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<document id="9A8571521AEFB411A4E8F2B5604C123E" ID="10.11646/zootaxa.4248.1.1SLASH10685" ID-CLB-Dataset="33324" ID-DOI="10.5281/zenodo.439444" ID-GBIF-Dataset="cf486ccc-d298-4179-8ccf-e29a9c894ab4" ID-ISSN="1175-5326" ID-Zenodo-Dep="439444" ID-ZooBank="71F0AC03-C1FB-40AB-8532-99A638FC91E9" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1490945156927" checkinUser="plazi" docAuthor="Génier, François &amp; Moretto, Philippe" docDate="2017" docId="3D5987B70768FF84FF17A00E04CCFCD9" docLanguage="en" docName="zootaxa.4248.1.1SLASH10685.pdf" docOrigin="Zootaxa 4248 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Digitonthophagus catta" docType="treatment" docVersion="15" lastPageNumber="24" masterDocId="C160FFCF077FFF93FF80A249006FFFCA" masterDocTitle="Digitonthophagus Balthasar, 1959: taxonomy, systematics, and morphological phylogeny of the genus revealing an African species complex (Coleoptera: Scarabaeidae: Scarabaeinae)" masterLastPageNumber="110" masterPageNumber="1" pageNumber="24" updateTime="1698438591535" updateUser="ExternalLinkService">
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<mods:title id="C79572755FCEB4A13068C5D067D10C3B">Digitonthophagus Balthasar, 1959: taxonomy, systematics, and morphological phylogeny of the genus revealing an African species complex (Coleoptera: Scarabaeidae: Scarabaeinae)</mods:title>
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<mods:namePart id="8FA853AC37430AC827C7EBE5C165B7B4">Génier, François</mods:namePart>
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<mods:namePart id="78F0FC9FD37BCE8847080805C695BA96">Moretto, Philippe</mods:namePart>
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<treatment id="3D5987B70768FF84FF17A00E04CCFCD9" ID-DOI="http://doi.org/10.5281/zenodo.5490068" ID-GBIF-Taxon="128090238" ID-Zenodo-Dep="5490068" LSID="urn:lsid:plazi:treatment:3D5987B70768FF84FF17A00E04CCFCD9" httpUri="http://treatment.plazi.org/id/3D5987B70768FF84FF17A00E04CCFCD9" lastPageNumber="24" pageId="23" pageNumber="24">
<subSubSection id="FDEA652A0768FF84FF17A00E020DFDA8" box="[151,610,583,610]" pageId="23" pageNumber="24" type="nomenclature">
<paragraph id="B54F36A10768FF84FF17A00E020DFDA8" blockId="23.[151,610,583,610]" box="[151,610,583,610]" pageId="23" pageNumber="24">
<heading id="EE0781CD0768FF84FF17A00E020DFDA8" bold="true" box="[151,610,583,610]" fontSize="11" level="1" pageId="23" pageNumber="24" reason="1">
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<taxonomicName id="72F04D220768FF84FF17A00E01DDFDAB" ID-CoL="363WH" box="[151,434,583,609]" class="Insecta" family="Scarabaeidae" genus="Digitonthophagus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="catta">
<emphasis id="8784EAB30768FF84FF17A00E01DDFDAB" bold="true" box="[151,434,583,609]" italics="true" pageId="23" pageNumber="24">Digitonthophagus catta</emphasis>
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species group
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</paragraph>
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<subSubSection id="FDEA652A0768FF84FF17A0C704CCFCD9" pageId="23" pageNumber="24" type="diagnosis">
<paragraph id="B54F36A10768FF84FF17A0C704CCFCD9" blockId="23.[151,1436,654,787]" pageId="23" pageNumber="24">
<emphasis id="8784EAB30768FF84FF17A0C70165FD6D" bold="true" box="[151,266,654,679]" pageId="23" pageNumber="24">Diagnosis</emphasis>
. Pronotal punctures scattered and small on posterior half (
<figureCitation id="2DCB2A240768FF84FC18A0C70399FD6D" box="[920,1014,654,679]" captionStart="FIGURES 55 62" captionStartId="95.[151,264,1926,1948]" captionTargetBox="[211,1375,193,1905]" captionTargetId="figure@95.[211,1375,193,1905]" captionTargetPageId="95" captionText="FIGURES 55 62. Male pronotum, dorsal view. 55. Digitonthophagus bonasus, alloreferent (Tamil Nadu State, India); 56. D. uks, holotype; 57. D. sahelicus, holotype; 58. D. catta, alloreferent (Tamil Nadu State, India); 59. D. eucatta, holotype; 60. D. falciger, holotype; 61. D. biflagellatus, holotype; 62. D. gazella, non type (Central Province, Zambia)." httpUri="https://zenodo.org/record/439459/files/figure.png" pageId="23" pageNumber="24">Figs. 58</figureCitation>
, 93); surface behind anterior pronotal angles concave; male with two erected cephalic horns (
<figureCitation id="2DCB2A240768FF84FC85A0FB030BFD01" box="[773,868,690,715]" captionStart="FIGURES 23 30. 23, 25, 27, 29" captionStartId="91.[151,264,1926,1948]" captionTargetBox="[211,1375,193,1905]" captionTargetId="figure@91.[211,1375,193,1905]" captionTargetPageId="91" captionText="FIGURES 23 30. 23, 25, 27, 29. Male head, dorsal view; 24, 26, 28, 30. Male head, frontal view. 23 24. Digitonthophagus bonasus, alloreferent (Tamil Nadu State, India); 25 26. D. uks, holotype; 27 28. D. sahelicus, holotype; 29 30. D. catta, alloreferent (Tamil Nadu State, India)." httpUri="https://zenodo.org/record/439455/files/figure.png" pageId="23" pageNumber="24">Figs. 30</figureCitation>
,
<figureCitation id="2DCB2A240768FF84FCEFA0FA03E0FD01" box="[879,911,691,715]" captionStart="FIGURES 31 38. 31, 33, 35, 37" captionStartId="92.[151,264,1926,1948]" captionTargetBox="[211,1375,193,1905]" captionTargetId="figure@92.[211,1375,193,1905]" captionTargetPageId="92" captionText="FIGURES 31 38. 31, 33, 35, 37. Male head, dorsal view; 32, 34, 36, 38. Male head, frontal view. 31 32. Digitonthophagus eucatta, alloreferent; 33 34. D. falciger, holotype; 35 36. D. biflagellatus, holotype; 37 38. D. gazella, non type (Central Province, Zambia)." httpUri="https://zenodo.org/record/439456/files/figure.png" pageId="23" pageNumber="24">32</figureCitation>
); females with anterior pronotal tubercles well developed and deeply obliquely concave (Figs. 94, 96); subapicodorsal lobe of FLP sclerite enlarged and sclerotized (
<figureCitation id="2DCB2A240768FF84FE9FA0B301E4FCD9" box="[287,395,762,787]" captionStart="FIGURES 135 146" captionStartId="104.[151,264,1926,1948]" captionTargetBox="[230,1287,220,1880]" captionTargetId="figure@104.[211,1375,193,1905]" captionTargetPageId="104" captionText="FIGURES 135 146. Internal sac sclerites (FLP, A + SA). 135, 138, 141, 144. Dorsal view; 136, 139, 142, 145. Ventral view. 137, 140, 143, 146. Right lateral view. 135 137. Digitonthophagus bonasus, alloreferent (Tamil Nadu State, India); 138 140. D. uks, holotype; 141 143. D. sahelicus, holotype; 144 146. D. catta, alloreferent." httpUri="https://zenodo.org/record/439464/files/figure.png" pageId="23" pageNumber="24">Figs. 144</figureCitation>
,
<figureCitation id="2DCB2A240768FF84FE17A0B201ABFCD9" box="[407,452,763,787]" captionStart="FIGURES 147 158" captionStartId="105.[151,264,1926,1948]" captionTargetBox="[213,1330,208,1899]" captionTargetId="figure@105.[211,1375,193,1905]" captionTargetPageId="105" captionText="FIGURES 147 158. Internal sac sclerites (FLP, A + SA). 147, 150, 153, 156. Dorsal view; 148, 151, 154, 157. Ventral view. 149, 152, 155, 158. Right lateral view. 147 149. Digitonthophagus eucatta, alloreferent (Tamil Nadu State, India); 150 152. D. falciger, holotype; 153 155. D. biflagellatus, holotype; 156 158. D. gazella, non type (Antisiranana Province, Madagascar)." httpUri="https://zenodo.org/record/439465/files/figure.png" pageId="23" pageNumber="24">147</figureCitation>
); FLP left surface with a conspicuous apophysis (
<figureCitation id="2DCB2A240768FF84FC71A0B30433FCD9" box="[1009,1116,762,787]" captionStart="FIGURES 135 146" captionStartId="104.[151,264,1926,1948]" captionTargetBox="[230,1287,220,1880]" captionTargetId="figure@104.[211,1375,193,1905]" captionTargetPageId="104" captionText="FIGURES 135 146. Internal sac sclerites (FLP, A + SA). 135, 138, 141, 144. Dorsal view; 136, 139, 142, 145. Ventral view. 137, 140, 143, 146. Right lateral view. 135 137. Digitonthophagus bonasus, alloreferent (Tamil Nadu State, India); 138 140. D. uks, holotype; 141 143. D. sahelicus, holotype; 144 146. D. catta, alloreferent." httpUri="https://zenodo.org/record/439464/files/figure.png" pageId="23" pageNumber="24">Figs. 144</figureCitation>
,
<figureCitation id="2DCB2A240768FF84FBE8A0B204F9FCD9" box="[1128,1174,763,787]" captionStart="FIGURES 147 158" captionStartId="105.[151,264,1926,1948]" captionTargetBox="[213,1330,208,1899]" captionTargetId="figure@105.[211,1375,193,1905]" captionTargetPageId="105" captionText="FIGURES 147 158. Internal sac sclerites (FLP, A + SA). 147, 150, 153, 156. Dorsal view; 148, 151, 154, 157. Ventral view. 149, 152, 155, 158. Right lateral view. 147 149. Digitonthophagus eucatta, alloreferent (Tamil Nadu State, India); 150 152. D. falciger, holotype; 153 155. D. biflagellatus, holotype; 156 158. D. gazella, non type (Antisiranana Province, Madagascar)." httpUri="https://zenodo.org/record/439465/files/figure.png" pageId="23" pageNumber="24">147</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
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