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<mods:title id="31200FBCED009ACA9CDA4F831C276A38">Digitonthophagus Balthasar, 1959: taxonomy, systematics, and morphological phylogeny of the genus revealing an African species complex (Coleoptera: Scarabaeidae: Scarabaeinae)</mods:title>
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<treatment id="3D5987B70771FF9DFF17A7A601EDF9B9" ID-DOI="http://doi.org/10.5281/zenodo.5490060" ID-GBIF-Taxon="128090232" ID-Zenodo-Dep="5490060" LSID="urn:lsid:plazi:treatment:3D5987B70771FF9DFF17A7A601EDF9B9" httpUri="http://treatment.plazi.org/id/3D5987B70771FF9DFF17A7A601EDF9B9" lastPageNumber="15" pageId="14" pageNumber="15">
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<paragraph id="B54F36A10771FF9DFF17A7A602E3F9C0" blockId="14.[151,652,1519,1546]" box="[151,652,1519,1546]" pageId="14" pageNumber="15">
<heading id="EE0781CD0771FF9DFF17A7A602E3F9C0" bold="true" box="[151,652,1519,1546]" fontSize="11" level="1" pageId="14" pageNumber="15" reason="1">
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<taxonomicName id="72F04D220771FF9DFF17A7A601B4F9C3" ID-CoL="363WG" box="[151,475,1519,1545]" class="Insecta" family="Scarabaeidae" genus="Digitonthophagus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="bonasus">
<emphasis id="8784EAB30771FF9DFF17A7A601B4F9C3" bold="true" box="[151,475,1519,1545]" italics="true" pageId="14" pageNumber="15">Digitonthophagus bonasus</emphasis>
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species group
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<subSubSection id="FDEA652A0771FF9DFF17A47F01EDF9B9" pageId="14" pageNumber="15" type="diagnosis">
<paragraph id="B54F36A10771FF9DFF17A47F01EDF9B9" blockId="14.[151,1436,1590,1651]" pageId="14" pageNumber="15">
<emphasis id="8784EAB30771FF9DFF17A47F0165F985" bold="true" box="[151,266,1590,1615]" pageId="14" pageNumber="15">Diagnosis</emphasis>
. Male cephalic horns long, obliquely divergent in frontal view (
<figureCitation id="2DCB2A240771FF9DFC5AA47F04E0F985" box="[986,1167,1590,1615]" captionStart="FIGURES 23 30. 23, 25, 27, 29" captionStartId="91.[151,264,1926,1948]" captionTargetBox="[211,1375,193,1905]" captionTargetId="figure@91.[211,1375,193,1905]" captionTargetPageId="91" captionText="FIGURES 23 30. 23, 25, 27, 29. Male head, dorsal view; 24, 26, 28, 30. Male head, frontal view. 23 24. Digitonthophagus bonasus, alloreferent (Tamil Nadu State, India); 25 26. D. uks, holotype; 27 28. D. sahelicus, holotype; 29 30. D. catta, alloreferent (Tamil Nadu State, India)." httpUri="https://zenodo.org/record/439455/files/figure.png" pageId="14" pageNumber="15">Figs. 24, 26, 28</figureCitation>
); females with horns on vertex (
<figureCitation id="2DCB2A240771FF9DFF6CA4130119F9B9" box="[236,374,1626,1651]" captionStart="FIGURES 71 78" captionStartId="97.[151,264,1926,1948]" captionTargetBox="[211,1375,193,1905]" captionTargetId="figure@97.[211,1375,193,1905]" captionTargetPageId="97" captionText="FIGURES 71 78. Female head, dorsal view. 71. Digitonthophagus bonasus, non type (Tamil Nadu State, India); 72. D. uks, allotype; 73. D. sahelicus, allotype; 74. D. catta, non type (India); 75. D. eucatta, allotype; 76. D. falciger, allotype; 77. D. biflagellatus, allotype; 78. D. gazella, non type (Antisiranana Province, Madagascar)." httpUri="https://zenodo.org/record/439461/files/figure.png" pageId="14" pageNumber="15">Figs. 7173</figureCitation>
).
</paragraph>
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</treatment>
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