treatments-xml/data/44/C6/8B/44C68B7595C95AE795395E0E5282D178.xml

<document ID-DOI="http://dx.doi.org/10.3897/zookeys.873.35421" ID-GBIF-Dataset="ad4b6ae3-173c-494f-b761-56211fc3fdeb" ID-PMC="PMC6728363" ID-Pensoft-Pub="1313-2970-873-25" ID-Pensoft-UUID="AB7ECFA7FA38588E89AA94B67B6C6026" ID-PubMed="31534383" ID-ZooBank="6149E4233C2847BB89FC927209B5D2DE" ModsDocID="1313-2970-873-25" checkinTime="1567102845756" checkinUser="pensoft" docAuthor="Hosie, Andrew M., Fromont, Jane, Munyard, Kylie &amp; Jones, Diana S." docDate="2019" docId="44C68B7595C95AE795395E0E5282D178" docLanguage="en" docName="ZooKeys 873: 25-42" docOrigin="ZooKeys 873" docSource="http://dx.doi.org/10.3897/zookeys.873.35421" docTitle="Membranobalanus porphyrophilus Hosie &amp; Jones, sp. nov." docType="treatment" docUuid="170E814F-E3ED-4120-8622-EC4230353C65" docUuidSource="ZooBank" docVersion="5" id="AB7ECFA7FA38588E89AA94B67B6C6026" lastPageNumber="36" masterDocId="AB7ECFA7FA38588E89AA94B67B6C6026" masterDocTitle="Description of a new species of Membranobalanus (Crustacea, Cirripedia) from southern Australia" masterLastPageNumber="42" masterPageNumber="25" pageNumber="29" updateTime="1668167761432" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>Description of a new species of Membranobalanus (Crustacea, Cirripedia) from southern Australia</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Hosie, Andrew M.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Fromont, Jane</mods:namePart>
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<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Munyard, Kylie</mods:namePart>
</mods:name>
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<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Jones, Diana S.</mods:namePart>
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<mods:title>ZooKeys</mods:title>
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<mods:date>2019</mods:date>
<mods:detail type="volume">
<mods:number>873</mods:number>
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<mods:start>25</mods:start>
<mods:end>42</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zookeys.873.35421</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.873.35421</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-873-25</mods:identifier>
<mods:identifier type="ZooBank">6149E4233C2847BB89FC927209B5D2DE</mods:identifier>
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<treatment ID-GBIF-Taxon="159319362" LSID="urn:lsid:zoobank.org:act:170E814F-E3ED-4120-8622-EC4230353C65" httpUri="http://treatment.plazi.org/id/44C68B7595C95AE795395E0E5282D178" lastPageId="11" lastPageNumber="36" pageId="4" pageNumber="29">
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<paragraph pageId="4" pageNumber="29">
<taxonomicName LSID="44C68B7595C95AE795395E0E5282D178" authority="Hosie &amp; Jones" class="Maxillopoda" family="Archaeobalanidae" genus="Membranobalanus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Membranobalanus porphyrophilus" order="Sessilia" pageId="4" pageNumber="29" phylum="Arthropoda" rank="species" species="porphyrophilus">Membranobalanus porphyrophilus Hosie &amp; Jones</taxonomicName>
<taxonomicNameLabel pageId="4" pageNumber="29">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Interactive 3 D, μCT derived volume reconstruction of Membranobalanus porphyrophilus sp. nov. holotype (WAM C 66803). Only the well-calcified plates are illustrated as scanning limitations prevented the softer prosoma from being differentiated. Note: To enable the interactive function of this figure, open the PDF in Adobe Reader program or web plug-in." figureDoi="10.3897/zookeys.873.35421.figure1" httpUri="https://binary.pensoft.net/fig/328935" pageId="4" pageNumber="29">Figures 1</figureCitation>
, 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Membranobalanus porphyrophilus sp. nov. parietes A holotype (WAM C 66803) B-J paratype (SAMA C 12708) K paratype (SAMA C 12706). A Whole shell lateral view B whole shell lateral view C, D rostrum, external and internal view E, F left lateral plate, external, and internal view G, H left carinolateral, external and internal view I, J carina external and internal view K internal view of articulated right carina, carinolateral and lateral plates. Scale bars: 5 mm (A-J); 2 mm (K)." figureDoi="10.3897/zookeys.873.35421.figure2" httpUri="https://binary.pensoft.net/fig/328936" pageId="4" pageNumber="29">2</figureCitation>
, 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Membranobalanus porphyrophilus sp. nov. opercular plates A-H holotype (WAM C 66803) I-L paratype (SAMA C 12708). A Left scutum, external view B left scutum, internal view C right scutum, external view D right scutum, internal view E left tergum, external view F left tergum, internal view G right tergum, external view H right tergum, internal view I left scutum, external view J left scutum, internal view K left tergum, external view L left tergum, internal view. Scale bars: 2 mm." figureDoi="10.3897/zookeys.873.35421.figure3" httpUri="https://binary.pensoft.net/fig/328937" pageId="4" pageNumber="29">3</figureCitation>
, 
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Membranobalanus porphyrophilus sp. nov. mouthparts A, B, G, H, I, L, N holotype (WAM C 66803) C, D, J, O paratype (SAMA C 12708) E, F, K, M paratype (SAMA C 12706). A, C Labrum and right mandiblular palp (left removed, damaged) B, D close up of A and D, arrows indicate teeth on labrum E left mandibular palp F detail of serrulate setae on mandibular palp ventral face G-I mandibles J maxillule K, L maxilla. Note: G, H, L damaged, setae lost during sonication in A &amp; K. Scale bars: 200 μm (A, C, E, G, I-O); 40 μm (B, F); 100 μm (D); 50 μm (H)." figureDoi="10.3897/zookeys.873.35421.figure4" httpUri="https://binary.pensoft.net/fig/328938" pageId="4" pageNumber="29">4</figureCitation>
, 
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Membranobalanus porphyrophilus sp. nov. cirri A-F holotype (WAM C 66803) G-I paratype (SAMA C 12706) J-L paratype (SAMA C 12708). A-C Right cirrus I-III lateral view D right cirrus III intermediate segments E right cirrus IV lateral view F same, anterior ramus intermediate segments G left cirrus IV anterior ramus intermediate segments H right cirrus IV anterodistal angle of distal segment of pedical showing ctenoid scales I, J Ieft cirrus IV basal segments of rami showing posterior spines K close up of spines on posterior margin of J L close up of spines and posterodistal spines basal segments of J. Note: B, C, E damaged. Scale bars: 400 μm (A-C, E); 100 μm (D, F, L); 150 μm (G, I); 50 μm (H, K); 200 μm (J)." figureDoi="10.3897/zookeys.873.35421.figure5" httpUri="https://binary.pensoft.net/fig/328939" pageId="4" pageNumber="29">5</figureCitation>
, 
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Membranobalanus porphyrophilus sp. nov. cirri A, B, D, E, H holotype (WAM C 66803) C, F, G paratype (SAMA C 12706). A Right cirrus V lateral view B basal segments of cirrus V rami C intermediate segments of cirrus V anterior ramus D right cirrus VI mesial view (damaged) E same, intermediate segments F example of long serrulate setae on cirrus VI G basal segment of pedicel of right cirrus VI, mesial view showing denticles on anterior and posterior margin H basidorsal point of penis. Scale bars: 400 μm (A, D); 100 μm (B, C); 50 μm (F); 150 μm (G, H); 80 μm (E)." figureDoi="10.3897/zookeys.873.35421.figure6" httpUri="https://binary.pensoft.net/fig/328940" pageId="4" pageNumber="29">6</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="29" type="materials_examined">
<paragraph pageId="4" pageNumber="29">Material examined.</paragraph>
<paragraph pageId="4" pageNumber="29">
<emphasis bold="true" pageId="4" pageNumber="29">Holotype.</emphasis>
AUSTRALIA • WAM C66803, 1 hermaphrodite; 9 mm rostro-carinal diameter; Western Australia SE of Rottnest Island, Wallace Island, The Count; 
<geoCoordinate degrees="32" direction="south" minutes="0.89" orientation="latitude" precision="9" value="-32.01483">32°0.89'S</geoCoordinate>
, 
<geoCoordinate degrees="115" direction="east" minutes="33.53" orientation="longitude" precision="9" value="115.55883">115°33.53'E</geoCoordinate>
; 12 m; coll. A.M. Hosie; 23 Feb 2017; host: WAM Z86929, 
<taxonomicName class="Demospongiae" family="Clionaidae" genus="Spheciospongia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Spheciospongia purpurea" order="Clionaida" pageId="4" pageNumber="29" phylum="Porifera" rank="species" species="purpurea">
<emphasis italics="true" pageId="4" pageNumber="29">Spheciospongia purpurea</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph pageId="4" pageNumber="29">
<emphasis bold="true" pageId="4" pageNumber="29">Paratypes.</emphasis>
AUSTRALIA • WAM C71852, 1 hermaphrodite; 8 mm rostro-carinal diameter; empty shell; same as data as for holotype. • WAM C71853, 1 hermaphrodite; same as data as for holotype. • WAM C71881, 1 hermaphrodite; same as data as for holotype. • SAMA C12706, 1 hermaphrodite; South Australia, Kangaroo Island, off Second Gully between Western River Cove and Snug Cove; 32 m; coll. J. Thiselton; 19 Nov 2002; host: SAMA S2910, 
<taxonomicName lsidName="S. purpurea" pageId="4" pageNumber="29" rank="species" species="purpurea">
<emphasis italics="true" pageId="4" pageNumber="29">S. purpurea</emphasis>
</taxonomicName>
. • SAMA C12707, 1 hermaphrodite; same data as for previous. • SAMA C12708, 1 hermaphrodite; 14 mm rostro-carinal diameter; same data as for previous.
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="29" type="diagnosis">
<paragraph pageId="4" pageNumber="29">Diagnosis.</paragraph>
<paragraph pageId="4" pageNumber="29">Shell wall robust, cylindrical, growth ridges weak; orifice toothed, large; rostrum basal margin broadly rounded, extending below basal plane of remaining parietes. Tergum narrow, beaked, spur narrow, separated from basiscutal angle by half its own width; scutum with faint, external longitudinal striations; basitergal angle broadly rounded. Cirri III and IV with row of strong, erect spines on anterodistal margin of anterior ramus; cirrus IV pedicel without erect spines; cirri IV and V with row of stout spines on posterior margin of anterior ramus basal segment.</paragraph>
</subSubSection>
<subSubSection lastPageId="11" lastPageNumber="36" pageId="4" pageNumber="29" type="description">
<paragraph pageId="4" pageNumber="29">Description.</paragraph>
<paragraph pageId="4" pageNumber="29">
All shell plates, prosoma, and internal organs stained purple 
<emphasis italics="true" pageId="4" pageNumber="29">in vivo</emphasis>
, otherwise white. Shell walls (
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Interactive 3 D, μCT derived volume reconstruction of Membranobalanus porphyrophilus sp. nov. holotype (WAM C 66803). Only the well-calcified plates are illustrated as scanning limitations prevented the softer prosoma from being differentiated. Note: To enable the interactive function of this figure, open the PDF in Adobe Reader program or web plug-in." figureDoi="10.3897/zookeys.873.35421.figure1" httpUri="https://binary.pensoft.net/fig/328935" pageId="4" pageNumber="29">Figs 1</figureCitation>
, 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Membranobalanus porphyrophilus sp. nov. parietes A holotype (WAM C 66803) B-J paratype (SAMA C 12708) K paratype (SAMA C 12706). A Whole shell lateral view B whole shell lateral view C, D rostrum, external and internal view E, F left lateral plate, external, and internal view G, H left carinolateral, external and internal view I, J carina external and internal view K internal view of articulated right carina, carinolateral and lateral plates. Scale bars: 5 mm (A-J); 2 mm (K)." figureDoi="10.3897/zookeys.873.35421.figure2" httpUri="https://binary.pensoft.net/fig/328936" pageId="4" pageNumber="29">2</figureCitation>
) parallel, except bowed rostrum; parietes externally with horizontal growth lines raised, ridge-like, giving shell a roughened appearance; radii prominent, summits oblique, sutural edges roughened, alae wide, summits oblique. Internally parietes smooth, sheath occupying approximately half of shell height, with horizontal growth lines, basal margin adpressed. Carina approximately as wide as lateral plate, carinolateral narrowest plate, approximately one-third width of lateral plate. Rostrum elongate, extending below basal margin of other parietes, almost twice length and twice width of lateral plate, basal margin broadly rounded below basal margin of latera. Basal margins of carina and carinolatera more or less perpendicular to shell vertical axis, latera basal margins curving basally to form contiguous rim connecting latera with rostrum. Basis membranous, follows contour of basal rim, not depending below parietes.
</paragraph>
<caption doi="10.3897/zookeys.873.35421.figure1" httpUri="https://binary.pensoft.net/fig/328935" pageId="4" pageNumber="29" start="Figure 1" startId="F1">
<paragraph pageId="4" pageNumber="29">
Figure 1. Interactive 3D, 
<normalizedToken originalValue="μCT">μCT</normalizedToken>
derived volume reconstruction of 
<taxonomicName class="Maxillopoda" family="Archaeobalanidae" genus="Membranobalanus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Membranobalanus porphyrophilus" order="Sessilia" pageId="4" pageNumber="29" phylum="Arthropoda" rank="species" species="porphyrophilus">
<emphasis italics="true" pageId="4" pageNumber="29">Membranobalanus porphyrophilus</emphasis>
</taxonomicName>
sp. nov. holotype (WAM C66803). Only the well-calcified plates are illustrated as scanning limitations prevented the softer prosoma from being differentiated. Note: To enable the interactive function of this figure, open the PDF in Adobe Reader program or web plug-in.
</paragraph>
</caption>
<caption doi="10.3897/zookeys.873.35421.figure2" httpUri="https://binary.pensoft.net/fig/328936" pageId="4" pageNumber="29" start="Figure 2" startId="F2">
<paragraph pageId="4" pageNumber="29">
Figure 2. 
<taxonomicName class="Maxillopoda" family="Archaeobalanidae" genus="Membranobalanus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Membranobalanus porphyrophilus" order="Sessilia" pageId="4" pageNumber="29" phylum="Arthropoda" rank="species" species="porphyrophilus">
<emphasis italics="true" pageId="4" pageNumber="29">Membranobalanus porphyrophilus</emphasis>
</taxonomicName>
sp. nov. parietes 
<emphasis bold="true" pageId="4" pageNumber="29">A</emphasis>
holotype (WAM C66803) 
<emphasis bold="true" pageId="4" pageNumber="29">
<normalizedToken originalValue="B–J">B-J</normalizedToken>
</emphasis>
paratype (SAMA C12708) 
<emphasis bold="true" pageId="4" pageNumber="29">K</emphasis>
paratype (SAMA C12706). 
<emphasis bold="true" pageId="4" pageNumber="29">A</emphasis>
Whole shell lateral view 
<emphasis bold="true" pageId="4" pageNumber="29">B</emphasis>
whole shell lateral view 
<emphasis bold="true" pageId="4" pageNumber="29">C, D</emphasis>
rostrum, external and internal view 
<emphasis bold="true" pageId="4" pageNumber="29">E, F</emphasis>
left lateral plate, external, and internal view 
<emphasis bold="true" pageId="4" pageNumber="29">G, H</emphasis>
left carinolateral, external and internal view 
<emphasis bold="true" pageId="4" pageNumber="29">I, J</emphasis>
carina external and internal view 
<emphasis bold="true" pageId="4" pageNumber="29">K</emphasis>
internal view of articulated right carina, carinolateral and lateral plates. Scale bars: 5 mm (
<emphasis bold="true" pageId="4" pageNumber="29">
<normalizedToken originalValue="A–J">A-J</normalizedToken>
</emphasis>
); 2 mm (
<emphasis bold="true" pageId="4" pageNumber="29">K</emphasis>
).
</paragraph>
</caption>
<paragraph lastPageId="5" lastPageNumber="30" pageId="4" pageNumber="29">
Scutum (
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Membranobalanus porphyrophilus sp. nov. opercular plates A-H holotype (WAM C 66803) I-L paratype (SAMA C 12708). A Left scutum, external view B left scutum, internal view C right scutum, external view D right scutum, internal view E left tergum, external view F left tergum, internal view G right tergum, external view H right tergum, internal view I left scutum, external view J left scutum, internal view K left tergum, external view L left tergum, internal view. Scale bars: 2 mm." figureDoi="10.3897/zookeys.873.35421.figure3" httpUri="https://binary.pensoft.net/fig/328937" pageId="4" pageNumber="29">
Fig. 3 
<normalizedToken originalValue="A–D">A-D</normalizedToken>
, I, J
</figureCitation>
) triangular, height 1.3-1.8 times width, growth lines prominent with faint longitudinal striations; basal and tergal margins separated by 
<pageBreakToken pageId="5" pageNumber="30" start="start">broadly</pageBreakToken>
rounded basitergal angle. Internal surface, slightly ridged apically near tergal margin, otherwise smooth; adductor muscle pit distinct; lateral depressor muscle pit distinct, without depressor muscle crests, extending one-third distance to beginning of articular ridge bounded by low, rounded adductor muscle ridge. Articular groove deep, narrow; articular ridge barely projecting beyond articular margin, basal margin curved.
</paragraph>
<caption doi="10.3897/zookeys.873.35421.figure3" httpUri="https://binary.pensoft.net/fig/328937" pageId="5" pageNumber="30" start="Figure 3" startId="F3">
<paragraph pageId="5" pageNumber="30">
Figure 3. 
<taxonomicName class="Maxillopoda" family="Archaeobalanidae" genus="Membranobalanus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Membranobalanus porphyrophilus" order="Sessilia" pageId="5" pageNumber="30" phylum="Arthropoda" rank="species" species="porphyrophilus">
<emphasis italics="true" pageId="5" pageNumber="30">Membranobalanus porphyrophilus</emphasis>
</taxonomicName>
sp. nov. opercular plates 
<emphasis bold="true" pageId="5" pageNumber="30">
<normalizedToken originalValue="A–H">A-H</normalizedToken>
</emphasis>
holotype (WAM C66803) 
<emphasis bold="true" pageId="5" pageNumber="30">
<normalizedToken originalValue="I–L">I-L</normalizedToken>
</emphasis>
paratype (SAMA C12708). 
<emphasis bold="true" pageId="5" pageNumber="30">A</emphasis>
Left scutum, external view 
<emphasis bold="true" pageId="5" pageNumber="30">B</emphasis>
left scutum, internal view 
<emphasis bold="true" pageId="5" pageNumber="30">C</emphasis>
right scutum, external view 
<emphasis bold="true" pageId="5" pageNumber="30">D</emphasis>
right scutum, internal view 
<emphasis bold="true" pageId="5" pageNumber="30">E</emphasis>
left tergum, external view 
<emphasis bold="true" pageId="5" pageNumber="30">F</emphasis>
left tergum, internal view 
<emphasis bold="true" pageId="5" pageNumber="30">G</emphasis>
right tergum, external view 
<emphasis bold="true" pageId="5" pageNumber="30">H</emphasis>
right tergum, internal view 
<emphasis bold="true" pageId="5" pageNumber="30">I</emphasis>
left scutum, external view 
<emphasis bold="true" pageId="5" pageNumber="30">J</emphasis>
left scutum, internal view 
<emphasis bold="true" pageId="5" pageNumber="30">K</emphasis>
left tergum, external view 
<emphasis bold="true" pageId="5" pageNumber="30">L</emphasis>
left tergum, internal view. Scale bars: 2 mm.
</paragraph>
</caption>
<paragraph lastPageId="7" lastPageNumber="32" pageId="6" pageNumber="31">
<pageBreakToken pageId="6" pageNumber="31" start="start">Tergum</pageBreakToken>
(
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Membranobalanus porphyrophilus sp. nov. opercular plates A-H holotype (WAM C 66803) I-L paratype (SAMA C 12708). A Left scutum, external view B left scutum, internal view C right scutum, external view D right scutum, internal view E left tergum, external view F left tergum, internal view G right tergum, external view H right tergum, internal view I left scutum, external view J left scutum, internal view K left tergum, external view L left tergum, internal view. Scale bars: 2 mm." figureDoi="10.3897/zookeys.873.35421.figure3" httpUri="https://binary.pensoft.net/fig/328937" pageId="6" pageNumber="31">
Fig. 3 
<normalizedToken originalValue="E–H">E-H</normalizedToken>
, K, L
</figureCitation>
) narrow, height more than twice width, articular and carinal margins arcuate with beaked apex; external growth lines conspicuous, but less raised than those of scutum; spur furrow indicated by shift in growth lines and slight depression; basiscutal angle sloping into spur; spur narrow, separated from basiscutal 
<pageBreakToken pageId="7" pageNumber="32" start="start">angle</pageBreakToken>
by half its width, basally truncate. Internally smooth, depressor muscle crests weak (may be absent in small specimens), articular groove wide, open, shallow; articular ridge low, becoming confluent with scutal margin basally.
</paragraph>
<paragraph pageId="7" pageNumber="32">
Labrum (
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Membranobalanus porphyrophilus sp. nov. mouthparts A, B, G, H, I, L, N holotype (WAM C 66803) C, D, J, O paratype (SAMA C 12708) E, F, K, M paratype (SAMA C 12706). A, C Labrum and right mandiblular palp (left removed, damaged) B, D close up of A and D, arrows indicate teeth on labrum E left mandibular palp F detail of serrulate setae on mandibular palp ventral face G-I mandibles J maxillule K, L maxilla. Note: G, H, L damaged, setae lost during sonication in A &amp; K. Scale bars: 200 μm (A, C, E, G, I-O); 40 μm (B, F); 100 μm (D); 50 μm (H)." figureDoi="10.3897/zookeys.873.35421.figure4" httpUri="https://binary.pensoft.net/fig/328938" pageId="7" pageNumber="32">
Fig. 4 
<normalizedToken originalValue="A–F">A-F</normalizedToken>
</figureCitation>
) bilobed, with deep medial notch bounded by rounded crests, each with two or three marginal teeth and numerous fine setae.
</paragraph>
<caption doi="10.3897/zookeys.873.35421.figure4" httpUri="https://binary.pensoft.net/fig/328938" pageId="7" pageNumber="32" start="Figure 4" startId="F4">
<paragraph pageId="7" pageNumber="32">
Figure 4. 
<taxonomicName class="Maxillopoda" family="Archaeobalanidae" genus="Membranobalanus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Membranobalanus porphyrophilus" order="Sessilia" pageId="7" pageNumber="32" phylum="Arthropoda" rank="species" species="porphyrophilus">
<emphasis italics="true" pageId="7" pageNumber="32">Membranobalanus porphyrophilus</emphasis>
</taxonomicName>
sp. nov. mouthparts 
<emphasis bold="true" pageId="7" pageNumber="32">A, B, G, H, I, L, N</emphasis>
holotype (WAM C66803) 
<emphasis bold="true" pageId="7" pageNumber="32">C, D, J, O</emphasis>
paratype (SAMA C12708) 
<emphasis bold="true" pageId="7" pageNumber="32">E, F, K, M</emphasis>
paratype (SAMA C12706). 
<emphasis bold="true" pageId="7" pageNumber="32">A, C</emphasis>
Labrum and right mandiblular palp (left removed, damaged) 
<emphasis bold="true" pageId="7" pageNumber="32">B, D</emphasis>
close up of A and D, arrows indicate teeth on labrum 
<emphasis bold="true" pageId="7" pageNumber="32">E</emphasis>
left mandibular palp 
<emphasis bold="true" pageId="7" pageNumber="32">F</emphasis>
detail of serrulate setae on mandibular palp ventral face 
<emphasis bold="true" pageId="7" pageNumber="32">
<normalizedToken originalValue="G–I">G-I</normalizedToken>
</emphasis>
mandibles 
<emphasis bold="true" pageId="7" pageNumber="32">J</emphasis>
maxillule 
<emphasis bold="true" pageId="7" pageNumber="32">K, L</emphasis>
maxilla. Note: G, H, L damaged, setae lost during sonication in A &amp; K. Scale bars: 200 
<normalizedToken originalValue="μm">μm</normalizedToken>
(
<emphasis bold="true" pageId="7" pageNumber="32">
A, C, E, G, 
<normalizedToken originalValue="I–O">I-O</normalizedToken>
</emphasis>
); 40 
<normalizedToken originalValue="μm">μm</normalizedToken>
(
<emphasis bold="true" pageId="7" pageNumber="32">B, F</emphasis>
); 100 
<normalizedToken originalValue="μm">μm</normalizedToken>
(
<emphasis bold="true" pageId="7" pageNumber="32">D</emphasis>
); 50 
<normalizedToken originalValue="μm">μm</normalizedToken>
(
<emphasis bold="true" pageId="7" pageNumber="32">H</emphasis>
).
</paragraph>
</caption>
<paragraph pageId="8" pageNumber="33">
<pageBreakToken pageId="8" pageNumber="33" start="start">Mandibular</pageBreakToken>
palp (
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Membranobalanus porphyrophilus sp. nov. mouthparts A, B, G, H, I, L, N holotype (WAM C 66803) C, D, J, O paratype (SAMA C 12708) E, F, K, M paratype (SAMA C 12706). A, C Labrum and right mandiblular palp (left removed, damaged) B, D close up of A and D, arrows indicate teeth on labrum E left mandibular palp F detail of serrulate setae on mandibular palp ventral face G-I mandibles J maxillule K, L maxilla. Note: G, H, L damaged, setae lost during sonication in A &amp; K. Scale bars: 200 μm (A, C, E, G, I-O); 40 μm (B, F); 100 μm (D); 50 μm (H)." figureDoi="10.3897/zookeys.873.35421.figure4" httpUri="https://binary.pensoft.net/fig/328938" pageId="8" pageNumber="33">Fig. 4A, C, E, G, H</figureCitation>
) rhomboid, apex obliquely truncate, anterior margin concave, posterior margin straight; setae (
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Membranobalanus porphyrophilus sp. nov. mouthparts A, B, G, H, I, L, N holotype (WAM C 66803) C, D, J, O paratype (SAMA C 12708) E, F, K, M paratype (SAMA C 12706). A, C Labrum and right mandiblular palp (left removed, damaged) B, D close up of A and D, arrows indicate teeth on labrum E left mandibular palp F detail of serrulate setae on mandibular palp ventral face G-I mandibles J maxillule K, L maxilla. Note: G, H, L damaged, setae lost during sonication in A &amp; K. Scale bars: 200 μm (A, C, E, G, I-O); 40 μm (B, F); 100 μm (D); 50 μm (H)." figureDoi="10.3897/zookeys.873.35421.figure4" httpUri="https://binary.pensoft.net/fig/328938" pageId="8" pageNumber="33">Fig. 4F</figureCitation>
) heavily serrulate, becoming longer and denser distally.
</paragraph>
<paragraph pageId="9" pageNumber="34">
<pageBreakToken pageId="9" pageNumber="34" start="start">Mandible</pageBreakToken>
(
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Membranobalanus porphyrophilus sp. nov. mouthparts A, B, G, H, I, L, N holotype (WAM C 66803) C, D, J, O paratype (SAMA C 12708) E, F, K, M paratype (SAMA C 12706). A, C Labrum and right mandiblular palp (left removed, damaged) B, D close up of A and D, arrows indicate teeth on labrum E left mandibular palp F detail of serrulate setae on mandibular palp ventral face G-I mandibles J maxillule K, L maxilla. Note: G, H, L damaged, setae lost during sonication in A &amp; K. Scale bars: 200 μm (A, C, E, G, I-O); 40 μm (B, F); 100 μm (D); 50 μm (H)." figureDoi="10.3897/zookeys.873.35421.figure4" httpUri="https://binary.pensoft.net/fig/328938" pageId="9" pageNumber="34">
Fig. 4 
<normalizedToken originalValue="I–K">I-K</normalizedToken>
</figureCitation>
) with four distinct teeth, 2nd and 3rd tooth bifid, 4th tooth much smaller than preceding three, 5th tooth obsolescent, confluent with molariform inferior angle; short setae covering inner and outer faces, longer fine setae on inferior and superior margins.
</paragraph>
<paragraph pageId="9" pageNumber="34">
Maxillule (
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Membranobalanus porphyrophilus sp. nov. mouthparts A, B, G, H, I, L, N holotype (WAM C 66803) C, D, J, O paratype (SAMA C 12708) E, F, K, M paratype (SAMA C 12706). A, C Labrum and right mandiblular palp (left removed, damaged) B, D close up of A and D, arrows indicate teeth on labrum E left mandibular palp F detail of serrulate setae on mandibular palp ventral face G-I mandibles J maxillule K, L maxilla. Note: G, H, L damaged, setae lost during sonication in A &amp; K. Scale bars: 200 μm (A, C, E, G, I-O); 40 μm (B, F); 100 μm (D); 50 μm (H)." figureDoi="10.3897/zookeys.873.35421.figure4" httpUri="https://binary.pensoft.net/fig/328938" pageId="9" pageNumber="34">Fig. 4l, M</figureCitation>
) with 10 robust setae on cutting margin, first, second, and ninth longer and more robust than remaining setae, inferior angle with several short robust setae. Cutting margin straight, with very slight notch below second seta. Dense, short setae regularly spaced on inner and outer faces, longer fine setae on inferior and superior margin
</paragraph>
<paragraph pageId="9" pageNumber="34">
Maxilla (
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Membranobalanus porphyrophilus sp. nov. mouthparts A, B, G, H, I, L, N holotype (WAM C 66803) C, D, J, O paratype (SAMA C 12708) E, F, K, M paratype (SAMA C 12706). A, C Labrum and right mandiblular palp (left removed, damaged) B, D close up of A and D, arrows indicate teeth on labrum E left mandibular palp F detail of serrulate setae on mandibular palp ventral face G-I mandibles J maxillule K, L maxilla. Note: G, H, L damaged, setae lost during sonication in A &amp; K. Scale bars: 200 μm (A, C, E, G, I-O); 40 μm (B, F); 100 μm (D); 50 μm (H)." figureDoi="10.3897/zookeys.873.35421.figure4" httpUri="https://binary.pensoft.net/fig/328938" pageId="9" pageNumber="34">Fig. 4N, O</figureCitation>
) bilobed, basal lobe ovate, serrulate setae arranged on anterior margin; distal lobe elongate, serrulate setae on anterior margin becoming more dense at apex, longer than those on basal lobe.
</paragraph>
<paragraph pageId="9" pageNumber="34">
Cirrus I (
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Membranobalanus porphyrophilus sp. nov. cirri A-F holotype (WAM C 66803) G-I paratype (SAMA C 12706) J-L paratype (SAMA C 12708). A-C Right cirrus I-III lateral view D right cirrus III intermediate segments E right cirrus IV lateral view F same, anterior ramus intermediate segments G left cirrus IV anterior ramus intermediate segments H right cirrus IV anterodistal angle of distal segment of pedical showing ctenoid scales I, J Ieft cirrus IV basal segments of rami showing posterior spines K close up of spines on posterior margin of J L close up of spines and posterodistal spines basal segments of J. Note: B, C, E damaged. Scale bars: 400 μm (A-C, E); 100 μm (D, F, L); 150 μm (G, I); 50 μm (H, K); 200 μm (J)." figureDoi="10.3897/zookeys.873.35421.figure5" httpUri="https://binary.pensoft.net/fig/328939" pageId="9" pageNumber="34">Fig. 5A</figureCitation>
) with unequal rami, anterior ramus twice length of posterior ramus, both rami bearing serrulate and simple setae. Posterior ramus segments with protuberant anterior margins, more densely setose than anterior ramus, arranged in tufts on anterior margins.
</paragraph>
<caption doi="10.3897/zookeys.873.35421.figure5" httpUri="https://binary.pensoft.net/fig/328939" pageId="9" pageNumber="34" start="Figure 5" startId="F5">
<paragraph pageId="9" pageNumber="34">
Figure 5. 
<taxonomicName class="Maxillopoda" family="Archaeobalanidae" genus="Membranobalanus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Membranobalanus porphyrophilus" order="Sessilia" pageId="9" pageNumber="34" phylum="Arthropoda" rank="species" species="porphyrophilus">
<emphasis italics="true" pageId="9" pageNumber="34">Membranobalanus porphyrophilus</emphasis>
</taxonomicName>
sp. nov. cirri 
<emphasis bold="true" pageId="9" pageNumber="34">
<normalizedToken originalValue="A–F">A-F</normalizedToken>
</emphasis>
holotype (WAM C66803) 
<emphasis bold="true" pageId="9" pageNumber="34">
<normalizedToken originalValue="G–I">G-I</normalizedToken>
</emphasis>
paratype (SAMA C12706) 
<emphasis bold="true" pageId="9" pageNumber="34">
<normalizedToken originalValue="J–L">J-L</normalizedToken>
</emphasis>
paratype (SAMA C12708). 
<emphasis bold="true" pageId="9" pageNumber="34">
<normalizedToken originalValue="A–C">A-C</normalizedToken>
</emphasis>
Right cirrus 
<normalizedToken originalValue="I–III">I-III</normalizedToken>
lateral view 
<emphasis bold="true" pageId="9" pageNumber="34">D</emphasis>
right cirrus III intermediate segments 
<emphasis bold="true" pageId="9" pageNumber="34">E</emphasis>
right cirrus IV lateral view 
<emphasis bold="true" pageId="9" pageNumber="34">F</emphasis>
same, anterior ramus intermediate segments 
<emphasis bold="true" pageId="9" pageNumber="34">G</emphasis>
left cirrus IV anterior ramus intermediate segments 
<emphasis bold="true" pageId="9" pageNumber="34">H</emphasis>
right cirrus IV anterodistal angle of distal segment of pedical showing ctenoid scales 
<emphasis bold="true" pageId="9" pageNumber="34">I, J</emphasis>
Ieft cirrus IV basal segments of rami showing posterior spines 
<emphasis bold="true" pageId="9" pageNumber="34">K</emphasis>
close up of spines on posterior margin of J 
<emphasis bold="true" pageId="9" pageNumber="34">L</emphasis>
close up of spines and posterodistal spines basal segments of J. Note: B, C, E damaged. Scale bars: 400 
<normalizedToken originalValue="μm">μm</normalizedToken>
(
<emphasis bold="true" pageId="9" pageNumber="34">
<normalizedToken originalValue="A–C">A-C</normalizedToken>
, E
</emphasis>
); 100 
<normalizedToken originalValue="μm">μm</normalizedToken>
(
<emphasis bold="true" pageId="9" pageNumber="34">D, F, L</emphasis>
); 150 
<normalizedToken originalValue="μm">μm</normalizedToken>
(
<emphasis bold="true" pageId="9" pageNumber="34">G, I</emphasis>
); 50 
<normalizedToken originalValue="μm">μm</normalizedToken>
(
<emphasis bold="true" pageId="9" pageNumber="34">H, K</emphasis>
); 200 
<normalizedToken originalValue="μm">μm</normalizedToken>
(
<emphasis bold="true" pageId="9" pageNumber="34">J</emphasis>
).
</paragraph>
</caption>
<paragraph pageId="9" pageNumber="34">
Cirrus II (
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Membranobalanus porphyrophilus sp. nov. cirri A-F holotype (WAM C 66803) G-I paratype (SAMA C 12706) J-L paratype (SAMA C 12708). A-C Right cirrus I-III lateral view D right cirrus III intermediate segments E right cirrus IV lateral view F same, anterior ramus intermediate segments G left cirrus IV anterior ramus intermediate segments H right cirrus IV anterodistal angle of distal segment of pedical showing ctenoid scales I, J Ieft cirrus IV basal segments of rami showing posterior spines K close up of spines on posterior margin of J L close up of spines and posterodistal spines basal segments of J. Note: B, C, E damaged. Scale bars: 400 μm (A-C, E); 100 μm (D, F, L); 150 μm (G, I); 50 μm (H, K); 200 μm (J)." figureDoi="10.3897/zookeys.873.35421.figure5" httpUri="https://binary.pensoft.net/fig/328939" pageId="9" pageNumber="34">Fig. 5B</figureCitation>
) shorter than other cirri, rami subequal; anterior margins of both rami slightly protuberant with serrulate and simple setae.
</paragraph>
<paragraph pageId="9" pageNumber="34">
Cirrus III (
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Membranobalanus porphyrophilus sp. nov. cirri A-F holotype (WAM C 66803) G-I paratype (SAMA C 12706) J-L paratype (SAMA C 12708). A-C Right cirrus I-III lateral view D right cirrus III intermediate segments E right cirrus IV lateral view F same, anterior ramus intermediate segments G left cirrus IV anterior ramus intermediate segments H right cirrus IV anterodistal angle of distal segment of pedical showing ctenoid scales I, J Ieft cirrus IV basal segments of rami showing posterior spines K close up of spines on posterior margin of J L close up of spines and posterodistal spines basal segments of J. Note: B, C, E damaged. Scale bars: 400 μm (A-C, E); 100 μm (D, F, L); 150 μm (G, I); 50 μm (H, K); 200 μm (J)." figureDoi="10.3897/zookeys.873.35421.figure5" httpUri="https://binary.pensoft.net/fig/328939" pageId="9" pageNumber="34">Fig. 5C, D</figureCitation>
) pedicel with plumose setae on anterior and posterior margins of both segments; rami equal; anterior ramus with erect spines and ctenoid scales on anterodistal portion of medial segments, tuft of long serrulate setae on rounded anterior margin up to approximately three times as long as segment, tuft of serrulate setae as long as segment at posterodistal angle.
</paragraph>
<paragraph pageId="9" pageNumber="34">
Cirrus IV (
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Membranobalanus porphyrophilus sp. nov. cirri A-F holotype (WAM C 66803) G-I paratype (SAMA C 12706) J-L paratype (SAMA C 12708). A-C Right cirrus I-III lateral view D right cirrus III intermediate segments E right cirrus IV lateral view F same, anterior ramus intermediate segments G left cirrus IV anterior ramus intermediate segments H right cirrus IV anterodistal angle of distal segment of pedical showing ctenoid scales I, J Ieft cirrus IV basal segments of rami showing posterior spines K close up of spines on posterior margin of J L close up of spines and posterodistal spines basal segments of J. Note: B, C, E damaged. Scale bars: 400 μm (A-C, E); 100 μm (D, F, L); 150 μm (G, I); 50 μm (H, K); 200 μm (J)." figureDoi="10.3897/zookeys.873.35421.figure5" httpUri="https://binary.pensoft.net/fig/328939" pageId="9" pageNumber="34">
Fig. 5 
<normalizedToken originalValue="E–L">E-L</normalizedToken>
</figureCitation>
) pedicel without erect teeth on anterodistal margins, pedicel with numerous small denticles on posterior and anterior margins of mesial face, tuft of short setae at posterodistal angle, small ctenoid scale-like denticles at anterodistal margin. Rami subequal in length; basal-most segment with up to 13 stout spines along posterior margin (holotype with six), basal segments with one or two stout spines at posterodistal angle. Anterior ramus intermediate segments with row of erect spines on anterodistal portion of all but most distal segments, much reduced in posterior ramus, without recurved teeth on anterior faces, posterodistal angles with tuft of setae.
</paragraph>
<paragraph pageId="9" pageNumber="34">
Cirrus V (
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Membranobalanus porphyrophilus sp. nov. cirri A, B, D, E, H holotype (WAM C 66803) C, F, G paratype (SAMA C 12706). A Right cirrus V lateral view B basal segments of cirrus V rami C intermediate segments of cirrus V anterior ramus D right cirrus VI mesial view (damaged) E same, intermediate segments F example of long serrulate setae on cirrus VI G basal segment of pedicel of right cirrus VI, mesial view showing denticles on anterior and posterior margin H basidorsal point of penis. Scale bars: 400 μm (A, D); 100 μm (B, C); 50 μm (F); 150 μm (G, H); 80 μm (E)." figureDoi="10.3897/zookeys.873.35421.figure6" httpUri="https://binary.pensoft.net/fig/328940" pageId="9" pageNumber="34">
Fig. 6 
<normalizedToken originalValue="A–C">A-C</normalizedToken>
</figureCitation>
) characters intermediate between cirri IV and VI; pedicel with numerous small denticles on posterior and anterior margins of mesial face, tuft of short setae at posterodistal angle, small ctenoid scale-like denticles on anterodistal margin. Rami equal, segments becoming elongate distally from half as long as wide to three times longer than wide, both rami with four pairs of serrulate setae on anterior margin, distal most pair longest, approximately 3 times length of segment; tuft of simple setae at anterodistal angle, setae up to half length of segment; erect teeth on anterior margins of intermediate segments less pronounced than those on cirrus IV; stout spines on posterior margins of rami similar to cirrus IV.
</paragraph>
<caption doi="10.3897/zookeys.873.35421.figure6" httpUri="https://binary.pensoft.net/fig/328940" pageId="9" pageNumber="34" start="Figure 6" startId="F6">
<paragraph pageId="9" pageNumber="34">
Figure 6. 
<taxonomicName class="Maxillopoda" family="Archaeobalanidae" genus="Membranobalanus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Membranobalanus porphyrophilus" order="Sessilia" pageId="9" pageNumber="34" phylum="Arthropoda" rank="species" species="porphyrophilus">
<emphasis italics="true" pageId="9" pageNumber="34">Membranobalanus porphyrophilus</emphasis>
</taxonomicName>
sp. nov. cirri 
<emphasis bold="true" pageId="9" pageNumber="34">A, B, D, E, H</emphasis>
holotype (WAM C66803) 
<emphasis bold="true" pageId="9" pageNumber="34">C, F, G</emphasis>
paratype (SAMA C12706). 
<emphasis bold="true" pageId="9" pageNumber="34">A</emphasis>
Right cirrus V lateral view 
<emphasis bold="true" pageId="9" pageNumber="34">B</emphasis>
basal segments of cirrus V rami 
<emphasis bold="true" pageId="9" pageNumber="34">C</emphasis>
intermediate segments of cirrus V anterior ramus 
<emphasis bold="true" pageId="9" pageNumber="34">D</emphasis>
right cirrus VI mesial view (damaged) 
<emphasis bold="true" pageId="9" pageNumber="34">E</emphasis>
same, intermediate segments 
<emphasis bold="true" pageId="9" pageNumber="34">F</emphasis>
example of long serrulate setae on cirrus VI 
<emphasis bold="true" pageId="9" pageNumber="34">G</emphasis>
basal segment of pedicel of right cirrus VI, mesial view showing denticles on anterior and posterior margin 
<emphasis bold="true" pageId="9" pageNumber="34">H</emphasis>
basidorsal point of penis. Scale bars: 400 
<normalizedToken originalValue="μm">μm</normalizedToken>
(
<emphasis bold="true" pageId="9" pageNumber="34">A, D</emphasis>
); 100 
<normalizedToken originalValue="μm">μm</normalizedToken>
(
<emphasis bold="true" pageId="9" pageNumber="34">B, C)</emphasis>
; 50 
<normalizedToken originalValue="μm">μm</normalizedToken>
(
<emphasis bold="true" pageId="9" pageNumber="34">F</emphasis>
); 150 
<normalizedToken originalValue="μm">μm</normalizedToken>
(
<emphasis bold="true" pageId="9" pageNumber="34">G, H</emphasis>
); 80 
<normalizedToken originalValue="μm">μm</normalizedToken>
(
<emphasis bold="true" pageId="9" pageNumber="34">E</emphasis>
).
</paragraph>
</caption>
<paragraph lastPageId="10" lastPageNumber="35" pageId="9" pageNumber="34">
Cirrus VI (
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Membranobalanus porphyrophilus sp. nov. cirri A, B, D, E, H holotype (WAM C 66803) C, F, G paratype (SAMA C 12706). A Right cirrus V lateral view B basal segments of cirrus V rami C intermediate segments of cirrus V anterior ramus D right cirrus VI mesial view (damaged) E same, intermediate segments F example of long serrulate setae on cirrus VI G basal segment of pedicel of right cirrus VI, mesial view showing denticles on anterior and posterior margin H basidorsal point of penis. Scale bars: 400 μm (A, D); 100 μm (B, C); 50 μm (F); 150 μm (G, H); 80 μm (E)." figureDoi="10.3897/zookeys.873.35421.figure6" httpUri="https://binary.pensoft.net/fig/328940" pageId="9" pageNumber="34">
Fig. 6 
<normalizedToken originalValue="D–G">D-G</normalizedToken>
</figureCitation>
) rami equal, pedicel with numerous small denticles on posteromesial face, tuft of simple setae at posterodistal angle; intermediate segments of both rami becoming elongate distally from 1.5 times wider than long to three times longer 
<pageBreakToken pageId="10" pageNumber="35" start="start">than</pageBreakToken>
wide, both rami with four pairs of serrulate setae on anterior margin, distal most pair longest, approximately three times the length of segment, basal pair 0.3 times length of segment; tuft of simple setae at anterodistal angle, setae up to 0.5 length of segment; anterior margins of all segments without erect teeth, posterior margins without stout spines.
</paragraph>
<paragraph pageId="10" pageNumber="35">Cirral segment counts as follows (anterior ramus, posterior ramus):</paragraph>
<paragraph pageId="10" pageNumber="35">
<table pageId="10" pageNumber="35">
<tr pageId="10" pageNumber="35">
<td colspan="1" isEmpty="true" pageId="10" pageNumber="35" rowspan="1">-</td>
<td colspan="1" pageId="10" pageNumber="35" rowspan="1">
<emphasis bold="true" pageId="10" pageNumber="35">Cirrus</emphasis>
</td>
<td colspan="1" pageId="10" pageNumber="35" rowspan="1">
<emphasis bold="true" pageId="10" pageNumber="35">I</emphasis>
</td>
<td colspan="1" pageId="10" pageNumber="35" rowspan="1">
<emphasis bold="true" pageId="10" pageNumber="35">II</emphasis>
</td>
<td colspan="1" pageId="10" pageNumber="35" rowspan="1">
<emphasis bold="true" pageId="10" pageNumber="35">III</emphasis>
</td>
<td colspan="1" pageId="10" pageNumber="35" rowspan="1">
<emphasis bold="true" pageId="10" pageNumber="35">IV</emphasis>
</td>
<td colspan="1" pageId="10" pageNumber="35" rowspan="1">
<emphasis bold="true" pageId="10" pageNumber="35">V</emphasis>
</td>
<td colspan="1" pageId="10" pageNumber="35" rowspan="1">
<emphasis bold="true" pageId="10" pageNumber="35">VI</emphasis>
</td>
</tr>
<tr pageId="10" pageNumber="35">
<td colspan="1" pageId="10" pageNumber="35" rowspan="2">WAM C66803</td>
<td colspan="1" pageId="10" pageNumber="35" rowspan="1">L</td>
<td colspan="1" pageId="10" pageNumber="35" rowspan="1">21, 9</td>
<td colspan="1" pageId="10" pageNumber="35" rowspan="1">9, 11</td>
<td colspan="1" pageId="10" pageNumber="35" rowspan="1">17, 13</td>
<td colspan="1" pageId="10" pageNumber="35" rowspan="1">27, 30</td>
<td colspan="1" pageId="10" pageNumber="35" rowspan="1">34, 32</td>
<td colspan="1" pageId="10" pageNumber="35" rowspan="1">33, 33</td>
</tr>
<tr pageId="10" pageNumber="35">
<td colspan="1" pageId="10" pageNumber="35" rowspan="1">R</td>
<td colspan="1" pageId="10" pageNumber="35" rowspan="1">20, 9</td>
<td colspan="1" pageId="10" pageNumber="35" rowspan="1">11, 9</td>
<td colspan="1" pageId="10" pageNumber="35" rowspan="1">17, 14</td>
<td colspan="1" pageId="10" pageNumber="35" rowspan="1">30, 30</td>
<td colspan="1" pageId="10" pageNumber="35" rowspan="1">32, 34</td>
<td colspan="1" pageId="10" pageNumber="35" rowspan="1">36, 34</td>
</tr>
</table>
</paragraph>
<paragraph pageId="11" pageNumber="36">
<pageBreakToken pageId="11" pageNumber="36" start="start">Penis</pageBreakToken>
longer than CVI, annulated, sparsely setose along length; basidorsal point (
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Membranobalanus porphyrophilus sp. nov. cirri A, B, D, E, H holotype (WAM C 66803) C, F, G paratype (SAMA C 12706). A Right cirrus V lateral view B basal segments of cirrus V rami C intermediate segments of cirrus V anterior ramus D right cirrus VI mesial view (damaged) E same, intermediate segments F example of long serrulate setae on cirrus VI G basal segment of pedicel of right cirrus VI, mesial view showing denticles on anterior and posterior margin H basidorsal point of penis. Scale bars: 400 μm (A, D); 100 μm (B, C); 50 μm (F); 150 μm (G, H); 80 μm (E)." figureDoi="10.3897/zookeys.873.35421.figure6" httpUri="https://binary.pensoft.net/fig/328940" pageId="11" pageNumber="36">Fig. 6H</figureCitation>
) prominent, triangular, height twice width.
</paragraph>
</subSubSection>
<subSubSection pageId="11" pageNumber="36" type="distribution">
<paragraph pageId="11" pageNumber="36">Distribution.</paragraph>
<paragraph pageId="11" pageNumber="36">Southern Australia, from Perth to Adelaide.</paragraph>
</subSubSection>
<subSubSection pageId="11" pageNumber="36" type="etymology">
<paragraph pageId="11" pageNumber="36">Etymology.</paragraph>
<paragraph pageId="11" pageNumber="36">
From Greek 
<emphasis italics="true" pageId="11" pageNumber="36">porphyra</emphasis>
, purple and 
<emphasis italics="true" pageId="11" pageNumber="36">philos</emphasis>
, indicating a love of; gender masculine. In reference to the vibrant purple colour of the only known host.
</paragraph>
</subSubSection>
<subSubSection pageId="11" pageNumber="36" type="remarks">
<paragraph pageId="11" pageNumber="36">Remarks.</paragraph>
<paragraph pageId="11" pageNumber="36">
<taxonomicName class="Maxillopoda" family="Archaeobalanidae" genus="Membranobalanus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Membranobalanus porphyrophilus" order="Sessilia" pageId="11" pageNumber="36" phylum="Arthropoda" rank="species" species="porphyrophilus">
<emphasis italics="true" pageId="11" pageNumber="36">Membranobalanus porphyrophilus</emphasis>
</taxonomicName>
sp. nov. is most readily distinguished from its congeners by the narrow, beaked tergum. The absence of recurved teeth on cirrus IV, the finely striated scutal growth lines, and the weak articular structure of the opercular plates further separates the newly described species from the predominantly American group of species, viz. 
<taxonomicName lsidName="M. costatus" pageId="11" pageNumber="36" rank="species" species="costatus">
<emphasis italics="true" pageId="11" pageNumber="36">M. costatus</emphasis>
</taxonomicName>
, 
<taxonomicName lsidName="M. declivis" pageId="11" pageNumber="36" rank="species" species="declivis">
<emphasis italics="true" pageId="11" pageNumber="36">M. declivis</emphasis>
</taxonomicName>
, 
<taxonomicName lsidName="M. koreanus" pageId="11" pageNumber="36" rank="species" species="koreanus">
<emphasis italics="true" pageId="11" pageNumber="36">M. koreanus</emphasis>
</taxonomicName>
, 
<taxonomicName lsidName="M. nebrias" pageId="11" pageNumber="36" rank="species" species="nebrias">
<emphasis italics="true" pageId="11" pageNumber="36">M. nebrias</emphasis>
</taxonomicName>
, 
<taxonomicName lsidName="M. orcutti" pageId="11" pageNumber="36" rank="species" species="orcutti">
<emphasis italics="true" pageId="11" pageNumber="36">M. orcutti</emphasis>
</taxonomicName>
, and 
<taxonomicName lsidName="M. robinae" pageId="11" pageNumber="36" rank="species" species="robinae">
<emphasis italics="true" pageId="11" pageNumber="36">M. robinae</emphasis>
</taxonomicName>
. The remaining species in the genus, 
<taxonomicName lsidName="M. brachialis" pageId="11" pageNumber="36" rank="species" species="brachialis">
<emphasis italics="true" pageId="11" pageNumber="36">M. brachialis</emphasis>
</taxonomicName>
, 
<taxonomicName lsidName="M. cuneiformis" pageId="11" pageNumber="36" rank="species" species="cuneiformis">
<emphasis italics="true" pageId="11" pageNumber="36">M. cuneiformis</emphasis>
</taxonomicName>
, and 
<taxonomicName lsidName="M. longirostrum" pageId="11" pageNumber="36" rank="species" species="longirostrum">
<emphasis italics="true" pageId="11" pageNumber="36">M. longirostrum</emphasis>
</taxonomicName>
all bear prominent, erect spines on the pedicel of cirrus IV and lack the stout spines on the posterior margins of the rami of cirri 
<normalizedToken originalValue="IV–VI">IV-VI</normalizedToken>
. This is only the second member of the genus reported from Australian waters. The other, 
<taxonomicName lsidName="M. cuneiformis" pageId="11" pageNumber="36" rank="species" species="cuneiformis">
<emphasis italics="true" pageId="11" pageNumber="36">M. cuneiformis</emphasis>
</taxonomicName>
, is known from near Darwin and was reported by 
<bibRefCitation author="Hiro, F" journalOrPublisher="&quot; Japan Journal of Zoology" pageId="14" pageNumber="39" pagination="621 - 636" refId="B11" refString="Hiro, F, 1936. Report on the Cirripedia collected in Malayan waters by the Ship &quot;Zuihomaru. . &quot; Japan Journal of Zoology 3: 621 - 636" title="Report on the Cirripedia collected in Malayan waters by the Ship &quot; Zuihomaru." volume="3" year="1936">Hiro (1936)</bibRefCitation>
as commensal with an unidentified species of 
<taxonomicName class="Demospongiae" family="Clionaidae" genus="Cliona" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cliona" order="Clionaida" pageId="11" pageNumber="36" phylum="Porifera" rank="genus">
<emphasis italics="true" pageId="11" pageNumber="36">Cliona</emphasis>
</taxonomicName>
that was bioeroding the shell of 
<taxonomicName class="Bivalvia" family="Margaritidae" genus="Pinctada" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pinctada maxima" order="Ostreida" pageId="11" pageNumber="36" phylum="Mollusca" rank="species" species="maxima">
<emphasis italics="true" pageId="11" pageNumber="36">Pinctada maxima</emphasis>
</taxonomicName>
(Jameson, 1901). Externally, 
<taxonomicName lsidName="M. cuneiformis" pageId="11" pageNumber="36" rank="species" species="cuneiformis">
<emphasis italics="true" pageId="11" pageNumber="36">M. cuneiformis</emphasis>
</taxonomicName>
can be separated from 
<taxonomicName lsidName="M. porphyrophilus" pageId="11" pageNumber="36" rank="species" species="porphyrophilus">
<emphasis italics="true" pageId="11" pageNumber="36">M. porphyrophilus</emphasis>
</taxonomicName>
by the conical shape of the shell, resulting in a small orifice and absent radii.
</paragraph>
</subSubSection>
</treatment>
</document>