="4.[160,225,1736,1759]" captionTargetBox="[163,1153,142,1704]" captionTargetId="figure-3@4.[163,1153,142,1704]" captionTargetPageId="4" captionText="Figure 2. Vichai cyanalepou sp. nov., holotype, adult J CW 38.5 mm, FMNH 7589, from Marojejy National Park, Sava region, northern Madagascar: (a) frontal view of carapace; (b) frontal view of right chela; (c) frontal view of left chela; (d) right cheliped merus, underside; (e) right cheliped merus, dorsal view; (f) right cheliped carpus, dorsal view; (g) third maxilliped, frontal view; (h) third maxilliped, posterior view; (i) mandible frontal view. Scale bar: a = 5.3 mm; b, c = 8.0 mm; d = 5.3 mm; e, f = 6.7 mm; g, h = 3.3 mm; i = 2.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7859825" httpUri="https://zenodo.org/record/7859825/files/figure.png" pageId="2" pageNumber="465">Figure 2</figureCitation>
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(a)); anterior lobe on terminal article of mandibular palp small (MPAL/MPTA = 0.4) (
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<figureCitation box="[760,855,1703,1728]" captionStart="Figure 2" captionStartId="4.[160,225,1736,1759]" captionTargetBox="[163,1153,142,1704]" captionTargetId="figure-3@4.[163,1153,142,1704]" captionTargetPageId="4" captionText="Figure 2. Vichai cyanalepou sp. nov., holotype, adult J CW 38.5 mm, FMNH 7589, from Marojejy National Park, Sava region, northern Madagascar: (a) frontal view of carapace; (b) frontal view of right chela; (c) frontal view of left chela; (d) right cheliped merus, underside; (e) right cheliped merus, dorsal view; (f) right cheliped carpus, dorsal view; (g) third maxilliped, frontal view; (h) third maxilliped, posterior view; (i) mandible frontal view. Scale bar: a = 5.3 mm; b, c = 8.0 mm; d = 5.3 mm; e, f = 6.7 mm; g, h = 3.3 mm; i = 2.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7859825" httpUri="https://zenodo.org/record/7859825/files/figure.png" pageId="2" pageNumber="465">Figure 2</figureCitation>
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(i)). Ambulatory legs (P2–5) stout, not elongated (P2–5/CW = 5.4) (
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<figureCitation box="[613,703,142,167]" captionStart="Figure 1" captionStartId="3.[160,225,1754,1777]" captionTargetBox="[172,1144,142,1722]" captionTargetId="figure-3@3.[172,1144,142,1722]" captionTargetPageId="3" captionText="Figure 1. Vichai cyanalepou sp. nov., holotype, adult J CW 38.5 mm, FMNH 7589, from Marojejy National Park,Sava region,northernMadagascar,whole animal:(a) dorsal view;(b) cephalothorax,ventral view.Scale bar: a = 10.8 mm; b = 5.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7859821" httpUri="https://zenodo.org/record/7859821/files/figure.png" pageId="5" pageNumber="468">Figure 1</figureCitation>
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(a);
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<tableCitation box="[752,838,142,167]" captionStart="Table 1" captionStartId="1.[160,216,1316,1339]" captionTargetPageId="1" captionText="Table 1. Vichai cyanalepou sp. nov. male holotype (FMNH 7589). Length in mm of the five articles (ischium, merus, carpus, propodus, dactylus) of the cheliped (P1) and the four ambulatory legs (P2–5), and the leg length in relation to the carapace width (CW). The sum of the length of each article of the four ambulatory legs divided by the carapace width is also shown (Total of 5 articles/CW), as is the sum of the relative length of the four ambulatory legs (ΣP2–5/CW = 5.35), placing this species in the short ambulatory leg size class (short: 5.0–5.9; medium: 6.0–6.9; long:>7.0)." httpUri="http://table.plazi.org/id/8CED6FA8676FB141FF2DA025EE1C5DA2" pageId="5" pageNumber="468" tableUuid="8CED6FA8676FB141FF2DA025EE1C5DA2">Table 1</tableCitation>
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); cheliped ischium margins smooth, rounded (
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<figureCitation box="[368,460,177,202]" captionStart="Figure 2" captionStartId="4.[160,225,1736,1759]" captionTargetBox="[163,1153,142,1704]" captionTargetId="figure-3@4.[163,1153,142,1704]" captionTargetPageId="4" captionText="Figure 2. Vichai cyanalepou sp. nov., holotype, adult J CW 38.5 mm, FMNH 7589, from Marojejy National Park, Sava region, northern Madagascar: (a) frontal view of carapace; (b) frontal view of right chela; (c) frontal view of left chela; (d) right cheliped merus, underside; (e) right cheliped merus, dorsal view; (f) right cheliped carpus, dorsal view; (g) third maxilliped, frontal view; (h) third maxilliped, posterior view; (i) mandible frontal view. Scale bar: a = 5.3 mm; b, c = 8.0 mm; d = 5.3 mm; e, f = 6.7 mm; g, h = 3.3 mm; i = 2.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7859825" httpUri="https://zenodo.org/record/7859825/files/figure.png" pageId="5" pageNumber="468">Figure 2</figureCitation>
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(d)); third maxilliped exopod with long flagellum (~0.5× merus length) (
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<figureCitation box="[258,354,212,237]" captionStart="Figure 2" captionStartId="4.[160,225,1736,1759]" captionTargetBox="[163,1153,142,1704]" captionTargetId="figure-3@4.[163,1153,142,1704]" captionTargetPageId="4" captionText="Figure 2. Vichai cyanalepou sp. nov., holotype, adult J CW 38.5 mm, FMNH 7589, from Marojejy National Park, Sava region, northern Madagascar: (a) frontal view of carapace; (b) frontal view of right chela; (c) frontal view of left chela; (d) right cheliped merus, underside; (e) right cheliped merus, dorsal view; (f) right cheliped carpus, dorsal view; (g) third maxilliped, frontal view; (h) third maxilliped, posterior view; (i) mandible frontal view. Scale bar: a = 5.3 mm; b, c = 8.0 mm; d = 5.3 mm; e, f = 6.7 mm; g, h = 3.3 mm; i = 2.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7859825" httpUri="https://zenodo.org/record/7859825/files/figure.png" pageId="5" pageNumber="468">Figure 2</figureCitation>
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(g,h)). G1TA medium length (G1TA/G1SA = 0.3), slim, basal two-thirds directed outward, distal third curving upward, tapering to blunt open tip; G1TA ventral side with faint sulcus between G1TA-G1SA junction (
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<figureCitation box="[762,855,281,306]" captionStart="Figure 3" captionStartId="6.[160,225,1101,1124]" captionTargetBox="[163,1153,142,1069]" captionTargetId="figure-190@6.[163,1153,142,1069]" captionTargetPageId="6" captionText="Figure 3. Vichai cyanalepou sp. nov., holotype, adult J CW 38.5 mm, FMNH 7589, from Marojejy National Park, Sava region, northern Madagascar: (a) right G1, ventral view; (b) right G1, dorsal view; (c) right G1, superior view; (d) right G2, ventral view. Scale bar: a, b = 1.4 mm; c = 1.1 mm; d = 1.7 mm." figureDoi="http://doi.org/10.5281/zenodo.7859827" httpUri="https://zenodo.org/record/7859827/files/figure.png" pageId="5" pageNumber="468">Figure 3</figureCitation>
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(a)); G1TA dorsal side with broad, trapezoid dorsal membrane (DM) at G1TA-G1SA junction; DM superior margin arrowhead-shaped, inferior margin U-shaped, lateral margin short, mesial margin elongated (
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<figureCitation box="[167,260,385,410]" captionStart="Figure 3" captionStartId="6.[160,225,1101,1124]" captionTargetBox="[163,1153,142,1069]" captionTargetId="figure-190@6.[163,1153,142,1069]" captionTargetPageId="6" captionText="Figure 3. Vichai cyanalepou sp. nov., holotype, adult J CW 38.5 mm, FMNH 7589, from Marojejy National Park, Sava region, northern Madagascar: (a) right G1, ventral view; (b) right G1, dorsal view; (c) right G1, superior view; (d) right G2, ventral view. Scale bar: a, b = 1.4 mm; c = 1.1 mm; d = 1.7 mm." figureDoi="http://doi.org/10.5281/zenodo.7859827" httpUri="https://zenodo.org/record/7859827/files/figure.png" pageId="5" pageNumber="468">Figure 3</figureCitation>
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(b)); G1SA lacking raised rounded shoulder on external margin near G1TA-G1SA junction (
|
||
<figureCitation box="[266,357,420,445]" captionStart="Figure 3" captionStartId="6.[160,225,1101,1124]" captionTargetBox="[163,1153,142,1069]" captionTargetId="figure-190@6.[163,1153,142,1069]" captionTargetPageId="6" captionText="Figure 3. Vichai cyanalepou sp. nov., holotype, adult J CW 38.5 mm, FMNH 7589, from Marojejy National Park, Sava region, northern Madagascar: (a) right G1, ventral view; (b) right G1, dorsal view; (c) right G1, superior view; (d) right G2, ventral view. Scale bar: a, b = 1.4 mm; c = 1.1 mm; d = 1.7 mm." figureDoi="http://doi.org/10.5281/zenodo.7859827" httpUri="https://zenodo.org/record/7859827/files/figure.png" pageId="5" pageNumber="468">Figure 3</figureCitation>
|
||
(a,b)); G2TA distinctly elongated, flagellum-like (G2TA/G2SA = 0.85) with pointed tip (
|
||
<figureCitation box="[297,389,454,479]" captionStart="Figure 3" captionStartId="6.[160,225,1101,1124]" captionTargetBox="[163,1153,142,1069]" captionTargetId="figure-190@6.[163,1153,142,1069]" captionTargetPageId="6" captionText="Figure 3. Vichai cyanalepou sp. nov., holotype, adult J CW 38.5 mm, FMNH 7589, from Marojejy National Park, Sava region, northern Madagascar: (a) right G1, ventral view; (b) right G1, dorsal view; (c) right G1, superior view; (d) right G2, ventral view. Scale bar: a, b = 1.4 mm; c = 1.1 mm; d = 1.7 mm." figureDoi="http://doi.org/10.5281/zenodo.7859827" httpUri="https://zenodo.org/record/7859827/files/figure.png" pageId="5" pageNumber="468">Figure 3</figureCitation>
|
||
(d)).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.7859821" ID-Zenodo-Dep="7859821" httpUri="https://zenodo.org/record/7859821/files/figure.png" pageId="3" pageNumber="466" startId="3.[160,225,1754,1777]" targetBox="[172,1144,142,1722]" targetPageId="3" targetType="figure">
|
||
<paragraph blockId="3.[159,1156,1754,1835]" pageId="3" pageNumber="466">
|
||
<emphasis bold="true" box="[160,251,1754,1777]" pageId="3" pageNumber="466">Figure 1.</emphasis>
|
||
<taxonomicName authority="Cumberlidge & Daniels & Soma & Leever, 2023" authorityName="Cumberlidge & Daniels & Soma & Leever" authorityYear="2023" box="[258,423,1754,1777]" class="Malacostraca" family="Potamonautidae" genus="Vichai" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="466" phylum="Arthropoda" rank="species" species="cyanalepou" status="sp. nov.">
|
||
<emphasis box="[258,423,1754,1777]" italics="true" pageId="3" pageNumber="466">Vichai cyanalepou</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[429,505,1754,1777]" pageId="3" pageNumber="466" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, holotype, adult J CW 38.5 mm, FMNH 7589, from
|
||
<taxonomicName authorityName="Cumberlidge, Boyko and Harvey" authorityYear="2002" box="[990,1071,1754,1777]" class="Malacostraca" family="Potamonautidae" genus="Marojejy" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="466" phylum="Arthropoda" rank="genus">Marojejy</taxonomicName>
|
||
National Park,Sava region, northern Madagascar, whole animal: (a) dorsal view; (b) cephalothorax,ventral view. Scale bar: a = 10.8 mm; b = 5.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.7859825" ID-Zenodo-Dep="7859825" httpUri="https://zenodo.org/record/7859825/files/figure.png" lastPageId="5" lastPageNumber="468" pageId="4" pageNumber="467" startId="4.[160,225,1736,1759]" targetBox="[163,1153,142,1704]" targetPageId="4" targetType="figure">
|
||
<paragraph blockId="4.[160,1156,1736,1788]" lastBlockId="5.[160,1157,1627,1738]" lastPageId="5" lastPageNumber="468" pageId="4" pageNumber="467">
|
||
<emphasis bold="true" box="[160,253,1736,1759]" pageId="4" pageNumber="467">Figure 2.</emphasis>
|
||
<taxonomicName authority="Cumberlidge & Daniels & Soma & Leever, 2023" authorityName="Cumberlidge & Daniels & Soma & Leever" authorityYear="2023" box="[263,440,1736,1759]" class="Malacostraca" family="Potamonautidae" genus="Vichai" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="467" phylum="Arthropoda" rank="species" species="cyanalepou" status="sp. nov.">
|
||
<emphasis box="[263,440,1736,1759]" italics="true" pageId="4" pageNumber="467">Vichai cyanalepou</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[449,532,1736,1759]" pageId="4" pageNumber="467" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, holotype, adult J CW 38.5 mm, FMNH 7589, from
|
||
<taxonomicName authorityName="Cumberlidge, Boyko and Harvey" authorityYear="2002" box="[1071,1156,1736,1759]" class="Malacostraca" family="Potamonautidae" genus="Marojejy" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="467" phylum="Arthropoda" rank="genus">Marojejy</taxonomicName>
|
||
National Park, Sava region, northern Madagascar: (a) frontal view of carapace; (b) frontal view of right chela; (c) frontal view of left chela; (d) right cheliped merus, underside; (e) right cheliped merus, dorsal view; (f) right cheliped carpus, dorsal view; (g) third maxilliped, frontal view; (h) third maxilliped, posterior view; (i) mandible frontal view. Scale bar: a =
|
||
<quantity box="[705,782,1685,1708]" metricMagnitude="-3" metricUnit="m" metricValue="5.3" pageId="5" pageNumber="468" unit="mm" value="5.3">5.3 mm</quantity>
|
||
; b, c = 8.0 mm; d =
|
||
<quantity box="[982,1058,1685,1708]" metricMagnitude="-3" metricUnit="m" metricValue="5.3" pageId="5" pageNumber="468" unit="mm" value="5.3">5.3 mm</quantity>
|
||
; e, f =
|
||
<quantity metricMagnitude="-3" metricUnit="m" metricValue="6.7" pageId="5" pageNumber="468" unit="mm" value="6.7">6.7 mm</quantity>
|
||
; g, h =
|
||
<quantity box="[279,356,1715,1738]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="5" pageNumber="468" unit="mm" value="3.3">3.3 mm</quantity>
|
||
; i =
|
||
<quantity box="[401,478,1715,1738]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="5" pageNumber="468" unit="mm" value="2.2">2.2 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection pageId="5" pageNumber="468" type="distribution">
|
||
<paragraph blockId="5.[160,1155,523,619]" box="[160,300,523,549]" pageId="5" pageNumber="468">
|
||
<emphasis bold="true" box="[160,300,523,549]" italics="true" pageId="5" pageNumber="468">Distribution</emphasis>
|
||
</paragraph>
|
||
<paragraph blockId="5.[160,1155,523,619]" pageId="5" pageNumber="468">
|
||
Endemic to
|
||
<collectingCountry box="[295,432,559,584]" name="Madagascar" pageId="5" pageNumber="468">Madagascar</collectingCountry>
|
||
.
|
||
<taxonomicName authority="Cumberlidge & Daniels & Soma & Leever, 2023" authorityName="Cumberlidge & Daniels & Soma & Leever" authorityYear="2023" box="[442,508,559,584]" class="Malacostraca" family="Potamonautidae" genus="Vichai" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="468" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis box="[442,508,559,584]" italics="true" pageId="5" pageNumber="468">Vichai</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[515,620,559,584]" pageId="5" pageNumber="468" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
is currently known only from
|
||
<taxonomicName authorityName="Cumberlidge, Boyko and Harvey" authorityYear="2002" box="[957,1054,559,584]" class="Malacostraca" family="Potamonautidae" genus="Marojejy" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="468" phylum="Arthropoda" rank="genus">Marojejy</taxonomicName>
|
||
National Park in the Sava Region in northern
|
||
<collectingCountry box="[568,705,594,619]" name="Madagascar" pageId="5" pageNumber="468">Madagascar</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="5" pageNumber="468" type="etymology">
|
||
<paragraph blockId="5.[160,1156,663,931]" box="[160,285,663,689]" pageId="5" pageNumber="468">
|
||
<emphasis bold="true" box="[160,285,663,689]" italics="true" pageId="5" pageNumber="468">Etymology</emphasis>
|
||
</paragraph>
|
||
<paragraph blockId="5.[160,1156,663,931]" pageId="5" pageNumber="468">
|
||
The genus is named in honour of
|
||
<taxonomicName authority="Srivaddhanaprabha" authorityName="Srivaddhanaprabha" box="[558,857,698,723]" class="Malacostraca" family="Potamonautidae" genus="Vichai" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="468" phylum="Arthropoda" rank="genus">Vichai Srivaddhanaprabha</taxonomicName>
|
||
, the owner of the English Premier League football club Leicester City (LCFC) from 2010 until his death in a tragic helicopter crash at the King Power Stadium in
|
||
<date box="[737,902,768,793]" pageId="5" pageNumber="468" value="2018-10">October 2018</date>
|
||
.
|
||
<taxonomicName box="[917,986,768,793]" class="Malacostraca" family="Potamonautidae" genus="Vichai" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="473" phylum="Arthropoda" rank="genus">Vichai</taxonomicName>
|
||
was not only extremely mindful of the Leicester community and the LCFC supporters, but under his leadership the Football Club won the 2015–16 English Premier League title against all expectations, after starting the season as 5000/1 rank outsiders. The generic name,
|
||
<taxonomicName box="[1083,1153,872,897]" class="Malacostraca" family="Potamonautidae" genus="Vichai" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="473" phylum="Arthropoda" rank="genus">Vichai</taxonomicName>
|
||
, is used as a Latin noun in nominative singular and treated as masculine.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="7" lastPageNumber="470" pageId="5" pageNumber="468" type="discussion">
|
||
<paragraph blockId="5.[160,1156,976,1556]" box="[160,262,976,1002]" pageId="5" pageNumber="468">
|
||
<emphasis bold="true" box="[160,262,976,1002]" italics="true" pageId="5" pageNumber="468">Remarks</emphasis>
|
||
</paragraph>
|
||
<paragraph blockId="5.[160,1156,976,1556]" lastBlockId="6.[160,1156,1247,1723]" lastPageId="6" lastPageNumber="469" pageId="5" pageNumber="468">
|
||
The genus
|
||
<taxonomicName box="[285,351,1011,1036]" class="Malacostraca" family="Potamonautidae" genus="Vichai" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="468" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[285,351,1011,1036]" italics="true" pageId="5" pageNumber="468">Vichai</emphasis>
|
||
</taxonomicName>
|
||
is established for a new species,
|
||
<taxonomicName authorityName="Cumberlidge & Daniels & Soma & Leever" authorityYear="2023" box="[729,881,1011,1036]" class="Malacostraca" family="Potamonautidae" genus="Vichai" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="468" phylum="Arthropoda" rank="species" species="cyanalepou">
|
||
<emphasis box="[729,881,1011,1036]" italics="true" pageId="5" pageNumber="468">V. cyanalepou</emphasis>
|
||
</taxonomicName>
|
||
(see later). The recognition of the present new genus is based on phylogenetic and morphological evidence. The molecular phylogeny in
|
||
<bibRefCitation author="Cumberlidge N & Soma JB & Leever EM & Daniels SR" box="[436,715,1081,1106]" pageId="5" pageNumber="468" pagination="584 - 599" refId="ref6294" refString="Cumberlidge N, Soma JB, Leever EM, Daniels SR. 2020. New lineages within the Malagasy freshwater crab fauna: agora n. gen. for Thelphusa goudoti H. Milne Edwards, 1853, and a phytotelmic new species of Malagasya Cumberlidge & Sternberg, 2002 (Brachyura: Potamonautidae: Deckeniidae). J Crust Biol. 40 (5): 584 - 599. doi: 10.1093 / jcbiol / ruaa 050." type="journal article" year="2020">Cumberlidge et al. (2020</bibRefCitation>
|
||
, fig. 1, ′New genus E̍) indicates that
|
||
<taxonomicName authority="Cumberlidge & Daniels & Soma & Leever, 2023" authorityName="Cumberlidge & Daniels & Soma & Leever" authorityYear="2023" class="Malacostraca" family="Potamonautidae" genus="Vichai" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="468" phylum="Arthropoda" rank="species" species="cyanalepou" status="sp. nov.">
|
||
<emphasis italics="true" pageId="5" pageNumber="468">V. cyanalepou</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[291,377,1115,1140]" pageId="5" pageNumber="468" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is part of a unique basal clade that is clearly genetically separate from all other Malagasy genera. Mophologically,
|
||
<taxonomicName authority="Cumberlidge & Daniels & Soma & Leever, 2023" authorityName="Cumberlidge & Daniels & Soma & Leever" authorityYear="2023" box="[641,707,1150,1175]" class="Malacostraca" family="Potamonautidae" genus="Vichai" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="468" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis box="[641,707,1150,1175]" italics="true" pageId="5" pageNumber="468">Vichai</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[713,817,1150,1175]" pageId="5" pageNumber="468" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
can be distinguished from the three other genera found in the
|
||
<taxonomicName authorityName="Cumberlidge, Boyko and Harvey" authorityYear="2002" box="[526,623,1185,1210]" class="Malacostraca" family="Potamonautidae" genus="Marojejy" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="468" phylum="Arthropoda" rank="genus">Marojejy</taxonomicName>
|
||
National Park (i.e.
|
||
<taxonomicName authority="Reed and Cumberlidge, 2006" authorityName="Reed and Cumberlidge" authorityYear="2006" class="Malacostraca" family="Potamonautidae" genus="Foza" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="468" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[832,883,1185,1210]" italics="true" pageId="5" pageNumber="468">Foza</emphasis>
|
||
<bibRefCitation author="Reed SK & Cumberlidge N" pageId="5" pageNumber="468" pagination="58 - 66" refId="ref6666" refString="Reed SK, Cumberlidge N. 2006. Foza raimundi, a new genus and species of potamonautid freshwater crab (Crustacea: Decapoda: Potamoidea) from northern Madagascar. Proc Biol Soc Wash. 119 (1): 58 - 66. doi: 10.2988 / 0006 - 324 X (2006) 119 [58: FRANGA] 2.0. CO; 2." type="journal article" year="2006">Reed and Cumberlidge, 2006</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authority="A. Milne-Edwards, 1872" authorityName="A. Milne-Edwards" authorityYear="1872" box="[232,693,1219,1244]" class="Malacostraca" family="Potamonautidae" genus="Hydrothelphusa" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="468" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[232,405,1219,1244]" italics="true" pageId="5" pageNumber="468">Hydrothelphusa</emphasis>
|
||
A. Milne-Edwards, 1872
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName authority="Cumberlidge, Boyko and Harvey, 2002" authorityName="Cumberlidge, Boyko and Harvey" authorityYear="2002" class="Malacostraca" family="Potamonautidae" genus="Marojejy" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="468" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[762,856,1219,1244]" italics="true" pageId="5" pageNumber="468">Marojejy</emphasis>
|
||
<bibRefCitation author="Cumberlidge N & Boyko CB & Harvey AW" pageId="5" pageNumber="468" pagination="65 - 77" refId="ref6025" refString="Cumberlidge N, Boyko CB, Harvey AW. 2002. A new genus and species of freshwater crab (Decapoda, Crustacea, Potamoidea) from northern Madagascar, and a second new species associated with Pandanus leaf axils. J Nat Hist. 36 (1): 65 - 77. doi: 10.1080 / 00222930010003800." type="journal article" year="2002">Cumberlidge, Boyko and Harvey, 2002</bibRefCitation>
|
||
</taxonomicName>
|
||
) as follows. The G1TA of
|
||
<taxonomicName authority="Cumberlidge & Daniels & Soma & Leever, 2023" authorityName="Cumberlidge & Daniels & Soma & Leever" authorityYear="2023" box="[625,692,1254,1279]" class="Malacostraca" family="Potamonautidae" genus="Vichai" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="468" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis box="[625,692,1254,1279]" italics="true" pageId="5" pageNumber="468">Vichai</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[703,813,1254,1279]" pageId="5" pageNumber="468" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
is medium in length (G1TA/ G1SA = 0.3) and slim where the basal two-thirds are directed outward and the distal third curves upward (
|
||
<figureCitation box="[397,487,1323,1348]" captionStart="Figure 3" captionStartId="6.[160,225,1101,1124]" captionTargetBox="[163,1153,142,1069]" captionTargetId="figure-190@6.[163,1153,142,1069]" captionTargetPageId="6" captionText="Figure 3. Vichai cyanalepou sp. nov., holotype, adult J CW 38.5 mm, FMNH 7589, from Marojejy National Park, Sava region, northern Madagascar: (a) right G1, ventral view; (b) right G1, dorsal view; (c) right G1, superior view; (d) right G2, ventral view. Scale bar: a, b = 1.4 mm; c = 1.1 mm; d = 1.7 mm." figureDoi="http://doi.org/10.5281/zenodo.7859827" httpUri="https://zenodo.org/record/7859827/files/figure.png" pageId="5" pageNumber="468">Figure 3</figureCitation>
|
||
(a–c)) vs a G1TA that is short (G1 TA/ SA = 0.25), straight, and broadly conical in
|
||
<taxonomicName authorityName="Reed and Cumberlidge" authorityYear="2006" box="[368,419,1358,1383]" class="Malacostraca" family="Potamonautidae" genus="Foza" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="468" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[368,419,1358,1383]" italics="true" pageId="5" pageNumber="468">Foza</emphasis>
|
||
</taxonomicName>
|
||
(see
|
||
<bibRefCitation author="Leever EM & Daniels SR & Soma JB & Cumberlidge N" box="[480,679,1358,1383]" pageId="5" pageNumber="468" pagination="241 - 263" refId="ref6477" refString="Leever EM, Daniels SR, Soma JB, Cumberlidge N. 2022. Two new genera and a new species of freshwater crabs from northern Madagascar: vahatra gen. nov. for Foza ambohitra Cumberlidge and Meyer, 2009, and Toamasina gen. nov. for Toamasina clarki sp. nov. (Brachyura: Potamoidea: Deckeniidae). J Nat Hist. 56 (1 - 4): 241 - 263. doi: 10.1080 / 00222933.2022.2049389." type="journal article" year="2022">Leever et al. 2022</bibRefCitation>
|
||
, fig.
|
||
<quantity box="[734,772,1358,1383]" metricMagnitude="-3" metricUnit="kg" metricValue="7.0" pageId="5" pageNumber="468" unit="g" value="7.0">7 g</quantity>
|
||
,h). The exorbital and epibranchial teeth in
|
||
<taxonomicName authority="Cumberlidge & Daniels & Soma & Leever, 2023" authorityName="Cumberlidge & Daniels & Soma & Leever" authorityYear="2023" box="[259,325,1393,1418]" class="Malacostraca" family="Potamonautidae" genus="Vichai" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="468" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis box="[259,325,1393,1418]" italics="true" pageId="5" pageNumber="468">Vichai</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[334,440,1393,1418]" pageId="5" pageNumber="468" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
are separated by a shallow notch (
|
||
<figureCitation box="[848,937,1393,1418]" captionStart="Figure 1" captionStartId="3.[160,225,1754,1777]" captionTargetBox="[172,1144,142,1722]" captionTargetId="figure-3@3.[172,1144,142,1722]" captionTargetPageId="3" captionText="Figure 1. Vichai cyanalepou sp. nov., holotype, adult J CW 38.5 mm, FMNH 7589, from Marojejy National Park,Sava region,northernMadagascar,whole animal:(a) dorsal view;(b) cephalothorax,ventral view.Scale bar: a = 10.8 mm; b = 5.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7859821" httpUri="https://zenodo.org/record/7859821/files/figure.png" pageId="5" pageNumber="468">Figure 1</figureCitation>
|
||
(a)), whereas these teeth are separated by a deep cleft in
|
||
<taxonomicName authority="A. Milne-Edwards, 1872" authorityName="A. Milne-Edwards" authorityYear="1872" box="[634,1098,1427,1452]" class="Malacostraca" family="Potamonautidae" genus="Hydrothelphusa" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="468" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[634,809,1427,1452]" italics="true" pageId="5" pageNumber="468">Hydrothelphusa</emphasis>
|
||
A. Milne-Edwards, 1872
|
||
</taxonomicName>
|
||
(see
|
||
<bibRefCitation author="Cumberlidge N & Sternberg R" box="[160,541,1462,1487]" pageId="5" pageNumber="468" pagination="41 - 79" refId="ref6374" refString="Cumberlidge N, Sternberg R. 2002. The freshwater crabs of Madagascar (Crustacea, Decapoda, Potamoidea). Zoosystema. 24: 41 - 79." type="journal article" year="2002">Cumberlidge and Sternberg 2002</bibRefCitation>
|
||
, fig. 1a–c;
|
||
<bibRefCitation author="Cumberlidge N & Marijnissen SAE & Thompson J" box="[664,938,1462,1487]" pageId="5" pageNumber="468" pagination="61 - 68" refId="ref6140" refString="Cumberlidge N, Marijnissen SAE, Thompson J. 2007. Hydrothelphusa vencesi, a new species of freshwater crab (Brachyura: Potamoidea: Potamonautidae) from southeastern Madagascar. Zootaxa. 1524 (1): 61 - 68. doi: 10.11646 / zootaxa. 1524.1.6." type="journal article" year="2007">Cumberlidge et al. 2007</bibRefCitation>
|
||
, figs 1, 16). Finally, the ambulatory legs of
|
||
<taxonomicName authority="Cumberlidge & Daniels & Soma & Leever, 2023" authorityName="Cumberlidge & Daniels & Soma & Leever" authorityYear="2023" box="[420,486,1497,1522]" class="Malacostraca" family="Potamonautidae" genus="Vichai" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="468" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis box="[420,486,1497,1522]" italics="true" pageId="5" pageNumber="468">Vichai</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[493,597,1497,1522]" pageId="5" pageNumber="468" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
are short and stout (ΣP2–5/CW = 5.4) (
|
||
<figureCitation box="[1030,1117,1497,1522]" captionStart="Figure 1" captionStartId="3.[160,225,1754,1777]" captionTargetBox="[172,1144,142,1722]" captionTargetId="figure-3@3.[172,1144,142,1722]" captionTargetPageId="3" captionText="Figure 1. Vichai cyanalepou sp. nov., holotype, adult J CW 38.5 mm, FMNH 7589, from Marojejy National Park,Sava region,northernMadagascar,whole animal:(a) dorsal view;(b) cephalothorax,ventral view.Scale bar: a = 10.8 mm; b = 5.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7859821" httpUri="https://zenodo.org/record/7859821/files/figure.png" pageId="5" pageNumber="468">Figure 1</figureCitation>
|
||
(a);
|
||
<tableCitation box="[160,240,1531,1556]" captionStart="Table 1" captionStartId="1.[160,216,1316,1339]" captionTargetPageId="1" captionText="Table 1. Vichai cyanalepou sp. nov. male holotype (FMNH 7589). Length in mm of the five articles (ischium, merus, carpus, propodus, dactylus) of the cheliped (P1) and the four ambulatory legs (P2–5), and the leg length in relation to the carapace width (CW). The sum of the length of each article of the four ambulatory legs divided by the carapace width is also shown (Total of 5 articles/CW), as is the sum of the relative length of the four ambulatory legs (ΣP2–5/CW = 5.35), placing this species in the short ambulatory leg size class (short: 5.0–5.9; medium: 6.0–6.9; long:>7.0)." httpUri="http://table.plazi.org/id/8CED6FA8676FB141FF2DA025EE1C5DA2" pageId="5" pageNumber="468" tableUuid="8CED6FA8676FB141FF2DA025EE1C5DA2">Table 1</tableCitation>
|
||
) and its eyestalks and corneas are of normal length (
|
||
<figureCitation box="[819,915,1531,1556]" captionStart="Figure 1" captionStartId="3.[160,225,1754,1777]" captionTargetBox="[172,1144,142,1722]" captionTargetId="figure-3@3.[172,1144,142,1722]" captionTargetPageId="3" captionText="Figure 1. Vichai cyanalepou sp. nov., holotype, adult J CW 38.5 mm, FMNH 7589, from Marojejy National Park,Sava region,northernMadagascar,whole animal:(a) dorsal view;(b) cephalothorax,ventral view.Scale bar: a = 10.8 mm; b = 5.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7859821" httpUri="https://zenodo.org/record/7859821/files/figure.png" pageId="5" pageNumber="468">Figures 1</figureCitation>
|
||
(a), 2(a)), whereas the ambulatory legs of
|
||
<taxonomicName authority="Cumberlidge et al. (2002)" authorityName="Cumberlidge" authorityYear="2002" box="[377,761,1247,1272]" class="Malacostraca" family="Potamonautidae" genus="Marojejy" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="469" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[377,470,1247,1272]" italics="true" pageId="6" pageNumber="469">Marojejy</emphasis>
|
||
Cumberlidge et al. (2002)
|
||
</taxonomicName>
|
||
are elongated and slender (ΣP2–5/ CW = 7.0) and its eyestalks are short, taper distally, and have a reduced cornea (see Cumberlidge et al. 2002, fig. 2f).
|
||
</paragraph>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.7859827" ID-Zenodo-Dep="7859827" httpUri="https://zenodo.org/record/7859827/files/figure.png" pageId="6" pageNumber="469" startId="6.[160,225,1101,1124]" targetBox="[163,1153,142,1069]" targetPageId="6" targetType="figure">
|
||
<paragraph blockId="6.[160,1157,1101,1182]" pageId="6" pageNumber="469">
|
||
<emphasis bold="true" box="[160,253,1101,1124]" pageId="6" pageNumber="469">Figure 3.</emphasis>
|
||
<taxonomicName authority="Cumberlidge & Daniels & Soma & Leever, 2023" authorityName="Cumberlidge & Daniels & Soma & Leever" authorityYear="2023" box="[263,440,1101,1124]" class="Malacostraca" family="Potamonautidae" genus="Vichai" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="469" phylum="Arthropoda" rank="species" species="cyanalepou" status="sp. nov.">
|
||
<emphasis box="[263,440,1101,1124]" italics="true" pageId="6" pageNumber="469">Vichai cyanalepou</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[449,532,1101,1124]" pageId="6" pageNumber="469" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, holotype, adult J CW 38.5 mm, FMNH 7589, from
|
||
<taxonomicName authorityName="Cumberlidge, Boyko and Harvey" authorityYear="2002" box="[1071,1156,1101,1124]" class="Malacostraca" family="Potamonautidae" genus="Marojejy" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="469" phylum="Arthropoda" rank="genus">Marojejy</taxonomicName>
|
||
National Park, Sava region, northern Madagascar: (a) right G1, ventral view; (b) right G1, dorsal view; (c) right G1, superior view; (d) right G2, ventral view. Scale bar: a, b = 1.4 mm; c = 1.1 mm; d = 1.7 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph blockId="6.[160,1156,1247,1723]" pageId="6" pageNumber="469">
|
||
The medium-sized anterior lobe on the terminal article of the mandibular palp of
|
||
<taxonomicName authority="Cumberlidge & Daniels & Soma & Leever, 2023" authorityName="Cumberlidge & Daniels & Soma & Leever" authorityYear="2023" box="[1089,1155,1351,1376]" class="Malacostraca" family="Potamonautidae" genus="Vichai" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="469" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis box="[1089,1155,1351,1376]" italics="true" pageId="6" pageNumber="469">Vichai</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[160,269,1386,1411]" pageId="6" pageNumber="469" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
(MPAL/MPTA = 0.4) (
|
||
<figureCitation box="[523,623,1386,1411]" captionStart="Figure 2" captionStartId="4.[160,225,1736,1759]" captionTargetBox="[163,1153,142,1704]" captionTargetId="figure-3@4.[163,1153,142,1704]" captionTargetPageId="4" captionText="Figure 2. Vichai cyanalepou sp. nov., holotype, adult J CW 38.5 mm, FMNH 7589, from Marojejy National Park, Sava region, northern Madagascar: (a) frontal view of carapace; (b) frontal view of right chela; (c) frontal view of left chela; (d) right cheliped merus, underside; (e) right cheliped merus, dorsal view; (f) right cheliped carpus, dorsal view; (g) third maxilliped, frontal view; (h) third maxilliped, posterior view; (i) mandible frontal view. Scale bar: a = 5.3 mm; b, c = 8.0 mm; d = 5.3 mm; e, f = 6.7 mm; g, h = 3.3 mm; i = 2.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7859825" httpUri="https://zenodo.org/record/7859825/files/figure.png" pageId="6" pageNumber="469">Figure 2</figureCitation>
|
||
(i)) distinguishes it from three other Malagasy genera:
|
||
<taxonomicName authority="(Cumberlidge, 2010)" baseAuthorityName="Cumberlidge" baseAuthorityYear="2010" box="[268,715,1421,1446]" class="Malacostraca" family="Potamonautidae" genus="Boreathelphusa" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="469" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[268,442,1421,1446]" italics="true" pageId="6" pageNumber="469">Boreathelphusa</emphasis>
|
||
(
|
||
<bibRefCitation author="Cumberlidge N" box="[470,706,1421,1446]" pageId="6" pageNumber="469" pagination="68" refId="ref5974" refString="Cumberlidge N. 2010. Boreathelphusa, a replacement name for Boreas Cumberlidge and Sternberg, 2002 (Crustacea: Brachyura: Potamonautidae), preoccupied by Boreas Morris, 19870 (Bryozoa: Hippothoidae). Zootaxa. 2450: 68. doi: 10.11646 / zootaxa. 2450.1.7." type="journal article" year="2010">Cumberlidge, 2010</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authority="Bott, 1955" authorityName="Bott" authorityYear="1955" box="[741,1090,1421,1446]" class="Malacostraca" family="Potamonautidae" genus="Madagapotamon" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="469" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[741,936,1421,1446]" italics="true" pageId="6" pageNumber="469">Madagapotamon</emphasis>
|
||
Bott, 1955
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName authority="Ng and Takeda (1994)" authorityName="Ng and Takeda" authorityYear="1994" box="[160,573,1455,1480]" class="Malacostraca" family="Potamonautidae" genus="Skelosophusa" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="469" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[160,307,1455,1480]" italics="true" pageId="6" pageNumber="469">Skelosophusa</emphasis>
|
||
<bibRefCitation author="Ng PKL & Takeda M" box="[317,573,1455,1480]" pageId="6" pageNumber="469" pagination="161 - 172" refId="ref6616" refString="Ng PKL, Takeda M. 1994. Skelosophusa (Crustacea, Decapoda, Brachyura), a new genus of potamonautid freshwater crab from Madagascar, with descriptions of two new species. Bull Nat Sci Mus Ser A (Zool). 20: 161 - 172." type="journal article" year="1994">Ng and Takeda (1994)</bibRefCitation>
|
||
</taxonomicName>
|
||
. In all three of these genera, the anterior lobe on the mandibular palp is noticeably small (MPAL/MPTA = 0.2), and ledge-like (see
|
||
<bibRefCitation author="Cumberlidge N & Sternberg R" box="[160,548,1525,1550]" pageId="6" pageNumber="469" pagination="41 - 79" refId="ref6374" refString="Cumberlidge N, Sternberg R. 2002. The freshwater crabs of Madagascar (Crustacea, Decapoda, Potamoidea). Zoosystema. 24: 41 - 79." type="journal article" year="2002">Cumberlidge and Sternberg 2002</bibRefCitation>
|
||
, fig. 4 h–l) rather than a broad rounded lobe as in
|
||
<taxonomicName authority="Cumberlidge & Daniels & Soma & Leever, 2023" authorityName="Cumberlidge & Daniels & Soma & Leever" authorityYear="2023" box="[160,226,1559,1584]" class="Malacostraca" family="Potamonautidae" genus="Vichai" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="469" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis box="[160,226,1559,1584]" italics="true" pageId="6" pageNumber="469">Vichai</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[233,338,1559,1584]" pageId="6" pageNumber="469" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
(
|
||
<figureCitation box="[352,448,1559,1584]" captionStart="Figure 2" captionStartId="4.[160,225,1736,1759]" captionTargetBox="[163,1153,142,1704]" captionTargetId="figure-3@4.[163,1153,142,1704]" captionTargetPageId="4" captionText="Figure 2. Vichai cyanalepou sp. nov., holotype, adult J CW 38.5 mm, FMNH 7589, from Marojejy National Park, Sava region, northern Madagascar: (a) frontal view of carapace; (b) frontal view of right chela; (c) frontal view of left chela; (d) right cheliped merus, underside; (e) right cheliped merus, dorsal view; (f) right cheliped carpus, dorsal view; (g) third maxilliped, frontal view; (h) third maxilliped, posterior view; (i) mandible frontal view. Scale bar: a = 5.3 mm; b, c = 8.0 mm; d = 5.3 mm; e, f = 6.7 mm; g, h = 3.3 mm; i = 2.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7859825" httpUri="https://zenodo.org/record/7859825/files/figure.png" pageId="6" pageNumber="469">Figure 2</figureCitation>
|
||
(i)).
|
||
</paragraph>
|
||
<paragraph blockId="6.[160,1156,1247,1723]" lastBlockId="7.[160,1156,142,1242]" lastPageId="7" lastPageNumber="470" pageId="6" pageNumber="469">
|
||
<taxonomicName authority="Cumberlidge & Daniels & Soma & Leever, 2023" authorityName="Cumberlidge & Daniels & Soma & Leever" authorityYear="2023" box="[192,258,1594,1619]" class="Malacostraca" family="Potamonautidae" genus="Vichai" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="469" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis box="[192,258,1594,1619]" italics="true" pageId="6" pageNumber="469">Vichai</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[265,370,1594,1619]" pageId="6" pageNumber="469" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
can be distinguished from the remaining Malagasy genera as follows. The postfrontal crest of
|
||
<taxonomicName authority="Cumberlidge & Daniels & Soma & Leever, 2023" authorityName="Cumberlidge & Daniels & Soma & Leever" authorityYear="2023" box="[426,492,1629,1654]" class="Malacostraca" family="Potamonautidae" genus="Vichai" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="469" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis box="[426,492,1629,1654]" italics="true" pageId="6" pageNumber="469">Vichai</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[498,602,1629,1654]" pageId="6" pageNumber="469" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
is incomplete and faint and does not traverse the entire carapace (
|
||
<figureCitation box="[345,431,1663,1688]" captionStart="Figure 1" captionStartId="3.[160,225,1754,1777]" captionTargetBox="[172,1144,142,1722]" captionTargetId="figure-3@3.[172,1144,142,1722]" captionTargetPageId="3" captionText="Figure 1. Vichai cyanalepou sp. nov., holotype, adult J CW 38.5 mm, FMNH 7589, from Marojejy National Park,Sava region,northernMadagascar,whole animal:(a) dorsal view;(b) cephalothorax,ventral view.Scale bar: a = 10.8 mm; b = 5.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7859821" httpUri="https://zenodo.org/record/7859821/files/figure.png" pageId="6" pageNumber="469">Figure 1</figureCitation>
|
||
(a)), whereas the postfrontal crest is well defined and completely traverses the carapace in
|
||
<taxonomicName authorityName="Cumberlidge, Soma, Leever & Daniels" authorityYear="2020" box="[441,506,1698,1723]" class="Malacostraca" family="Potamonautidae" genus="Agora" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="469" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[441,506,1698,1723]" italics="true" pageId="6" pageNumber="469">Agora</emphasis>
|
||
</taxonomicName>
|
||
(see
|
||
<bibRefCitation author="Cumberlidge N & Soma JB & Leever EM & Daniels SR" box="[564,829,1698,1723]" pageId="6" pageNumber="469" pagination="584 - 599" refId="ref6294" refString="Cumberlidge N, Soma JB, Leever EM, Daniels SR. 2020. New lineages within the Malagasy freshwater crab fauna: agora n. gen. for Thelphusa goudoti H. Milne Edwards, 1853, and a phytotelmic new species of Malagasya Cumberlidge & Sternberg, 2002 (Brachyura: Potamonautidae: Deckeniidae). J Crust Biol. 40 (5): 584 - 599. doi: 10.1093 / jcbiol / ruaa 050." type="journal article" year="2020">Cumberlidge et al. 2020</bibRefCitation>
|
||
). The anterolateral regions of the carapace surface of
|
||
<taxonomicName authority="Cumberlidge & Daniels & Soma & Leever, 2023" authorityName="Cumberlidge & Daniels & Soma & Leever" authorityYear="2023" box="[440,506,142,167]" class="Malacostraca" family="Potamonautidae" genus="Vichai" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="470" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis box="[440,506,142,167]" italics="true" pageId="7" pageNumber="470">Vichai</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[516,624,142,167]" pageId="7" pageNumber="470" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
are mostly smooth and only faintly carinated (
|
||
<figureCitation box="[168,259,177,202]" captionStart="Figure 1" captionStartId="3.[160,225,1754,1777]" captionTargetBox="[172,1144,142,1722]" captionTargetId="figure-3@3.[172,1144,142,1722]" captionTargetPageId="3" captionText="Figure 1. Vichai cyanalepou sp. nov., holotype, adult J CW 38.5 mm, FMNH 7589, from Marojejy National Park,Sava region,northernMadagascar,whole animal:(a) dorsal view;(b) cephalothorax,ventral view.Scale bar: a = 10.8 mm; b = 5.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7859821" httpUri="https://zenodo.org/record/7859821/files/figure.png" pageId="7" pageNumber="470">Figure 1</figureCitation>
|
||
(a)), vs anterolateral regions of the carapace that have fields of well-defined carinae in
|
||
<taxonomicName authorityName="Cumberlidge, Soma, Leever, Clark & Daniels" authorityYear="2021" box="[274,394,212,237]" class="Malacostraca" family="Potamonautidae" genus="Crosnautes" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="470" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[274,394,212,237]" italics="true" pageId="7" pageNumber="470">Crosnautes</emphasis>
|
||
</taxonomicName>
|
||
(see
|
||
<bibRefCitation author="Cumberlidge N & Soma J & Leever EM & Clark PF & Daniels SR" box="[452,718,212,237]" pageId="7" pageNumber="470" pagination="1 - 10" refId="ref6232" refString="Cumberlidge N, Soma J, Leever EM, Clark PF, Daniels SR. 2021. Description of a new genus and two new species of freshwater crabs (Decapoda: Brachyura: Potamonautidae) from southeastern Madagascar. J Crust Biol. 41 (3): 1 - 10. doi: 10.1093 / jcbiol / ruab 034." type="journal article" year="2021">Cumberlidge et al. 2021</bibRefCitation>
|
||
, fig. 1a,b). The G1TA of
|
||
<taxonomicName authority="Cumberlidge & Daniels & Soma & Leever, 2023" authorityName="Cumberlidge & Daniels & Soma & Leever" authorityYear="2023" box="[980,1046,212,237]" class="Malacostraca" family="Potamonautidae" genus="Vichai" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="470" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis box="[980,1046,212,237]" italics="true" pageId="7" pageNumber="470">Vichai</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1053,1156,212,237]" pageId="7" pageNumber="470" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
is slim, tapers evenly, and curves upward distally (
|
||
<figureCitation box="[710,802,246,271]" captionStart="Figure 3" captionStartId="6.[160,225,1101,1124]" captionTargetBox="[163,1153,142,1069]" captionTargetId="figure-190@6.[163,1153,142,1069]" captionTargetPageId="6" captionText="Figure 3. Vichai cyanalepou sp. nov., holotype, adult J CW 38.5 mm, FMNH 7589, from Marojejy National Park, Sava region, northern Madagascar: (a) right G1, ventral view; (b) right G1, dorsal view; (c) right G1, superior view; (d) right G2, ventral view. Scale bar: a, b = 1.4 mm; c = 1.1 mm; d = 1.7 mm." figureDoi="http://doi.org/10.5281/zenodo.7859827" httpUri="https://zenodo.org/record/7859827/files/figure.png" pageId="7" pageNumber="470">Figure 3</figureCitation>
|
||
(a,b)), vs a G1TA that is broadest at the midpoint and is straight along its length in
|
||
<taxonomicName authority="Meyer et al. (2014)" authorityName="Meyer" authorityYear="2014" box="[718,1104,281,306]" class="Malacostraca" family="Potamonautidae" genus="Glabrithelphusa" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="470" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[718,890,281,306]" italics="true" pageId="7" pageNumber="470">Glabrithelphusa</emphasis>
|
||
<bibRefCitation author="Meyer KS & Cumberlidge N & Koppin JC" box="[897,1104,281,306]" pageId="7" pageNumber="470" pagination="65 - 72" refId="ref6561" refString="Meyer KS, Cumberlidge N, Koppin JC. 2014. A new genus and species of freshwater crab from Madagascar (Decapoda, Brachyura, Potamoidea, Potamonautidae). Potamoidea, Potamonautidae). Zootaxa. 3884 (1): 65 - 72. doi: 10.11646 / zootaxa. 3884.1.5." type="journal article" year="2014">Meyer et al. (2014)</bibRefCitation>
|
||
</taxonomicName>
|
||
(see
|
||
<bibRefCitation author="Meyer KS & Cumberlidge N & Koppin JC" box="[160,351,316,341]" pageId="7" pageNumber="470" pagination="65 - 72" refId="ref6561" refString="Meyer KS, Cumberlidge N, Koppin JC. 2014. A new genus and species of freshwater crab from Madagascar (Decapoda, Brachyura, Potamoidea, Potamonautidae). Potamoidea, Potamonautidae). Zootaxa. 3884 (1): 65 - 72. doi: 10.11646 / zootaxa. 3884.1.5." type="journal article" year="2014">Meyer et al. 2014</bibRefCitation>
|
||
, figs 1a, 3a). The lateral carapace margin of
|
||
<taxonomicName authority="Cumberlidge & Daniels & Soma & Leever, 2023" authorityName="Cumberlidge & Daniels & Soma & Leever" authorityYear="2023" box="[838,904,316,341]" class="Malacostraca" family="Potamonautidae" genus="Vichai" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="470" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis box="[838,904,316,341]" italics="true" pageId="7" pageNumber="470">Vichai</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[911,1015,316,341]" pageId="7" pageNumber="470" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
is lined with small granules (Figure (1a)), vs a lateral margin that is lined with either small or medium-sized teeth in
|
||
<taxonomicName authority="Cumberlidge and Sternberg (2002)" authorityName="Cumberlidge and Sternberg" authorityYear="2002" box="[336,882,385,410]" class="Malacostraca" family="Potamonautidae" genus="Malagasya" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="470" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[336,457,385,410]" italics="true" pageId="7" pageNumber="470">Malagasya</emphasis>
|
||
<bibRefCitation author="Cumberlidge N & Sternberg R" box="[469,882,385,410]" pageId="7" pageNumber="470" pagination="41 - 79" refId="ref6374" refString="Cumberlidge N, Sternberg R. 2002. The freshwater crabs of Madagascar (Crustacea, Decapoda, Potamoidea). Zoosystema. 24: 41 - 79." type="journal article" year="2002">Cumberlidge and Sternberg (2002)</bibRefCitation>
|
||
</taxonomicName>
|
||
(see
|
||
<bibRefCitation author="Cumberlidge N & Sternberg R" pageId="7" pageNumber="470" pagination="41 - 79" refId="ref6374" refString="Cumberlidge N, Sternberg R. 2002. The freshwater crabs of Madagascar (Crustacea, Decapoda, Potamoidea). Zoosystema. 24: 41 - 79." type="journal article" year="2002">Cumberlidge and Sternberg 2002</bibRefCitation>
|
||
, fig. 1e,f;
|
||
<bibRefCitation author="Cumberlidge N & Soma JB & Leever EM & Daniels SR" box="[441,711,420,445]" pageId="7" pageNumber="470" pagination="584 - 599" refId="ref6294" refString="Cumberlidge N, Soma JB, Leever EM, Daniels SR. 2020. New lineages within the Malagasy freshwater crab fauna: agora n. gen. for Thelphusa goudoti H. Milne Edwards, 1853, and a phytotelmic new species of Malagasya Cumberlidge & Sternberg, 2002 (Brachyura: Potamonautidae: Deckeniidae). J Crust Biol. 40 (5): 584 - 599. doi: 10.1093 / jcbiol / ruaa 050." type="journal article" year="2020">Cumberlidge et al. 2020</bibRefCitation>
|
||
, fig. 10a,c,e).
|
||
</paragraph>
|
||
<paragraph blockId="7.[160,1156,142,1242]" pageId="7" pageNumber="470">
|
||
<taxonomicName authority="Cumberlidge & Daniels & Soma & Leever, 2023" authorityName="Cumberlidge & Daniels & Soma & Leever" authorityYear="2023" box="[192,258,454,479]" class="Malacostraca" family="Potamonautidae" genus="Vichai" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="470" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis box="[192,258,454,479]" italics="true" pageId="7" pageNumber="470">Vichai</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[269,377,454,479]" pageId="7" pageNumber="470" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
can be also distinguished from
|
||
<emphasis box="[760,880,454,479]" italics="true" pageId="7" pageNumber="470">
|
||
<collectingRegion box="[760,880,454,479]" country="Madagascar" name="Toamasina" pageId="7" pageNumber="470">Toamasina</collectingRegion>
|
||
</emphasis>
|
||
<bibRefCitation author="Leever EM & Daniels SR & Soma JB & Cumberlidge N" box="[891,1117,454,479]" pageId="7" pageNumber="470" pagination="241 - 263" refId="ref6477" refString="Leever EM, Daniels SR, Soma JB, Cumberlidge N. 2022. Two new genera and a new species of freshwater crabs from northern Madagascar: vahatra gen. nov. for Foza ambohitra Cumberlidge and Meyer, 2009, and Toamasina gen. nov. for Toamasina clarki sp. nov. (Brachyura: Potamoidea: Deckeniidae). J Nat Hist. 56 (1 - 4): 241 - 263. doi: 10.1080 / 00222933.2022.2049389." type="journal article" year="2022">Leever et al. (2022)</bibRefCitation>
|
||
, as follows. The margins of the cheliped ischium of
|
||
<taxonomicName authority="Cumberlidge & Daniels & Soma & Leever, 2023" authorityName="Cumberlidge & Daniels & Soma & Leever" authorityYear="2023" box="[694,760,489,514]" class="Malacostraca" family="Potamonautidae" genus="Vichai" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="470" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis box="[694,760,489,514]" italics="true" pageId="7" pageNumber="470">Vichai</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[767,870,489,514]" pageId="7" pageNumber="470" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
are smooth and rounded (
|
||
<figureCitation box="[168,261,524,549]" captionStart="Figure 2" captionStartId="4.[160,225,1736,1759]" captionTargetBox="[163,1153,142,1704]" captionTargetId="figure-3@4.[163,1153,142,1704]" captionTargetPageId="4" captionText="Figure 2. Vichai cyanalepou sp. nov., holotype, adult J CW 38.5 mm, FMNH 7589, from Marojejy National Park, Sava region, northern Madagascar: (a) frontal view of carapace; (b) frontal view of right chela; (c) frontal view of left chela; (d) right cheliped merus, underside; (e) right cheliped merus, dorsal view; (f) right cheliped carpus, dorsal view; (g) third maxilliped, frontal view; (h) third maxilliped, posterior view; (i) mandible frontal view. Scale bar: a = 5.3 mm; b, c = 8.0 mm; d = 5.3 mm; e, f = 6.7 mm; g, h = 3.3 mm; i = 2.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7859825" httpUri="https://zenodo.org/record/7859825/files/figure.png" pageId="7" pageNumber="470">Figure 2</figureCitation>
|
||
(d,e)), vs cheliped ischium margins that are lined with small teeth (see
|
||
<bibRefCitation author="Leever EM & Daniels SR & Soma JB & Cumberlidge N" pageId="7" pageNumber="470" pagination="241 - 263" refId="ref6477" refString="Leever EM, Daniels SR, Soma JB, Cumberlidge N. 2022. Two new genera and a new species of freshwater crabs from northern Madagascar: vahatra gen. nov. for Foza ambohitra Cumberlidge and Meyer, 2009, and Toamasina gen. nov. for Toamasina clarki sp. nov. (Brachyura: Potamoidea: Deckeniidae). J Nat Hist. 56 (1 - 4): 241 - 263. doi: 10.1080 / 00222933.2022.2049389." type="journal article" year="2022">Leever et al. 2022</bibRefCitation>
|
||
, fig.
|
||
<quantity box="[301,339,558,583]" metricMagnitude="-3" metricUnit="kg" metricValue="4.0" pageId="7" pageNumber="470" unit="g" value="4.0">4 g</quantity>
|
||
,h); the exopod of the third maxilliped of
|
||
<taxonomicName authority="Cumberlidge & Daniels & Soma & Leever, 2023" authorityName="Cumberlidge & Daniels & Soma & Leever" authorityYear="2023" box="[803,869,558,583]" class="Malacostraca" family="Potamonautidae" genus="Vichai" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="470" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis box="[803,869,558,583]" italics="true" pageId="7" pageNumber="470">Vichai</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[876,981,558,583]" pageId="7" pageNumber="470" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
has a mediumlength flagellum (~0.5× merus length) (
|
||
<figureCitation box="[595,687,593,618]" captionStart="Figure 2" captionStartId="4.[160,225,1736,1759]" captionTargetBox="[163,1153,142,1704]" captionTargetId="figure-3@4.[163,1153,142,1704]" captionTargetPageId="4" captionText="Figure 2. Vichai cyanalepou sp. nov., holotype, adult J CW 38.5 mm, FMNH 7589, from Marojejy National Park, Sava region, northern Madagascar: (a) frontal view of carapace; (b) frontal view of right chela; (c) frontal view of left chela; (d) right cheliped merus, underside; (e) right cheliped merus, dorsal view; (f) right cheliped carpus, dorsal view; (g) third maxilliped, frontal view; (h) third maxilliped, posterior view; (i) mandible frontal view. Scale bar: a = 5.3 mm; b, c = 8.0 mm; d = 5.3 mm; e, f = 6.7 mm; g, h = 3.3 mm; i = 2.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7859825" httpUri="https://zenodo.org/record/7859825/files/figure.png" pageId="7" pageNumber="470">Figure 2</figureCitation>
|
||
(g,h)), vs a third maxilliped exopod with an elongated flagellum (equal to the merus length) (see
|
||
<bibRefCitation author="Leever EM & Daniels SR & Soma JB & Cumberlidge N" box="[801,1012,628,653]" pageId="7" pageNumber="470" pagination="241 - 263" refId="ref6477" refString="Leever EM, Daniels SR, Soma JB, Cumberlidge N. 2022. Two new genera and a new species of freshwater crabs from northern Madagascar: vahatra gen. nov. for Foza ambohitra Cumberlidge and Meyer, 2009, and Toamasina gen. nov. for Toamasina clarki sp. nov. (Brachyura: Potamoidea: Deckeniidae). J Nat Hist. 56 (1 - 4): 241 - 263. doi: 10.1080 / 00222933.2022.2049389." type="journal article" year="2022">Leever et al. 2022</bibRefCitation>
|
||
, fig. 2 f). In addition, the ambulatory legs of
|
||
<taxonomicName authority="Cumberlidge & Daniels & Soma & Leever, 2023" authorityName="Cumberlidge & Daniels & Soma & Leever" authorityYear="2023" box="[551,617,662,687]" class="Malacostraca" family="Potamonautidae" genus="Vichai" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="470" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis box="[551,617,662,687]" italics="true" pageId="7" pageNumber="470">Vichai</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[628,737,662,687]" pageId="7" pageNumber="470" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
(P2–5) are short and stout (ΣP2–5/ CW = 5.4) (
|
||
<figureCitation box="[298,391,697,722]" captionStart="Figure 1" captionStartId="3.[160,225,1754,1777]" captionTargetBox="[172,1144,142,1722]" captionTargetId="figure-3@3.[172,1144,142,1722]" captionTargetPageId="3" captionText="Figure 1. Vichai cyanalepou sp. nov., holotype, adult J CW 38.5 mm, FMNH 7589, from Marojejy National Park,Sava region,northernMadagascar,whole animal:(a) dorsal view;(b) cephalothorax,ventral view.Scale bar: a = 10.8 mm; b = 5.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7859821" httpUri="https://zenodo.org/record/7859821/files/figure.png" pageId="7" pageNumber="470">Figure 1</figureCitation>
|
||
(a);
|
||
<tableCitation box="[441,528,697,722]" captionStart="Table 1" captionStartId="1.[160,216,1316,1339]" captionTargetPageId="1" captionText="Table 1. Vichai cyanalepou sp. nov. male holotype (FMNH 7589). Length in mm of the five articles (ischium, merus, carpus, propodus, dactylus) of the cheliped (P1) and the four ambulatory legs (P2–5), and the leg length in relation to the carapace width (CW). The sum of the length of each article of the four ambulatory legs divided by the carapace width is also shown (Total of 5 articles/CW), as is the sum of the relative length of the four ambulatory legs (ΣP2–5/CW = 5.35), placing this species in the short ambulatory leg size class (short: 5.0–5.9; medium: 6.0–6.9; long:>7.0)." httpUri="http://table.plazi.org/id/8CED6FA8676FB141FF2DA025EE1C5DA2" pageId="7" pageNumber="470" tableUuid="8CED6FA8676FB141FF2DA025EE1C5DA2">Table 1</tableCitation>
|
||
), vs ambulatory legs (P2–5) that are medium length (ΣP2–5/CW = 6.4) in
|
||
<emphasis box="[392,512,732,757]" italics="true" pageId="7" pageNumber="470">
|
||
<collectingRegion box="[392,512,732,757]" country="Madagascar" name="Toamasina" pageId="7" pageNumber="470">Toamasina</collectingRegion>
|
||
</emphasis>
|
||
(see
|
||
<bibRefCitation author="Leever EM & Daniels SR & Soma JB & Cumberlidge N" box="[572,770,732,757]" pageId="7" pageNumber="470" pagination="241 - 263" refId="ref6477" refString="Leever EM, Daniels SR, Soma JB, Cumberlidge N. 2022. Two new genera and a new species of freshwater crabs from northern Madagascar: vahatra gen. nov. for Foza ambohitra Cumberlidge and Meyer, 2009, and Toamasina gen. nov. for Toamasina clarki sp. nov. (Brachyura: Potamoidea: Deckeniidae). J Nat Hist. 56 (1 - 4): 241 - 263. doi: 10.1080 / 00222933.2022.2049389." type="journal article" year="2022">Leever et al. 2022</bibRefCitation>
|
||
, table 3).
|
||
</paragraph>
|
||
<paragraph blockId="7.[160,1156,142,1242]" pageId="7" pageNumber="470">
|
||
Finally,
|
||
<taxonomicName authority="Cumberlidge & Daniels & Soma & Leever, 2023" authorityName="Cumberlidge & Daniels & Soma & Leever" authorityYear="2023" box="[282,349,766,791]" class="Malacostraca" family="Potamonautidae" genus="Vichai" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="470" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis box="[282,349,766,791]" italics="true" pageId="7" pageNumber="470">Vichai</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[359,468,766,791]" pageId="7" pageNumber="470" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
can be distinguished from
|
||
<taxonomicName authority="Leever et al. (2022)" authorityName="Leever" authorityYear="2022" box="[794,1118,766,791]" class="Malacostraca" family="Deckeniidae" genus="Vahatra" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="470" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[794,883,766,791]" italics="true" pageId="7" pageNumber="470">Vahatra</emphasis>
|
||
<bibRefCitation author="Leever EM & Daniels SR & Soma JB & Cumberlidge N" box="[893,1118,766,791]" pageId="7" pageNumber="470" pagination="241 - 263" refId="ref6477" refString="Leever EM, Daniels SR, Soma JB, Cumberlidge N. 2022. Two new genera and a new species of freshwater crabs from northern Madagascar: vahatra gen. nov. for Foza ambohitra Cumberlidge and Meyer, 2009, and Toamasina gen. nov. for Toamasina clarki sp. nov. (Brachyura: Potamoidea: Deckeniidae). J Nat Hist. 56 (1 - 4): 241 - 263. doi: 10.1080 / 00222933.2022.2049389." type="journal article" year="2022">Leever et al. (2022)</bibRefCitation>
|
||
</taxonomicName>
|
||
by the characters of the chelipeds. The inner margin of the cheliped carpus of
|
||
<taxonomicName authority="Cumberlidge & Daniels & Soma & Leever, 2023" authorityName="Cumberlidge & Daniels & Soma & Leever" authorityYear="2023" box="[1088,1155,801,826]" class="Malacostraca" family="Potamonautidae" genus="Vichai" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="470" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis box="[1088,1155,801,826]" italics="true" pageId="7" pageNumber="470">Vichai</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[160,270,836,861]" pageId="7" pageNumber="470" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
has a relatively small pointed distal tooth, a smaller acute proximal tooth, and granules between these teeth (
|
||
<figureCitation box="[594,694,870,895]" captionStart="Figure 2" captionStartId="4.[160,225,1736,1759]" captionTargetBox="[163,1153,142,1704]" captionTargetId="figure-3@4.[163,1153,142,1704]" captionTargetPageId="4" captionText="Figure 2. Vichai cyanalepou sp. nov., holotype, adult J CW 38.5 mm, FMNH 7589, from Marojejy National Park, Sava region, northern Madagascar: (a) frontal view of carapace; (b) frontal view of right chela; (c) frontal view of left chela; (d) right cheliped merus, underside; (e) right cheliped merus, dorsal view; (f) right cheliped carpus, dorsal view; (g) third maxilliped, frontal view; (h) third maxilliped, posterior view; (i) mandible frontal view. Scale bar: a = 5.3 mm; b, c = 8.0 mm; d = 5.3 mm; e, f = 6.7 mm; g, h = 3.3 mm; i = 2.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7859825" httpUri="https://zenodo.org/record/7859825/files/figure.png" pageId="7" pageNumber="470">Figure 2</figureCitation>
|
||
(f)), vs a cheliped carpus inner margin with a relatively larger, pointed distal tooth and a significantly smaller, acute proximal tooth, with no granules between the teeth in
|
||
<taxonomicName box="[779,868,940,965]" class="Malacostraca" family="Deckeniidae" genus="Vahatra" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="470" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[779,868,940,965]" italics="true" pageId="7" pageNumber="470">Vahatra</emphasis>
|
||
</taxonomicName>
|
||
(see
|
||
<bibRefCitation author="Leever EM & Daniels SR & Soma JB & Cumberlidge N" box="[936,1150,940,965]" pageId="7" pageNumber="470" pagination="241 - 263" refId="ref6477" refString="Leever EM, Daniels SR, Soma JB, Cumberlidge N. 2022. Two new genera and a new species of freshwater crabs from northern Madagascar: vahatra gen. nov. for Foza ambohitra Cumberlidge and Meyer, 2009, and Toamasina gen. nov. for Toamasina clarki sp. nov. (Brachyura: Potamoidea: Deckeniidae). J Nat Hist. 56 (1 - 4): 241 - 263. doi: 10.1080 / 00222933.2022.2049389." type="journal article" year="2022">Leever et al. 2022</bibRefCitation>
|
||
, fig. 4f). The cutting edge of the fixed finger (pollex of propodus) of the major (left) chela of
|
||
<taxonomicName authority="Cumberlidge & Daniels & Soma & Leever, 2023" authorityName="Cumberlidge & Daniels & Soma & Leever" authorityYear="2023" box="[262,329,1009,1034]" class="Malacostraca" family="Potamonautidae" genus="Vichai" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="470" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis box="[262,329,1009,1034]" italics="true" pageId="7" pageNumber="470">Vichai</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[338,446,1009,1034]" pageId="7" pageNumber="470" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
has one large molar tooth flanked by small teeth proximally (
|
||
<figureCitation box="[168,265,1044,1069]" captionStart="Figure 2" captionStartId="4.[160,225,1736,1759]" captionTargetBox="[163,1153,142,1704]" captionTargetId="figure-3@4.[163,1153,142,1704]" captionTargetPageId="4" captionText="Figure 2. Vichai cyanalepou sp. nov., holotype, adult J CW 38.5 mm, FMNH 7589, from Marojejy National Park, Sava region, northern Madagascar: (a) frontal view of carapace; (b) frontal view of right chela; (c) frontal view of left chela; (d) right cheliped merus, underside; (e) right cheliped merus, dorsal view; (f) right cheliped carpus, dorsal view; (g) third maxilliped, frontal view; (h) third maxilliped, posterior view; (i) mandible frontal view. Scale bar: a = 5.3 mm; b, c = 8.0 mm; d = 5.3 mm; e, f = 6.7 mm; g, h = 3.3 mm; i = 2.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7859825" httpUri="https://zenodo.org/record/7859825/files/figure.png" pageId="7" pageNumber="470">Figure 2</figureCitation>
|
||
(e)), whereas the fixed finger of the major (right) chela of
|
||
<taxonomicName box="[960,1049,1044,1069]" class="Malacostraca" family="Deckeniidae" genus="Vahatra" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="470" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[960,1049,1044,1069]" italics="true" pageId="7" pageNumber="470">Vahatra</emphasis>
|
||
</taxonomicName>
|
||
has four large molars proximally (see
|
||
<bibRefCitation author="Leever EM & Daniels SR & Soma JB & Cumberlidge N" box="[502,712,1078,1103]" pageId="7" pageNumber="470" pagination="241 - 263" refId="ref6477" refString="Leever EM, Daniels SR, Soma JB, Cumberlidge N. 2022. Two new genera and a new species of freshwater crabs from northern Madagascar: vahatra gen. nov. for Foza ambohitra Cumberlidge and Meyer, 2009, and Toamasina gen. nov. for Toamasina clarki sp. nov. (Brachyura: Potamoidea: Deckeniidae). J Nat Hist. 56 (1 - 4): 241 - 263. doi: 10.1080 / 00222933.2022.2049389." type="journal article" year="2022">Leever et al. 2022</bibRefCitation>
|
||
, fig.
|
||
<quantity box="[774,815,1078,1103]" metricMagnitude="-3" metricUnit="kg" metricValue="4.0" pageId="7" pageNumber="470" unit="g" value="4.0">4 g</quantity>
|
||
,h). Further, the cutting edge of the movable finger (dactylus) of the major (left) chela of
|
||
<taxonomicName authority="Cumberlidge & Daniels & Soma & Leever, 2023" authorityName="Cumberlidge & Daniels & Soma & Leever" authorityYear="2023" box="[874,941,1113,1138]" class="Malacostraca" family="Potamonautidae" genus="Vichai" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="470" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis box="[874,941,1113,1138]" italics="true" pageId="7" pageNumber="470">Vichai</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[952,1061,1113,1138]" pageId="7" pageNumber="470" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
is lined with small teeth only and lacks large teeth (
|
||
<figureCitation box="[677,772,1148,1173]" captionStart="Figure 2" captionStartId="4.[160,225,1736,1759]" captionTargetBox="[163,1153,142,1704]" captionTargetId="figure-3@4.[163,1153,142,1704]" captionTargetPageId="4" captionText="Figure 2. Vichai cyanalepou sp. nov., holotype, adult J CW 38.5 mm, FMNH 7589, from Marojejy National Park, Sava region, northern Madagascar: (a) frontal view of carapace; (b) frontal view of right chela; (c) frontal view of left chela; (d) right cheliped merus, underside; (e) right cheliped merus, dorsal view; (f) right cheliped carpus, dorsal view; (g) third maxilliped, frontal view; (h) third maxilliped, posterior view; (i) mandible frontal view. Scale bar: a = 5.3 mm; b, c = 8.0 mm; d = 5.3 mm; e, f = 6.7 mm; g, h = 3.3 mm; i = 2.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7859825" httpUri="https://zenodo.org/record/7859825/files/figure.png" pageId="7" pageNumber="470">Figure 2</figureCitation>
|
||
(e)), whereas the cutting edge of the movable finger of the major (right) chela of
|
||
<taxonomicName box="[797,888,1182,1207]" class="Malacostraca" family="Deckeniidae" genus="Vahatra" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="470" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[797,888,1182,1207]" italics="true" pageId="7" pageNumber="470">Vahatra</emphasis>
|
||
</taxonomicName>
|
||
has two large teeth proximally and one large tooth midway (see
|
||
<bibRefCitation author="Leever EM & Daniels SR & Soma JB & Cumberlidge N" box="[697,908,1217,1242]" pageId="7" pageNumber="470" pagination="241 - 263" refId="ref6477" refString="Leever EM, Daniels SR, Soma JB, Cumberlidge N. 2022. Two new genera and a new species of freshwater crabs from northern Madagascar: vahatra gen. nov. for Foza ambohitra Cumberlidge and Meyer, 2009, and Toamasina gen. nov. for Toamasina clarki sp. nov. (Brachyura: Potamoidea: Deckeniidae). J Nat Hist. 56 (1 - 4): 241 - 263. doi: 10.1080 / 00222933.2022.2049389." type="journal article" year="2022">Leever et al. 2022</bibRefCitation>
|
||
, fig.
|
||
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|
||
,h).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |